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<document id="2BC8799C8AE381DC80D00A41D0B6FA61" ID-CLB-Dataset="46809" ID-DOI="10.11646/zootaxa.4078.1.16" ID-GBIF-Dataset="9382c912-1e52-41c8-8499-1c515191c8ae" ID-ISSN="1175-5326" ID-Zenodo-Dep="261493" ID-ZooBank="0102DC85-29D8-4B7A-BD24-984C115189AA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1460361037286" checkinUser="plazi" docAuthor="Barão, Kim Ribeiro, Garbelotto, Thereza De Almeida, Campos, Luiz Alexandre &amp; Grazia, Jocelia" docDate="2016" docId="03BF87AB1B27FFC49AC49AAD34ABFDF1" docLanguage="en" docName="zootaxa.4078.1.16.pdf" docOrigin="Zootaxa 4078 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Braunus brettlizorum Thomas 1997, comb. nov." docType="treatment" docVersion="11" lastPageNumber="181" masterDocId="FF86FFD31B2BFFC99A539C4A366FFFE9" masterDocTitle="Unusual looking pentatomids: reassessing the taxonomy of Braunus Distant and Lojus McDonald (Hemiptera: Heteroptera: Pentatomidae)" masterLastPageNumber="186" masterPageNumber="168" pageNumber="180" updateTime="1698294510363" updateUser="ExternalLinkService">
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<mods:title id="4F98458659CC967C9E2AA0D0626449B8">Unusual looking pentatomids: reassessing the taxonomy of Braunus Distant and Lojus McDonald (Hemiptera: Heteroptera: Pentatomidae)</mods:title>
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<mods:namePart id="F1924FB5E2AAF0B125D12C9E878764CE">Barão, Kim Ribeiro</mods:namePart>
<mods:affiliation id="F80316C467FB94D614BEBA638C1C1AC3">Programa de Pós-Graduação em Biologia Animal &amp; Departamento de Zoologia, Universidade Federal do Rio Grande do Sul, Av. Bento Gonçalves 9500, Porto Alegre, Rio Grande do Sul, Brazil 91501 - 970</mods:affiliation>
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<mods:namePart id="86493864152F8761CC40E4D8EB058A26">Garbelotto, Thereza De Almeida</mods:namePart>
<mods:affiliation id="87A88A6A0B83AE7F54A28C742B255157">Programa de Pós-Graduação em Biologia Animal &amp; Departamento de Zoologia, Universidade Federal do Rio Grande do Sul, Av. Bento Gonçalves 9500, Porto Alegre, Rio Grande do Sul, Brazil 91501 - 970</mods:affiliation>
<mods:nameIdentifier id="91BAC81FE2AAE5F3D776ABD6D445CE11" type="email">therezagarbelotto@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="3D56B7CE83BC3300EDB1B6F3E325485B">Campos, Luiz Alexandre</mods:namePart>
<mods:affiliation id="269817FE6DA250ECA497E4599955B3B9">Programa de Pós-Graduação em Biologia Animal &amp; Departamento de Zoologia, Universidade Federal do Rio Grande do Sul, Av. Bento Gonçalves 9500, Porto Alegre, Rio Grande do Sul, Brazil 91501 - 970</mods:affiliation>
<mods:nameIdentifier id="55D8D96F975CBCE6E8DE6B472D3F768B" type="email">luiz.campos@ufrgs.br</mods:nameIdentifier>
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<mods:namePart id="38617BC5536C3CEA32D614F7485B0276">Grazia, Jocelia</mods:namePart>
<mods:affiliation id="22E54F7F91255148528454FE634B37FF">Programa de Pós-Graduação em Biologia Animal &amp; Departamento de Zoologia, Universidade Federal do Rio Grande do Sul, Av. Bento Gonçalves 9500, Porto Alegre, Rio Grande do Sul, Brazil 91501 - 970</mods:affiliation>
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<mods:date id="2F4A6C997D75FD704DE87CCBD86C4723">2016</mods:date>
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<mods:number id="EF90D12987789CA35F546F8FFD92CF47">4078</mods:number>
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<treatment id="03BF87AB1B27FFC49AC49AAD34ABFDF1" ID-DOI="http://doi.org/10.5281/zenodo.6081316" ID-GBIF-Taxon="119409739" ID-Zenodo-Dep="6081316" LSID="urn:lsid:plazi:treatment:03BF87AB1B27FFC49AC49AAD34ABFDF1" httpUri="http://treatment.plazi.org/id/03BF87AB1B27FFC49AC49AAD34ABFDF1" lastPageId="13" lastPageNumber="181" pageId="12" pageNumber="180">
<subSubSection id="C30C65361B27FFC59AC49AAD349CF8E8" box="[151,755,1767,1793]" pageId="12" pageNumber="180" type="nomenclature">
<paragraph id="8BA936BD1B27FFC59AC49AAD349CF8E8" blockId="12.[151,755,1767,1827]" box="[151,755,1767,1793]" pageId="12" pageNumber="180">
<heading id="D0E181D11B27FFC59AC49AAD349CF8E8" bold="true" box="[151,755,1767,1793]" fontSize="11" level="1" pageId="12" pageNumber="180" reason="1">
<emphasis id="B962EAAF1B27FFC59AC49AAD349CF8E8" bold="true" box="[151,755,1767,1793]" pageId="12" pageNumber="180">
<taxonomicName id="4C164D3E1B27FFC59AC49AAD340AF8E8" authority="Thomas, 1997" authorityName="Thomas" authorityYear="1997" box="[151,613,1767,1793]" class="Insecta" family="Pentatomidae" genus="Braunus" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="180" phylum="Arthropoda" rank="species" species="brettlizorum" status="comb. nov.">
<emphasis id="B962EAAF1B27FFC59AC49AAD37F3F8E8" bold="true" box="[151,412,1767,1793]" italics="true" pageId="12" pageNumber="180">Braunus brettlizorum</emphasis>
(
<bibRefCitation id="EF874B4C1B27FFC59BFF9AAD3432F8E8" author="Thomas" box="[428,605,1767,1793]" pageId="12" pageNumber="180" refString="Thomas, D. B. (1997) The anocellate, flightless genus Lojus McDonald (Heteroptera: Pentatomidae). Annals of the Entomological Society of America, 90, 569 - 574. http: // dx. doi. org / 10.1093 / aesa / 90.5.569" type="journal article" year="1997">Thomas, 1997</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A25157D41B27FFC5983E9AAD349CF8E8" box="[621,755,1767,1793]" pageId="12" pageNumber="180">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C30C65361B27FFC59AC49B403725F8CB" box="[151,330,1802,1827]" pageId="12" pageNumber="180" type="description">
<paragraph id="8BA936BD1B27FFC59AC49B403725F8CB" blockId="12.[151,755,1767,1827]" box="[151,330,1802,1827]" pageId="12" pageNumber="180">
(
<figureCitation id="132D2A381B27FFC59ACD9B403776F8CB" box="[158,281,1802,1827]" captionStart="FIGURES 1 11" captionStartId="3.[151,264,1812,1835]" captionTargetBox="[151,1436,949,1788]" captionTargetId="figure@3.[151,1436,945,1789]" captionTargetPageId="3" captionText="FIGURES 1 11. Schematic drawings of pronotum of Braunus Distant, 1899 in dorsal view. 1 3. B. sciocorinus (Walker, 1867); 4. B. machadoi Barão &amp; Garbelotto sp. nov.; 5. B. gibbus Garbelotto &amp; Barão sp. nov.; 6. B. prionotus Barão &amp; Garbelotto sp. nov.; 7. B. asperus (McDonald, 1982) comb. nov.; 8. B. ateuchus (Brailovski &amp; Rolston, 1986) comb. nov.; 9. B. bordoni (Thomas, 1997) comb. nov.; 10. B. brettlizorum (Thomas, 1997) comb. nov.; 11. B. ocellatus (Thomas, 2001) comb. nov. Scale bars = 1 mm." httpUri="https://zenodo.org/record/261495/files/figure.png" pageId="12" pageNumber="180">Figures 10</figureCitation>
,
<figureCitation id="132D2A381B27FFC59B769B40372DF8CB" box="[293,322,1802,1826]" captionStart="FIGURES 78 82" captionStartId="13.[151,264,1110,1133]" captionTargetBox="[151,1436,591,1086]" captionTargetId="figure@13.[151,1436,591,1086]" captionTargetPageId="13" captionText="FIGURES 78 82. Distribution of species of Braunus Distant, 1899 and localities of some collection events. 78. map of localities; 79. a collection locality of B. sciocorinus (Walker, 1867) and B. ocellatus (Thomas, 2001) comb. nov. (ECUADOR, Napo, Yanayacu Biological Station; photo by D. Forero); 80. type locality of B. machadoi Barão &amp; Garbelotto sp. nov. (PERU, Cuzco, Wayqecha Research Center; photo by C. Weirauch); 81 82. a collection locality of B. ocellatus (Thomas, 2001) comb. nov. (ECUADOR, Napo, 1.3 km SW of Baeza; photos by J. Skuhrovec)." httpUri="https://zenodo.org/record/261505/files/figure.png" pageId="12" pageNumber="180">78</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30C65361B27FFC59AC49B1934A2F881" box="[151,717,1874,1896]" pageId="12" pageNumber="180" type="reference_group">
<paragraph id="8BA936BD1B27FFC59AC49B1934A2F881" blockId="12.[151,717,1874,1896]" box="[151,717,1874,1896]" pageId="12" pageNumber="180">
<treatmentCitationGroup id="AB0611931B27FFC59AC49B1934A2F881" box="[151,717,1874,1896]" pageId="12" pageNumber="180">
<taxonomicName id="4C164D3E1B27FFC59AC49B193472F881" authority="Thomas, 1997: 570" authorityName="Thomas" authorityPageNumber="570" authorityYear="1997" box="[151,541,1874,1896]" class="Insecta" family="Pentatomidae" genus="Lojus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="180" phylum="Arthropoda" rank="species" species="brettlizorum">
<emphasis id="B962EAAF1B27FFC59AC49B19373EF881" box="[151,337,1874,1896]" italics="true" pageId="12" pageNumber="180">Lojus brettlizorum</emphasis>
<treatmentCitation id="0AB710AC1B27FFC59B0B9B183472F881" author="Thomas" box="[344,541,1874,1896]" page="570" pageId="12" pageNumber="180" year="1997">
<bibRefCitation id="EF874B4C1B27FFC59B0B9B183783F881" author="Thomas" box="[344,492,1874,1896]" pageId="12" pageNumber="180" refString="Thomas, D. B. (1997) The anocellate, flightless genus Lojus McDonald (Heteroptera: Pentatomidae). Annals of the Entomological Society of America, 90, 569 - 574. http: // dx. doi. org / 10.1093 / aesa / 90.5.569" type="journal article" year="1997">Thomas, 1997</bibRefCitation>
: 570
</treatmentCitation>
</taxonomicName>
572, figs 15, 8.
</treatmentCitationGroup>
</paragraph>
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<subSubSection id="C30C65361B27FFC59AC49BDE3327F838" pageId="12" pageNumber="180" type="materials_examined">
<paragraph id="8BA936BD1B27FFC59AC49BDE3327F838" blockId="12.[151,1436,1940,2037]" pageId="12" pageNumber="180">
<materialsCitation id="3B7E3CE01B27FFC59AC49BDE34ABF838" ID-GBIF-Occurrence="2636403313" collectingDate="1995-05-20" collectingDateMax="1995-05-21" collectingDateMin="1995-05-20" collectionCode="UNSM" collectorName="B. Ratcliffe &amp; M. Jameson" country="Costa Rica" elevation="1280" latitude="9.7" location="Ref. Nac. Tapanti" longLatPrecision="1299" longitude="-83.76667" pageId="12" pageNumber="180" specimenCount="1" specimenCount-male="1" stateProvince="Cartago" typeStatus="holotype">
<emphasis id="B962EAAF1B27FFC59AC49BDE37EBF844" bold="true" box="[151,388,1940,1965]" pageId="12" pageNumber="180">Material examined.</emphasis>
<typeStatus id="54AD881F1B27FFC59BDD9BDE3799F844" box="[398,502,1940,1965]" pageId="12" pageNumber="180" type="holotype">Holotype</typeStatus>
<specimenCount id="9D10FD341B27FFC598539BDF3478F845" box="[512,535,1941,1964]" pageId="12" pageNumber="180" type="male"></specimenCount>
,
<collectingCountry id="F301762D1B27FFC598759BDF34BFF844" box="[550,720,1941,1965]" name="Costa Rica" pageId="12" pageNumber="180" value="Costa Rica">COSTA RICA</collectingCountry>
,
<collectingRegion id="49D2F85F1B27FFC5988C9BDF3555F844" box="[735,826,1941,1965]" pageId="12" pageNumber="180">Cartago</collectingRegion>
,
<location id="8EC960661B27FFC5991A9BDE3277F844" LSID="urn:lsid:plazi:treatment:03BF87AB1B27FFC49AC49AAD34ABFDF1:8EC960661B27FFC5991A9BDE3277F844" box="[841,1048,1940,1965]" country="Costa Rica" latitude="9.7" longLatPrecision="1299" longitude="-83.76667" name="Ref. Nac. Tapanti" pageId="12" pageNumber="180" stateProvince="Cartago">Ref. Nac. Tapanti</location>
,
<geoCoordinate id="EE22507A1B27FFC59E759BDF3216F844" box="[1062,1145,1941,1965]" degrees="9" direction="north" minutes="42" orientation="latitude" pageId="12" pageNumber="180" precision="925" value="9.7">9°42N</geoCoordinate>
<geoCoordinate id="EE22507A1B27FFC59ED09BDE3285F844" box="[1155,1258,1940,1965]" degrees="83" direction="west" minutes="46" orientation="longitude" pageId="12" pageNumber="180" precision="925" value="-83.76667">83°46W</geoCoordinate>
,
<date id="FFA8107D1B27FFC59EAA9BDE33F7F844" box="[1273,1432,1940,1965]" pageId="12" pageNumber="180" value="1995-05-20" valueMax="1995-05-21" valueMin="1995-05-20">
<collectingDate id="EFECE9951B27FFC59EAA9BDE33F7F844" box="[1273,1432,1940,1965]" pageId="12" pageNumber="180" value="1995-05-20" valueMax="1995-05-21" valueMin="1995-05-20">2021.V.1995</collectingDate>
</date>
,
<quantity id="4CEE9B581B27FFC59AC49BF33685F838" box="[151,234,1976,2001]" metricMagnitude="3" metricUnit="m" metricValue="1.28016" pageId="12" pageNumber="180" unit="ft" value="4200.0">
<elevation id="003BD18E1B27FFC59AC49BF33685F838" box="[151,234,1976,2001]" metricMagnitude="3" metricUnit="m" metricValue="1.28016" pageId="12" pageNumber="180" unit="ft" value="4200.0">4200 ft</elevation>
</quantity>
,
<collectorName id="26E3536B1B27FFC59AA69BF3344CF839" box="[245,547,1976,2001]" pageId="12" pageNumber="180">B. Ratcliffe &amp; M. Jameson</collectorName>
leg. (
<collectionCode id="ED07AE781B27FFC598329BF334D7F838" LSID="urn:lsid:biocol.org:col:34868" box="[609,696,1977,2001]" collectionName="USA, Nebraska, Lincoln, University of Nebraska State Museum" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34868" name="University of Nebraska State Museum" pageId="12" pageNumber="180">UNSM</collectionCode>
).
</materialsCitation>
<materialsCitation id="3B7E3CE01B27FFC598989BF3332BF838" ID-GBIF-Occurrence="2636403301" box="[715,1348,1976,2001]" collectingDate="1995-05-20" collectingDateMax="1995-05-21" collectingDateMin="1995-05-20" collectionCode="DBTC" collectorName="B. Ratcliffe &amp; M. Jameson" country="Costa Rica" elevation="1280" latitude="9.7" location="Ref. Nac. Tapanti" longLatPrecision="1299" longitude="-83.76667" pageId="12" pageNumber="180" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Cartago" typeStatus="paratype">
<typeStatus id="54AD881F1B27FFC598989BF33557F838" box="[715,824,1977,2001]" pageId="12" pageNumber="180" type="paratype">Paratypes</typeStatus>
<specimenCount id="9D10FD341B27FFC5996C9BF33504F839" box="[831,875,1977,2001]" pageId="12" pageNumber="180" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D10FD341B27FFC599249BF335F4F838" box="[887,923,1977,2001]" pageId="12" pageNumber="180" type="female">1♀</specimenCount>
: same label data as holotype (
<collectionCode id="ED07AE781B27FFC59EB89BF33353F838" box="[1259,1340,1977,2001]" collectionName="Donald B. Thomas" pageId="12" pageNumber="180">DBTC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30C65361B27FFC49A949B963567FE1C" lastPageId="13" lastPageNumber="181" pageId="12" pageNumber="180" type="diagnosis">
<paragraph id="8BA936BD1B27FFC49A949B963567FE1C" blockId="12.[151,1436,1940,2037]" lastBlockId="13.[151,1437,151,536]" lastPageId="13" lastPageNumber="181" pageId="12" pageNumber="180">
<emphasis id="B962EAAF1B27FFC59A949B96372EF81C" bold="true" box="[199,321,2012,2037]" pageId="12" pageNumber="180">Diagnosis.</emphasis>
Body color dark brown. Anterior angles of pronotum each with small, inconspicuous, obtuse projection; each anterolateral margin serrated; humeral angles projecting laterad, each bilobed apically, lobes acute; posterior margin of pronotum concave (
<figureCitation id="132D2A381B26FFC498029CF634CBFF3D" box="[593,676,188,213]" captionStart="FIGURES 1 11" captionStartId="3.[151,264,1812,1835]" captionTargetBox="[151,1436,949,1788]" captionTargetId="figure@3.[151,1436,945,1789]" captionTargetPageId="3" captionText="FIGURES 1 11. Schematic drawings of pronotum of Braunus Distant, 1899 in dorsal view. 1 3. B. sciocorinus (Walker, 1867); 4. B. machadoi Barão &amp; Garbelotto sp. nov.; 5. B. gibbus Garbelotto &amp; Barão sp. nov.; 6. B. prionotus Barão &amp; Garbelotto sp. nov.; 7. B. asperus (McDonald, 1982) comb. nov.; 8. B. ateuchus (Brailovski &amp; Rolston, 1986) comb. nov.; 9. B. bordoni (Thomas, 1997) comb. nov.; 10. B. brettlizorum (Thomas, 1997) comb. nov.; 11. B. ocellatus (Thomas, 2001) comb. nov. Scale bars = 1 mm." httpUri="https://zenodo.org/record/261495/files/figure.png" pageId="13" pageNumber="181">Fig. 10</figureCitation>
). Apex of scutellum reaching anterior margin of abdominal segment VII on males and anterior margin of abdominal segment VI on females. Apex of hemelytral radial vein calloused; corium reaching posterior margin of abdominal segment VI; membranal veins linear, not reticulated; membrane fully developed on males, surpassing abdominal apex, and reduced to a thin stripe on females. Male genitalia: inferior layer of ventral rim of pygophore emarginated, with two small projections medially. Phallus: limit of sclerotized and membranous area of phallotheca straight. Each lateroventral conjunctival process hatchet-like, strongly sclerotized. Vesica cup-like ventrally. Process of gonopore bilobed apically. Female genitalia: sutural margins of each gonocoxite 8 parallel basally, divergent and straight apically; posterior margins of each gonocoxite 8 and laterotergite 8 straight; laterotergites 9 digitiform.
</paragraph>
</subSubSection>
<subSubSection id="C30C65361B26FFC49A949DB534ABFDF1" box="[199,708,511,536]" pageId="13" pageNumber="181" type="distribution">
<paragraph id="8BA936BD1B26FFC49A949DB534ABFDF1" blockId="13.[151,1437,151,536]" box="[199,708,511,536]" pageId="13" pageNumber="181">
<emphasis id="B962EAAF1B26FFC49A949DB5370FFDF1" bold="true" box="[199,352,511,536]" pageId="13" pageNumber="181">Distribution.</emphasis>
Costa Rica (Cartago) (
<figureCitation id="132D2A381B26FFC498309DB534D8FDF1" box="[611,695,511,536]" captionStart="FIGURES 78 82" captionStartId="13.[151,264,1110,1133]" captionTargetBox="[151,1436,591,1086]" captionTargetId="figure@13.[151,1436,591,1086]" captionTargetPageId="13" captionText="FIGURES 78 82. Distribution of species of Braunus Distant, 1899 and localities of some collection events. 78. map of localities; 79. a collection locality of B. sciocorinus (Walker, 1867) and B. ocellatus (Thomas, 2001) comb. nov. (ECUADOR, Napo, Yanayacu Biological Station; photo by D. Forero); 80. type locality of B. machadoi Barão &amp; Garbelotto sp. nov. (PERU, Cuzco, Wayqecha Research Center; photo by C. Weirauch); 81 82. a collection locality of B. ocellatus (Thomas, 2001) comb. nov. (ECUADOR, Napo, 1.3 km SW of Baeza; photos by J. Skuhrovec)." httpUri="https://zenodo.org/record/261505/files/figure.png" pageId="13" pageNumber="181">Fig. 78</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>