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<document id="B1A58BEB5887CCD5278CEAC4B6200770" ID-DOI="10.5281/zenodo.213050" ID-GBIF-Dataset="25b27738-26cd-4231-8a21-bdabd68d7797" ID-ISSN="1175-5326" ID-Zenodo-Dep="213050" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460500756554" checkinUser="plazi" docAuthor="Rifkind, Jacques" docDate="2012" docId="1E4A8038FF8EFFB1FF77FA814B0445C6" docLanguage="en" docName="zt03397p027.pdf" docOrigin="Zootaxa 3397" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Enoclerus hespenheidei Rifkind, n. sp." docType="treatment" docVersion="8" lastPageNumber="12" masterDocId="E273F840FF84FFBAFFE0FFBE4E3B424C" masterDocTitle="New Central American and Mexican Enoclerus Gahan (Coleoptera: Cleridae: Clerinae): Part II" masterLastPageNumber="27" masterPageNumber="1" pageNumber="11" updateTime="1698311763485" updateUser="plazi">
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<mods:title id="A27059965917EA411DD4C940C7C56E37">New Central American and Mexican Enoclerus Gahan (Coleoptera: Cleridae: Clerinae): Part II</mods:title>
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<mods:namePart id="18CE25917C87405B4882964C7FD299D8">Rifkind, Jacques</mods:namePart>
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<mods:date id="938E5898B32481B8E1DD0E3E29E4DE4F">2012</mods:date>
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<mods:identifier id="52DF93AFDBFA8377998AF2D34E7E64D9" type="DOI">10.5281/zenodo.213050</mods:identifier>
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<treatment id="1E4A8038FF8EFFB1FF77FA814B0445C6" ID-DOI="http://doi.org/10.5281/zenodo.6168345" ID-GBIF-Taxon="119521294" ID-Zenodo-Dep="6168345" LSID="urn:lsid:plazi:treatment:1E4A8038FF8EFFB1FF77FA814B0445C6" httpUri="http://treatment.plazi.org/id/1E4A8038FF8EFFB1FF77FA814B0445C6" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<subSubSection id="DEF962A5FF8EFFB0FF77FA814EC34736" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="965C312EFF8EFFB0FF77FA814C4B4715" blockId="10.[151,624,1343,1402]" box="[151,624,1343,1369]" pageId="10" pageNumber="11">
<heading id="CD148642FF8EFFB0FF77FA814C4B4715" bold="true" box="[151,624,1343,1369]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
<emphasis id="A497ED3CFF8EFFB0FF77FA814C4B4715" bold="true" box="[151,624,1343,1369]" pageId="10" pageNumber="11">
<taxonomicName id="51E34AADFF8EFFB0FF77FA814C1A4715" authority="Rifkind" authorityName="Rifkind" box="[151,545,1343,1369]" class="Insecta" family="Cleridae" genus="Enoclerus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="hespenheidei" status="sp. nov.">
<emphasis id="A497ED3CFF8EFFB0FF77FA814F834715" bold="true" box="[151,440,1343,1369]" italics="true" pageId="10" pageNumber="11">Enoclerus hespenheidei</emphasis>
Rifkind
</taxonomicName>
,
<taxonomicNameLabel id="BFA45047FF8EFFB0FDCEFA814C4B4715" box="[558,624,1343,1369]" pageId="10" pageNumber="11" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="965C312EFF8EFFB0FF77FADC4EC34736" blockId="10.[151,624,1343,1402]" box="[151,248,1378,1402]" pageId="10" pageNumber="11">
(
<figureCitation id="0ED82DABFF8EFFB0FF7FFADC4ED44736" box="[159,239,1378,1402]" captionStart="FIGURES 10 18" captionStartId="20.[151,264,1876,1899]" captionTargetBox="[264,1345,234,1808]" captionTargetId="figure@20.[212,1382,193,1858]" captionTargetPageId="20" captionText="FIGURES 10 18. Habitus of: 10. Enoclerus cavei Rifkind n. sp. (holotype male). 11. Enoclerus hespenheidei Rifkind n. sp. (holotype female). 12. Enoclerus melissae Rifkind n. sp. (holotype male). 13. Enoclerus opitzi Rifkind n. sp. (holotype male). 14. Enoclerus urbanus Rifkind n. sp. (holotype male (? )). 15. Enoclerus crinitus Rifkind n. sp. (holotype male). 16. Enoclerus mocho Rifkind n. sp. (holotype female). 17. Enoclerus regnadkcin Rifkind n. sp. (holotype female). 18. Enoclerus citricornis Rifkind n. sp. (holotype male)." httpUri="https://zenodo.org/record/213052/files/figure.png" pageId="10" pageNumber="11">Fig. 11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DEF962A5FF8EFFB0FF77FA154D7645EB" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="965C312EFF8EFFB0FF77FA154D7645EB" blockId="10.[151,1437,1450,2035]" pageId="10" pageNumber="11">
<emphasis id="A497ED3CFF8EFFB0FF77FA154F61478F" bold="true" box="[151,346,1450,1475]" pageId="10" pageNumber="11">
<typeStatus id="49588F8CFF8EFFB0FF77FA154EE9478F" box="[151,210,1451,1475]" pageId="10" pageNumber="11">Type</typeStatus>
specimens.
</emphasis>
<typeStatus id="49588F8CFF8EFFB0FE82FA144FF1478E" box="[354,458,1450,1474]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
female:
<collectingCountry id="EEF471BEFF8EFFB0FDCEFA144C97478E" box="[558,684,1450,1474]" name="Costa Rica" pageId="10" pageNumber="11">Costa Rica</collectingCountry>
, Puntarenas Province, Peninsula de Osa, R [ancho] Quemado, Rio Riyito,
<quantity id="511B9CCBFF8EFFB0FF0DFA714F0247AB" box="[237,313,1487,1511]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="11" unit="m" value="200.0">200 m</quantity>
,
<date id="E25D17EEFF8EFFB0FEA7FA714F9D47AB" box="[327,422,1487,1511]" bridgedPair="-" pageId="10" pageNumber="11" value="1990-11">xi1990</date>
, Hanson &amp; Godoy.
<typeStatus id="49588F8CFF8EFFB0FD6DFA714CCE47AB" box="[653,757,1487,1511]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
deposited in
<collectionCode id="F0F2A9EBFF8EFFB0FC73FA714DDB47AB" LSID="urn:lsid:biocol.org:col:33156" box="[915,992,1487,1511]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="10" pageNumber="11">CSCA</collectionCode>
.
<typeStatus id="49588F8CFF8EFFB0FC0CFA6E4A6647A4" box="[1004,1117,1488,1512]" pageId="10" pageNumber="11" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="EEF471BEFF8EFFB0FB8AFA714B2F47AB" box="[1130,1300,1487,1511]" name="Costa Rica" pageId="10" pageNumber="11">COSTA RICA</collectingCountry>
: Puntarenas Province:
<quantity id="511B9CCBFF8EFFB0FEEBFA4B4F4A4441" box="[267,369,1525,1549]" metricMagnitude="3" metricUnit="m" metricValue="5.27" pageId="10" pageNumber="11" unit="km" value="5.27">5, 27 km</quantity>
S. Pto. Jiménez, Rio Piro,
<quantity id="511B9CCBFF8EFFB0FD49FA4B4CDD4441" box="[681,742,1525,1549]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="10" pageNumber="11" unit="m" value="75.0">75 m</quantity>
,
<date id="E25D17EEFF8EFFB0FD14FA4B4D694441" box="[756,850,1525,1549]" bridgedPair="-" pageId="10" pageNumber="11" value="1990-11">xi1990</date>
, Hanson &amp; Godoy; 3, Cerro Rincón,
<quantity id="511B9CCBFF8EFFB0FAE0FA4B4B724441" box="[1280,1353,1525,1549]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="11" unit="m" value="200.0">200 m</quantity>
S. hito [marker],
<quantity id="511B9CCBFF8EFFB0FEE6F9A44F74447E" box="[262,335,1562,1586]" metricMagnitude="2" metricUnit="m" metricValue="7.45" pageId="10" pageNumber="11" unit="m" value="745.0">745 m</quantity>
,
<date id="E25D17EEFF8EFFB0FEBBF9A44F82447E" box="[347,441,1562,1586]" bridgedPair="-" pageId="10" pageNumber="11" value="1990-11">xi1990</date>
, Hanson &amp; Godoy; 1, R. Piro,
<quantity id="511B9CCBFF8EFFB0FCF0F9A44D6C447E" box="[784,855,1562,1586]" metricMagnitude="4" metricUnit="m" metricValue="2.7" pageId="10" pageNumber="11" unit="km" value="27.0">27 km</quantity>
S. Puerto Jiménez, Peninsula Osa,
<date id="E25D17EEFF8EFFB0FB01F9A44B04447E" box="[1249,1343,1562,1586]" bridgedPair="-" pageId="10" pageNumber="11" value="1990-11">xi1990</date>
, P. Hanson, C. Godoy, colls., Malaise trap; 1, Peninsula Osa, Cerro Rincón,
<geoCoordinate id="F3D757E9FF8EFFB0FC51F9814A2E441B" box="[945,1045,1599,1623]" direction="north" orientation="latitude" pageId="10" pageNumber="11" precision="925" value="8.516666">8° 31' N</geoCoordinate>
,
<geoCoordinate id="F3D757E9FF8EFFB0FBC2F9814AA3441B" box="[1058,1176,1599,1623]" direction="west" orientation="longitude" pageId="10" pageNumber="11" precision="925" value="-83.46667">83° 28' W</geoCoordinate>
,
<quantity id="511B9CCBFF8EFFB0FB46F9814ACA441B" box="[1190,1265,1599,1623]" metricMagnitude="2" metricUnit="m" metricValue="7.45" pageId="10" pageNumber="11" unit="m" value="745.0">745 m</quantity>
, Virgin Forest, Malaise trap,
<date id="E25D17EEFF8EFFB0FED4F9DB4FBF4431" box="[308,388,1637,1661]" bridgedPair="-" pageId="10" pageNumber="11" value="1991-01">i1991</date>
, P. Hanson, coll.; 2, same data as previous except
<date id="E25D17EEFF8EFFB0FC29F9DB4A244431" box="[969,1055,1637,1661]" bridgedPair="-" pageId="10" pageNumber="11" value="1990-10">x1990</date>
, Hanson &amp; Godoy, colls.; 1, Estación Sirena,
<quantity id="511B9CCBFF8EFFB0FED5F9344FA744EE" box="[309,412,1674,1698]" metricMagnitude="1" metricUnit="m" metricValue="5.05" metricValueMax="10.0" metricValueMin="0.1" pageId="10" pageNumber="11" unit="m" value="50.5" valueMax="100.0" valueMin="1.0">1100 m</quantity>
,
<date id="E25D17EEFF8EFFB0FE4AF9344C5A44EE" box="[426,609,1674,1698]" pageId="10" pageNumber="11" value="1994-12">December 1994</date>
, G. Fonseca,
<collectionCode id="F0F2A9EBFF8EFFB0FD1CF9344D7944EE" box="[764,834,1674,1698]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="10" pageNumber="11">INBio</collectionCode>
reference # CR1002, 164769, Labeled # LS
<date id="E25D17EEFF8EFFB0FAA0F9344BAC44EE" box="[1344,1431,1674,1698]" pageId="10" pageNumber="11" value="00-05-27">270500</date>
, 508300 #4367; 1, P. N. Corcovado, Est. Sirena,
<quantity id="511B9CCBFF8EFFB0FD33F9114D07448B" box="[723,828,1711,1735]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="10" pageNumber="11" unit="m" value="50.0" valueMax="100.0" valueMin="0.0">0100 m</quantity>
, 1991, Tp. Malaise, LS
<date id="E25D17EEFF8EFFB0FB90F9114AFC448B" box="[1136,1223,1711,1735]" pageId="10" pageNumber="11" value="00-05-27">270500</date>
, 508300,
<collectionCode id="F0F2A9EBFF8EFFB0FADDF9114BB8448B" box="[1341,1411,1711,1735]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="10" pageNumber="11">INBio</collectionCode>
# CR1001, 292989; 1, Peninsula Osa, Est. Sirena,
<date id="E25D17EEFF8EFFB0FD0AF9684D5144A1" box="[746,874,1749,1774]" pageId="10" pageNumber="11" value="1994-08">Ago. 1994</date>
, M. Segura, LS
<date id="E25D17EEFF8EFFB0FBCEF96B4ABF44A1" box="[1070,1156,1749,1773]" pageId="10" pageNumber="11" value="00-05-27">270500</date>
_508300, #3219,
<collectionCode id="F0F2A9EBFF8EFFB0FAB6F96B4BA744A1" box="[1366,1436,1749,1773]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="10" pageNumber="11">INBio</collectionCode>
CR1002, 007800; 1, P. N. Corcovado, Est. Sirena,
<quantity id="511B9CCBFF8EFFB0FD39F9444D7B455E" box="[729,832,1786,1810]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="10" pageNumber="11" unit="m" value="50.0" valueMax="100.0" valueMin="0.0">0100 m</quantity>
,
<date id="E25D17EEFF8EFFB0FCADF9444DFB455E" box="[845,960,1786,1810]" pageId="10" pageNumber="11" value="1989-10">Oct. 1989</date>
, G. Fonseca, LS
<date id="E25D17EEFF8EFFB0FB6FF9444ADD455E" box="[1167,1254,1786,1810]" pageId="10" pageNumber="11" value="00-05-27">270500</date>
, 508300,
<collectionCode id="F0F2A9EBFF8EFFB0FAB6F9444BA7455E" box="[1366,1436,1786,1810]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="10" pageNumber="11">INBio</collectionCode>
CR1000, 660128; 1, Golfo Dulce,
<quantity id="511B9CCBFF8EFFB0FDF9F8A14C6A457B" box="[537,593,1823,1847]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="11" unit="km" value="3.0">3 km</quantity>
SW Rincón,
<quantity id="511B9CCBFF8EFFB0FD07F8A14D19457B" box="[743,802,1823,1847]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="11" unit="m" value="10.0">10 m</quantity>
,
<date id="E25D17EEFF8EFFB0FCCDF8A14DAA457B" box="[813,913,1823,1847]" bridgedPair="-" pageId="10" pageNumber="11" value="1989-12">xii1989</date>
<date id="E25D17EEFF8EFFB0FC4FF8A14A37457B" box="[943,1036,1823,1847]" bridgedPair="-" pageId="10" pageNumber="11" value="1990-03">iii1990</date>
, Hanson; 1, Golfo Dulce,
<quantity id="511B9CCBFF8EFFB0FAD0F8A14B4C457B" box="[1328,1399,1823,1847]" metricMagnitude="4" metricUnit="m" metricValue="2.4" pageId="10" pageNumber="11" unit="km" value="24.0">24 km</quantity>
W. Piedras Blancas,
<quantity id="511B9CCBFF8EFFB0FEBAF8FB4F9F4511" box="[346,420,1861,1885]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="11" unit="m" value="200.0">200 m</quantity>
,
<date id="E25D17EEFF8EFFB0FE51F8FB4C534511" box="[433,616,1861,1885]" pageId="10" pageNumber="11" value="1990-12">December 1990</date>
, Hanson, coll.; 1 Est. Agujas, Sendero Zamia, Rio Agujas,
<quantity id="511B9CCBFF8EFFB0FAEDF8FB4B6C4511" box="[1293,1367,1861,1885]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="11" unit="m" value="300.0">300 m</quantity>
,
<date id="E25D17EEFF8EFFB0FA84F8FB4F3045CE" pageId="10" pageNumber="11" value="1996-01-02" valueMax="1996-01-15" valueMin="1996-01-02">215 Ene. 1996</date>
, A. Azofeifa, Intersection L_S 276750_526550, #7208,
<collectionCode id="F0F2A9EBFF8EFFB0FC9CF8D44DF945CE" box="[892,962,1898,1922]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="10" pageNumber="11">INBio</collectionCode>
CR1002, 396657.
<typeStatus id="49588F8CFF8EFFB0FB75F8D54B3945CF" box="[1173,1282,1899,1923]" pageId="10" pageNumber="11" type="paratype">Paratypes</typeStatus>
are deposited in
<collectionCode id="F0F2A9EBFF8EFFB0FF54F8314F3B45EB" LSID="urn:lsid:biocol.org:col:33156" box="[180,256,1935,1959]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="10" pageNumber="11">CSCA</collectionCode>
,
<collectionCode id="F0F2A9EBFF8EFFB0FEEBF8314F6A45EB" box="[267,337,1935,1959]" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="10" pageNumber="11">INBC</collectionCode>
,
<collectionCode id="F0F2A9EBFF8EFFB0FEBCF8314F8B45EB" LSID="urn:lsid:biocol.org:col:34802" box="[348,432,1935,1959]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="10" pageNumber="11">LACM</collectionCode>
,
<collectionCode id="F0F2A9EBFF8EFFB0FE5BF8314C2B45EB" box="[443,528,1935,1959]" httpUri="http://grbio.org/cool/ww9x-1v7b" name="Museo de Insectos" pageId="10" pageNumber="11">MUCR</collectionCode>
,
<collectionCode id="F0F2A9EBFF8EFFB0FDFBF82E4C4845EB" box="[539,627,1936,1959]" httpUri="http://grbio.org/cool/brtj-zfcn" name="W.F. Barr Entomological Collection" pageId="10" pageNumber="11">WFBM</collectionCode>
,
<collectionCode id="F0F2A9EBFF8EFFB0FD9FF8314CF045EB" box="[639,715,1935,1959]" pageId="10" pageNumber="11">FWSC</collectionCode>
and
<collectionCode id="F0F2A9EBFF8EFFB0FCE1F8314D7245EB" box="[769,841,1935,1959]" pageId="10" pageNumber="11">JNRC</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="DEF962A5FF8EFFB1FF27F80B4B3B43F9" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="965C312EFF8EFFB1FF27F80B4B3B43F9" blockId="10.[151,1437,1450,2035]" lastBlockId="11.[151,1437,151,1930]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<emphasis id="A497ED3CFF8EFFB0FF27F80B4F7A4581" bold="true" box="[199,321,1973,1998]" pageId="10" pageNumber="11">Diagnosis.</emphasis>
Among its Middle American congeners,
<taxonomicName id="51E34AADFF8EFFB0FCFBF8084DF54581" box="[795,974,1973,1997]" class="Insecta" family="Cleridae" genus="Enoclerus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="hespenheidei">
<emphasis id="A497ED3CFF8EFFB0FCFBF8084DF54581" box="[795,974,1973,1997]" italics="true" pageId="10" pageNumber="11">E. hespenheidei</emphasis>
</taxonomicName>
is the only one to possess the following combination of characteristics: pronotum with anterior margin orangetestaceous, posterior margin with a triangular patch of pale vestiture; elytra with subbasal swellings shallow, each crowned with a short, longitudinal, noneburneous carina, terminating posteriorly in a shallowly elevated spine; dorsal vestiture with a subtle but definite velvety aspect, reflecting slight cupreous tints in certain lights; abdomen orangish. Similar, known or potentially sympatric species are separable as follows:
<taxonomicName id="51E34AADFF8FFFB1FD5BFEB74DBE4353" authority="Rifkind" authorityName="Rifkind" box="[699,901,263,288]" class="Insecta" family="Cleridae" genus="Enoclerus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="laselva">
<emphasis id="A497ED3CFF8FFFB1FD5BFEB74D12436C" box="[699,809,264,288]" italics="true" pageId="11" pageNumber="12">E. laselva</emphasis>
Rifkind
</taxonomicName>
and
<taxonomicName id="51E34AADFF8FFFB1FC5AFEB74AB84353" authority="Rifkind" authorityName="Rifkind" box="[954,1155,263,288]" class="Insecta" family="Cleridae" genus="Enoclerus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="duttoni">
<emphasis id="A497ED3CFF8FFFB1FC5AFEB74A1C436C" box="[954,1063,264,288]" italics="true" pageId="11" pageNumber="12">E. duttoni</emphasis>
Rifkind
</taxonomicName>
have the subbasal elytral carinae eburneous, the pronotum and abdomen unicolorous (black);
<taxonomicName id="51E34AADFF8FFFB1FC74FE904A484309" authority="Wolcott" authorityName="Wolcott" box="[916,1139,301,325]" class="Insecta" family="Cleridae" genus="Enoclerus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="fugitivus">
<emphasis id="A497ED3CFF8FFFB1FC74FE904A2F4309" box="[916,1044,301,325]" italics="true" pageId="11" pageNumber="12">E. fugitivus</emphasis>
Wolcott
</taxonomicName>
(and related species) have the subbasal elytral tumescences strongly elevated and cristate above, the abdomen black or bicolorous;
<taxonomicName id="51E34AADFF8FFFB1FAACFEEA4F4843C3" authority="Spinola" authorityName="Spinola" class="Insecta" family="Cleridae" genus="Enoclerus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="crabronarius">
<emphasis id="A497ED3CFF8FFFB1FAACFEEA4F3E43DC" italics="true" pageId="11" pageNumber="12">E. crabronarius</emphasis>
(Spinola)
</taxonomicName>
(and related species) have the subbasal carinae absent, the abdomen black;
<taxonomicName id="51E34AADFF8FFFB1FB22FEC74B4B43C3" authority="Barr" authorityName="Barr" box="[1218,1392,376,400]" class="Insecta" family="Cleridae" genus="Enoclerus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="signifer">
<emphasis id="A497ED3CFF8FFFB1FB22FEC74B0C43DC" box="[1218,1335,376,400]" italics="true" pageId="11" pageNumber="12">E. signifer</emphasis>
Barr
</taxonomicName>
has the pronotum unicolorous (black), the elytra shining and inconspicuously setose, the abdomen black.
</paragraph>
</subSubSection>
<subSubSection id="DEF962A5FF8FFFB1FF27FE7C4F5444E6" pageId="11" pageNumber="12" type="description">
<paragraph id="965C312EFF8FFFB1FF27FE7C4A8D47A3" blockId="11.[151,1437,151,1930]" pageId="11" pageNumber="12">
<emphasis id="A497ED3CFF8FFFB1FF27FE7C4FE44397" bold="true" box="[199,479,450,475]" pageId="11" pageNumber="12">
Description (
<typeStatus id="49588F8CFF8FFFB1FE80FE7C4FE94397" box="[352,466,450,475]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
).
</emphasis>
Length: 8.40 mm. Color: piceus; apices of maxillary palpi, antennomeres 1 and 2 (in part), trochanters, basal 1/3 of metafemora, metathoracic intercoxal process, mesepimeron and anterior margin of pronotum (narrowly at sides, triangularly extended toward posterior at middle), testaceous to reddishtestaceous; abdomen orangish; anterior margin of elytra at middle (exclusive of scutellum), and extending posteriorly to subbasal spine, castaneous; elytra with a pair of truncated, oblique eburneous fasciae at middle, narrowed at anterior margin laterally, rounded internally, broadly interrupted before suture; elytral posterior 1/5 yellowtestaceous anteriorly, coffeecolored posteriorly and internally at suture. Head: antenna moderate in length; club composed of terminal 3 antennomeres; surface finely, densely but shallowly, granulatepunctate, set with mostly suberect and a few longer, erect pale setae, rather thinly arrayed on front, more densely so on cranium. Pronotum: about as broad as long, slightly narrower than elytra at base; transverse impression broadly Ushaped at middle, rather shallow; anterior margin subtruncate; disk subflattened above; posterior slope moderately precipitous; collar not particularly broad; surface sculpturing as on head; anterior pale region clothed with very short and short, suberect pale setae; disk with a dense but inconspicuous covering of very short, erect dark setae, intermixed with longer, erect dark setae; posterior pronotal 3/7 with a triangular patch of pale pilosity. Elytra: elongate, almost
<date id="E25D17EEFF8FFFB1FB52FC194AEF41F3" box="[1202,1236,935,959]" pageId="11" pageNumber="12">2X</date>
as long as broad, broadest at posterior 2/5; anterior margin feebly bisinuate; humeri rounded; sides gradually constricted behind humeri to anterior 1/3, then gradually expanded to posterior 2/5, from where they gradually, arcuately converge to separately rounded, dehiscent apices; eplipleural fold distinctively broad on anterior 1/3; subbasal tumescences moderate, each crowned with a short carina, terminating posteriorly in a small knob or obtuse spine; disk slightly concave at middle; posterior slope rather gradual; surface densely, deeply and quite coarsely punctate on anterior 1/ 3 (except finely punctate at anterior margin and between subbasal tumescences), punctures less densely arrayed and somewhat less coarse toward middle, then increasingly shallow and inconspicuous posterior to pale median fascia; vestiture on darkened area of disk with a dense but inconspicuous covering of very short, erect dark setae, intermixed with longer, erect dark setae, imparting a slight velvety aspect with cupreous reflections; base thinly set with subrecumbent and erect pale setae; disk posterior to tumescences bearing a sagittate patch of subrecumbent setae; eburneous fascia densely but inconspicuously covered with pale suberect and erect setae; posterior 1/5 with a dense covering of pale, mostly recumbent setae of moderate length. Mesosternum: posterior median process distinctly elevated, slightly emarginate at apex. Metasternum: anterior median process broadly concave at base, spiculate at sides and apically. Abdomen: shining, very sparsely setose; visible sternite 5 with posterior margin transverse; visible sternite 6 with sides oblique, posterior margin rounded. Legs: rather robust.
</paragraph>
<paragraph id="965C312EFF8FFFB1FF27FA434F5444E6" blockId="11.[151,1437,151,1930]" pageId="11" pageNumber="12">
<emphasis id="A497ED3CFF8FFFB1FF27FA434F004459" bold="true" box="[199,315,1533,1558]" pageId="11" pageNumber="12">Variation.</emphasis>
Length of available specimens ranges from 7.20 mm to 10.00 mm. Males have visible sternite 5 broadly, feebly emarginate at posterior margin, visible sternite 6 short, hidden beneath 5 and truncate at posterior margin, surpassed by visible tergite 6, which has the sides oblique and the posterior margin rounded. Setal daggers are absent in the male; the tegmen is well sclerotized. Color and integumental sculpturing are rather uniform within the
<typeStatus id="49588F8CFF8FFFB1FF20F92D4F1B44E7" box="[192,288,1683,1707]" pageId="11" pageNumber="12" type="paratype">paratype</typeStatus>
series.
</paragraph>
</subSubSection>
<subSubSection id="DEF962A5FF8FFFB1FF27F9064A8E44B9" pageId="11" pageNumber="12" type="etymology">
<paragraph id="965C312EFF8FFFB1FF27F9064A8E44B9" blockId="11.[151,1437,151,1930]" pageId="11" pageNumber="12">
<emphasis id="A497ED3CFF8FFFB1FF27F9064F75449C" bold="true" box="[199,334,1720,1744]" pageId="11" pageNumber="12">Etymology.</emphasis>
The specific epithet honors Henry H. Hespenheide, whose work on mimicry complexes among neotropical insects has greatly informed our understanding of patterns of mimicry in
<taxonomicName id="51E34AADFF8FFFB1FBAEF9634A8A44B9" box="[1102,1201,1757,1781]" class="Insecta" family="Cleridae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="family">Cleridae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="DEF962A5FF8FFFB1FF27F8BC4B0A4556" box="[199,1329,1794,1819]" pageId="11" pageNumber="12" type="distribution">
<paragraph id="965C312EFF8FFFB1FF27F8BC4B0A4556" blockId="11.[151,1437,151,1930]" box="[199,1329,1794,1819]" pageId="11" pageNumber="12">
<emphasis id="A497ED3CFF8FFFB1FF27F8BC4F5B4557" bold="true" box="[199,352,1794,1819]" pageId="11" pageNumber="12">Distribution.</emphasis>
Known only from lowland rainforest in the Osa Peninsula of southwestern
<collectingCountry id="EEF471BEFF8FFFB1FB4FF8BC4B164556" box="[1199,1325,1794,1818]" name="Costa Rica" pageId="11" pageNumber="12">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="DEF962A5FF8FFFB1FF27F8994B0445C6" pageId="11" pageNumber="12" type="biology_ecology">
<paragraph id="965C312EFF8FFFB1FF27F8994B0445C6" blockId="11.[151,1437,151,1930]" pageId="11" pageNumber="12">
<emphasis id="A497ED3CFF8FFFB1FF27F8994F1C450C" bold="true" box="[199,295,1831,1856]" pageId="11" pageNumber="12">Biology.</emphasis>
Like several other
<taxonomicName id="51E34AADFF8FFFB1FDE0F8964C4A450C" box="[512,625,1832,1856]" class="Insecta" family="Cleridae" genus="Enoclerus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="A497ED3CFF8FFFB1FDE0F8964C4A450C" box="[512,625,1832,1856]" italics="true" pageId="11" pageNumber="12">Enoclerus</emphasis>
</taxonomicName>
species that inhabit primary rainforest (e.g.
<taxonomicName id="51E34AADFF8FFFB1FB84F8974AE8450C" box="[1124,1235,1832,1856]" class="Insecta" family="Cleridae" genus="Enoclerus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="laselva">
<emphasis id="A497ED3CFF8FFFB1FB84F8974AE8450C" box="[1124,1235,1832,1856]" italics="true" pageId="11" pageNumber="12">E. laselva</emphasis>
</taxonomicName>
), the new species has been collected almost exclusively in Malaise traps. It may be that these species occupy a spatial niche above the understory, as they do not generally turn up in beating or sweeping samples from low lying vegetation.
</paragraph>
</subSubSection>
</treatment>
</document>