treatments-xml/data/AD/52/3B/AD523BFAF6185B938D3AC34CFE592D15.xml
2024-06-21 12:47:39 +02:00

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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.911.48703" ID-GBIF-Dataset="f1f5ae71-a561-482f-b674-1fadfddcd51d" ID-PMC="PMC7031386" ID-Pensoft-Pub="1313-2970-911-51" ID-Pensoft-UUID="0384A7E91F0C5CAE89798D12AE96B1BD" ID-PubMed="32104138" ID-ZooBank="377165315DC8498EA6358C128EB345AA" ModsDocID="1313-2970-911-51" checkinTime="1581612239756" checkinUser="pensoft" docAuthor="Yu, Li, Liu, Fengxiang, Zhang, Zengtao, Wang, Yan, Li, Daiqin &amp; Xu, Xin" docDate="2020" docId="AD523BFAF6185B938D3AC34CFE592D15" docLanguage="en" docName="ZooKeys 911: 51-66" docOrigin="ZooKeys 911" docSource="http://dx.doi.org/10.3897/zookeys.911.48703" docTitle="Qiongthela baoting Yu &amp; Liu &amp; Zhang &amp; Wang &amp; Li &amp; Xu 2020, sp. nov." docType="treatment" docUuid="C104261D-DBFB-4A70-84FD-5CF0BD15B82E" docUuidSource="ZooBank" docVersion="4" id="0384A7E91F0C5CAE89798D12AE96B1BD" lastPageNumber="51" masterDocId="0384A7E91F0C5CAE89798D12AE96B1BD" masterDocTitle="Four new species of the primitively segmented spider genus Qiongthela from Hainan Island, China (Mesothelae, Liphistiidae)" masterLastPageNumber="66" masterPageNumber="51" pageNumber="51" updateTime="1668168443875" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Four new species of the primitively segmented spider genus Qiongthela from Hainan Island, China (Mesothelae, Liphistiidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yu, Li</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liu, Fengxiang</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhang, Zengtao</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wang, Yan</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Daiqin</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Xu, Xin</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>911</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>51</mods:start>
<mods:end>66</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.911.48703</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.911.48703</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-911-51</mods:identifier>
<mods:identifier type="ZooBank">377165315DC8498EA6358C128EB345AA</mods:identifier>
<mods:identifier type="Pensoft-UUID">0384A7E91F0C5CAE89798D12AE96B1BD</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="162136152" LSID="urn:lsid:zoobank.org:act:C104261D-DBFB-4A70-84FD-5CF0BD15B82E" httpUri="http://treatment.plazi.org/id/AD523BFAF6185B938D3AC34CFE592D15" lastPageNumber="51" pageId="0" pageNumber="51">
<subSubSection pageId="0" pageNumber="51" type="nomenclature">
<paragraph pageId="0" pageNumber="51">
<taxonomicName LSID="AD523BFA-F618-5B93-8D3A-C34CFE592D15" authority="Yu &amp; Liu &amp; Zhang &amp; Wang &amp; Li &amp; Xu, 2020" authorityName="Yu &amp; Liu &amp; Zhang &amp; Wang &amp; Li &amp; Xu" authorityYear="2020" class="Arachnida" family="Liphistiidae" genus="Qiongthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Qiongthela baoting" order="Araneae" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="baoting" status="sp. nov.">Qiongthela baoting</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="51">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male and female genital anatomy of Qiongthela baoting sp. nov. A palp prolateral view B palp ventral view C palp retrolateral view D-F palp distal view G vulva dorsal view H vulva ventral view. A-F XUX- 2017 - 195 (holotype) G-H XUX- 2017 - 196. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.911.48703.figure2" httpUri="https://binary.pensoft.net/fig/379565" pageId="0" pageNumber="51">Figure 2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="type material">
<paragraph pageId="0" pageNumber="51">Type material.</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" pageId="0" pageNumber="51">Holotype</emphasis>
: CHINA · 1 ♂; Hainan Province, Baoting County, Maogan Town, Zaye Village;
<geoCoordinate degrees="18.60" direction="north" orientation="latitude" precision="555" value="18.6">18.60°N</geoCoordinate>
,
<geoCoordinate degrees="109.57" direction="east" orientation="longitude" precision="555" value="109.57">109.57°E</geoCoordinate>
; alt. 410 m; 21 August 2017; D. Li, F.X. Liu and X. Xu leg.; XUX-2017-195 (matured on 25 August 2018 at HNU).
<emphasis bold="true" pageId="0" pageNumber="51">Paratype</emphasis>
: CHINA · 1 ♀; same data as for holotype; XUX-2017-196.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="diagnosis">
<paragraph pageId="0" pageNumber="51">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="51">
Male of
<taxonomicName lsidName="Q. baoting" pageId="0" pageNumber="51" rank="species" species="baoting">
<emphasis italics="true" pageId="0" pageNumber="51">Q. baoting</emphasis>
</taxonomicName>
sp. nov. can be distinguished from that of
<taxonomicName lsidName="Q. baishensis" pageId="0" pageNumber="51" rank="species" species="baishensis">
<emphasis italics="true" pageId="0" pageNumber="51">Q. baishensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="Q. jianfeng" pageId="0" pageNumber="51" rank="species" species="jianfeng">
<emphasis italics="true" pageId="0" pageNumber="51">Q. jianfeng</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="Q. nui" pageId="0" pageNumber="51" rank="species" species="nui">
<emphasis italics="true" pageId="0" pageNumber="51">Q. nui</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="Q. wuzhi" pageId="0" pageNumber="51" rank="species" species="wuzhi">
<emphasis italics="true" pageId="0" pageNumber="51">Q. wuzhi</emphasis>
</taxonomicName>
, and the other three new species by the conductor with a pointed apex (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male and female genital anatomy of Qiongthela baoting sp. nov. A palp prolateral view B palp ventral view C palp retrolateral view D-F palp distal view G vulva dorsal view H vulva ventral view. A-F XUX- 2017 - 195 (holotype) G-H XUX- 2017 - 196. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.911.48703.figure2" httpUri="https://binary.pensoft.net/fig/379565" pageId="0" pageNumber="51">2A-D</figureCitation>
); from all the other
<taxonomicName authorityName="Xu &amp; Kuntner" authorityYear="2015" class="Arachnida" family="Liphistiidae" genus="Qiongthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Qiongthela" order="Araneae" pageId="0" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="51">Qiongthela</emphasis>
</taxonomicName>
species by the contrategulum with four edges distally (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male and female genital anatomy of Qiongthela baoting sp. nov. A palp prolateral view B palp ventral view C palp retrolateral view D-F palp distal view G vulva dorsal view H vulva ventral view. A-F XUX- 2017 - 195 (holotype) G-H XUX- 2017 - 196. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.911.48703.figure2" httpUri="https://binary.pensoft.net/fig/379565" pageId="0" pageNumber="51">2A, D</figureCitation>
), and by the marginal apophysis of the tegulum with a flake-like, semi-translucent apex (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male and female genital anatomy of Qiongthela baoting sp. nov. A palp prolateral view B palp ventral view C palp retrolateral view D-F palp distal view G vulva dorsal view H vulva ventral view. A-F XUX- 2017 - 195 (holotype) G-H XUX- 2017 - 196. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.911.48703.figure2" httpUri="https://binary.pensoft.net/fig/379565" pageId="0" pageNumber="51">2A, D, F</figureCitation>
). Female of
<taxonomicName lsidName="Q. baoting" pageId="0" pageNumber="51" rank="species" species="baoting">
<emphasis italics="true" pageId="0" pageNumber="51">Q. baoting</emphasis>
</taxonomicName>
sp. nov. differs from that of
<taxonomicName lsidName="Q. baishensis" pageId="0" pageNumber="51" rank="species" species="baishensis">
<emphasis italics="true" pageId="0" pageNumber="51">Q. baishensis</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="Q. nui" pageId="0" pageNumber="51" rank="species" species="nui">
<emphasis italics="true" pageId="0" pageNumber="51">Q. nui</emphasis>
</taxonomicName>
by the base of the lateral receptacular clusters close to the inners, and by the genital stalks of the inners thicker than those of the laterals (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male and female genital anatomy of Qiongthela baoting sp. nov. A palp prolateral view B palp ventral view C palp retrolateral view D-F palp distal view G vulva dorsal view H vulva ventral view. A-F XUX- 2017 - 195 (holotype) G-H XUX- 2017 - 196. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.911.48703.figure2" httpUri="https://binary.pensoft.net/fig/379565" pageId="0" pageNumber="51">2G, H</figureCitation>
); from the other
<taxonomicName authorityName="Xu &amp; Kuntner" authorityYear="2015" class="Arachnida" family="Liphistiidae" genus="Qiongthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Qiongthela" order="Araneae" pageId="0" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="51">Qiongthela</emphasis>
</taxonomicName>
species by two paired receptacular clusters all along the anterior margin of the bursa copulatrix, with distinct genital stalks, and the inners larger than the laterals (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male and female genital anatomy of Qiongthela baoting sp. nov. A palp prolateral view B palp ventral view C palp retrolateral view D-F palp distal view G vulva dorsal view H vulva ventral view. A-F XUX- 2017 - 195 (holotype) G-H XUX- 2017 - 196. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.911.48703.figure2" httpUri="https://binary.pensoft.net/fig/379565" pageId="0" pageNumber="51">2G, H</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.911.48703.figure2" httpUri="https://binary.pensoft.net/fig/379565" pageId="0" pageNumber="51" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" pageId="0" pageNumber="51">Figure 2.</emphasis>
Male and female genital anatomy of
<taxonomicName authorityName="Yu &amp; Liu &amp; Zhang &amp; Wang &amp; Li &amp; Xu" authorityYear="2020" class="Arachnida" family="Liphistiidae" genus="Qiongthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Qiongthela baoting" order="Araneae" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="baoting">
<emphasis italics="true" pageId="0" pageNumber="51">Qiongthela baoting</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="51">A</emphasis>
palp prolateral view
<emphasis bold="true" pageId="0" pageNumber="51">B</emphasis>
palp ventral view
<emphasis bold="true" pageId="0" pageNumber="51">C</emphasis>
palp retrolateral view
<emphasis bold="true" pageId="0" pageNumber="51">D-F</emphasis>
palp distal view
<emphasis bold="true" pageId="0" pageNumber="51">G</emphasis>
vulva dorsal view
<emphasis bold="true" pageId="0" pageNumber="51">H</emphasis>
vulva ventral view.
<emphasis bold="true" pageId="0" pageNumber="51">A-F</emphasis>
XUX-2017-195 (holotype)
<emphasis bold="true" pageId="0" pageNumber="51">G-H</emphasis>
XUX-2017-196. Scale bars: 0.5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="description">
<paragraph pageId="0" pageNumber="51">Description.</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Male</emphasis>
(holotype, Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. General somatic morphology of Qiongthela baoting sp. nov. and a map showing the type localities of seven known Qiongthela species and all sites of four new Qiongthela species in southern Vietnam and Hainan Island, China. A female (XUX- 2017 - 196) B male (XUX- 2017 - 195) C geographical map. Seven known species are indicated in brown solid circles, and four new species are indicated in red, blue, green, and black solid circles." figureDoi="10.3897/zookeys.911.48703.figure1" httpUri="https://binary.pensoft.net/fig/379564" pageId="0" pageNumber="51">1B</figureCitation>
). Carapace dark brown; opisthosoma light brown, with 12 dark brown tergites, close to each other, the first 2-7 larger than others, and the fourth largest; sternum narrow, much longer than wide; a few fine pointed hairs running over the ocular area; chelicerae with promargin of cheliceral groove bearing 9 denticles of variable size; legs with firm hairs and spines; 7 spinnerets. Measurements: BL 12.39, CL 6.17, CW 5.62, OL 6.44, OW 4.52; ALE&gt; PLE&gt; PME&gt; AME; leg I 17.60 (5.15 + 1.65 + 3.82 + 4.28 + 2.69), leg II 16.95 (4.71 + 1.48 + 3.67 + 4.30 + 2.79), leg III 19.26 (4.56 + 1.31 + 3.73 + 6.04 + 3.63), leg IV 25.77 (6.52 + 1.61 + 5.32 + 7.68 + 4.64).
</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis italics="true" pageId="0" pageNumber="51">Palp.</emphasis>
Cymbium with a short, thick projection dorsally (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male and female genital anatomy of Qiongthela baoting sp. nov. A palp prolateral view B palp ventral view C palp retrolateral view D-F palp distal view G vulva dorsal view H vulva ventral view. A-F XUX- 2017 - 195 (holotype) G-H XUX- 2017 - 196. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.911.48703.figure2" httpUri="https://binary.pensoft.net/fig/379565" pageId="0" pageNumber="51">2F</figureCitation>
); paracymbium unpigmented and unsclerotised prolaterally, with numerous setae at the tip (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male and female genital anatomy of Qiongthela baoting sp. nov. A palp prolateral view B palp ventral view C palp retrolateral view D-F palp distal view G vulva dorsal view H vulva ventral view. A-F XUX- 2017 - 195 (holotype) G-H XUX- 2017 - 196. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.911.48703.figure2" httpUri="https://binary.pensoft.net/fig/379565" pageId="0" pageNumber="51">2A, B</figureCitation>
). Contrategulum with an irregular dentate edge proximally and four edges distally: the inner edge sharp, very short; the middle two edges serrate, one towards the proximal portion of contrategulum, the other ended at the centre of the contrategulum; the outer edge short, smooth, slightly sclerotised (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male and female genital anatomy of Qiongthela baoting sp. nov. A palp prolateral view B palp ventral view C palp retrolateral view D-F palp distal view G vulva dorsal view H vulva ventral view. A-F XUX- 2017 - 195 (holotype) G-H XUX- 2017 - 196. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.911.48703.figure2" httpUri="https://binary.pensoft.net/fig/379565" pageId="0" pageNumber="51">2A, D, F</figureCitation>
). The marginal apophysis of tegulum long, wide basally, with a flake-like and semi-translucent apex distally (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male and female genital anatomy of Qiongthela baoting sp. nov. A palp prolateral view B palp ventral view C palp retrolateral view D-F palp distal view G vulva dorsal view H vulva ventral view. A-F XUX- 2017 - 195 (holotype) G-H XUX- 2017 - 196. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.911.48703.figure2" httpUri="https://binary.pensoft.net/fig/379565" pageId="0" pageNumber="51">2A, D</figureCitation>
), a proximally directed terminal apophysis of tegulum with smooth margin, narrowing to a slightly hooked apex (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male and female genital anatomy of Qiongthela baoting sp. nov. A palp prolateral view B palp ventral view C palp retrolateral view D-F palp distal view G vulva dorsal view H vulva ventral view. A-F XUX- 2017 - 195 (holotype) G-H XUX- 2017 - 196. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.911.48703.figure2" httpUri="https://binary.pensoft.net/fig/379565" pageId="0" pageNumber="51">2A-E</figureCitation>
). Conductor situated ventro-proximally on embolus, basal portion fused with embolus, distal free narrowing to a pointed apex (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male and female genital anatomy of Qiongthela baoting sp. nov. A palp prolateral view B palp ventral view C palp retrolateral view D-F palp distal view G vulva dorsal view H vulva ventral view. A-F XUX- 2017 - 195 (holotype) G-H XUX- 2017 - 196. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.911.48703.figure2" httpUri="https://binary.pensoft.net/fig/379565" pageId="0" pageNumber="51">2A-E</figureCitation>
). Embolus largely sclerotised, with a wide, flat opening of sperm duct distally (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male and female genital anatomy of Qiongthela baoting sp. nov. A palp prolateral view B palp ventral view C palp retrolateral view D-F palp distal view G vulva dorsal view H vulva ventral view. A-F XUX- 2017 - 195 (holotype) G-H XUX- 2017 - 196. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.911.48703.figure2" httpUri="https://binary.pensoft.net/fig/379565" pageId="0" pageNumber="51">2A, D, E</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis bold="true" italics="true" pageId="0" pageNumber="51">Female</emphasis>
(Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. General somatic morphology of Qiongthela baoting sp. nov. and a map showing the type localities of seven known Qiongthela species and all sites of four new Qiongthela species in southern Vietnam and Hainan Island, China. A female (XUX- 2017 - 196) B male (XUX- 2017 - 195) C geographical map. Seven known species are indicated in brown solid circles, and four new species are indicated in red, blue, green, and black solid circles." figureDoi="10.3897/zookeys.911.48703.figure1" httpUri="https://binary.pensoft.net/fig/379564" pageId="0" pageNumber="51">1A</figureCitation>
). Carapace dark brown; opisthosoma reddish brown, with 12 red-brown tergites, close to each other, the first 2-7 larger than the others, and the fourth largest; sternum narrow, nearly twice as long as wide; a few fine pointed hairs running over the ocular area; chelicerae robust with promargin of cheliceral groove containing 10 denticles of variable size; legs with firm hairs and spines; 7 spinnerets. Measurements: BL 16.35, CL 7.30, CW 6.12, OL 7.59, OW 6.25; ALE&gt; PLE&gt; PME&gt; AME; palp 10.09 (3.66 + 1.01 + 2.60 + 2.82), leg I 11.78 (3.69 + 1.16 + 3.02 + 2.35 + 1.57), leg II 12.34 (3.94 + 1.44 + 2.69 + 2.58 + 1.70), leg III 10.99 (3.21 + 1.04 + 2.35 +2.90 + 1.47), leg IV 20.26 (5.85 + 1.93 + 4.17 + 5.45 + 2.86).
</paragraph>
<paragraph pageId="0" pageNumber="51">
<emphasis italics="true" pageId="0" pageNumber="51">Female genitalia.</emphasis>
Two pairs of receptacular clusters along the anterior margin of the bursa copulatrix, close to each other, the inner ones distinctly larger than the laterals, with genital stalks thicker than those of the laterals (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Male and female genital anatomy of Qiongthela baoting sp. nov. A palp prolateral view B palp ventral view C palp retrolateral view D-F palp distal view G vulva dorsal view H vulva ventral view. A-F XUX- 2017 - 195 (holotype) G-H XUX- 2017 - 196. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.911.48703.figure2" httpUri="https://binary.pensoft.net/fig/379565" pageId="0" pageNumber="51">2G, H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="etymology">
<paragraph pageId="0" pageNumber="51">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="51">The species epithet, a noun in apposition, refers to the type locality.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="distribution">
<paragraph pageId="0" pageNumber="51">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="51">Hainan (Baoting), China.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="51" type="genbank accession number">
<paragraph pageId="0" pageNumber="51">GenBank accession number.</paragraph>
<paragraph pageId="0" pageNumber="51">Holotype (XUX-2017-195): MN911989.</paragraph>
</subSubSection>
</treatment>
</document>