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<document ID-DOI="http://dx.doi.org/10.3897/asp.79.e67185" ID-GBIF-Dataset="0f8f850f-4b02-4477-b780-bf054d7c6131" ID-GBIF-Taxon="182075311" ID-Pensoft-Pub="1864-8312-79-115" ID-Pensoft-UUID="A4EA6692C24753F7B3CB5DFCAF1EFFAD" ID-ZooBank="6D857849A9ED4DA3A5B834FDAAB0A318" LSID="urn:lsid:zoobank.org:act:EA121C9A-5C41-4D42-B238-5213A549EA4F" ModsDocID="1864-8312-79-115" checkinTime="1621063390318" checkinUser="pensoft" docAuthor="da Silveira, Luiz Felipe Lima, Roza, Andre Silva, Vaz, Stephanie &amp; Mermudes, Jose Ricardo M." docDate="2021" docId="13DC2F171EA05C6280790D2A421393B3" docLanguage="en" docName="ArthropodSystPhyl 79: 115-150" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 79" docPubDate="2021-05-14" docSource="http://dx.doi.org/10.3897/asp.79.e67185" docTitle="Costalampys gen. nov. Silveira, Roza, Vaz &amp; Mermudes" docType="treatment" docVersion="3" id="A4EA6692C24753F7B3CB5DFCAF1EFFAD" lastPageNumber="115" masterDocId="A4EA6692C24753F7B3CB5DFCAF1EFFAD" masterDocTitle="Description and phylogenetic analysis of a new firefly genus from the Atlantic Rainforest, with five new species and new combinations (Coleoptera: Lampyridae: Lampyrinae)" masterLastPageNumber="150" masterPageNumber="115" pageId="0" pageNumber="115" updateTime="1632022650993" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Description and phylogenetic analysis of a new firefly genus from the Atlantic Rainforest, with five new species and new combinations (Coleoptera: Lampyridae: Lampyrinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>da Silveira, Luiz Felipe Lima</mods:namePart>
<mods:affiliation>Biology Department, Western Carolina University, 122 Central Dr, Cullowhee, NC 28723, United States of America; Luiz Felipe Lima da Silveira [silveira. lfl @ gmail. com]</mods:affiliation>
<mods:nameIdentifier type="email">silveira.lfl@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Roza, Andre Silva</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0886-5159</mods:nameIdentifier>
<mods:affiliation>Laboratorio de Entomologia, Departamento de Zoologia, Instituto de Biologia, Universidade Federal do Rio de Janeiro, A 1 - 107, Bloco A, Av. Carlos Chagas Filho, 373, Cidade Universitaria, Ilha do Fundao, Rio de Janeiro, Brasil; Andre Silva Roza [andreroza 1993 @ gmail. com]; Jose Ricardo Miras Mermudes [jrmermudes @ gmail. com]</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vaz, Stephanie</mods:namePart>
<mods:affiliation>Laboratorio de Ecologia de Insetos, Departamento de Ecologia, Instituto de Biologia, Universidade Federal do Rio de Janeiro, A 0 - 111, Bloco A, Av. Carlos Chagas Filho, 373, Cidade Universitaria, Ilha do Fundao, Rio de Janeiro, Brasil; Stephanie Vaz [anievaz @ gmail. com]</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mermudes, Jose Ricardo M.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-2030-7483</mods:nameIdentifier>
<mods:affiliation>Laboratorio de Entomologia, Departamento de Zoologia, Instituto de Biologia, Universidade Federal do Rio de Janeiro, A 1 - 107, Bloco A, Av. Carlos Chagas Filho, 373, Cidade Universitaria, Ilha do Fundao, Rio de Janeiro, Brasil; Andre Silva Roza [andreroza 1993 @ gmail. com]; Jose Ricardo Miras Mermudes [jrmermudes @ gmail. com]</mods:affiliation>
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<mods:title>Arthropod Systematics &amp; amp; Phylogeny</mods:title>
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<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-05-14</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>79</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>115</mods:start>
<mods:end>150</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/asp.79.e67185</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.79.e67185</mods:identifier>
<mods:identifier type="Pensoft-Pub">1864-8312-79-115</mods:identifier>
<mods:identifier type="ZooBank">6D857849A9ED4DA3A5B834FDAAB0A318</mods:identifier>
<mods:identifier type="Pensoft-UUID">A4EA6692C24753F7B3CB5DFCAF1EFFAD</mods:identifier>
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<treatment ID-GBIF-Taxon="182075311" LSID="urn:lsid:zoobank.org:act:EA121C9A-5C41-4D42-B238-5213A549EA4F" httpUri="http://treatment.plazi.org/id/13DC2F171EA05C6280790D2A421393B3" lastPageNumber="115" pageId="0" pageNumber="115">
<subSubSection pageId="0" pageNumber="115" type="nomenclature">
<paragraph pageId="0" pageNumber="115">
<taxonomicName LSID="http://zoobank.org/EA121C9A-5C41-4D42-B238-5213A549EA4F" authority="gen. nov. Silveira, Roza, Vaz &amp; Mermudes" authorityName="gen. nov. Silveira, Roza, Vaz &amp; Mermudes" class="Insecta" family="Lampyridae" genus="Costalampys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Costalampys" order="Coleoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">Costalampys gen. nov. Silveira, Roza, Vaz &amp; Mermudes</taxonomicName>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Costalampys bella sp. nov. A, Male habitus, dorsal view; B, Male habitus, ventral view; C, Female habitus, dorsal view; D, Female habitus, ventral view; E, Male antena, lateral view; F, Male pronotum, dorsal view; G, Male terminal segments, ventral view; H, Pygidium, dorsal view; I, Syntergite, dorsal view; J, Sternum IX, Ventral view; K, Aedeagus, dorsal view; L, Aedeagus, lateral view; M, Aedeagus, ventral view. Scale bar: 2 mm (A-D); Scale bar: 1.0 mm (E; G); Scale bar: 0.5 mm (F; H-M)." figureDoi="10.3897/asp.79.e67185.figure6" httpUri="https://binary.pensoft.net/fig/543209" pageId="0" pageNumber="115">Figs 6</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Costalampys bisbinotata (Pic) comb. nov. A, Male habitus, dorsal view; B, Male habitus, ventral view; C, Female habitus, dorsal view; D, Female habitus, ventral view; E, Male antena, lateral view; F, Male pronotum, dorsal view; G, Male terminal segments, ventral view; H, Pygidium, dorsal view; I, Syntergite, dorsal view; J, Sternum IX, Ventral view; K, Aedeagus, dorsal view; L, Aedeagus, lateral view; M, Aedeagus, ventral view. Scale bar: 2 mm (A-D); Scale bar: 1.0 mm (E; G); Scale bar: 0.5 mm (F; H-M)." figureDoi="10.3897/asp.79.e67185.figure7" httpUri="https://binary.pensoft.net/fig/543210" pageId="0" pageNumber="115">, 7</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Costalampys capixaba sp. nov. A, Male habitus, dorsal view; B, Male habitus, ventral view; C, Female habitus, dorsal view; D, Female habitus, ventral view; E, Male antena, lateral view; F, Male pronotum, dorsal view; G, Male terminal segments, ventral view; H, Pygidium, dorsal view; I, Syntergite, dorsal view; J, Sternum IX, Ventral view; K, Aedeagus, dorsal view; L, Aedeagus, lateral view; M, Aedeagus, ventral view. Scale bar: 2 mm (A-D); Scale bar: 1.0 mm (E-M)." figureDoi="10.3897/asp.79.e67185.figure8" httpUri="https://binary.pensoft.net/fig/543211" pageId="0" pageNumber="115">, 8</figureCitation>
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Costalampys decorata (Olivier) comb. nov. A, Male habitus, dorsal view; B, Male habitus, ventral view; C, Female habitus, dorsal view; D, Female habitus, ventral view; E, Male antena, lateral view; F, Male pronotum, dorsal view; G, Male terminal segments, ventral view; H, Pygidium, dorsal view; I, Syntergite, dorsal view; J, Sternum IX, Ventral view; K, Aedeagus, dorsal view; L, Aedeagus, lateral view; M, Aedeagus, ventral view. Scale bar: 2 mm (A-D; G-H); Scale bar: 0.5 mm (I-M)." figureDoi="10.3897/asp.79.e67185.figure9" httpUri="https://binary.pensoft.net/fig/543212" pageId="0" pageNumber="115">, 9</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Costalampys decorata (Olivier) comb. nov., male in natural environment (in Brazil: SP, Votorantim). Photo author: Enio Branco." figureDoi="10.3897/asp.79.e67185.figure10" httpUri="https://binary.pensoft.net/fig/543213" pageId="0" pageNumber="115">, 10</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Costalampys delicata sp. nov. A, Male habitus, dorsal view; B, Male habitus, ventral view. Scale bar: 2 mm (A-B)." figureDoi="10.3897/asp.79.e67185.figure11" httpUri="https://binary.pensoft.net/fig/543214" pageId="0" pageNumber="115">, 11</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Costalampys delicata sp. nov., male head. A, Head capsule, dorsal view; B, Head capsule, ventral view; C, Head capsule, lateral view; D, Head capsule, frontal view; E, Head capsule, posterior view; F, Head capsule, occipital view; G, Mandible, dorsal view; H, Mandible, ventral view; I, Tentorium, dorsal view; J, Tentorium, frontal view; K, Tentorium, lateral view; L, antenna, lateral view. Scale bar: 0.5 mm (A-K); Scale bar: 1.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure12" httpUri="https://binary.pensoft.net/fig/543215" pageId="0" pageNumber="115">, 12</figureCitation>
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Costalampys delicata sp. nov. A, Male prothorax, dorsal view; B, Male prothorax, ventral view; C, Male prothorax, lateral view; D, Male prothorax, posterior view; E, Mesoscutellum and alinotum, dorsal view; F, Pterothoracical sclerites, lateral view; G, Pterothoracical sclerites, ventral view; H, Pterothoracical sclerites, lateral view; I, Proleg, lateral view; J, Mesoleg, lateral view; K, Metaleg, lateral view; L, Right wing, dorsal. Scale bar: 1 mm (A-K); Scale bar: 2.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure13" httpUri="https://binary.pensoft.net/fig/543216" pageId="0" pageNumber="115">, 13</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Costalampys delicata sp. nov. A, Abdominal sclerites, dorsal view; B, Abdominal sclerites, ventral view; C, Pygidium, dorsal view; D, Syntergite, dorsal view; E, Sternite IX, ventral view; F, Aedeagus, dorsal view; G, Aedeagus, lateral view; H, Aedeagus, ventral view. Scale bar: 0.5 mm (A-H)." figureDoi="10.3897/asp.79.e67185.figure14" httpUri="https://binary.pensoft.net/fig/543217" pageId="0" pageNumber="115">, 14</figureCitation>
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Costalampys delicata sp. nov. A, Female habitus, dorsal view; B, Female habitus, ventral view; C, Female antenna, lateral view; D, Sternum VIII, ventral view; E, Pygidium, Dorsal view; F, Ovipositor, dorsal view; G, Ovipositor, lateral view; H, Internal genitalia, dorsal view; I, Internal genitalia, ventral view. Scale bar: 2 mm (A-C); Scale bar: 1.0 mm (D-I)." figureDoi="10.3897/asp.79.e67185.figure15" httpUri="https://binary.pensoft.net/fig/543218" pageId="0" pageNumber="115">, 15</figureCitation>
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Costalampys joanae sp. nov. A, Male habitus, dorsal view; B, Male habitus, ventral view; C, Male antenna, lateral view; D, Male pronotum, dorsal view; E, Sternum IX, ventral view; F, Pygidium, dorsal view; G, Syntergite, dorsal view; H, Sternum IX, Ventral view; I, Aedeagus, dorsal view, blue arrow pointing to the transverse keel on the dorsal plate; J, Aedeagus, lateral view; K, Aedeagus, ventral view; L, Aedeagus, oblique view, red arrow pointing to the ventral plate. Scale bar: 2 mm (A-C); Scale bar: 1.0 mm (D-L)." figureDoi="10.3897/asp.79.e67185.figure16" httpUri="https://binary.pensoft.net/fig/543219" pageId="0" pageNumber="115">, 16</figureCitation>
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Costalampys klugii (Motschulsky) comb. nov. A, Male habitus, dorsal view; B, Male habitus, ventral view; C, Female habitus, dorsal view; D, Female habitus, ventral view; E, Male antenna, lateral view; F, Male pronotum, dorsal view; G, Sternum IX, ventral view; H, Pygidium, dorsal view; I, Syntergite, dorsal view; J, Sternum IX, Ventral view; K, Aedeagus, dorsal view; L, Aedeagus, lateral view; M, Aedeagus, ventral view. Scale bar: 2 mm (A-D); Scale bar: 1.0 mm (E-M)." figureDoi="10.3897/asp.79.e67185.figure17" httpUri="https://binary.pensoft.net/fig/543220" pageId="0" pageNumber="115">, 17</figureCitation>
<figureCitation captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Costalampys minima sp. nov. A, Male habitus, dorsal view; B, Male habitus, ventral view; C, Male antenna, lateral view; D, Male pronotum, dorsal view; E, Male terminal segments, ventral view; F, Pygidium, dorsal view; G, Syntergite, dorsal view; H, Sternum IX, Ventral view; I, Aedeagus, dorsal view; J, Aedeagus, lateral view; K, Aedeagus, ventral view. Scale bar: 2 mm (A-B); Scale bar: 1.0 mm (C; E); Scale bar: 0.5 mm (D; F-K)." figureDoi="10.3897/asp.79.e67185.figure18" httpUri="https://binary.pensoft.net/fig/543221" pageId="0" pageNumber="115">, 18</figureCitation>
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Costalampys pauper (Olivier) comb. nov. A, Male habitus, dorsal view; B, Male habitus, ventral view; C, Female habitus, dorsal view; D, Female habitus, ventral view; E, Male antenna, lateral view; F, Male pronotum, dorsal view; G, Sternum IX, ventral view; H, Pygidium, dorsal view; I, Syntergite, dorsal view; J, Sternum IX, Ventral view; K, Aedeagus, dorsal view; L, Aedeagus, lateral view; M, Aedeagus, ventral view. Scale bar: 2 mm (A-D); Scale bar: 1.0 mm (E); Scale bar: 0.5 mm (G-K)." figureDoi="10.3897/asp.79.e67185.figure19" httpUri="https://binary.pensoft.net/fig/543222" pageId="0" pageNumber="115">, 19</figureCitation>
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Costalampys quadriguttata (Gorham) comb. nov. A, Male habitus, dorsal view; B, Male habitus, ventral view; C, Female habitus, dorsal view; D, Female habitus, ventral view; E, Male antenna, lateral view; F, Male pronotum, dorsal view; G, Sternum IX, ventral view; H, Pygidium, dorsal view; I, Syntergite, dorsal view; J, Sternum IX, Ventral view; K, Aedeagus, dorsal view; L, Aedeagus, lateral view; M, Aedeagus, ventral view. Scale bar: 2 mm (A-D); Scale bar: 1.0 mm (E-M)." figureDoi="10.3897/asp.79.e67185.figure20" httpUri="https://binary.pensoft.net/fig/543223" pageId="0" pageNumber="115">, 20</figureCitation>
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Costalampys tricolor (Gorham) comb. nov. A, Male habitus, dorsal view; B, Male habitus, ventral view; C, Female habitus, dorsal view; D, Female habitus, ventral view; E, Male antenna, lateral view; F, Male pronotum, dorsal view; G, Sternum IX, ventral view; H, Pygidium, dorsal view; I, Syntergite, dorsal view; J, Sternum IX, Ventral view; K, Aedeagus, dorsal view; L, Aedeagus, lateral view; M, Aedeagus, ventral view. Scale bar: 2 mm (A-D); Scale bar: 1.0 mm (E; G); Scale bar: 0.5 mm (F; H-M)." figureDoi="10.3897/asp.79.e67185.figure21" httpUri="https://binary.pensoft.net/fig/543224" pageId="0" pageNumber="115">, 21</figureCitation>
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Distribution map of Costalampys gen. nov. species." figureDoi="10.3897/asp.79.e67185.figure22" httpUri="https://binary.pensoft.net/fig/543225" pageId="0" pageNumber="115">, 22</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="115" type="type species">
<paragraph pageId="0" pageNumber="115">Type species.</paragraph>
<paragraph pageId="0" pageNumber="115">
<taxonomicName authority="Silveira, Roza, Vaz &amp; Mermudes, 2021-14" authorityName="Silveira, Roza, Vaz &amp; Mermudes, 2021-14" class="Insecta" family="Lampyridae" genus="Costalampys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Costalampys delicata" order="Hymenoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="delicata">
<emphasis italics="true" pageId="0" pageNumber="115">Costalampys delicata</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="115">sp. nov.</emphasis>
, by original designation.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="115" type="etymology">
<paragraph pageId="0" pageNumber="115">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="115">
<taxonomicName authority="Silveira, Roza, Vaz &amp; Mermudes, 2021-14" authorityName="Silveira, Roza, Vaz &amp; Mermudes, 2021-14" class="Insecta" family="Lampyridae" genus="Costalampys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Costalampys" order="Hymenoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Costalampys</emphasis>
</taxonomicName>
is proposed in honor of our dear Professor and friend, the entomologist Dr. Cleide Costa, which greatly influenced our lives and the study of beetles. Gender feminine.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="115" type="diagnosis">
<paragraph pageId="0" pageNumber="115">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="115">
Antenna with 11 antennomeres, covered in decumbent, short and thin bristles (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Costalampys delicata sp. nov. A, Male habitus, dorsal view; B, Male habitus, ventral view. Scale bar: 2 mm (A-B)." figureDoi="10.3897/asp.79.e67185.figure11" httpUri="https://binary.pensoft.net/fig/543214" pageId="0" pageNumber="115">11A-B</figureCitation>
,
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Costalampys delicata sp. nov., male head. A, Head capsule, dorsal view; B, Head capsule, ventral view; C, Head capsule, lateral view; D, Head capsule, frontal view; E, Head capsule, posterior view; F, Head capsule, occipital view; G, Mandible, dorsal view; H, Mandible, ventral view; I, Tentorium, dorsal view; J, Tentorium, frontal view; K, Tentorium, lateral view; L, antenna, lateral view. Scale bar: 0.5 mm (A-K); Scale bar: 1.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure12" httpUri="https://binary.pensoft.net/fig/543215" pageId="0" pageNumber="115">12L</figureCitation>
), antennal socket wide, about half as wide as eye, close-set, reniform, antennifer process conspicuous (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Costalampys delicata sp. nov., male head. A, Head capsule, dorsal view; B, Head capsule, ventral view; C, Head capsule, lateral view; D, Head capsule, frontal view; E, Head capsule, posterior view; F, Head capsule, occipital view; G, Mandible, dorsal view; H, Mandible, ventral view; I, Tentorium, dorsal view; J, Tentorium, frontal view; K, Tentorium, lateral view; L, antenna, lateral view. Scale bar: 0.5 mm (A-K); Scale bar: 1.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure12" httpUri="https://binary.pensoft.net/fig/543215" pageId="0" pageNumber="115">12A-F</figureCitation>
). Maxillary palpomere IV fusiform (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Costalampys delicata sp. nov., male head. A, Head capsule, dorsal view; B, Head capsule, ventral view; C, Head capsule, lateral view; D, Head capsule, frontal view; E, Head capsule, posterior view; F, Head capsule, occipital view; G, Mandible, dorsal view; H, Mandible, ventral view; I, Tentorium, dorsal view; J, Tentorium, frontal view; K, Tentorium, lateral view; L, antenna, lateral view. Scale bar: 0.5 mm (A-K); Scale bar: 1.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure12" httpUri="https://binary.pensoft.net/fig/543215" pageId="0" pageNumber="115">12A-F</figureCitation>
). Labial palpomere III securiform, with sides divergent posteriorly (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Costalampys delicata sp. nov., male head. A, Head capsule, dorsal view; B, Head capsule, ventral view; C, Head capsule, lateral view; D, Head capsule, frontal view; E, Head capsule, posterior view; F, Head capsule, occipital view; G, Mandible, dorsal view; H, Mandible, ventral view; I, Tentorium, dorsal view; J, Tentorium, frontal view; K, Tentorium, lateral view; L, antenna, lateral view. Scale bar: 0.5 mm (A-K); Scale bar: 1.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure12" httpUri="https://binary.pensoft.net/fig/543215" pageId="0" pageNumber="115">12A-F</figureCitation>
). Pronotum roughly semilunar, with a marginal row of wider, deep punctures; disc convex in lateral view; hypomeron with ventral margin sinuose (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Costalampys delicata sp. nov. A, Male prothorax, dorsal view; B, Male prothorax, ventral view; C, Male prothorax, lateral view; D, Male prothorax, posterior view; E, Mesoscutellum and alinotum, dorsal view; F, Pterothoracical sclerites, lateral view; G, Pterothoracical sclerites, ventral view; H, Pterothoracical sclerites, lateral view; I, Proleg, lateral view; J, Mesoleg, lateral view; K, Metaleg, lateral view; L, Right wing, dorsal. Scale bar: 1 mm (A-K); Scale bar: 2.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure13" httpUri="https://binary.pensoft.net/fig/543216" pageId="0" pageNumber="115">13A-D</figureCitation>
). Abdominal terga with posterior angles almost right-angled to slightly produced (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Costalampys delicata sp. nov. A, Abdominal sclerites, dorsal view; B, Abdominal sclerites, ventral view; C, Pygidium, dorsal view; D, Syntergite, dorsal view; E, Sternite IX, ventral view; F, Aedeagus, dorsal view; G, Aedeagus, lateral view; H, Aedeagus, ventral view. Scale bar: 0.5 mm (A-H)." figureDoi="10.3897/asp.79.e67185.figure14" httpUri="https://binary.pensoft.net/fig/543217" pageId="0" pageNumber="115">14A</figureCitation>
). Tibial spurs present. Male with antennomeres III-X cylindrical and basally flabellate, lamellae up to five times longer than antennomere, sternum IX exposed, apically rounded (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Costalampys delicata sp. nov. A, Abdominal sclerites, dorsal view; B, Abdominal sclerites, ventral view; C, Pygidium, dorsal view; D, Syntergite, dorsal view; E, Sternite IX, ventral view; F, Aedeagus, dorsal view; G, Aedeagus, lateral view; H, Aedeagus, ventral view. Scale bar: 0.5 mm (A-H)." figureDoi="10.3897/asp.79.e67185.figure14" httpUri="https://binary.pensoft.net/fig/543217" pageId="0" pageNumber="115">14C</figureCitation>
). Phallus with dorsal and ventral plates; dorsal plate basally fused to parameres, with a subapical transversal keel, without transverse groove, apically rounded or acute, not indented (Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Costalampys joanae sp. nov. A, Male habitus, dorsal view; B, Male habitus, ventral view; C, Male antenna, lateral view; D, Male pronotum, dorsal view; E, Sternum IX, ventral view; F, Pygidium, dorsal view; G, Syntergite, dorsal view; H, Sternum IX, Ventral view; I, Aedeagus, dorsal view, blue arrow pointing to the transverse keel on the dorsal plate; J, Aedeagus, lateral view; K, Aedeagus, ventral view; L, Aedeagus, oblique view, red arrow pointing to the ventral plate. Scale bar: 2 mm (A-C); Scale bar: 1.0 mm (D-L)." figureDoi="10.3897/asp.79.e67185.figure16" httpUri="https://binary.pensoft.net/fig/543219" pageId="0" pageNumber="115">16I-L</figureCitation>
). Female with antennae compressed and serrate (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Costalampys delicata sp. nov. A, Female habitus, dorsal view; B, Female habitus, ventral view; C, Female antenna, lateral view; D, Sternum VIII, ventral view; E, Pygidium, Dorsal view; F, Ovipositor, dorsal view; G, Ovipositor, lateral view; H, Internal genitalia, dorsal view; I, Internal genitalia, ventral view. Scale bar: 2 mm (A-C); Scale bar: 1.0 mm (D-I)." figureDoi="10.3897/asp.79.e67185.figure15" httpUri="https://binary.pensoft.net/fig/543218" pageId="0" pageNumber="115">15C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="115" type="description">
<paragraph pageId="0" pageNumber="115">
Description. Head Figs
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Costalampys delicata sp. nov. A, Male habitus, dorsal view; B, Male habitus, ventral view. Scale bar: 2 mm (A-B)." figureDoi="10.3897/asp.79.e67185.figure11" httpUri="https://binary.pensoft.net/fig/543214" pageId="0" pageNumber="115">11A, B</figureCitation>
,
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Costalampys delicata sp. nov., male head. A, Head capsule, dorsal view; B, Head capsule, ventral view; C, Head capsule, lateral view; D, Head capsule, frontal view; E, Head capsule, posterior view; F, Head capsule, occipital view; G, Mandible, dorsal view; H, Mandible, ventral view; I, Tentorium, dorsal view; J, Tentorium, frontal view; K, Tentorium, lateral view; L, antenna, lateral view. Scale bar: 0.5 mm (A-K); Scale bar: 1.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure12" httpUri="https://binary.pensoft.net/fig/543215" pageId="0" pageNumber="115">12A-F</figureCitation>
);
</paragraph>
<paragraph pageId="0" pageNumber="115">
Head capsule about a 1/3
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide (in dorsal view, Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Costalampys delicata sp. nov., male head. A, Head capsule, dorsal view; B, Head capsule, ventral view; C, Head capsule, lateral view; D, Head capsule, frontal view; E, Head capsule, posterior view; F, Head capsule, occipital view; G, Mandible, dorsal view; H, Mandible, ventral view; I, Tentorium, dorsal view; J, Tentorium, frontal view; K, Tentorium, lateral view; L, antenna, lateral view. Scale bar: 0.5 mm (A-K); Scale bar: 1.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure12" httpUri="https://binary.pensoft.net/fig/543215" pageId="0" pageNumber="115">12A</figureCitation>
), lateral margins slightly convergent posteriorly (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Costalampys delicata sp. nov., male head. A, Head capsule, dorsal view; B, Head capsule, ventral view; C, Head capsule, lateral view; D, Head capsule, frontal view; E, Head capsule, posterior view; F, Head capsule, occipital view; G, Mandible, dorsal view; H, Mandible, ventral view; I, Tentorium, dorsal view; J, Tentorium, frontal view; K, Tentorium, lateral view; L, antenna, lateral view. Scale bar: 0.5 mm (A-K); Scale bar: 1.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure12" httpUri="https://binary.pensoft.net/fig/543215" pageId="0" pageNumber="115">12A</figureCitation>
), slightly longer than tall (in lateral view, Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Costalampys delicata sp. nov., male head. A, Head capsule, dorsal view; B, Head capsule, ventral view; C, Head capsule, lateral view; D, Head capsule, frontal view; E, Head capsule, posterior view; F, Head capsule, occipital view; G, Mandible, dorsal view; H, Mandible, ventral view; I, Tentorium, dorsal view; J, Tentorium, frontal view; K, Tentorium, lateral view; L, antenna, lateral view. Scale bar: 0.5 mm (A-K); Scale bar: 1.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure12" httpUri="https://binary.pensoft.net/fig/543215" pageId="0" pageNumber="115">12 C</figureCitation>
). Frons slightly prominent and swollen (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Costalampys delicata sp. nov., male head. A, Head capsule, dorsal view; B, Head capsule, ventral view; C, Head capsule, lateral view; D, Head capsule, frontal view; E, Head capsule, posterior view; F, Head capsule, occipital view; G, Mandible, dorsal view; H, Mandible, ventral view; I, Tentorium, dorsal view; J, Tentorium, frontal view; K, Tentorium, lateral view; L, antenna, lateral view. Scale bar: 0.5 mm (A-K); Scale bar: 1.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure12" httpUri="https://binary.pensoft.net/fig/543215" pageId="0" pageNumber="115">12C</figureCitation>
). Antennal sockets reniform, 1/2 as wide as eye; antennifer process conspicuous (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Costalampys delicata sp. nov., male head. A, Head capsule, dorsal view; B, Head capsule, ventral view; C, Head capsule, lateral view; D, Head capsule, frontal view; E, Head capsule, posterior view; F, Head capsule, occipital view; G, Mandible, dorsal view; H, Mandible, ventral view; I, Tentorium, dorsal view; J, Tentorium, frontal view; K, Tentorium, lateral view; L, antenna, lateral view. Scale bar: 0.5 mm (A-K); Scale bar: 1.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure12" httpUri="https://binary.pensoft.net/fig/543215" pageId="0" pageNumber="115">12A-D</figureCitation>
). Vertex somewhat convex, posterior margin bisinuose (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Costalampys delicata sp. nov., male head. A, Head capsule, dorsal view; B, Head capsule, ventral view; C, Head capsule, lateral view; D, Head capsule, frontal view; E, Head capsule, posterior view; F, Head capsule, occipital view; G, Mandible, dorsal view; H, Mandible, ventral view; I, Tentorium, dorsal view; J, Tentorium, frontal view; K, Tentorium, lateral view; L, antenna, lateral view. Scale bar: 0.5 mm (A-K); Scale bar: 1.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure12" httpUri="https://binary.pensoft.net/fig/543215" pageId="0" pageNumber="115">12A, B</figureCitation>
). Antenna with 11 antennomeres, scape constricted proximally, pedicel almost as long as wide and constricted medially, antennomeres III-X subequal in length, with decumbent bristles; Frontoclypeus slightly curved anteriorly (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Costalampys delicata sp. nov., male head. A, Head capsule, dorsal view; B, Head capsule, ventral view; C, Head capsule, lateral view; D, Head capsule, frontal view; E, Head capsule, posterior view; F, Head capsule, occipital view; G, Mandible, dorsal view; H, Mandible, ventral view; I, Tentorium, dorsal view; J, Tentorium, frontal view; K, Tentorium, lateral view; L, antenna, lateral view. Scale bar: 0.5 mm (A-K); Scale bar: 1.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure12" httpUri="https://binary.pensoft.net/fig/543215" pageId="0" pageNumber="115">12C</figureCitation>
). Labrum connected to frontoclypeus by membrane; about 3
<normalizedToken originalValue="×">x</normalizedToken>
wider than long, anterior margin straight to somewhat rounded, sclerotized or evanescent (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Costalampys delicata sp. nov., male head. A, Head capsule, dorsal view; B, Head capsule, ventral view; C, Head capsule, lateral view; D, Head capsule, frontal view; E, Head capsule, posterior view; F, Head capsule, occipital view; G, Mandible, dorsal view; H, Mandible, ventral view; I, Tentorium, dorsal view; J, Tentorium, frontal view; K, Tentorium, lateral view; L, antenna, lateral view. Scale bar: 0.5 mm (A-K); Scale bar: 1.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure12" httpUri="https://binary.pensoft.net/fig/543215" pageId="0" pageNumber="115">12D</figureCitation>
). Mandibles long and slender, monotonically arcuate, apex acute, internal tooth absent, external margin very sparsely setose at basal 1/2, bristles very thin and bright, barely visible (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Costalampys delicata sp. nov., male head. A, Head capsule, dorsal view; B, Head capsule, ventral view; C, Head capsule, lateral view; D, Head capsule, frontal view; E, Head capsule, posterior view; F, Head capsule, occipital view; G, Mandible, dorsal view; H, Mandible, ventral view; I, Tentorium, dorsal view; J, Tentorium, frontal view; K, Tentorium, lateral view; L, antenna, lateral view. Scale bar: 0.5 mm (A-K); Scale bar: 1.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure12" httpUri="https://binary.pensoft.net/fig/543215" pageId="0" pageNumber="115">12A-D</figureCitation>
). Maxilla with cardo well-sclerotized, triangular; stipe oblong in ventral view, internal margin slightly curved, posterior margins truncate, palp with four palpomeres; palpomere III triangular (dorsal view); IV lanceolate, with internal margin covered with minute, dense bristles, 2-3
<normalizedToken originalValue="×">x</normalizedToken>
longer than III (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Costalampys delicata sp. nov., male head. A, Head capsule, dorsal view; B, Head capsule, ventral view; C, Head capsule, lateral view; D, Head capsule, frontal view; E, Head capsule, posterior view; F, Head capsule, occipital view; G, Mandible, dorsal view; H, Mandible, ventral view; I, Tentorium, dorsal view; J, Tentorium, frontal view; K, Tentorium, lateral view; L, antenna, lateral view. Scale bar: 0.5 mm (A-K); Scale bar: 1.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure12" httpUri="https://binary.pensoft.net/fig/543215" pageId="0" pageNumber="115">12A-D</figureCitation>
). Labium with mentum well-sclerotized and bristled, completely divided sagittally, forming two plates, each plate elliptic, with inner margin almost straight, outer margin rounded; submentum sclerotized and bristled, subcordiform, elongate; palp with three palpomeres, palpomere III securiform, sides divergent, apical margin slightly curved (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Costalampys delicata sp. nov., male head. A, Head capsule, dorsal view; B, Head capsule, ventral view; C, Head capsule, lateral view; D, Head capsule, frontal view; E, Head capsule, posterior view; F, Head capsule, occipital view; G, Mandible, dorsal view; H, Mandible, ventral view; I, Tentorium, dorsal view; J, Tentorium, frontal view; K, Tentorium, lateral view; L, antenna, lateral view. Scale bar: 0.5 mm (A-K); Scale bar: 1.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure12" httpUri="https://binary.pensoft.net/fig/543215" pageId="0" pageNumber="115">12A-D</figureCitation>
). Gular sutures conspicuous; gular bar transverse and strongly emarginate posteriorly, as wide as submentum greater width. Occiput pyriform, about a 1/3
<normalizedToken originalValue="×">x</normalizedToken>
as wide as head posterior width (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Costalampys delicata sp. nov., male head. A, Head capsule, dorsal view; B, Head capsule, ventral view; C, Head capsule, lateral view; D, Head capsule, frontal view; E, Head capsule, posterior view; F, Head capsule, occipital view; G, Mandible, dorsal view; H, Mandible, ventral view; I, Tentorium, dorsal view; J, Tentorium, frontal view; K, Tentorium, lateral view; L, antenna, lateral view. Scale bar: 0.5 mm (A-K); Scale bar: 1.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure12" httpUri="https://binary.pensoft.net/fig/543215" pageId="0" pageNumber="115">12F</figureCitation>
). Tentorium long and slender, just slightly curved backwards, projected internally at basal 1/3, projection internally rounded (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Costalampys delicata sp. nov., male head. A, Head capsule, dorsal view; B, Head capsule, ventral view; C, Head capsule, lateral view; D, Head capsule, frontal view; E, Head capsule, posterior view; F, Head capsule, occipital view; G, Mandible, dorsal view; H, Mandible, ventral view; I, Tentorium, dorsal view; J, Tentorium, frontal view; K, Tentorium, lateral view; L, antenna, lateral view. Scale bar: 0.5 mm (A-K); Scale bar: 1.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure12" httpUri="https://binary.pensoft.net/fig/543215" pageId="0" pageNumber="115">12I-K</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="115">
Thorax (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Costalampys delicata sp. nov. A, Male prothorax, dorsal view; B, Male prothorax, ventral view; C, Male prothorax, lateral view; D, Male prothorax, posterior view; E, Mesoscutellum and alinotum, dorsal view; F, Pterothoracical sclerites, lateral view; G, Pterothoracical sclerites, ventral view; H, Pterothoracical sclerites, lateral view; I, Proleg, lateral view; J, Mesoleg, lateral view; K, Metaleg, lateral view; L, Right wing, dorsal. Scale bar: 1 mm (A-K); Scale bar: 2.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure13" httpUri="https://binary.pensoft.net/fig/543216" pageId="0" pageNumber="115">13A-L</figureCitation>
).
</emphasis>
Pronotum semilunar, posterior angles somewhat acute; disc slightly wider than long in dorsal view, convex, regularly punctured, punctures small and bristled; with a line of distinct deep marginal punctures; pronotal expansions well developed, anterior expansion maximal length almost half as long as disc, posterior expansion slightly sinuose, especially where disc meets lateral expansion, and by median line; slightly wider than humeral distance (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Costalampys delicata sp. nov. A, Male prothorax, dorsal view; B, Male prothorax, ventral view; C, Male prothorax, lateral view; D, Male prothorax, posterior view; E, Mesoscutellum and alinotum, dorsal view; F, Pterothoracical sclerites, lateral view; G, Pterothoracical sclerites, ventral view; H, Pterothoracical sclerites, lateral view; I, Proleg, lateral view; J, Mesoleg, lateral view; K, Metaleg, lateral view; L, Right wing, dorsal. Scale bar: 1 mm (A-K); Scale bar: 2.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure13" httpUri="https://binary.pensoft.net/fig/543216" pageId="0" pageNumber="115">13A-D</figureCitation>
); lateral expansions straight in posterior view (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Costalampys delicata sp. nov. A, Male prothorax, dorsal view; B, Male prothorax, ventral view; C, Male prothorax, lateral view; D, Male prothorax, posterior view; E, Mesoscutellum and alinotum, dorsal view; F, Pterothoracical sclerites, lateral view; G, Pterothoracical sclerites, ventral view; H, Pterothoracical sclerites, lateral view; I, Proleg, lateral view; J, Mesoleg, lateral view; K, Metaleg, lateral view; L, Right wing, dorsal. Scale bar: 1 mm (A-K); Scale bar: 2.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure13" httpUri="https://binary.pensoft.net/fig/543216" pageId="0" pageNumber="115">13C-D</figureCitation>
). Hypomeron slightly over 2
<normalizedToken originalValue="×">x</normalizedToken>
longer than tall (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Costalampys delicata sp. nov. A, Male prothorax, dorsal view; B, Male prothorax, ventral view; C, Male prothorax, lateral view; D, Male prothorax, posterior view; E, Mesoscutellum and alinotum, dorsal view; F, Pterothoracical sclerites, lateral view; G, Pterothoracical sclerites, ventral view; H, Pterothoracical sclerites, lateral view; I, Proleg, lateral view; J, Mesoleg, lateral view; K, Metaleg, lateral view; L, Right wing, dorsal. Scale bar: 1 mm (A-K); Scale bar: 2.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure13" httpUri="https://binary.pensoft.net/fig/543216" pageId="0" pageNumber="115">13C</figureCitation>
). Prosternum about 4
<normalizedToken originalValue="×">x</normalizedToken>
as wide as its major length (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Costalampys delicata sp. nov. A, Male prothorax, dorsal view; B, Male prothorax, ventral view; C, Male prothorax, lateral view; D, Male prothorax, posterior view; E, Mesoscutellum and alinotum, dorsal view; F, Pterothoracical sclerites, lateral view; G, Pterothoracical sclerites, ventral view; H, Pterothoracical sclerites, lateral view; I, Proleg, lateral view; J, Mesoleg, lateral view; K, Metaleg, lateral view; L, Right wing, dorsal. Scale bar: 1 mm (A-K); Scale bar: 2.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure13" httpUri="https://binary.pensoft.net/fig/543216" pageId="0" pageNumber="115">13B</figureCitation>
). Proendosternite apically clavate, as long as core prosternum major length (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Costalampys delicata sp. nov., male head. A, Head capsule, dorsal view; B, Head capsule, ventral view; C, Head capsule, lateral view; D, Head capsule, frontal view; E, Head capsule, posterior view; F, Head capsule, occipital view; G, Mandible, dorsal view; H, Mandible, ventral view; I, Tentorium, dorsal view; J, Tentorium, frontal view; K, Tentorium, lateral view; L, antenna, lateral view. Scale bar: 0.5 mm (A-K); Scale bar: 1.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure12" httpUri="https://binary.pensoft.net/fig/543215" pageId="0" pageNumber="115">12B, D</figureCitation>
). Mesoscutellum with posterior margin rounded (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Costalampys delicata sp. nov. A, Male prothorax, dorsal view; B, Male prothorax, ventral view; C, Male prothorax, lateral view; D, Male prothorax, posterior view; E, Mesoscutellum and alinotum, dorsal view; F, Pterothoracical sclerites, lateral view; G, Pterothoracical sclerites, ventral view; H, Pterothoracical sclerites, lateral view; I, Proleg, lateral view; J, Mesoleg, lateral view; K, Metaleg, lateral view; L, Right wing, dorsal. Scale bar: 1 mm (A-K); Scale bar: 2.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure13" httpUri="https://binary.pensoft.net/fig/543216" pageId="0" pageNumber="115">13E</figureCitation>
). Elytra ellipsoid, 3-4
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide, pubescent, secondary pubescence present, with a line of conspicuous punctures all over sutural and lateral margins (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Costalampys delicata sp. nov. A, Male habitus, dorsal view; B, Male habitus, ventral view. Scale bar: 2 mm (A-B)." figureDoi="10.3897/asp.79.e67185.figure11" httpUri="https://binary.pensoft.net/fig/543214" pageId="0" pageNumber="115">11 A, B</figureCitation>
). Hind wing well-developed, posterior margin sinuose, slightly over 2
<normalizedToken originalValue="×">x</normalizedToken>
wider than long, r3 almost as long as r4, radial cell 2.5
<normalizedToken originalValue="×">x</normalizedToken>
wider than long, almost reaching anterior margin, costal row of setae inconspicuous (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Costalampys delicata sp. nov. A, Male prothorax, dorsal view; B, Male prothorax, ventral view; C, Male prothorax, lateral view; D, Male prothorax, posterior view; E, Mesoscutellum and alinotum, dorsal view; F, Pterothoracical sclerites, lateral view; G, Pterothoracical sclerites, ventral view; H, Pterothoracical sclerites, lateral view; I, Proleg, lateral view; J, Mesoleg, lateral view; K, Metaleg, lateral view; L, Right wing, dorsal. Scale bar: 1 mm (A-K); Scale bar: 2.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure13" httpUri="https://binary.pensoft.net/fig/543216" pageId="0" pageNumber="115">13L</figureCitation>
); CuA2 crossvein absent, mp-cu crossvein present; RP+MP1+2 as long as 3/4
<normalizedToken originalValue="×">x</normalizedToken>
r4 length, reaching distal margin, J indistinct (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Costalampys delicata sp. nov. A, Male prothorax, dorsal view; B, Male prothorax, ventral view; C, Male prothorax, lateral view; D, Male prothorax, posterior view; E, Mesoscutellum and alinotum, dorsal view; F, Pterothoracical sclerites, lateral view; G, Pterothoracical sclerites, ventral view; H, Pterothoracical sclerites, lateral view; I, Proleg, lateral view; J, Mesoleg, lateral view; K, Metaleg, lateral view; L, Right wing, dorsal. Scale bar: 1 mm (A-K); Scale bar: 2.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure13" httpUri="https://binary.pensoft.net/fig/543216" pageId="0" pageNumber="115">13L</figureCitation>
). Alinotum slightly wider than long, lateral margins slightly convergent posteriorly, posterior margin slightly curved; prescutum extending slightly beyond than half metascutum length (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Costalampys delicata sp. nov. A, Male prothorax, dorsal view; B, Male prothorax, ventral view; C, Male prothorax, lateral view; D, Male prothorax, posterior view; E, Mesoscutellum and alinotum, dorsal view; F, Pterothoracical sclerites, lateral view; G, Pterothoracical sclerites, ventral view; H, Pterothoracical sclerites, lateral view; I, Proleg, lateral view; J, Mesoleg, lateral view; K, Metaleg, lateral view; L, Right wing, dorsal. Scale bar: 1 mm (A-K); Scale bar: 2.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure13" httpUri="https://binary.pensoft.net/fig/543216" pageId="0" pageNumber="115">13E</figureCitation>
); rounded area of scutum weakly sclerotized, scutum-prescutal plates sclerotized and extending ridges to alinotum posterior margin; metascutellum glabrous. Mesosternum weakly sclerotized, posterior margin medially acute (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Costalampys delicata sp. nov. A, Male prothorax, dorsal view; B, Male prothorax, ventral view; C, Male prothorax, lateral view; D, Male prothorax, posterior view; E, Mesoscutellum and alinotum, dorsal view; F, Pterothoracical sclerites, lateral view; G, Pterothoracical sclerites, ventral view; H, Pterothoracical sclerites, lateral view; I, Proleg, lateral view; J, Mesoleg, lateral view; K, Metaleg, lateral view; L, Right wing, dorsal. Scale bar: 1 mm (A-K); Scale bar: 2.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure13" httpUri="https://binary.pensoft.net/fig/543216" pageId="0" pageNumber="115">13F, G</figureCitation>
). Mesoepimeron connate to metasternum (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Costalampys delicata sp. nov. A, Male prothorax, dorsal view; B, Male prothorax, ventral view; C, Male prothorax, lateral view; D, Male prothorax, posterior view; E, Mesoscutellum and alinotum, dorsal view; F, Pterothoracical sclerites, lateral view; G, Pterothoracical sclerites, ventral view; H, Pterothoracical sclerites, lateral view; I, Proleg, lateral view; J, Mesoleg, lateral view; K, Metaleg, lateral view; L, Right wing, dorsal. Scale bar: 1 mm (A-K); Scale bar: 2.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure13" httpUri="https://binary.pensoft.net/fig/543216" pageId="0" pageNumber="115">13G</figureCitation>
). Mesosternum-mesepisternum suture barely visible (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Costalampys delicata sp. nov. A, Male prothorax, dorsal view; B, Male prothorax, ventral view; C, Male prothorax, lateral view; D, Male prothorax, posterior view; E, Mesoscutellum and alinotum, dorsal view; F, Pterothoracical sclerites, lateral view; G, Pterothoracical sclerites, ventral view; H, Pterothoracical sclerites, lateral view; I, Proleg, lateral view; J, Mesoleg, lateral view; K, Metaleg, lateral view; L, Right wing, dorsal. Scale bar: 1 mm (A-K); Scale bar: 2.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure13" httpUri="https://binary.pensoft.net/fig/543216" pageId="0" pageNumber="115">13G</figureCitation>
). Mesepisternum-mesepimeron suture conspicuous (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Costalampys delicata sp. nov. A, Male prothorax, dorsal view; B, Male prothorax, ventral view; C, Male prothorax, lateral view; D, Male prothorax, posterior view; E, Mesoscutellum and alinotum, dorsal view; F, Pterothoracical sclerites, lateral view; G, Pterothoracical sclerites, ventral view; H, Pterothoracical sclerites, lateral view; I, Proleg, lateral view; J, Mesoleg, lateral view; K, Metaleg, lateral view; L, Right wing, dorsal. Scale bar: 1 mm (A-K); Scale bar: 2.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure13" httpUri="https://binary.pensoft.net/fig/543216" pageId="0" pageNumber="115">13F, G</figureCitation>
). Metasternum strongly depressed by mesocoxae, anterior medial keel prominent up to anterior one third, discrimen reaching basal 1/2 of metasternum length, lateral margins divergent posteriorly up to lateral-most part of metacoxa, then convergent posteriorly, posterior margin bisinuose (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Costalampys delicata sp. nov. A, Male prothorax, dorsal view; B, Male prothorax, ventral view; C, Male prothorax, lateral view; D, Male prothorax, posterior view; E, Mesoscutellum and alinotum, dorsal view; F, Pterothoracical sclerites, lateral view; G, Pterothoracical sclerites, ventral view; H, Pterothoracical sclerites, lateral view; I, Proleg, lateral view; J, Mesoleg, lateral view; K, Metaleg, lateral view; L, Right wing, dorsal. Scale bar: 1 mm (A-K); Scale bar: 2.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure13" httpUri="https://binary.pensoft.net/fig/543216" pageId="0" pageNumber="115">13F, G</figureCitation>
). Profemur about as long as protibia, meso and metafemora slightly shorter than respective tibiae (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Costalampys delicata sp. nov. A, Male prothorax, dorsal view; B, Male prothorax, ventral view; C, Male prothorax, lateral view; D, Male prothorax, posterior view; E, Mesoscutellum and alinotum, dorsal view; F, Pterothoracical sclerites, lateral view; G, Pterothoracical sclerites, ventral view; H, Pterothoracical sclerites, lateral view; I, Proleg, lateral view; J, Mesoleg, lateral view; K, Metaleg, lateral view; L, Right wing, dorsal. Scale bar: 1 mm (A-K); Scale bar: 2.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure13" httpUri="https://binary.pensoft.net/fig/543216" pageId="0" pageNumber="115">13I-K</figureCitation>
). Tibial spurs present (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Costalampys delicata sp. nov. A, Male prothorax, dorsal view; B, Male prothorax, ventral view; C, Male prothorax, lateral view; D, Male prothorax, posterior view; E, Mesoscutellum and alinotum, dorsal view; F, Pterothoracical sclerites, lateral view; G, Pterothoracical sclerites, ventral view; H, Pterothoracical sclerites, lateral view; I, Proleg, lateral view; J, Mesoleg, lateral view; K, Metaleg, lateral view; L, Right wing, dorsal. Scale bar: 1 mm (A-K); Scale bar: 2.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure13" httpUri="https://binary.pensoft.net/fig/543216" pageId="0" pageNumber="115">13I-K</figureCitation>
). Tarsomere I about 2
<normalizedToken originalValue="×">x</normalizedToken>
longer than II, II 2
<normalizedToken originalValue="×">x</normalizedToken>
longer than III, III subequal in length to core IV, IV bilobed, lobes reaching two thirds V length, V with claws simple, without inner teeth (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Costalampys delicata sp. nov. A, Male prothorax, dorsal view; B, Male prothorax, ventral view; C, Male prothorax, lateral view; D, Male prothorax, posterior view; E, Mesoscutellum and alinotum, dorsal view; F, Pterothoracical sclerites, lateral view; G, Pterothoracical sclerites, ventral view; H, Pterothoracical sclerites, lateral view; I, Proleg, lateral view; J, Mesoleg, lateral view; K, Metaleg, lateral view; L, Right wing, dorsal. Scale bar: 1 mm (A-K); Scale bar: 2.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure13" httpUri="https://binary.pensoft.net/fig/543216" pageId="0" pageNumber="115">13I-K</figureCitation>
). Mesendosternum with two parasagittal projections directed outwards, irregularly alate (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Costalampys delicata sp. nov. A, Male prothorax, dorsal view; B, Male prothorax, ventral view; C, Male prothorax, lateral view; D, Male prothorax, posterior view; E, Mesoscutellum and alinotum, dorsal view; F, Pterothoracical sclerites, lateral view; G, Pterothoracical sclerites, ventral view; H, Pterothoracical sclerites, lateral view; I, Proleg, lateral view; J, Mesoleg, lateral view; K, Metaleg, lateral view; L, Right wing, dorsal. Scale bar: 1 mm (A-K); Scale bar: 2.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure13" httpUri="https://binary.pensoft.net/fig/543216" pageId="0" pageNumber="115">13I-K</figureCitation>
). Metendosternum spatulate, roughly rhomboid, diamond-shaped (2
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide, median projection acute anteriorly and posteriorly, with two acute lateral laminae) (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Costalampys delicata sp. nov. A, Male prothorax, dorsal view; B, Male prothorax, ventral view; C, Male prothorax, lateral view; D, Male prothorax, posterior view; E, Mesoscutellum and alinotum, dorsal view; F, Pterothoracical sclerites, lateral view; G, Pterothoracical sclerites, ventral view; H, Pterothoracical sclerites, lateral view; I, Proleg, lateral view; J, Mesoleg, lateral view; K, Metaleg, lateral view; L, Right wing, dorsal. Scale bar: 1 mm (A-K); Scale bar: 2.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure13" httpUri="https://binary.pensoft.net/fig/543216" pageId="0" pageNumber="115">13H</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="115">
Abdomen (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Costalampys delicata sp. nov. A, Abdominal sclerites, dorsal view; B, Abdominal sclerites, ventral view; C, Pygidium, dorsal view; D, Syntergite, dorsal view; E, Sternite IX, ventral view; F, Aedeagus, dorsal view; G, Aedeagus, lateral view; H, Aedeagus, ventral view. Scale bar: 0.5 mm (A-H)." figureDoi="10.3897/asp.79.e67185.figure14" httpUri="https://binary.pensoft.net/fig/543217" pageId="0" pageNumber="115">14A-H</figureCitation>
).
</emphasis>
Tergum I with anterior margin membranous (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Costalampys delicata sp. nov. A, Abdominal sclerites, dorsal view; B, Abdominal sclerites, ventral view; C, Pygidium, dorsal view; D, Syntergite, dorsal view; E, Sternite IX, ventral view; F, Aedeagus, dorsal view; G, Aedeagus, lateral view; H, Aedeagus, ventral view. Scale bar: 0.5 mm (A-H)." figureDoi="10.3897/asp.79.e67185.figure14" httpUri="https://binary.pensoft.net/fig/543217" pageId="0" pageNumber="115">14A</figureCitation>
), laterotergite membranous, roughly rectangular, with sparse bristles (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Costalampys delicata sp. nov. A, Abdominal sclerites, dorsal view; B, Abdominal sclerites, ventral view; C, Pygidium, dorsal view; D, Syntergite, dorsal view; E, Sternite IX, ventral view; F, Aedeagus, dorsal view; G, Aedeagus, lateral view; H, Aedeagus, ventral view. Scale bar: 0.5 mm (A-H)." figureDoi="10.3897/asp.79.e67185.figure14" httpUri="https://binary.pensoft.net/fig/543217" pageId="0" pageNumber="115">14A</figureCitation>
); spiracle elliptical, obliquely attached to thorax, more vertically so (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Costalampys delicata sp. nov. A, Abdominal sclerites, dorsal view; B, Abdominal sclerites, ventral view; C, Pygidium, dorsal view; D, Syntergite, dorsal view; E, Sternite IX, ventral view; F, Aedeagus, dorsal view; G, Aedeagus, lateral view; H, Aedeagus, ventral view. Scale bar: 0.5 mm (A-H)." figureDoi="10.3897/asp.79.e67185.figure14" httpUri="https://binary.pensoft.net/fig/543217" pageId="0" pageNumber="115">14A</figureCitation>
). Terga II-VII with posterior corners almost right-angled to slightly projected, somewhat rounded, posterior margins somewhat rounded (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Costalampys delicata sp. nov. A, Abdominal sclerites, dorsal view; B, Abdominal sclerites, ventral view; C, Pygidium, dorsal view; D, Syntergite, dorsal view; E, Sternite IX, ventral view; F, Aedeagus, dorsal view; G, Aedeagus, lateral view; H, Aedeagus, ventral view. Scale bar: 0.5 mm (A-H)." figureDoi="10.3897/asp.79.e67185.figure14" httpUri="https://binary.pensoft.net/fig/543217" pageId="0" pageNumber="115">14A, B</figureCitation>
). Sterna II-IX visible (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Costalampys delicata sp. nov. A, Abdominal sclerites, dorsal view; B, Abdominal sclerites, ventral view; C, Pygidium, dorsal view; D, Syntergite, dorsal view; E, Sternite IX, ventral view; F, Aedeagus, dorsal view; G, Aedeagus, lateral view; H, Aedeagus, ventral view. Scale bar: 0.5 mm (A-H)." figureDoi="10.3897/asp.79.e67185.figure14" httpUri="https://binary.pensoft.net/fig/543217" pageId="0" pageNumber="115">14B</figureCitation>
), VI often bearing a rounded lantern, of variable size. Spiracles dorsal, at sternum anterior th (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Costalampys delicata sp. nov. A, Abdominal sclerites, dorsal view; B, Abdominal sclerites, ventral view; C, Pygidium, dorsal view; D, Syntergite, dorsal view; E, Sternite IX, ventral view; F, Aedeagus, dorsal view; G, Aedeagus, lateral view; H, Aedeagus, ventral view. Scale bar: 0.5 mm (A-H)." figureDoi="10.3897/asp.79.e67185.figure14" httpUri="https://binary.pensoft.net/fig/543217" pageId="0" pageNumber="115">14A, B</figureCitation>
). Sternum VIII with lateral
<normalizedToken originalValue="“larval”">&quot;larval&quot;</normalizedToken>
lanterns, posterior margins medially sinuose or mucronate (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Costalampys delicata sp. nov. A, Abdominal sclerites, dorsal view; B, Abdominal sclerites, ventral view; C, Pygidium, dorsal view; D, Syntergite, dorsal view; E, Sternite IX, ventral view; F, Aedeagus, dorsal view; G, Aedeagus, lateral view; H, Aedeagus, ventral view. Scale bar: 0.5 mm (A-H)." figureDoi="10.3897/asp.79.e67185.figure14" httpUri="https://binary.pensoft.net/fig/543217" pageId="0" pageNumber="115">14A, B</figureCitation>
). Pygidium with posterior margin centrally rounded (Fig.
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Costalampys decorata (Olivier) comb. nov. A, Male habitus, dorsal view; B, Male habitus, ventral view; C, Female habitus, dorsal view; D, Female habitus, ventral view; E, Male antena, lateral view; F, Male pronotum, dorsal view; G, Male terminal segments, ventral view; H, Pygidium, dorsal view; I, Syntergite, dorsal view; J, Sternum IX, Ventral view; K, Aedeagus, dorsal view; L, Aedeagus, lateral view; M, Aedeagus, ventral view. Scale bar: 2 mm (A-D; G-H); Scale bar: 0.5 mm (I-M)." figureDoi="10.3897/asp.79.e67185.figure9" httpUri="https://binary.pensoft.net/fig/543212" pageId="0" pageNumber="115">9H</figureCitation>
) or almost truncate (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Costalampys klugii (Motschulsky) comb. nov. A, Male habitus, dorsal view; B, Male habitus, ventral view; C, Female habitus, dorsal view; D, Female habitus, ventral view; E, Male antenna, lateral view; F, Male pronotum, dorsal view; G, Sternum IX, ventral view; H, Pygidium, dorsal view; I, Syntergite, dorsal view; J, Sternum IX, Ventral view; K, Aedeagus, dorsal view; L, Aedeagus, lateral view; M, Aedeagus, ventral view. Scale bar: 2 mm (A-D); Scale bar: 1.0 mm (E-M)." figureDoi="10.3897/asp.79.e67185.figure17" httpUri="https://binary.pensoft.net/fig/543220" pageId="0" pageNumber="115">17H</figureCitation>
), sometimes posteriorly mucronate (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Costalampys delicata sp. nov. A, Abdominal sclerites, dorsal view; B, Abdominal sclerites, ventral view; C, Pygidium, dorsal view; D, Syntergite, dorsal view; E, Sternite IX, ventral view; F, Aedeagus, dorsal view; G, Aedeagus, lateral view; H, Aedeagus, ventral view. Scale bar: 0.5 mm (A-H)." figureDoi="10.3897/asp.79.e67185.figure14" httpUri="https://binary.pensoft.net/fig/543217" pageId="0" pageNumber="115">14C</figureCitation>
) posterior corners weekly produced or barely present.
<emphasis bold="true" pageId="0" pageNumber="115">Male.</emphasis>
Antennomeres III-X cylindrical and flabellate, with lamellae long and slender, of variable length, apical antennomere at least slightly longer than subapical one (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Costalampys delicata sp. nov., male head. A, Head capsule, dorsal view; B, Head capsule, ventral view; C, Head capsule, lateral view; D, Head capsule, frontal view; E, Head capsule, posterior view; F, Head capsule, occipital view; G, Mandible, dorsal view; H, Mandible, ventral view; I, Tentorium, dorsal view; J, Tentorium, frontal view; K, Tentorium, lateral view; L, antenna, lateral view. Scale bar: 0.5 mm (A-K); Scale bar: 1.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure12" httpUri="https://binary.pensoft.net/fig/543215" pageId="0" pageNumber="115">12L</figureCitation>
). Syntergite consisting of paired lateral plates convergent posteriorly (putatively tergite IX or paraproct), median transversal suture absent, anterior margin mildly to strongly convergent (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Costalampys delicata sp. nov. A, Abdominal sclerites, dorsal view; B, Abdominal sclerites, ventral view; C, Pygidium, dorsal view; D, Syntergite, dorsal view; E, Sternite IX, ventral view; F, Aedeagus, dorsal view; G, Aedeagus, lateral view; H, Aedeagus, ventral view. Scale bar: 0.5 mm (A-H)." figureDoi="10.3897/asp.79.e67185.figure14" httpUri="https://binary.pensoft.net/fig/543217" pageId="0" pageNumber="115">14D, E</figureCitation>
); not posteriorly fused to sternum IX. Sternum IX roughly symmetric, posterior margin rounded. Phallus (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Costalampys delicata sp. nov. A, Abdominal sclerites, dorsal view; B, Abdominal sclerites, ventral view; C, Pygidium, dorsal view; D, Syntergite, dorsal view; E, Sternite IX, ventral view; F, Aedeagus, dorsal view; G, Aedeagus, lateral view; H, Aedeagus, ventral view. Scale bar: 0.5 mm (A-H)." figureDoi="10.3897/asp.79.e67185.figure14" httpUri="https://binary.pensoft.net/fig/543217" pageId="0" pageNumber="115">14F-H</figureCitation>
) with a dorsal and a ventral plate; dorsal plate basally fused to parameres, without a transverse groove apically rounded or acute, not indented), curved dorsally or straight (in lateral view), with a subapical transversal keel; ventral plate elongate, with sides straight. Paramere slightly longer than dorsal plate, with a ventral subapical tooth.
<emphasis bold="true" pageId="0" pageNumber="115">
Female (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Costalampys delicata sp. nov. A, Female habitus, dorsal view; B, Female habitus, ventral view; C, Female antenna, lateral view; D, Sternum VIII, ventral view; E, Pygidium, Dorsal view; F, Ovipositor, dorsal view; G, Ovipositor, lateral view; H, Internal genitalia, dorsal view; I, Internal genitalia, ventral view. Scale bar: 2 mm (A-C); Scale bar: 1.0 mm (D-I)." figureDoi="10.3897/asp.79.e67185.figure15" httpUri="https://binary.pensoft.net/fig/543218" pageId="0" pageNumber="115">15D-I</figureCitation>
).
</emphasis>
Antenna compressed and serrate, with apical corners almost right-angled or projected and acute (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Costalampys delicata sp. nov. A, Female habitus, dorsal view; B, Female habitus, ventral view; C, Female antenna, lateral view; D, Sternum VIII, ventral view; E, Pygidium, Dorsal view; F, Ovipositor, dorsal view; G, Ovipositor, lateral view; H, Internal genitalia, dorsal view; I, Internal genitalia, ventral view. Scale bar: 2 mm (A-C); Scale bar: 1.0 mm (D-I)." figureDoi="10.3897/asp.79.e67185.figure15" httpUri="https://binary.pensoft.net/fig/543218" pageId="0" pageNumber="115">15A-C</figureCitation>
). Sternum VIII as long as wide, spiculum ventrale long and slender, 3/4
<normalizedToken originalValue="×">x</normalizedToken>
as long as sternum (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Costalampys delicata sp. nov. A, Female habitus, dorsal view; B, Female habitus, ventral view; C, Female antenna, lateral view; D, Sternum VIII, ventral view; E, Pygidium, Dorsal view; F, Ovipositor, dorsal view; G, Ovipositor, lateral view; H, Internal genitalia, dorsal view; I, Internal genitalia, ventral view. Scale bar: 2 mm (A-C); Scale bar: 1.0 mm (D-I)." figureDoi="10.3897/asp.79.e67185.figure15" httpUri="https://binary.pensoft.net/fig/543218" pageId="0" pageNumber="115">15D, E</figureCitation>
). Internal genitalia with a slightly sclerotized, elongate spermatophore-digesting gland, almost twice as long and slightly wider than the spermatheca, which is membranous and globose (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Costalampys delicata sp. nov. A, Female habitus, dorsal view; B, Female habitus, ventral view; C, Female antenna, lateral view; D, Sternum VIII, ventral view; E, Pygidium, Dorsal view; F, Ovipositor, dorsal view; G, Ovipositor, lateral view; H, Internal genitalia, dorsal view; I, Internal genitalia, ventral view. Scale bar: 2 mm (A-C); Scale bar: 1.0 mm (D-I)." figureDoi="10.3897/asp.79.e67185.figure15" httpUri="https://binary.pensoft.net/fig/543218" pageId="0" pageNumber="115">15F-I</figureCitation>
). Ovipositor short, valvifers free and slightly curved, slightly over 2
<normalizedToken originalValue="×">x</normalizedToken>
longer than coxite; proximal plates sclerotized, fused medially in a somewhat C-shaped structure (with sides straight and convergent apically, medially straight anteriorly, posterior margin slightly curved), coxites apically free (i.e. not fused), coxital baculi well-developed, sclerotized, apically convergent; styli as long as 1/4
<normalizedToken originalValue="×">x</normalizedToken>
coxite length, well-sclerotized; proctiger well-developed, formed by two elongate plates, convergent apically, laterotergite rudimentary, subquadrate in ventral view, weakly sclerotized, barely attached to valvifers (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Costalampys delicata sp. nov. A, Female habitus, dorsal view; B, Female habitus, ventral view; C, Female antenna, lateral view; D, Sternum VIII, ventral view; E, Pygidium, Dorsal view; F, Ovipositor, dorsal view; G, Ovipositor, lateral view; H, Internal genitalia, dorsal view; I, Internal genitalia, ventral view. Scale bar: 2 mm (A-C); Scale bar: 1.0 mm (D-I)." figureDoi="10.3897/asp.79.e67185.figure15" httpUri="https://binary.pensoft.net/fig/543218" pageId="0" pageNumber="115">15 F,G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="115" type="biology_ecology">
<paragraph pageId="0" pageNumber="115">Biology.</paragraph>
<paragraph pageId="0" pageNumber="115">
Five species of
<taxonomicName authority="Silveira, Roza, Vaz &amp; Mermudes, 2021-14" authorityName="Silveira, Roza, Vaz &amp; Mermudes, 2021-14" class="Insecta" family="Lampyridae" genus="Costalampys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Costalampys" order="Hymenoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Costalampys</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="115">gen. nov.</emphasis>
have been observed active during daytime, namely
<taxonomicName genus="C." lsidName="C. decorata" pageId="0" pageNumber="115" rank="species" species="decorata">
<emphasis italics="true" pageId="0" pageNumber="115">C. decorata</emphasis>
</taxonomicName>
(Olivier, 1888)
<emphasis bold="true" pageId="0" pageNumber="115">comb. nov.</emphasis>
,
<taxonomicName genus="C." lsidName="C. delicata" pageId="0" pageNumber="115" rank="species" species="delicata">
<emphasis italics="true" pageId="0" pageNumber="115">C. delicata</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="115">sp. nov.</emphasis>
,
<taxonomicName genus="C." lsidName="C. joanae" pageId="0" pageNumber="115" rank="species" species="joanae">
<emphasis italics="true" pageId="0" pageNumber="115">C. joanae</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="115">sp. nov.</emphasis>
,
<taxonomicName genus="C." lsidName="C. pauper" pageId="0" pageNumber="115" rank="species" species="pauper">
<emphasis italics="true" pageId="0" pageNumber="115">C. pauper</emphasis>
</taxonomicName>
(Olivier, 1899)
<emphasis bold="true" pageId="0" pageNumber="115">comb. nov.</emphasis>
, and
<taxonomicName genus="C." lsidName="C. tricolor" pageId="0" pageNumber="115" rank="species" species="tricolor">
<emphasis italics="true" pageId="0" pageNumber="115">C. tricolor</emphasis>
</taxonomicName>
(Gorham, 1880)
<emphasis bold="true" pageId="0" pageNumber="115">comb. nov.</emphasis>
Males and females were observed perching on leaves of understory bushes and growing trees (as in Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Costalampys decorata (Olivier) comb. nov., male in natural environment (in Brazil: SP, Votorantim). Photo author: Enio Branco." figureDoi="10.3897/asp.79.e67185.figure10" httpUri="https://binary.pensoft.net/fig/543213" pageId="0" pageNumber="115">10</figureCitation>
). Males will sustain the antennae erect and pointed out towards the wind current, very similar to other firefly taxa with flabellate antennae, like many
<taxonomicName class="Insecta" family="Lampyridae" genus="Ethra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ethra" order="Coleoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Ethra</emphasis>
</taxonomicName>
Laporte, 1833,
<taxonomicName class="Insecta" family="Lampyridae" genus="Amydetes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amydetes" order="Coleoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Amydetes</emphasis>
</taxonomicName>
Illiger, 1807 and
<taxonomicName class="Insecta" family="Lampyridae" genus="Cladodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cladodes" order="Coleoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Cladodes</emphasis>
</taxonomicName>
species (L. Silveira pers. ob.). Given the similarity in overall morphology, it is likely that other
<taxonomicName authority="Silveira, Roza, Vaz &amp; Mermudes, 2021-14" authorityName="Silveira, Roza, Vaz &amp; Mermudes, 2021-14" class="Insecta" family="Lampyridae" genus="Costalampys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Costalampys" order="Hymenoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Costalampys</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="115">gen. nov.</emphasis>
species in the genus are also diurnal. Where observed, lanterns are functional, and individuals will glow if disturbed. Other roles of bioluminescence (e.g., in courtship) are unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="115" type="distribution">
<paragraph pageId="0" pageNumber="115">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="115">In the Atlantic rainforest, with occasional records in adjacent Caatinga patches.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="115" type="remarks">
<paragraph pageId="0" pageNumber="115">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="115">
<taxonomicName authority="Silveira, Roza, Vaz &amp; Mermudes, 2021-14" authorityName="Silveira, Roza, Vaz &amp; Mermudes, 2021-14" class="Insecta" family="Lampyridae" genus="Costalampys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Costalampys" order="Hymenoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Costalampys</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="115">gen. nov.</emphasis>
superficially resembles the neotropical genera
<taxonomicName authorityName="Blanchard" authorityYear="1846" class="Hexapoda" family="Lampyridae" genus="Psilocladus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Psilocladus" order="Coleoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Psilocladus</emphasis>
</taxonomicName>
(type genus of
<taxonomicName authorityName="McDermott" authorityYear="1964" lsidName="" pageId="0" pageNumber="115" rank="subFamily" subFamily="Psilocladinae">Psilocladinae</taxonomicName>
),
<taxonomicName class="Insecta" family="Lampyridae" genus="Amydetes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amydetes" order="Coleoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Amydetes</emphasis>
</taxonomicName>
(type genus of
<taxonomicName authorityName="E.Olivier" authorityYear="1907" lsidName="" pageId="0" pageNumber="115" rank="subFamily" subFamily="Amydetinae">Amydetinae</taxonomicName>
), and
<taxonomicName class="Insecta" family="Lampyridae" genus="Ethra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ethra" order="Coleoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Ethra</emphasis>
</taxonomicName>
(
<taxonomicName lsidName="" pageId="0" pageNumber="115" rank="subFamily" subFamily="Lampyrinae">Lampyrinae</taxonomicName>
) due to the branched antennae.
<taxonomicName authority="Silveira, Roza, Vaz &amp; Mermudes, 2021-14" authorityName="Silveira, Roza, Vaz &amp; Mermudes, 2021-14" class="Insecta" family="Lampyridae" genus="Costalampys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Costalampys" order="Hymenoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Costalampys</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="115">gen. nov.</emphasis>
was found closer to lampyrine taxa in all our analyses, and never clustered with
<taxonomicName class="Insecta" family="Lampyridae" genus="Amydetes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amydetes" order="Coleoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Amydetes</emphasis>
</taxonomicName>
(
<taxonomicName authorityName="E.Olivier" authorityYear="1907" lsidName="" pageId="0" pageNumber="115" rank="subFamily" subFamily="Amydetinae">Amydetinae</taxonomicName>
) or
<taxonomicName authorityName="Blanchard" authorityYear="1846" class="Hexapoda" family="Lampyridae" genus="Psilocladus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Psilocladus" order="Coleoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Psilocladus</emphasis>
</taxonomicName>
(
<taxonomicName authorityName="McDermott" authorityYear="1964" lsidName="" pageId="0" pageNumber="115" rank="subFamily" subFamily="Psilocladinae">Psilocladinae</taxonomicName>
). Therefore,
<taxonomicName authority="Silveira, Roza, Vaz &amp; Mermudes, 2021-14" authorityName="Silveira, Roza, Vaz &amp; Mermudes, 2021-14" class="Insecta" family="Lampyridae" genus="Costalampys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Costalampys" order="Hymenoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Costalampys</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="115">gen. nov.</emphasis>
is here tentatively placed as
<taxonomicName lsidName="" pageId="0" pageNumber="115" rank="subFamily" subFamily="Lampyrinae">Lampyrinae</taxonomicName>
, notwithstanding the lack of definitive diagnostic characters for the
<taxonomicName lsidName="" pageId="0" pageNumber="115" rank="subFamily" subFamily="Lampyrinae">Lampyrinae</taxonomicName>
(cf.
<bibRefCitation author="Jeng, ML" journalOrPublisher="Bioinformatics (Oxford, England)" pageId="0" pageNumber="115" refId="B23" refString="Jeng, ML, 2008. Comprehensive phylogenetics, systematics, and evolution of neoteny of Lampyridae (Insecta: Coleoptera). Ph.D. thesis. University of Kansas." title="Comprehensive phylogenetics, systematics, and evolution of neoteny of Lampyridae (Insecta: Coleoptera). Ph. D. thesis. University of Kansas." year="2008">Jeng 2008</bibRefCitation>
,
<bibRefCitation DOI="https://doi.org/10.1093/isd/ixz024" author="Martin, GJ" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="115" refId="B31" refString="Martin, GJ, Stanger-Hall, KF, Branham, MA, Silveira, LFL, Lower, SE, Hall, DW, Li, X-Y, Lemmon, AR, Moriarty Lemmon, E, Bybee, SM, 2019. Higher-level phylogeny and reclassification of Lampyridae (Coleoptera: Elateroidea). Insect Systematics and Diversity 3. https://doi.org/10.1093/isd/ixz024" title="Higher-level phylogeny and reclassification of Lampyridae (Coleoptera: Elateroidea). Insect Systematics and Diversity 3." url="https://doi.org/10.1093/isd/ixz024" year="2019">Martin et al. 2019</bibRefCitation>
). While the tribal classification within
<taxonomicName lsidName="" pageId="0" pageNumber="115" rank="subFamily" subFamily="Lampyrinae">Lampyrinae</taxonomicName>
remains unsteady (
<bibRefCitation DOI="https://doi.org/10.1093/isd/ixz024" author="Martin, GJ" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="115" refId="B31" refString="Martin, GJ, Stanger-Hall, KF, Branham, MA, Silveira, LFL, Lower, SE, Hall, DW, Li, X-Y, Lemmon, AR, Moriarty Lemmon, E, Bybee, SM, 2019. Higher-level phylogeny and reclassification of Lampyridae (Coleoptera: Elateroidea). Insect Systematics and Diversity 3. https://doi.org/10.1093/isd/ixz024" title="Higher-level phylogeny and reclassification of Lampyridae (Coleoptera: Elateroidea). Insect Systematics and Diversity 3." url="https://doi.org/10.1093/isd/ixz024" year="2019">Martin et al. 2019</bibRefCitation>
), we place
<taxonomicName authority="Silveira, Roza, Vaz &amp; Mermudes, 2021-14" authorityName="Silveira, Roza, Vaz &amp; Mermudes, 2021-14" class="Insecta" family="Lampyridae" genus="Costalampys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Costalampys" order="Hymenoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Costalampys</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="115">gen. nov.</emphasis>
as
<emphasis italics="true" pageId="0" pageNumber="115">incertae sedis</emphasis>
within the subfamily.
</paragraph>
<paragraph pageId="0" pageNumber="115">
Males of
<taxonomicName authority="Silveira, Roza, Vaz &amp; Mermudes, 2021-14" authorityName="Silveira, Roza, Vaz &amp; Mermudes, 2021-14" class="Insecta" family="Lampyridae" genus="Costalampys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Costalampys" order="Hymenoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Costalampys</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="115">gen. nov.</emphasis>
especially resemble those of
<taxonomicName class="Insecta" family="Lampyridae" genus="Ethra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ethra" order="Coleoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Ethra</emphasis>
</taxonomicName>
(
<taxonomicName lsidName="" pageId="0" pageNumber="115" rank="subFamily" subFamily="Lampyrinae">Lampyrinae</taxonomicName>
), and specimens we now recognise as
<taxonomicName authority="Silveira, Roza, Vaz &amp; Mermudes, 2021-14" authorityName="Silveira, Roza, Vaz &amp; Mermudes, 2021-14" class="Insecta" family="Lampyridae" genus="Costalampys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Costalampys" order="Hymenoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Costalampys</emphasis>
</taxonomicName>
were often found in collections identified as
<taxonomicName class="Insecta" family="Lampyridae" genus="Ethra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ethra" order="Coleoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Ethra</emphasis>
</taxonomicName>
(L. Silveira, unpublished results). However,
<taxonomicName authority="Silveira, Roza, Vaz &amp; Mermudes, 2021-14" authorityName="Silveira, Roza, Vaz &amp; Mermudes, 2021-14" class="Insecta" family="Lampyridae" genus="Costalampys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Costalampys" order="Hymenoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Costalampys</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="115">gen. nov.</emphasis>
can be distinguished from
<taxonomicName class="Insecta" family="Lampyridae" genus="Ethra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ethra" order="Coleoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Ethra</emphasis>
</taxonomicName>
by: antennal lamellae (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Costalampys delicata sp. nov., male head. A, Head capsule, dorsal view; B, Head capsule, ventral view; C, Head capsule, lateral view; D, Head capsule, frontal view; E, Head capsule, posterior view; F, Head capsule, occipital view; G, Mandible, dorsal view; H, Mandible, ventral view; I, Tentorium, dorsal view; J, Tentorium, frontal view; K, Tentorium, lateral view; L, antenna, lateral view. Scale bar: 0.5 mm (A-K); Scale bar: 1.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure12" httpUri="https://binary.pensoft.net/fig/543215" pageId="0" pageNumber="115">12L</figureCitation>
) basally inserted (apically inserted in
<taxonomicName class="Insecta" family="Lampyridae" genus="Ethra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ethra" order="Coleoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Ethra</emphasis>
</taxonomicName>
); labial palpi (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Costalampys delicata sp. nov., male head. A, Head capsule, dorsal view; B, Head capsule, ventral view; C, Head capsule, lateral view; D, Head capsule, frontal view; E, Head capsule, posterior view; F, Head capsule, occipital view; G, Mandible, dorsal view; H, Mandible, ventral view; I, Tentorium, dorsal view; J, Tentorium, frontal view; K, Tentorium, lateral view; L, antenna, lateral view. Scale bar: 0.5 mm (A-K); Scale bar: 1.0 mm (L)." figureDoi="10.3897/asp.79.e67185.figure12" httpUri="https://binary.pensoft.net/fig/543215" pageId="0" pageNumber="115">12A-F</figureCitation>
) with sides apically divergent (parallel-sided in
<taxonomicName class="Insecta" family="Lampyridae" genus="Ethra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ethra" order="Coleoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Ethra</emphasis>
</taxonomicName>
); and pygidium with sides rounded (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Costalampys delicata sp. nov. A, Abdominal sclerites, dorsal view; B, Abdominal sclerites, ventral view; C, Pygidium, dorsal view; D, Syntergite, dorsal view; E, Sternite IX, ventral view; F, Aedeagus, dorsal view; G, Aedeagus, lateral view; H, Aedeagus, ventral view. Scale bar: 0.5 mm (A-H)." figureDoi="10.3897/asp.79.e67185.figure14" httpUri="https://binary.pensoft.net/fig/543217" pageId="0" pageNumber="115">14C</figureCitation>
) (straight in
<taxonomicName class="Insecta" family="Lampyridae" genus="Ethra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ethra" order="Coleoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Ethra</emphasis>
</taxonomicName>
). Females of all
<taxonomicName class="Insecta" family="Lampyridae" genus="Ethra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ethra" order="Coleoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Ethra</emphasis>
</taxonomicName>
spp. remain unknown.
</paragraph>
<paragraph pageId="0" pageNumber="115">
<taxonomicName authority="Silveira, Roza, Vaz &amp; Mermudes, 2021-14" authorityName="Silveira, Roza, Vaz &amp; Mermudes, 2021-14" class="Insecta" family="Lampyridae" genus="Costalampys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Costalampys" order="Hymenoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Costalampys</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="115">gen. nov</emphasis>
. shares several similarities with
<taxonomicName class="Insecta" family="Lampyridae" genus="Scissicauda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scissicauda" order="Coleoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Scissicauda</emphasis>
</taxonomicName>
<bibRefCitation author="McDermott, FA" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="115" refId="B33" refString="McDermott, FA, 1964. The taxonomy of the Lampyridae (Coleoptera). Transactions of the American Entomological Society (1890-) 90: 1-72." title="The taxonomy of the Lampyridae (Coleoptera). Transactions of the American Entomological Society (1890 -) 90: 1 - 72." year="1964">McDermott 1964</bibRefCitation>
, such as: antennal sockets large and close-set; maxillary palp with palpomere IV large and fusiform, submentum cordiform, pronotum semilunar, proendosternite apically clavate, elytral outline ellipsoid; metaendostenrite elongate, rhomboid; abdominal sternum II bearing paired vitreous spots medially; aedeagus with phallus bearing dorsal and ventral plates, dorsal plate enlarged apically, projecting ventrally and partially embracing the internal sac. It is also noteworthy that the genotypic
<taxonomicName genus="S." lsidName="S. disjuncta" pageId="0" pageNumber="115" rank="species" species="disjuncta">
<emphasis italics="true" pageId="0" pageNumber="115">S. disjuncta</emphasis>
</taxonomicName>
also has basally flabellate antennae, although its bristles are long and erect (
<bibRefCitation DOI="https://doi.org/10.1080/14772000.2016.1153006" author="Silveira, LFL" journalOrPublisher="Systematics and Biodiversity" pageId="0" pageNumber="115" pagination="371 - 384" refId="B66" refString="Silveira, LFL, Khattar, G, Souto, P, Mermudes, JRM, Takiya, DM, Monteiro, RF, 2016b. Integrative taxonomy of new firefly taxa from the Atlantic Rainforest. Systematics and Biodiversity 14: 371 - 384, DOI: https://doi.org/10.1080/14772000.2016.1153006" title="Integrative taxonomy of new firefly taxa from the Atlantic Rainforest." url="https://doi.org/10.1080/14772000.2016.1153006" volume="14" year="2016 b">Silveira et al. 2016b</bibRefCitation>
). However,
<taxonomicName authority="Silveira, Roza, Vaz &amp; Mermudes, 2021-14" authorityName="Silveira, Roza, Vaz &amp; Mermudes, 2021-14" class="Insecta" family="Lampyridae" genus="Costalampys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Costalampys" order="Hymenoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Costalampys</emphasis>
</taxonomicName>
differs from
<taxonomicName class="Insecta" family="Lampyridae" genus="Scissicauda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scissicauda" order="Coleoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Scissicauda</emphasis>
</taxonomicName>
by: pronotal disc strongly convex in lateral view (almost flat in
<taxonomicName class="Insecta" family="Lampyridae" genus="Scissicauda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scissicauda" order="Coleoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Scissicauda</emphasis>
</taxonomicName>
); hypomeron strongly sinuose (mildly sinuose in
<taxonomicName class="Insecta" family="Lampyridae" genus="Scissicauda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scissicauda" order="Coleoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Scissicauda</emphasis>
</taxonomicName>
); sternum VIII as long as VII, posterior margin medially mucronate or slightly sinuose (at least a 1/5
<normalizedToken originalValue="×">x</normalizedToken>
longer, and posterior margin strongly sinuose in
<taxonomicName class="Insecta" family="Lampyridae" genus="Scissicauda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scissicauda" order="Coleoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Scissicauda</emphasis>
</taxonomicName>
); sternum IX visible (concealed in
<taxonomicName class="Insecta" family="Lampyridae" genus="Scissicauda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scissicauda" order="Coleoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Scissicauda</emphasis>
</taxonomicName>
); male pygidium rounded posteriorly, sometimes mucronate (with sides divergent and posterior margin strongly indented in
<taxonomicName class="Insecta" family="Lampyridae" genus="Scissicauda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scissicauda" order="Coleoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Scissicauda</emphasis>
</taxonomicName>
); phallus with dorsal plate without a transverse medial groove (present in in
<taxonomicName class="Insecta" family="Lampyridae" genus="Scissicauda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scissicauda" order="Coleoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Scissicauda</emphasis>
</taxonomicName>
); internal female genitalia with spermatheca (absent in
<taxonomicName class="Insecta" family="Lampyridae" genus="Scissicauda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scissicauda" order="Coleoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Scissicauda</emphasis>
</taxonomicName>
[based on the type species
<taxonomicName genus="S." lsidName="S. disjuncta" pageId="0" pageNumber="115" rank="species" species="disjuncta">
<emphasis italics="true" pageId="0" pageNumber="115">S. disjuncta</emphasis>
</taxonomicName>
]), ovipositor in
<taxonomicName class="Insecta" family="Lampyridae" genus="Scissicauda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Scissicauda" order="Coleoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="115">Scissicauda</emphasis>
</taxonomicName>
[based on the type species
<taxonomicName genus="S." lsidName="S. disjuncta" pageId="0" pageNumber="115" rank="species" species="disjuncta">
<emphasis italics="true" pageId="0" pageNumber="115">S. disjuncta</emphasis>
</taxonomicName>
]).
</paragraph>
</subSubSection>
</treatment>
</document>
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