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<document id="7DD80ADD812E88FBE5B92B130CFF9FAD" ID-CLB-Dataset="4388" ID-DOI="10.1111/j.1096-3642.2003.00102.x" ID-GBIF-Dataset="02fb64c3-0755-4e55-8d37-5a5dfcdcfda2" ID-ISSN="0024-4082" ID-Zenodo-Dep="5428881" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630419726431" checkinUser="diego" docAuthor="Modesto, Sean P. &amp; Sues, Hans-Dieter" docDate="2004" docId="9B60DA23723FFFDCB0513B6E4325FC93" docLanguage="en" docName="j.1096-3642.2003.00102.x.pdf" docOrigin="Zoological Journal of the Linnean Society 140 (3)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2003.00102.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Prolacerta broomi Parrington 1935" docType="treatment" docVersion="7" lastPageNumber="337" masterDocId="6759A25B723EFFDEB3293E5E4275FF83" masterDocTitle="The skull of the Early Triassic archosauromorph reptile Prolacerta broomi and its phylogenetic significance" masterLastPageNumber="351" masterPageNumber="335" pageNumber="336" updateTime="1699267248586" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="99A952C96F59E128182AAC5589FBA891">The skull of the Early Triassic archosauromorph reptile Prolacerta broomi and its phylogenetic significance</mods:title>
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<treatment id="9B60DA23723FFFDCB0513B6E4325FC93" ID-DOI="http://doi.org/10.5281/zenodo.5488621" ID-GBIF-Taxon="185435729" ID-Zenodo-Dep="5488621" LSID="urn:lsid:plazi:treatment:9B60DA23723FFFDCB0513B6E4325FC93" httpUri="http://treatment.plazi.org/id/9B60DA23723FFFDCB0513B6E4325FC93" lastPageId="2" lastPageNumber="337" pageId="1" pageNumber="336">
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<paragraph id="13766B35723FFFDFB0513B6E4722FAC4" blockId="1.[888,1367,1231,1351]" box="[888,1367,1327,1351]" pageId="1" pageNumber="336">
<heading id="483EDC59723FFFDFB0513B6E4722FAC4" box="[888,1367,1327,1351]" centered="true" fontSize="9" level="2" pageId="1" pageNumber="336" reason="2">
<taxonomicName id="D4C910B6723FFFDFB0513B6E4722FAC4" authority="PARRINGTON, 1935" authorityName="Parrington" authorityYear="1935" box="[888,1367,1327,1351]" class="Reptilia" family="Protorosauridae" genus="Prolacerta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Protorosauria" pageId="1" pageNumber="350" phylum="Chordata" rank="species" species="broomi">
<emphasis id="21BDB727723FFFDFB0513B6E4618FAC5" box="[888,1133,1328,1351]" italics="true" pageId="1" pageNumber="336">PROLACERTA BROOMI</emphasis>
<bibRefCitation id="775816C4723FFFDFB7543B714722FAC4" author="Parrington FR" box="[1149,1367,1327,1351]" pageId="1" pageNumber="336" pagination="197 - 205" refId="ref11810" refString="Parrington FR. 1935. On Prolacerta broomi, gen. et sp. n. and the origin of lizards. Annals and Magazine of Natural History 16: 197 - 205." type="journal article" year="1935">PARRINGTON, 1935</bibRefCitation>
</taxonomicName>
</heading>
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<paragraph id="13766B35723FFFDFB01D3B3F46D8F9AE" blockId="1.[820,1436,1376,1582]" pageId="1" pageNumber="336">
<emphasis id="21BDB727723FFFDFB01D3B3F41D1FAF5" box="[820,932,1377,1398]" italics="true" pageId="1" pageNumber="336">Diagnosis</emphasis>
. Distinguished by the presence of septomaxillae, conspicuous posterolateral exposure of the lacrimal duct openings (condition unknown in some other possibly related taxa, e.g.
<taxonomicName id="D4C910B6723FFFDFB7413BE3477CFA51" box="[1128,1289,1469,1490]" class="Reptilia" family="Protorosauridae" genus="Protorosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Protorosauria" pageId="1" pageNumber="336" phylum="Chordata" rank="genus">
<emphasis id="21BDB727723FFFDFB7413BE3477CFA51" box="[1128,1289,1469,1490]" italics="true" pageId="1" pageNumber="336">Protorosaurus</emphasis>
</taxonomicName>
), absence of postparietals, and by extensive contact between the surangular and the prearticular in the articular region of the mandibular ramus.
</paragraph>
<paragraph id="13766B35723FFFDFB01D381041D0F900" blockId="1.[820,1435,1614,1667]" pageId="1" pageNumber="336">
<materialsCitation id="A3A16168723FFFDFB01D381041D0F900" ID-GBIF-Occurrence="3341467302" collectionCode="UMZC" pageId="1" pageNumber="336" specimenCode="UMZC 2003.40" specimenCount="1" typeStatus="holotype">
<emphasis id="21BDB727723FFFDFB01D381041EDF9E0" box="[820,920,1614,1635]" italics="true" pageId="1" pageNumber="336">
<typeStatus id="CC72D597723FFFDFB01D381041EDF9E0" box="[820,920,1614,1635]" pageId="1" pageNumber="336" type="holotype">Holotype</typeStatus>
</emphasis>
.
<specimenCode id="436FC34E723FFFDFB09F38104602F9E0" box="[950,1143,1614,1635]" collectionCode="UMZC" pageId="1" pageNumber="336">UMZC 2003.40</specimenCode>
, a partial skull and mandible.
</materialsCitation>
</paragraph>
<paragraph id="13766B35723FFFDCB01D38FD4359FE95" blockId="1.[820,1436,1699,1905]" lastBlockId="2.[143,757,195,278]" lastPageId="2" lastPageNumber="337" pageId="1" pageNumber="336">
<emphasis id="21BDB727723FFFDFB01D38FD4679F93B" box="[820,1036,1699,1720]" italics="true" pageId="1" pageNumber="336">Material examined</emphasis>
.
<materialsCitation id="A3A16168723FFFDCB73738FD435CFE95" ID-GBIF-Occurrence="3341467301" collectionCode="UCMP" lastPageId="2" lastPageNumber="337" pageId="1" pageNumber="336" specimenCode="UCMP 37151" specimenCount="1" typeStatus="holotype">
BP/1/471 (
<typeStatus id="CC72D597723FFFDFB7B038FD468EF93A" box="[1177,1275,1699,1721]" pageId="1" pageNumber="336" type="holotype">holotype</typeStatus>
of subjective junior synonym
<taxonomicName id="D4C910B6723FFFDFB0DA389C4118F975" authority="Broom &amp; Robinson, 1948" authorityName="Broom &amp; Robinson" authorityYear="1948" class="Monogenea" family="Thoracocotylidae" genus="Pricea" kingdom="Animalia" order="Mazocraeidea" pageId="1" pageNumber="336" phylum="Platyhelminthes" rank="species" species="longiceps">
<emphasis id="21BDB727723FFFDFB0DA389C46D8F955" box="[1011,1197,1729,1751]" italics="true" pageId="1" pageNumber="336">Pricea longiceps</emphasis>
<bibRefCitation id="775816C4723FFFDFB79E389C4118F975" author="Broom R &amp; Robinson JT" pageId="1" pageNumber="336" pagination="392 - 407" refId="ref10779" refString="Broom R, Robinson JT. 1948. Some new fossil reptiles from the Karroo beds of South Africa. Proceedings of the Zoological Society of London B 118: 392 - 407." type="journal article" year="1948">Broom &amp; Robinson, 1948</bibRefCitation>
</taxonomicName>
), a complete skull with attached mandible; BP/1/ 2675, a nearly complete skull, now mostly disarticulated, with postcranial skeleton; BP/1/4504a, a skull of a small individual; BP/1/5066, a partial, flattened skull; BP/1/5375, a skull, complete from mid-snout to occiput with accompanying portions of the mandible;
<specimenCode id="436FC34E723CFFDCB3A63EBF435EFF74" box="[143,299,225,247]" collectionCode="UCMP" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="2" pageNumber="337" type="University or college">UCMP 37151</specimenCode>
, a complete skull with articulated cervical vertebrae
</materialsCitation>
.
</paragraph>
<paragraph id="13766B35723CFFDCB3A63F714325FC93" blockId="2.[143,759,302,784]" pageId="2" pageNumber="337">
<emphasis id="21BDB727723CFFDCB3A63F714035FEC7" box="[143,576,302,324]" italics="true" pageId="2" pageNumber="337">Horizon and geographical provenance</emphasis>
.
<taxonomicName id="D4C910B6723CFFDCB1733F7040A5FEC0" box="[602,720,302,323]" class="Reptilia" family="Protorosauridae" genus="Prolacerta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Protorosauria" pageId="2" pageNumber="337" phylum="Chordata" rank="genus">
<emphasis id="21BDB727723CFFDCB1733F7040A5FEC0" box="[602,720,302,323]" italics="true" pageId="2" pageNumber="337">Prolacerta</emphasis>
</taxonomicName>
is currently known only from the Lower Triassic
<taxonomicName id="D4C910B6723CFFDCB1813F1343C7FE02" authority="Assemblage Zone" authorityName="Assemblage Zone" class="Reptilia" family="Lystrosauridae" genus="Lystrosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Therapsida" pageId="2" pageNumber="337" phylum="Chordata" rank="genus">
<emphasis id="21BDB727723CFFDCB1813F1342A9FE02" italics="true" pageId="2" pageNumber="337">Lystrosaurus</emphasis>
Assemblage Zone
</taxonomicName>
(Beaufort Group) of
<collectingCountry id="6BDE2BA5723CFFDCB19B3F3542A1FE23" name="South Africa" pageId="2" pageNumber="337">South Africa</collectingCountry>
(
<bibRefCitation id="775816C4723CFFDCB3CD3FD4404FFE23" author="Groenewald GH &amp; Kitching JW" box="[228,570,394,416]" pageId="2" pageNumber="337" pagination="35 - 39" refId="ref11354" refString="Groenewald GH, Kitching JW. 1995. Biostratigraphy of the Lystrosaurus Assemblage Zone. In: Rubidge BS, ed. Biostratigraphy of the Beaufort Group (Karoo Supergroup). Pretoria: South African Committee for Stratigraphy, Biostratigraphic Series 1: 35 - 39." type="book chapter" year="1995">Groenewald &amp; Kitching, 1995</bibRefCitation>
) and correlative strata of the Fremouw Formation in the Transantarctic Mountains of
<collectingCountry id="6BDE2BA5723CFFDCB27A3F9943BEFE5E" box="[339,459,455,477]" name="Antarctica" pageId="2" pageNumber="337">Antarctica</collectingCountry>
(
<bibRefCitation id="775816C4723CFFDCB2F23F99400CFE5E" author="Colbert EH" box="[475,633,455,477]" pageId="2" pageNumber="337" pagination="1 - 19" refId="ref10922" refString="Colbert EH. 1987. The Triassic reptile Prolacerta in Antarctica. American Museum Novitates 2882: 1 - 19." type="journal article" year="1987">Colbert, 1987</bibRefCitation>
). The
<typeStatus id="CC72D597723CFFDCB1963F9942B5FE78" pageId="2" pageNumber="337" type="holotype">holotype</typeStatus>
of
<taxonomicName id="D4C910B6723CFFDCB3CF3FB94325FE78" authorityName="Parrington" authorityYear="1935" box="[230,336,486,508]" class="Reptilia" family="Protorosauridae" genus="Prolacerta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Protorosauria" pageId="2" pageNumber="337" phylum="Chordata" rank="species" species="broomi">
<emphasis id="21BDB727723CFFDCB3CF3FB94325FE78" box="[230,336,486,508]" italics="true" pageId="2" pageNumber="337">P. broomi</emphasis>
</taxonomicName>
came from Harrismith Commonage, Harrismith District,
<collectingRegion id="D10DA5D7723CFFDCB2A53C5B4078FD99" box="[396,525,517,538]" country="South Africa" name="Free State" pageId="2" pageNumber="337">Free State</collectingRegion>
,
<collectingCountry id="6BDE2BA5723CFFDCB1093C5B40CFFD98" box="[544,698,517,539]" name="South Africa" pageId="2" pageNumber="337">South Africa</collectingCountry>
. The specimens used in this study were collected from the following South African localities: BP/1/471, Honingkrans (Hueningkrans), Burgersdorp District; BP/1/ 2675, Harrismith Commonage; BP/1/4504a, Fairydale, Bethulie District; BP/1/5066, Queens Hill, Harrismith District; BP/1/5375, Rietport, Dewetsdorp District; UCMP 37151, Big Bank, Harrismith District (UCMP locality
<accessionNumber id="0C9AF6D6723CFFDCB3C33CA44330FC93" box="[234,325,762,784]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/V36115" pageId="2" pageNumber="337" type="EnaNcbi">V36115</accessionNumber>
).
</paragraph>
</subSubSection>
</treatment>
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