treatments-xml/data/26/75/87/267587D7FF9482277A617F6ED98BFB33.xml
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<document id="2D0BFF68031E17A80F3313B365815209" ID-DOI="10.11646/zootaxa.1263.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5065414" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625285053292" checkinUser="felipe" docAuthor="Hodgson, Chris &amp; Foldi, Imre" docDate="2006" docId="267587D7FF9482277A617F6ED98BFB33" docLanguage="en" docName="zt01263p120.pdf" docOrigin="Zootaxa 1263 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1263.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Porphyrophora hamelii Brandt. Letters 1833" docType="treatment" docVersion="3" lastPageNumber="250" masterDocId="DA4CFFAFFFC3827A7B697937DB14FF95" masterDocTitle="A review of the Margarodidae sensu Morrison (Hemiptera: Coccoidea) and some related taxa based on the morphology of adult males" masterLastPageNumber="250" masterPageNumber="1" pageNumber="1" updateTime="1698986938651" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="EA67D18C458D3101DCF0399954FF3384">A review of the Margarodidae sensu Morrison (Hemiptera: Coccoidea) and some related taxa based on the morphology of adult males</mods:title>
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<mods:namePart id="02168F60EDC23DEC69FB7FA957E664D4">Hodgson, Chris</mods:namePart>
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<mods:namePart id="3148C013AD27F2A3C3DA222B20904F18">Foldi, Imre</mods:namePart>
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<mods:title id="859A388D18B0AE8D0CB8046544670F0B">Zootaxa</mods:title>
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<mods:date id="5393986C3F7DB8A3A607E718E4E8FFB4">2006</mods:date>
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<treatment id="267587D7FF9482277A617F6ED98BFB33" LSID="urn:lsid:plazi:treatment:267587D7FF9482277A617F6ED98BFB33" httpUri="http://treatment.plazi.org/id/267587D7FF9482277A617F6ED98BFB33" lastPageId="93" pageId="87">
<subSubSection id="E6C6654AFF94822D7A617F6ED991F9E6" box="[264,645,1625,1651]" pageId="87" type="nomenclature">
<paragraph id="AE6336C1FF94822D7A617F6ED991F9E6" blockId="87.[264,645,1625,1686]" box="[264,645,1625,1651]" pageId="87">
<heading id="F52B81ADFF94822D7A617F6ED991F9E6" bold="true" box="[264,645,1625,1651]" fontSize="11" level="1" pageId="87" reason="1">
<taxonomicName id="69DC4D42FF94822D7A617F6ED991F9E6" ID-CoL="4LZ9M" authority="Brandt" authorityName="Brandt. Letters" authorityYear="1833" box="[264,645,1625,1651]" class="Insecta" family="Margarodidae" genus="Porphyrophora" kingdom="Animalia" order="Hemiptera" pageId="87" phylum="Arthropoda" rank="species" species="hamelii">
<emphasis id="9CA8EAD3FF94822D7A617F6ED991F9E6" bold="true" box="[264,645,1625,1651]" pageId="87">
<emphasis id="9CA8EAD3FF94822D7A617F6ED931F9E6" bold="true" box="[264,549,1625,1651]" italics="true" pageId="87">Porphyrophora hamelii</emphasis>
Brandt
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E6C6654AFF9482277A617F4BD98BFB33" lastPageId="93" pageId="87" type="description">
<paragraph id="AE6336C1FF94822D7A617F4BDA7AF903" blockId="87.[264,645,1625,1686]" box="[264,366,1660,1686]" pageId="87">
(
<figureCitation id="36E72A44FF94822D7A787F4BDA71F903" box="[273,357,1660,1686]" captionStart="FIGURE 13" captionStartId="89.[264,367,1817,1841]" captionTargetBox="[295,1269,341,1800]" captionTargetId="figure-42@89.[264,1323,325,1817]" captionTargetPageId="89" captionText="FIGURE 13. Male Porphyrophora hamelii Brandt. Letters as for Fig. 6 except K = peg­like seta, and U = segment VI of antenna." figureDoi="http://doi.org/10.5281/zenodo.5065450" httpUri="https://zenodo.org/record/5065450/files/figure.png" pageId="87">Fig. 13</figureCitation>
)
</paragraph>
<paragraph id="AE6336C1FF94822D7A617FFBD8C7F976" blockId="87.[264,979,1740,1763]" box="[264,979,1740,1763]" pageId="87">
<taxonomicName id="69DC4D42FF94822D7A617FFBD941F976" authority="Brandt" authorityName="Brandt. Letters" authorityYear="1833" box="[264,597,1740,1763]" class="Insecta" family="Margarodidae" genus="Porphyrophora" kingdom="Animalia" order="Hemiptera" pageId="87" phylum="Arthropoda" rank="species" species="hamelii">
<emphasis id="9CA8EAD3FF94822D7A617FFBD917F976" box="[264,515,1740,1763]" italics="true" pageId="87">Porphyrophora hamelii</emphasis>
Brandt
</taxonomicName>
,
<emphasis id="9CA8EAD3FF94822D79097FFBD961F976" box="[608,629,1740,1763]" italics="true" pageId="87">in</emphasis>
Brandt &amp; Ratzeburg, 1833: 356.
</paragraph>
<paragraph id="AE6336C1FF94822D7A617E23DE3EF8C3" blockId="87.[264,1322,1812,1878]" pageId="87">
<emphasis id="9CA8EAD3FF94822D7A617E23DADFF8B8" box="[264,459,1812,1837]" italics="true" pageId="87">Material studied</emphasis>
<materialsCitation id="1EB43C9CFF94822D7A517E0BDE3EF8C3" box="[312,1322,1852,1878]" collectingDate="1999-08-05" collectionCode="BMNH" collectorName="H. Vahedi" country="Iran" location="Kermenshah" pageId="87" specimenCount="1">
<emphasis id="9CA8EAD3FF94822D7A517E0BDA97F8C3" bold="true" box="[312,387,1852,1878]" pageId="87">
<collectingCountry id="D6CB7651FF94822D7A517E0BDA97F8C3" box="[312,387,1852,1878]" name="Iran" pageId="87">IRAN</collectingCountry>
</emphasis>
,
<location id="AB03601AFF94822D7AFF7E0BD939F8C3" LSID="urn:lsid:plazi:treatment:267587D7FF9482277A617F6ED98BFB33:AB03601AFF94822D7AFF7E0BD939F8C3" box="[406,557,1852,1878]" country="Iran" name="Kermenshah" pageId="87">Kermenshah</location>
,
<collectedFrom id="1DC0753CFF94822D79557E0BD82BF8C0" box="[572,831,1852,1878]" pageId="87">
on
<taxonomicName id="69DC4D42FF94822D790C7E0BD82BF8C0" box="[613,831,1852,1877]" class="Actinopterygii" family="Cynodontidae" genus="Cynodon" kingdom="Animalia" order="Characiformes" pageId="87" phylum="Chordata" rank="species" species="dactylon">
<emphasis id="9CA8EAD3FF94822D790C7E0BD82BF8C0" box="[613,831,1852,1877]" italics="true" pageId="87">Cynodon dactylon</emphasis>
</taxonomicName>
</collectedFrom>
,
<date id="DA621001FF94822D783B7E0BD8C7F8C3" box="[850,979,1852,1878]" pageId="87" value="1999-08-05">
<collectingDate id="CA26E9E9FF94822D783B7E0BD8C7F8C3" box="[850,979,1852,1878]" pageId="87" value="1999-08-05">5.viii.1999</collectingDate>
</date>
,
<collectorName id="03295317FF94822D788D7E0BDF49F8C3" box="[996,1117,1852,1878]" pageId="87">H. Vahedi</collectorName>
(
<collectionCode id="C8CDAE04FF94822D7F187E0BDFC4F8C3" box="[1137,1232,1852,1878]" pageId="87">BMNH</collectionCode>
): 10/10
</materialsCitation>
</paragraph>
<paragraph id="AE6336C1FF9B82227A61782AD97BFEA3" blockId="88.[264,623,284,310]" box="[264,623,284,310]" pageId="88">
<specimenCount id="B8DAFD48FF9B82227A61782ADA3BFEBA" box="[264,303,285,303]" pageId="88" type="male">♂♂</specimenCount>
(in fair to good condition).
</paragraph>
<paragraph id="AE6336C1FF9B82227A61785BDACEFE10" blockId="88.[264,1325,364,870]" box="[264,474,364,389]" pageId="88">
<emphasis id="9CA8EAD3FF9B82227A61785BDACEFE10" box="[264,474,364,389]" italics="true" pageId="88">Mounted material</emphasis>
</paragraph>
<paragraph id="AE6336C1FF9B82227A5178A3DF04FCF3" blockId="88.[264,1325,364,870]" pageId="88">
Large, 3.34.3 mm long, 0.921.15 mm wide across prealare. With a pair of large compound eyes which nearly meet ventrally. Body with few setae, none very long; all setae hair­like, apparently with shallow basal sockets (hs); loculate pores (lp) present on abdomen only, probably in dorsopleural region; each
<quantity id="69249B24FF9B822278F07B3BD8EFFDB3" box="[921,1019,524,550]" metricMagnitude="1" metricUnit="m" metricValue="1.15" metricValueMax="1.4" metricValueMin="0.9" pageId="88" unit="m" value="11.5" valueMax="14.0" valueMin="9.0">914 m</quantity>
wide, mainly with 720 loculi. Collared setae (cs), hairs (hrs) and convex pores (cp) absent. Antennae long, with long setae randomly distributed, often with satellite setae (sats). Some sclerotised areas with nodulations. Legs well developed but anterior pair fossorial; anterior femur rather setose, with many bifurcated setae (bs), legs otherwise with few setae; spur­like setae present on all tibiae and tarsi; tarsi 1 segmented; claws without a denticle; claw digitules acute. Abdominal segment I not visible ventrally; abdomen without lateral caudal extensions; segments VI and VII with groups of tubular ducts (tdc) dorsally. Penial sheath strongly sclerotised and quadrate; with a very long, thin aedeagus.
</paragraph>
<paragraph id="AE6336C1FF9B82227A617AABDA53FC20" blockId="88.[264,1323,924,1870]" box="[264,327,924,949]" pageId="88">
<emphasis id="9CA8EAD3FF9B82227A617AABDA53FC20" box="[264,327,924,949]" italics="true" pageId="88">Head</emphasis>
</paragraph>
<paragraph id="AE6336C1FF9B82227A517AF3DF88F8DB" blockId="88.[264,1323,924,1870]" pageId="88">
Triangular in dorsal view, length
<quantity id="69249B24FF9B8222786A7AF3D880FC4B" box="[771,916,964,990]" metricMagnitude="2" metricUnit="m" metricValue="2.775" metricValueMax="2.9" metricValueMin="2.65" pageId="88" unit="m" value="277.5" valueMax="290.0" valueMin="265.0">265290 m</quantity>
, width across compound eyes
<quantity id="69249B24FF9B82227A617ADBDA8EFB93" box="[264,410,1004,1030]" metricMagnitude="2" metricUnit="m" metricValue="4.425" metricValueMax="4.75" metricValueMin="4.1" pageId="88" unit="m" value="442.5" valueMax="475.0" valueMin="410.0">
410
<specimenCount id="B8DAFD48FF9B82227A2A7ADBDA8EFB93" box="[323,410,1004,1030]" pageId="88" type="generic">475 m</specimenCount>
</quantity>
.
<emphasis id="9CA8EAD3FF9B82227AC17ADBD91BFB90" box="[424,527,1004,1029]" italics="true" pageId="88">Dorsally</emphasis>
with a well­developed postoccipital suture (pos) extending across posterior part of epicranium but not reaching eyes laterally; postocciput (poc) possibly absent; rest of dorsal surface of head (dorsomedial part of epicranium (dmep)) barely sclerotised. Midcranial ridge (mcr) absent. Dmep with 03 hs dorsal head setae (dhs) on each side, each
<quantity id="69249B24FF9B82227AAD7DBBD90CFB33" box="[452,536,1164,1190]" metricMagnitude="0" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="88" unit="m" value="6.0" valueMax="7.0" valueMin="5.0">57 m</quantity>
long. Pores absent.
<emphasis id="9CA8EAD3FF9B822278607DBBD860FB30" box="[777,884,1164,1189]" italics="true" pageId="88">Laterally</emphasis>
with a pair of compound eyes (cde), each more or less oval in shape, but much broader ventrally, where they nearly meet; each cde about
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long and each with about 80100 ommatidia. Each compound eye (cde) with a narrow, lightly sclerotised, ocular sclerite (ocs) along postero­dorsal and posterior margins, each with a single ocellus (o) dorsally,
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wide. Preocular ridge (procr) probably present on dorsal margin of cde. Postocular ridge (pocr) present posterior to each ocular sclerite (ocs), extending from near each ocellus to posterior margin of each compound eye; sclerotised ventral projection (p) probably present.
<emphasis id="9CA8EAD3FF9B82227F4D7C93DF9AFA28" box="[1060,1166,1444,1469]" italics="true" pageId="88">Ventrally</emphasis>
with a series of strongly sclerotised ridges forming a four­armed cross, composed of: (i) midcranial ridge (vmcr) anteriorly, (ii) a pair of preocular ridges (procr) laterally and (iii) a pair of preoral ridges (pror) posteriorly; each pror extending posteriorly to fuse with ventral plate (vp) which lies medially along anterior margin of prothorax. Ventral part of epicranium (vmep) otherwise membranous except for some lesser sclerotisation in each angle between mid­cranial ridge (mcr) and preocular ridges (procr); setae few ­ with just 35 hs on each side anterior to preocular ridges. Cranial apophysis (ca) not detected. No structures representing tentorial arms, tentorial bridge or tendon­like apodeme detected but mouth (m) clearly visible in centre of ventral plate (vp). Ventral plate (vp) rather quadrate and sclerotised, extending posteriorly onto prothorax. Ventral sclerites (vs) absent.
</paragraph>
<caption id="FAA36649FF9A82237A617E2ED94AF8C2" ID-DOI="http://doi.org/10.5281/zenodo.5065450" ID-Zenodo-Dep="5065450" httpUri="https://zenodo.org/record/5065450/files/figure.png" pageId="89" startId="89.[264,367,1817,1841]" targetBox="[295,1269,341,1800]" targetPageId="89">
<paragraph id="AE6336C1FF9A82237A617E2ED94AF8C2" blockId="89.[264,1323,1817,1879]" pageId="89">
<emphasis id="9CA8EAD3FF9A82237A617E2EDA8DF8A4" bold="true" box="[264,409,1817,1841]" pageId="89">FIGURE 13.</emphasis>
Male
<taxonomicName id="69DC4D42FF9A82237A897E2DD892F8A5" authority="Brandt. Letters" authorityName="Brandt. Letters" authorityYear="1833" box="[480,902,1817,1841]" class="Insecta" family="Margarodidae" genus="Porphyrophora" kingdom="Animalia" order="Hemiptera" pageId="89" phylum="Arthropoda" rank="species" species="hamelii">
<emphasis id="9CA8EAD3FF9A82237A897E2DD9C9F8A4" box="[480,733,1818,1841]" italics="true" pageId="89">Porphyrophora hamelii</emphasis>
Brandt. Letters
</taxonomicName>
as for Fig. 6 except K = peg­like seta, and U = segment VI of antenna.
</paragraph>
</caption>
<paragraph id="AE6336C1FF9982207A51782CDA62FC23" blockId="90.[264,1323,283,950]" pageId="90">
<emphasis id="9CA8EAD3FF9982207A51782CDAA4FEA0" bold="true" box="[312,432,283,309]" pageId="90">Antennae</emphasis>
: usually 14­segmented (
<specimenCount id="B8DAFD48FF998220798C782BD869FEA3" box="[741,893,284,310]" pageId="90" type="generic">2 specimens</specimenCount>
with 13 segments and 1 with 15 segments); scape without a sclerotised articulatory socket with head; length 1.72.0 mm (ratio of total­body length to antennal length 1:0.49). Scape (scp)
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long,
<quantity id="69249B24FF9982207A6178A3DA83FE3B" box="[264,407,404,430]" metricMagnitude="2" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="90" unit="m" value="130.0" valueMax="140.0" valueMin="120.0">120140 m</quantity>
wide, sclerotised, with about 610 large stiff setae, each about
<quantity id="69249B24FF9982207F1178A3DFFCFE3B" box="[1144,1256,404,430]" metricMagnitude="1" metricUnit="m" metricValue="5.75" metricValueMax="6.0" metricValueMin="5.5" pageId="90" unit="m" value="57.5" valueMax="60.0" valueMin="55.0">5560 m</quantity>
long, + 35 minute hs. Pedicel (pdc)
<quantity id="69249B24FF99822079EB788BD9E0FE43" box="[642,756,444,470]" metricMagnitude="1" metricUnit="m" metricValue="7.9" metricValueMax="8.4" metricValueMin="7.4" pageId="90" unit="m" value="79.0" valueMax="84.0" valueMin="74.0">7484 m</quantity>
long,
<quantity id="69249B24FF998220782B788BD8D5FE43" box="[834,961,444,470]" metricMagnitude="1" metricUnit="m" metricValue="9.75" metricValueMax="10.5" metricValueMin="9.0" pageId="90" unit="m" value="97.5" valueMax="105.0" valueMin="90.0">90105 m</quantity>
wide; with 1220 hs only; no campaniform sensilla (camp) or coeloconic sensilla (cos) detected on any specimen. Each flagellar segment (III to apex)
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wide; each segment with 1223 long setae, mostly on outer margin, particularly on more apical segments (setae mostly about
<quantity id="69249B24FF9982207A617B6BDA93FDE3" box="[264,391,604,630]" metricMagnitude="2" metricUnit="m" metricValue="1.03" metricValueMax="1.1" metricValueMin="0.96" pageId="90" unit="m" value="103.0" valueMax="110.0" valueMin="96.0">96110 m</quantity>
long); each long seta with 0 or 1 (usually 0, rarely 2) satellite setae (sats); most segments also with a few short hs; segments IIIXI each with 05 bristles (ab) on outer surface; segments VIXI with a group of peg­like setae, also on outer surface, while segment VI (and sometimes IV and apical segment) have a group of coeloconic sensilla (cos). Apical segment elongate oval (partially divided on
<specimenCount id="B8DAFD48FF99822078BE7BCBDF71FC83" box="[983,1125,764,790]" pageId="90" type="generic">1 specimen</specimenCount>
), with a diffuse group of 1420 long seta; with a few basiconic sensilla (bass); capitate setae (caps) absent. Segment lengths (m): III 140145, IV 153170, V 145155, VI 150155, VII 135153, VIII 130153, IX 125135, X 115135, XI 115125, XII 110125, XIII 100130, XIV 125169.
</paragraph>
<paragraph id="AE6336C1FF9982207A617ADBDA48FB90" blockId="90.[264,1323,1004,1892]" box="[264,348,1004,1029]" pageId="90">
<emphasis id="9CA8EAD3FF9982207A617ADBDA48FB90" box="[264,348,1004,1029]" italics="true" pageId="90">Thorax</emphasis>
</paragraph>
<paragraph id="AE6336C1FF9982207A517D23DF02FAEF" blockId="90.[264,1323,1004,1892]" pageId="90">
<emphasis id="9CA8EAD3FF9982207A517D23DAD5FBBB" bold="true" box="[312,449,1044,1070]" pageId="90">Prothorax:</emphasis>
neck broad, with no indications of a cervical groove.
<emphasis id="9CA8EAD3FF9982207F2B7D23DFBCFBB8" box="[1090,1192,1044,1069]" italics="true" pageId="90">Dorsally</emphasis>
: pronotum (prn) represented by a single sclerotised bar dorsally more or less overlying ventral plate. Post­tergites (pt) small.
<emphasis id="9CA8EAD3FF99822079497D53D998FBE8" box="[544,652,1124,1149]" italics="true" pageId="90">Laterally</emphasis>
with a pair of strong cervical sclerites (cv), which extend anteriorly to ocular sclerite (ocs); with a large proepimeron (epm
<subScript id="32583484FF9982207F4D7DABDF38FB3F" attach="left" box="[1060,1068,1180,1194]" fontSize="6" pageId="90">1</subScript>
). Pleural ridge (plr
<subScript id="32583484FF9982207E707DABDE35FB3F" attach="left" box="[1305,1313,1180,1194]" fontSize="6" pageId="90">1</subScript>
) short, extending dorsally from articulation with coxa; pleural apophysis (pla
<subScript id="32583484FF9982207FC67DF0DFA3FB40" attach="right" box="[1199,1207,1223,1237]" fontSize="6" pageId="90">1</subScript>
) distinct.
<emphasis id="9CA8EAD3FF9982207A617DD5DA66FB6E" box="[264,370,1250,1275]" italics="true" pageId="90">Ventrally</emphasis>
: prosternum (stn
<subScript id="32583484FF99822079557DC5D950FA95" attach="left" box="[572,580,1266,1280]" fontSize="6" pageId="90">1</subScript>
) membranous, without a sternite or prosternal apophysis; with no transverse ridge. Pores absent. Setae as follows: median pronotal setae (mpns) absent; with a small group of 14 hs propleural setae (pl
<subScript id="32583484FF99822078337C72D876FAC6" attach="right" box="[858,866,1349,1363]" fontSize="6" pageId="90">1</subScript>
s) on each side; with 15 post­tergital setae (pts) associated with each post­tergite; setae absent ventrally.
</paragraph>
<paragraph id="AE6336C1FF9982217A517CB0D929FD62" blockId="90.[264,1323,1004,1892]" lastBlockId="91.[264,1324,284,1896]" lastPageId="91" pageId="90">
<emphasis id="9CA8EAD3FF9982207A517CB0DAD9FA34" bold="true" box="[312,461,1415,1441]" pageId="90">Mesothorax</emphasis>
:
<emphasis id="9CA8EAD3FF9982207AB67CBFD92BFA34" box="[479,575,1416,1441]" italics="true" pageId="90">dorsally</emphasis>
: prescutum (prsc) large and oval (length
<quantity id="69249B24FF9982207F467CBFDFD5FA37" box="[1071,1217,1416,1442]" metricMagnitude="2" metricUnit="m" metricValue="1.85" metricValueMax="2.1" metricValueMin="1.6" pageId="90" unit="m" value="185.0" valueMax="210.0" valueMin="160.0">160210 m</quantity>
, greatest width
<quantity id="69249B24FF9982207A307C87DAFEFA5F" box="[345,490,1456,1482]" metricMagnitude="2" metricUnit="m" metricValue="2.575" metricValueMax="2.7" metricValueMin="2.45" pageId="90" unit="m" value="257.5" valueMax="270.0" valueMin="245.0">245270 m</quantity>
), with a nodulated surface; prescutal ridges (pscr) short; prescutal sutures (pscs) present, separating prescutum from scutum; prescutum without prescutal setae (prscs). Scutum (sct) sclerotised throughout and nodulated on anterolateral margin; with a group of 2125 hs scutal setae (scts) on each side medially. Scutellum (scl) triangular,
<quantity id="69249B24FF9982207AE37F67D90CF9FF" box="[394,536,1616,1642]" metricMagnitude="2" metricUnit="m" metricValue="3.2" metricValueMax="3.8" metricValueMin="2.6" pageId="90" unit="m" value="320.0" valueMax="380.0" valueMin="260.0">260380 m</quantity>
long,
<quantity id="69249B24FF998220790E7F67D9E1F9FF" box="[615,757,1616,1642]" metricMagnitude="2" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="90" unit="m" value="600.0" valueMax="700.0" valueMin="500.0">500700 m</quantity>
wide; anterior margin of scutoscutellar sutures (scuts) extending close to prescutum anteriorly; ridge (rd) along posterior margin slightly weaker; without oval membranous areas laterally; without scutellar setae (scls).
<emphasis id="9CA8EAD3FF9982207FDE7F97DE36F92C" box="[1207,1314,1696,1721]" italics="true" pageId="90">Laterally</emphasis>
: prealare (pra) elongate. Tegula (teg) large and sclerotised, with a group of 1630 short hs tegular setae (tegs). Mesopleural ridge (plr
<subScript id="32583484FF99822079907E37D815F89B" attach="right" box="[761,769,1792,1806]" fontSize="6" pageId="90">2</subScript>
) well developed, with a deep pleural apophysis (pla
<subScript id="32583484FF9982207A5F7E1CDA2AF8AC" attach="right" box="[310,318,1835,1849]" fontSize="6" pageId="90">2</subScript>
). Mesepisternum (eps
<subScript id="32583484FF998220792F7E1CD95AF8AC" attach="right" box="[582,590,1835,1849]" fontSize="6" pageId="90">2</subScript>
) nodulated. Mesothoracic spiracles (sp
<subScript id="32583484FF9982207F767E1CDF33F8AC" attach="left" box="[1055,1063,1835,1849]" fontSize="6" pageId="90">2</subScript>
): width of peritremes
<quantity id="69249B24FF9982207A617E71DA9FF8F5" box="[264,395,1862,1888]" metricMagnitude="2" metricUnit="m" metricValue="1.025" metricValueMax="1.2" metricValueMin="0.85" pageId="90" unit="m" value="102.5" valueMax="120.0" valueMin="85.0">
85
<specimenCount id="B8DAFD48FF9982207A5D7E71DA9FF8F5" box="[308,395,1862,1888]" pageId="90" type="generic">120 m</specimenCount>
</quantity>
.
<emphasis id="9CA8EAD3FF9982207AF67E71D91DF8CA" box="[415,521,1862,1887]" italics="true" pageId="90">Ventrally</emphasis>
: basisternum (stn
<subScript id="32583484FF99822079837E61D9E6F8F1" attach="left" box="[746,754,1878,1892]" fontSize="6" pageId="90">2</subScript>
) large, nodulated; length
<quantity id="69249B24FF9982207F2B7E71DFC7F8F5" box="[1090,1235,1862,1888]" metricMagnitude="2" metricUnit="m" metricValue="5.075" metricValueMax="5.7" metricValueMin="4.45" pageId="90" unit="m" value="507.5" valueMax="570.0" valueMin="445.0">445570 m</quantity>
, width
<quantity id="69249B24FF9882217A61782BDA8EFEA3" box="[264,410,284,310]" metricMagnitude="2" metricUnit="m" metricValue="6.35" metricValueMax="7.0" metricValueMin="5.7" pageId="91" unit="m" value="635.0" valueMax="700.0" valueMin="570.0">570700 m</quantity>
; with a total of 3570 hs basisternal setae (stn
<subScript id="32583484FF9882217885781BD8E0FEAF" attach="both" box="[1004,1012,300,314]" fontSize="6" pageId="91">2</subScript>
s) distributed throughout; median ridge (mdr) absent; anteriorly without a distinct marginal ridge (mr) but ridge well developed between basisternum and lateropleurite (lpl); posteriorly with well­developed precoxal ridges (pcr
<subScript id="32583484FF9882217A9E7890DAEBFE20" attach="left" box="[503,511,423,437]" fontSize="6" pageId="91">2</subScript>
); furca (f) moderately narrow posteriorly, waisted, with rather long arms which diverge strongly; lateropleurite (lpl) approximately triangular in shape, bounded ventrally by a broad marginal ridge, posteriorly by a broad precoxal ridge and laterally by an extension of subepisternal ridge (ser); each lpl with a group of 918 hs lateropleurite setae (lpls); 1 or 2 hs also occasionally present on marginal and subepisternal ridges; subepisternal ridge (ser) distinct. Postmesospiracular setae (pm
<subScript id="32583484FF9882217E647B45DE01FD15" attach="both" box="[1293,1301,626,640]" fontSize="6" pageId="91">2</subScript>
s) absent.
<emphasis id="9CA8EAD3FF9882217A0A7BBAD91EFD33" box="[355,522,653,678]" italics="true" pageId="91">Wing sclerites</emphasis>
: apparently similar to other members of this group: costal complex of veins (ccx) distinct; axillary sclerites 1ax, 2ax and 3ax showing nothing distinctive and additional sclerite present.
</paragraph>
<paragraph id="AE6336C1FF9882217A517A33DA7FFB55" blockId="91.[264,1324,284,1896]" pageId="91">
<emphasis id="9CA8EAD3FF9882217A517A33DAD8FC8B" bold="true" box="[312,460,772,798]" pageId="91">Metathorax</emphasis>
:
<emphasis id="9CA8EAD3FF9882217AB47A32D929FC8B" box="[477,573,773,798]" italics="true" pageId="91">dorsally</emphasis>
: metapostnotum (pn
<subScript id="32583484FF988221785D7A22D828FCB6" attach="left" box="[820,828,789,803]" fontSize="6" pageId="91">3</subScript>
) present as a pair of large sclerites, each with a broad arm which extends pleurally; with 410 hs metatergal setae (mts) extending across segment.
<emphasis id="9CA8EAD3FF9882217AB97A6FD92FFCE4" box="[464,571,856,881]" italics="true" pageId="91">Laterally</emphasis>
: dorsospiracular setae (dss) absent. Suspensorial sclerites (ss) well developed. Pleural ridge (plr
<subScript id="32583484FF98822179F07AA7D9B5FC0B" attach="right" box="[665,673,912,926]" fontSize="6" pageId="91">3</subScript>
) well developed; precoxal ridge (pcr
<subScript id="32583484FF9882217F357AA7DF70FC0B" attach="right" box="[1116,1124,912,926]" fontSize="6" pageId="91">3</subScript>
) well developed and extending about
<quantity id="69249B24FF98822179677A9DD988FC51" box="[526,668,938,964]" metricMagnitude="2" metricUnit="m" metricValue="2.31" metricValueMax="2.6" metricValueMin="2.02" pageId="91" unit="m" value="231.0" valueMax="260.0" valueMin="202.0">202260 m</quantity>
ventrally; with a moderately deep pleural apophysis (pla
<subScript id="32583484FF9882217A5F7AD4DA2AFC64" attach="right" box="[310,318,995,1009]" fontSize="6" pageId="91">3</subScript>
). Metepisternum (eps
<subScript id="32583484FF988221792C7AD4D959FC64" attach="left" box="[581,589,995,1009]" fontSize="6" pageId="91">3</subScript>
) present as a sclerotisation on ventral margin of metapleural ridge (plr
<subScript id="32583484FF9882217A157D3ADA90FB8E" attach="right" box="[380,388,1037,1051]" fontSize="6" pageId="91">3</subScript>
); metepimeron (epm
<subScript id="32583484FF98822179EF7D3AD99AFB8E" attach="left" box="[646,654,1037,1051]" fontSize="6" pageId="91">3</subScript>
) absent. Posterior spiracles (sp
<subScript id="32583484FF9882217F7A7D3ADF0FFB8E" attach="right" box="[1043,1051,1037,1051]" fontSize="6" pageId="91">3</subScript>
): width of peritremes about
<quantity id="69249B24FF9882217A397D1EDAC6FBD6" box="[336,466,1065,1091]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.15" metricValueMin="0.85" pageId="91" unit="m" value="100.0" valueMax="115.0" valueMin="85.0">
85
<specimenCount id="B8DAFD48FF9882217A147D1EDAC6FBD6" box="[381,466,1065,1091]" pageId="91" type="generic">115 m</specimenCount>
</quantity>
.
<emphasis id="9CA8EAD3FF9882217AB77D1ED95CFBD7" box="[478,584,1065,1090]" italics="true" pageId="91">Ventrally</emphasis>
: metasternum (stn
<subScript id="32583484FF988221784B7D0ED83EFBD2" attach="right" box="[802,810,1081,1095]" fontSize="6" pageId="91">3</subScript>
) large and sclerotised, with well­developed lateral apophyses (stn
<subScript id="32583484FF988221794C7D54D939FBE4" attach="both" box="[549,557,1123,1137]" fontSize="6" pageId="91">3</subScript>
a). Postmesoprecoxal ridge setae (ppcr
<subScript id="32583484FF9882217F587D54DF2DFBE4" attach="both" box="[1073,1081,1123,1137]" fontSize="6" pageId="91">2</subScript>
s) absent; anterior metasternal setae (amss): total of 3 or 4 hs; posterior metasternal setae (pmss): total of 913 hs.
</paragraph>
<paragraph id="AE6336C1FF9882217A517DF9D933F9C5" blockId="91.[264,1324,284,1896]" pageId="91">
<emphasis id="9CA8EAD3FF9882217A517DF9DA93FB7D" bold="true" box="[312,391,1230,1256]" pageId="91">Wings</emphasis>
: large and well developed; length 3.23.3 mm; width 1.21.4 mm (ratio of length to width 1:0.4; ratio of total­body length to wing length 1:0.86). Subcostal thickening (sclt) well developed; wing anterior to subcostal thickening only lightly sclerotised; rest of wing membranous, sometimes with faint reticulations; with a line of 914 circular sensoria (sens) and 2027 fine setae (als) along sclt; radius (rad) and media (med) veins apparently represented by light sclerotisations; alar fold (af) not detected. Alar lobe (al) present but unsclerotised. Hamulohalteres (h) mainly sclerotised and quite broad, with a strong vein along anterior margin, which has a strong indentation proximally; length
<quantity id="69249B24FF9882217A317F39DAFCF9BD" box="[344,488,1550,1576]" metricMagnitude="2" metricUnit="m" metricValue="3.3" metricValueMax="3.4" metricValueMin="3.2" pageId="91" unit="m" value="330.0" valueMax="340.0" valueMin="320.0">320340 m</quantity>
, width
<quantity id="69249B24FF98822179547F39D9A6F9BD" box="[573,690,1550,1576]" metricMagnitude="1" metricUnit="m" metricValue="6.0" metricValueMax="6.5" metricValueMin="5.5" pageId="91" unit="m" value="60.0" valueMax="65.0" valueMin="55.0">5565 m</quantity>
; with 1 blunt apical hamulus (ham), about
<quantity id="69249B24FF9882217FCD7F39DFFDF9BD" box="[1188,1257,1550,1576]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="91" unit="m" value="25.0">25 m</quantity>
long, which bends posteriorly.
</paragraph>
<paragraph id="AE6336C1FF9882267A517F69DFE6FBC3" blockId="91.[264,1324,284,1896]" lastBlockId="92.[264,1323,284,1110]" lastPageId="92" pageId="91">
<emphasis id="9CA8EAD3FF9882217A517F69DA66F9ED" bold="true" box="[312,370,1630,1656]" pageId="91">Legs</emphasis>
.
<emphasis id="9CA8EAD3FF9882217AEB7F68D951F9ED" box="[386,581,1631,1656]" italics="true" pageId="91">Prothoracic legs</emphasis>
: fossorial; lengths (m): coxae (cx) 387445; trochanter (tr) + femur (fm) 673785; tibia (ti) 220260 and tarsus (ta) 225260; coxa (cx) with a group of 3035 small hs near coxal base + about 1820 longer setae distally. Trochanter (tr) with about 12 hs but without a long flagellate seta; each with a group of 819 campaniform sensilla (camp) on each lateral surface near coxal articulation. Femur (fm) with about 110130 bifurcating setae (bs), 60 long hs (
<quantity id="69249B24FF98822178767E11D8BAF8D5" box="[799,942,1830,1856]" metricMagnitude="2" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="91" unit="m" value="105.0" valueMax="110.0" valueMin="100.0">100110 m</quantity>
long) plus about 60+ short hs. Tibia (ti) with 46 spur­like setae along ventral margin; bifurcating setae (bs) absent; with 2540 hs elsewhere; spurs (tibs) at distal end of tibia (tibs) largest, each about
<quantity id="69249B24FF9F82267FC2782BDE3FFEA3" box="[1195,1323,284,310]" metricMagnitude="1" metricUnit="m" metricValue="8.0" metricValueMax="10.0" metricValueMin="6.0" pageId="92" unit="m" value="80.0" valueMax="100.0" valueMin="60.0">60100 m</quantity>
long. Tarsi (ta) 1 segmented, with a group of 819 campaniform sensilla (camp) on proximal dorsal surface; without bifurcating setae; with 07 spinose setae along ventral margin plus about 1012 hs setae on other surfaces; tarsal spurs (tabs) barely differentiated, each
<quantity id="69249B24FF9F82267A92788BD979FE43" box="[507,621,444,470]" metricMagnitude="1" metricUnit="m" metricValue="4.75" metricValueMax="5.5" metricValueMin="4.0" pageId="92" unit="m" value="47.5" valueMax="55.0" valueMin="40.0">4055 m</quantity>
long; tarsal digitules (tdgt) apparently absent. Claws (c) broadly fused to base of tarsus, without any articulation; length
<quantity id="69249B24FF9F8226789F78D3DF92FE6B" box="[1014,1158,484,510]" metricMagnitude="2" metricUnit="m" metricValue="1.55" metricValueMax="1.8" metricValueMin="1.3" pageId="92" unit="m" value="155.0" valueMax="180.0" valueMin="130.0">130180 m</quantity>
; without claw digitules (cdgt).
<emphasis id="9CA8EAD3FF9F82267AA67B3BD83AFDB0" box="[463,814,524,549]" italics="true" pageId="92">Meso­ and metathoracic legs</emphasis>
: lengths (m): coxae (cx) II 330385; III
<quantity id="69249B24FF9F82267A617B03DA8EFDDB" box="[264,410,564,590]" metricMagnitude="2" metricUnit="m" metricValue="3.8" metricValueMax="4.15" metricValueMin="3.45" pageId="92" unit="m" value="380.0" valueMax="415.0" valueMin="345.0">345415 m</quantity>
; metacoxae with a group of 1926 small hs near base + 3155 longer setae distally. Trochanter (tr) + femur (fm): II 645700; III
<quantity id="69249B24FF9F822678F27B6BDF3DFDE3" box="[923,1065,604,630]" metricMagnitude="2" metricUnit="m" metricValue="7.025" metricValueMax="7.6" metricValueMin="6.45" pageId="92" unit="m" value="702.5" valueMax="760.0" valueMin="645.0">645760 m</quantity>
long; metatrochanter with 1425 hs but without either a long flagellate seta or bifurcate setae (bs); trochanter with a group of 1014 campaniform sensilla (camp) on each lateral surface near coxal articulation. Metafemur with 2224 spinose setae along posteroventral margin, plus 4560 hs. Tibia (ti): II 660725; III
<quantity id="69249B24FF9F822679127BCBD81FFC83" box="[635,779,764,790]" metricMagnitude="2" metricUnit="m" metricValue="7.4" metricValueMax="7.65" metricValueMin="7.15" pageId="92" unit="m" value="740.0" valueMax="765.0" valueMin="715.0">715765 m</quantity>
long; metatibiae with 1935 spur­like setae along ventral margin; bifurcating setae (bs) absent; with 4560 hs elsewhere; each tibia with 2 tibial spurs (tibs) distally, longest about
<quantity id="69249B24FF9F8226783D7A7BD8D1FCF3" box="[852,965,844,870]" metricMagnitude="1" metricUnit="m" metricValue="4.25" metricValueMax="5.0" metricValueMin="3.5" pageId="92" unit="m" value="42.5" valueMax="50.0" valueMin="35.0">3550 m</quantity>
long. Tarsi (ta) 1 segmented lengths: II 260275; III
<quantity id="69249B24FF9F822679437A43D9ADFC1B" box="[554,697,884,910]" metricMagnitude="2" metricUnit="m" metricValue="2.975" metricValueMax="3.15" metricValueMin="2.8" pageId="92" unit="m" value="297.5" valueMax="315.0" valueMin="280.0">280315 m</quantity>
long; with a group of 2 (5?) campaniform sensilla (camp) on proximal dorsal surface and perhaps more distally; without bifurcating setae (bs); with 712 spur­like setae along ventral margin plus about 2530 hs setae on other surfaces; tarsal spurs (tabs) barely differentiated,
<quantity id="69249B24FF9F8226780E7ADBD8CCFB93" box="[871,984,1004,1030]" metricMagnitude="1" metricUnit="m" metricValue="3.3" metricValueMax="3.8" metricValueMin="2.8" pageId="92" unit="m" value="33.0" valueMax="38.0" valueMin="28.0">2838 m</quantity>
long; tarsal digitules (tdgt) apparently absent. Claws (c) broad basally, without a denticle (cd); claw III about
<quantity id="69249B24FF9F82267A617D0BDA83FBC3" box="[264,407,1084,1110]" metricMagnitude="2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="92" unit="m" value="110.0" valueMax="120.0" valueMin="100.0">100120 m</quantity>
long, with 2 strongly spinose digitules (cdgt), each about
<quantity id="69249B24FF9F82267F577D0BDFA4FBC3" box="[1086,1200,1084,1110]" metricMagnitude="1" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="92" unit="m" value="22.5" valueMax="25.0" valueMin="20.0">2025 m</quantity>
long.
</paragraph>
<paragraph id="AE6336C1FF9F82267A617DBBDA63FB30" blockId="92.[264,1323,1164,1870]" box="[264,375,1164,1189]" pageId="92">
<emphasis id="9CA8EAD3FF9F82267A617DBBDA63FB30" box="[264,375,1164,1189]" italics="true" pageId="92">Abdomen</emphasis>
</paragraph>
<paragraph id="AE6336C1FF9F82267A517D83DA7DF8B3" blockId="92.[264,1323,1164,1870]" pageId="92">
Without caudal extensions. Small tergites (at) present mediolaterally on anterior borders of all segments; large sternites (as) present medially on segments IIVIII. Dorsal abdominal setae (ads) absent medially on dorsum of segments IVII, but with 2 or 3 on each side of VIII; ventral abdominal setae (avs) in broad bands of about 1330 hs across posterior margin of sternites IIVII and with a small diffuse group of 3 or 4 on either side of VIII. Loculate pores (lp) absent from tergites and sternites of segments IVII. Segments VI and VII each with a median group of 1527 large, sclerotised tubular ducts (tdc) which secrete long wax filaments (each filament perhaps
<quantity id="69249B24FF9F8226783E7CFBD8A3FA73" box="[855,951,1484,1510]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="92" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">45 mm</quantity>
long); each group of ducts with a band of minute setae along anterior margin. Pleurites only represented by 2 lightly sclerotised areas laterally on segment VIII. Pleural setae probably divided into dorsal (dps) and ventral pleural setae (vps), latter represented by a single hs per segment; dps very few (14 per segment) associated with a group of lp, with 13 lp on segments IIV, 38 lp on V &amp; VI, 713 lp on VII and 1820 lp across VIII dorsally; each loculate pore
<quantity id="69249B24FF9F82267FD27FA3DE3FF93B" box="[1211,1323,1684,1710]" metricMagnitude="1" metricUnit="m" metricValue="1.2" metricValueMax="1.4" metricValueMin="1.0" pageId="92" unit="m" value="12.0" valueMax="14.0" valueMin="10.0">1014 m</quantity>
wide, with 720 loculi in 13 rings but without a central loculus. Long setae entirely absent from abdomen. Abdominal spiracles (asp) apparently absent (also absent on adult female).
</paragraph>
<paragraph id="AE6336C1FF9F82277A517E03D87AFDDB" blockId="92.[264,1323,1164,1870]" lastBlockId="93.[264,1323,284,590]" lastPageId="93" pageId="92">
<emphasis id="9CA8EAD3FF9F82267A517E03D910F8DB" bold="true" box="[312,516,1844,1870]" pageId="92">Genital segment</emphasis>
: anus (an) not detected. Penial sheath (ps) strongly sclerotised and quadrate, with rather parallel sides, about
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long,
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wide; most of sclerotised area covered in numerous short hs (pss). Penial sheath with a long ventral slit, whose sides are more membranous than elsewhere on penial sheath. Attached to anterior margin of ventral slit is a broad, rather flat, heavily sclerotised basal rod (bra) from anterior end of which emerges a long, thin aedeagus, perhaps 0.91.0 mm long, possibly in a membranous sheath along part of its length. Aedeagus (aed) appears to emerge through a small projection on ventral, posterior margin of penial sheath, which has a few minute sensilla (psp); no eversible endophallus (eph) found.
</paragraph>
<paragraph id="AE6336C1FF9E82277A617BB3D844FD08" blockId="93.[264,1323,644,1190]" box="[264,848,644,669]" pageId="93">
<emphasis id="9CA8EAD3FF9E82277A617BB3D844FD08" box="[264,848,644,669]" italics="true" pageId="93">Characters typical of male hypogeal margarodids</emphasis>
</paragraph>
<paragraph id="AE6336C1FF9E82277A517B9BD98BFB33" blockId="93.[264,1323,644,1190]" pageId="93">(i) fossorial anterior legs; (ii) thorax broadening rapidly behind head, with no neck; (iii) tarsi all 1 segmented; (iv) hamulohalteres each with 1 hamulus; (iv) each trochanter with a group of round campaniform sensilla; (v) abdominal segments VI and VII each with a median dorsal group of tubular ducts; (vi) scutellum strongly triangular, without a pair of membranous areas laterally; (vii) some species with satellite setae on antennae; (viii) some species with bifurcated setae, particularly on prothoracic legs; (ix) some species with loculate pores; (x) penial sheath generally short and squat; (xi) hamulohalteres with an indentation on anterior margin; (xii) most species with setae on or near each lateropleurite; (xiii) all coxae with a group of very small setae near basal articulation; (xiv) compound eyes particularly large, almost meeting ventrally; (xv) absence of collared setae; (xvi) absence of convex pores; (xvii) scutum without a median membranous area; (xviii) basisternum with marginal ridge poorly demarcated along anterior margin, and (xix) sternites generally well developed.</paragraph>
</subSubSection>
</treatment>
</document>