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<mods:title id="067F4D4F628F4A99674D17D85D7D1ACB">Sciapodinae from the Himalayan region with description of nine new species from Tibet (Diptera: Dolichopodidae)</mods:title>
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<treatment id="03D987E7FFE2FF9E31CDFAA63C1DFE2F" ID-DOI="http://doi.org/10.5281/zenodo.5930453" ID-GBIF-Taxon="154553431" ID-Zenodo-Dep="5930453" LSID="urn:lsid:plazi:treatment:03D987E7FFE2FF9E31CDFAA63C1DFE2F" httpUri="http://treatment.plazi.org/id/03D987E7FFE2FF9E31CDFAA63C1DFE2F" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<subSubSection id="C36A657AFFE2FF9F31CDFAA63D6DFAD7" box="[151,555,1297,1323]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="8BCF36F1FFE2FF9F31CDFAA63D6DFAD7" blockId="18.[151,555,1297,1323]" box="[151,555,1297,1323]" pageId="18" pageNumber="19">
<heading id="D087819DFFE2FF9F31CDFAA63D6DFAD7" bold="true" box="[151,555,1297,1323]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
<emphasis id="B904EAE3FFE2FF9F31CDFAA63D6DFAD7" bold="true" box="[151,555,1297,1323]" pageId="18" pageNumber="19">
Genus
<taxonomicName id="4C704D72FFE2FF9F31B4FAA63D6DFAD7" ID-CoL="3T3R" ID-ENA="192486" authority="Bigot, 1859" authorityName="Bigot" authorityYear="1859" box="[238,555,1297,1323]" class="Insecta" family="Dolichopodidae" genus="Condylostylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B904EAE3FFE2FF9F31B4FAA63EDEFAD7" bold="true" box="[238,408,1297,1323]" italics="true" pageId="18" pageNumber="19">Condylostylus</emphasis>
Bigot, 1859
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36A657AFFE2FF9E31CDFAEF3F99FE77" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="reference_group">
<paragraph id="8BCF36F1FFE2FF9F31CDFAEF3DB9F991" blockId="18.[151,1437,1367,2005]" pageId="18" pageNumber="19">
<taxonomicName id="4C704D72FFE2FF9F31CDFAEF3E70FA8C" ID-CoL="3T3R" authorityName="Bigot" authorityYear="1859" box="[151,310,1368,1392]" class="Insecta" family="Dolichopodidae" genus="Condylostylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B904EAE3FFE2FF9F31CDFAEF3E70FA8C" box="[151,310,1368,1392]" italics="true" pageId="18" pageNumber="19">Condylostylus</emphasis>
</taxonomicName>
is the second largest genus in
<taxonomicName id="4C704D72FFE2FF9F33C3FAE03C63FA8C" ID-CoL="8YQ4J" box="[665,805,1367,1392]" class="Insecta" family="Dolichopodidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="subFamily" subFamily="Sciapodinae">Sciapodinae</taxonomicName>
, with a high diversity in the Neotropical region, where 70% of the species occur (
<bibRefCitation id="EFE14B00FFE2FF9F308EFACB3D21FA68" author="Bickel, D. J." box="[468,615,1404,1429]" pageId="18" pageNumber="19" pagination="1 - 394" refId="ref24202" refString="Bickel, D. J. (1994) The Australian Sciapodinae (Diptera: Dolichopodidae), with a review of the Oriental and Australasian faunas, and a world conspectus of the subfamily. Records of the Australian Museum Supplement, 21, 1 - 394. https: // doi. org / 10.3853 / j. 0812 - 7387.21.1994.50" type="journal article" year="1994">Bickel 1994</bibRefCitation>
). Though the genus includes over 250 species, only eight species are recorded in the Himalayas region, of which 2 pairs of species are suggested to be synonyms.
<taxonomicName id="4C704D72FFE2FF9F35A7FA173E21FA21" ID-CoL="XL3V" authority="Becker" authorityName="Becker" authorityYear="1922" class="Insecta" family="Dolichopodidae" genus="Condylostylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="conspectus">
<emphasis id="B904EAE3FFE2FF9F35A7FA173E54FA20" italics="true" pageId="18" pageNumber="19">Condylostylus conspectus</emphasis>
Becker
</taxonomicName>
and
<taxonomicName id="4C704D72FFE2FF9F30C6FA733D73FA20" ID-CoL="XL33" authority="Parent" authorityName="Parent" authorityYear="1934" box="[412,565,1476,1500]" class="Insecta" family="Dolichopodidae" genus="Condylostylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="caii">
<emphasis id="B904EAE3FFE2FF9F30C6FA733EA0FA20" box="[412,486,1476,1500]" italics="true" pageId="18" pageNumber="19">C. caii</emphasis>
Parent
</taxonomicName>
were examined by
<bibRefCitation id="EFE14B00FFE2FF9F3257FA733CE1FA20" author="Bickel, D. J." box="[781,935,1476,1501]" pageId="18" pageNumber="19" pagination="1 - 394" refId="ref24202" refString="Bickel, D. J. (1994) The Australian Sciapodinae (Diptera: Dolichopodidae), with a review of the Oriental and Australasian faunas, and a world conspectus of the subfamily. Records of the Australian Museum Supplement, 21, 1 - 394. https: // doi. org / 10.3853 / j. 0812 - 7387.21.1994.50" type="journal article" year="1994">Bickel (1994)</bibRefCitation>
who suggested the species to be very similar and probably conspecific. No additional
<taxonomicName id="4C704D72FFE2FF9F3326FA5F3D8BFA03" ID-CoL="XL33" authorityName="Parent" authorityYear="1934" box="[636,717,1512,1535]" class="Insecta" family="Dolichopodidae" genus="Condylostylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="caii">
<emphasis id="B904EAE3FFE2FF9F3326FA5F3D8BFA03" box="[636,717,1512,1535]" italics="true" pageId="18" pageNumber="19">C. caii</emphasis>
</taxonomicName>
have been reported since
<bibRefCitation id="EFE14B00FFE2FF9F3554FA503BE9F9FC" author="Bickel, D. J." box="[1038,1199,1511,1536]" pageId="18" pageNumber="19" pagination="1 - 394" refId="ref24202" refString="Bickel, D. J. (1994) The Australian Sciapodinae (Diptera: Dolichopodidae), with a review of the Oriental and Australasian faunas, and a world conspectus of the subfamily. Records of the Australian Museum Supplement, 21, 1 - 394. https: // doi. org / 10.3853 / j. 0812 - 7387.21.1994.50" type="journal article" year="1994">Bickel (1994)</bibRefCitation>
and the differences between the two species mentioned by
<bibRefCitation id="EFE14B00FFE2FF9F3328F9BA3C54F9D8" author="Parent, O." box="[626,786,1548,1572]" pageId="18" pageNumber="19" pagination="113 - 140" refId="ref24991" refString="Parent, O. (1934) Especes nouvelles de Dipteres Dolichopodides. Encyclopedie Entomologique (B II) Diptera, 7, 113 - 140." type="journal article" year="1934">Parent (1934)</bibRefCitation>
are actually not sound based on our inspection of
<emphasis id="B904EAE3FFE2FF9F34D8F9BB3E5FF9B4" italics="true" pageId="18" pageNumber="19">
<taxonomicName id="4C704D72FFE2FF9F34D8F9BB3E53F9B4" ID-CoL="XL3V" authorityName="Becker" authorityYear="1922" class="Insecta" family="Dolichopodidae" genus="Condylostylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="conspectus">C. conspectus</taxonomicName>
.
</emphasis>
We agree with Bickels assessment that they are possibly synonyms; however, we do not make the decision since we have not studied the
<typeStatus id="54CB8853FFE2FF9F3311F9E23D3DF991" box="[587,635,1621,1645]" pageId="18" pageNumber="19">type</typeStatus>
specimens.
</paragraph>
<paragraph id="8BCF36F1FFE2FF9F319DF9CF3D57F8E3" blockId="18.[151,1437,1367,2005]" pageId="18" pageNumber="19">
<taxonomicName id="4C704D72FFE2FF9F319DF9CF3D70F96C" ID-CoL="XL7N" authority="Becker" authorityName="Becker" box="[199,566,1655,1680]" class="Insecta" family="Dolichopodidae" genus="Condylostylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="nebulosus">
<emphasis id="B904EAE3FFE2FF9F319DF9CF3E98F96C" box="[199,478,1656,1680]" italics="true" pageId="18" pageNumber="19">Condylostylus nebulosus</emphasis>
Becker
</taxonomicName>
and
<taxonomicName id="4C704D72FFE2FF9F332BF9CF3C0DF973" ID-CoL="5ZRPJ" authority="Parent" authorityName="Parent" authorityYear="1929" box="[625,843,1656,1680]" class="Insecta" family="Dolichopodidae" genus="Condylostylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="luteicoxa">
<emphasis id="B904EAE3FFE2FF9F332BF9CF3DBDF96C" box="[625,763,1656,1680]" italics="true" pageId="18" pageNumber="19">C. luteicoxa</emphasis>
Parent
</taxonomicName>
only differ in the mid tarsus and the description by Parent has some discrepancies with the
<typeStatus id="54CB8853FFE2FF9F3305F92B3D82F949" box="[607,708,1692,1717]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
, which led
<bibRefCitation id="EFE14B00FFE2FF9F3213F92B3CA3F948" author="Bickel, D. J." box="[841,997,1692,1717]" pageId="18" pageNumber="19" pagination="1 - 394" refId="ref24202" refString="Bickel, D. J. (1994) The Australian Sciapodinae (Diptera: Dolichopodidae), with a review of the Oriental and Australasian faunas, and a world conspectus of the subfamily. Records of the Australian Museum Supplement, 21, 1 - 394. https: // doi. org / 10.3853 / j. 0812 - 7387.21.1994.50" type="journal article" year="1994">Bickel (1994)</bibRefCitation>
and
<bibRefCitation id="EFE14B00FFE2FF9F3545F92A3BA8F948" author="Yang, D. &amp; Zhang, L. &amp; Wang, M. &amp; Zhu, Y." box="[1055,1262,1692,1717]" pageId="18" pageNumber="19" pagination="111 - 291" refId="ref25610" refString="Yang, D., Zhang, L., Wang, M. &amp; Zhu, Y. (2011) Fauna Sinica Insecta Uol. 53. Diptera Dolichopodidae. Science Press, Beijing, pp. 111 - 291." type="book chapter" year="2011">
Yang
<emphasis id="B904EAE3FFE2FF9F3538F92A3BD2F948" box="[1122,1172,1692,1716]" italics="true" pageId="18" pageNumber="19">et al</emphasis>
. (2011)
</bibRefCitation>
to suggest that they belong to the same species. But here they are still treated as separate species because the
<typeStatus id="54CB8853FFE2FF9F359CF9083A75F924" box="[1222,1331,1727,1752]" pageId="18" pageNumber="19" type="holotype">holotypes</typeStatus>
have not been examined and there are differences in the descriptions. Similar to
<taxonomicName id="4C704D72FFE2FF9F32E9F9533BF2F901" ID-CoL="9H6V2" authority=", Nyingchi" authorityName="Nyingchi" box="[947,1204,1764,1789]" class="Insecta" family="Dolichopodidae" genus="Chrysosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B904EAE3FFE2FF9F32E9F9533B79F900" box="[947,1087,1764,1788]" italics="true" pageId="18" pageNumber="19">Chrysosoma</emphasis>
, Nyingchi
</taxonomicName>
is the only region in
<collectingRegion id="49B4F813FFE2FF9F31CDF8B03F97F8DC" box="[151,209,1799,1824]" country="China" name="Xizang" pageId="18" pageNumber="19">Tibet</collectingRegion>
we collected
<taxonomicName id="4C704D72FFE2FF9F3036F8BF3D4DF8DC" ID-CoL="3T3R" authorityName="Bigot" authorityYear="1859" box="[364,523,1800,1824]" class="Insecta" family="Dolichopodidae" genus="Condylostylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="B904EAE3FFE2FF9F3036F8BF3D4DF8DC" box="[364,523,1800,1824]" italics="true" pageId="18" pageNumber="19">Condylostylus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BCF36F1FFE2FF9E319DF89B3F99FE77" blockId="18.[151,1437,1367,2005]" lastBlockId="19.[151,1436,151,467]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
Species of this genus can be distinguished by the following features: Head: frons with 1 strong vt arising from strongly raised tubercle on frons in both sexes; face sometimes bulging beneath antenna; male clypeus broad and close to eye margin; arista-like stylus usually dorsal. Thorax usually with 23 pairs of long acr, 5 strong dc; strong lateral and median sc. Legs: Fore coxa with 3 strong black distolateral bristles; fore tibia usually without strong bristles; TII with obvious ad and pd; male usually with crocheted hairs on basal half of mid tibia and tarsus; hind tarsus sometimes with flattened pad-like tarsomeres. Wing often with dark brown bands, sometimes with enclosed window on crossvein dm-cu. M
<subScript id="17F434B4FFE3FF9E30A3FF7C3D47FF25" attach="left" box="[505,513,203,217]" fontSize="6" pageId="19" pageNumber="20">1</subScript>
characteristically with sharp, almost right angle bend at branch with M
<subScript id="17F434B4FFE3FF9E3446FF7C3A62FF25" attach="right" box="[1308,1316,203,217]" fontSize="6" pageId="19" pageNumber="20">2</subScript>
; crossvein dm-cu usually straight, sometimes slightly sinuate. Abdomen: Tergum VII well developed, sternum VII strongly reduced, membranous; hypopygium often relatively small compared with other
<taxonomicName id="4C704D72FFE3FF9E3528FEB13BB8FEE3" ID-CoL="8YQ4J" box="[1138,1278,262,287]" class="Insecta" family="Dolichopodidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="subFamily" subFamily="Sciapodinae">Sciapodinae</taxonomicName>
, and basally encapsulated within segment VII; hypandrium often short and broad, lateral arm of hypandrium reduced or absent; cercus usually simple, commonly elongate and filamentous, seldom clavate and expanded (
<bibRefCitation id="EFE14B00FFE3FF9E35D7FEF93A5FFE9B" author="Bickel, D. J." box="[1165,1305,334,359]" pageId="19" pageNumber="20" pagination="1 - 394" refId="ref24202" refString="Bickel, D. J. (1994) The Australian Sciapodinae (Diptera: Dolichopodidae), with a review of the Oriental and Australasian faunas, and a world conspectus of the subfamily. Records of the Australian Museum Supplement, 21, 1 - 394. https: // doi. org / 10.3853 / j. 0812 - 7387.21.1994.50" type="journal article" year="1994">Bickel 1994</bibRefCitation>
;
<bibRefCitation id="EFE14B00FFE3FF9E347FFEE73F95FE77" author="Yang, D. &amp; Zhang, L. &amp; Wang, M. &amp; Zhu, Y." pageId="19" pageNumber="20" pagination="111 - 291" refId="ref25610" refString="Yang, D., Zhang, L., Wang, M. &amp; Zhu, Y. (2011) Fauna Sinica Insecta Uol. 53. Diptera Dolichopodidae. Science Press, Beijing, pp. 111 - 291." type="book chapter" year="2011">
Yang
<emphasis id="B904EAE3FFE3FF9E343EFEE73AD3FE9B" box="[1380,1429,335,359]" italics="true" pageId="19" pageNumber="20">et al</emphasis>
. 2011
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36A657AFFE3FF9E319DFE203C1DFE2F" pageId="19" pageNumber="20" type="description">
<paragraph id="8BCF36F1FFE3FF9E319DFE203C1DFE2F" blockId="19.[151,1436,151,467]" pageId="19" pageNumber="20">
A key to
<taxonomicName id="4C704D72FFE3FF9E3074FE203E8BFE53" authorityName="Bigot" authorityYear="1859" box="[302,461,407,431]" class="Insecta" family="Dolichopodidae" genus="Condylostylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B904EAE3FFE3FF9E3074FE203E8BFE53" box="[302,461,407,431]" italics="true" pageId="19" pageNumber="20">Condylostylus</emphasis>
</taxonomicName>
from the Himalayan region is provided.
<taxonomicName id="4C704D72FFE3FF9E32CDFE203BB6FE53" authority="Parent" authorityName="Parent" authorityYear="1932" box="[919,1264,407,431]" class="Insecta" family="Dolichopodidae" genus="Condylostylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="albifrons">
<emphasis id="B904EAE3FFE3FF9E32CDFE203BE7FE53" box="[919,1185,407,431]" italics="true" pageId="19" pageNumber="20">Condylostylus albifrons</emphasis>
Parent
</taxonomicName>
is not included because this species record in based only on female specimens.
</paragraph>
</subSubSection>
</treatment>
</document>