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<document id="D46E7EB3C96531EF2429202707E1ACAB" ID-DOI="10.5281/zenodo.5415296" ID-ISSN="0253-116X" ID-Zenodo-Dep="5415296" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630357985480" checkinUser="marcus" docAuthor="Smetana, Aleš" docDate="2015" docId="074E878C744DFF92FF4C8D52DE724131" docLanguage="en" docName="LinzerbiolBeitr.2015.47.1.897-903.pdf" docOrigin="Linzer biologische Beiträge 47 (1)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Indoquedius frater Smetana 2015, nov.sp." docType="treatment" docVersion="4" lastPageNumber="900" masterDocId="FB77FFF4744FFF91FFFE8C07DF764223" masterDocTitle="Contributions to the knowledge of the Quediina (Coleoptera: Staphylinidae: Staphylinini) of China. Part 53. Genus Indoquedius BLACKWELDER, 1952. Section 2" masterLastPageNumber="903" masterPageNumber="897" pageNumber="899" updateTime="1699246318473" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="C524C208106FF354921DE502F0EF5D89">Contributions to the knowledge of the Quediina (Coleoptera: Staphylinidae: Staphylinini) of China. Part 53. Genus Indoquedius BLACKWELDER, 1952. Section 2</mods:title>
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<mods:namePart id="582C9EA56B35968191CD1B38818052B7">Smetana, Aleš</mods:namePart>
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<subSubSection id="C7FD6511744DFF93FF4C8D52DC8243BA" pageId="2" pageNumber="899" type="nomenclature">
<paragraph id="8F58369A744DFF93FF4C8D52DD424348" blockId="2.[178,564,341,363]" box="[178,564,341,363]" pageId="2" pageNumber="899">
<heading id="D41081F6744DFF93FF4C8D52DD424348" box="[178,564,341,363]" fontSize="8" level="2" pageId="2" pageNumber="899" reason="2">
<emphasis id="BD93EA88744DFF93FF4C8D52DD424348" bold="true" box="[178,564,341,363]" pageId="2" pageNumber="899">
<taxonomicName id="48E74D19744DFF93FF4C8D52DE004348" ID-CoL="3PNK3" authority="Smetana, 2015" authorityName="Smetana" authorityYear="2015" box="[178,374,341,363]" class="Insecta" family="Staphylinidae" genus="Indoquedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="899" phylum="Arthropoda" rank="species" species="frater" status="sp. nov.">
<emphasis id="BD93EA88744DFF93FF4C8D52DE004348" bold="true" box="[178,374,341,363]" italics="true" pageId="2" pageNumber="899">Indoquedius frater</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A6A057F3744DFF93FE828D51DEBF4348" box="[380,457,342,363]" pageId="2" pageNumber="899" rank="species">nov.sp.</taxonomicNameLabel>
(
<figureCitation id="17DC2A1F744DFF93FE2B8D52DD5A4348" box="[469,556,341,363]" captionStart="Figs 1-6" captionStartId="6.[178,218,1203,1225]" captionTargetBox="[178,1082,223,1191]" captionTargetId="figure-1@6.[178,1084,220,1194]" captionTargetPageId="6" captionText="Figs 1-6: Indoquedius frater sp. nov. (1) apical portion of male sternite 8; (2) tergite 10 of male genital segment; (3) sternite 9 of male genital segment; (4) aedoeagus, ventral view; (5) apical portion of ventral side of paramere with sensory peg setae; (6) tergite 10 of female genital segment." figureDoi="http://doi.org/10.5281/zenodo.5415298" httpUri="https://zenodo.org/record/5415298/files/figure.png" pageId="2" pageNumber="899">Figs 1-6</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="8F58369A744DFF93FF4C8D83DC8243BA" blockId="2.[178,1085,387,489]" box="[178,1012,387,410]" pageId="2" pageNumber="899">
T y p e l o c a l i t y.
<collectingCountry id="F7F0760A744DFF93FE5B8D83DEAB43B9" box="[421,477,388,410]" name="China" pageId="2" pageNumber="899">China</collectingCountry>
:
<collectingRegion id="4D23F878744DFF93FE1A8D83DD5B43BA" box="[484,557,388,409]" country="China" name="Yunnan" pageId="2" pageNumber="899">Yunnan</collectingRegion>
: SE Pingbian,
<geoCoordinate id="EAD3505D744DFF93FD558D83DC5643BA" box="[683,800,387,409]" degrees="22" direction="north" minutes="54" orientation="latitude" pageId="2" pageNumber="899" precision="15" seconds="31" value="22.908611">22° 54’31’’N</geoCoordinate>
<geoCoordinate id="EAD3505D744DFF93FCDA8D83DCD543BA" box="[804,931,388,409]" degrees="103" direction="east" minutes="41" orientation="longitude" pageId="2" pageNumber="899" precision="15" seconds="44" value="103.69556">103° 41’44’’E</geoCoordinate>
,
<quantity id="481F9B7F744DFF93FC528D83DC8643BA" box="[940,1008,388,409]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="2" pageNumber="899" unit="m" value="2100.0">2100 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C7FD6511744DFF92FF4C8DA3DE724131" lastPageId="3" lastPageNumber="900" pageId="2" pageNumber="899" type="description">
<paragraph id="8F58369A744DFF93FF4C8DA3DEB743CA" blockId="2.[178,1085,387,489]" pageId="2" pageNumber="899">
T y p e m a t e r i a l.
<typeStatus id="505C8838744DFF93FE4B8DA3DD704399" box="[437,518,420,442]" pageId="2" pageNumber="899" type="holotype">Holotype</typeStatus>
(Ƌ) and
<typeStatus id="505C8838744DFF93FD9A8DA3DDDC4399" box="[612,682,420,442]" pageId="2" pageNumber="899" type="allotype">allotype</typeStatus>
(
<specimenCount id="99E1FD13744DFF93FD448DAEDDBB439D" box="[698,717,425,446]" pageId="2" pageNumber="899" type="female">♀</specimenCount>
):
<collectingCountry id="F7F0760A744DFF93FD1D8DA3DC5A439A" box="[739,812,420,441]" name="China" pageId="2" pageNumber="899">CHINA</collectingCountry>
: &quot;
<collectingCountry id="F7F0760A744DFF93FCC18DA3DCF1439A" box="[831,903,420,441]" name="China" pageId="2" pageNumber="899">CHINA</collectingCountry>
[22a]-
<collectingRegion id="4D23F878744DFF93FC318DA3DB61439A" box="[975,1047,420,441]" country="China" name="Yunnan" pageId="2" pageNumber="899">Yunnan</collectingRegion>
, SE Pingbian, primary forest,
<geoCoordinate id="EAD3505D744DFF93FE438DBBDD4443F2" box="[445,562,443,465]" degrees="22" direction="north" minutes="54" orientation="latitude" pageId="2" pageNumber="899" precision="15" seconds="31" value="22.908611">22° 54’31’’N</geoCoordinate>
<geoCoordinate id="EAD3505D744DFF93FDC98DBBDDC343F2" box="[567,693,444,465]" degrees="103" direction="east" minutes="41" orientation="longitude" pageId="2" pageNumber="899" precision="15" seconds="44" value="103.69556">103° 41’44’’E</geoCoordinate>
,
<quantity id="481F9B7F744DFF93FD418DBBDC7543F2" box="[703,771,444,465]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="2" pageNumber="899" unit="m" value="2100.0">2100 m</quantity>
,
<date id="FB59105A744DFF93FCF28DBBDC0B43F2" box="[780,893,444,465]" pageId="2" pageNumber="899" value="2014-08-28">28.VIII.2014</date>
V. Assing&quot;.
<typeStatus id="505C8838744DFF93FC128DBCDB4B43F2" box="[1004,1085,443,465]" pageId="2" pageNumber="899" type="holotype">Holotype</typeStatus>
in ASC,
<typeStatus id="505C8838744DFF93FED58DD4DE0743CA" box="[299,369,467,489]" pageId="2" pageNumber="899" type="allotype">allotype</typeStatus>
in VAC.
</paragraph>
<paragraph id="8F58369A744DFF93FF4C8DFFDB734707" blockId="2.[178,1086,504,1316]" pageId="2" pageNumber="899">D e s c r i p t i o n. Body entirely black; maxillary and labial palpi piceous-black, with last segment paler, antennae black, with last four segments milky white., legs black with paler tarsi. Head rounded, slightly wider than long (ratio 1.17); eyes very large and convex, tempora about half as long as eyes seen from above (ratio 0.50); two punctures along medial margin of each eye between anterior and posterior frontal punctures; posterior frontal puncture situated close to posteriomedial margin of eye, separated from it by distance slightly shorter than diameter of puncture; one puncture between it and posterior margin of head; temporal puncture small, situated closer to posterior margin of eye than to posterior margin of head, two fine punctures posteriomediad of it, tempora impunctate; surface of head without microsculpture. Antenna long, hardly thickened toward apex, segment 3 markedly longer than segment 2 (ratio 1.57), following segments distinctly longer than wide but becoming gradually shorter toward antennal apex, last segment asymmetrically acuminate, slightly longer than preceding segment. Pronotum feebly wider than at midline long (ratio 1.11), broadly rounded at base, narrowed anteriad, transversely convex, lateral pronotal groove smooth, without microsculpture; dorsal rows each with three punctures, sublateral rows absent; large lateral puncture separated from lateral pronotal groove by distance about equal to diameter of puncture; anterior angles of pronotum impunctate; surface of pronotum without microsculpture. Scutellum large, coarsely and densely punctuate and setose. Elytra moderately long, at base slightly narrower than pronotum at widest point, at suture vaguely (ratio 1.08), at sides slightly (ratio 1.12) longer than pronotum at midline; punctuation coarse and dense, transverse interspaces between punctures shorter than diameters of punctures, surface between punctures without microsculpture. Wings fully developed. Abdomen with tergite 7 (fifth visible) with distinct whitish apical seam of palisade fringe; tergite 2 (in front of first fully visible tergite) impunctate; punctuation of abdominal tergites considerably finer than that on elytra, moderately dense, about evenly covering each tergite, but becoming gradually sparser toward apex of abdomen; pubescence black; surface between punctures with excessively fine microsculpture of transverse striae.</paragraph>
<paragraph id="8F58369A744DFF92FF4C8933DD7B432D" blockId="2.[178,1086,1332,1588]" lastBlockId="3.[178,1085,219,270]" lastPageId="3" lastPageNumber="900" pageId="2" pageNumber="899">
Male. First four segments of front tarsus markedly dilated, subbilobed, each densely covered with tenent setae ventrally; segment 2 wider than apex of tibia (ratio 1. 21), segment 4 narrower than preceding segments. Sternite 8 with one long seta at each side, with inconspicuous, shallow. arcuate medioapical emargiantion, small triangular area before emargination flattened and smooth (
<figureCitation id="17DC2A1F744DFF93FD9589AEDDD1479C" box="[619,679,1449,1471]" captionStart="Figs 1-6" captionStartId="6.[178,218,1203,1225]" captionTargetBox="[178,1082,223,1191]" captionTargetId="figure-1@6.[178,1084,220,1194]" captionTargetPageId="6" captionText="Figs 1-6: Indoquedius frater sp. nov. (1) apical portion of male sternite 8; (2) tergite 10 of male genital segment; (3) sternite 9 of male genital segment; (4) aedoeagus, ventral view; (5) apical portion of ventral side of paramere with sensory peg setae; (6) tergite 10 of female genital segment." figureDoi="http://doi.org/10.5281/zenodo.5415298" httpUri="https://zenodo.org/record/5415298/files/figure.png" pageId="2" pageNumber="899">Fig. 1</figureCitation>
). Genital segment with tergite 10 wide, markedly narrowed toward emarginated apex, setose as in
<figureCitation id="17DC2A1F744DFF93FCE289C1DC2A47F8" box="[796,860,1478,1500]" captionStart="Figs 1-6" captionStartId="6.[178,218,1203,1225]" captionTargetBox="[178,1082,223,1191]" captionTargetId="figure-1@6.[178,1084,220,1194]" captionTargetPageId="6" captionText="Figs 1-6: Indoquedius frater sp. nov. (1) apical portion of male sternite 8; (2) tergite 10 of male genital segment; (3) sternite 9 of male genital segment; (4) aedoeagus, ventral view; (5) apical portion of ventral side of paramere with sensory peg setae; (6) tergite 10 of female genital segment." figureDoi="http://doi.org/10.5281/zenodo.5415298" httpUri="https://zenodo.org/record/5415298/files/figure.png" pageId="2" pageNumber="899">Fig. 2</figureCitation>
; sternite 9 with short basal portion, apical portion deeply emarginated apically, with transparent &quot;mirror&quot; at each lateral margin, setose as in
<figureCitation id="17DC2A1F744DFF93FDFD8A07DD344435" box="[515,578,1536,1558]" captionStart="Figs 1-6" captionStartId="6.[178,218,1203,1225]" captionTargetBox="[178,1082,223,1191]" captionTargetId="figure-1@6.[178,1084,220,1194]" captionTargetPageId="6" captionText="Figs 1-6: Indoquedius frater sp. nov. (1) apical portion of male sternite 8; (2) tergite 10 of male genital segment; (3) sternite 9 of male genital segment; (4) aedoeagus, ventral view; (5) apical portion of ventral side of paramere with sensory peg setae; (6) tergite 10 of female genital segment." figureDoi="http://doi.org/10.5281/zenodo.5415298" httpUri="https://zenodo.org/record/5415298/files/figure.png" pageId="2" pageNumber="899">Fig. 3</figureCitation>
. Aedoeagus (
<figureCitation id="17DC2A1F744DFF93FD338A07DC464435" box="[717,816,1536,1558]" captionStart="Figs 1-6" captionStartId="6.[178,218,1203,1225]" captionTargetBox="[178,1082,223,1191]" captionTargetId="figure-1@6.[178,1084,220,1194]" captionTargetPageId="6" captionText="Figs 1-6: Indoquedius frater sp. nov. (1) apical portion of male sternite 8; (2) tergite 10 of male genital segment; (3) sternite 9 of male genital segment; (4) aedoeagus, ventral view; (5) apical portion of ventral side of paramere with sensory peg setae; (6) tergite 10 of female genital segment." figureDoi="http://doi.org/10.5281/zenodo.5415298" httpUri="https://zenodo.org/record/5415298/files/figure.png" pageId="2" pageNumber="899">Figs. 4, 5</figureCitation>
) quite similar to that of
<taxonomicName id="48E74D19744DFF93FBD08A06DF8D4410" authorityName="SMETANA" authorityYear="2014" class="Insecta" family="Staphylinidae" genus="Indoquedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="899" phylum="Arthropoda" rank="species" species="jendeki">
<emphasis id="BD93EA88744DFF93FBD08A06DF8D4410" italics="true" pageId="2" pageNumber="899">I. jendeki</emphasis>
</taxonomicName>
, but markedly larger and more elongate, paramere markedly longer, particularly the subparallelsided apical portion; internal sac similar to that of
<taxonomicName id="48E74D19744CFF92FCBB8CDCDCD542D3" authorityName="SMETANA" authorityYear="2014" box="[837,931,219,240]" class="Insecta" family="Staphylinidae" genus="Indoquedius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="900" phylum="Arthropoda" rank="species" species="jendeki">
<emphasis id="BD93EA88744CFF92FCBB8CDCDCD542D3" box="[837,931,219,240]" italics="true" pageId="3" pageNumber="900">I. jendeki</emphasis>
</taxonomicName>
, but composed of much finer spine-like elements.
</paragraph>
<paragraph id="8F58369A744CFF92FF4C8D1ADD1D434D" blockId="3.[178,1085,285,366]" pageId="3" pageNumber="900">
Female. First four segments of front tarsus markedly dilated, not appreciably different from those of male. Genital segment with tergite 10 wide, markedly narrowed toward deeply emarginate apex, setose as in
<figureCitation id="17DC2A1F744CFF92FDD78D5FDD10434D" box="[553,614,344,366]" captionStart="Figs 1-6" captionStartId="6.[178,218,1203,1225]" captionTargetBox="[178,1082,223,1191]" captionTargetId="figure-1@6.[178,1084,220,1194]" captionTargetPageId="6" captionText="Figs 1-6: Indoquedius frater sp. nov. (1) apical portion of male sternite 8; (2) tergite 10 of male genital segment; (3) sternite 9 of male genital segment; (4) aedoeagus, ventral view; (5) apical portion of ventral side of paramere with sensory peg setae; (6) tergite 10 of female genital segment." figureDoi="http://doi.org/10.5281/zenodo.5415298" httpUri="https://zenodo.org/record/5415298/files/figure.png" pageId="3" pageNumber="900">Fig. 6</figureCitation>
.
</paragraph>
<paragraph id="8F58369A744CFF92FF4C8D7ADEE043B0" blockId="3.[178,406,381,403]" box="[178,406,381,403]" pageId="3" pageNumber="900">
Length 12.0-
<quantity id="481F9B7F744CFF92FECA8D7ADEE443B0" box="[308,402,381,403]" metricMagnitude="-2" metricUnit="m" metricValue="1.24" pageId="3" pageNumber="900" unit="mm" value="12.4">12.4 mm</quantity>
.
</paragraph>
<paragraph id="8F58369A744CFF92FF4C8DA4DC1143F6" blockId="3.[178,1085,418,470]" pageId="3" pageNumber="900">
E t y m o l o g y. The specific epithet is the Latin noun
<emphasis id="BD93EA88744CFF92FCE08DA5DC214394" box="[798,855,418,439]" italics="true" pageId="3" pageNumber="900">frater</emphasis>
, -
<emphasis id="BD93EA88744CFF92FC8C8DA4DCE2439B" box="[882,916,419,440]" italics="true" pageId="3" pageNumber="900">tris</emphasis>
, m (brother) in apposition. It denotes the close similarity of the species to
<taxonomicName id="48E74D19744CFF92FCFC8DC7DC1743F6" authorityName="SMETANA" authorityYear="2014" box="[770,865,448,469]" class="Insecta" family="Staphylinidae" genus="Indoquedius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="900" phylum="Arthropoda" rank="species" species="jendeki">
<emphasis id="BD93EA88744CFF92FCFC8DC7DC1743F6" box="[770,865,448,469]" italics="true" pageId="3" pageNumber="900">I. jendeki</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8F58369A744CFF92FF4C8DE2DDE0403B" blockId="3.[178,1085,485,536]" pageId="3" pageNumber="900">
G e o g r a p h i c a l D i s t r i b u t i o n.
<taxonomicName id="48E74D19744CFF92FD698DE2DC2D43D9" authorityName="Smetana" authorityYear="2015" box="[663,859,485,506]" class="Insecta" family="Staphylinidae" genus="Indoquedius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="900" phylum="Arthropoda" rank="species" species="frater">
<emphasis id="BD93EA88744CFF92FD698DE2DC2D43D9" box="[663,859,485,506]" italics="true" pageId="3" pageNumber="900">Indoquedius frater</emphasis>
</taxonomicName>
is at present known only from the
<typeStatus id="505C8838744CFF92FEBC8E04DE18403B" box="[322,366,515,536]" pageId="3" pageNumber="900">type</typeStatus>
locality in southern
<collectingRegion id="4D23F878744CFF92FDC08E04DDE7403B" box="[574,657,515,536]" country="China" name="Yunnan" pageId="3" pageNumber="900">Yunnan</collectingRegion>
.
</paragraph>
<paragraph id="8F58369A744CFF92FF4C8E2FDD6F4078" blockId="3.[178,1085,552,603]" pageId="3" pageNumber="900">
B i o n o m i c s. The
<specimenCount id="99E1FD13744CFF92FE548E2FDD35401D" box="[426,579,552,574]" pageId="3" pageNumber="900" type="generic">two specimens</specimenCount>
of the original series were taken in a &quot;primary forest&quot;, but no details are available.
</paragraph>
<paragraph id="8F58369A744CFF92FF4C8E6CDE724131" blockId="3.[178,1085,618,786]" pageId="3" pageNumber="900">
R e c o g n i t i o n a n d c o m m e n t s.
<taxonomicName id="48E74D19744CFF92FD5E8E6DDC17405C" authorityName="Smetana" authorityYear="2015" box="[672,865,618,639]" class="Insecta" family="Staphylinidae" genus="Indoquedius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="900" phylum="Arthropoda" rank="species" species="frater">
<emphasis id="BD93EA88744CFF92FD5E8E6DDC17405C" box="[672,865,618,639]" italics="true" pageId="3" pageNumber="900">Indoquedius frater</emphasis>
</taxonomicName>
is obviously closely related and similar to
<taxonomicName id="48E74D19744CFF92FE6B8E8FDE8340BF" authorityName="SMETANA" authorityYear="2014" box="[405,501,647,669]" class="Insecta" family="Staphylinidae" genus="Indoquedius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="900" phylum="Arthropoda" rank="species" species="jendeki">
<emphasis id="BD93EA88744CFF92FE6B8E8FDE8340BF" box="[405,501,647,669]" italics="true" pageId="3" pageNumber="900">I. jendeki</emphasis>
</taxonomicName>
, as documented by the similar sclerites of both the male and female genital segments, and by the shape of the aedoeagus, but it may be externally easily distinguished by the uniformly deep black coloration of the body, by the more robust body shape, and by the coarser punctuation of both the elytra and abdominal tergites.
</paragraph>
</subSubSection>
</treatment>
</document>
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