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<document id="82A5D1A2C8EC9983AEA9EF8E6CB1FACA" ID-DOI="10.11646/zootaxa.5419.2.8" ID-ISSN="1175-5326" ID-Zenodo-Dep="10792148" ID-ZooBank="9E853730-34F6-4990-99DB-30B7E1F402EF" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1709795875585" checkinUser="plazi" docAuthor="Lv, Shasha, Yang, Lin &amp; Chen, Xiangsheng" docDate="2024" docId="03C4A67B3031FFBFFF05F987FD09FE1B" docLanguage="en" docName="zootaxa.5419.2.8.pdf" docOrigin="Zootaxa 5419 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5419.2.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Metanigrus angularus Lv &amp; Yang &amp; Chen 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="295" masterDocId="FFFDDE033032FFB9FF92FF94FFFAFF93" masterDocTitle="Two new species of the planthopper genus Metanigrus Tsaur, Yang &amp; Wilson from China (Hemiptera: Fulgoromorpha: Meenoplidae), with an updated checklist and key to species" masterLastPageNumber="295" masterPageNumber="289" pageNumber="292" updateTime="1709796508692" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="8CE3B288F1CE680E4A4C90675E7B4F57">Two new species of the planthopper genus Metanigrus Tsaur, Yang &amp; Wilson from China (Hemiptera: Fulgoromorpha: Meenoplidae), with an updated checklist and key to species</mods:title>
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<mods:namePart id="48F96F9886F109E1D316B40D4A6B1B4F">Lv, Shasha</mods:namePart>
<mods:affiliation id="974640975F94E43633EBEA5F89BB259E">Institute of Entomology, Guizhou University, Guiyang, Guizhou 550025, P. R. China. &amp; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources of Guizhou, Guizhou University, Guiyang, Guizhou, 550025 P. R. China.</mods:affiliation>
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<mods:namePart id="3359778465403FF95B3C74C4F7134B6F">Yang, Lin</mods:namePart>
<mods:affiliation id="02F9E085AAF013FE610A8B12EBE527AA">Institute of Entomology, Guizhou University, Guiyang, Guizhou 550025, P. R. China. &amp; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources of Guizhou, Guizhou University, Guiyang, Guizhou, 550025 P. R. China.</mods:affiliation>
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<mods:namePart id="94ED3A8460141EB7D7DCE71D4C02B1C6">Chen, Xiangsheng</mods:namePart>
<mods:affiliation id="1F645B2E3089E087295CBDF6C8B5517D">Institute of Entomology, Guizhou University, Guiyang, Guizhou 550025, P. R. China. &amp; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources of Guizhou, Guizhou University, Guiyang, Guizhou, 550025 P. R. China.</mods:affiliation>
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<mods:date id="D02727AE503176205BDF5D1C64CE7E3A">2024</mods:date>
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<subSubSection id="C37744E63031FFBAFF05F987FDF6F9BE" box="[151,524,1554,1581]" pageId="3" pageNumber="292" type="nomenclature">
<paragraph id="8BD2176D3031FFBAFF05F987FDF6F9BE" blockId="3.[151,524,1554,1617]" box="[151,524,1554,1581]" pageId="3" pageNumber="292">
<heading id="D09AA0013031FFBAFF05F987FDF6F9BE" bold="true" box="[151,524,1554,1581]" fontSize="11" level="1" pageId="3" pageNumber="292" reason="1">
<emphasis id="B919CB7F3031FFBAFF05F987FDF6F9BE" bold="true" box="[151,524,1554,1581]" pageId="3" pageNumber="292">
<taxonomicName id="4C6D6CEE3031FFBAFF05F987FE5DF9BE" authority="Lv &amp; Yang &amp; Chen, 2024" authorityName="Lv &amp; Yang &amp; Chen" authorityYear="2024" box="[151,423,1554,1581]" class="Insecta" family="Meenoplidae" genus="Metanigrus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="292" phylum="Arthropoda" rank="species" species="angularus" status="sp. nov.">
<emphasis id="B919CB7F3031FFBAFF05F987FE5DF9BE" bold="true" box="[151,423,1554,1581]" italics="true" pageId="3" pageNumber="292">Metanigrus angularus</emphasis>
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<taxonomicNameLabel id="A22A76043031FFBAFE3CF987FDF6F9BE" box="[430,524,1555,1581]" pageId="3" pageNumber="292" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C37744E63031FFBAFF05F9A3FAC5F90A" pageId="3" pageNumber="292" type="description">
<paragraph id="8BD2176D3031FFBAFF05F9A3FEEFF9C2" blockId="3.[151,524,1554,1617]" box="[151,277,1591,1617]" pageId="3" pageNumber="292">
<figureCitation id="13560BE83031FFBAFF05F9A3FEEFF9C2" box="[151,277,1591,1617]" captionStart="FIGURES 1426" captionStartId="4.[151,264,1786,1810]" captionTargetBox="[158,1429,188,1756]" captionTargetId="figure-23@4.[151,1436,181,1763]" captionTargetPageId="4" captionText="FIGURES 1426. Metanigrus angularus sp. nov., male 14. Habitus, dorsal view; 15. Habitus, lateral view; 16. Head and thorax, dorsal view; 17. Frons, ventral view; 18. Forewing; 19. Pygofer, lateral view; 20. Pygofer, ventral view; 21. Hindwing; 22. Anal tube, dorsal view; 23. Gonostyli, lateral view; 24. Gonostyli, dorsal view; 25. Aedeagus, ventral view; 26. Aedeagus, lateral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10792154" httpUri="https://zenodo.org/record/10792154/files/figure.png" pageId="3" pageNumber="292">Figs 1426</figureCitation>
</paragraph>
<paragraph id="8BD2176D3031FFBAFF05F9EBFAC5F90A" blockId="3.[151,1437,1662,2013]" box="[151,1343,1662,1689]" pageId="3" pageNumber="292">
<emphasis id="B919CB7F3031FFBAFF05F9EBFEB4F90A" bold="true" box="[151,334,1663,1689]" pageId="3" pageNumber="292">Measurements.</emphasis>
Length (including forewing): male 3.804.11 mm (N = 5), female 4.534.85 mm (N = 4).
</paragraph>
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<subSubSection id="C37744E63031FFBCFF55F936FBC0F8AD" lastPageId="5" lastPageNumber="294" pageId="3" pageNumber="292" type="diagnosis">
<paragraph id="8BD2176D3031FFBAFF55F936FC0EF8DE" blockId="3.[151,1437,1662,2013]" pageId="3" pageNumber="292">
<emphasis id="B919CB7F3031FFBAFF55F936FEB8F92F" bold="true" box="[199,322,1698,1724]" pageId="3" pageNumber="292">Diagnosis.</emphasis>
Upper part of frons (
<figureCitation id="13560BE83031FFBAFDABF937FD76F92E" box="[569,652,1699,1725]" captionStart="FIGURES 1426" captionStartId="4.[151,264,1786,1810]" captionTargetBox="[158,1429,188,1756]" captionTargetId="figure-23@4.[151,1436,181,1763]" captionTargetPageId="4" captionText="FIGURES 1426. Metanigrus angularus sp. nov., male 14. Habitus, dorsal view; 15. Habitus, lateral view; 16. Head and thorax, dorsal view; 17. Frons, ventral view; 18. Forewing; 19. Pygofer, lateral view; 20. Pygofer, ventral view; 21. Hindwing; 22. Anal tube, dorsal view; 23. Gonostyli, lateral view; 24. Gonostyli, dorsal view; 25. Aedeagus, ventral view; 26. Aedeagus, lateral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10792154" httpUri="https://zenodo.org/record/10792154/files/figure.png" pageId="3" pageNumber="292">Fig. 17</figureCitation>
) grayish yellow, middle and basal parts yellowish brown; forewings (
<figureCitation id="13560BE83031FFBAFF0DF953FF08F972" box="[159,242,1735,1761]" captionStart="FIGURES 1426" captionStartId="4.[151,264,1786,1810]" captionTargetBox="[158,1429,188,1756]" captionTargetId="figure-23@4.[151,1436,181,1763]" captionTargetPageId="4" captionText="FIGURES 1426. Metanigrus angularus sp. nov., male 14. Habitus, dorsal view; 15. Habitus, lateral view; 16. Head and thorax, dorsal view; 17. Frons, ventral view; 18. Forewing; 19. Pygofer, lateral view; 20. Pygofer, ventral view; 21. Hindwing; 22. Anal tube, dorsal view; 23. Gonostyli, lateral view; 24. Gonostyli, dorsal view; 25. Aedeagus, ventral view; 26. Aedeagus, lateral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10792154" httpUri="https://zenodo.org/record/10792154/files/figure.png" pageId="3" pageNumber="292">Fig. 18</figureCitation>
) with black spots, veins brownish black; apex of pygofer (
<figureCitation id="13560BE83031FFBAFC19F953FC27F972" box="[907,989,1735,1761]" captionStart="FIGURES 1426" captionStartId="4.[151,264,1786,1810]" captionTargetBox="[158,1429,188,1756]" captionTargetId="figure-23@4.[151,1436,181,1763]" captionTargetPageId="4" captionText="FIGURES 1426. Metanigrus angularus sp. nov., male 14. Habitus, dorsal view; 15. Habitus, lateral view; 16. Head and thorax, dorsal view; 17. Frons, ventral view; 18. Forewing; 19. Pygofer, lateral view; 20. Pygofer, ventral view; 21. Hindwing; 22. Anal tube, dorsal view; 23. Gonostyli, lateral view; 24. Gonostyli, dorsal view; 25. Aedeagus, ventral view; 26. Aedeagus, lateral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10792154" httpUri="https://zenodo.org/record/10792154/files/figure.png" pageId="3" pageNumber="292">Fig. 20</figureCitation>
) each with an obtuse process in ventral view; dorsolateral and middle parts of anal tube (
<figureCitation id="13560BE83031FFBAFD2CF97FFCEAF896" box="[702,784,1771,1797]" captionStart="FIGURES 1426" captionStartId="4.[151,264,1786,1810]" captionTargetBox="[158,1429,188,1756]" captionTargetId="figure-23@4.[151,1436,181,1763]" captionTargetPageId="4" captionText="FIGURES 1426. Metanigrus angularus sp. nov., male 14. Habitus, dorsal view; 15. Habitus, lateral view; 16. Head and thorax, dorsal view; 17. Frons, ventral view; 18. Forewing; 19. Pygofer, lateral view; 20. Pygofer, ventral view; 21. Hindwing; 22. Anal tube, dorsal view; 23. Gonostyli, lateral view; 24. Gonostyli, dorsal view; 25. Aedeagus, ventral view; 26. Aedeagus, lateral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10792154" httpUri="https://zenodo.org/record/10792154/files/figure.png" pageId="3" pageNumber="292">Fig. 19</figureCitation>
) each with a fingerlike process in lateral view, basolateral fingerlike process wide, apex curved dorsad; aedeagus (
<figureCitation id="13560BE83031FFBAFC90F89BFC7AF8BA" box="[770,896,1807,1833]" captionStart="FIGURES 1426" captionStartId="4.[151,264,1786,1810]" captionTargetBox="[158,1429,188,1756]" captionTargetId="figure-23@4.[151,1436,181,1763]" captionTargetPageId="4" captionText="FIGURES 1426. Metanigrus angularus sp. nov., male 14. Habitus, dorsal view; 15. Habitus, lateral view; 16. Head and thorax, dorsal view; 17. Frons, ventral view; 18. Forewing; 19. Pygofer, lateral view; 20. Pygofer, ventral view; 21. Hindwing; 22. Anal tube, dorsal view; 23. Gonostyli, lateral view; 24. Gonostyli, dorsal view; 25. Aedeagus, ventral view; 26. Aedeagus, lateral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10792154" httpUri="https://zenodo.org/record/10792154/files/figure.png" pageId="3" pageNumber="292">Figs 19, 26</figureCitation>
) slender and long, apex curved ventrad in lateral view; outer process of gonostyli (
<figureCitation id="13560BE83031FFBAFD9FF8A7FD77F8DE" box="[525,653,1843,1869]" captionStart="FIGURES 1426" captionStartId="4.[151,264,1786,1810]" captionTargetBox="[158,1429,188,1756]" captionTargetId="figure-23@4.[151,1436,181,1763]" captionTargetPageId="4" captionText="FIGURES 1426. Metanigrus angularus sp. nov., male 14. Habitus, dorsal view; 15. Habitus, lateral view; 16. Head and thorax, dorsal view; 17. Frons, ventral view; 18. Forewing; 19. Pygofer, lateral view; 20. Pygofer, ventral view; 21. Hindwing; 22. Anal tube, dorsal view; 23. Gonostyli, lateral view; 24. Gonostyli, dorsal view; 25. Aedeagus, ventral view; 26. Aedeagus, lateral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10792154" httpUri="https://zenodo.org/record/10792154/files/figure.png" pageId="3" pageNumber="292">Figs 19, 23</figureCitation>
) angular at apex in lateral view.
</paragraph>
<paragraph id="8BD2176D3031FFBAFF55F8C2FC87F84E" blockId="3.[151,1437,1662,2013]" pageId="3" pageNumber="292">
<emphasis id="B919CB7F3031FFBAFF55F8C2FEB5F8E3" bold="true" box="[199,335,1878,1904]" pageId="3" pageNumber="292">Coloration.</emphasis>
General color grayish brown (
<figureCitation id="13560BE83031FFBAFD22F8C3FCCFF8E2" box="[688,821,1878,1905]" captionStart="FIGURES 1426" captionStartId="4.[151,264,1786,1810]" captionTargetBox="[158,1429,188,1756]" captionTargetId="figure-23@4.[151,1436,181,1763]" captionTargetPageId="4" captionText="FIGURES 1426. Metanigrus angularus sp. nov., male 14. Habitus, dorsal view; 15. Habitus, lateral view; 16. Head and thorax, dorsal view; 17. Frons, ventral view; 18. Forewing; 19. Pygofer, lateral view; 20. Pygofer, ventral view; 21. Hindwing; 22. Anal tube, dorsal view; 23. Gonostyli, lateral view; 24. Gonostyli, dorsal view; 25. Aedeagus, ventral view; 26. Aedeagus, lateral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10792154" httpUri="https://zenodo.org/record/10792154/files/figure.png" pageId="3" pageNumber="292">Figs 14, 15</figureCitation>
), with thin white wax powder. Head yellowish white. Upper part of frons grayish yellow, middle and basal parts yellowish brown. Clypeus yellowish brown. Eyes black, ocelli grayish white. Pronotum white, middle aera with 2 faint yellowish spots. Mesonotum brownish yellow. Forewings with black spots, veins brownish black. Hindwings hyaline.
</paragraph>
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<paragraph id="8BD2176D3036FFBDFF05F96EFE19F8ED" blockId="4.[151,1437,1786,1919]" pageId="4" pageNumber="293">
<emphasis id="B919CB7F3036FFBDFF05F96EFEADF881" bold="true" box="[151,343,1786,1810]" pageId="4" pageNumber="293">FIGURES 1426.</emphasis>
<taxonomicName id="4C6D6CEE3036FFBDFEF2F96FFDBFF880" authority="Lv &amp; Yang &amp; Chen, 2024" authorityName="Lv &amp; Yang &amp; Chen" authorityYear="2024" box="[352,581,1787,1811]" class="Insecta" family="Meenoplidae" genus="Metanigrus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="293" phylum="Arthropoda" rank="species" species="angularus" status="sp. nov.">
<emphasis id="B919CB7F3036FFBDFEF2F96FFDBFF880" box="[352,581,1787,1811]" italics="true" pageId="4" pageNumber="293">Metanigrus angularus</emphasis>
</taxonomicName>
<emphasis id="B919CB7F3036FFBDFDDDF96FFD58F880" bold="true" box="[591,674,1787,1811]" pageId="4" pageNumber="293">
<taxonomicNameLabel id="A22A76043036FFBDFDDDF96FFD58F880" box="[591,674,1787,1811]" pageId="4" pageNumber="293" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male 14. Habitus, dorsal view; 15. Habitus, lateral view; 16. Head and thorax, dorsal view; 17. Frons, ventral view; 18. Forewing; 19. Pygofer, lateral view; 20. Pygofer, ventral view; 21. Hindwing; 22. Anal tube, dorsal view; 23. Gonostyli, lateral view; 24. Gonostyli, dorsal view; 25. Aedeagus, ventral view; 26. Aedeagus, lateral view. Scale bars: 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BD2176D3036FFBDFF55F839FA66F854" blockId="4.[151,1436,1965,2028]" box="[199,1436,1965,1991]" pageId="4" pageNumber="293">
<emphasis id="B919CB7F3036FFBDFF55F839FE67F854" bold="true" box="[199,413,1965,1991]" pageId="4" pageNumber="293">Head and thorax.</emphasis>
Head (
<figureCitation id="13560BE83036FFBDFE61F839FD82F854" box="[499,632,1965,1991]" captionStart="FIGURES 1426" captionStartId="4.[151,264,1786,1810]" captionTargetBox="[158,1429,188,1756]" captionTargetId="figure-23@4.[151,1436,181,1763]" captionTargetPageId="4" captionText="FIGURES 1426. Metanigrus angularus sp. nov., male 14. Habitus, dorsal view; 15. Habitus, lateral view; 16. Head and thorax, dorsal view; 17. Frons, ventral view; 18. Forewing; 19. Pygofer, lateral view; 20. Pygofer, ventral view; 21. Hindwing; 22. Anal tube, dorsal view; 23. Gonostyli, lateral view; 24. Gonostyli, dorsal view; 25. Aedeagus, ventral view; 26. Aedeagus, lateral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10792154" httpUri="https://zenodo.org/record/10792154/files/figure.png" pageId="4" pageNumber="293">Figs 14, 16</figureCitation>
) significantly narrower than pronotum, without median carina. Vertex
</paragraph>
<paragraph id="8BD2176D3036FFBCFF05F845FC2FFE66" blockId="4.[151,1436,1965,2028]" lastBlockId="5.[151,1436,151,501]" lastPageId="5" lastPageNumber="294" pageId="4" pageNumber="293">
(
<figureCitation id="13560BE83036FFBDFF0DF845FEE4F878" box="[159,286,2001,2027]" captionStart="FIGURES 1426" captionStartId="4.[151,264,1786,1810]" captionTargetBox="[158,1429,188,1756]" captionTargetId="figure-23@4.[151,1436,181,1763]" captionTargetPageId="4" captionText="FIGURES 1426. Metanigrus angularus sp. nov., male 14. Habitus, dorsal view; 15. Habitus, lateral view; 16. Head and thorax, dorsal view; 17. Frons, ventral view; 18. Forewing; 19. Pygofer, lateral view; 20. Pygofer, ventral view; 21. Hindwing; 22. Anal tube, dorsal view; 23. Gonostyli, lateral view; 24. Gonostyli, dorsal view; 25. Aedeagus, ventral view; 26. Aedeagus, lateral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10792154" httpUri="https://zenodo.org/record/10792154/files/figure.png" pageId="4" pageNumber="293">Figs 14, 16</figureCitation>
) shorter in middle line than wide basally (1: 2.52), with two posterolateral areolets at base not meeting medially. Frons (
<figureCitation id="13560BE83037FFBCFEC4FF03FE52FF22" box="[342,424,151,177]" captionStart="FIGURES 1426" captionStartId="4.[151,264,1786,1810]" captionTargetBox="[158,1429,188,1756]" captionTargetId="figure-23@4.[151,1436,181,1763]" captionTargetPageId="4" captionText="FIGURES 1426. Metanigrus angularus sp. nov., male 14. Habitus, dorsal view; 15. Habitus, lateral view; 16. Head and thorax, dorsal view; 17. Frons, ventral view; 18. Forewing; 19. Pygofer, lateral view; 20. Pygofer, ventral view; 21. Hindwing; 22. Anal tube, dorsal view; 23. Gonostyli, lateral view; 24. Gonostyli, dorsal view; 25. Aedeagus, ventral view; 26. Aedeagus, lateral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10792154" httpUri="https://zenodo.org/record/10792154/files/figure.png" pageId="5" pageNumber="294">Fig. 17</figureCitation>
) subrectangular, without median carina, longer in middle line than wide at widest portion (about 1.65: 1), lateral carinae with a row of dense sensory pits along outer margin. Postclypeus (
<figureCitation id="13560BE83037FFBCFB4AFF2FFAD0FF46" box="[1240,1322,187,213]" captionStart="FIGURES 1426" captionStartId="4.[151,264,1786,1810]" captionTargetBox="[158,1429,188,1756]" captionTargetId="figure-23@4.[151,1436,181,1763]" captionTargetPageId="4" captionText="FIGURES 1426. Metanigrus angularus sp. nov., male 14. Habitus, dorsal view; 15. Habitus, lateral view; 16. Head and thorax, dorsal view; 17. Frons, ventral view; 18. Forewing; 19. Pygofer, lateral view; 20. Pygofer, ventral view; 21. Hindwing; 22. Anal tube, dorsal view; 23. Gonostyli, lateral view; 24. Gonostyli, dorsal view; 25. Aedeagus, ventral view; 26. Aedeagus, lateral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10792154" httpUri="https://zenodo.org/record/10792154/files/figure.png" pageId="5" pageNumber="294">Fig. 17</figureCitation>
) with distinct median and lateral carinae. Frontoclypeal suture nearly straight. Rostrum elongate, in repose well surpassing hind-coxae. Pronotum (
<figureCitation id="13560BE83037FFBCFE32FE97FE0BFE8E" box="[416,497,259,285]" captionStart="FIGURES 1426" captionStartId="4.[151,264,1786,1810]" captionTargetBox="[158,1429,188,1756]" captionTargetId="figure-23@4.[151,1436,181,1763]" captionTargetPageId="4" captionText="FIGURES 1426. Metanigrus angularus sp. nov., male 14. Habitus, dorsal view; 15. Habitus, lateral view; 16. Head and thorax, dorsal view; 17. Frons, ventral view; 18. Forewing; 19. Pygofer, lateral view; 20. Pygofer, ventral view; 21. Hindwing; 22. Anal tube, dorsal view; 23. Gonostyli, lateral view; 24. Gonostyli, dorsal view; 25. Aedeagus, ventral view; 26. Aedeagus, lateral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10792154" httpUri="https://zenodo.org/record/10792154/files/figure.png" pageId="5" pageNumber="294">Fig. 16</figureCitation>
) wider than maximum width of head (including eyes) (1.53: 1), with median carina and sinuate lateral carinae. Mesonotum (
<figureCitation id="13560BE83037FFBCFDF7FEB3FD42FED2" box="[613,696,295,321]" captionStart="FIGURES 1426" captionStartId="4.[151,264,1786,1810]" captionTargetBox="[158,1429,188,1756]" captionTargetId="figure-23@4.[151,1436,181,1763]" captionTargetPageId="4" captionText="FIGURES 1426. Metanigrus angularus sp. nov., male 14. Habitus, dorsal view; 15. Habitus, lateral view; 16. Head and thorax, dorsal view; 17. Frons, ventral view; 18. Forewing; 19. Pygofer, lateral view; 20. Pygofer, ventral view; 21. Hindwing; 22. Anal tube, dorsal view; 23. Gonostyli, lateral view; 24. Gonostyli, dorsal view; 25. Aedeagus, ventral view; 26. Aedeagus, lateral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10792154" httpUri="https://zenodo.org/record/10792154/files/figure.png" pageId="5" pageNumber="294">Fig. 16</figureCitation>
) about 5.22 times longer than pronotum in midline, with distinct median carina. Forewings (
<figureCitation id="13560BE83037FFBCFE5CFEDFFDDBFEF6" box="[462,545,331,357]" captionStart="FIGURES 1426" captionStartId="4.[151,264,1786,1810]" captionTargetBox="[158,1429,188,1756]" captionTargetId="figure-23@4.[151,1436,181,1763]" captionTargetPageId="4" captionText="FIGURES 1426. Metanigrus angularus sp. nov., male 14. Habitus, dorsal view; 15. Habitus, lateral view; 16. Head and thorax, dorsal view; 17. Frons, ventral view; 18. Forewing; 19. Pygofer, lateral view; 20. Pygofer, ventral view; 21. Hindwing; 22. Anal tube, dorsal view; 23. Gonostyli, lateral view; 24. Gonostyli, dorsal view; 25. Aedeagus, ventral view; 26. Aedeagus, lateral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10792154" httpUri="https://zenodo.org/record/10792154/files/figure.png" pageId="5" pageNumber="294">Fig. 18</figureCitation>
) slender, longer than maximal width (2.76: 1), almost as wide at postnodal as prenodal area, veins ScP + R + MP with several sensory pits, MP3 two branched; five postnodal closed cells with C1 smaller than C2, C5 placed next to C3, C4 distally displaced after nodal line, im, r-m and anterior two-thirds of margin with white areas. Hindwings (
<figureCitation id="13560BE83037FFBCFDADFE23FD6AFE42" box="[575,656,439,465]" captionStart="FIGURES 1426" captionStartId="4.[151,264,1786,1810]" captionTargetBox="[158,1429,188,1756]" captionTargetId="figure-23@4.[151,1436,181,1763]" captionTargetPageId="4" captionText="FIGURES 1426. Metanigrus angularus sp. nov., male 14. Habitus, dorsal view; 15. Habitus, lateral view; 16. Head and thorax, dorsal view; 17. Frons, ventral view; 18. Forewing; 19. Pygofer, lateral view; 20. Pygofer, ventral view; 21. Hindwing; 22. Anal tube, dorsal view; 23. Gonostyli, lateral view; 24. Gonostyli, dorsal view; 25. Aedeagus, ventral view; 26. Aedeagus, lateral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10792154" httpUri="https://zenodo.org/record/10792154/files/figure.png" pageId="5" pageNumber="294">Fig. 21</figureCitation>
) with lateral margin notched at CuA1, CuP, and A2, more strongly at A1; A2 not reaching posterior margin. Metatibiotarsal formula: (3+5)-6-5.
</paragraph>
<caption id="DF1247E53037FFBCFF05FAC5FCFFFAF9" ID-DOI="http://doi.org/10.5281/zenodo.10792156" ID-Zenodo-Dep="10792156" box="[151,773,1361,1386]" httpUri="https://zenodo.org/record/10792156/files/figure.png" pageId="5" pageNumber="294" startId="5.[151,250,1361,1385]" targetBox="[157,1429,531,1332]" targetPageId="5" targetType="figure">
<paragraph id="8BD2176D3037FFBCFF05FAC5FCFFFAF9" blockId="5.[151,773,1361,1386]" box="[151,773,1361,1386]" pageId="5" pageNumber="294">
<emphasis id="B919CB7F3037FFBCFF05FAC5FEDAFAFA" bold="true" box="[151,288,1361,1385]" pageId="5" pageNumber="294">FIGURE 27.</emphasis>
Geographic distribution of
<taxonomicName id="4C6D6CEE3037FFBCFDABFAC6FCFFFAF9" box="[569,773,1362,1386]" class="Insecta" family="Meenoplidae" genus="Metanigrus" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B919CB7F3037FFBCFDABFAC6FD54FAFA" box="[569,686,1362,1385]" italics="true" pageId="5" pageNumber="294">Metanigrus</emphasis>
species.
</taxonomicName>
</paragraph>
</caption>
<paragraph id="8BD2176D3037FFBCFF55FA0CFBC0F8AD" blockId="5.[151,1437,1431,2034]" pageId="5" pageNumber="294">
<emphasis id="B919CB7F3037FFBCFF55FA0CFE82FA22" bold="true" box="[199,376,1431,1458]" pageId="5" pageNumber="294">Male genitalia.</emphasis>
Pygofer (
<figureCitation id="13560BE83037FFBCFE76FA0CFD98FA21" box="[484,610,1432,1458]" captionStart="FIGURES 1426" captionStartId="4.[151,264,1786,1810]" captionTargetBox="[158,1429,188,1756]" captionTargetId="figure-23@4.[151,1436,181,1763]" captionTargetPageId="4" captionText="FIGURES 1426. Metanigrus angularus sp. nov., male 14. Habitus, dorsal view; 15. Habitus, lateral view; 16. Head and thorax, dorsal view; 17. Frons, ventral view; 18. Forewing; 19. Pygofer, lateral view; 20. Pygofer, ventral view; 21. Hindwing; 22. Anal tube, dorsal view; 23. Gonostyli, lateral view; 24. Gonostyli, dorsal view; 25. Aedeagus, ventral view; 26. Aedeagus, lateral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10792154" httpUri="https://zenodo.org/record/10792154/files/figure.png" pageId="5" pageNumber="294">Figs 19, 20</figureCitation>
) symmetrical, gradually narrowed towards apex, with sinuate anterior and posterior margins, pygopher with internal sulcus; in ventral view, lateral margins slightly curved, apex each with an obtuse process. Anal tube (
<figureCitation id="13560BE83037FFBCFE51FA74FDB8FA69" box="[451,578,1504,1530]" captionStart="FIGURES 1426" captionStartId="4.[151,264,1786,1810]" captionTargetBox="[158,1429,188,1756]" captionTargetId="figure-23@4.[151,1436,181,1763]" captionTargetPageId="4" captionText="FIGURES 1426. Metanigrus angularus sp. nov., male 14. Habitus, dorsal view; 15. Habitus, lateral view; 16. Head and thorax, dorsal view; 17. Frons, ventral view; 18. Forewing; 19. Pygofer, lateral view; 20. Pygofer, ventral view; 21. Hindwing; 22. Anal tube, dorsal view; 23. Gonostyli, lateral view; 24. Gonostyli, dorsal view; 25. Aedeagus, ventral view; 26. Aedeagus, lateral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10792154" httpUri="https://zenodo.org/record/10792154/files/figure.png" pageId="5" pageNumber="294">Figs 19, 22</figureCitation>
) in lateral view, hand-shaped, anal style slender, lateral lobes distally shorter than anal style, dorsolateral fingerlike process spindly, apex rounded, middle fingerlike process tapering towards end, acuminate apically, basolateral fingerlike process wide, apex curved dorsad; symmetrical in dorsal view, with two fingerlike processes on each side, middle of basal fingerlike process thinner than two ends, rounded at apex. Aedeagus (
<figureCitation id="13560BE83037FFBCFE85F9E4FE30F919" box="[279,458,1648,1675]" captionStart="FIGURES 1426" captionStartId="4.[151,264,1786,1810]" captionTargetBox="[158,1429,188,1756]" captionTargetId="figure-23@4.[151,1436,181,1763]" captionTargetPageId="4" captionText="FIGURES 1426. Metanigrus angularus sp. nov., male 14. Habitus, dorsal view; 15. Habitus, lateral view; 16. Head and thorax, dorsal view; 17. Frons, ventral view; 18. Forewing; 19. Pygofer, lateral view; 20. Pygofer, ventral view; 21. Hindwing; 22. Anal tube, dorsal view; 23. Gonostyli, lateral view; 24. Gonostyli, dorsal view; 25. Aedeagus, ventral view; 26. Aedeagus, lateral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10792154" httpUri="https://zenodo.org/record/10792154/files/figure.png" pageId="5" pageNumber="294">Figs 19, 25, 26</figureCitation>
) slender, long, apex curved ventrad in lateral view, with tubular periandrium, lateral lobes swelling on both sides, surface beset with scale-like productions, ventral lobe short, dorsal lobe fingerlike; in ventral view, wider at base. Gonostyli (
<figureCitation id="13560BE83037FFBCFDDFF92CFCDBF941" box="[589,801,1720,1746]" captionStart="FIGURES 1426" captionStartId="4.[151,264,1786,1810]" captionTargetBox="[158,1429,188,1756]" captionTargetId="figure-23@4.[151,1436,181,1763]" captionTargetPageId="4" captionText="FIGURES 1426. Metanigrus angularus sp. nov., male 14. Habitus, dorsal view; 15. Habitus, lateral view; 16. Head and thorax, dorsal view; 17. Frons, ventral view; 18. Forewing; 19. Pygofer, lateral view; 20. Pygofer, ventral view; 21. Hindwing; 22. Anal tube, dorsal view; 23. Gonostyli, lateral view; 24. Gonostyli, dorsal view; 25. Aedeagus, ventral view; 26. Aedeagus, lateral view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10792154" httpUri="https://zenodo.org/record/10792154/files/figure.png" pageId="5" pageNumber="294">Figs 19, 20, 23, 24</figureCitation>
) slender, divided into two parts, inner process extremely narrow, curved apically, tapering into a process; in lateral view, outer process convex, apex angular, inner margin with long setae; in ventral view, divergent, top of outer process fan-shaped, inner process sharpened downward; in dorsal view, lateral margins curved, outer process angular convex at apex medially.
</paragraph>
</subSubSection>
<subSubSection id="C37744E63037FFBCFF55F8DCFEB9F839" pageId="5" pageNumber="294" type="materials_examined">
<paragraph id="8BD2176D3037FFBCFF55F8DCFEB9F839" blockId="5.[151,1437,1431,2034]" pageId="5" pageNumber="294">
<emphasis id="B919CB7F3037FFBCFF55F8DCFE89F8F2" bold="true" box="[199,371,1863,1890]" pageId="5" pageNumber="294">Type material.</emphasis>
<materialsCitation id="3B051D303037FFBCFEEAF8DCFF11F815" collectingDate="2023-06-21" country="China" county="Mengla County" location="Gou" municipality="Menglun Town" pageId="5" pageNumber="294" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54D6A9CF3037FFBCFEEAF8DCFE1FF8F1" box="[376,485,1864,1890]" pageId="5" pageNumber="294" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D6BDCE43037FFBCFE7FF8DCFDFEF8F1" box="[493,516,1864,1890]" count="1" pageId="5" pageNumber="294" type="male"></specimenCount>
,
<collectingCountry id="F37A57FD3037FFBCFD9DF8DCFD96F8F1" box="[527,620,1864,1890]" name="China" pageId="5" pageNumber="294">CHINA</collectingCountry>
:
<collectingMunicipality id="6BB68D173037FFBCFDE6F8DCFCDAF8F1" box="[628,800,1864,1890]" pageId="5" pageNumber="294">Menglun Town</collectingMunicipality>
,
<collectingCounty id="62B36FE13037FFBCFCBBF8DCFC22F8F1" box="[809,984,1864,1890]" pageId="5" pageNumber="294">Mengla County</collectingCounty>
,
<collectingRegion id="49A9D98F3037FFBCFC72F8DCFB5BF8F1" box="[992,1185,1864,1890]" country="China" name="Yunnan" pageId="5" pageNumber="294">Yunnan Province</collectingRegion>
,
<date id="FFD331AD3037FFBCFB38F8DCFAC5F8F1" box="[1194,1343,1864,1890]" pageId="5" pageNumber="294" value="2023-06-21">
<collectingDate id="EF97C8453037FFBCFB38F8DCFAC5F8F1" box="[1194,1343,1864,1890]" pageId="5" pageNumber="294" value="2023-06-21">21 June 2023</collectingDate>
</date>
, Guang-Li
<location id="8EB241B63037FFBCFF25F8F8FF11F815" LSID="urn:lsid:plazi:treatment:03C4A67B3031FFBFFF05F987FD09FE1B:8EB241B63037FFBCFF25F8F8FF11F815" box="[183,235,1900,1926]" country="China" county="Mengla County" municipality="Menglun Town" name="Gou" pageId="5" pageNumber="294" stateProvince="Yunnan">Gou</location>
</materialsCitation>
.
<materialsCitation id="3B051D303037FFBCFF66F8F8FEC5F839" collectingDate="2023-06-21" country="China" county="Mengla County" location="Yan-Qiong Yang" municipality="Menglun Town" pageId="5" pageNumber="294" specimenCount="8" specimenCount-female="4" specimenCount-male="4" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="54D6A9CF3037FFBCFF66F8F8FE9FF815" box="[244,357,1900,1926]" pageId="5" pageNumber="294" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D6BDCE43037FFBCFEE2F8F8FE50F815" box="[368,426,1900,1926]" count="4" pageId="5" pageNumber="294" type="male">4♂♂</specimenCount>
,
<specimenCount id="9D6BDCE43037FFBCFE27F8F8FE14F815" box="[437,494,1900,1926]" count="4" pageId="5" pageNumber="294" type="female">4♀♀</specimenCount>
,
<collectingMunicipality id="6BB68D173037FFBCFE69F8F8FD52F815" box="[507,680,1900,1926]" pageId="5" pageNumber="294">Menglun Town</collectingMunicipality>
,
<collectingCounty id="62B36FE13037FFBCFD21F8F8FC99F815" box="[691,867,1900,1926]" pageId="5" pageNumber="294">Mengla County</collectingCounty>
,
<collectingRegion id="49A9D98F3037FFBCFCFEF8F8FBD4F815" box="[876,1070,1900,1926]" country="China" name="Yunnan" pageId="5" pageNumber="294">Yunnan Province</collectingRegion>
,
<date id="FFD331AD3037FFBCFBA8F8F8FB2BF815" box="[1082,1233,1900,1926]" pageId="5" pageNumber="294" value="2023-06-21">
<collectingDate id="EF97C8453037FFBCFBA8F8F8FB2BF815" box="[1082,1233,1900,1926]" pageId="5" pageNumber="294" value="2023-06-21">21 June 2023</collectingDate>
</date>
,
<location id="8EB241B63037FFBCFB49F8F8FA62F815" LSID="urn:lsid:plazi:treatment:03C4A67B3031FFBFFF05F987FD09FE1B:8EB241B63037FFBCFB49F8F8FA62F815" box="[1243,1432,1900,1926]" country="China" county="Mengla County" municipality="Menglun Town" name="Yan-Qiong Yang" pageId="5" pageNumber="294" stateProvince="Yunnan">Yan-Qiong Yang</location>
, Guang-Li
<location id="8EB241B63037FFBCFE99F804FEC5F839" LSID="urn:lsid:plazi:treatment:03C4A67B3031FFBFFF05F987FD09FE1B:8EB241B63037FFBCFE99F804FEC5F839" box="[267,319,1936,1962]" country="China" county="Mengla County" municipality="Menglun Town" name="Gou" pageId="5" pageNumber="294" stateProvince="Yunnan">Gou</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37744E63037FFBCFF55F820FD87F861" pageId="5" pageNumber="294" type="etymology">
<paragraph id="8BD2176D3037FFBCFF55F820FD87F861" blockId="5.[151,1437,1431,2034]" pageId="5" pageNumber="294">
<emphasis id="B919CB7F3037FFBCFF55F820FEB4F85D" bold="true" box="[199,334,1972,1998]" pageId="5" pageNumber="294">Etymology.</emphasis>
The specific name is derived from the Latin adjective
<taxonomicName id="4C6D6CEE3037FFBCFC2EF820FBC0F85D" authorityName="Lv &amp; Yang &amp; Chen" authorityYear="2024" box="[956,1082,1972,1998]" class="Insecta" family="Meenoplidae" genus="Metanigrus" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="angularus">
<emphasis id="B919CB7F3037FFBCFC5AF820FBC0F85D" box="[968,1082,1972,1998]" italics="true" pageId="5" pageNumber="294">angularus</emphasis>
</taxonomicName>
”, referring to the outer process of gonostyli angular at apex in lateral view.
</paragraph>
</subSubSection>
<subSubSection id="C37744E63034FFBFFF55FF02FE11FEF7" pageId="6" pageNumber="295" type="discussion">
<paragraph id="8BD2176D3034FFBFFF55FF02FE11FEF7" blockId="6.[151,1436,150,392]" pageId="6" pageNumber="295">
<emphasis id="B919CB7F3034FFBFFF55FF02FEC1FF23" bold="true" box="[199,315,150,176]" pageId="6" pageNumber="295">Remarks.</emphasis>
This species is similar to
<taxonomicName id="4C6D6CEE3034FFBFFDC8FF03FC5EFF23" authority="Lv &amp; Chen, 2021" authorityName="Lv &amp; Chen" authorityYear="2021" box="[602,932,150,177]" class="Insecta" family="Meenoplidae" genus="Metanigrus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="295" phylum="Arthropoda" rank="species" species="guttatus">
<emphasis id="B919CB7F3034FFBFFDC8FF03FD23FF23" box="[602,729,151,176]" italics="true" pageId="6" pageNumber="295">M. guttatus</emphasis>
Lv &amp; Chen, 2021
</taxonomicName>
, but differs from the latter in: (1) dorsolateral and middle parts of anal tube each with a fingerlike process in lateral view (dorsolateral part of anal tube with a fingerlike process, middle part without fingerlike process in lateral view in
<taxonomicName id="4C6D6CEE3034FFBFFC79FF4BFB91FF6B" authorityName="Lv &amp; Chen" authorityYear="2021" box="[1003,1131,223,248]" class="Insecta" family="Meenoplidae" genus="Metanigrus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="295" phylum="Arthropoda" rank="species" species="guttatus">
<emphasis id="B919CB7F3034FFBFFC79FF4BFB91FF6B" box="[1003,1131,223,248]" italics="true" pageId="6" pageNumber="295">M. guttatus</emphasis>
</taxonomicName>
); (2) aedeagus slender and long, apex curved ventrad in lateral view (aedeagus stout, apex not curved ventrad in lateral view in
<taxonomicName id="4C6D6CEE3034FFBFFA99FE97FA71FE8F" authorityName="Lv &amp; Chen" authorityYear="2021" box="[1291,1419,259,284]" class="Insecta" family="Meenoplidae" genus="Metanigrus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="295" phylum="Arthropoda" rank="species" species="guttatus">
<emphasis id="B919CB7F3034FFBFFA99FE97FA71FE8F" box="[1291,1419,259,284]" italics="true" pageId="6" pageNumber="295">M. guttatus</emphasis>
</taxonomicName>
); (3) outer process of gonostyli angular at apex in lateral view (outer process of gonostyli nearly fan-shaped at apex in lateral view in
<taxonomicName id="4C6D6CEE3034FFBFFEC9FEDFFE20FEF7" authorityName="Lv &amp; Chen" authorityYear="2021" box="[347,474,331,356]" class="Insecta" family="Meenoplidae" genus="Metanigrus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="295" phylum="Arthropoda" rank="species" species="guttatus">
<emphasis id="B919CB7F3034FFBFFEC9FEDFFE20FEF7" box="[347,474,331,356]" italics="true" pageId="6" pageNumber="295">M. guttatus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C37744E63034FFBFFF55FEFAFD09FE1B" box="[199,755,366,392]" pageId="6" pageNumber="295" type="distribution">
<paragraph id="8BD2176D3034FFBFFF55FEFAFD09FE1B" blockId="6.[151,1436,150,392]" box="[199,755,366,392]" pageId="6" pageNumber="295">
<emphasis id="B919CB7F3034FFBFFF55FEFAFE9BFE1B" bold="true" box="[199,353,366,392]" pageId="6" pageNumber="295">Distribution.</emphasis>
<collectingCountry id="F37A57FD3034FFBFFEF5FEFAFE50FE1B" box="[359,426,366,392]" name="China" pageId="6" pageNumber="295">China</collectingCountry>
(
<collectingRegion id="49A9D98F3034FFBFFE2BFEFAFD81FE1B" box="[441,635,366,392]" country="China" name="Yunnan" pageId="6" pageNumber="295">Yunnan province</collectingRegion>
) (
<figureCitation id="13560BE83034FFBFFD00FEFAFD1CFE1B" box="[658,742,366,392]" captionStart="FIGURE 27" captionStartId="5.[151,250,1361,1385]" captionTargetBox="[157,1429,531,1332]" captionTargetId="figure-193@5.[151,1435,525,1338]" captionTargetPageId="5" captionText="FIGURE 27. Geographic distribution of Metanigrus species." figureDoi="http://doi.org/10.5281/zenodo.10792156" httpUri="https://zenodo.org/record/10792156/files/figure.png" pageId="6" pageNumber="295">Fig. 27</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>