399 lines
30 KiB
XML
399 lines
30 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.852.28964" ID-GBIF-Dataset="f29916c9-895c-46ac-97ee-fd94ac9cea74" ID-PMC="PMC6562051" ID-Pensoft-Pub="1313-2970-852-1" ID-PubMed="31210739" ID-ZBK="0CAC34BD1C75415F973DC37E4557D06F" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1313-2970-852-1" ModsDocOrigin="ZooKeys 852" ModsDocTitle="Two new species of Acanthobothrium Blanchard, 1848 (Onchobothriidae) in Narcineentemedor Jordan & Starks, 1895 (Narcinidae) from Acapulco, Guerrero, Mexico" checkinTime="1559826557769" checkinUser="pensoft" docAuthor="Zaragoza-Tapia, Francisco, Pulido-Flores, Griselda, Violante-Gonzalez, Juan & Monks, Scott" docDate="2019" docId="9DD1F1DD33E097C5F34CBCEA0A22A92C" docLanguage="en" docName="ZooKeys 852: 1-21" docOrigin="ZooKeys 852" docSource="http://dx.doi.org/10.3897/zookeys.852.28964" docTitle="Acanthobothrium vidali Zaragoza-Tapia, Pulido-Flores, Violante-Gonzalez & Monks, 2019, sp. nov." docType="treatment" docUuid="9D9106EF-7772-4E61-A44F-B2D151782329" docUuidSource="ZooBank" docVersion="5" lastPageNumber="1" masterDocId="FFC4FFC8FFD67550FFD1FFA8EE643021" masterDocTitle="Two new species of Acanthobothrium Blanchard, 1848 (Onchobothriidae) in Narcineentemedor Jordan & Starks, 1895 (Narcinidae) from Acapulco, Guerrero, Mexico" masterLastPageNumber="21" masterPageNumber="1" pageNumber="1" updateTime="1668167339479" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Two new species of Acanthobothrium Blanchard, 1848 (Onchobothriidae) in Narcineentemedor Jordan & Starks, 1895 (Narcinidae) from Acapulco, Guerrero, Mexico</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zaragoza-Tapia, Francisco</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Pulido-Flores, Griselda</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Violante-Gonzalez, Juan</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Monks, Scott</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>852</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>21</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.852.28964</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.852.28964</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-852-1</mods:identifier>
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<mods:identifier type="ZBK">0CAC34BD1C75415F973DC37E4557D06F</mods:identifier>
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<mods:identifier type="ZooBank">0CAC34BD1C75415F973DC37E4557D06F</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="157422764" LSID="urn:lsid:zoobank.org:act:9D9106EF-7772-4E61-A44F-B2D151782329" httpUri="http://treatment.plazi.org/id/9DD1F1DD33E097C5F34CBCEA0A22A92C" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/9D9106EF-7772-4E61-A44F-B2D151782329" class="Cestoda" family="Onchobothriidae" genus="Acanthobothrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthobothrium vidali" order="Onchoproteocephalidea" pageId="0" pageNumber="1" phylum="Platyhelminthes" rank="species" species="vidali">Acanthobothrium vidali</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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Figures 3
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<normalizedToken originalValue="A–D">A-D</normalizedToken>
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; 4
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<normalizedToken originalValue="A–D">A-D</normalizedToken>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">Holotype (CNHE-11134), 7 paratypes (CNHE-11135), 3 paratypes (HWML-139979, 139980, 139981), and 7 paratypes (CHE-P00082).</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Other material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Cestoda" family="Onchobothriidae" genus="Acanthobothrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthobothrium franus" order="Onchoproteocephalidea" pageId="0" pageNumber="1" phylum="Platyhelminthes" rank="species" species="franus">Acanthobothrium franus</taxonomicName>
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Marques, Centritto & Stewart, 1997 (
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<normalizedToken originalValue="CNHE–">CNHE-</normalizedToken>
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3140, Costa Rica) paratype;
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<taxonomicName lsidName="A. inbiorium" pageId="0" pageNumber="1" rank="species" species="inbiorium">A. inbiorium</taxonomicName>
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Marques, Centritto & Stewart, 1997 (
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<normalizedToken originalValue="CNHE–">CNHE-</normalizedToken>
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3138, Costa Rica) paratype;
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<taxonomicName lsidName="A. obuncus" pageId="0" pageNumber="1" rank="species" species="obuncus">A. obuncus</taxonomicName>
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Marques, Brooks & Barriga, 1997 (
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<normalizedToken originalValue="CNHE–">CNHE-</normalizedToken>
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3032A, 3167B, Ecuador) holotype;
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<taxonomicName lsidName="A. soberoni" pageId="0" pageNumber="1" rank="species" species="soberoni">A. soberoni</taxonomicName>
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Ghoshroy & Caira, 2001 (
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<normalizedToken originalValue="CNHE–">CNHE-</normalizedToken>
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4042,
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<normalizedToken originalValue="México">Mexico</normalizedToken>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type host">
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<paragraph pageId="0" pageNumber="1">Type host.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Elasmobranchii" family="Narcinidae" genus="Narcine" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Narcine entemedor" order="Torpediniformes" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="entemedor">Narcine entemedor</taxonomicName>
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Jordan & Starks, 1895 (
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<taxonomicName lsidName="Elasmobranchii" pageId="0" pageNumber="1" rank="subclass" subclass="Elasmobranchii">Elasmobranchii</taxonomicName>
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:
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<taxonomicName lsidName="Elasmobranchii" order="Torpediniformes" pageId="0" pageNumber="1" rank="order">Torpediniformes</taxonomicName>
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:
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<taxonomicName family="Narcinidae" lsidName="" pageId="0" pageNumber="1" rank="family">Narcinidae</taxonomicName>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type locality">
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<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<normalizedToken originalValue="Bahía">Bahia</normalizedToken>
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de Acapulco (Playa Las Hamacas:
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="16.853056">16°51'11"N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-99.89972">99°53'59"W</geoCoordinate>
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), Guerrero,
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<normalizedToken originalValue="México">Mexico</normalizedToken>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="site of infection">
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<paragraph pageId="0" pageNumber="1">Site of infection.</paragraph>
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<paragraph pageId="0" pageNumber="1">Spiral intestine.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="quantitative descriptors of parasite populations (bush et al. 1997)">
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<paragraph pageId="0" pageNumber="1">
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Quantitative descriptors of parasite populations (
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<bibRefCitation author="Bush, AO" journalOrPublisher="Journal of Parasitology" pageId="0" pageNumber="1" pagination="575 - 583" title="Parasitology meets ecology on its own terms: Margolis et al. revisited." url="https://doi.org/10.2307/3284227" volume="83" year="1997">Bush et al. 1997</bibRefCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="1">Prevalence = 36.36% (4 of 11 rays were infected); abundance = 1.91 (21 helminths in 11 rays); mean intensity = 5.25 (21 helminths in 4 infected rays).</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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The species is named in honor of Dr Victor Vidal
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<normalizedToken originalValue="Martínez">Martinez</normalizedToken>
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(Departamento de Recursos del Mar, CINVESTAV-IPN, Merida, Yucatan, Mexico), for his contribution to our knowledge of helminths of fishes from Mexico.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Cestoda" family="Onchobothriidae" genus="Acanthobothrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthobothrium vidali" order="Onchoproteocephalidea" pageId="0" pageNumber="1" phylum="Platyhelminthes" rank="species" species="vidali">Acanthobothrium vidali</taxonomicName>
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sp. nov. is a Category 6 species. This species is large, with a range of 164-214 craspedote proglottids, with a range of 50-76 testes per proglottid, and the arms of ovary unequal in length (asymmetrical). This new species also can be distinguished from similar congeners by total length, number of proglottids, diameter of accessory sucker, the length of the cirrus sac, number of testes per proglottid, and size of the hooks.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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[Based on 5 complete worms and 16 partial specimens] Worms 26.5-70.9 mm (n = 5) long, greatest width at level of mature proglottids, euapolytic; 164-214 (n = 5) proglottids per worm. Scolex 880-1,400 (n = 20) long by 680-1,170 (n = 20) wide, with four bothridia; maximum width of scolex at level of posterior margin of anterior loculus (Figs 3A, 4A). Bothridia free posteriorly, tri-locular, 770-1,230 (n = 20) long by 320-570 (n = 20) wide, with anterior muscular pad (Figs 3A, 4A). Muscular pad 250-325 (n = 18) wide, with apical sucker 75-150 (n = 19) and one pair of bipronged hooks at posterior margin (Figs 3A, 4A). Anterior loculus of bothridia 400-650 (n = 20) long; middle loculus 170-310 (n = 20) long; posterior loculus 150-340 (n = 20) long (Figs 3A, 4A); loculus length ratio (anterior:middle:posterior) 1:0.48:0.50. Velum between medial margins of bothridia in dorsal or ventral pairs not seen (Figs 3A, 4A). Hooks bipronged, hollow, with tubercle on proximal surface of axial prong; internal channels of axial and abaxial prongs continuous, smooth, base and anterior part of each hook embedded in musculature of scolex, tips of prongs free (Figs 3A, B, 4A). Bases (handles) of medial and lateral hooks articulate with one another (Figs 3B, 4A). Lateral hook measurements (n = 15): A 140-170, B 200-285, C 140-305, D 360-465; Medial hook measurements (n = 15):
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<normalizedToken originalValue="A’">A'</normalizedToken>
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100-165,
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<normalizedToken originalValue="B’">B'</normalizedToken>
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225-300,
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<normalizedToken originalValue="C’">C'</normalizedToken>
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200-270,
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<normalizedToken originalValue="D’">D'</normalizedToken>
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300-425. Cephalic peduncle 2.38-9.13 mm (n = 15) long by 0.15-0.23 mm (n = 15) wide, microtriches not seen on the scolex or cephalic peduncle (Figs 3A, 4A, B, C). Proglottids craspedote. Immature proglottids 50-230 (n = 17) long by 240-520 (n = 17) wide, mature proglottids 260-700 (n = 10) long by 300-790 (n = 10) wide (Fig. 3C), terminal proglottids 1,120 (n = 1) long by 480 (n = 1) wide. Genital pore marginal, irregularly alternating, 49%-63% (n = 9) of proglottid length from anterior end in mature proglottids; genital atrium present (Figs 3C, D, 4D). Testes arranged in two to three irregular columns on each side of the uterus, in frontal view testes wider than long in mature proglottids, 50-125 (n = 6) long by 40-50 (n = 6) wide (Fig. 3C). In terminal proglottids, anteriormost testes wider than long and posteriormost testes longer than wide. Total number of testes 50-76 (n = 10), aporal 26-40 (n = 10), poral 23-36 (n = 10), preporal 17-26 (n = 10), postporal 5-11 (n = 10). All testes located anterior to ovarian isthmus. Cirrus sac pyriform, 125-175 (n = 6) long by 30-75 (n = 6) wide in mature proglottids (Figs 3C, D, 4D). Cirrus armed. Vagina anterior to cirrus sac (Figs 3C, D, 4D),walls relatively thick, covered with gland cells. Vagina extending laterally from common genital atrium, following anterior margin of cirrus sac, weakly sinuous posteriorly along medial line of proglottid to
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<normalizedToken originalValue="oötype">ooetype</normalizedToken>
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(Fig. 3C); vaginal sphincter absent. Seminal receptacle not seen. Ovary in mature proglottids H-shaped in frontal view (Fig. 3C); posterior lobes wider than anterior lobes. Ovarian isthmus approximately 2/3 of the distance from anterior end of ovary. Arms of ovary unequal in length, aporal arm always longer than poral arm (Fig. 3C). Aporal arm 150-260 (n = 8) long, reaching to posterior margin of cirrus sac, poral arm 125-225 (n = 8) long in mature proglottids, not reaching posterior margin of cirrus sac.
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<normalizedToken originalValue="Mehlis’">Mehlis'</normalizedToken>
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gland posterior to ovarian isthmus. Vitiellarium follicular, forming lateral bands, extending from near anterior margin of proglottid to near posterior margin of proglottid (Fig. 3C); follicles 15-20 (n = 8) long by 10-15 (n = 8) wide. Uterus thin-walled, saccate, extending from anterior margin of proglottid to near posterior margin of proglottid. Excretory ducts lateral. Gravid proglottids and eggs not seen.
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</paragraph>
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<caption pageId="0" pageNumber="1">
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<paragraph pageId="0" pageNumber="1">
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Figure 3. Holotype of
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<taxonomicName class="Cestoda" family="Onchobothriidae" genus="Acanthobothrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthobothrium vidali" order="Onchoproteocephalidea" pageId="0" pageNumber="1" phylum="Platyhelminthes" rank="species" species="vidali">Acanthobothrium vidali</taxonomicName>
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sp. nov. (CNHE-11134). A Scolex B hooks C mature proglottid D genitalia. Scale bars: 400
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<normalizedToken originalValue="µm">µm</normalizedToken>
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(A); 200
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<normalizedToken originalValue="µm">µm</normalizedToken>
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(
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<normalizedToken originalValue="B–D">B-D</normalizedToken>
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).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="remarks">
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<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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There are 14 Category 6 species of
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<taxonomicName class="Cestoda" family="Onchobothriidae" genus="Acanthobothrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthobothrium" order="Onchoproteocephalidea" pageId="0" pageNumber="1" phylum="Platyhelminthes" rank="genus">Acanthobothrium</taxonomicName>
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that have been described worldwide. Of these, seven species have been found in the Pacific Ocean, four of which are amphi-American (Table 2).
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<taxonomicName class="Cestoda" family="Onchobothriidae" genus="Acanthobothrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthobothrium vidali" order="Onchoproteocephalidea" pageId="0" pageNumber="1" phylum="Platyhelminthes" rank="species" species="vidali">Acanthobothrium vidali</taxonomicName>
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sp. nov. also is a Category 6 species, bringing the total number to 15. Category 6 species have a total length>15 mm (the length of
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<taxonomicName lsidName="A. vidali" pageId="0" pageNumber="1" rank="species" species="vidali">A. vidali</taxonomicName>
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sp. nov. is 26.5-70.9 mm), a strobila made up of> 50 proglottids (
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<taxonomicName lsidName="A. vidali" pageId="0" pageNumber="1" rank="species" species="vidali">A. vidali</taxonomicName>
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sp. nov. has 164-214 proglottids), number of testes per proglottids ≤ 80 (
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<taxonomicName lsidName="A. vidali" pageId="0" pageNumber="1" rank="species" species="vidali">A. vidali</taxonomicName>
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sp. nov. has 50-76 testes per proglottid), and the arms of the ovary are asymmetrical.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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As presented in Table 2, the new species can be distinguished from similar Category 6 species of
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<taxonomicName class="Cestoda" family="Onchobothriidae" genus="Acanthobothrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthobothrium" order="Onchoproteocephalidea" pageId="0" pageNumber="1" phylum="Platyhelminthes" rank="genus">Acanthobothrium</taxonomicName>
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that have been described from the Pacific Coast of the Americas (amphi-American species), and from others parts of Pacific Ocean by the measurementes given in Table 2. The total length of the new species (26.5-70.9 mm) is longer than that of
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<taxonomicName lsidName="A. aetiobatidis" pageId="0" pageNumber="1" rank="species" species="aetiobatidis">A. aetiobatidis</taxonomicName>
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(Shipley, 1900) Southwell, 1925. The number of proglottids of the new species (164-214) is greater than that of
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<taxonomicName lsidName="A. gonzalesmugaburoi" pageId="0" pageNumber="1" rank="species" species="gonzalesmugaburoi">A. gonzalesmugaburoi</taxonomicName>
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Severino & Sarmiento, 1979. The diameter of accessory sucker of the new species (75-150) is larger than that of
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<taxonomicName lsidName="A. obuncus" pageId="0" pageNumber="1" rank="species" species="obuncus">A. obuncus</taxonomicName>
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(33-48) and
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<taxonomicName lsidName="A. soberoni" pageId="0" pageNumber="1" rank="species" species="soberoni">A. soberoni</taxonomicName>
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(40-65). The length of the cirrus sac of the new species (125-175) is shorter than that of
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<taxonomicName lsidName="A. obuncus" pageId="0" pageNumber="1" rank="species" species="obuncus">A. obuncus</taxonomicName>
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(258-322),
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<taxonomicName lsidName="A. aetiobatidis" pageId="0" pageNumber="1" rank="species" species="aetiobatidis">A. aetiobatidis</taxonomicName>
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(200-250), and
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<taxonomicName lsidName="A. rodmani" pageId="0" pageNumber="1" rank="species" species="rodmani">A. rodmani</taxonomicName>
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Fyler, Caira & Jensen, 2009 (190-234). The number of testes per proglottid of the new species (50-76) is greater than that of
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<taxonomicName lsidName="A. aetiobatidis" pageId="0" pageNumber="1" rank="species" species="aetiobatidis">A. aetiobatidis</taxonomicName>
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(23-28),
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<taxonomicName lsidName="A. arlenae" pageId="0" pageNumber="1" rank="species" species="arlenae">A. arlenae</taxonomicName>
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Campbell & Beverage, 2002 (17-26), and
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<taxonomicName lsidName="A. rodmani" pageId="0" pageNumber="1" rank="species" species="rodmani">A. rodmani</taxonomicName>
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(17-26). Finally, the measurements of the hooks of the eight species can be found in Table 2.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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The new species is the fourth species of
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<taxonomicName class="Cestoda" family="Onchobothriidae" genus="Acanthobothrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthobothrium" order="Onchoproteocephalidea" pageId="0" pageNumber="1" phylum="Platyhelminthes" rank="genus">Acanthobothrium</taxonomicName>
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reported from
|
||
<taxonomicName lsidName="N. entemedor" pageId="0" pageNumber="1" rank="species" species="entemedor">N. entemedor</taxonomicName>
|
||
, preceded by
|
||
<taxonomicName lsidName="A. franus" pageId="0" pageNumber="1" rank="species" species="franus">A. franus</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="A. inbiorium" pageId="0" pageNumber="1" rank="species" species="inbiorium">A. inbiorium</taxonomicName>
|
||
(Category 5 species), and
|
||
<taxonomicName lsidName="A. soniae" pageId="0" pageNumber="1" rank="species" species="soniae">A. soniae</taxonomicName>
|
||
sp. nov. (Category 2 species) described above. All species have been reported from the Pacific Coast of the Americas.
|
||
<taxonomicName class="Cestoda" family="Onchobothriidae" genus="Acanthobothrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthobothrium vidali" order="Onchoproteocephalidea" pageId="0" pageNumber="1" phylum="Platyhelminthes" rank="species" species="vidali">Acanthobothrium vidali</taxonomicName>
|
||
sp. nov. can be distinguished from these other species by number of proglottids (164-214) is greater than that of
|
||
<taxonomicName lsidName="A. franus" pageId="0" pageNumber="1" rank="species" species="franus">A. franus</taxonomicName>
|
||
(68-141) and
|
||
<taxonomicName lsidName="A. soniae" pageId="0" pageNumber="1" rank="species" species="soniae">A. soniae</taxonomicName>
|
||
sp. nov. (10-13). The total length of the lateral hook of
|
||
<taxonomicName lsidName="A. vidali" pageId="0" pageNumber="1" rank="species" species="vidali">A. vidali</taxonomicName>
|
||
sp. nov. (360-465) is longer than that of
|
||
<taxonomicName lsidName="A. inbiorium" pageId="0" pageNumber="1" rank="species" species="inbiorium">A. inbiorium</taxonomicName>
|
||
(95-120
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
). The length of the axial prong of the lateral hook of
|
||
<taxonomicName lsidName="A. vidali" pageId="0" pageNumber="1" rank="species" species="vidali">A. vidali</taxonomicName>
|
||
sp. nov. (200-285) is longer than that of
|
||
<taxonomicName lsidName="A. inbiorium" pageId="0" pageNumber="1" rank="species" species="inbiorium">A. inbiorium</taxonomicName>
|
||
(65-75
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
) (Table 3).
|
||
</paragraph>
|
||
<caption pageId="0" pageNumber="1">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Figure 4. Light microscope photographs of holotype of
|
||
<taxonomicName class="Cestoda" family="Onchobothriidae" genus="Acanthobothrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthobothrium vidali" order="Onchoproteocephalidea" pageId="0" pageNumber="1" phylum="Platyhelminthes" rank="species" species="vidali">Acanthobothrium vidali</taxonomicName>
|
||
sp. nov. (CNHE-11134). A Scolex B details of bothridium C peduncle cephalic D genitalia. Scale bars: 400
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
(A); 40
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
(B);100
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
(C, D). Abbreviations: vg vagina; cs cirrus sac; gp genital pore.
|
||
</paragraph>
|
||
</caption>
|
||
<caption pageId="0" pageNumber="1">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Table 2. Comparison of
|
||
<taxonomicName class="Cestoda" family="Onchobothriidae" genus="Acanthobothrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthobothrium vidali" order="Onchoproteocephalidea" pageId="0" pageNumber="1" phylum="Platyhelminthes" rank="species" species="vidali">Acanthobothrium vidali</taxonomicName>
|
||
sp. nov. vs. Category 6 species of the genus from the Pacific Ocean. Abbreviations: No. Number; A Base (handle) length; B Axial prong length; C Abaxial prong length; D Total hook length. Note: the use of
|
||
<normalizedToken originalValue="“–”">"-"</normalizedToken>
|
||
without numerical values are measurement ranges that overlap those of
|
||
<taxonomicName lsidName="A. vidali" pageId="0" pageNumber="1" rank="species" species="vidali">A. vidali</taxonomicName>
|
||
sp. nov.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<table pageId="0" pageNumber="1">
|
||
<tr pageId="0" pageNumber="1">
|
||
<th colspan="1" pageId="0" pageNumber="1" rowspan="2">
|
||
Species of
|
||
<taxonomicName class="Cestoda" family="Onchobothriidae" genus="Acanthobothrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthobothrium" order="Onchoproteocephalidea" pageId="0" pageNumber="1" phylum="Platyhelminthes" rank="genus">Acanthobothrium</taxonomicName>
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="1" rowspan="2">Total length (mm)</th>
|
||
<th colspan="1" pageId="0" pageNumber="1" rowspan="2">No. of proglottids</th>
|
||
<th colspan="1" pageId="0" pageNumber="1" rowspan="2">Diameter of accesory sucker (μm)</th>
|
||
<th colspan="1" pageId="0" pageNumber="1" rowspan="2">Length of cirrus sac (μm)</th>
|
||
<th colspan="1" pageId="0" pageNumber="1" rowspan="2">No. of testes per proglottid</th>
|
||
<th colspan="4" pageId="0" pageNumber="1" rowspan="1">Measurements of hook (μm)</th>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">A</th>
|
||
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">B</th>
|
||
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">C</th>
|
||
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">D</th>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="10" pageId="0" pageNumber="1" rowspan="1">Pacific coast of the Americas</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName lsidName="A. vidali" pageId="0" pageNumber="1" rank="species" species="vidali">A. vidali</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName lsidName="A. gonzalesmugaburoi" pageId="0" pageNumber="1" rank="species" species="gonzalesmugaburoi">A. gonzalesmugaburoi</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName lsidName="A. maculatum" pageId="0" pageNumber="1" rank="species" species="maculatum">A. maculatum</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName lsidName="A. obuncus" pageId="0" pageNumber="1" rank="species" species="obuncus">A. obuncus</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName lsidName="A. soberoni" pageId="0" pageNumber="1" rank="species" species="soberoni">A. soberoni</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="10" pageId="0" pageNumber="1" rowspan="1">Other parts of the Pacific Ocean</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName lsidName="A. aetiobatidis" pageId="0" pageNumber="1" rank="species" species="aetiobatidis">A. aetiobatidis</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName lsidName="A. arlenae" pageId="0" pageNumber="1" rank="species" species="arlenae">A. arlenae</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName lsidName="A. rodmani" pageId="0" pageNumber="1" rank="species" species="rodmani">A. rodmani</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption pageId="0" pageNumber="1">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Table 3. Comparison of species of
|
||
<taxonomicName class="Cestoda" family="Onchobothriidae" genus="Acanthobothrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthobothrium" order="Onchoproteocephalidea" pageId="0" pageNumber="1" phylum="Platyhelminthes" rank="genus">Acanthobothrium</taxonomicName>
|
||
that have been reported from
|
||
<taxonomicName class="Elasmobranchii" family="Narcinidae" genus="Narcine" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Narcine entemedor" order="Torpediniformes" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="entemedor">Narcine entemedor</taxonomicName>
|
||
. Abbreviations: No. Number; A Base (handle) length; B Axial prong length; C Abaxial prong length; D Total hook length. Note: Information taken from the original descriptions and this study.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<table pageId="0" pageNumber="1">
|
||
<tr pageId="0" pageNumber="1">
|
||
<th colspan="1" pageId="0" pageNumber="1" rowspan="2">
|
||
Species of
|
||
<taxonomicName class="Cestoda" family="Onchobothriidae" genus="Acanthobothrium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acanthobothrium" order="Onchoproteocephalidea" pageId="0" pageNumber="1" phylum="Platyhelminthes" rank="genus">Acanthobothrium</taxonomicName>
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="1" rowspan="2">Total length (mm)</th>
|
||
<th colspan="1" pageId="0" pageNumber="1" rowspan="2">No. of proglottids</th>
|
||
<th colspan="1" pageId="0" pageNumber="1" rowspan="2">Diameter of accesory sucker (μm)</th>
|
||
<th colspan="1" pageId="0" pageNumber="1" rowspan="2">Length of cirrus sac (μm)</th>
|
||
<th colspan="1" pageId="0" pageNumber="1" rowspan="2">No. of testes per proglottid</th>
|
||
<th colspan="4" pageId="0" pageNumber="1" rowspan="1">Measurements of hook (μm)</th>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">A</th>
|
||
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">B</th>
|
||
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">C</th>
|
||
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">D</th>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="10" pageId="0" pageNumber="1" rowspan="1">
|
||
Reported from
|
||
<taxonomicName class="Elasmobranchii" family="Narcinidae" genus="Narcine" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Narcine entemedor" order="Torpediniformes" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="entemedor">Narcine entemedor</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName lsidName="A. franus" pageId="0" pageNumber="1" rank="species" species="franus">A. franus</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName lsidName="A. inbiorium" pageId="0" pageNumber="1" rank="species" species="inbiorium">A. inbiorium</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName lsidName="A. soniae" pageId="0" pageNumber="1" rank="species" species="soniae">A. soniae</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName lsidName="A. vidali" pageId="0" pageNumber="1" rank="species" species="vidali">A. vidali</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |