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<document id="70614A8528DD94294F173AD189DA2B2B" ID-CLB-Dataset="45558" ID-DOI="10.5281/zenodo.279913" ID-GBIF-Dataset="95a0854d-800c-4830-92b6-03f7dfce7c17" ID-ISSN="1175-5326" ID-Zenodo-Dep="279913" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460433707244" checkinUser="plazi" docAuthor="Budaeva, Nataliya" docDate="2012" docId="03D5AA06FFAFCF69FF65FC6B1956FCE8" docLanguage="en" docName="zt03176p060.pdf" docOrigin="Zootaxa 3176" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Leptoecia midatlantica Budaeva, 2012, new species" docType="treatment" docVersion="9" lastPageNumber="57" masterDocId="FFECD27EFFA9CF65FFF2FFC91D62FFB4" masterDocTitle="Leptoecia midatlantica, a new species of the deep-sea quill-worms (Polychaeta: Onuphidae: Hyalinoeciinae) from the Mid-Atlantic Ridge" masterLastPageNumber="60" masterPageNumber="45" pageNumber="51" updateTime="1698331748596" updateUser="ExternalLinkService">
<mods:mods id="D4F31AD9EC6D45ADB5D70EB8649E27DA" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D0B828CFE9AEB35EF2B8627B0B29FB59">
<mods:title id="56CE2D60086697E1FE96C8EF976D2720">Leptoecia midatlantica, a new species of the deep-sea quill-worms (Polychaeta: Onuphidae: Hyalinoeciinae) from the Mid-Atlantic Ridge</mods:title>
</mods:titleInfo>
<mods:name id="DEC8602976303F952C8A023FE1BCD5E5" type="personal">
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<mods:namePart id="83CF4CCB3AB81F236A6ABE48937297A6">Budaeva, Nataliya</mods:namePart>
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<mods:title id="6AA3ABCE7A7C36A3C90459F563CDF07C">Zootaxa</mods:title>
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<mods:part id="3ADAF6A174D78458359D483ED1B97083">
<mods:date id="1698751716BAE273F8B1E9B25E11F980">2012</mods:date>
<mods:detail id="F56A8AFEEF8B193C8B57FD2A1BAC6EE9" type="volume">
<mods:number id="C53AB42A79AA9C3D49EF468A6B8C6FA7">3176</mods:number>
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<mods:classification id="5EA18F689C5CF75ED91E45C7070AFF60">journal article</mods:classification>
<mods:identifier id="162D4DB91A55BC0DEF8D1434B3B90CEE" type="CLB-Dataset">45558</mods:identifier>
<mods:identifier id="F3D1FB5F7504D2473372464CAA9854F6" type="DOI">10.5281/zenodo.279913</mods:identifier>
<mods:identifier id="DDDEA315D01AF8FE29B77C40E1599AC2" type="GBIF-Dataset">95a0854d-800c-4830-92b6-03f7dfce7c17</mods:identifier>
<mods:identifier id="7F9EC06787F089EC7B2D9147A3BBD315" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="4404C14C56EA455D055BED014EBC10AC" type="Zenodo-Dep">279913</mods:identifier>
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<treatment id="03D5AA06FFAFCF69FF65FC6B1956FCE8" ID-DOI="http://doi.org/10.5281/zenodo.5695879" ID-GBIF-Taxon="119431172" ID-Zenodo-Dep="5695879" LSID="urn:lsid:plazi:treatment:03D5AA06FFAFCF69FF65FC6B1956FCE8" httpUri="http://treatment.plazi.org/id/03D5AA06FFAFCF69FF65FC6B1956FCE8" lastPageId="12" lastPageNumber="57" pageId="6" pageNumber="51">
<subSubSection id="C366489BFFAFCF63FF65FC6B1C30FC68" pageId="6" pageNumber="51" type="nomenclature">
<paragraph id="8BC31B10FFAFCF63FF65FC6B1F55FC0F" blockId="6.[151,567,930,988]" box="[151,567,930,955]" pageId="6" pageNumber="51">
<heading id="D08BAC7CFFAFCF63FF65FC6B1F55FC0F" bold="true" box="[151,567,930,955]" fontSize="10" level="2" pageId="6" pageNumber="51" reason="2">
<emphasis id="B908C702FFAFCF63FF65FC6B1F55FC0F" bold="true" box="[151,567,930,955]" pageId="6" pageNumber="51">
<taxonomicName id="4C7C6093FFAFCF63FF65FC6B1CFCFC0F" ID-CoL="3TNYW" box="[151,414,930,955]" class="Polychaeta" family="Onuphidae" genus="Leptoecia" kingdom="Animalia" order="Eunicida" pageId="6" pageNumber="51" phylum="Annelida" rank="species" species="midatlantica" status="sp. nov.">
<emphasis id="B908C702FFAFCF63FF65FC6B1CFCFC0F" bold="true" box="[151,414,930,955]" italics="true" pageId="6" pageNumber="51">Leptoecia midatlantica</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A23B7A79FFAFCF63FE5FFC6A1F55FC0F" box="[429,567,930,955]" pageId="6" pageNumber="51" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC31B10FFAFCF63FF65FC0D1C30FC68" blockId="6.[151,567,930,988]" box="[151,338,964,988]" pageId="6" pageNumber="51">
<figureCitation id="13470795FFAFCF63FF65FC0D1C60FC68" box="[151,258,964,988]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStartId-0="5.[151,250,1394,1417]" captionStartId-1="7.[151,250,1744,1767]" captionStartId-2="8.[151,250,1892,1915]" captionStartId-3="9.[151,250,1524,1547]" captionStartId-4="10.[151,250,1624,1647]" captionStartId-5="11.[151,250,1073,1096]" captionStartId-6="13.[151,250,680,703]" captionTargetBox-0="[151,1436,181,1373]" captionTargetBox-1="[151,1436,193,1722]" captionTargetBox-2="[151,1436,193,1871]" captionTargetBox-3="[151,1436,193,1502]" captionTargetBox-4="[151,1436,193,1596]" captionTargetBox-5="[219,1369,327,1053]" captionTargetBox-6="[151,1419,191,649]" captionTargetId-0="figure@5.[151,1436,181,1373]" captionTargetId-1="figure@7.[151,1436,193,1723]" captionTargetId-2="figure@8.[151,1436,193,1871]" captionTargetId-3="figure@9.[151,1436,193,1503]" captionTargetId-4="figure@10.[151,1436,193,1598]" captionTargetId-5="figure@11.[219,1370,327,1053]" captionTargetId-6="figure@13.[151,1436,191,649]" captionTargetPageId-0="5" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionTargetPageId-4="10" captionTargetPageId-5="11" captionTargetPageId-6="13" captionText-0="FIGURE 1. Distribution of Leptoecia midatlantica sp. nov. The black arrow indicates the type location; 2500 m, 3000 m and 4500 m isobaths shown." captionText-1="FIGURE 2. Leptoecia midatlantica sp. nov. A, anterior region of the holotype, ventral view; B, same, dorsal view; C – E shape of the prostomium, ventral view; F, maxillae; G, mandibles; H, pygidium of the holotype, ventral view; I, same, dorsal view; J – L, pygidia, dorsal view." captionText-2="FIGURE 3. Leptoecia midatlantica sp. nov. A, C, anterior region of the body, ventral view; B, same, dorsal view; D, F prostomium and peristomium, ventral view; E, same, dorsal view; G, sensory organs on prostomium, dorsal view; H, posterior region of the body, ventral view; I, pygidium, dorsal view; J pygidial glandular buds; K, posterior chaetigers, dorsal view. ac—anal cirrus, dc—dorsal cirrus, fl—frontal lip, la—lateral antenna, ll—lower lip, ma—median antenna, mnd—mandibles, p—palp, per—peristomium, pg—pygidium, pl—postchaetal lobe, prl—prechaetal lobe, ul—upper lip, vc—ventral cirrus." captionText-3="FIGURE 4. Leptoecia midatlantica sp. nov. A, parapodium of chaetiger 1, anterior view; B, same, posterior view; C, parapodium of chaetiger 2, anterior view; D, parapodium of chaetiger 3, anterior view; E, parapodium of chaetiger 6, posterior view; F, parapodium of chaetiger 12, anterior view; G, parapodium of chaetiger 33, anterior view; H, parapodium of chaetiger 34, anterior view; I, parapodium of chaetiger 35, anterior view; J, parapodium of chaetiger 66, anterior view." captionText-4="FIGURE 5. Leptoecia midatlantica sp. nov. A, parapodium of chaetiger 2, ventral view; B, parapodium of chaetiger 3, ventral view; C, parapodia of chaetigers 8 – 10, dorsal view; D – G, simple bidentate falcigers from chaetiger 1; H, simple unidentate falciger from chaetiger 1; I, limbate chaetae; J – L, pectinate chaetae; M, N, simple subacicular hooks from anterior region of the body; O, simple subacicular hooks from the posterior region of the body; P, compound subacicular hook from the last chaetiger. dc—dorsal cirrus, pl—postchaetal lobe, vgp—ventral glandular pad, vc—ventral cirrus." captionText-5="FIGURE 6. Leptoecia benthaliana. A – C, anterior falcigers (the black arrows indicate subdistal reduction in the diameter of the falciger, modified from Orensanz 1990); Leptoecia midatlantica sp. nov. D, E, simple unidentate falcigers from chaetiger 1; F, G, simple bidentate falcigers from chaetiger 1; H, simple bidentate hook from chaetiger 35." captionText-6="FIGURE 7. Leptoecia midatlantica sp. nov. A, relationship between body width of specimens (N = 67) and origin of subacicular hooks; B, average body width of specimens with bilobed and conical prostomia." httpUri-0="https://zenodo.org/record/279914/files/figure.png" httpUri-1="https://zenodo.org/record/279915/files/figure.png" httpUri-2="https://zenodo.org/record/279916/files/figure.png" httpUri-3="https://zenodo.org/record/279917/files/figure.png" httpUri-4="https://zenodo.org/record/279918/files/figure.png" httpUri-5="https://zenodo.org/record/279919/files/figure.png" httpUri-6="https://zenodo.org/record/279920/files/figure.png" pageId="6" pageNumber="51">Figs. 1–7</figureCitation>
,
<tableCitation id="C6FE2EABFFAFCF63FEFFFC0D1C30FC68" box="[269,338,964,988]" captionStart="TABLE 3" captionStartId="11.[151,235,1202,1223]" captionTargetBox="[151,1436,1282,1800]" captionTargetPageId="11" captionText="TABLE 3. Intraspecific variability of the main morphological characters of Leptoecia midatlantica sp. nov. SD corresponds to standard deviation; N represents the number of the examined specimens." httpUri="http://table.plazi.org/id/DF034B98FFA2CF6EFF65FB7B1E15FB52" pageId="6" pageNumber="51" tableUuid="DF034B98FFA2CF6EFF65FB7B1E15FB52">Tab. 3</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C366489BFFAFCF63FF65FBCE1CC7F98E" pageId="6" pageNumber="51" type="materials_examined">
<paragraph id="8BC31B10FFAFCF63FF65FBCE1DBAFB42" blockId="6.[151,1437,1030,2026]" pageId="6" pageNumber="51">
<emphasis id="B908C702FFAFCF63FF65FBCE1C25FBAB" bold="true" box="[151,327,1030,1055]" pageId="6" pageNumber="51">
<typeStatus id="54C7A5B2FFAFCF63FF65FBCE1DB0FBAB" box="[151,210,1031,1055]" pageId="6" pageNumber="51">Type</typeStatus>
material.
</emphasis>
<collectionCode id="ED6D83D5FFAFCF63FEA2FBCE1CD8FBAA" LSID="urn:lsid:biocol.org:col:34665" box="[336,442,1031,1054]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="6" pageNumber="51">NHMUK</collectionCode>
2011.351 St. JC037/70, 4% formalin transferred to 70% ethanol (
<typeStatus id="54C7A5B2FFAFCF63FB46FBCF187EFBAA" box="[1204,1308,1030,1054]" pageId="6" pageNumber="51" type="holotype">holotype</typeStatus>
);
<collectionCode id="ED6D83D5FFAFCF63FAC0FBCE18FEFBAA" LSID="urn:lsid:biocol.org:col:34665" box="[1330,1436,1031,1054]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="6" pageNumber="51">NHMUK</collectionCode>
2011.352–361 St. JC037/70, 4% formalin transferred to 70% ethanol (11
<typeStatus id="54C7A5B2FFAFCF63FC34FBE31954FBF6" box="[966,1078,1066,1090]" pageId="6" pageNumber="51" type="paratype">paratypes</typeStatus>
);
<collectionCode id="ED6D83D5FFAFCF63FBBBFBE219D1FBF6" LSID="urn:lsid:biocol.org:col:34665" box="[1097,1203,1067,1090]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="6" pageNumber="51">NHMUK</collectionCode>
2011.362 St. JC037/ 70, 96% ethanol (1
<typeStatus id="54C7A5B2FFAFCF63FE89FB871C82FBD2" box="[379,480,1102,1126]" pageId="6" pageNumber="51" type="paratype">paratype</typeStatus>
);
<collectionCode id="ED6D83D5FFAFCF63FE04FB861F02FBD2" LSID="urn:lsid:biocol.org:col:34665" box="[502,608,1103,1126]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="6" pageNumber="51">NHMUK</collectionCode>
2011.363–370 St. JC037/61, 4% formalin transferred to 70% ethanol (8
<typeStatus id="54C7A5B2FFAFCF63FF65FBBB1C6AFB3E" box="[151,264,1138,1162]" pageId="6" pageNumber="51" type="paratype">paratypes</typeStatus>
);
<collectionCode id="ED6D83D5FFAFCF63FEE8FBBA1CE6FB3E" LSID="urn:lsid:biocol.org:col:34665" box="[282,388,1139,1162]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="6" pageNumber="51">NHMUK</collectionCode>
2011.371, St. JC037/61, 96% ethanol (1
<typeStatus id="54C7A5B2FFAFCF63FCA1FBBB1EDBFB3E" box="[851,953,1138,1162]" pageId="6" pageNumber="51" type="paratype">paratype</typeStatus>
);
<collectionCode id="ED6D83D5FFAFCF63FC39FBBA197EFB3E" box="[971,1052,1139,1162]" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="6" pageNumber="51">ZMBN</collectionCode>
87901 St. JC037/61, 4% formalin transferred to 70% ethanol (2
<typeStatus id="54C7A5B2FFAFCF63FE15FB5F1F35FB1A" box="[487,599,1174,1198]" pageId="6" pageNumber="51" type="paratype">paratypes</typeStatus>
);
<collectionCode id="ED6D83D5FFAFCF63FD98FB5E1FD9FB1A" box="[618,699,1175,1198]" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="6" pageNumber="51">ZMBN</collectionCode>
87902 St. JC037/61, 96% ethanol (1
<typeStatus id="54C7A5B2FFAFCF63FB92FB5F19A7FB1A" box="[1120,1221,1174,1198]" pageId="6" pageNumber="51" type="paratype">paratype</typeStatus>
);
<collectionCode id="ED6D83D5FFAFCF63FB2BFB5E1848FB1A" box="[1241,1322,1175,1198]" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="6" pageNumber="51">ZMBN</collectionCode>
87903 St. JC037/70, 4% formalin transferred to 70% ethanol (5
<typeStatus id="54C7A5B2FFAFCF63FD1CFB731E3DFB66" box="[750,863,1210,1234]" pageId="6" pageNumber="51" type="paratype">paratypes</typeStatus>
);
<collectionCode id="ED6D83D5FFAFCF63FC83FB721EA0FB66" box="[881,962,1211,1234]" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="6" pageNumber="51">ZMBN</collectionCode>
87904 St. JC037/70, 96% ethanol (1
<typeStatus id="54C7A5B2FFAFCF63FA90FB721DAEFB42" pageId="6" pageNumber="51" type="paratype">paratype</typeStatus>
).
</paragraph>
<paragraph id="8BC31B10FFAFCF63FF37FACA1CB8FA46" blockId="6.[151,1437,1030,2026]" pageId="6" pageNumber="51">
<emphasis id="B908C702FFAFCF63FF37FACA1F45FAAF" bold="true" box="[197,551,1282,1307]" pageId="6" pageNumber="51">
Non
<typeStatus id="54C7A5B2FFAFCF63FEF3FACA1C56FAAF" box="[257,308,1283,1307]" pageId="6" pageNumber="51">type</typeStatus>
material examined.
</emphasis>
<collectionCode id="ED6D83D5FFAFCF63FDC3FACA1FE7FAAE" box="[561,645,1283,1306]" httpUri="http://grbio.org/cool/0tqf-efed" name="Museum of Zoology at the University of Bergen, Invertebrate Collection" pageId="6" pageNumber="51">ZMBN</collectionCode>
, RV
<emphasis id="B908C702FFAFCF63FD30FACA1E4BFAAE" box="[706,809,1283,1306]" italics="true" pageId="6" pageNumber="51">
G.
<collectingCountry id="F36B5B80FFAFCF63FD28FACA1F8CFAAE" box="[730,750,1283,1306]" name="Iceland" pageId="6" pageNumber="51">O</collectingCountry>
Sars
</emphasis>
, St. 368, 4% formalin transferred to 70% ethanol (1);
<collectionCode id="ED6D83D5FFAFCF63FF65FAEF1D86FA8A" box="[151,228,1318,1342]" pageId="6" pageNumber="51">NOCS</collectionCode>
,
<collectionCode id="ED6D83D5FFAFCF63FF02FAEF1C47FA8A" box="[240,293,1318,1342]" pageId="6" pageNumber="51">RRS</collectionCode>
<emphasis id="B908C702FFAFCF63FEDEFAE11CD6FA8A" box="[300,436,1318,1343]" italics="true" pageId="6" pageNumber="51">James Cook</emphasis>
: St. JC011/101, 4% formalin transferred to 70% ethanol (5); St. JC011/111, 4% formalin transferred to 70% ethanol (2); St. JC037/19, 96% ethanol (5); St. JC037/61, 4% formalin transferred to 70% ethanol (73); St. JC037/61, 96% ethanol (12); St. JC037/67, 4% formalin transferred to 70% ethanol (6); St. JC037/67, 96% ethanol (2); St. JC037/70, 4% formalin transferred to 70% ethanol (61); St. JC037/70, 96% ethanol (14); St. JC037/79, 4% formalin transferred to 70% ethanol (1);
<collectionCode id="ED6D83D5FFAFCF63FCF0FA7F1E50FA7A" LSID="urn:lsid:biocol.org:col:14844" box="[770,818,1462,1486]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14844" name="Scripps Institution of Oceanography" pageId="6" pageNumber="51">SIO</collectionCode>
,
<collectionCode id="ED6D83D5FFAFCF63FCCFFA7F1E10FA7A" box="[829,882,1462,1486]" pageId="6" pageNumber="51">RRS</collectionCode>
<emphasis id="B908C702FFAFCF63FC8BFA711963FA7A" box="[889,1025,1462,1487]" italics="true" pageId="6" pageNumber="51">James Cook</emphasis>
: St. JC037/61, 96% ethanol (10); St. JC037/70, 96% ethanol (20).
</paragraph>
<paragraph id="8BC31B10FFAFCF63FF37FA361CC7F98E" blockId="6.[151,1437,1030,2026]" pageId="6" pageNumber="51">
<emphasis id="B908C702FFAFCF63FF37FA361C00F9A3" bold="true" box="[197,354,1534,1559]" pageId="6" pageNumber="51">
<typeStatus id="54C7A5B2FFAFCF63FF37FA361C62F9A3" box="[197,256,1535,1559]" pageId="6" pageNumber="51">Type</typeStatus>
locality.
</emphasis>
North Atlantic, Mid-Atlantic Ridge, Charlie-Gibbs Fracture Zone,
<geoCoordinate id="EE487DD7FFAFCF63FBA3FA3719CFF9A2" box="[1105,1197,1534,1558]" direction="north" orientation="latitude" pageId="6" pageNumber="51" precision="925" value="54.216667">54º13'N</geoCoordinate>
,
<geoCoordinate id="EE487DD7FFAFCF63FB4AFA371878F9A2" box="[1208,1306,1534,1558]" direction="west" orientation="longitude" pageId="6" pageNumber="51" precision="925" value="-36.066666">36º04'W</geoCoordinate>
–
<geoCoordinate id="EE487DD7FFAFCF63FADBFA3718F5F9A2" box="[1321,1431,1534,1558]" direction="north" orientation="latitude" pageId="6" pageNumber="51" precision="92" value="54.175">54º10.5'N</geoCoordinate>
,
<geoCoordinate id="EE487DD7FFAFCF63FF65F9EB1D95F98E" box="[151,247,1570,1594]" direction="west" orientation="longitude" pageId="6" pageNumber="51" precision="925" value="-36.083332">36º05'W</geoCoordinate>
,
<quantity id="4C84B6F5FFAFCF63FEF0F9EB1CC2F98E" box="[258,416,1570,1594]" metricMagnitude="3" metricUnit="m" metricValue="2.6094999999999997" metricValueMax="2.6149999999999998" metricValueMin="2.604" pageId="6" pageNumber="51" unit="m" value="2609.5" valueMax="2615.0" valueMin="2604.0">2604–2615 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C366489BFFAFCF63FF37F98F19D7F97E" pageId="6" pageNumber="51" type="diagnosis">
<paragraph id="8BC31B10FFAFCF63FF37F98F19D7F97E" blockId="6.[151,1437,1030,2026]" pageId="6" pageNumber="51">
<emphasis id="B908C702FFAFCF63FF37F98F1C5DF9EB" bold="true" box="[197,319,1606,1631]" pageId="6" pageNumber="51">Diagnosis.</emphasis>
Both uni- and bidentate simple falcigers exclusively on the first pair of parapodia, rarely only bidentate falcigers present; falcigers straight, without subdistal reduction in diameter; subacicular hooks from chaetiger 28–39; dorsal cirri on all chaetigers; digitiform postchaetal lobes till chaetiger 8–16; peristomium half as long as first chaetiger; tubes always dorsoventrally flattened with longitudinal ribs along both sides
</paragraph>
</subSubSection>
<subSubSection id="C366489BFFAFCF69FF37F91F19AAFF60" lastPageId="12" lastPageNumber="57" pageId="6" pageNumber="51" type="description">
<paragraph id="8BC31B10FFAFCF63FF37F91F198DF882" blockId="6.[151,1437,1030,2026]" pageId="6" pageNumber="51">
<emphasis id="B908C702FFAFCF63FF37F91F1C34F95B" bold="true" box="[197,342,1750,1775]" pageId="6" pageNumber="51">Description.</emphasis>
<typeStatus id="54C7A5B2FFAFCF63FE93F91F1CAEF95A" box="[353,460,1750,1774]" pageId="6" pageNumber="51" type="holotype">Holotype</typeStatus>
, a complete specimen consisting of 66 chaetigers,
<quantity id="4C84B6F5FFAFCF63FBD2F91F1911F95A" box="[1056,1139,1750,1774]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="6" pageNumber="51" unit="mm" value="31.0">31 mm</quantity>
long and 0.93 mm wide. Other examined specimens varying from 0.69 mm to 1.31 mm in width and from
<quantity id="4C84B6F5FFAFCF63FBD9F9331919F8A6" box="[1067,1147,1786,1810]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="6" pageNumber="51" unit="mm" value="19.0">19 mm</quantity>
to
<quantity id="4C84B6F5FFAFCF63FB6CF933198FF8A6" box="[1182,1261,1786,1810]" metricMagnitude="-2" metricUnit="m" metricValue="4.1" pageId="6" pageNumber="51" unit="mm" value="41.0">41 mm</quantity>
in length. Longest examined specimen consists of 81 chaetigers. All specimens yellowish, lacking colour pattern.
</paragraph>
<paragraph id="8BC31B10FFAFCF63FF37F88B1909F85E" blockId="6.[151,1437,1030,2026]" pageId="6" pageNumber="51">
Prostomium of variable shape, from distally pointed through conical or slightly bilobed, to distinctly bilobed with two hemispherical frontal lips (
<figureCitation id="13470795FFAFCF63FDCDF8AF1FE6F8CA" box="[575,644,1894,1918]" captionStart="FIGURE 2" captionStartId="7.[151,250,1744,1767]" captionTargetBox="[151,1436,193,1722]" captionTargetId="figure@7.[151,1436,193,1723]" captionTargetPageId="7" captionText="FIGURE 2. Leptoecia midatlantica sp. nov. A, anterior region of the holotype, ventral view; B, same, dorsal view; C – E shape of the prostomium, ventral view; F, maxillae; G, mandibles; H, pygidium of the holotype, ventral view; I, same, dorsal view; J – L, pygidia, dorsal view." httpUri="https://zenodo.org/record/279915/files/figure.png" pageId="6" pageNumber="51">Fig. 2</figureCitation>
A–E, 3A–F). Eyes absent. Nuchal groves absent (
<figureCitation id="13470795FFAFCF63FB3AF8AF186EF8CA" box="[1224,1292,1894,1918]" captionStart="FIGURE 3" captionStartId="8.[151,250,1892,1915]" captionTargetBox="[151,1436,193,1871]" captionTargetId="figure@8.[151,1436,193,1871]" captionTargetPageId="8" captionText="FIGURE 3. Leptoecia midatlantica sp. nov. A, C, anterior region of the body, ventral view; B, same, dorsal view; D, F prostomium and peristomium, ventral view; E, same, dorsal view; G, sensory organs on prostomium, dorsal view; H, posterior region of the body, ventral view; I, pygidium, dorsal view; J pygidial glandular buds; K, posterior chaetigers, dorsal view. ac—anal cirrus, dc—dorsal cirrus, fl—frontal lip, la—lateral antenna, ll—lower lip, ma—median antenna, mnd—mandibles, p—palp, per—peristomium, pg—pygidium, pl—postchaetal lobe, prl—prechaetal lobe, ul—upper lip, vc—ventral cirrus." httpUri="https://zenodo.org/record/279916/files/figure.png" pageId="6" pageNumber="51">Fig. 3</figureCitation>
E, G). Palps reaching chaetiger 1; lateral antennae reaching chaetiger 8 (5–13); median antenna longer and thicker than lateral antennae, reaching chaetiger 9 (7–16). Ceratophores consisting of 2 (1–3) rings equal in length (
<figureCitation id="13470795FFAFCF63FB21F8671875F872" box="[1235,1303,1966,1990]" captionStart="FIGURE 2" captionStartId="7.[151,250,1744,1767]" captionTargetBox="[151,1436,193,1722]" captionTargetId="figure@7.[151,1436,193,1723]" captionTargetPageId="7" captionText="FIGURE 2. Leptoecia midatlantica sp. nov. A, anterior region of the holotype, ventral view; B, same, dorsal view; C – E shape of the prostomium, ventral view; F, maxillae; G, mandibles; H, pygidium of the holotype, ventral view; I, same, dorsal view; J – L, pygidia, dorsal view." httpUri="https://zenodo.org/record/279915/files/figure.png" pageId="6" pageNumber="51">Fig. 2</figureCitation>
B, 3E). Peristomium about half as long as the first chaetiger; peristomial cirri absent (
<figureCitation id="13470795FFAFCF63FC2BF81B197EF85E" box="[985,1052,2002,2026]" captionStart="FIGURE 2" captionStartId="7.[151,250,1744,1767]" captionTargetBox="[151,1436,193,1722]" captionTargetId="figure@7.[151,1436,193,1723]" captionTargetPageId="7" captionText="FIGURE 2. Leptoecia midatlantica sp. nov. A, anterior region of the holotype, ventral view; B, same, dorsal view; C – E shape of the prostomium, ventral view; F, maxillae; G, mandibles; H, pygidium of the holotype, ventral view; I, same, dorsal view; J – L, pygidia, dorsal view." httpUri="https://zenodo.org/record/279915/files/figure.png" pageId="6" pageNumber="51">Fig. 2</figureCitation>
B, 3E).
</paragraph>
<caption id="DF034B98FFAECF62FF65F9191CE4F897" httpUri="https://zenodo.org/record/279915/files/figure.png" pageId="7" pageNumber="52" targetBox="[151,1436,193,1722]" targetPageId="7">
<paragraph id="8BC31B10FFAECF62FF65F9191CE4F897" blockId="7.[151,1437,1744,1827]" pageId="7" pageNumber="52">
<emphasis id="B908C702FFAECF62FF65F9191C71F951" bold="true" box="[151,275,1744,1767]" pageId="7" pageNumber="52">FIGURE 2.</emphasis>
<taxonomicName id="4C7C6093FFAECF62FEEBF9181F61F952" box="[281,515,1744,1766]" class="Polychaeta" family="Onuphidae" genus="Leptoecia" kingdom="Animalia" order="Eunicida" pageId="7" pageNumber="52" phylum="Annelida" rank="species" species="midatlantica" status="sp. nov.">
<emphasis id="B908C702FFAECF62FEEBF9181F61F952" box="[281,515,1744,1766]" italics="true" pageId="7" pageNumber="52">Leptoecia midatlantica</emphasis>
</taxonomicName>
<emphasis id="B908C702FFAECF62FDF8F9181F38F952" bold="true" box="[522,602,1745,1766]" pageId="7" pageNumber="52">
<taxonomicNameLabel id="A23B7A79FFAECF62FDF8F9181F38F952" box="[522,602,1745,1766]" pageId="7" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, anterior region of the holotype, ventral view; B, same, dorsal view; C–E shape of the prostomium, ventral view; F, maxillae; G, mandibles; H, pygidium of the holotype, ventral view; I, same, dorsal view; J– L, pygidia, dorsal view.
</paragraph>
</caption>
<caption id="DF034B98FFA1CF6DFF65F8AD1868F841" httpUri="https://zenodo.org/record/279916/files/figure.png" pageId="8" pageNumber="53" targetBox="[151,1436,193,1871]" targetPageId="8">
<paragraph id="8BC31B10FFA1CF6DFF65F8AD1868F841" blockId="8.[151,1436,1892,2037]" pageId="8" pageNumber="53">
<emphasis id="B908C702FFA1CF6DFF65F8AD1C70F8CE" bold="true" box="[151,274,1892,1915]" pageId="8" pageNumber="53">FIGURE 3.</emphasis>
<taxonomicName id="4C7C6093FFA1CF6DFEEBF8AF1F61F8CF" box="[281,515,1893,1915]" class="Polychaeta" family="Onuphidae" genus="Leptoecia" kingdom="Animalia" order="Eunicida" pageId="8" pageNumber="53" phylum="Annelida" rank="species" species="midatlantica" status="sp. nov.">
<emphasis id="B908C702FFA1CF6DFEEBF8AF1F61F8CF" box="[281,515,1893,1915]" italics="true" pageId="8" pageNumber="53">Leptoecia midatlantica</emphasis>
</taxonomicName>
<emphasis id="B908C702FFA1CF6DFDFBF8AC1F3BF8CE" bold="true" box="[521,601,1893,1914]" pageId="8" pageNumber="53">
<taxonomicNameLabel id="A23B7A79FFA1CF6DFDFBF8AC1F3BF8CE" box="[521,601,1893,1914]" pageId="8" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, C, anterior region of the body, ventral view; B, same, dorsal view; D, F prostomium and peristomium, ventral view; E, same, dorsal view; G, sensory organs on prostomium, dorsal view; H, posterior region of the body, ventral view; I, pygidium, dorsal view; J pygidial glandular buds; K, posterior chaetigers, dorsal view. ac—anal cirrus, dc—dorsal cirrus, fl—frontal lip, la—lateral antenna, ll—lower lip, ma—median antenna, mnd—mandibles, p—palp, per—peristomium, pg—pygidium, pl—postchaetal lobe, prl—prechaetal lobe, ul—upper lip, vc—ventral cirrus.
</paragraph>
</caption>
<caption id="DF034B98FFA0CF6CFF65FA3D19DFF9D2" httpUri="https://zenodo.org/record/279917/files/figure.png" pageId="9" pageNumber="54" targetBox="[151,1436,193,1502]" targetPageId="9">
<paragraph id="8BC31B10FFA0CF6CFF65FA3D19DFF9D2" blockId="9.[151,1436,1524,1638]" pageId="9" pageNumber="54">
<emphasis id="B908C702FFA0CF6CFF65FA3D1C71F9BE" bold="true" box="[151,275,1524,1547]" pageId="9" pageNumber="54">FIGURE 4.</emphasis>
<taxonomicName id="4C7C6093FFA0CF6CFEE8FA3F1F66F9BF" box="[282,516,1525,1547]" class="Polychaeta" family="Onuphidae" genus="Leptoecia" kingdom="Animalia" order="Eunicida" pageId="9" pageNumber="54" phylum="Annelida" rank="species" species="midatlantica" status="sp. nov.">
<emphasis id="B908C702FFA0CF6CFEE8FA3F1F66F9BF" box="[282,516,1525,1547]" italics="true" pageId="9" pageNumber="54">Leptoecia midatlantica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B7A79FFA0CF6CFDFEFA3C1F3FF9BE" box="[524,605,1525,1546]" pageId="9" pageNumber="54" rank="species">
<emphasis id="B908C702FFA0CF6CFDFEFA3C1F3FF9BE" bold="true" box="[524,605,1525,1546]" pageId="9" pageNumber="54">sp. nov.</emphasis>
</taxonomicNameLabel>
A, parapodium of chaetiger 1, anterior view; B, same, posterior view; C, parapodium of chaetiger 2, anterior view; D, parapodium of chaetiger 3, anterior view; E, parapodium of chaetiger 6, posterior view; F, parapodium of chaetiger 12, anterior view; G, parapodium of chaetiger 33, anterior view; H, parapodium of chaetiger 34, anterior view; I, parapodium of chaetiger 35, anterior view; J, parapodium of chaetiger 66, anterior view.
</paragraph>
</caption>
<paragraph id="8BC31B10FFA0CF6CFF37F97E1946F813" blockId="9.[151,1436,1719,1959]" pageId="9" pageNumber="54">
First anterior pair of parapodia modified, prolonged, projecting lateroventrally and directing anteriorly, with auricular prechaetal lobes, knob-like postchaetal lobes and subulate dorsal and ventral cirri (
<figureCitation id="13470795FFA0CF6CFB5FF9121992F947" box="[1197,1264,1755,1779]" captionStart="FIGURE 3" captionStartId="8.[151,250,1892,1915]" captionTargetBox="[151,1436,193,1871]" captionTargetId="figure@8.[151,1436,193,1871]" captionTargetPageId="8" captionText="FIGURE 3. Leptoecia midatlantica sp. nov. A, C, anterior region of the body, ventral view; B, same, dorsal view; D, F prostomium and peristomium, ventral view; E, same, dorsal view; G, sensory organs on prostomium, dorsal view; H, posterior region of the body, ventral view; I, pygidium, dorsal view; J pygidial glandular buds; K, posterior chaetigers, dorsal view. ac—anal cirrus, dc—dorsal cirrus, fl—frontal lip, la—lateral antenna, ll—lower lip, ma—median antenna, mnd—mandibles, p—palp, per—peristomium, pg—pygidium, pl—postchaetal lobe, prl—prechaetal lobe, ul—upper lip, vc—ventral cirrus." httpUri="https://zenodo.org/record/279916/files/figure.png" pageId="9" pageNumber="54">Fig. 3</figureCitation>
F, 4A, B). Second pair of parapodia with short and rounded prechaetal lobes, subulate postchaetal lobes and dorsal and ventral cirri (
<figureCitation id="13470795FFA0CF6CFF27F8EA1C7AF88F" box="[213,280,1827,1851]" captionStart="FIGURE 4" captionStartId="9.[151,250,1524,1547]" captionTargetBox="[151,1436,193,1502]" captionTargetId="figure@9.[151,1436,193,1503]" captionTargetPageId="9" captionText="FIGURE 4. Leptoecia midatlantica sp. nov. A, parapodium of chaetiger 1, anterior view; B, same, posterior view; C, parapodium of chaetiger 2, anterior view; D, parapodium of chaetiger 3, anterior view; E, parapodium of chaetiger 6, posterior view; F, parapodium of chaetiger 12, anterior view; G, parapodium of chaetiger 33, anterior view; H, parapodium of chaetiger 34, anterior view; I, parapodium of chaetiger 35, anterior view; J, parapodium of chaetiger 66, anterior view." httpUri="https://zenodo.org/record/279917/files/figure.png" pageId="9" pageNumber="54">Fig. 4</figureCitation>
C, 5A). Subulate ventral cirri replaced by flattened round glandular pads from chaetiger 3 (
<figureCitation id="13470795FFA0CF6CFAFCF8EA1832F88F" box="[1294,1360,1827,1851]" captionStart="FIGURE 2" captionStartId="7.[151,250,1744,1767]" captionTargetBox="[151,1436,193,1722]" captionTargetId="figure@7.[151,1436,193,1723]" captionTargetPageId="7" captionText="FIGURE 2. Leptoecia midatlantica sp. nov. A, anterior region of the holotype, ventral view; B, same, dorsal view; C – E shape of the prostomium, ventral view; F, maxillae; G, mandibles; H, pygidium of the holotype, ventral view; I, same, dorsal view; J – L, pygidia, dorsal view." httpUri="https://zenodo.org/record/279915/files/figure.png" pageId="9" pageNumber="54">Fig. 2</figureCitation>
A, 4D, 5B). Dorsal cirri present on all chaetigers including the posteriormost ones, subulate on anterior parapodia and decreasing in length posteriorly (
<figureCitation id="13470795FFA0CF6CFDFAF8A21F28F837" box="[520,586,1899,1923]" captionStart="FIGURE 2" captionStartId="7.[151,250,1744,1767]" captionTargetBox="[151,1436,193,1722]" captionTargetId="figure@7.[151,1436,193,1723]" captionTargetPageId="7" captionText="FIGURE 2. Leptoecia midatlantica sp. nov. A, anterior region of the holotype, ventral view; B, same, dorsal view; C – E shape of the prostomium, ventral view; F, maxillae; G, mandibles; H, pygidium of the holotype, ventral view; I, same, dorsal view; J – L, pygidia, dorsal view." httpUri="https://zenodo.org/record/279915/files/figure.png" pageId="9" pageNumber="54">
Fig.
<date id="FFC23DD0FFA0CF6CFDCEF8A21F28F837" box="[572,586,1899,1923]" pageId="9" pageNumber="54">2</date>
</figureCitation>
I–L,
<date id="FFC23DD0FFA0CF6CFD72F8A21FF8F837" box="[640,666,1899,1923]" pageId="9" pageNumber="54">3I</date>
, K). Postchaetal lobes subulate to digitiform on first 13 (8–16) chaetigers (
<figureCitation id="13470795FFA0CF6CFF16F8461C45F813" box="[228,295,1935,1959]" captionStart="FIGURE 5" captionStartId="10.[151,250,1624,1647]" captionTargetBox="[151,1436,193,1596]" captionTargetId="figure@10.[151,1436,193,1598]" captionTargetPageId="10" captionText="FIGURE 5. Leptoecia midatlantica sp. nov. A, parapodium of chaetiger 2, ventral view; B, parapodium of chaetiger 3, ventral view; C, parapodia of chaetigers 8 – 10, dorsal view; D – G, simple bidentate falcigers from chaetiger 1; H, simple unidentate falciger from chaetiger 1; I, limbate chaetae; J – L, pectinate chaetae; M, N, simple subacicular hooks from anterior region of the body; O, simple subacicular hooks from the posterior region of the body; P, compound subacicular hook from the last chaetiger. dc—dorsal cirrus, pl—postchaetal lobe, vgp—ventral glandular pad, vc—ventral cirrus." httpUri="https://zenodo.org/record/279918/files/figure.png" pageId="9" pageNumber="54">Fig. 5</figureCitation>
C), completely disappearing posteriorly. Branchiae absent (
<figureCitation id="13470795FFA0CF6CFC4DF8461963F813" box="[959,1025,1935,1959]" captionStart="FIGURE 3" captionStartId="8.[151,250,1892,1915]" captionTargetBox="[151,1436,193,1871]" captionTargetId="figure@8.[151,1436,193,1871]" captionTargetPageId="8" captionText="FIGURE 3. Leptoecia midatlantica sp. nov. A, C, anterior region of the body, ventral view; B, same, dorsal view; D, F prostomium and peristomium, ventral view; E, same, dorsal view; G, sensory organs on prostomium, dorsal view; H, posterior region of the body, ventral view; I, pygidium, dorsal view; J pygidial glandular buds; K, posterior chaetigers, dorsal view. ac—anal cirrus, dc—dorsal cirrus, fl—frontal lip, la—lateral antenna, ll—lower lip, ma—median antenna, mnd—mandibles, p—palp, per—peristomium, pg—pygidium, pl—postchaetal lobe, prl—prechaetal lobe, ul—upper lip, vc—ventral cirrus." httpUri="https://zenodo.org/record/279916/files/figure.png" pageId="9" pageNumber="54">Fig. 3</figureCitation>
B).
</paragraph>
<caption id="DF034B98FFA3CF6FFF65F9911970F940" httpUri="https://zenodo.org/record/279918/files/figure.png" pageId="10" pageNumber="55" targetBox="[151,1436,193,1596]" targetPageId="10">
<paragraph id="8BC31B10FFA3CF6FFF65F9911970F940" blockId="10.[151,1437,1624,1780]" pageId="10" pageNumber="55">
<emphasis id="B908C702FFA3CF6FFF65F9911C70F9DA" bold="true" box="[151,274,1624,1647]" pageId="10" pageNumber="55">FIGURE 5.</emphasis>
<taxonomicName id="4C7C6093FFA3CF6FFEEBF9931F61F9DB" box="[281,515,1625,1647]" class="Polychaeta" family="Onuphidae" genus="Leptoecia" kingdom="Animalia" order="Eunicida" pageId="10" pageNumber="55" phylum="Annelida" rank="species" species="midatlantica" status="sp. nov.">
<emphasis id="B908C702FFA3CF6FFEEBF9931F61F9DB" box="[281,515,1625,1647]" italics="true" pageId="10" pageNumber="55">Leptoecia midatlantica</emphasis>
</taxonomicName>
<emphasis id="B908C702FFA3CF6FFDFBF9901F3BF9DA" bold="true" box="[521,601,1625,1646]" pageId="10" pageNumber="55">
<taxonomicNameLabel id="A23B7A79FFA3CF6FFDFBF9901F3BF9DA" box="[521,601,1625,1646]" pageId="10" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, parapodium of chaetiger 2, ventral view; B, parapodium of chaetiger 3, ventral view; C, parapodia of chaetigers 8–10, dorsal view; D–G, simple bidentate falcigers from chaetiger 1; H, simple unidentate falciger from chaetiger 1; I, limbate chaetae; J–L, pectinate chaetae; M, N, simple subacicular hooks from anterior region of the body; O, simple subacicular hooks from the posterior region of the body; P, compound subacicular hook from the last chaetiger. dc—dorsal cirrus, pl—postchaetal lobe, vgp—ventral glandular pad, vc—ventral cirrus.
</paragraph>
</caption>
<paragraph id="8BC31B10FFA3CF6EFF37F8D71CCBFEA8" blockId="10.[151,1436,1822,2026]" lastBlockId="11.[151,1437,151,284]" lastPageId="11" lastPageNumber="56" pageId="10" pageNumber="55">
First pair of parapodia with simple uni- and bidentate falcigers, rarely only bidentate falcigers present; capillary, limbate or pectinate chaetae absent. Anterior falcigers with short, distally blunt paired hoods and without subdistal reduction in diameter (
<figureCitation id="13470795FFA3CF6FFE12F8AF1F41F8CA" box="[480,547,1894,1918]" captionStart="FIGURE 5" captionStartId="10.[151,250,1624,1647]" captionTargetBox="[151,1436,193,1596]" captionTargetId="figure@10.[151,1436,193,1598]" captionTargetPageId="10" captionText="FIGURE 5. Leptoecia midatlantica sp. nov. A, parapodium of chaetiger 2, ventral view; B, parapodium of chaetiger 3, ventral view; C, parapodia of chaetigers 8 – 10, dorsal view; D – G, simple bidentate falcigers from chaetiger 1; H, simple unidentate falciger from chaetiger 1; I, limbate chaetae; J – L, pectinate chaetae; M, N, simple subacicular hooks from anterior region of the body; O, simple subacicular hooks from the posterior region of the body; P, compound subacicular hook from the last chaetiger. dc—dorsal cirrus, pl—postchaetal lobe, vgp—ventral glandular pad, vc—ventral cirrus." httpUri="https://zenodo.org/record/279918/files/figure.png" pageId="10" pageNumber="55">Fig. 5</figureCitation>
D–H, 6D–G). Limbate (
<figureCitation id="13470795FFA3CF6FFCC5F8AF1E19F8CA" box="[823,891,1894,1918]" captionStart="FIGURE 5" captionStartId="10.[151,250,1624,1647]" captionTargetBox="[151,1436,193,1596]" captionTargetId="figure@10.[151,1436,193,1598]" captionTargetPageId="10" captionText="FIGURE 5. Leptoecia midatlantica sp. nov. A, parapodium of chaetiger 2, ventral view; B, parapodium of chaetiger 3, ventral view; C, parapodia of chaetigers 8 – 10, dorsal view; D – G, simple bidentate falcigers from chaetiger 1; H, simple unidentate falciger from chaetiger 1; I, limbate chaetae; J – L, pectinate chaetae; M, N, simple subacicular hooks from anterior region of the body; O, simple subacicular hooks from the posterior region of the body; P, compound subacicular hook from the last chaetiger. dc—dorsal cirrus, pl—postchaetal lobe, vgp—ventral glandular pad, vc—ventral cirrus." httpUri="https://zenodo.org/record/279918/files/figure.png" pageId="10" pageNumber="55">
Fig.
<date id="FFC23DD0FFA3CF6FFC9FF8AF1E19F8CA" box="[877,891,1894,1918]" pageId="10" pageNumber="55">5</date>
</figureCitation>
I) and pectinate (
<figureCitation id="13470795FFA3CF6FFBCCF8AF19E3F8CA" box="[1086,1153,1894,1918]" captionStart="FIGURE 5" captionStartId="10.[151,250,1624,1647]" captionTargetBox="[151,1436,193,1596]" captionTargetId="figure@10.[151,1436,193,1598]" captionTargetPageId="10" captionText="FIGURE 5. Leptoecia midatlantica sp. nov. A, parapodium of chaetiger 2, ventral view; B, parapodium of chaetiger 3, ventral view; C, parapodia of chaetigers 8 – 10, dorsal view; D – G, simple bidentate falcigers from chaetiger 1; H, simple unidentate falciger from chaetiger 1; I, limbate chaetae; J – L, pectinate chaetae; M, N, simple subacicular hooks from anterior region of the body; O, simple subacicular hooks from the posterior region of the body; P, compound subacicular hook from the last chaetiger. dc—dorsal cirrus, pl—postchaetal lobe, vgp—ventral glandular pad, vc—ventral cirrus." httpUri="https://zenodo.org/record/279918/files/figure.png" pageId="10" pageNumber="55">Fig. 5</figureCitation>
J–L) chaetae arranged in two fascicles starting from chaetiger 2: dorsal fascicle with 5–8 pectinate chaetae and 1–2 limbate chaetae, ventral fascicle with 3–4 limbate chaetae (
<figureCitation id="13470795FFA3CF6FFDECF8671F03F872" box="[542,609,1966,1990]" captionStart="FIGURE 4" captionStartId="9.[151,250,1524,1547]" captionTargetBox="[151,1436,193,1502]" captionTargetId="figure@9.[151,1436,193,1503]" captionTargetPageId="9" captionText="FIGURE 4. Leptoecia midatlantica sp. nov. A, parapodium of chaetiger 1, anterior view; B, same, posterior view; C, parapodium of chaetiger 2, anterior view; D, parapodium of chaetiger 3, anterior view; E, parapodium of chaetiger 6, posterior view; F, parapodium of chaetiger 12, anterior view; G, parapodium of chaetiger 33, anterior view; H, parapodium of chaetiger 34, anterior view; I, parapodium of chaetiger 35, anterior view; J, parapodium of chaetiger 66, anterior view." httpUri="https://zenodo.org/record/279917/files/figure.png" pageId="10" pageNumber="55">Fig. 4</figureCitation>
C–G). Simple bidentate subacicular hooks (
<figureCitation id="13470795FFA3CF6FFBB9F86719EFF872" box="[1099,1165,1966,1990]" captionStart="FIGURE 5" captionStartId="10.[151,250,1624,1647]" captionTargetBox="[151,1436,193,1596]" captionTargetId="figure@10.[151,1436,193,1598]" captionTargetPageId="10" captionText="FIGURE 5. Leptoecia midatlantica sp. nov. A, parapodium of chaetiger 2, ventral view; B, parapodium of chaetiger 3, ventral view; C, parapodia of chaetigers 8 – 10, dorsal view; D – G, simple bidentate falcigers from chaetiger 1; H, simple unidentate falciger from chaetiger 1; I, limbate chaetae; J – L, pectinate chaetae; M, N, simple subacicular hooks from anterior region of the body; O, simple subacicular hooks from the posterior region of the body; P, compound subacicular hook from the last chaetiger. dc—dorsal cirrus, pl—postchaetal lobe, vgp—ventral glandular pad, vc—ventral cirrus." httpUri="https://zenodo.org/record/279918/files/figure.png" pageId="10" pageNumber="55">Fig. 5</figureCitation>
M–
<collectingCountry id="F36B5B80FFA3CF6FFB45F86719AEF872" box="[1207,1228,1966,1990]" name="Iceland" pageId="10" pageNumber="55">O</collectingCountry>
, 6H) starting from chaetiger 38 (28–39) within the ventral fascicle of limbate chaetae (
<figureCitation id="13470795FFA3CF6FFC5AF81B1E8CF85E" box="[936,1006,2002,2026]" captionStart="FIGURE 4" captionStartId="9.[151,250,1524,1547]" captionTargetBox="[151,1436,193,1502]" captionTargetId="figure@9.[151,1436,193,1503]" captionTargetPageId="9" captionText="FIGURE 4. Leptoecia midatlantica sp. nov. A, parapodium of chaetiger 1, anterior view; B, same, posterior view; C, parapodium of chaetiger 2, anterior view; D, parapodium of chaetiger 3, anterior view; E, parapodium of chaetiger 6, posterior view; F, parapodium of chaetiger 12, anterior view; G, parapodium of chaetiger 33, anterior view; H, parapodium of chaetiger 34, anterior view; I, parapodium of chaetiger 35, anterior view; J, parapodium of chaetiger 66, anterior view." httpUri="https://zenodo.org/record/279917/files/figure.png" pageId="10" pageNumber="55">Fig. 4</figureCitation>
H–J). Appearance of first subacicular hooks not size-dependent in the specimens examined (
<figureCitation id="13470795FFA2CF6EFCF7FF5E1E2BFF1B" box="[773,841,151,175]" captionStart="FIGURE 7" captionStartId="13.[151,250,680,703]" captionTargetBox="[151,1419,191,649]" captionTargetId="figure@13.[151,1436,191,649]" captionTargetPageId="13" captionText="FIGURE 7. Leptoecia midatlantica sp. nov. A, relationship between body width of specimens (N = 67) and origin of subacicular hooks; B, average body width of specimens with bilobed and conical prostomia." httpUri="https://zenodo.org/record/279920/files/figure.png" pageId="11" pageNumber="56">Fig. 7</figureCitation>
A). Compound subacicular hooks representing juvenile condition in few posteriormost chaetigers (
<figureCitation id="13470795FFA2CF6EFD58FF751F8FFF60" box="[682,749,188,212]" captionStart="FIGURE 5" captionStartId="10.[151,250,1624,1647]" captionTargetBox="[151,1436,193,1596]" captionTargetId="figure@10.[151,1436,193,1598]" captionTargetPageId="10" captionText="FIGURE 5. Leptoecia midatlantica sp. nov. A, parapodium of chaetiger 2, ventral view; B, parapodium of chaetiger 3, ventral view; C, parapodia of chaetigers 8 – 10, dorsal view; D – G, simple bidentate falcigers from chaetiger 1; H, simple unidentate falciger from chaetiger 1; I, limbate chaetae; J – L, pectinate chaetae; M, N, simple subacicular hooks from anterior region of the body; O, simple subacicular hooks from the posterior region of the body; P, compound subacicular hook from the last chaetiger. dc—dorsal cirrus, pl—postchaetal lobe, vgp—ventral glandular pad, vc—ventral cirrus." httpUri="https://zenodo.org/record/279918/files/figure.png" pageId="11" pageNumber="56">Fig. 5</figureCitation>
P). Ventral limbate chaetae present in all posterior parapodia. Pectinate chaetae flat with straight distal margins and up to ten denticles. Aciculae yellow with pointed tips, 1–3 per parapodium (
<figureCitation id="13470795FFA2CF6EFEA5FECD1CFEFEA8" box="[343,412,260,284]" captionStart="FIGURE 4" captionStartId="9.[151,250,1524,1547]" captionTargetBox="[151,1436,193,1502]" captionTargetId="figure@9.[151,1436,193,1503]" captionTargetPageId="9" captionText="FIGURE 4. Leptoecia midatlantica sp. nov. A, parapodium of chaetiger 1, anterior view; B, same, posterior view; C, parapodium of chaetiger 2, anterior view; D, parapodium of chaetiger 3, anterior view; E, parapodium of chaetiger 6, posterior view; F, parapodium of chaetiger 12, anterior view; G, parapodium of chaetiger 33, anterior view; H, parapodium of chaetiger 34, anterior view; I, parapodium of chaetiger 35, anterior view; J, parapodium of chaetiger 66, anterior view." httpUri="https://zenodo.org/record/279917/files/figure.png" pageId="11" pageNumber="56">Fig. 4</figureCitation>
).
</paragraph>
<caption id="DF034B98FFA2CF6EFF65FBF81950FB31" httpUri="https://zenodo.org/record/279919/files/figure.png" pageId="11" pageNumber="56" targetBox="[219,1369,327,1053]" targetPageId="11">
<paragraph id="8BC31B10FFA2CF6EFF65FBF81950FB31" blockId="11.[151,1436,1073,1157]" pageId="11" pageNumber="56">
<emphasis id="B908C702FFA2CF6EFF65FBF81C70FBF3" bold="true" box="[151,274,1073,1096]" pageId="11" pageNumber="56">FIGURE 6.</emphasis>
<taxonomicName id="4C7C6093FFA2CF6EFEEAFBFA1C99FBFC" box="[280,507,1074,1096]" class="Polychaeta" family="Onuphidae" genus="Leptoecia" kingdom="Animalia" order="Eunicida" pageId="11" pageNumber="56" phylum="Annelida" rank="species" species="benthaliana">
<emphasis id="B908C702FFA2CF6EFEEAFBFA1C99FBFC" box="[280,507,1074,1096]" italics="true" pageId="11" pageNumber="56">Leptoecia benthaliana</emphasis>
</taxonomicName>
. A–C, anterior falcigers (the black arrows indicate subdistal reduction in the diameter of the falciger, modified from Orensanz 1990);
<taxonomicName id="4C7C6093FFA2CF6EFDCAFB9B1E41FBD2" box="[568,803,1104,1127]" class="Polychaeta" family="Onuphidae" genus="Leptoecia" kingdom="Animalia" order="Eunicida" pageId="11" pageNumber="56" phylum="Annelida" rank="species" species="midatlantica" status="sp. nov.">
<emphasis id="B908C702FFA2CF6EFDCAFB9B1E41FBD2" box="[568,803,1104,1127]" italics="true" pageId="11" pageNumber="56">Leptoecia midatlantica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23B7A79FFA2CF6EFCDBFB981E18FBD2" box="[809,890,1105,1126]" pageId="11" pageNumber="56" rank="species">
<emphasis id="B908C702FFA2CF6EFCDBFB981E18FBD2" bold="true" box="[809,890,1105,1126]" pageId="11" pageNumber="56">sp. nov.</emphasis>
</taxonomicNameLabel>
D, E, simple unidentate falcigers from chaetiger 1; F, G, simple bidentate falcigers from chaetiger 1; H, simple bidentate hook from chaetiger 35.
</paragraph>
</caption>
<caption id="DF034B98FFA2CF6EFF65FB7B1E15FB52" ID-Table-UUID="DF034B98FFA2CF6EFF65FB7B1E15FB52" httpUri="http://table.plazi.org/id/DF034B98FFA2CF6EFF65FB7B1E15FB52" pageId="11" pageNumber="56" targetBox="[151,1436,1282,1800]" targetIsTable="true" targetPageId="11">
<paragraph id="8BC31B10FFA2CF6EFF65FB7B1E15FB52" blockId="11.[151,1436,1202,1254]" pageId="11" pageNumber="56">
<emphasis id="B908C702FFA2CF6EFF65FB7B1C66FB73" bold="true" box="[151,260,1202,1223]" pageId="11" pageNumber="56">TABLE 3.</emphasis>
Intraspecific variability of the main morphological characters of
<taxonomicName id="4C7C6093FFA2CF6EFC6BFB7A19E6FB7C" box="[921,1156,1202,1224]" class="Polychaeta" family="Onuphidae" genus="Leptoecia" kingdom="Animalia" order="Eunicida" pageId="11" pageNumber="56" phylum="Annelida" rank="species" species="midatlantica" status="sp. nov.">
<emphasis id="B908C702FFA2CF6EFC6BFB7A19E6FB7C" box="[921,1156,1202,1224]" italics="true" pageId="11" pageNumber="56">Leptoecia midatlantica</emphasis>
</taxonomicName>
<emphasis id="B908C702FFA2CF6EFB78FB7B19B8FB73" bold="true" box="[1162,1242,1202,1223]" pageId="11" pageNumber="56">
<taxonomicNameLabel id="A23B7A79FFA2CF6EFB78FB7B19B8FB73" box="[1162,1242,1202,1223]" pageId="11" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
SD corresponds to standard deviation; N represents the number of the examined specimens.
</paragraph>
</caption>
<paragraph id="8BC31B10FFA2CF6EFF6DFACB1845F8B3" pageId="11" pageNumber="56">
<table id="F97CE9B0FFA2309AFF65FACB18FEF8BC" box="[151,1436,1282,1800]" gridcols="5" gridrows="5" pageId="11" pageNumber="56">
<tr id="354C1952FFA2309AFF65FACB18FEFA27" box="[151,1436,1282,1427]" gridrow="0" pageId="11" pageNumber="56">
<th id="769D702EFFA2309AFF65FACB1FADFA27" box="[151,719,1282,1427]" gridcol="0" gridrow="0" pageId="11" pageNumber="56">Character Width (at chaetiger 10), mm Length, mm Number of chaetigers</th>
<th id="769D702EFFA2309AFCEBFACB1E1CFA27" box="[793,894,1282,1427]" gridcol="1" gridrow="0" pageId="11" pageNumber="56">Range 0.69–1.31 19–43 52–81</th>
<th id="769D702EFFA2309AFC13FACB195EFA27" box="[993,1084,1282,1427]" gridcol="2" gridrow="0" pageId="11" pageNumber="56">Mean 1.07 26.80 69.31</th>
<th id="769D702EFFA2309AFB63FACB19DFFA27" box="[1169,1213,1282,1427]" gridcol="3" gridrow="0" pageId="11" pageNumber="56">SD 0.13 4.67 5.36</th>
<th id="769D702EFFA2309AFAFCFACB18FEFA27" box="[1294,1436,1282,1427]" gridcol="4" gridrow="0" pageId="11" pageNumber="56">N 74 74 74</th>
</tr>
<tr id="354C1952FFA2309AFF65FA6E18FEFA52" box="[151,1436,1447,1510]" gridrow="1" pageId="11" pageNumber="56">
<th id="769D702EFFA2309AFF65FA6E1FADFA52" box="[151,719,1447,1510]" gridcol="0" gridrow="1" pageId="11" pageNumber="56">Palps reach chaetiger Lateral antennae reach chaetiger</th>
<td id="769D702EFFA2309AFCEBFA6E1E1CFA52" box="[793,894,1447,1510]" gridcol="1" gridrow="1" pageId="11" pageNumber="56">1 5–13</td>
<td id="769D702EFFA2309AFC13FA6E195EFA52" box="[993,1084,1447,1510]" gridcol="2" gridrow="1" pageId="11" pageNumber="56">invariant 8.49</td>
<td id="769D702EFFA2309AFB63FA6E19DFFA52" box="[1169,1213,1447,1510]" gridcol="3" gridrow="1" pageId="11" pageNumber="56">1.60</td>
<td id="769D702EFFA2309AFAFCFA6E18FEFA52" box="[1294,1436,1447,1510]" gridcol="4" gridrow="1" pageId="11" pageNumber="56">74 71</td>
</tr>
<tr id="354C1952FFA2309AFF65FA3318FEF93F" box="[151,1436,1530,1675]" gridrow="2" pageId="11" pageNumber="56">
<th id="769D702EFFA2309AFF65FA331FADF93F" box="[151,719,1530,1675]" gridcol="0" gridrow="2" pageId="11" pageNumber="56">Median antenna reaches chaetiger Maximal number of rings in lateral ceratophores Number of chaetigers with developed postchaetal lobes Number of subulate ventral cirri</th>
<td id="769D702EFFA2309AFCEBFA331E1CF93F" box="[793,894,1530,1675]" gridcol="1" gridrow="2" pageId="11" pageNumber="56">7–16 1–3 8–16 2</td>
<td id="769D702EFFA2309AFC13FA33195EF93F" box="[993,1084,1530,1675]" gridcol="2" gridrow="2" pageId="11" pageNumber="56">12.34 2.18 11.32 invariant</td>
<td id="769D702EFFA2309AFB63FA3319DFF93F" box="[1169,1213,1530,1675]" gridcol="3" gridrow="2" pageId="11" pageNumber="56">2.08 0.58 2.02</td>
<td id="769D702EFFA2309AFAFCFA3318FEF93F" box="[1294,1436,1530,1675]" gridcol="4" gridrow="2" pageId="11" pageNumber="56">71 74 73 74</td>
</tr>
<tr id="354C1952FFA2309AFF65F95618FEF96A" box="[151,1436,1695,1758]" gridrow="3" pageId="11" pageNumber="56">
<th id="769D702EFFA2309AFF65F9561FADF96A" box="[151,719,1695,1758]" gridcol="0" gridrow="3" pageId="11" pageNumber="56">Number of chaetigers with falcigers Subacicular hooks first present on chaetiger</th>
<td id="769D702EFFA2309AFCEBF9561E1CF96A" box="[793,894,1695,1758]" gridcol="1" gridrow="3" pageId="11" pageNumber="56">1 28–39</td>
<td id="769D702EFFA2309AFC13F956195EF96A" box="[993,1084,1695,1758]" gridcol="2" gridrow="3" pageId="11" pageNumber="56">invariant 34.13</td>
<td id="769D702EFFA2309AFB63F95619DFF96A" box="[1169,1213,1695,1758]" gridcol="3" gridrow="3" pageId="11" pageNumber="56">2.49</td>
<td id="769D702EFFA2309AFAFCF95618FEF96A" box="[1294,1436,1695,1758]" gridcol="4" gridrow="3" pageId="11" pageNumber="56">74 67</td>
</tr>
<tr id="354C1952FFA2309AFF65F93B18FEF8BC" box="[151,1436,1778,1800]" gridrow="4" pageId="11" pageNumber="56">
<th id="769D702EFFA2309AFF65F93B1FADF8BC" box="[151,719,1778,1800]" gridcol="0" gridrow="4" pageId="11" pageNumber="56">Number of anal cirri</th>
<td id="769D702EFFA2309AFCEBF93B1E1CF8BC" box="[793,894,1778,1800]" gridcol="1" gridrow="4" pageId="11" pageNumber="56">2–5</td>
<td id="769D702EFFA2309AFC13F93B195EF8BC" box="[993,1084,1778,1800]" gridcol="2" gridrow="4" pageId="11" pageNumber="56">2.19</td>
<td id="769D702EFFA2309AFB63F93B19DFF8BC" box="[1169,1213,1778,1800]" gridcol="3" gridrow="4" pageId="11" pageNumber="56">0.59</td>
<td id="769D702EFFA2309AFAFCF93B18FEF8BC" box="[1294,1436,1778,1800]" gridcol="4" gridrow="4" pageId="11" pageNumber="56">74</td>
</tr>
</table>
</paragraph>
<paragraph id="8BC31B10FFA2CF6EFF37F88E19FEF813" blockId="11.[151,1437,1863,2031]" pageId="11" pageNumber="56">
Mandibles with almost straight distal margins (
<figureCitation id="13470795FFA2CF6EFD28F88E1E7EF8EB" box="[730,796,1863,1887]" captionStart="FIGURE 2" captionStartId="7.[151,250,1744,1767]" captionTargetBox="[151,1436,193,1722]" captionTargetId="figure@7.[151,1436,193,1723]" captionTargetPageId="7" captionText="FIGURE 2. Leptoecia midatlantica sp. nov. A, anterior region of the holotype, ventral view; B, same, dorsal view; C – E shape of the prostomium, ventral view; F, maxillae; G, mandibles; H, pygidium of the holotype, ventral view; I, same, dorsal view; J – L, pygidia, dorsal view." httpUri="https://zenodo.org/record/279915/files/figure.png" pageId="11" pageNumber="56">Fig. 2</figureCitation>
G, 3F). Maxillae weakly sclerotized; carriers longer than wide; distal parts of MxI pointed, dark brown in colour (
<figureCitation id="13470795FFA2CF6EFCC7F8A21E1EF837" box="[821,892,1899,1923]" captionStart="FIGURE 2" captionStartId="7.[151,250,1744,1767]" captionTargetBox="[151,1436,193,1722]" captionTargetId="figure@7.[151,1436,193,1723]" captionTargetPageId="7" captionText="FIGURE 2. Leptoecia midatlantica sp. nov. A, anterior region of the holotype, ventral view; B, same, dorsal view; C – E shape of the prostomium, ventral view; F, maxillae; G, mandibles; H, pygidium of the holotype, ventral view; I, same, dorsal view; J – L, pygidia, dorsal view." httpUri="https://zenodo.org/record/279915/files/figure.png" pageId="11" pageNumber="56">Fig. 2</figureCitation>
F). Maxillary formula (based on
<specimenCount id="9D7AD099FFA2CF6EFB0FF8A218F3F837" box="[1277,1425,1899,1923]" pageId="11" pageNumber="56" type="generic">3 specimens</specimenCount>
): MxI=1+1; MxII=13–18+13–17; MxIII=10–13+0; MxIV=10–12+11–17. Maxillae V absent.
</paragraph>
<paragraph id="8BC31B10FFA2CF6EFF37F87A185BF85B" blockId="11.[151,1437,1863,2031]" pageId="11" pageNumber="56">
Pygidium conical, covered with glandular buds (
<figureCitation id="13470795FFA2CF6EFD13F87A1E40F87F" box="[737,802,1971,1995]" captionStart="FIGURE 3" captionStartId="8.[151,250,1892,1915]" captionTargetBox="[151,1436,193,1871]" captionTargetId="figure@8.[151,1436,193,1871]" captionTargetPageId="8" captionText="FIGURE 3. Leptoecia midatlantica sp. nov. A, C, anterior region of the body, ventral view; B, same, dorsal view; D, F prostomium and peristomium, ventral view; E, same, dorsal view; G, sensory organs on prostomium, dorsal view; H, posterior region of the body, ventral view; I, pygidium, dorsal view; J pygidial glandular buds; K, posterior chaetigers, dorsal view. ac—anal cirrus, dc—dorsal cirrus, fl—frontal lip, la—lateral antenna, ll—lower lip, ma—median antenna, mnd—mandibles, p—palp, per—peristomium, pg—pygidium, pl—postchaetal lobe, prl—prechaetal lobe, ul—upper lip, vc—ventral cirrus." httpUri="https://zenodo.org/record/279916/files/figure.png" pageId="11" pageNumber="56">Fig. 3</figureCitation>
H–J), with anus opening dorsally (
<figureCitation id="13470795FFA2CF6EFB50F87A1981F87F" box="[1186,1251,1971,1995]" captionStart="FIGURE 2" captionStartId="7.[151,250,1744,1767]" captionTargetBox="[151,1436,193,1722]" captionTargetId="figure@7.[151,1436,193,1723]" captionTargetPageId="7" captionText="FIGURE 2. Leptoecia midatlantica sp. nov. A, anterior region of the holotype, ventral view; B, same, dorsal view; C – E shape of the prostomium, ventral view; F, maxillae; G, mandibles; H, pygidium of the holotype, ventral view; I, same, dorsal view; J – L, pygidia, dorsal view." httpUri="https://zenodo.org/record/279915/files/figure.png" pageId="11" pageNumber="56">
Fig.
<date id="FFC23DD0FFA2CF6EFB27F87A1981F87F" box="[1237,1251,1971,1995]" pageId="11" pageNumber="56">2</date>
</figureCitation>
I–L). Number of anal cirri varying from 2 to 5, but most examined specimens (90%) with two anal cirri (
<figureCitation id="13470795FFA2CF6EFB9EF81E19CCF85B" box="[1132,1198,2007,2031]" captionStart="FIGURE 2" captionStartId="7.[151,250,1744,1767]" captionTargetBox="[151,1436,193,1722]" captionTargetId="figure@7.[151,1436,193,1723]" captionTargetPageId="7" captionText="FIGURE 2. Leptoecia midatlantica sp. nov. A, anterior region of the holotype, ventral view; B, same, dorsal view; C – E shape of the prostomium, ventral view; F, maxillae; G, mandibles; H, pygidium of the holotype, ventral view; I, same, dorsal view; J – L, pygidia, dorsal view." httpUri="https://zenodo.org/record/279915/files/figure.png" pageId="11" pageNumber="56">Fig. 2</figureCitation>
H–L, 3H, I).
</paragraph>
<paragraph id="8BC31B10FFA5CF69FF37FF5E19AAFF60" blockId="12.[151,1437,151,860]" pageId="12" pageNumber="57">
Tubes translucent, thin-walled, dorsoventrally flattened with distinct longitudinal thickened ribs along both lateral margins.
<typeStatus id="54C7A5B2FFA5CF69FEDDFF751CF5FF60" box="[303,407,188,212]" pageId="12" pageNumber="57" type="holotype">Holotype</typeStatus>
containing spherical oocytes in its body cavity around chaetigers 19–28.
</paragraph>
</subSubSection>
<subSubSection id="C366489BFFA5CF69FF37FF2919AEFD78" pageId="12" pageNumber="57" type="discussion">
<paragraph id="8BC31B10FFA5CF69FF37FF2919AEFD78" blockId="12.[151,1437,151,860]" pageId="12" pageNumber="57">
<emphasis id="B908C702FFA5CF69FF37FF291C5BFF4C" bold="true" box="[197,313,224,248]" pageId="12" pageNumber="57">Remarks.</emphasis>
<taxonomicName id="4C7C6093FFA5CF69FEB0FF281F25FF4C" box="[322,583,224,248]" class="Polychaeta" family="Onuphidae" genus="Leptoecia" kingdom="Animalia" order="Eunicida" pageId="12" pageNumber="57" phylum="Annelida" rank="species" species="midatlantica" status="sp. nov.">
<emphasis id="B908C702FFA5CF69FEB0FF281F25FF4C" box="[322,583,224,248]" italics="true" pageId="12" pageNumber="57">Leptoecia midatlantica</emphasis>
</taxonomicName>
<emphasis id="B908C702FFA5CF69FDA2FF291FD1FF4C" bold="true" box="[592,691,224,248]" pageId="12" pageNumber="57">
<taxonomicNameLabel id="A23B7A79FFA5CF69FDA2FF291FCFFF4C" box="[592,685,224,248]" pageId="12" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
can be distinguished from its congeners by the presence of both uni- and bidentate simple falcigers on the first pair of parapodia. However this character was variable in the studied material. Some specimens displayed the presence of only bidentate falcigers, but never only unidentate falcigers.
<taxonomicName id="4C7C6093FFA5CF69FA77FEE01C47FED0" class="Polychaeta" family="Onuphidae" genus="Leptoecia" kingdom="Animalia" order="Eunicida" pageId="12" pageNumber="57" phylum="Annelida" rank="species" species="midatlantica" status="sp. nov.">
<emphasis id="B908C702FFA5CF69FA77FEE01C47FED0" italics="true" pageId="12" pageNumber="57">L. midatlantica</emphasis>
</taxonomicName>
<emphasis id="B908C702FFA5CF69FEDEFE851CECFED0" bold="true" box="[300,398,332,356]" pageId="12" pageNumber="57">
<taxonomicNameLabel id="A23B7A79FFA5CF69FEDEFE851CEAFED0" box="[300,392,332,356]" pageId="12" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
is similar to
<taxonomicName id="4C7C6093FFA5CF69FDD1FE841FA5FED0" box="[547,711,332,356]" class="Polychaeta" family="Onuphidae" genus="Leptoecia" kingdom="Animalia" order="Eunicida" pageId="12" pageNumber="57" phylum="Annelida" rank="species" species="benthaliana">
<emphasis id="B908C702FFA5CF69FDD1FE841FA5FED0" box="[547,711,332,356]" italics="true" pageId="12" pageNumber="57">L. benthaliana</emphasis>
</taxonomicName>
in having falcigers on the first chaetiger only, dorsal cirri on all parapodia, and subacicular hooks starting in the middle of the body (around chaetiger 35).
<bibRefCitation id="EFED66E1FFA5CF69FB61FEA6182DFE33" author="Orensanz" box="[1171,1359,367,391]" pageId="12" pageNumber="57" refString="Orensanz, J. M. (1990) The Eunicemorph polychaete annelids from Antarctic and Subantarctic Seas. With addenda to the Eunicemorpha of Argentina, Chile, New Zealand, Australia, and the Southern Indian Ocean. Antarctic Research Series Biology of the Antarctic Seas XXI, 52, 1 - 183." type="journal article" year="1990">Orensanz (1990)</bibRefCitation>
examined ca.
<specimenCount id="9D7AD099FFA5CF69FF06FE5D1CE9FE18" box="[244,395,404,428]" pageId="12" pageNumber="57" type="generic">60 specimens</specimenCount>
of
<taxonomicName id="4C7C6093FFA5CF69FE5DFE5C1F30FE18" box="[431,594,404,428]" class="Polychaeta" family="Onuphidae" genus="Leptoecia" kingdom="Animalia" order="Eunicida" pageId="12" pageNumber="57" phylum="Annelida" rank="species" species="benthaliana">
<emphasis id="B908C702FFA5CF69FE5DFE5C1F30FE18" box="[431,594,404,428]" italics="true" pageId="12" pageNumber="57">L. benthaliana</emphasis>
</taxonomicName>
describing the variation in the shape of the prostomium (slightly bilobed or pointed) and shape of the tube (circular in cross section or with lateral ribs on one or both sides). The examined specimens of
<taxonomicName id="4C7C6093FFA5CF69FEC2FE141CB9FE40" box="[304,475,476,500]" class="Polychaeta" family="Onuphidae" genus="Leptoecia" kingdom="Animalia" order="Eunicida" pageId="12" pageNumber="57" phylum="Annelida" rank="species" species="midatlantica" status="sp. nov.">
<emphasis id="B908C702FFA5CF69FEC2FE141CB9FE40" box="[304,475,476,500]" italics="true" pageId="12" pageNumber="57">L. midatlantica</emphasis>
</taxonomicName>
<emphasis id="B908C702FFA5CF69FE10FE151F21FE40" bold="true" box="[482,579,476,500]" pageId="12" pageNumber="57">
<taxonomicNameLabel id="A23B7A79FFA5CF69FE10FE151F5FFE40" box="[482,573,476,500]" pageId="12" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
also demonstrated the variation in the shape of the prostomium and the variable degree of frontal and upper lips development. Among
<specimenCount id="9D7AD099FFA5CF69FCDCFE361EA4FDA3" box="[814,966,511,535]" pageId="12" pageNumber="57" type="generic">74 specimens</specimenCount>
examined, 42 had a slightly or noticeably bilobed prostomium (
<figureCitation id="13470795FFA5CF69FE7EFDED1CADFD88" box="[396,463,548,572]" captionStart="FIGURE 2" captionStartId="7.[151,250,1744,1767]" captionTargetBox="[151,1436,193,1722]" captionTargetId="figure@7.[151,1436,193,1723]" captionTargetPageId="7" captionText="FIGURE 2. Leptoecia midatlantica sp. nov. A, anterior region of the holotype, ventral view; B, same, dorsal view; C – E shape of the prostomium, ventral view; F, maxillae; G, mandibles; H, pygidium of the holotype, ventral view; I, same, dorsal view; J – L, pygidia, dorsal view." httpUri="https://zenodo.org/record/279915/files/figure.png" pageId="12" pageNumber="57">Fig. 2</figureCitation>
A, B, E, 3E, F) and 32 had a conical or pointed prostomium (
<figureCitation id="13470795FFA5CF69FB7DFDED19B6FD88" box="[1167,1236,548,572]" captionStart="FIGURE 2" captionStartId="7.[151,250,1744,1767]" captionTargetBox="[151,1436,193,1722]" captionTargetId="figure@7.[151,1436,193,1723]" captionTargetPageId="7" captionText="FIGURE 2. Leptoecia midatlantica sp. nov. A, anterior region of the holotype, ventral view; B, same, dorsal view; C – E shape of the prostomium, ventral view; F, maxillae; G, mandibles; H, pygidium of the holotype, ventral view; I, same, dorsal view; J – L, pygidia, dorsal view." httpUri="https://zenodo.org/record/279915/files/figure.png" pageId="12" pageNumber="57">Fig. 2</figureCitation>
C, D, 3A, D). All examined specimens had flattened tubes; no tubes circular in cross-section were found in the studied material.
<taxonomicName id="4C7C6093FFA5CF69FA77FD801C47FD30" class="Polychaeta" family="Onuphidae" genus="Leptoecia" kingdom="Animalia" order="Eunicida" pageId="12" pageNumber="57" phylum="Annelida" rank="species" species="midatlantica" status="sp. nov.">
<emphasis id="B908C702FFA5CF69FA77FD801C47FD30" italics="true" pageId="12" pageNumber="57">L. midatlantica</emphasis>
</taxonomicName>
<emphasis id="B908C702FFA5CF69FED9FDA51CE9FD30" bold="true" box="[299,395,620,644]" pageId="12" pageNumber="57">
<taxonomicNameLabel id="A23B7A79FFA5CF69FED9FDA51CE7FD30" box="[299,389,620,644]" pageId="12" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
can be distinguished from
<taxonomicName id="4C7C6093FFA5CF69FD4AFDA41E38FD30" box="[696,858,620,644]" class="Polychaeta" family="Onuphidae" genus="Leptoecia" kingdom="Animalia" order="Eunicida" pageId="12" pageNumber="57" phylum="Annelida" rank="species" species="benthaliana">
<emphasis id="B908C702FFA5CF69FD4AFDA41E38FD30" box="[696,858,620,644]" italics="true" pageId="12" pageNumber="57">L. benthaliana</emphasis>
</taxonomicName>
by the shape of anterior falcigers. In
<taxonomicName id="4C7C6093FFA5CF69FB0BFDA418F9FD30" box="[1273,1435,620,644]" class="Polychaeta" family="Onuphidae" genus="Leptoecia" kingdom="Animalia" order="Eunicida" pageId="12" pageNumber="57" phylum="Annelida" rank="species" species="benthaliana">
<emphasis id="B908C702FFA5CF69FB0BFDA418F9FD30" box="[1273,1435,620,644]" italics="true" pageId="12" pageNumber="57">L. benthaliana</emphasis>
</taxonomicName>
all falcigers independently on their size have subdistal reduction in their diameter (
<bibRefCitation id="EFED66E1FFA5CF69FBBEFD46199BFD13" author="Orensanz" box="[1100,1273,655,679]" pageId="12" pageNumber="57" refString="Orensanz, J. M. (1990) The Eunicemorph polychaete annelids from Antarctic and Subantarctic Seas. With addenda to the Eunicemorpha of Argentina, Chile, New Zealand, Australia, and the Southern Indian Ocean. Antarctic Research Series Biology of the Antarctic Seas XXI, 52, 1 - 183." type="journal article" year="1990">Orensanz 1990</bibRefCitation>
) (
<figureCitation id="13470795FFA5CF69FAE1FD461835FD13" box="[1299,1367,655,679]" captionStart="FIGURE 6" captionStartId="11.[151,250,1073,1096]" captionTargetBox="[219,1369,327,1053]" captionTargetId="figure@11.[219,1370,327,1053]" captionTargetPageId="11" captionText="FIGURE 6. Leptoecia benthaliana. A – C, anterior falcigers (the black arrows indicate subdistal reduction in the diameter of the falciger, modified from Orensanz 1990); Leptoecia midatlantica sp. nov. D, E, simple unidentate falcigers from chaetiger 1; F, G, simple bidentate falcigers from chaetiger 1; H, simple bidentate hook from chaetiger 35." httpUri="https://zenodo.org/record/279919/files/figure.png" pageId="12" pageNumber="57">Fig. 6</figureCitation>
A–C), whereas in
<taxonomicName id="4C7C6093FFA5CF69FEE4FD7C1CA3FD78" box="[278,449,692,716]" class="Polychaeta" family="Onuphidae" genus="Leptoecia" kingdom="Animalia" order="Eunicida" pageId="12" pageNumber="57" phylum="Annelida" rank="species" species="midatlantica" status="sp. nov.">
<emphasis id="B908C702FFA5CF69FEE4FD7C1CA3FD78" box="[278,449,692,716]" italics="true" pageId="12" pageNumber="57">L. midatlantica</emphasis>
</taxonomicName>
<emphasis id="B908C702FFA5CF69FE3AFD7D1F4BFD78" bold="true" box="[456,553,692,716]" pageId="12" pageNumber="57">
<taxonomicNameLabel id="A23B7A79FFA5CF69FE3AFD7D1F41FD78" box="[456,547,692,716]" pageId="12" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
falcigers uniform in their diameter throughout (
<figureCitation id="13470795FFA5CF69FBB0FD7D19E7FD78" box="[1090,1157,692,716]" captionStart="FIGURE 6" captionStartId="11.[151,250,1073,1096]" captionTargetBox="[219,1369,327,1053]" captionTargetId="figure@11.[219,1370,327,1053]" captionTargetPageId="11" captionText="FIGURE 6. Leptoecia benthaliana. A – C, anterior falcigers (the black arrows indicate subdistal reduction in the diameter of the falciger, modified from Orensanz 1990); Leptoecia midatlantica sp. nov. D, E, simple unidentate falcigers from chaetiger 1; F, G, simple bidentate falcigers from chaetiger 1; H, simple bidentate hook from chaetiger 35." httpUri="https://zenodo.org/record/279919/files/figure.png" pageId="12" pageNumber="57">Fig. 6</figureCitation>
D–G).
</paragraph>
</subSubSection>
<subSubSection id="C366489BFFA5CF69FF37FD1E1871FCA0" pageId="12" pageNumber="57" type="distribution">
<paragraph id="8BC31B10FFA5CF69FF37FD1E1871FCA0" blockId="12.[151,1437,151,860]" pageId="12" pageNumber="57">
<emphasis id="B908C702FFA5CF69FF37FD1E1C3CFD44" bold="true" box="[197,350,727,752]" pageId="12" pageNumber="57">Distribution.</emphasis>
<taxonomicName id="4C7C6093FFA5CF69FE96FD101F05FD44" box="[356,615,728,752]" class="Polychaeta" family="Onuphidae" genus="Leptoecia" kingdom="Animalia" order="Eunicida" pageId="12" pageNumber="57" phylum="Annelida" rank="species" species="midatlantica" status="sp. nov.">
<emphasis id="B908C702FFA5CF69FE96FD101F05FD44" box="[356,615,728,752]" italics="true" pageId="12" pageNumber="57">Leptoecia midatlantica</emphasis>
</taxonomicName>
<emphasis id="B908C702FFA5CF69FD9FFD111FACFD44" bold="true" box="[621,718,728,752]" pageId="12" pageNumber="57">
<taxonomicNameLabel id="A23B7A79FFA5CF69FD9FFD111FAAFD44" box="[621,712,728,752]" pageId="12" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
is known from the northern part of the Mid-Atlantic Ridge from the north off the Azores to the southern tip of the Reykjanes ridge (
<figureCitation id="13470795FFA5CF69FC75FD351EAAFCA0" box="[903,968,764,788]" captionStart="FIGURE 1" captionStartId="5.[151,250,1394,1417]" captionTargetBox="[151,1436,181,1373]" captionTargetId="figure@5.[151,1436,181,1373]" captionTargetPageId="5" captionText="FIGURE 1. Distribution of Leptoecia midatlantica sp. nov. The black arrow indicates the type location; 2500 m, 3000 m and 4500 m isobaths shown." httpUri="https://zenodo.org/record/279914/files/figure.png" pageId="12" pageNumber="57">Fig. 1</figureCitation>
). Depth range
<quantity id="4C84B6F5FFA5CF69FB83FD35186DFCA0" box="[1137,1295,764,788]" metricMagnitude="3" metricUnit="m" metricValue="2.4305" metricValueMax="2.754" metricValueMin="2.107" pageId="12" pageNumber="57" unit="m" value="2430.5" valueMax="2754.0" valueMin="2107.0">2107–2754 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C366489BFFA5CF69FF37FCE91956FCE8" pageId="12" pageNumber="57" type="etymology">
<paragraph id="8BC31B10FFA5CF69FF37FCE91956FCE8" blockId="12.[151,1437,151,860]" pageId="12" pageNumber="57">
<emphasis id="B908C702FFA5CF69FF37FCE91C2EFC8C" bold="true" box="[197,332,800,824]" pageId="12" pageNumber="57">Etymology.</emphasis>
The specific name,
<taxonomicName id="4C7C6093FFA5CF69FDCEFCE91FABFC8C" box="[572,713,800,824]" class="Polychaeta" family="Onuphidae" genus="Leptoecia" kingdom="Animalia" order="Eunicida" pageId="12" pageNumber="57" phylum="Annelida" rank="species" species="midatlantica">
<emphasis id="B908C702FFA5CF69FDCEFCE91FABFC8C" box="[572,713,800,824]" italics="true" pageId="12" pageNumber="57">midatlantica</emphasis>
</taxonomicName>
, refers to the geographical range of the new species that was reported from the deep-sea habitats of the northern part of the Mid-Atlantic Ridge.
</paragraph>
</subSubSection>
</treatment>
</document>
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