270 lines
29 KiB
XML
270 lines
29 KiB
XML
<document id="FD341DD8E5F42533865DDD5B5B403EE1" ID-CLB-Dataset="29513" ID-DOI="10.11646/zootaxa.4483.2.10" ID-GBIF-Dataset="5b0dfb7c-9e7e-4e3e-b438-5e25073759ba" ID-ISSN="1175-5326" ID-Zenodo-Dep="1437631" ID-ZooBank="0618503B-5824-4A23-980D-317A4B5DFAF4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538122456990" checkinUser="plazi" docAuthor="Liu, Qin, Zhong, Guang-Hui, Wang, Ping, Liu, Yang & Guo, Peng" docDate="2018" docId="03D987CCFFF6FFE7B080FD35396DFEA7" docLanguage="en" docName="zootaxa.4483.2.10.pdf" docOrigin="Zootaxa 4483 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hebius yanbianensis Liu & Zhong & Wang & Liu & Guo 2018, sp. nov." docType="treatment" docVersion="5" lastPageNumber="392" masterDocId="FFE0FFB4FFF5FFE0B017FFCF3D6CFFCF" masterDocTitle="A new species of the genus Hebius (Squamata: Colubridae) from Sichuan, China" masterLastPageNumber="394" masterPageNumber="385" pageNumber="388" updateTime="1698556628485" updateUser="ExternalLinkService">
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<mods:titleInfo id="6AF1DFE61B50D8279CE8448881340787">
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<mods:title id="FE411599963BAE15A121C7345BA6DBDF">A new species of the genus Hebius (Squamata: Colubridae) from Sichuan, China</mods:title>
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<mods:namePart id="68EE9C705965F918CDF23AD99A8298B7">Liu, Qin</mods:namePart>
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<mods:roleTerm id="6959F0F41F9ADAB6362CF0E50F3AF5A3">Author</mods:roleTerm>
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<mods:namePart id="C8998224A22B21395E83B39BB55921D5">Zhong, Guang-Hui</mods:namePart>
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<mods:namePart id="273EFA6486CF43334C4F9158F6CF9A5A">Wang, Ping</mods:namePart>
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<mods:namePart id="7FF4423091C3CFCF9BECA266BFEAC7EB">Liu, Yang</mods:namePart>
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<mods:namePart id="9095A1336463F1125F57991D5C6C5902">Guo, Peng</mods:namePart>
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<mods:title id="C86DC42C3EB17072A5376A4F0C2CBC43">Zootaxa</mods:title>
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<mods:part id="A04C277F475978460F4B9649076CBD6C">
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<mods:date id="F72F54E4E3FA388BA2A9472AB2869894">2018</mods:date>
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<mods:number id="D3DFF04BEC0B9E07D52E14D5C17008CA">2018-09-21</mods:number>
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<mods:number id="0360BD82B5A18BAC3BC6AD4FEC227CA4">4483</mods:number>
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<treatment id="03D987CCFFF6FFE7B080FD35396DFEA7" ID-GBIF-Taxon="148403585" LSID="urn:lsid:plazi:treatment:03D987CCFFF6FFE7B080FD35396DFEA7" httpUri="http://treatment.plazi.org/id/03D987CCFFF6FFE7B080FD35396DFEA7" lastPageId="7" lastPageNumber="392" pageId="3" pageNumber="388">
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<subSubSection id="C36A6551FFF6FFE3B080FD353C7DFCF9" pageId="3" pageNumber="388" type="nomenclature">
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<paragraph id="8BCF36DAFFF6FFE3B080FD353C9BFCDB" blockId="3.[151,503,762,822]" box="[151,503,762,788]" pageId="3" pageNumber="388">
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<heading id="D08781B6FFF6FFE3B080FD353C9BFCDB" bold="true" box="[151,503,762,788]" fontSize="11" level="1" pageId="3" pageNumber="388" reason="1">
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<emphasis id="B904EAC8FFF6FFE3B080FD353C9BFCDB" bold="true" box="[151,503,762,788]" pageId="3" pageNumber="388">
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<taxonomicName id="4C704D59FFF6FFE3B080FD353CFFFCDB" authority="Liu & Zhong & Wang & Liu & Guo, 2018" authorityName="Liu & Zhong & Wang & Liu & Guo" authorityYear="2018" box="[151,403,762,788]" class="Reptilia" family="Colubridae" genus="Hebius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="388" phylum="Chordata" rank="species" species="yanbianensis" status="sp. nov.">
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<emphasis id="B904EAC8FFF6FFE3B080FD353CFFFCDB" bold="true" box="[151,403,762,788]" italics="true" pageId="3" pageNumber="388">Hebius yanbianensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23757B3FFF6FFE3B18DFD353C9BFCDB" box="[410,503,762,788]" pageId="3" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BCF36DAFFF6FFE3B080FCD23C7DFCF9" blockId="3.[151,503,762,822]" box="[151,273,797,822]" pageId="3" pageNumber="388">
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(
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<figureCitation id="134B2A5FFFF6FFE3B088FCD23C65FCF9" box="[159,265,797,822]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStart-3="FIGURE 6" captionStart-4="FIGURE 7" captionStartId-0="3.[151,250,2004,2026]" captionStartId-1="4.[151,250,1521,1543]" captionStartId-2="5.[151,250,1431,1453]" captionStartId-3="6.[151,250,1983,2005]" captionStartId-4="7.[151,250,1401,1423]" captionTargetBox-0="[150,1436,1018,1982]" captionTargetBox-1="[161,1431,205,1498]" captionTargetBox-2="[301,1290,208,1404]" captionTargetBox-3="[300,1294,603,1956]" captionTargetBox-4="[150,1436,675,1379]" captionTargetId-0="figure@3.[150,1436,1018,1982]" captionTargetId-1="figure@4.[155,1432,193,1499]" captionTargetId-2="figure@5.[289,1298,193,1410]" captionTargetId-3="figure@6.[288,1299,591,1961]" captionTargetId-4="figure@7.[150,1436,415,1379]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionText-0="FIGURE 3. General view of the holotype of Hebius yanbianensis sp. nov. (YBU 15018) in life. Photo by G.H. Zhong." captionText-1="FIGURE 4. Dorsal (top) and ventral (bottom) views of the holotype of Hebius yanbianensis sp. nov. (YBU15018) in preservative. Photo by P. Guo." captionText-2="FIGURE 5. Dorsal (A), ventral (B), and lateral (B) views of the head of the holotype of Hebius yanbianensis sp. nov. in preservative. Photo by P. Guo." captionText-3="FIGURE 6. Dorsal (A), ventral (B) and lateral (C) views of CT scanned skull of the holotype of Hebius yanbianensis sp. nov." captionText-4="FIGURE 7. Habitat of Hebius yanbianensis sp. nov. Photo by G.H. Zhong." httpUri-0="https://zenodo.org/record/1437637/files/figure.png" httpUri-1="https://zenodo.org/record/1437639/files/figure.png" httpUri-2="https://zenodo.org/record/1437643/files/figure.png" httpUri-3="https://zenodo.org/record/1437645/files/figure.png" httpUri-4="https://zenodo.org/record/1437647/files/figure.png" pageId="3" pageNumber="388">Figs. 3–7</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C36A6551FFF6FFE3B080FCAA3CFBFC09" pageId="3" pageNumber="388" type="materials_examined">
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<paragraph id="8BCF36DAFFF6FFE3B080FCAA3CFBFC09" blockId="3.[151,1436,869,966]" pageId="3" pageNumber="388">
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<materialsCitation id="3B183C87FFF6FFE3B080FCAA3CFBFC09" ID-GBIF-Occurrence="1913860762" collectingDate="2015-03" collectorName="Guanghui Zhong" country="China" county="Yanbian County" elevation="1974" latitude="101.584564" location="Zemulong Town" longLatPrecision="1" longitude="27.231112" pageId="3" pageNumber="388" specimenCount="1" stateProvince="South Sichuan Province" typeStatus="holotype">
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<emphasis id="B904EAC8FFF6FFE3B080FCAA3C67FCB2" bold="true" box="[151,267,869,893]" pageId="3" pageNumber="388">
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<typeStatus id="54CB8878FFF6FFE3B080FCAA3C6AFCB2" box="[151,262,869,893]" pageId="3" pageNumber="388" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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YBU 15018, an adult male, collected by
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<collectorName id="2685530CFFF6FFE3B2F5FCAA3ECEFCB1" box="[738,930,869,894]" pageId="3" pageNumber="388">Guanghui Zhong</collectorName>
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and Ping Wang in
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<collectingDate id="EF8AE9F2FFF6FFE3B46CFCAA3868FCB1" box="[1147,1284,869,894]" pageId="3" pageNumber="388" value="2015-03">
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<date id="FFCE101AFFF6FFE3B46CFCAA3868FCB1" box="[1147,1284,869,894]" pageId="3" pageNumber="388" value="2015-03">
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March
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||
<quantity id="4C889B3FFFF6FFE3B4DBFCAA3868FCB1" box="[1228,1284,869,894]" metricMagnitude="1" metricUnit="m" metricValue="5.1181" pageId="3" pageNumber="388" unit="in" value="2015.0">2015</quantity>
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</date>
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||
</collectingDate>
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||
in
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<location id="8EAF6001FFF6FFE3B53EFCAA3DB5FC6E" LSID="urn:lsid:plazi:treatment:03D987CCFFF6FFE7B080FD35396DFEA7:8EAF6001FFF6FFE3B53EFCAA3DB5FC6E" country="China" county="Yanbian County" latitude="101.584564" longLatPrecision="1" longitude="27.231112" name="Zemulong Town" pageId="3" pageNumber="388" stateProvince="South Sichuan Province">Zemulong Town</location>
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,
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<collectingCounty id="62AE4E56FFF6FFE3B0F1FC463CF2FC6D" box="[230,414,905,930]" pageId="3" pageNumber="388">Yanbian County</collectingCounty>
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(
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||
<geoCoordinate id="EE44501DFFF6FFE3B1B9FC453F1DFC6D" box="[430,625,905,930]" direction="north" orientation="latitude" pageId="3" pageNumber="388" precision="1" value="101.584564">N 101°35′04.43″</geoCoordinate>
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,
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<geoCoordinate id="EE44501DFFF6FFE3B269FC453E5CFC6D" box="[638,816,905,930]" direction="east" orientation="longitude" pageId="3" pageNumber="388" precision="1" value="27.231112">E 27°13′52.00″</geoCoordinate>
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),
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<collectingRegion id="49B4F838FFF6FFE3B352FC463935FC6D" box="[837,1113,905,930]" inferredBySuffix="true" pageId="3" pageNumber="388">
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South
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<collectingRegion id="49B4F838FFF6FFE3B398FC463935FC6D" box="[911,1113,905,930]" country="China" name="Sichuan" pageId="3" pageNumber="388">Sichuan Province</collectingRegion>
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</collectingRegion>
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,
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<collectingCountry id="F367764AFFF6FFE3B472FC4639C7FC6D" box="[1125,1195,905,930]" name="China" pageId="3" pageNumber="388">China</collectingCountry>
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, at an elevation of
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<elevation id="005DD1E9FFF6FFE3B590FC453CFFFC09" metricMagnitude="3" metricUnit="m" metricValue="1.974" pageId="3" pageNumber="388" unit="m" value="1974.0">
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<quantity id="4C889B3FFFF6FFE3B590FC453DB2FC0A" metricMagnitude="3" metricUnit="m" metricValue="1.974" pageId="3" pageNumber="388" unit="m" value="1974.0">1, 974 m</quantity>
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above sea level
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</elevation>
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.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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||
<caption id="DF0F6652FFF6FFE3B080F81B3827F825" box="[151,1355,2004,2026]" httpUri="https://zenodo.org/record/1437637/files/figure.png" pageId="3" pageNumber="388" startId="3.[151,250,2004,2026]" targetBox="[150,1436,1018,1982]" targetPageId="3">
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<paragraph id="8BCF36DAFFF6FFE3B080F81B3827F825" blockId="3.[151,1355,2004,2026]" box="[151,1355,2004,2026]" pageId="3" pageNumber="388">
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<emphasis id="B904EAC8FFF6FFE3B080F81B3C7EF826" bold="true" box="[151,274,2004,2026]" pageId="3" pageNumber="388">FIGURE 3.</emphasis>
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General view of the holotype of
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<taxonomicName id="4C704D59FFF6FFE3B275F81B3E58F826" authority="Liu & Zhong & Wang & Liu & Guo, 2018" authorityName="Liu & Zhong & Wang & Liu & Guo" authorityYear="2018" box="[610,820,2004,2025]" class="Reptilia" family="Colubridae" genus="Hebius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="388" phylum="Chordata" rank="species" species="yanbianensis" status="sp. nov.">
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<emphasis id="B904EAC8FFF6FFE3B275F81B3E58F826" box="[610,820,2004,2025]" italics="true" pageId="3" pageNumber="388">Hebius yanbianensis</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="A23757B3FFF6FFE3B32DF81A3EE7F825" box="[826,907,2005,2026]" pageId="3" pageNumber="388" rank="species">
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||
<emphasis id="B904EAC8FFF6FFE3B32DF81A3EE7F825" bold="true" box="[826,907,2005,2026]" pageId="3" pageNumber="388">sp. nov.</emphasis>
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</taxonomicNameLabel>
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(YBU 15018) in life. Photo by G.H. Zhong.
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</paragraph>
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||
</caption>
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||
<caption id="DF0F6652FFF1FFE4B080FA3E3CA7F9EA" httpUri="https://zenodo.org/record/1437639/files/figure.png" pageId="4" pageNumber="389" startId="4.[151,250,1521,1543]" targetBox="[161,1431,205,1498]" targetPageId="4">
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<paragraph id="8BCF36DAFFF1FFE4B080FA3E3CA7F9EA" blockId="4.[151,1436,1521,1573]" pageId="4" pageNumber="389">
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||
<emphasis id="B904EAC8FFF1FFE4B080FA3E3C74F9C9" bold="true" box="[151,280,1521,1543]" pageId="4" pageNumber="389">FIGURE 4.</emphasis>
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Dorsal (top) and ventral (bottom) views of the holotype of
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<taxonomicName id="4C704D59FFF1FFE4B3A7FA3E39E4F9C9" authority="Liu & Zhong & Wang & Liu & Guo, 2018" authorityName="Liu & Zhong & Wang & Liu & Guo" authorityYear="2018" box="[944,1160,1521,1542]" class="Reptilia" family="Colubridae" genus="Hebius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="389" phylum="Chordata" rank="species" species="yanbianensis" status="sp. nov.">
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||
<emphasis id="B904EAC8FFF1FFE4B3A7FA3E39E4F9C9" box="[944,1160,1521,1542]" italics="true" pageId="4" pageNumber="389">Hebius yanbianensis</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="A23757B3FFF1FFE4B483FA3E3986F9C9" box="[1172,1258,1521,1542]" pageId="4" pageNumber="389" rank="species">
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||
<emphasis id="B904EAC8FFF1FFE4B483FA3E3986F9C9" bold="true" box="[1172,1258,1521,1542]" pageId="4" pageNumber="389">sp. nov.</emphasis>
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</taxonomicNameLabel>
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(YBU15018) in preservative. Photo by P. Guo.
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</paragraph>
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||
</caption>
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<subSubSection id="C36A6551FFF1FFE4B0D0F99E3E53F878" pageId="4" pageNumber="389" type="diagnosis">
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<paragraph id="8BCF36DAFFF1FFE4B0D0F99E3E53F878" blockId="4.[151,1436,1617,2011]" pageId="4" pageNumber="389">
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||
<emphasis id="B904EAC8FFF1FFE4B0D0F99E3C2DF9A5" bold="true" box="[199,321,1617,1642]" pageId="4" pageNumber="389">Diagnosis.</emphasis>
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<taxonomicName id="4C704D59FFF1FFE4B15FF99D3F5CF9A5" authority="Liu & Zhong & Wang & Liu & Guo, 2018" authorityName="Liu & Zhong & Wang & Liu & Guo" authorityYear="2018" box="[328,560,1618,1642]" class="Reptilia" family="Colubridae" genus="Hebius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="389" phylum="Chordata" rank="species" species="yanbianensis" status="sp. nov.">
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||
<emphasis id="B904EAC8FFF1FFE4B15FF99D3F5CF9A5" box="[328,560,1618,1642]" italics="true" pageId="4" pageNumber="389">Hebius yanbianensis</emphasis>
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||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23757B3FFF1FFE4B22FF99D3FFDF9A5" box="[568,657,1618,1642]" pageId="4" pageNumber="389" rank="species">
|
||
<emphasis id="B904EAC8FFF1FFE4B22FF99D3FFDF9A5" bold="true" box="[568,657,1618,1642]" pageId="4" pageNumber="389">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
can be distinguished from its congeners on the basis of the following combination of characters: 1) TL/SVL ratio 0.35; 2) three postoculars; 3) six temporals in formula 1/(1+1) +1+2; 4) eight supralabials, 4th and 5th in contact with the eye, 6th largest;5) ten infralabials, the first five bordering the anterior chinshields; 6) 172 ventrals (plus one preventral); 7) cloacal plate divided; 8) 90 pairs of subcaudals; 9) dorsal scales in 19-19-17 rows, all weakly keeled except the outer two; 10) reduction of dorsal scale rows from 19 to 17 scale (VS19TO17) at the position above 99th ventral scale; 11) reduction of the dorsal tail scales from 8 to 6 scale rows (SC8TO6) at the ventral position of 18th subcaudal, and from 8 to 6 (SC6TO4) at the position above 37th subcaudal; 12) postorbital bones not touching frontals, the parietal ridge weakly developed; the end of the supratemporal bones extending beyond the braincase; maxillary teeth 23-25, the last two distinctly enlarged without diastema between the last two and the anterior teeth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A6551FFF1FFE7B0D0F80D3E52FF1A" lastPageId="7" lastPageNumber="392" pageId="4" pageNumber="389" type="description">
|
||
<paragraph id="8BCF36DAFFF1FFE4B0D0F80D380BF814" blockId="4.[151,1436,1617,2011]" box="[199,1383,1986,2011]" pageId="4" pageNumber="389">
|
||
<emphasis id="B904EAC8FFF1FFE4B0D0F80D3C88F814" bold="true" box="[199,484,1986,2011]" pageId="4" pageNumber="389">
|
||
Description of
|
||
<typeStatus id="54CB8878FFF1FFE4B161F80C3CB3F814" box="[374,479,1987,2011]" pageId="4" pageNumber="389" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
A male specimen with SVL
|
||
<quantity id="4C889B3FFFF1FFE4B33EF80C3EE5F815" box="[809,905,1987,2011]" metricMagnitude="-1" metricUnit="m" metricValue="4.2" pageId="4" pageNumber="389" unit="mm" value="420.0">420 mm</quantity>
|
||
; tail complete, TL
|
||
<quantity id="4C889B3FFFF1FFE4B44AF80D39D0F815" box="[1117,1212,1986,2011]" metricMagnitude="-1" metricUnit="m" metricValue="1.45" pageId="4" pageNumber="389" unit="mm" value="145.0">145 mm</quantity>
|
||
, TL/SVL 0.35.
|
||
</paragraph>
|
||
<caption id="DF0F6652FFF0FFE5B080FA583CA7FA03" httpUri="https://zenodo.org/record/1437643/files/figure.png" pageId="5" pageNumber="390" startId="5.[151,250,1431,1453]" targetBox="[301,1290,208,1404]" targetPageId="5">
|
||
<paragraph id="8BCF36DAFFF0FFE5B080FA583CA7FA03" blockId="5.[151,1436,1431,1484]" pageId="5" pageNumber="390">
|
||
<emphasis id="B904EAC8FFF0FFE5B080FA583C79FA63" bold="true" box="[151,277,1431,1453]" pageId="5" pageNumber="390">FIGURE 5.</emphasis>
|
||
Dorsal (A), ventral (B), and lateral (B) views of the head of the holotype of
|
||
<taxonomicName id="4C704D59FFF0FFE5B45BFA58384DFA63" authority="Liu & Zhong & Wang & Liu & Guo, 2018" authorityName="Liu & Zhong & Wang & Liu & Guo" authorityYear="2018" box="[1100,1313,1431,1452]" class="Reptilia" family="Colubridae" genus="Hebius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="390" phylum="Chordata" rank="species" species="yanbianensis" status="sp. nov.">
|
||
<emphasis id="B904EAC8FFF0FFE5B45BFA58384DFA63" box="[1100,1313,1431,1452]" italics="true" pageId="5" pageNumber="390">Hebius yanbianensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23757B3FFF0FFE5B53CFA583813FA63" box="[1323,1407,1431,1452]" pageId="5" pageNumber="390" rank="species">
|
||
<emphasis id="B904EAC8FFF0FFE5B53CFA583813FA63" bold="true" box="[1323,1407,1431,1452]" pageId="5" pageNumber="390">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
in preservative. Photo by P. Guo.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCF36DAFFF0FFE5B0D0FA373994F897" blockId="5.[151,1436,1528,1991]" pageId="5" pageNumber="390">
|
||
Head distinct from neck. Rostral visible dorsally. Internasals subtriangular, wider than long. Prefrontals large, extending laterally onto sides of head. Frontal longer than wide. Supraoculars broadly in contact with prefrontals, with supraoculars two-thirds as broad as frontals. Single loreal long, rectangular. Preoculars 2/2, postoculars 3/3. Six temporals, formula 1/(1+1) +1+2 on both sides, anteriorly one long, above 2 short + 1 long + 2 posterior temporals. Supralabials 8/8, the 4th and 5th bordering orbit. Infralabials 10/10, the first
|
||
<quantity id="4C889B3FFFF0FFE5B46EF94439CAF96B" box="[1145,1190,1675,1700]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="5" pageNumber="390" unit="in" value="5.0">5 in</quantity>
|
||
contact with anterior chinshields. Nostrils lateral, nasals undivided. In life, dorsum and upper surface of taildark gray with numerous dorsal scales edged in bright yellow, especially on the sides of the forepart of the body, making the body strongly flecked with yellow; the upper surface of the body less flecked; in posterior part of the body, yellow flecking less conspicuous and limited to aligned blotches on the sides. The abdomen milk-white with a large, black, triangular blotch near the outer end of each ventral, these spots forming a strongly marked ventrolateral series.
|
||
</paragraph>
|
||
<paragraph id="8BCF36DAFFF0FFE5B0D0F8AC39B6F808" blockId="5.[151,1436,1528,1991]" pageId="5" pageNumber="390">
|
||
Dorsal scale rows 19-19-17, all feebly keeled except the outer two rows. Reduction from 19 to 17 dorsal scale rows at the position of the 2nd and 3rd dorsal scale rows and between the 96th and 101
|
||
<taxonomicName id="4C704D59FFF0FFE5B45CF84939D4F86C" box="[1099,1208,1926,1955]" pageId="5" pageNumber="390" rank="subSpecies" subSpecies="ventrals">st ventrals</taxonomicName>
|
||
. Ventrals 172 plus 1 pre-ventral. Subcaudals 90, paired. Cloacal divided. The scale rows reduction formula on the tail:
|
||
</paragraph>
|
||
<paragraph id="8BCF36DAFFF3FFE6B0D7FF6F3F53FF72" blockId="6.[152,600,160,239]" box="[192,575,160,189]" pageId="6" pageNumber="391">5(2 + 3) 18(3 + 4) 36(2 + 3)</paragraph>
|
||
<paragraph id="8BCF36DAFFF3FFE6B08FFF7B3F34FF1E" blockId="6.[152,600,160,239]" box="[152,600,180,209]" pageId="6" pageNumber="391">
|
||
<date id="FFCE101AFFF3FFE6B08FFF7B3CABFF1E" box="[152,455,180,209]" pageId="6" pageNumber="391" value="2010-06-08">10 8 6</date>
|
||
4
|
||
</paragraph>
|
||
<paragraph id="8BCF36DAFFF3FFE6B0AAFF023F2FFF20" blockId="6.[152,600,160,239]" box="[189,579,201,239]" pageId="6" pageNumber="391">
|
||
6
|
||
<emphasis id="B904EAC8FFF3FFE6B0C7FF063DB7FF20" bold="true" box="[208,219,201,239]" pageId="6" pageNumber="391">(</emphasis>
|
||
2 + 3
|
||
<emphasis id="B904EAC8FFF3FFE6B10DFF063C49FF20" bold="true" box="[282,293,201,239]" pageId="6" pageNumber="391">)</emphasis>
|
||
17
|
||
<emphasis id="B904EAC8FFF3FFE6B149FF063C05FF20" bold="true" box="[350,361,201,239]" pageId="6" pageNumber="391">(</emphasis>
|
||
3 + 4
|
||
<emphasis id="B904EAC8FFF3FFE6B1B0FF063CDEFF20" bold="true" box="[423,434,201,239]" pageId="6" pageNumber="391">)</emphasis>
|
||
38
|
||
<emphasis id="B904EAC8FFF3FFE6B1F9FF063C95FF20" bold="true" box="[494,505,201,239]" pageId="6" pageNumber="391">(</emphasis>
|
||
2 + 3
|
||
<emphasis id="B904EAC8FFF3FFE6B22FFF063F2FFF20" bold="true" box="[568,579,201,239]" pageId="6" pageNumber="391">)</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCF36DAFFF3FFE6B0D0FEE53CC6FE38" blockId="6.[151,1437,298,503]" pageId="6" pageNumber="391">
|
||
The skull of the
|
||
<typeStatus id="54CB8878FFF3FFE6B19FFEE53C86FE8C" box="[392,490,298,323]" pageId="6" pageNumber="391" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="134B2A5FFFF3FFE6B1EBFEE53F2EFE8C" box="[508,578,298,323]" captionStart="FIGURE 6" captionStartId="6.[151,250,1983,2005]" captionTargetBox="[300,1294,603,1956]" captionTargetId="figure@6.[288,1299,591,1961]" captionTargetPageId="6" captionText="FIGURE 6. Dorsal (A), ventral (B) and lateral (C) views of CT scanned skull of the holotype of Hebius yanbianensis sp. nov." httpUri="https://zenodo.org/record/1437645/files/figure.png" pageId="6" pageNumber="391">Fig. 6</figureCitation>
|
||
) displays a generalized colubrid pattern (
|
||
<bibRefCitation id="EFE14B2BFFF3FFE6B408FEE539D3FE8C" box="[1055,1215,298,323]" pageId="6" pageNumber="391" refId="ref3444" refString="Cundall, D. (1981) Cranial osteology of the colubrid snake genus Opheodrys. Copeia, 2, 353 - 371. https: // doi. org / 10.2307 / 1444224" type="journal article">Cundall 1981</bibRefCitation>
|
||
). Nasals tend to be triangular. Prefrontals are large with medial dorsal process. Frontal is about twice longer than wide. Large postorbits do not touch frontals. Parietals show to be bowed outward, with its end extending beyond the braincase. The quadrates are much broader anteriorly than posteriorly. There are 23–25 maxillary teeth on both sides, the last two are distinctly enlarged, without diastema between the last two and the anterior teeth. There are about 30 teeth on both sides of dentary.
|
||
</paragraph>
|
||
<caption id="DF0F6652FFF3FFE6B080F87038FBF81A" box="[151,1431,1983,2005]" httpUri="https://zenodo.org/record/1437645/files/figure.png" pageId="6" pageNumber="391" startId="6.[151,250,1983,2005]" targetBox="[300,1294,603,1956]" targetPageId="6">
|
||
<paragraph id="8BCF36DAFFF3FFE6B080F87038FBF81A" blockId="6.[151,1431,1983,2005]" box="[151,1431,1983,2005]" pageId="6" pageNumber="391">
|
||
<emphasis id="B904EAC8FFF3FFE6B080F8703C7EF81B" bold="true" box="[151,274,1983,2005]" pageId="6" pageNumber="391">FIGURE 6.</emphasis>
|
||
Dorsal (A), ventral (B) and lateral (C) views of CT scanned skull of the holotype of
|
||
<taxonomicName id="4C704D59FFF3FFE6B47AF80F382CF81A" authority="Liu & Zhong & Wang & Liu & Guo, 2018" authorityName="Liu & Zhong & Wang & Liu & Guo" authorityYear="2018" box="[1133,1344,1984,2005]" class="Reptilia" family="Colubridae" genus="Hebius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="391" phylum="Chordata" rank="species" species="yanbianensis" status="sp. nov.">
|
||
<emphasis id="B904EAC8FFF3FFE6B47AF80F382CF81A" box="[1133,1344,1984,2005]" italics="true" pageId="6" pageNumber="391">Hebius yanbianensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23757B3FFF3FFE6B551F80F38FBF81A" box="[1350,1431,1984,2005]" pageId="6" pageNumber="391" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCF36DAFFF2FFE7B0D0FF573E52FF1A" blockId="7.[151,1436,151,360]" pageId="7" pageNumber="392">
|
||
<emphasis id="B904EAC8FFF2FFE7B0D0FF573CE9FF7F" bold="true" box="[199,389,151,176]" pageId="7" pageNumber="392">Natural history.</emphasis>
|
||
The specimen was collected on an ecological transition belt between farmland and evergreen forest (
|
||
<figureCitation id="134B2A5FFFF2FFE7B0F1FF733C46FF1B" box="[230,298,188,213]" captionStart="FIGURE 7" captionStartId="7.[151,250,1401,1423]" captionTargetBox="[150,1436,675,1379]" captionTargetId="figure@7.[150,1436,415,1379]" captionTargetPageId="7" captionText="FIGURE 7. Habitat of Hebius yanbianensis sp. nov. Photo by G.H. Zhong." httpUri="https://zenodo.org/record/1437647/files/figure.png" pageId="7" pageNumber="392">Fig. 7</figureCitation>
|
||
). No data on diet and reproduction is available.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A6551FFF2FFE7B0D0FF2F385DFEEF" pageId="7" pageNumber="392" type="etymology">
|
||
<paragraph id="8BCF36DAFFF2FFE7B0D0FF2F385DFEEF" blockId="7.[151,1436,151,360]" pageId="7" pageNumber="392">
|
||
<emphasis id="B904EAC8FFF2FFE7B0D0FF2F3C22FF37" bold="true" box="[199,334,224,248]" pageId="7" pageNumber="392">Etymology.</emphasis>
|
||
The specific name refers to the
|
||
<typeStatus id="54CB8878FFF2FFE7B2A2FF2F3F89FF37" box="[693,741,224,248]" pageId="7" pageNumber="392">type</typeStatus>
|
||
locality of this species, Yanbian County,
|
||
<collectingRegion id="49B4F838FFF2FFE7B4A3FF10387EFF37" box="[1204,1298,223,248]" country="China" name="Sichuan" pageId="7" pageNumber="392">Sichuan</collectingRegion>
|
||
. We suggest the following common names: Yanbian Fulianshe (
|
||
<emphasis id="B904EAC8FFF2FFE7B2CDFEC83E0AFEEF" bold="true" box="[730,870,263,288]" pageId="7" pageNumber="392">ÊṸȒřẘ</emphasis>
|
||
) and Yanbian keelback snake (English).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A6551FFF2FFE7B0D0FEE4396DFEA7" pageId="7" pageNumber="392" type="distribution">
|
||
<paragraph id="8BCF36DAFFF2FFE7B0D0FEE4396DFEA7" blockId="7.[151,1436,151,360]" pageId="7" pageNumber="392">
|
||
<emphasis id="B904EAC8FFF2FFE7B0D0FEE43C0CFE8B" bold="true" box="[199,352,299,324]" pageId="7" pageNumber="392">Distribution.</emphasis>
|
||
This species is currently known only from the
|
||
<typeStatus id="54CB8878FFF2FFE7B379FEE33EF2FE8B" box="[878,926,300,324]" pageId="7" pageNumber="392">type</typeStatus>
|
||
locality, Yanbian (
|
||
<figureCitation id="134B2A5FFFF2FFE7B466FEE439C0FE8B" box="[1137,1196,299,324]" captionStart="FIGURE 1" captionStartId="1.[151,250,1251,1273]" captionTargetBox="[166,1424,210,1216]" captionTargetId="figure@1.[151,1436,195,1227]" captionTargetPageId="1" captionText="FIGURE 1. Map showing the type locality (indicated by a red filled circle) of Hebius yanbianensis sp. nov. in Sichuan Province, China." httpUri="https://zenodo.org/record/1437633/files/figure.png" pageId="7" pageNumber="392">Fig.1</figureCitation>
|
||
) that is located in the Hengduan Mountains valley in
|
||
<collectingRegion id="49B4F838FFF2FFE7B1EFFE803FD3FEA7" box="[504,703,335,360]" country="China" name="Sichuan" pageId="7" pageNumber="392">Sichuan Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="F367764AFFF2FFE7B2DCFE803E62FEA7" box="[715,782,335,360]" name="China" pageId="7" pageNumber="392">China</collectingCountry>
|
||
with dry-hot climate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |