871 lines
94 KiB
XML
871 lines
94 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1190.114958" ID-Pensoft-Pub="1313-2970-1190-259" ID-Pensoft-UUID="76BAA6C26E1A559E8597D66C26B690E4" ID-ZooBank="A5D5D928CE844E13BC31ED2393B3403D" ModsDocID="1313-2970-1190-259" checkinTime="1706706784770" checkinUser="pensoft" docAuthor="Nguyen, Anh D., Vu, Tam T. T. & Eguchi, Katsuyuki" docDate="2024" docId="450DB17E1A665A2B8774486FA1123F88" docLanguage="en" docName="ZooKeys 1190: 259-280" docOrigin="ZooKeys 1190" docPubDate="2024-01-30" docSource="http://dx.doi.org/10.3897/zookeys.1190.114958" docTitle="Pacidesmus tuachua Nguyen & Vu & Eguchi 2024, sp. nov." docType="treatment" docUuid="CB8E9D35-3365-4416-81CA-589284A6D42F" docUuidSource="ZooBank" docVersion="2" id="76BAA6C26E1A559E8597D66C26B690E4" lastPageNumber="259" masterDocId="76BAA6C26E1A559E8597D66C26B690E4" masterDocTitle="The millipede family Polydesmidae Leach, 1816 (Diplopoda, Polydesmida) from Vietnam, with a description of a new cavernicolous species" masterLastPageNumber="280" masterPageNumber="259" pageNumber="259" updateTime="1706707086208" updateUser="ExternalLinkService">
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<mods:title id="3D997D0C8C23F46F8788D3C4825306F2">The millipede family Polydesmidae Leach, 1816 (Diplopoda, Polydesmida) from Vietnam, with a description of a new cavernicolous species</mods:title>
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<mods:namePart id="7FDB7195F4603D1BA71E700702FA3527">Nguyen, Anh D.</mods:namePart>
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<mods:affiliation id="C9593343E2EA62A849080574C0CF1C65">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18, Hoangquocviet Rd., Caugiay District, Hanoi, Vietnam & Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18, Hoangquocviet Rd., Caugiay District, Hanoi, Vietnam</mods:affiliation>
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<mods:nameIdentifier id="9251A09AFDBC85F291EEDCE4EB85FEE5" type="email">ducanh410@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="E7EDD9676D27001174D772BE31C8A4A2">Vu, Tam T. T.</mods:namePart>
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<mods:affiliation id="DE698FD283B5260A1274DC24E11152BC">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18, Hoangquocviet Rd., Caugiay District, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart id="40079ECF7763FF537424D085BF0BC80C">Eguchi, Katsuyuki</mods:namePart>
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<mods:affiliation id="2EC40D563587F188E002C467727DD035">Graduate School of Science, Tokyo Metropolitan University, Minami-osawa 1 - 1, Hachioji, Tokyo 192 - 0397, Japan & Department of International Health and Medical Anthropology, Institute of Tropical Medicine, 1 - 12 - 4 Sakamoto, Nagasaki University, Nagasaki, 852 - 8523, Japan</mods:affiliation>
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<mods:date id="2E0A63EF7B748C39A2DC60C9D0B9C758">2024</mods:date>
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<mods:number id="EDD71452C72424C64E4024FA076A42F8">2024-01-30</mods:number>
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<mods:number id="0E73F0BF84EAA4EA96810EFDB6037052">1190</mods:number>
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<mods:identifier id="D93728407D5C92FDA41EADEEBB12108F" type="DOI">http://dx.doi.org/10.3897/zookeys.1190.114958</mods:identifier>
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<treatment id="450DB17E1A665A2B8774486FA1123F88" LSID="urn:lsid:zoobank.org:act:CB8E9D35-3365-4416-81CA-589284A6D42F" httpUri="http://treatment.plazi.org/id/450DB17E1A665A2B8774486FA1123F88" lastPageNumber="259" pageId="0" pageNumber="259">
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<subSubSection id="1776DB39F17CDCC844BD46ADDEBE0418" pageId="0" pageNumber="259" type="nomenclature">
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<paragraph id="2CD2968DE2D9A584A40250631E5E9C3E" pageId="0" pageNumber="259">
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<taxonomicName id="D131D73EAFCC88A738E2C8669AB44849" LSID="https://zoobank.org/CB8E9D35-3365-4416-81CA-589284A6D42F" authority="Nguyen & Vu & Eguchi, 2024" authorityName="Nguyen & Vu & Eguchi" authorityYear="2024" class="Diplopoda" family="Polydesmidae" genus="Pacidesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pacidesmus tuachua" order="Polydesmida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="species" species="tuachua" status="sp. nov.">Pacidesmus tuachua</taxonomicName>
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<taxonomicNameLabel id="6FF36315B09D67B6AD8560EAAB3B2BF3" pageId="0" pageNumber="259">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="4F434811ED44AC173A09206D8F537F23" pageId="0" pageNumber="259" type="description">
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<paragraph id="DDA5BA972515ADD3F289815CE28BDE7C" pageId="0" pageNumber="259">
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<figureCitation id="AFA337C4FF0B19957FCD8FAD167CE777" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Records of polydesmid species in Vietnam and of all known Pacidesmus species." figureDoi="10.3897/zookeys.1190.114958.figure1" httpUri="https://binary.pensoft.net/fig/976107" pageId="0" pageNumber="259">Figs 1</figureCitation>
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<figureCitation id="2778AF362842024E6AAA2D0704ED683F" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Pacidesmus tuachua sp. nov., holotype ♂ (IEBR-Myr 951) A anterior part of body, dorsal view B anterior part of body, lateral view C anterior part of body, ventral view D segments 8 - 10, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1190.114958.figure9" httpUri="https://binary.pensoft.net/fig/976115" pageId="0" pageNumber="259">, 9</figureCitation>
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<figureCitation id="2B5D7F255F5B864A118772B4949B6EA5" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Pacidesmus tuachua sp. nov., holotype ♂ (IEBR-Myr 951) A segments 8 - 10, ventral view B-D posterior part of body, dorsal, vental and lateral views, respectively. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1190.114958.figure10" httpUri="https://binary.pensoft.net/fig/976116" pageId="0" pageNumber="259">, 10</figureCitation>
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||
<figureCitation id="9C299954D9DE71F222F2C859133BB4F1" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Pacidesmus tuachua sp. nov., holotype ♂ (IEBR-Myr 951) A telson, ventral view B telson, lateral view C right antenna, anterior view D gonopods in situ under UV light, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1190.114958.figure11" httpUri="https://binary.pensoft.net/fig/976117" pageId="0" pageNumber="259">, 11</figureCitation>
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||
<figureCitation id="4A53D95F01A2281132916891391C51E1" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pacidesmus tuachua sp. nov., holotype ♂ (IEBR-Myr 951) A anterior part of body, under the UV light, dorsal view B segments 8 - 10, dorsal view C posterior part of body, dorsal view D posterior part of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1190.114958.figure12" httpUri="https://binary.pensoft.net/fig/976118" pageId="0" pageNumber="259">, 12</figureCitation>
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||
<figureCitation id="E4379D5F1A8B3444F30BBA7252593599" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Pacidesmus tuachua sp. nov., holotype ♂ (IEBR-Myr 951) right gonopod, under UV light A mesal view B ventral view C lateral view D dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1190.114958.figure13" httpUri="https://binary.pensoft.net/fig/976119" pageId="0" pageNumber="259">, 13</figureCitation>
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||
<figureCitation id="32FE7404976A16017A56E4514FF93CFD" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Pacidesmus tuachua sp. nov., holotype ♂ (IEBR-Myr 951) right gonopod A mesal view B lateral view. Abbreviations: co = coxite; pref = prefemorite; ca = cannula; en = endomere; pu = puvillus; p 1 = first process; p 2 = second process. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1190.114958.figure14" httpUri="https://binary.pensoft.net/fig/976120" pageId="0" pageNumber="259">, 14</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="80367EF79CD3EFA6C87C0A2D1B149381" pageId="0" pageNumber="259" type="materials_examined">
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||
<paragraph id="7C4369A578ED0833EB48BF8B101252A5" pageId="0" pageNumber="259">Materials examined.</paragraph>
|
||
<paragraph id="0449FDC5E79E799D4372B1B25C802DCA" pageId="0" pageNumber="259">
|
||
<materialsCitation id="94A8DECD89423CA203AA7B80D4375C2B" collectingDate="2022-04-12" collectorName="Anh D. Nguyen" country="Vietnam" county="Tua Chua District" elevation="600" latitude="21.876945" location="Xa Nhe cave" longLatPrecision="21" longitude="103.41334" municipality="Xa Nhe" specimenCount="♂" specimenCount-male="1" stateProvince="Dien Bien Province" typeStatus="Holotype">
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<emphasis id="B43B703BD29AF8E2A236748816308001" bold="true" pageId="0" pageNumber="259">
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<emphasis id="48197A2CDD2E6049306CA98446914105" italics="true" pageId="0" pageNumber="259">
|
||
<typeStatus id="44A1F3FEEABF9BB77294FA630DD5EFB6" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
.
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||
</emphasis>
|
||
<collectingCountry id="20F494ACC3C1AEF571A17B0CB2FE9357" name="Vietnam">Vietnam</collectingCountry>
|
||
•
|
||
<specimenCount id="3E25F8A811CD974826EF6842D75085E2" count="1" type="male">♂</specimenCount>
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||
;
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||
<collectingRegion id="61902D91312B760E4397F876294FCFE6" country="Vietnam" name="Dien Bien">Dien Bien Province</collectingRegion>
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||
,
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||
<collectingCounty id="3E6D8CD311DAD504AB6CF240A34B11AA">Tua Chua District</collectingCounty>
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||
,
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||
<collectingMunicipality id="4363E3083E3F40171C773B76DFE39722">Xa Nhe</collectingMunicipality>
|
||
commune,
|
||
<location id="CE6145950DE634C29EF814AE81E9336D" LSID="urn:lsid:plazi:treatment:450DB17E1A665A2B8774486FA1123F88:CE6145950DE634C29EF814AE81E9336D" country="Vietnam" county="Tua Chua District" latitude="21.876945" longLatPrecision="21" longitude="103.41334" municipality="Xa Nhe" name="Xa Nhe cave" stateProvince="Dien Bien Province">Xa Nhe cave</location>
|
||
;
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||
<elevation id="45EF68C9409EEB434058B5639CF898B6" metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">
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<quantity id="3FA543C3E4ED2576138A15070C152E42" metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">600 m</quantity>
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||
a.s.l.
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</elevation>
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;
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||
<geoCoordinate id="97D7C192FD6877BA62030B1DFD28B53F" degrees="21" direction="north" minutes="52" orientation="latitude" precision="15" seconds="37" value="21.876945">21°52'37"N</geoCoordinate>
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||
,
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<geoCoordinate id="2F2C6E4DC23DF9A89948341E85C72EF5" degrees="103" direction="east" minutes="24" orientation="longitude" precision="15" seconds="48" value="103.41334">103°24'48"E</geoCoordinate>
|
||
;
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||
<collectingDate id="4925FBC8D53AD4FE331EF436305BBCA0" value="2022-04-12">12 April 2022</collectingDate>
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||
;
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||
<collectorName id="8CC7DAB75F01F40B17E47CE36CF2B127">Anh D. Nguyen</collectorName>
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leg.;
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||
<emphasis id="CDF86C5347E90051074E172712BC6E85" bold="true" pageId="0" pageNumber="259">IEBR-Myr 951H.</emphasis>
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||
</materialsCitation>
|
||
</paragraph>
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||
<caption id="7224EA7D65E12C4290D4AC53739CF861" doi="10.3897/zookeys.1190.114958.figure3" httpUri="https://binary.pensoft.net/fig/976109" pageId="0" pageNumber="259" start="Figure 3" startId="F3">
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<paragraph id="C4EFF76D180382A19086D5CE4A72721A" pageId="0" pageNumber="259">
|
||
<emphasis id="8DFD624842D53E2A2A129A8BED926D22" bold="true" pageId="0" pageNumber="259">Figure 3.</emphasis>
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||
<taxonomicName id="9B72873208D99A0A0FE286C6423A2D23" authorityName="Nguyen" authorityYear="2009" class="Diplopoda" family="Polydesmidae" genus="Polydesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polydesmus vietnamicus" order="Polydesmida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="species" species="vietnamicus">
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||
<emphasis id="B32EBBB9AC25AA60136B83C674974731" italics="true" pageId="0" pageNumber="259">Polydesmus vietnamicus</emphasis>
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||
</taxonomicName>
|
||
Nguyen, 2009 from
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||
<collectingRegion id="2C5A6C114B205A9764536BE15A55CE96" country="Vietnam" name="Ha Giang">Ha Giang Province</collectingRegion>
|
||
(IEBR-Myr 808)
|
||
<specimenCount id="AA77C10F4E7611656387958BBB242A27" count="1" type="male">♂</specimenCount>
|
||
<emphasis id="DC385606BAEFB0FA9AD4B5501681A4B5" bold="true" pageId="0" pageNumber="259">A</emphasis>
|
||
midbody segment 8-11, dorsal view
|
||
<emphasis id="EF96B0F0BE3BD770DD466FB2EE390572" bold="true" pageId="0" pageNumber="259">B</emphasis>
|
||
posterior part of body, dorsal view
|
||
<emphasis id="ECA2F9F09EFAD362F014B731FB566460" bold="true" pageId="0" pageNumber="259">C</emphasis>
|
||
posterior part of body, lateral view
|
||
<emphasis id="528E13F857D3C5223673261E2D9D0EE0" bold="true" pageId="0" pageNumber="259">D</emphasis>
|
||
posterior part of body, ventral view. Scale bars:
|
||
<quantity id="9C86E03CD7B1C2FDF2CB1F687530BDC1" metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="3B8BF246FF5524A77CB1C38FE7802A43" pageId="0" pageNumber="259">
|
||
<materialsCitation id="5D5AB944CF00DB86D10EE0AB707709E7" collectingDate="2021-01-12" collectorName="Anh D. Nguyen" country="Vietnam" county="Tua Chua District" elevation="600" latitude="21.876917" location="Kho Chua La cave" longLatPrecision="1" longitude="103.41331" municipality="Xa Nhe" specimenCount="15" specimenCount-female="10" specimenCount-male="5" stateProvince="Dien Bien Province" typeStatus="Paratypes">
|
||
<emphasis id="04AE28B5F80A200D102BC4F5E44E31DD" bold="true" pageId="0" pageNumber="259">
|
||
<emphasis id="472E6FCF8B613CE724F8AB6183BE46D5" italics="true" pageId="0" pageNumber="259">
|
||
<typeStatus id="D260FDE857A1AAA6706AC272D0B8435B" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
.
|
||
</emphasis>
|
||
<collectingCountry id="8C01D3EF572A5A0F9CF51B7DBFD5D697" name="Vietnam">Vietnam</collectingCountry>
|
||
-
|
||
<collectingRegion id="14D078769BC4FED31EE2463CCC649B32" country="Vietnam" name="Dien Bien">Dien Bien Province</collectingRegion>
|
||
•
|
||
<specimenCount id="237A66CB1174E0C39FC600D7B1D3146B" count="5" type="male">5 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="80526EF81384B9BEB99F282E926A952A" count="10" type="female">10 ♀♀</specimenCount>
|
||
;
|
||
<collectingCounty id="BB78941D7E620A95EC07AFAEF8810D4A">Tua Chua District</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="C3C15968F9AFF7180D591E8C3209A3A2">Xa Nhe</collectingMunicipality>
|
||
commune,
|
||
<location id="89D2178DAB49C359FFD3703E18A2B6BF" LSID="urn:lsid:plazi:treatment:450DB17E1A665A2B8774486FA1123F88:89D2178DAB49C359FFD3703E18A2B6BF" country="Vietnam" county="Tua Chua District" latitude="21.876917" longLatPrecision="1" longitude="103.41331" municipality="Xa Nhe" name="Kho Chua La cave" stateProvince="Dien Bien Province">Kho Chua La cave</location>
|
||
;
|
||
<elevation id="D2ABF413D84C09738F51D63B1BD28007" metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">
|
||
<quantity id="CA4FB95CEA6C6E8269FB193A7B420AE7" metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">600 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
;
|
||
<geoCoordinate id="372BE428722F68916C4F1F21121610CC" degrees="21" direction="north" minutes="52" orientation="latitude" precision="1" seconds="36.9" value="21.876917">21°52'36.9"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="7B97E35B7DFFC564873C2EB53E9D40C4" degrees="103" direction="east" minutes="24" orientation="longitude" precision="1" seconds="47.9" value="103.41331">103°24'47.9"E</geoCoordinate>
|
||
;
|
||
<collectingDate id="64DACEF960AB9526639F07839C01C5FD" value="2021-01-12">12 January 2021</collectingDate>
|
||
;
|
||
<collectorName id="7FAB318CC40D2F6BC35B6C75B957850E">Anh D. Nguyen</collectorName>
|
||
leg.; IEBR-Myr 899
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="F02FB1672AB6CD114286B8B0D4B28861" collectingDate="2021-01-12" collectorName="Anh D. Nguyen" country="Vietnam" county="Tua Chua District" elevation="600" latitude="21.876917" location="Kho Chua La cave" longLatPrecision="1" longitude="103.41331" municipality="Xa Nhe" specimenCount="7" specimenCount-female="4" specimenCount-male="3" stateProvince="Dien Bien Province" typeStatus="paratype">
|
||
<specimenCount id="E1F6ADE3BB46429F3D1EDA2522FF1310" count="3" type="male">3 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="72C212883AD741D6AE5428787E1204EC" count="4" type="female">4 ♀♀</specimenCount>
|
||
;
|
||
<collectingCounty id="0AABAB839172B1F35C2D5EB216F08C94">Tua Chua District</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="F31787A389DDFF503EA2498853938108">Xa Nhe</collectingMunicipality>
|
||
commune,
|
||
<location id="911FFB4A18DD2FBC9BFA3C99A17893F5" LSID="urn:lsid:plazi:treatment:450DB17E1A665A2B8774486FA1123F88:911FFB4A18DD2FBC9BFA3C99A17893F5" country="Vietnam" county="Tua Chua District" latitude="21.876917" longLatPrecision="1" longitude="103.41331" municipality="Xa Nhe" name="Kho Chua La cave" stateProvince="Dien Bien Province">Kho Chua La cave</location>
|
||
;
|
||
<elevation id="A6B437CB0B76961B9129193A97FC2C47" metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">
|
||
<quantity id="30D5EFE9DB6DE3D58BA879C122129523" metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">600 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
;
|
||
<geoCoordinate id="CD92093AFC4CFEB24204647EE761F74F" degrees="21" direction="north" minutes="52" orientation="latitude" precision="1" seconds="36.9" value="21.876917">21°52'36.9"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="99ADA781118B9690A5A9CA2FC200B74E" degrees="103" direction="east" minutes="24" orientation="longitude" precision="1" seconds="47.9" value="103.41331">103°24'47.9"E</geoCoordinate>
|
||
;
|
||
<collectingDate id="24F74AF937416E3620A6A7C601F04CC0" value="2021-01-12">12 January 2021</collectingDate>
|
||
;
|
||
<collectorName id="80B3CBA0440D4EE6477872AE7C87D22C">Anh D. Nguyen</collectorName>
|
||
leg.; IEBR-Myr 900
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="E59936772FC5263FC44ADBE0AB757510" collectingDate="2022-04-12" collectorName="Anh D. Nguyen" country="Vietnam" county="Tua Chua District" elevation="600" latitude="21.876917" location="Kho Chua La cave" longLatPrecision="1" longitude="103.41331" municipality="Xa Nhe" specimenCount="9" specimenCount-female="5" specimenCount-male="4" stateProvince="Dien Bien Province" typeStatus="paratype">
|
||
<specimenCount id="86EFA059A8B758FE68580FA523DAA92E" count="4" type="male">4 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="5A70D073D2F1A49FA421D43A8AB7C150" count="5" type="female">5 ♀♀</specimenCount>
|
||
;
|
||
<collectingCounty id="CDD003518BD39DC0EFFF58601F05E56B">Tua Chua District</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="82E46F747A3D3E1C087BB1E23E4A3DA3">Xa Nhe</collectingMunicipality>
|
||
commune,
|
||
<location id="8B185EDF6F4DFD3692064B1363A2EBAD" LSID="urn:lsid:plazi:treatment:450DB17E1A665A2B8774486FA1123F88:8B185EDF6F4DFD3692064B1363A2EBAD" country="Vietnam" county="Tua Chua District" latitude="21.876917" longLatPrecision="1" longitude="103.41331" municipality="Xa Nhe" name="Kho Chua La cave" stateProvince="Dien Bien Province">Kho Chua La cave</location>
|
||
;
|
||
<elevation id="BD45C3189D7FF0C16C02FAC3EDF4C7CE" metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">
|
||
<quantity id="0D4D1D4B717722EFD003A4843174AF6B" metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">600 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
;
|
||
<geoCoordinate id="E8D5380FB543CB2E65644C8F8A5BDB81" degrees="21" direction="north" minutes="52" orientation="latitude" precision="1" seconds="36.9" value="21.876917">21°52'36.9"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="B6D8ADA6B220C56F60AA78CEA6680537" degrees="103" direction="east" minutes="24" orientation="longitude" precision="1" seconds="47.9" value="103.41331">103°24'47.9"E</geoCoordinate>
|
||
;
|
||
<collectingDate id="B45E7FB1E9BE11CD6EA8811D99B3026D" value="2022-04-12">12 April 2022</collectingDate>
|
||
;
|
||
<collectorName id="5EE5FBEB2421734C5025767A640A9EBB">Anh D. Nguyen</collectorName>
|
||
leg.; IEBR-Myr 951P
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="77738FDB8BF5544D924341DEFDF48BD8" collectingDate="2022-04-12" collectorName="Anh D. Nguyen" country="Vietnam" county="Tua Chua District" elevation="600" latitude="21.876945" location="Xa Nhe cave" longLatPrecision="21" longitude="103.41334" municipality="Xa Nhe" specimenCount="9" specimenCount-female="5" specimenCount-male="4" stateProvince="Dien Bien Province" typeStatus="paratype">
|
||
<specimenCount id="787560C1D5F34F595A5DC6E92AD54884" count="4" type="male">4 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="4BC70C173D209A10F1954221DA1CD7F8" count="5" type="female">5 ♀♀</specimenCount>
|
||
;
|
||
<collectingCounty id="91A43DB71E12D41F50254AFAC426CA5F">Tua Chua District</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="507002150D471F42685175E2664E4D88">Xa Nhe</collectingMunicipality>
|
||
commune,
|
||
<location id="CBE92502CD9607FB3094E6F57323FBB5" LSID="urn:lsid:plazi:treatment:450DB17E1A665A2B8774486FA1123F88:CBE92502CD9607FB3094E6F57323FBB5" country="Vietnam" county="Tua Chua District" latitude="21.876945" longLatPrecision="21" longitude="103.41334" municipality="Xa Nhe" name="Xa Nhe cave" stateProvince="Dien Bien Province">Xa Nhe cave</location>
|
||
;
|
||
<elevation id="0E5DD30A2411E78B1CA6626A9E3BA483" metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">
|
||
<quantity id="8588F9B3C72B2E30EFF650C0D3CCB6F1" metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">600 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
;
|
||
<geoCoordinate id="5F8F9B931EB3CBD43D2799A14A3AA742" degrees="21" direction="north" minutes="52" orientation="latitude" precision="15" seconds="37" value="21.876945">21°52'37"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="41E5CF52B4E81B71AA1226E791617F82" degrees="103" direction="east" minutes="24" orientation="longitude" precision="15" seconds="48" value="103.41334">103°24'48"E</geoCoordinate>
|
||
;
|
||
<collectingDate id="CEC12537623B391AABC543F98806CF9F" value="2022-04-12">12 April 2022</collectingDate>
|
||
;
|
||
<collectorName id="B6DA45DF4759830F9D2E937E66C4D7F6">Anh D. Nguyen</collectorName>
|
||
leg.; IEBR-Myr 952
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="853489CA791F4FEC39799E6F925A0662" collectingDate="2022-04-12" collectorName="Anh D. Nguyen" country="Vietnam" county="Tua Chua District" elevation="600" latitude="21.876917" location="Kho Chua La cave" longLatPrecision="1" longitude="103.41331" municipality="Xa Nhe" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Dien Bien Province" typeStatus="paratype">
|
||
<specimenCount id="3D8A2FA46368A603B45D912301FDAA30" count="2" type="male">2 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="3624D355828941051D56B5C934B71655" count="4" type="female">4 ♀♀</specimenCount>
|
||
;
|
||
<collectingCounty id="95EE91F1DF1097BE5B5DB345177CF226">Tua Chua District</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="D719C83E1CFB7DC197243F57A05D6271">Xa Nhe</collectingMunicipality>
|
||
commune,
|
||
<location id="40A95A91F4B5E9EBA026A3172F715C21" LSID="urn:lsid:plazi:treatment:450DB17E1A665A2B8774486FA1123F88:40A95A91F4B5E9EBA026A3172F715C21" country="Vietnam" county="Tua Chua District" latitude="21.876917" longLatPrecision="1" longitude="103.41331" municipality="Xa Nhe" name="Kho Chua La cave" stateProvince="Dien Bien Province">Kho Chua La cave</location>
|
||
;
|
||
<elevation id="F2977F41D634E4CF4CE5AF55AF038C3C" metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">
|
||
<quantity id="952EAF852FF70C1AABEC5FE84CF5B917" metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">600 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
;
|
||
<geoCoordinate id="9E3E80885E8DD3CFE87E57BFB71A108F" degrees="21" direction="north" minutes="52" orientation="latitude" precision="1" seconds="36.9" value="21.876917">21°52'36.9"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="2408ACF1057655EB4687EC6392FE1445" degrees="103" direction="east" minutes="24" orientation="longitude" precision="1" seconds="47.9" value="103.41331">103°24'47.9"E</geoCoordinate>
|
||
;
|
||
<collectingDate id="6D8984E1CFE06B96560B69CF31BFFE0B" value="2022-04-12">12 April 2022</collectingDate>
|
||
;
|
||
<collectorName id="C7A62F3564B7411C846AD55C2A458C15">Anh D. Nguyen</collectorName>
|
||
leg.; IEBR-Myr 953
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="13838BFA8A11F7FFC8BB347722B3DA3E" doi="10.3897/zookeys.1190.114958.figure4" httpUri="https://binary.pensoft.net/fig/976110" pageId="0" pageNumber="259" start="Figure 4" startId="F4">
|
||
<paragraph id="8F22224539D2D621AB5F59CF00F31968" pageId="0" pageNumber="259">
|
||
<emphasis id="042D79A85120BC144F0627D00BB634A8" bold="true" pageId="0" pageNumber="259">Figure 4.</emphasis>
|
||
<taxonomicName id="8ADEAD638D6501402E17B8916DA7ADEA" authorityName="Nguyen" authorityYear="2009" class="Diplopoda" family="Polydesmidae" genus="Polydesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polydesmus vietnamicus" order="Polydesmida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="species" species="vietnamicus">
|
||
<emphasis id="EE96A5201C72AAB2C62566EDC94EDED6" italics="true" pageId="0" pageNumber="259">Polydesmus vietnamicus</emphasis>
|
||
</taxonomicName>
|
||
Nguyen, 2009 from
|
||
<collectingRegion id="2EFFB3367FE9DAC34BE52F539776215C" country="Vietnam" name="Ha Giang">Ha Giang Province</collectingRegion>
|
||
(IEBR-Myr 808)
|
||
<specimenCount id="D41F384CDB7024C98541991470183C7A" count="1" type="male">♂</specimenCount>
|
||
and
|
||
<specimenCount id="A47166FF79E34E970B48D36AA72B9E51" count="1" type="female">♀</specimenCount>
|
||
under the UV light
|
||
<emphasis id="60999F8A1274E161F63650401BC5F6DA" bold="true" pageId="0" pageNumber="259">A</emphasis>
|
||
<specimenCount id="FDF526FA97D1E958DF13B5AFB75214AD" count="1" type="male">♂</specimenCount>
|
||
, anterior part of body, dorsal view
|
||
<emphasis id="B255E76A85AC7F66C1D58EFEA456ACDE" bold="true" pageId="0" pageNumber="259">B</emphasis>
|
||
<specimenCount id="3E0228135FA7732D56507D33790B7A18" count="1" type="male">♂</specimenCount>
|
||
, anterior part of body, ventral view
|
||
<emphasis id="DBA885FEA70122FF838303CDECA373D5" bold="true" pageId="0" pageNumber="259">C</emphasis>
|
||
<specimenCount id="B412659791184102F73A65ABBD2E8524" count="1" type="female">♀</specimenCount>
|
||
, anterior part of body, ventral view
|
||
<emphasis id="A65732E40A51AC99DAF4CB70F73E9C92" bold="true" pageId="0" pageNumber="259">D</emphasis>
|
||
<specimenCount id="F68BABC26203D5C33E7CA2A5073C6D22" count="1" type="male">♂</specimenCount>
|
||
, midbody segment 8-10, dorsal view. Scale bars:
|
||
<quantity id="BB0396F0214E33EC827F71C2CF797CBD" metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="64C7E9D9D8D0E5830A920E59BB5DEF05" pageId="0" pageNumber="259" type="diagnosis">
|
||
<paragraph id="CE164B4C11804F589DA26468D16463BA" pageId="0" pageNumber="259">Diagnosis.</paragraph>
|
||
<paragraph id="A41D73712C8E9B9DF911D3EF28374C4A" pageId="0" pageNumber="259">The new species can be distinguished from its congeners by a combination of the following features: unpigmented colouration, small size (midbody width <4.0 mm), head narrower than collum, absence of sphaerotrichomes, lateral budges on male prefemora, subfalcate gonopod telopodite, absence of exomere, endomere with an acute triangular process distally and a broad triangular process medially, and endomere tip slightly and unequally bifid.</paragraph>
|
||
<caption id="9BBC43A2A01BE661A008F2F36DD8BFA8" doi="10.3897/zookeys.1190.114958.figure5" httpUri="https://binary.pensoft.net/fig/976111" pageId="0" pageNumber="259" start="Figure 5" startId="F5">
|
||
<paragraph id="177AF8B64A7DD8ECC9181075675324D1" pageId="0" pageNumber="259">
|
||
<emphasis id="2462DB902097692D84F3E28772FB9BF9" bold="true" pageId="0" pageNumber="259">Figure 5.</emphasis>
|
||
<taxonomicName id="676DC5E58370070AC4AFC28FE49CD6A0" authorityName="Nguyen" authorityYear="2009" class="Diplopoda" family="Polydesmidae" genus="Polydesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polydesmus vietnamicus" order="Polydesmida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="species" species="vietnamicus">
|
||
<emphasis id="913E94C97F5E5379A30309D5A890FEF2" italics="true" pageId="0" pageNumber="259">Polydesmus vietnamicus</emphasis>
|
||
</taxonomicName>
|
||
Nguyen, 2009 from Ha Giang Province (IEBR-Myr 808) ♂, under the UV light
|
||
<emphasis id="43089488A1834DE32A11580B7232995A" bold="true" pageId="0" pageNumber="259">A</emphasis>
|
||
posterior part of body, dorsal view
|
||
<emphasis id="79FAE20DC2F0760CC0C9EB07B7049FAE" bold="true" pageId="0" pageNumber="259">B</emphasis>
|
||
posterior part of body, ventral view
|
||
<emphasis id="ED88EBB1FD5DF6560CD1ECFFE6E0175C" bold="true" pageId="0" pageNumber="259">C</emphasis>
|
||
telson, dorsal view
|
||
<emphasis id="87A5B9F1DE335992485C220D3260C666" bold="true" pageId="0" pageNumber="259">D</emphasis>
|
||
telson, ventral view. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="F9092AAB165C4A65D6828509225D1FE2" pageId="0" pageNumber="259">
|
||
The species is truly cavernicolous, characterized by white or unpigmented colour and living within a cave. As a troglobiont species, it groups with all 12 troglobiont or troglophile congeners from China (Table
|
||
<tableCitation id="A85E9940696F60ACCBF47F7CF9D52786" captionStart="Table 1" captionStartId="T1" captionText="Table 1. List of all known species of the genus Pacidesmus Golovatch, 1991." httpUri="http://table.plazi.org/id/0C5F5CAECAFE591431E86E1E73C00D1C" pageId="0" pageNumber="259" tableUuid="0C5F5CAECAFE591431E86E1E73C00D1C">1</tableCitation>
|
||
). However, this species differs from all of these, except
|
||
<taxonomicName id="2B6B297617A0B47C4A6D15DC9C900ABF" lsidName="P. bifidus" pageId="0" pageNumber="259" rank="species" species="bifidus">
|
||
<emphasis id="C43A7AE30A08DA9CECFDE94112A93470" italics="true" pageId="0" pageNumber="259">P. bifidus</emphasis>
|
||
</taxonomicName>
|
||
from the Hengli Xin Don Cave, Guangxi Province in the absence of an exomere and the gonopod telopodite showing no additional processes; the endomere also has two additional processes, and the tip of the endomere bears two tiny teeth. The new species is similar to
|
||
<taxonomicName id="89DCC266EA45C9541AB6C0A6726F7D68" lsidName="P. bifidus" pageId="0" pageNumber="259" rank="species" species="bifidus">
|
||
<emphasis id="5B6A931F34AD06D05239189721B1B015" italics="true" pageId="0" pageNumber="259">P. bifidus</emphasis>
|
||
</taxonomicName>
|
||
in having a troglomorphic appearance, the absence of an exomere, and a bifid tip of the endomere, but it differs in having two tiny teeth at the tip of the endomere tip (vs two long processes in
|
||
<taxonomicName id="EB10D23FF1C25A568007A7779D64C0CD" lsidName="P. bifidus" pageId="0" pageNumber="259" rank="species" species="bifidus">
|
||
<emphasis id="10639CD7525603557F5DCCF2C520363A" italics="true" pageId="0" pageNumber="259">P. bifidus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<caption id="0C5F5CAECAFE591431E86E1E73C00D1C" ID-Table-UUID="0C5F5CAECAFE591431E86E1E73C00D1C" httpUri="http://table.plazi.org/id/0C5F5CAECAFE591431E86E1E73C00D1C" pageId="0" pageNumber="259" start="Table 1" startId="T1">
|
||
<paragraph id="29E30F510F1D311CF826483BD9861823" pageId="0" pageNumber="259">
|
||
<emphasis id="2716FDE5A7FAC7A0B0101673CFEB4FF0" bold="true" pageId="0" pageNumber="259">Table 1.</emphasis>
|
||
List of all known species of the genus
|
||
<taxonomicName id="52A16601417CEF91E1DD6D4F5D51A6F5" authorityName="Golovatch" authorityYear="1991" class="Diplopoda" family="Polydesmidae" genus="Pacidesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pacidesmus" order="Polydesmida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="02F23BC8BFAED5A0AEAE9D9F600C9771" italics="true" pageId="0" pageNumber="259">Pacidesmus</emphasis>
|
||
</taxonomicName>
|
||
Golovatch, 1991.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="166CB8CBF62838FA5BC8CB4BBF48BC70" pageId="0" pageNumber="259">
|
||
<table id="E686389ABA379A59F7176496140C3826" pageId="0" pageNumber="259">
|
||
<tr id="8C66FBA5A3338699975DEE374DD385B2" pageId="0" pageNumber="259">
|
||
<th id="BC620A8D22B3DFD58AE3E8F176FAD6E5" colspan="1" pageId="0" pageNumber="259" rowspan="1">Species</th>
|
||
<th id="1C9A9255906D9A3EBAFA62D24BFF0262" colspan="1" pageId="0" pageNumber="259" rowspan="1">Localities</th>
|
||
</tr>
|
||
<tr id="AE90F5D11FB7A30C57B02DE683C3098F" pageId="0" pageNumber="259">
|
||
<td id="ABD38AF05F945895148B0A0A0C1D6037" colspan="1" pageId="0" pageNumber="259" rowspan="1">
|
||
<taxonomicName id="EFE6BB24DF6B2FFB0026A0978C4742DE" authorityName="Golovatch, Geoffroy & Mauries" authorityYear="2010" class="Diplopoda" family="Polydesmidae" genus="Pacidesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pacidesmus armatus" order="Polydesmida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="species" species="armatus">
|
||
<emphasis id="21A36D4BCD1B04FCDA578AB78BD96891" italics="true" pageId="0" pageNumber="259">Pacidesmus armatus</emphasis>
|
||
</taxonomicName>
|
||
Golovatch, Geoffroy &
|
||
<normalizedToken id="D2E73A6D0DF9F2A6CFB52D27DDD23907" originalValue="Mauriès">Mauries</normalizedToken>
|
||
, 2010
|
||
</td>
|
||
<td id="4277936C30BF5CC60BBBE7AB5F0485DD" colspan="1" pageId="0" pageNumber="259" rowspan="1">
|
||
China, Guangxi Prov., Huanjiang, Cave Xiao Lan Dong (
|
||
<bibRefCitation id="E6A6DE797388FB29FEFD444C42338E85" author="Golovatch, SI" journalOrPublisher="Tropical Natural History" pageId="0" pageNumber="259" pagination="159 - 169" refId="B17" refString="Golovatch, SI, Geoffroy, JJ, Mauries, JP, 2010. Review of the Millipede Genus Pacidesmus Golovatch, 1991, with Descriptions of Three New Species from Caves in Southern China (Diplopoda: Polydesmida: Polydesmidae). Tropical Natural History 10: 159 - 169" title="Review of the Millipede Genus Pacidesmus Golovatch, 1991, with Descriptions of Three New Species from Caves in Southern China (Diplopoda: Polydesmida: Polydesmidae)." volume="10" year="2010">Golovatch et al. 2010</bibRefCitation>
|
||
)
|
||
</td>
|
||
</tr>
|
||
<tr id="CF83F1D2FD801E5EC8E4B8AE05030844" pageId="0" pageNumber="259">
|
||
<td id="29ACB626A7525E9A90DC4EBECCF196F6" colspan="1" pageId="0" pageNumber="259" rowspan="1">
|
||
<taxonomicName id="23E29628C0662BFB15D23F376B36DEB5" authorityName="Golovatch, Geoffroy & Mauries" authorityYear="2010" class="Diplopoda" family="Polydesmidae" genus="Pacidesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pacidesmus bedosae" order="Polydesmida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="species" species="bedosae">
|
||
<emphasis id="D1C73B7A34B16F82B6654A1518DED35E" italics="true" pageId="0" pageNumber="259">Pacidesmus bedosae</emphasis>
|
||
</taxonomicName>
|
||
Golovatch, Geoffroy &
|
||
<normalizedToken id="69D8C4BD12C3A8C60E0008F1C496EC09" originalValue="Mauriès">Mauries</normalizedToken>
|
||
, 2010
|
||
</td>
|
||
<td id="A51BF8FFCC837EE031934BEE04F217E3" colspan="1" pageId="0" pageNumber="259" rowspan="1">
|
||
China, Guangxi Prov., Huanjiang, Cave Dong Tu Dong (
|
||
<bibRefCitation id="91AC347ECA0AEBE61550DF38BD179A92" author="Golovatch, SI" journalOrPublisher="Tropical Natural History" pageId="0" pageNumber="259" pagination="159 - 169" refId="B17" refString="Golovatch, SI, Geoffroy, JJ, Mauries, JP, 2010. Review of the Millipede Genus Pacidesmus Golovatch, 1991, with Descriptions of Three New Species from Caves in Southern China (Diplopoda: Polydesmida: Polydesmidae). Tropical Natural History 10: 159 - 169" title="Review of the Millipede Genus Pacidesmus Golovatch, 1991, with Descriptions of Three New Species from Caves in Southern China (Diplopoda: Polydesmida: Polydesmidae)." volume="10" year="2010">Golovatch et al. 2010</bibRefCitation>
|
||
)
|
||
</td>
|
||
</tr>
|
||
<tr id="28C060E78DD4CC3BC6A14D4F0FE175E1" pageId="0" pageNumber="259">
|
||
<td id="51A688BC474A6540503632DCCA1E1530" colspan="1" pageId="0" pageNumber="259" rowspan="1">
|
||
<taxonomicName id="54612360FEE8635065D6C57ECF210D4C" authorityName="Golovatch & Geoffroy" authorityYear="2014" class="Diplopoda" family="Polydesmidae" genus="Pacidesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pacidesmus bifidus" order="Polydesmida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="species" species="bifidus">
|
||
<emphasis id="49A572562A1F685A3788D85ACEF39B79" italics="true" pageId="0" pageNumber="259">Pacidesmus bifidus</emphasis>
|
||
</taxonomicName>
|
||
Golovatch & Geoffroy, 2014
|
||
</td>
|
||
<td id="4239938F4094049FC1B1722420A2D37F" colspan="1" pageId="0" pageNumber="259" rowspan="1">
|
||
China, Guangxi Prov., Cave Hengli Xin Dong near Fengshan (Fengshan Xian) (
|
||
<bibRefCitation id="06F0D86182F8DC3185DFC246C459C969" DOI="https://doi.org/10.15298/rusentj.23.2.01" author="Golovatch, SI" journalOrPublisher="Russian Entomological Journal" pageId="0" pageNumber="259" pagination="91 - 105" refId="B13" refString="Golovatch, SI, Geoffroy, JJ, 2014. On some new or poorly-known species of the millipede family Polydesmidae from southern China (Diplopoda: Polydesmida). Russian Entomological Journal 23 (1): 91 - 105, DOI: https://doi.org/10.15298/rusentj.23.2.01" title="On some new or poorly-known species of the millipede family Polydesmidae from southern China (Diplopoda: Polydesmida)." url="https://doi.org/10.15298/rusentj.23.2.01" volume="23" year="2014">Golovatch and Geoffroy 2014</bibRefCitation>
|
||
)
|
||
</td>
|
||
</tr>
|
||
<tr id="75C7C68B406D2EB2CF68505BB3B18EFF" pageId="0" pageNumber="259">
|
||
<td id="937AE61BEC598F9E28C38B179497563F" colspan="1" pageId="0" pageNumber="259" rowspan="1">
|
||
<taxonomicName id="4016DDDB6BC881C5615F1594F6AB7910" authorityName="Golovatch & Geoffroy" authorityYear="2006" class="Diplopoda" family="Polydesmidae" genus="Pacidesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pacidesmus martensi" order="Polydesmida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="species" species="martensi">
|
||
<emphasis id="6E55364217EC83FC9DC43F3620C8DBA4" italics="true" pageId="0" pageNumber="259">Pacidesmus martensi</emphasis>
|
||
</taxonomicName>
|
||
Golovatch & Geoffroy, 2006
|
||
</td>
|
||
<td id="1A21BE6E71F947C83BE5591553243BFA" colspan="1" pageId="0" pageNumber="259" rowspan="1">
|
||
China, Guizhou Prov., Qianxi County, Hong Lin Town, Ishui Luo Dong Cave China, Guizhou Prov., Dafang County, Yangzhamba Village, Hei Dong Cave China, Guizhou Province, Qianxi County, Honglin Town, Jisha Village, I Dong Cave (
|
||
<bibRefCitation id="E0A5710D6D8D3D34022794ED23219B52" author="Golovatch, SI" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="259" pagination="81 - 89" refId="B16" refString="Golovatch, SI, Geoffroy, J-J, Mauries, J-P, 2007. Several new or poorly-known cavernicolous millipedes from southern China. Arthropoda Selecta 15: 81 - 89" title="Several new or poorly-known cavernicolous millipedes from southern China." volume="15" year="2007">Golovatch et al. 2007</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7A8D7E159C08CC6ED3551A959E8D7B4B" DOI="https://doi.org/10.11646/zootaxa.1325.1.24" author="Golovatch, SI" journalOrPublisher="Zootaxa" pageId="0" pageNumber="259" pagination="363 - 368" refId="B12" refString="Golovatch, SI, Geoffroy, J-J, 2006. Review of the Southeast Asian millipede genus Pacidesmus Golovatch, with the description of a new troglobitic species from southern China (Diplopoda: Polydesmida: Polydesmidae). Zootaxa 1325 (1): 363 - 368, DOI: https://doi.org/10.11646/zootaxa.1325.1.24" title="Review of the Southeast Asian millipede genus Pacidesmus Golovatch, with the description of a new troglobitic species from southern China (Diplopoda: Polydesmida: Polydesmidae)." url="https://doi.org/10.11646/zootaxa.1325.1.24" volume="1325" year="2006">Golovatch and Geoffroy 2006</bibRefCitation>
|
||
)
|
||
</td>
|
||
</tr>
|
||
<tr id="27E0B36FA2244591A34E00B85860DB36" pageId="0" pageNumber="259">
|
||
<td id="37722C4D402CB8CA35C5E25859617BFD" colspan="1" pageId="0" pageNumber="259" rowspan="1">
|
||
<taxonomicName id="35A335C63541F6EFE80F001396B245CB" authorityName="Golovatch" authorityYear="1991" class="Diplopoda" family="Polydesmidae" genus="Pacidesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pacidesmus shelleyi" order="Polydesmida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="species" species="shelleyi">
|
||
<emphasis id="AAC2353479372F3B1949360C9712AE74" italics="true" pageId="0" pageNumber="259">Pacidesmus shelleyi</emphasis>
|
||
</taxonomicName>
|
||
Goolovatch, 1991
|
||
</td>
|
||
<td id="A22B7BA2CB7F46D3E1787EA6DE55473A" colspan="1" pageId="0" pageNumber="259" rowspan="1">
|
||
Thailand, Chieng Mai Province, Doi Inthanon National Park (
|
||
<bibRefCitation id="03BB55F4D8A15DF3F86008291B910EBC" author="Golovatch, SI" journalOrPublisher="Steenstrupia (Copenhagen)" pageId="0" pageNumber="259" pagination="141 - 159" refId="B9" refString="Golovatch, SI, 1991. The millipede family Polydesmidae in Southeast Asia, with notes on phylogeny (Diplopoda: Polydesmida). Steenstrupia (Copenhagen) 17: 141 - 159" title="The millipede family Polydesmidae in Southeast Asia, with notes on phylogeny (Diplopoda: Polydesmida)." volume="17" year="1991">Golovatch 1991</bibRefCitation>
|
||
)
|
||
</td>
|
||
</tr>
|
||
<tr id="D8CE9AE47556C8F80787C8DD62C7D420" pageId="0" pageNumber="259">
|
||
<td id="3F60656E66C65C53AD23DEAAAA55C345" colspan="1" pageId="0" pageNumber="259" rowspan="1">
|
||
<taxonomicName id="4E51FA64D8A8B23FC6238DE4A54DEFBE" baseAuthorityName="Golovatch & Hoffman" baseAuthorityYear="1989" class="Diplopoda" family="Polydesmidae" genus="Pacidesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pacidesmus sinensis" order="Polydesmida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="species" species="sinensis">
|
||
<emphasis id="B575A0DCECFE74E2DA68591B72729531" italics="true" pageId="0" pageNumber="259">Pacidesmus sinensis</emphasis>
|
||
</taxonomicName>
|
||
(Golovatch & Hoffman, 1989)
|
||
</td>
|
||
<td id="3051FEAF233AFCDFDD33CE33EAD6D3F7" colspan="1" pageId="0" pageNumber="259" rowspan="1">
|
||
China, Guizhou Province, Ziyun County, Getuhe National Geopark, Suidao Dong Cave (
|
||
<bibRefCitation id="26F56AF0B9B23AE19E99C7F3511FCADD" author="Liu, WX" journalOrPublisher="The Raffles Bulletin of Zoology" pageId="0" pageNumber="259" pagination="78 - 87" refId="B24" refString="Liu, WX, Golovatch, SI, 2020. Three new cavernicolous species and three new records of the millipede genus Pacidesmus from southern China (Diplopoda, Polydesmida, Polydesmidae). The Raffles Bulletin of Zoology 35: 78 - 87" title="Three new cavernicolous species and three new records of the millipede genus Pacidesmus from southern China (Diplopoda, Polydesmida, Polydesmidae)." volume="35" year="2020">Liu and Golovatch 2020</bibRefCitation>
|
||
) A cave in Guizhou Province and Cave Kaikou Dong, Zhenning County, Guizhou Province, China (
|
||
<bibRefCitation id="09B90EE755E40AF5A83D9BBAE8A6093E" author="Loksa, I" journalOrPublisher="Acta Zoologica Academiae Scientiarum Hungaricae" pageId="0" pageNumber="259" pagination="135 - 148" refId="B25" refString="Loksa, I, 1960. Einige neue Diplopoden- und Chilopodenaren aus chinesischen Hoehlen. Acta Zoologica Academiae Scientiarum Hungaricae 6: 135 - 148" title="Einige neue Diplopoden- und Chilopodenaren aus chinesischen Hoehlen." volume="6" year="1960">Loksa 1960</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="938689C4BBB1608FA69FA9967A12281B" DOI="https://doi.org/10.1080/03946975.1989.10539436" author="Golovatch, SI" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="259" pagination="159 - 164" refId="B14" refString="Golovatch, SI, Hoffman, RL, 1989. Identity of Polydesmus hamatus Brandt, 1841, a Malagasy milliped (Diplopoda Polydesmida Dalodesmidae). Tropical Zoology 2 (2): 159 - 164, DOI: https://doi.org/10.1080/03946975.1989.10539436" title="Identity of Polydesmus hamatus Brandt, 1841, a Malagasy milliped (Diplopoda Polydesmida Dalodesmidae)." url="https://doi.org/10.1080/03946975.1989.10539436" volume="2" year="1989">Golovatch and Hoffman 1989</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="76CDF351D2BD628009BC23DB2E022988" author="Chen, JX" journalOrPublisher="Journal of Nanjing University" pageId="0" pageNumber="259" pagination="277 - 281" refId="B4" refString="Chen, JX, Meng, WX, 1990. A revision of the specific name of Polydesmus hamatus Loksa, 1960 with a redescription (Diplopoda: Polydesmida: Polydesmidae). Journal of Nanjing University 26: 277 - 281" title="A revision of the specific name of Polydesmus hamatus Loksa, 1960 with a redescription (Diplopoda: Polydesmida: Polydesmidae)." volume="26" year="1990">Chen and Meng 1990</bibRefCitation>
|
||
)
|
||
</td>
|
||
</tr>
|
||
<tr id="68CB2BE8044B0E17D20A449D46CFE81E" pageId="0" pageNumber="259">
|
||
<td id="DD32C7201E6ECBDE5FE8889E05F1A7AB" colspan="1" pageId="0" pageNumber="259" rowspan="1">
|
||
<taxonomicName id="89957D75522274600D0EE0D923C07186" authorityName="Golovatch, Geoffroy & Mauriès" authorityYear="2006" class="Diplopoda" family="Polydesmidae" genus="Pacidesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pacidesmus superdraco" order="Polydesmida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="species" species="superdraco">
|
||
<emphasis id="7F034BD88C2383171C7979CAB3A2BCF2" italics="true" pageId="0" pageNumber="259">Pacidesmus superdraco</emphasis>
|
||
</taxonomicName>
|
||
Golovatch, Geoffroy &
|
||
<normalizedToken id="089CD2C857F7015B14B5E0B0FC3FC587" originalValue="Mauriès">Mauries</normalizedToken>
|
||
, 2006
|
||
</td>
|
||
<td id="C49443DB0532D1248E63F042F218623C" colspan="1" pageId="0" pageNumber="259" rowspan="1">
|
||
Cave Laitai Dong, Libo County, Guizhou Province (
|
||
<bibRefCitation id="D337ED67AE2DF3AE40BD358C68557582" author="Golovatch, SI" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="259" pagination="81 - 89" refId="B16" refString="Golovatch, SI, Geoffroy, J-J, Mauries, J-P, 2007. Several new or poorly-known cavernicolous millipedes from southern China. Arthropoda Selecta 15: 81 - 89" title="Several new or poorly-known cavernicolous millipedes from southern China." volume="15" year="2007">Golovatch et al. 2007</bibRefCitation>
|
||
)
|
||
</td>
|
||
</tr>
|
||
<tr id="2C9B33B9721EF99936F6FFB72B391324" pageId="0" pageNumber="259">
|
||
<td id="57E4DAD66E29E44DB9E2F86501C105CE" colspan="1" pageId="0" pageNumber="259" rowspan="1">
|
||
<taxonomicName id="BE622B25DED658D1234D49A280AF124A" authorityName="Golovatch, Geoffroy & Mauries" authorityYear="2010" class="Diplopoda" family="Polydesmidae" genus="Pacidesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pacidesmus tiani" order="Polydesmida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="species" species="tiani">
|
||
<emphasis id="8DFE0F97DC915F3B34AD6C7A96F8F471" italics="true" pageId="0" pageNumber="259">Pacidesmus tiani</emphasis>
|
||
</taxonomicName>
|
||
Golovatch, Geoffroy &
|
||
<normalizedToken id="4D3605AC20985D0F00E79F62CA3F7189" originalValue="Mauriès">Mauries</normalizedToken>
|
||
, 2010
|
||
</td>
|
||
<td id="36C0EBE50A2190A2AE47B6D0CD3CBBC5" colspan="1" pageId="0" pageNumber="259" rowspan="1">
|
||
China, Guangxi Prov., Huanjiang, Cave Gang Lai Dong (
|
||
<bibRefCitation id="952D117EED603517A991EED50F9CCA2F" author="Golovatch, SI" journalOrPublisher="Tropical Natural History" pageId="0" pageNumber="259" pagination="159 - 169" refId="B17" refString="Golovatch, SI, Geoffroy, JJ, Mauries, JP, 2010. Review of the Millipede Genus Pacidesmus Golovatch, 1991, with Descriptions of Three New Species from Caves in Southern China (Diplopoda: Polydesmida: Polydesmidae). Tropical Natural History 10: 159 - 169" title="Review of the Millipede Genus Pacidesmus Golovatch, 1991, with Descriptions of Three New Species from Caves in Southern China (Diplopoda: Polydesmida: Polydesmidae)." volume="10" year="2010">Golovatch et al. 2010</bibRefCitation>
|
||
)
|
||
</td>
|
||
</tr>
|
||
<tr id="02E2E5B52136CC21BCD1AB75916F3DB6" pageId="0" pageNumber="259">
|
||
<td id="8E632A20EBC577E2A9FC1AF5D8B5CCEE" colspan="1" pageId="0" pageNumber="259" rowspan="1">
|
||
<taxonomicName id="29525E248B6AB15DE90B389F8BF8444F" authorityName="Golovatch & Geoffroy" authorityYear="2014" class="Diplopoda" family="Polydesmidae" genus="Pacidesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pacidesmus trifidus" order="Polydesmida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="species" species="trifidus">
|
||
<emphasis id="7618638616C152CC88E08302B34BB42F" italics="true" pageId="0" pageNumber="259">Pacidesmus trifidus</emphasis>
|
||
</taxonomicName>
|
||
Golovatch & Geoffroy, 2014
|
||
</td>
|
||
<td id="B20472FE3C2455C3820AB7A265E7A274" colspan="1" pageId="0" pageNumber="259" rowspan="1">
|
||
China, Guangxi Prov., Guilin County, Grotte des Squelettes (
|
||
<bibRefCitation id="C577B6CB9AFFA9BB9B5278FEF7E338E5" DOI="https://doi.org/10.15298/rusentj.23.2.01" author="Golovatch, SI" journalOrPublisher="Russian Entomological Journal" pageId="0" pageNumber="259" pagination="91 - 105" refId="B13" refString="Golovatch, SI, Geoffroy, JJ, 2014. On some new or poorly-known species of the millipede family Polydesmidae from southern China (Diplopoda: Polydesmida). Russian Entomological Journal 23 (1): 91 - 105, DOI: https://doi.org/10.15298/rusentj.23.2.01" title="On some new or poorly-known species of the millipede family Polydesmidae from southern China (Diplopoda: Polydesmida)." url="https://doi.org/10.15298/rusentj.23.2.01" volume="23" year="2014">Golovatch and Geoffroy 2014</bibRefCitation>
|
||
).
|
||
</td>
|
||
</tr>
|
||
<tr id="3697BB7610091289AC7FF42965797EAB" pageId="0" pageNumber="259">
|
||
<td id="7A5469E7146A98847E78BA653F4AF2E6" colspan="1" pageId="0" pageNumber="259" rowspan="1">
|
||
<taxonomicName id="E76C434427ABE628FA72222518EE4A88" authorityName="Liu & Golovatch" authorityYear="2020" class="Diplopoda" family="Polydesmidae" genus="Pacidesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pacidesmus trilobatus" order="Polydesmida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="species" species="trilobatus">
|
||
<emphasis id="FB46A16DD59CC192015DFA976C257631" italics="true" pageId="0" pageNumber="259">Pacidesmus trilobatus</emphasis>
|
||
</taxonomicName>
|
||
Liu & Golovatch, 2020
|
||
</td>
|
||
<td id="32638E6BA1F3E8AE4B960294045778E4" colspan="1" pageId="0" pageNumber="259" rowspan="1">
|
||
China, Yunnan Province, Maguan County, Pojiao Town, Dayan Dong Cave China, Yunnan Province, Wenshan County, Liujing Town, Laozhai Village, I Dong Cave (
|
||
<bibRefCitation id="436E5C94508CF02B764D0D39B1524C82" author="Liu, WX" journalOrPublisher="The Raffles Bulletin of Zoology" pageId="0" pageNumber="259" pagination="78 - 87" refId="B24" refString="Liu, WX, Golovatch, SI, 2020. Three new cavernicolous species and three new records of the millipede genus Pacidesmus from southern China (Diplopoda, Polydesmida, Polydesmidae). The Raffles Bulletin of Zoology 35: 78 - 87" title="Three new cavernicolous species and three new records of the millipede genus Pacidesmus from southern China (Diplopoda, Polydesmida, Polydesmidae)." volume="35" year="2020">Liu and Golovatch 2020</bibRefCitation>
|
||
).
|
||
</td>
|
||
</tr>
|
||
<tr id="54C84F8C36AB9C62F41F35729EC3D808" pageId="0" pageNumber="259">
|
||
<td id="ABFD9DF65AAB7EFC3FD8747FE5A95767" colspan="1" pageId="0" pageNumber="259" rowspan="1">
|
||
<taxonomicName id="9067D10DA3610583C300A60D3A1F760F" authorityName="Liu & Golovatch" authorityYear="2020" class="Diplopoda" family="Polydesmidae" genus="Pacidesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pacidesmus uncatus" order="Polydesmida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="species" species="uncatus">
|
||
<emphasis id="D5A674DF56387762EEBD2D90D205C9F9" italics="true" pageId="0" pageNumber="259">Pacidesmus uncatus</emphasis>
|
||
</taxonomicName>
|
||
Liu & Golovatch, 2020
|
||
</td>
|
||
<td id="F82522DC0195A035E741DC426CDDA7D0" colspan="1" pageId="0" pageNumber="259" rowspan="1">
|
||
China, Yunnan Province, Qujing City, Zhanyi County, Tianshengqiao Dong Cave (
|
||
<bibRefCitation id="9D363D6E68A1A7CEC0407BDA9E6E02AC" author="Liu, WX" journalOrPublisher="The Raffles Bulletin of Zoology" pageId="0" pageNumber="259" pagination="78 - 87" refId="B24" refString="Liu, WX, Golovatch, SI, 2020. Three new cavernicolous species and three new records of the millipede genus Pacidesmus from southern China (Diplopoda, Polydesmida, Polydesmidae). The Raffles Bulletin of Zoology 35: 78 - 87" title="Three new cavernicolous species and three new records of the millipede genus Pacidesmus from southern China (Diplopoda, Polydesmida, Polydesmidae)." volume="35" year="2020">Liu and Golovatch 2020</bibRefCitation>
|
||
)
|
||
</td>
|
||
</tr>
|
||
<tr id="CCD818C4C112C93C8B4AA97DDE5F1842" pageId="0" pageNumber="259">
|
||
<td id="B280FC3A68C240365D2092049DE98D3C" colspan="1" pageId="0" pageNumber="259" rowspan="1">
|
||
<taxonomicName id="2EE8FAF560894937B9C3B6F97796BF63" authorityName="Liu & Golovatch" authorityYear="2020" class="Diplopoda" family="Polydesmidae" genus="Pacidesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pacidesmus whitteni" order="Polydesmida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="species" species="whitteni">
|
||
<emphasis id="E1BCE678C21DC55AFE21129D50A105CA" italics="true" pageId="0" pageNumber="259">Pacidesmus whitteni</emphasis>
|
||
</taxonomicName>
|
||
Liu & Golovatch, 2020
|
||
</td>
|
||
<td id="D817EFA84AA21F15E008DD44AE1FEF52" colspan="1" pageId="0" pageNumber="259" rowspan="1">
|
||
China, Guangxi Zhuang Autonomous Region, Fengshan County, Jinya Town, Hangdong Village, I Dong Cave (
|
||
<bibRefCitation id="280222817131961C809BF39C4A580510" author="Liu, WX" journalOrPublisher="The Raffles Bulletin of Zoology" pageId="0" pageNumber="259" pagination="78 - 87" refId="B24" refString="Liu, WX, Golovatch, SI, 2020. Three new cavernicolous species and three new records of the millipede genus Pacidesmus from southern China (Diplopoda, Polydesmida, Polydesmidae). The Raffles Bulletin of Zoology 35: 78 - 87" title="Three new cavernicolous species and three new records of the millipede genus Pacidesmus from southern China (Diplopoda, Polydesmida, Polydesmidae)." volume="35" year="2020">Liu and Golovatch 2020</bibRefCitation>
|
||
)
|
||
</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="C41F1002A2D1C28298181DEAC24AE383" doi="10.3897/zookeys.1190.114958.figure6" httpUri="https://binary.pensoft.net/fig/976112" pageId="0" pageNumber="259" start="Figure 6" startId="F6">
|
||
<paragraph id="D0C16561895515E01754D8434D41FB7D" pageId="0" pageNumber="259">
|
||
<emphasis id="90B0A71BB984DF266D75FC0C9D1F37E6" bold="true" pageId="0" pageNumber="259">Figure 6.</emphasis>
|
||
<taxonomicName id="DD1A115A51F1F38D25B33C01C48384E3" authorityName="Nguyen" authorityYear="2009" class="Diplopoda" family="Polydesmidae" genus="Polydesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polydesmus vietnamicus" order="Polydesmida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="species" species="vietnamicus">
|
||
<emphasis id="78C024A5C59ADBBDDD9D018C45199FAF" italics="true" pageId="0" pageNumber="259">Polydesmus vietnamicus</emphasis>
|
||
</taxonomicName>
|
||
Nguyen, 2009 from Ha Giang Province (IEBR-Myr 808) ♂, segment 7 with gonopods under the normal (
|
||
<emphasis id="585676C6DDC50BE2DCC108DEFF0E22A2" bold="true" pageId="0" pageNumber="259">A, B</emphasis>
|
||
) and UV (
|
||
<emphasis id="75539660D1E8E993CF8A46CBBA838CCC" bold="true" pageId="0" pageNumber="259">C, D</emphasis>
|
||
) light
|
||
<emphasis id="A6781D3CE923F2D3318A3C9BDB7CAEF6" bold="true" pageId="0" pageNumber="259">A, C</emphasis>
|
||
posterior views
|
||
<emphasis id="98929B361D93783B7FAF970AE45B57CE" bold="true" pageId="0" pageNumber="259">B, D</emphasis>
|
||
anterior views. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="4731D5350FC2DD85718393B1B86D2A00" pageId="0" pageNumber="259">
|
||
The new species is assigned to
|
||
<taxonomicName id="91F948B8A72D9AE4417DF4000BA4DD57" authorityName="Golovatch" authorityYear="1991" class="Diplopoda" family="Polydesmidae" genus="Pacidesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pacidesmus" order="Polydesmida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E14375B78EE5E7A80E840E0C2A8FB566" italics="true" pageId="0" pageNumber="259">Pacidesmus</emphasis>
|
||
</taxonomicName>
|
||
because of the following characters: the seminal groove starts mesally, then recurves laterad at the base of a particularly prominent endomere branch to enter an accessory seminal chamber that opens on a setose pulvillus; endomere bears additional processes.
|
||
</paragraph>
|
||
<caption id="12749AB93FE3A355A1746669F9A1C773" doi="10.3897/zookeys.1190.114958.figure7" httpUri="https://binary.pensoft.net/fig/976113" pageId="0" pageNumber="259" start="Figure 7" startId="F7">
|
||
<paragraph id="5A409A47F9C18A7EF448FCE69BA708DC" pageId="0" pageNumber="259">
|
||
<emphasis id="D6CBC3BA9A6B39FF5580C33886D8D323" bold="true" pageId="0" pageNumber="259">Figure 7.</emphasis>
|
||
<taxonomicName id="4F2BAA716528AA2DF8BBC3D87F91648E" authorityName="Nguyen" authorityYear="2009" class="Diplopoda" family="Polydesmidae" genus="Polydesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polydesmus vietnamicus" order="Polydesmida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="species" species="vietnamicus">
|
||
<emphasis id="E498A6021AA4A11A31102AE13928B782" italics="true" pageId="0" pageNumber="259">Polydesmus vietnamicus</emphasis>
|
||
</taxonomicName>
|
||
Nguyen, 2009 from Ha Giang Province (IEBR-Myr 808) ♂, right gonopod under the UV light
|
||
<emphasis id="AE41FDCA20EE9C314C393E495EF7A337" bold="true" pageId="0" pageNumber="259">A</emphasis>
|
||
ventral view
|
||
<emphasis id="E99F75601AA0A923FD36451CAA9B3D01" bold="true" pageId="0" pageNumber="259">B</emphasis>
|
||
lateral view
|
||
<emphasis id="E362AA9C153CE7229515C3FD80BE7822" bold="true" pageId="0" pageNumber="259">C</emphasis>
|
||
dorsal view
|
||
<emphasis id="F231E26BD60331FFC7890ED69CA68BB0" bold="true" pageId="0" pageNumber="259">D</emphasis>
|
||
mesal view. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="C55EB4BD727FE48F2B6318374288B4C3" pageId="0" pageNumber="259" type="etymology">
|
||
<paragraph id="198A8C14272C5DCC731D58351701E1F7" pageId="0" pageNumber="259">Etymology.</paragraph>
|
||
<paragraph id="B251C7BA80F45D9A2FE4A0638211F918" pageId="0" pageNumber="259">The specific epithet is treated as a noun in apposition and is based on the "Tua Chua" district where the two caves are located.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="627318E393E63A27BEB360F16B049C8B" pageId="0" pageNumber="259" type="description">
|
||
<paragraph id="DA44A46C319A61F69175A81596FABD49" pageId="0" pageNumber="259">Description.</paragraph>
|
||
<paragraph id="6A604BFD12353A17A0455250B7FC555A" pageId="0" pageNumber="259">
|
||
Holotype length ca 16.3 mm, width of midbody pro- and metazonae 1.0 mm and 1.5 mm, respectively. In width, head <collum <segment 3 = 4 <2 <5 = 15, thereafter body gradually tapering towards telson (Figs
|
||
<figureCitation id="E6EE42B1AB0A2B307128773746E05E3B" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Pacidesmus tuachua sp. nov., holotype ♂ (IEBR-Myr 951) A anterior part of body, dorsal view B anterior part of body, lateral view C anterior part of body, ventral view D segments 8 - 10, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1190.114958.figure9" httpUri="https://binary.pensoft.net/fig/976115" pageId="0" pageNumber="259">9</figureCitation>
|
||
,
|
||
<figureCitation id="5DE6BAA4FE309B30CEF51BBF9659759A" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Pacidesmus tuachua sp. nov., holotype ♂ (IEBR-Myr 951) A segments 8 - 10, ventral view B-D posterior part of body, dorsal, vental and lateral views, respectively. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1190.114958.figure10" httpUri="https://binary.pensoft.net/fig/976116" pageId="0" pageNumber="259">10</figureCitation>
|
||
,
|
||
<figureCitation id="612709F2069EA4F3FA3C984F29F6AA4F" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pacidesmus tuachua sp. nov., holotype ♂ (IEBR-Myr 951) A anterior part of body, under the UV light, dorsal view B segments 8 - 10, dorsal view C posterior part of body, dorsal view D posterior part of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1190.114958.figure12" httpUri="https://binary.pensoft.net/fig/976118" pageId="0" pageNumber="259">12</figureCitation>
|
||
). Colouration in alcohol rather uniformly white (Figs
|
||
<figureCitation id="3B5B1F0D84A49A27D954EE013FF49A81" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Polydesmus vietnamicus Nguyen, 2009 from Ha Giang Province (IEBR-Myr 808) ♂, right gonopod A mesal view B lateral view. Abbreviations: co = coxite; pref = prefemorite; ca = cannula; en = endomere; sl = solenomere; mt = midway teeth; ap = additional process. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1190.114958.figure8" httpUri="https://binary.pensoft.net/fig/976114" pageId="0" pageNumber="259">8</figureCitation>
|
||
,
|
||
<figureCitation id="84ED7D82D6CDBF306EFB7BDDA9246358" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Pacidesmus tuachua sp. nov., holotype ♂ (IEBR-Myr 951) A anterior part of body, dorsal view B anterior part of body, lateral view C anterior part of body, ventral view D segments 8 - 10, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1190.114958.figure9" httpUri="https://binary.pensoft.net/fig/976115" pageId="0" pageNumber="259">9</figureCitation>
|
||
). Body with 20 segments. Antennae long and only slightly clavate, possibly reaching past segment 3 if stretching laterally; antennomere 3 longest, approximately 1.3
|
||
<normalizedToken id="5589A066B5947BEE6C8287B08EF7007D" originalValue="×">x</normalizedToken>
|
||
longer than subequal antennomeres 4-6; antennomeres 5 and 6 each with a small, compact, distodorsal group of bacilliform sensilla; antennomere 7 with a minute dorsoparabasal cone and a distodorsal group of microscopic sensilla (Figs
|
||
<figureCitation id="349045813B7C3BE2A91421F97E48E1B6" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Pacidesmus tuachua sp. nov., holotype ♂ (IEBR-Myr 951) A anterior part of body, dorsal view B anterior part of body, lateral view C anterior part of body, ventral view D segments 8 - 10, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1190.114958.figure9" httpUri="https://binary.pensoft.net/fig/976115" pageId="0" pageNumber="259">9A-C</figureCitation>
|
||
,
|
||
<figureCitation id="751CF9CC4FDCEF4C6B82E5C5FEA56AC9" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Pacidesmus tuachua sp. nov., holotype ♂ (IEBR-Myr 951) A telson, ventral view B telson, lateral view C right antenna, anterior view D gonopods in situ under UV light, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1190.114958.figure11" httpUri="https://binary.pensoft.net/fig/976117" pageId="0" pageNumber="259">11C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="FFC35F5C2BC485329F65D1D2E2308D0E" doi="10.3897/zookeys.1190.114958.figure8" httpUri="https://binary.pensoft.net/fig/976114" pageId="0" pageNumber="259" start="Figure 8" startId="F8">
|
||
<paragraph id="40BDD0DBE0ECAFF4952297F8E7B6727A" pageId="0" pageNumber="259">
|
||
<emphasis id="2F8253597A5D91926317DCB434D0D877" bold="true" pageId="0" pageNumber="259">Figure 8.</emphasis>
|
||
<taxonomicName id="CE1678B94C218BFAD91DA0BDB6A00557" authorityName="Nguyen" authorityYear="2009" class="Diplopoda" family="Polydesmidae" genus="Polydesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polydesmus vietnamicus" order="Polydesmida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="species" species="vietnamicus">
|
||
<emphasis id="641BC8B3A268B9677F3ABC4BC99E2123" italics="true" pageId="0" pageNumber="259">Polydesmus vietnamicus</emphasis>
|
||
</taxonomicName>
|
||
Nguyen, 2009 from Ha Giang Province (IEBR-Myr 808) ♂, right gonopod
|
||
<emphasis id="63FFE9791BED34B4269C18204DDA224C" bold="true" pageId="0" pageNumber="259">A</emphasis>
|
||
mesal view
|
||
<emphasis id="698078CBA808C2E47EA9D36DCBD6F876" bold="true" pageId="0" pageNumber="259">B</emphasis>
|
||
lateral view. Abbreviations:
|
||
<emphasis id="549FF7F88D82B79550BC4ED7E63E0D1B" italics="true" pageId="0" pageNumber="259">co</emphasis>
|
||
= coxite;
|
||
<emphasis id="553C1C5FB95D02089783D22966962FB3" italics="true" pageId="0" pageNumber="259">pref</emphasis>
|
||
= prefemorite;
|
||
<emphasis id="F089BFDD05794B19A647AC681AA341BE" italics="true" pageId="0" pageNumber="259">ca</emphasis>
|
||
= cannula;
|
||
<emphasis id="27E51228FE2A3033D186F16D287471CD" italics="true" pageId="0" pageNumber="259">en</emphasis>
|
||
= endomere;
|
||
<emphasis id="E5E90E32575DECDA86326D10659326A8" italics="true" pageId="0" pageNumber="259">sl</emphasis>
|
||
= solenomere;
|
||
<emphasis id="712889FDB19361BAC541EDE1C4D94E29" italics="true" pageId="0" pageNumber="259">mt</emphasis>
|
||
= midway teeth;
|
||
<emphasis id="5265E86C47569B5B4B698EAA2A8BEA89" italics="true" pageId="0" pageNumber="259">ap</emphasis>
|
||
= additional process. Scale bars: 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="F098BA1ACC3A7EC1E198C1DC823CBD50" doi="10.3897/zookeys.1190.114958.figure9" httpUri="https://binary.pensoft.net/fig/976115" pageId="0" pageNumber="259" start="Figure 9" startId="F9">
|
||
<paragraph id="52400DEDDBE655E9072D932D7AC18830" pageId="0" pageNumber="259">
|
||
<emphasis id="FE28D1AC73AD5F98484FE1896E23F3CF" bold="true" pageId="0" pageNumber="259">Figure 9.</emphasis>
|
||
<taxonomicName id="D3DEFB5B1F80D9330F33EDDA6A5EA09F" authorityName="Nguyen & Vu & Eguchi" authorityYear="2024" class="Diplopoda" family="Polydesmidae" genus="Pacidesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pacidesmus tuachua" order="Polydesmida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="species" species="tuachua">
|
||
<emphasis id="CF2775552DD9F28459D72744E359855C" italics="true" pageId="0" pageNumber="259">Pacidesmus tuachua</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype ♂ (IEBR-Myr 951)
|
||
<emphasis id="0250A047FF32C256B33AE6E7F700AB93" bold="true" pageId="0" pageNumber="259">A</emphasis>
|
||
anterior part of body, dorsal view
|
||
<emphasis id="F1A382F1B6399C143CAEFE1EE57A388E" bold="true" pageId="0" pageNumber="259">B</emphasis>
|
||
anterior part of body, lateral view
|
||
<emphasis id="A35904D064051E597E1FC6455F41744B" bold="true" pageId="0" pageNumber="259">C</emphasis>
|
||
anterior part of body, ventral view
|
||
<emphasis id="42799E992DB198AC06566C74481BA3A8" bold="true" pageId="0" pageNumber="259">D</emphasis>
|
||
segments 8-10, dorsal view. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="CA464E85CB5BF40F489597D4EF069D2F" doi="10.3897/zookeys.1190.114958.figure10" httpUri="https://binary.pensoft.net/fig/976116" pageId="0" pageNumber="259" start="Figure 10" startId="F10">
|
||
<paragraph id="DF507149E24214ACB7284115A58CA739" pageId="0" pageNumber="259">
|
||
<emphasis id="5B0C00A410A771DFA2B6F2C1567351FE" bold="true" pageId="0" pageNumber="259">Figure 10.</emphasis>
|
||
<taxonomicName id="DC86DA96966A7BE8DC4B6B96E016C01D" authorityName="Nguyen & Vu & Eguchi" authorityYear="2024" class="Diplopoda" family="Polydesmidae" genus="Pacidesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pacidesmus tuachua" order="Polydesmida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="species" species="tuachua">
|
||
<emphasis id="878EFCC99235E9D1E814AE9FA0756752" italics="true" pageId="0" pageNumber="259">Pacidesmus tuachua</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype ♂ (IEBR-Myr 951)
|
||
<emphasis id="0BC28120F9F64C524372A7FF518F0DAF" bold="true" pageId="0" pageNumber="259">A</emphasis>
|
||
segments 8-10, ventral view
|
||
<emphasis id="C745C9727C5E8060828262F3CCB8D5D0" bold="true" pageId="0" pageNumber="259">B-D</emphasis>
|
||
posterior part of body, dorsal, vental and lateral views, respectively. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E8D0E34885AC0DCD5A348C6E7D12D6C6" pageId="0" pageNumber="259">
|
||
Paraterga (Figs
|
||
<figureCitation id="F684833C2D1BD69BF16027B3E2E28F6E" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Pacidesmus tuachua sp. nov., holotype ♂ (IEBR-Myr 951) A anterior part of body, dorsal view B anterior part of body, lateral view C anterior part of body, ventral view D segments 8 - 10, dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1190.114958.figure9" httpUri="https://binary.pensoft.net/fig/976115" pageId="0" pageNumber="259">9</figureCitation>
|
||
,
|
||
<figureCitation id="B998890FC9DA92E645976111A1E37F81" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Pacidesmus tuachua sp. nov., holotype ♂ (IEBR-Myr 951) A segments 8 - 10, ventral view B-D posterior part of body, dorsal, vental and lateral views, respectively. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1190.114958.figure10" httpUri="https://binary.pensoft.net/fig/976116" pageId="0" pageNumber="259">10</figureCitation>
|
||
,
|
||
<figureCitation id="C0D3AC0C6FF80ABF7E01B3F32B7DC6BA" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pacidesmus tuachua sp. nov., holotype ♂ (IEBR-Myr 951) A anterior part of body, under the UV light, dorsal view B segments 8 - 10, dorsal view C posterior part of body, dorsal view D posterior part of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1190.114958.figure12" httpUri="https://binary.pensoft.net/fig/976118" pageId="0" pageNumber="259">12A-C</figureCitation>
|
||
) strongly developed, set high, starting with collum, dorsum faintly convex; paraterga mostly weakly upturned above dorsum. Caudolateral corner of paraterga acute, postcollum ones extending increasingly past rear tergal margin, especially so in segments 16-18. All poreless segments with three incisions, all pore-bearing ones with four minute incisions at lateral margin. Front margins of metaterga narrowly bordered and forming distinct shoulders.
|
||
</paragraph>
|
||
<paragraph id="2DAD55C08D4C3E0BC2B1D69BBCF2BC73" pageId="0" pageNumber="259">Ozopores evident, dorsal, located in front of posteriormost marginal incision of paraterga 5, 7, 9, 10, 12-13, 15-19.</paragraph>
|
||
<paragraph id="6456CB12AD54A90932E53353D7B1AEFB" pageId="0" pageNumber="259">
|
||
Metatergal sculpture typical, poorly developed, obliterate, with three transverse rows of typical (= polydesmid), setigerous, polygonal bosses. Tergal setae short, slightly longer only on collum, simple, often obliterate. Stricture between pro- and metazona wide, shallow and nearly smooth. Limbus exceedingly thin, microdenticulate (Fig.
|
||
<figureCitation id="3D32ED197B713F78BEC233F32E3BD92C" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pacidesmus tuachua sp. nov., holotype ♂ (IEBR-Myr 951) A anterior part of body, under the UV light, dorsal view B segments 8 - 10, dorsal view C posterior part of body, dorsal view D posterior part of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1190.114958.figure12" httpUri="https://binary.pensoft.net/fig/976118" pageId="0" pageNumber="259">12A-C</figureCitation>
|
||
). Pleurosternal carinae absent.
|
||
</paragraph>
|
||
<paragraph id="E46EEC50F3084B7BDB768A101C52241C" pageId="0" pageNumber="259">
|
||
Epiproct (Figs
|
||
<figureCitation id="AADC41FA4D5B18A942E7EB42256479D5" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Pacidesmus tuachua sp. nov., holotype ♂ (IEBR-Myr 951) A telson, ventral view B telson, lateral view C right antenna, anterior view D gonopods in situ under UV light, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1190.114958.figure11" httpUri="https://binary.pensoft.net/fig/976117" pageId="0" pageNumber="259">11A, B</figureCitation>
|
||
,
|
||
<figureCitation id="A92724D4A52E36F432456E38A05D4583" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pacidesmus tuachua sp. nov., holotype ♂ (IEBR-Myr 951) A anterior part of body, under the UV light, dorsal view B segments 8 - 10, dorsal view C posterior part of body, dorsal view D posterior part of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1190.114958.figure12" httpUri="https://binary.pensoft.net/fig/976118" pageId="0" pageNumber="259">12C, D</figureCitation>
|
||
) short, conical, pre-apical lateral papillae evident. Hypoproct (Figs
|
||
<figureCitation id="3D10776E78892C4DE52AD29AAC5D3C80" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Pacidesmus tuachua sp. nov., holotype ♂ (IEBR-Myr 951) A telson, ventral view B telson, lateral view C right antenna, anterior view D gonopods in situ under UV light, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1190.114958.figure11" httpUri="https://binary.pensoft.net/fig/976117" pageId="0" pageNumber="259">11A</figureCitation>
|
||
,
|
||
<figureCitation id="164BDFC729279F345D2B82F8C5FAB31D" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pacidesmus tuachua sp. nov., holotype ♂ (IEBR-Myr 951) A anterior part of body, under the UV light, dorsal view B segments 8 - 10, dorsal view C posterior part of body, dorsal view D posterior part of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1190.114958.figure12" httpUri="https://binary.pensoft.net/fig/976118" pageId="0" pageNumber="259">12D</figureCitation>
|
||
) subtriangular; distolateral setiferous knobs small, but distinct and well separated.
|
||
</paragraph>
|
||
<caption id="4B19D158A3E836CFF852230176FF6197" doi="10.3897/zookeys.1190.114958.figure11" httpUri="https://binary.pensoft.net/fig/976117" pageId="0" pageNumber="259" start="Figure 11" startId="F11">
|
||
<paragraph id="BF2651045C6D5F9CDFF20D2BD9038F2C" pageId="0" pageNumber="259">
|
||
<emphasis id="1BD5AF8227BE123230E4F210EEC74F81" bold="true" pageId="0" pageNumber="259">Figure 11.</emphasis>
|
||
<taxonomicName id="21BAD6E1EE8BF152601A7DAF79580934" authorityName="Nguyen & Vu & Eguchi" authorityYear="2024" class="Diplopoda" family="Polydesmidae" genus="Pacidesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pacidesmus tuachua" order="Polydesmida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="species" species="tuachua">
|
||
<emphasis id="7B9AAC6E248BF0243123C6FE5DFF3AD5" italics="true" pageId="0" pageNumber="259">Pacidesmus tuachua</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype ♂ (IEBR-Myr 951)
|
||
<emphasis id="1B88A28F03D314896043E69C11A51F56" bold="true" pageId="0" pageNumber="259">A</emphasis>
|
||
telson, ventral view
|
||
<emphasis id="E168CD9E1B48CA60BC33CF717C09B36C" bold="true" pageId="0" pageNumber="259">B</emphasis>
|
||
telson, lateral view
|
||
<emphasis id="7A322C283232BCE22C4750A6DB9B7B4A" bold="true" pageId="0" pageNumber="259">C</emphasis>
|
||
right antenna, anterior view
|
||
<emphasis id="F6DA11B6C96434F7383C1D4B77096563" bold="true" pageId="0" pageNumber="259">D</emphasis>
|
||
gonopods in situ under UV light, ventral view. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="2CE8657394D32428EA071E19010B3763" doi="10.3897/zookeys.1190.114958.figure12" httpUri="https://binary.pensoft.net/fig/976118" pageId="0" pageNumber="259" start="Figure 12" startId="F12">
|
||
<paragraph id="293E8B9F60E1AA0C766EFCEA4FDA49B5" pageId="0" pageNumber="259">
|
||
<emphasis id="686F245253762A88C83E9F8D402F8E57" bold="true" pageId="0" pageNumber="259">Figure 12.</emphasis>
|
||
<taxonomicName id="76E76E1490A5EE2EE3AD9715F42CDD16" authorityName="Nguyen & Vu & Eguchi" authorityYear="2024" class="Diplopoda" family="Polydesmidae" genus="Pacidesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pacidesmus tuachua" order="Polydesmida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="species" species="tuachua">
|
||
<emphasis id="B2DACB89983709894642AD8EA0983474" italics="true" pageId="0" pageNumber="259">Pacidesmus tuachua</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype ♂ (IEBR-Myr 951)
|
||
<emphasis id="0FCA3CB5420F1DE2852EA10E24420423" bold="true" pageId="0" pageNumber="259">A</emphasis>
|
||
anterior part of body, under the UV light, dorsal view
|
||
<emphasis id="91B2464033F513B0964AE0DED2B438B4" bold="true" pageId="0" pageNumber="259">B</emphasis>
|
||
segments 8-10, dorsal view
|
||
<emphasis id="F252BB9A9D87F281B0005F28F191FFC7" bold="true" pageId="0" pageNumber="259">C</emphasis>
|
||
posterior part of body, dorsal view
|
||
<emphasis id="A1D9BF62D52DDE29DB74D4799AAE08FB" bold="true" pageId="0" pageNumber="259">D</emphasis>
|
||
posterior part of body, ventral view. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="32522C128D2096E203A0539C7E6D2456" pageId="0" pageNumber="259">
|
||
Sterna without modifications, but setose (Fig.
|
||
<figureCitation id="06D0342E995A94FE4DF896122510C6ED" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Pacidesmus tuachua sp. nov., holotype ♂ (IEBR-Myr 951) A segments 8 - 10, ventral view B-D posterior part of body, dorsal, vental and lateral views, respectively. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1190.114958.figure10" httpUri="https://binary.pensoft.net/fig/976116" pageId="0" pageNumber="259">10A, C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="7182200633B8B3DE1FDBB17F8651FD34" pageId="0" pageNumber="259">
|
||
Legs generally long and slender, apparently slightly incrassate, approximately 1.7-1.8
|
||
<normalizedToken id="7EC35D2B28DB14B55D25EEF10F4C0555" originalValue="×">x</normalizedToken>
|
||
as long as midbody height, densely setose, almost all setae simple, poorly branching setae with minute, distal, side branchlets only on slender prefemora, latter devoid of lateral bulges.
|
||
</paragraph>
|
||
<paragraph id="20EBF1124D6682D7332025B62550624E" pageId="0" pageNumber="259">
|
||
Gonopods (Figs
|
||
<figureCitation id="C2CDDC5D8CD6BDEE465FB077B35F92E5" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Pacidesmus tuachua sp. nov., holotype ♂ (IEBR-Myr 951) A telson, ventral view B telson, lateral view C right antenna, anterior view D gonopods in situ under UV light, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1190.114958.figure11" httpUri="https://binary.pensoft.net/fig/976117" pageId="0" pageNumber="259">11D</figureCitation>
|
||
,
|
||
<figureCitation id="517A9E786FE3039827F13904584F194A" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Pacidesmus tuachua sp. nov., holotype ♂ (IEBR-Myr 951) A anterior part of body, under the UV light, dorsal view B segments 8 - 10, dorsal view C posterior part of body, dorsal view D posterior part of body, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1190.114958.figure12" httpUri="https://binary.pensoft.net/fig/976118" pageId="0" pageNumber="259">12</figureCitation>
|
||
,
|
||
<figureCitation id="9693A4EFA9EF97E414A4730C233DFFEF" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Pacidesmus tuachua sp. nov., holotype ♂ (IEBR-Myr 951) right gonopod, under UV light A mesal view B ventral view C lateral view D dorsal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1190.114958.figure13" httpUri="https://binary.pensoft.net/fig/976119" pageId="0" pageNumber="259">13</figureCitation>
|
||
) characteristically subfalcate (vs suberect in all other congeners). with large, rectangular coxites (
|
||
<emphasis id="0C7CEA433E08E8B8A5A88DAFD30A0E8C" bold="true" pageId="0" pageNumber="259">co</emphasis>
|
||
), with a few long setae ventrally; a long, simple, flagelliform cannula (
|
||
<emphasis id="6E022ACC056B9215D33E608819F13563" bold="true" pageId="0" pageNumber="259">ca</emphasis>
|
||
) as usual. Telopodite elongate, stout, strongly falcate or C-shaped; prefemorite (
|
||
<emphasis id="F1EFCA224AFCD47EE41C0E05BBE6D18A" bold="true" pageId="0" pageNumber="259">pref</emphasis>
|
||
) densely setose; seminal groove starting mesally, then recurving laterad to run to the opening on a hairy pulvillus. Endomere (
|
||
<emphasis id="669F59D6E472CAF97B95FF736BC74426" bold="true" pageId="0" pageNumber="259">en</emphasis>
|
||
) with two additional processes, a shorter, larger, broader triangular process at midlength (
|
||
<emphasis id="1FC433B1BA6531435B057494176FCE02" bold="true" pageId="0" pageNumber="259">p1</emphasis>
|
||
), and a longer, acuter, triangular process at
|
||
<normalizedToken id="EB46162452B5717274B10CF784878453" originalValue="¾">3/4</normalizedToken>
|
||
length (
|
||
<emphasis id="EEEB02AF0077363DD4FB9D9B673BBFE8" bold="true" pageId="0" pageNumber="259">p2</emphasis>
|
||
). Tip of endomere unequally bifid, a longer and a shorter branch. Neither an exomere nor a clivus.
|
||
</paragraph>
|
||
<caption id="35321DFDF18ECF4C4A2852B847DE10D0" doi="10.3897/zookeys.1190.114958.figure13" httpUri="https://binary.pensoft.net/fig/976119" pageId="0" pageNumber="259" start="Figure 13" startId="F13">
|
||
<paragraph id="2D7B67757737A99863295BE191A3D7AC" pageId="0" pageNumber="259">
|
||
<emphasis id="82622BA8FFC8466CB86BE72D6CC14723" bold="true" pageId="0" pageNumber="259">Figure 13.</emphasis>
|
||
<taxonomicName id="F77593DAA0279820DDBB7157FE9A5CFA" authorityName="Nguyen & Vu & Eguchi" authorityYear="2024" class="Diplopoda" family="Polydesmidae" genus="Pacidesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pacidesmus tuachua" order="Polydesmida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="species" species="tuachua">
|
||
<emphasis id="2177A6021282BC4B58202A9A7B4CC8DC" italics="true" pageId="0" pageNumber="259">Pacidesmus tuachua</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype ♂ (IEBR-Myr 951) right gonopod, under UV light
|
||
<emphasis id="9E494B0ED198CD51A0EAF0C56EDB89C0" bold="true" pageId="0" pageNumber="259">A</emphasis>
|
||
mesal view
|
||
<emphasis id="8343D0B0C72CEC0D5483EF975D1C2C11" bold="true" pageId="0" pageNumber="259">B</emphasis>
|
||
ventral view
|
||
<emphasis id="18E6C781D45F9869CE90252460FF2006" bold="true" pageId="0" pageNumber="259">C</emphasis>
|
||
lateral view
|
||
<emphasis id="EC4C0B5FB36F6359412B6E5F5E552528" bold="true" pageId="0" pageNumber="259">D</emphasis>
|
||
dorsal view. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5F7D8EFDFCD8AEB31A28C1ED61F4CCF8" pageId="0" pageNumber="259">
|
||
<emphasis id="FF5C6F2C0B48E87FB964D9B46FA8C5ED" bold="true" pageId="0" pageNumber="259">Female.</emphasis>
|
||
Slightly larger than male, length ca 16.8 mm, width of pro- and metazona about 1.1 mm and 1.7 mm, respectively. Paraterga slightly less strongly developed. Legs unmodified, somewhat shorter and more slender. Vulvae highly elevated. Epigynal ridge low.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4BAF9F1C8C775685D9FC07BF1013B40D" pageId="0" pageNumber="259" type="dna barcode">
|
||
<paragraph id="0784F3BA73EBB6B4A6D4044252913C84" pageId="0" pageNumber="259">DNA barcode.</paragraph>
|
||
<paragraph id="EE6A3D7BD50724535D1BBFB18BB18898" pageId="0" pageNumber="259">
|
||
Two COI sequences (661 bp) were uploaded to the GenBank with the accession numbers PP118040 and PP118041. The new species has a close COI identity to
|
||
<taxonomicName id="33D499A7AD153CDB27F6E69076284914" authorityName="Verhoeff" authorityYear="1937" class="Diplopoda" family="Polydesmidae" genus="Epanerchodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epanerchodus koreanus" order="Polydesmida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="species" species="koreanus">
|
||
<emphasis id="F947F12CC1EF05235197E927F4DBCBEF" italics="true" pageId="0" pageNumber="259">Epanerchodus koreanus</emphasis>
|
||
</taxonomicName>
|
||
(NC051495) at 88.72% (query coverage 97%).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EAA1DF1660DB28A2D0CDA7421C6436ED" pageId="0" pageNumber="259" type="habitat">
|
||
<paragraph id="0DFC57BB7DB9EF8AA7E87463DB866A59" pageId="0" pageNumber="259">Habitat.</paragraph>
|
||
<paragraph id="FFEAC0423C088B65F6DC05140AB38B90" pageId="0" pageNumber="259">
|
||
This species is to be considered a true troglobiont because it shows the typical morphological features of a cave-dweller. It was collected exclusively in the dark zone of the caves as described below. Kho Chua La and Xa Nhe caves are both located close together, approximately 500 m in distance. These caves are at the centre of the Xa Nhe commune, Dien Bien Province, northwestern Vietnam. The two caves are tunnel-like: they are high (15-20 m), wide (15-20 m), and long (1,000-1,500 m). The floor is mainly wet, with clay, and some small pools. Several other millipede species have been found in these caves, including
|
||
<taxonomicName id="6F1F92B2B8DC600CAD4CCBAE5CF4DB38" authorityName="Gervais" authorityYear="1847" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Glyphiulus" order="Spirostreptida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="386FF1DD81FDAC4B6BBE44D6F7DC7773" italics="true" pageId="0" pageNumber="259">Glyphiulus</emphasis>
|
||
</taxonomicName>
|
||
sp. (
|
||
<taxonomicName id="4A3F2DF2763D89F3CF138E3317BAD625" authorityName="Gervais" authorityYear="1847" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Spirostreptida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="order">Spirostreptida</taxonomicName>
|
||
,
|
||
<taxonomicName id="A38B770101AC933E2D175FE42B34B9A7" authorityName="Chamberlin" authorityYear="1922" class="Diplopoda" family="Glyphiulidae" kingdom="Animalia" lsidName="" order="Spirostreptida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="family">Glyphiulidae</taxonomicName>
|
||
) and
|
||
<taxonomicName id="795AF33834575B688A84B69B8C67FD6C" authorityName="Silvestri" authorityYear="1910" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eutrichodesmus" order="Polydesmida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="71B8DDBD8E0E0B5385AF50FB40315ADC" italics="true" pageId="0" pageNumber="259">Eutrichodesmus</emphasis>
|
||
</taxonomicName>
|
||
sp. (
|
||
<taxonomicName id="AE9A41493E0F8F86C3F3FD39D61B93CB" authorityName="Silvestri" authorityYear="1910" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Polydesmida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="order">Polydesmida</taxonomicName>
|
||
,:
|
||
<taxonomicName id="9EDB6ACD488319458895E44970F2F011" authorityName="Cook" authorityYear="1895" class="Diplopoda" family="Haplodesmidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Polydesmida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="family">Haplodesmidae</taxonomicName>
|
||
). The new species was found>1000 m from the entrance.
|
||
</paragraph>
|
||
<paragraph id="502D0D6259F4A552F94BBE9D166A276F" pageId="0" pageNumber="259">
|
||
Kho Chua La and Xa Nhe caves are located on the Tua Chua karst plateau in northeastern Dien Bien Province, northwestern Vietnam. The natural area is about 68,414 ha, and 70% of this area is composed of limestone mountains, which are known for their layers of majestic rugged rock and unique natural landscape. The karst region contains many stunning and well-known caves, such as Kho Chua La, Tham Khem, Hau Chua, Xa Nhe, and Pe Rang Ki (
|
||
<bibRefCitation id="B7258A6A2718DA6847ACC6790A563EBD" DOI="https://doi.org/10.30892/gtg.44414-946" author="Nguyen, NTP" journalOrPublisher="Geo Journal of Tourism and Geosites" pageId="0" pageNumber="259" pagination="1300 - 1305" refId="B31" refString="Nguyen, NTP, Tran, BD, Do, DL, 2022. Assessing the potential of community-based ecotourism toward sustainable development: A case study in Tua Chua karst plateau - Dien Bien - Vietnam. Geo Journal of Tourism and Geosites 44 (4): 1300 - 1305, DOI: https://doi.org/10.30892/gtg.44414-946" title="Assessing the potential of community-based ecotourism toward sustainable development: A case study in Tua Chua karst plateau - Dien Bien - Vietnam." url="https://doi.org/10.30892/gtg.44414-946" volume="44" year="2022">Nguyen et al. 2022</bibRefCitation>
|
||
). Furthermore, the Tua Chua karst plateau of northwestern Vietnam is close to the Yunnan and Sichuan karst regions of southwestern China. Given this, it is not surprising to discover the genus
|
||
<taxonomicName id="57F4817B8A166CEBB8195A28F5CC67DE" authorityName="Golovatch" authorityYear="1991" class="Diplopoda" family="Polydesmidae" genus="Pacidesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pacidesmus" order="Polydesmida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="FA4762F4AEDDA23ABEE15EA12789B6F4" italics="true" pageId="0" pageNumber="259">Pacidesmus</emphasis>
|
||
</taxonomicName>
|
||
in northwestern Vietnam. The distance between
|
||
<taxonomicName id="F4C0A5A1F7B070361C90E42694072BFD" authorityName="Nguyen & Vu & Eguchi" authorityYear="2024" class="Diplopoda" family="Polydesmidae" genus="Pacidesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pacidesmus tuachua" order="Polydesmida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="species" species="tuachua">
|
||
<emphasis id="5C1B71DC0507F1C2C95338962D26F272" italics="true" pageId="0" pageNumber="259">Pacidesmus tuachua</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. and
|
||
<taxonomicName id="38B35C18DD7EA0F5824DF0FA52E57F13" authorityName="Liu & Golovatch" authorityYear="2020" class="Diplopoda" family="Polydesmidae" genus="Pacidesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pacidesmus trilobatus" order="Polydesmida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="species" species="trilobatus">
|
||
<emphasis id="123D383E39A1100A182E721D038F3377" italics="true" pageId="0" pageNumber="259">Pacidesmus trilobatus</emphasis>
|
||
</taxonomicName>
|
||
Liu & Golovatch, 2020 from Guangxi Province, China, is about 150 km northeast-southwest (Fig.
|
||
<figureCitation id="C1D2781E749A4A66C1B21B1306D4C004" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Records of polydesmid species in Vietnam and of all known Pacidesmus species." figureDoi="10.3897/zookeys.1190.114958.figure1" httpUri="https://binary.pensoft.net/fig/976107" pageId="0" pageNumber="259">1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="A731B33A1F86061FEB0CCD83478D8A01" doi="10.3897/zookeys.1190.114958.figure14" httpUri="https://binary.pensoft.net/fig/976120" pageId="0" pageNumber="259" start="Figure 14" startId="F14">
|
||
<paragraph id="30995383D6931D2C2B29731191E565E3" pageId="0" pageNumber="259">
|
||
<emphasis id="EA5A0E41A5F9CF7A1ED494D60C67BAC4" bold="true" pageId="0" pageNumber="259">Figure 14.</emphasis>
|
||
<taxonomicName id="0EB9968ED7949CB02E7F237029CAF8AA" authorityName="Nguyen & Vu & Eguchi" authorityYear="2024" class="Diplopoda" family="Polydesmidae" genus="Pacidesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pacidesmus tuachua" order="Polydesmida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="species" species="tuachua">
|
||
<emphasis id="0D25082B67C5445ADDCB7DC6A7AF3E7D" italics="true" pageId="0" pageNumber="259">Pacidesmus tuachua</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype ♂ (IEBR-Myr 951) right gonopod
|
||
<emphasis id="5307D891E2667DB4777F9FC48CCB3FF2" bold="true" pageId="0" pageNumber="259">A</emphasis>
|
||
mesal view
|
||
<emphasis id="DDE9DC04C71107A69AD204657908BC05" bold="true" pageId="0" pageNumber="259">B</emphasis>
|
||
lateral view. Abbreviations:
|
||
<emphasis id="AF9B9F2B72525EB36F3AE5E8C7047E8D" italics="true" pageId="0" pageNumber="259">co</emphasis>
|
||
= coxite;
|
||
<emphasis id="C5BA3FA9252279DF86EAB89315A6E519" italics="true" pageId="0" pageNumber="259">pref</emphasis>
|
||
= prefemorite;
|
||
<emphasis id="1A558557FD80DD76EE915FDCF22D4FB2" italics="true" pageId="0" pageNumber="259">ca</emphasis>
|
||
= cannula;
|
||
<emphasis id="1B524A61221DB4A6A9B284C17F7557B1" italics="true" pageId="0" pageNumber="259">en</emphasis>
|
||
= endomere;
|
||
<emphasis id="ABD04D45644578E8D237494A25D20DB4" italics="true" pageId="0" pageNumber="259">pu</emphasis>
|
||
= puvillus;
|
||
<emphasis id="9C89229BA1D141A5D65252B7207B55C0" italics="true" pageId="0" pageNumber="259">p1</emphasis>
|
||
= first process;
|
||
<emphasis id="66761DB3D92DF9BDD21F1911A9AD8FCF" italics="true" pageId="0" pageNumber="259">p2</emphasis>
|
||
= second process. Scale bars: 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="5930107751D603E98C9EC813BAFB7A5E" pageId="0" pageNumber="259" type="remarks">
|
||
<paragraph id="5CDFB40E73CF4A0531ECC2ABD380A3C4" pageId="0" pageNumber="259">Remarks.</paragraph>
|
||
<paragraph id="DDA76538E8E84ED27EDB0790F9E4CE56" pageId="0" pageNumber="259">
|
||
While there remains a noticeable geographical gap between the mountainous northern Thailand species and the troglobionts of southern China, the discovery of a new species in northern Vietnam partially fills this gap. Like the more eastern species, the new species is also troglobiotic, and whether these species should be classified in a distinct genus related to a more restricted
|
||
<taxonomicName id="6A9552FD3202D77352E493E1454D6A74" authorityName="Golovatch" authorityYear="1991" class="Diplopoda" family="Polydesmidae" genus="Pacidesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pacidesmus" order="Polydesmida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="74DEA0676AEC9CB3B4494A983C6462A9" italics="true" pageId="0" pageNumber="259">Pacidesmus</emphasis>
|
||
</taxonomicName>
|
||
(including only the type species) needs exploration.
|
||
</paragraph>
|
||
<paragraph id="CAC771D85C43056E3FBFE124E3EA7C7D" pageId="0" pageNumber="259">
|
||
The discovery of a new species marks the first record of the genus
|
||
<taxonomicName id="982BEF314897683C688561E2B294C3E0" authorityName="Golovatch" authorityYear="1991" class="Diplopoda" family="Polydesmidae" genus="Pacidesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pacidesmus" order="Polydesmida" pageId="0" pageNumber="259" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6391AD1BDCF5066F76D0C4D1AD1FA17B" italics="true" pageId="0" pageNumber="259">Pacidesmus</emphasis>
|
||
</taxonomicName>
|
||
in Vietnam.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |