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<document id="062ED0775D8BF2D77BC594D6679B2C0E" ID-CLB-Dataset="46809" ID-DOI="10.11646/zootaxa.4078.1.16" ID-GBIF-Dataset="9382c912-1e52-41c8-8499-1c515191c8ae" ID-ISSN="1175-5326" ID-Zenodo-Dep="261493" ID-ZooBank="0102DC85-29D8-4B7A-BD24-984C115189AA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1460361037286" checkinUser="plazi" docAuthor="Barão, Kim Ribeiro, Garbelotto, Thereza De Almeida, Campos, Luiz Alexandre &amp; Grazia, Jocelia" docDate="2016" docId="03BF87AB1B21FFC39AC49FC03464FA34" docLanguage="en" docName="zootaxa.4078.1.16.pdf" docOrigin="Zootaxa 4078 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Braunus asperus" docType="treatment" docVersion="11" lastPageNumber="178" masterDocId="FF86FFD31B2BFFC99A539C4A366FFFE9" masterDocTitle="Unusual looking pentatomids: reassessing the taxonomy of Braunus Distant and Lojus McDonald (Hemiptera: Heteroptera: Pentatomidae)" masterLastPageNumber="186" masterPageNumber="168" pageNumber="178" updateTime="1698294510363" updateUser="ExternalLinkService">
<mods:mods id="4E959FA1FAC1B915E98445DF92899B28" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="C442BDB18F0A28E879ACD85B146ABCBC">Unusual looking pentatomids: reassessing the taxonomy of Braunus Distant and Lojus McDonald (Hemiptera: Heteroptera: Pentatomidae)</mods:title>
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<mods:namePart id="1907343BA42CA50E5503F1F217741991">Barão, Kim Ribeiro</mods:namePart>
<mods:affiliation id="B74DA07B42CC9D6A0C508EB041581C04">Programa de Pós-Graduação em Biologia Animal &amp; Departamento de Zoologia, Universidade Federal do Rio Grande do Sul, Av. Bento Gonçalves 9500, Porto Alegre, Rio Grande do Sul, Brazil 91501 - 970</mods:affiliation>
<mods:nameIdentifier id="BC18D0F99450CB882324FEBE44A578E9" type="email">kbarao@yahoo.com.br</mods:nameIdentifier>
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<mods:namePart id="0471719F37809870C8B1FE4E4685BBFF">Garbelotto, Thereza De Almeida</mods:namePart>
<mods:affiliation id="CA6B7C6F5E00BFBF7962D67E4C865C46">Programa de Pós-Graduação em Biologia Animal &amp; Departamento de Zoologia, Universidade Federal do Rio Grande do Sul, Av. Bento Gonçalves 9500, Porto Alegre, Rio Grande do Sul, Brazil 91501 - 970</mods:affiliation>
<mods:nameIdentifier id="5AEBB839B46D90D97B47B691FA1587CD" type="email">therezagarbelotto@hotmail.com</mods:nameIdentifier>
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<mods:name id="D0B99A39EECD1239F9AF0CE4F282E0AD" type="personal">
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<mods:namePart id="5DD6A5C3170C91D2562D0945E0EE3E37">Campos, Luiz Alexandre</mods:namePart>
<mods:affiliation id="127EB967425FA1F35928654F12DD48E4">Programa de Pós-Graduação em Biologia Animal &amp; Departamento de Zoologia, Universidade Federal do Rio Grande do Sul, Av. Bento Gonçalves 9500, Porto Alegre, Rio Grande do Sul, Brazil 91501 - 970</mods:affiliation>
<mods:nameIdentifier id="6016BE9232E88F6CF8CCF3837B3F64A9" type="email">luiz.campos@ufrgs.br</mods:nameIdentifier>
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<mods:namePart id="FDDF9FE53141115F8949E7D147EA8C84">Grazia, Jocelia</mods:namePart>
<mods:affiliation id="D4AACF534EB4743CF047FED44C90836D">Programa de Pós-Graduação em Biologia Animal &amp; Departamento de Zoologia, Universidade Federal do Rio Grande do Sul, Av. Bento Gonçalves 9500, Porto Alegre, Rio Grande do Sul, Brazil 91501 - 970</mods:affiliation>
<mods:nameIdentifier id="02EEC38A525F5E2452C2DD5478BAABF0" type="email">jocelia@ufrgs.br</mods:nameIdentifier>
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<mods:title id="D91D5A710D31B67E7D7F0B19AC3861F5">Zootaxa</mods:title>
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<mods:date id="E84BE7843EA0354D930354A4D80887B9">2016</mods:date>
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<mods:number id="65E1FA621CE034E865FAD316A4A1C428">2016-02-09</mods:number>
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<mods:number id="D5099D8E0CDE40C64F4D44D17CAEBA20">4078</mods:number>
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<mods:identifier id="257B135552297CADEFE1498597755671" type="DOI">10.11646/zootaxa.4078.1.16</mods:identifier>
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<treatment id="03BF87AB1B21FFC39AC49FC03464FA34" ID-DOI="http://doi.org/10.5281/zenodo.6081310" ID-GBIF-Taxon="119409743" ID-Zenodo-Dep="6081310" LSID="urn:lsid:plazi:treatment:03BF87AB1B21FFC39AC49FC03464FA34" httpUri="http://treatment.plazi.org/id/03BF87AB1B21FFC39AC49FC03464FA34" lastPageNumber="178" pageId="10" pageNumber="178">
<subSubSection id="C30C65361B21FFC39AC49FC034B5FC4C" box="[151,730,906,933]" pageId="10" pageNumber="178" type="nomenclature">
<paragraph id="8BA936BD1B21FFC39AC49FC034B5FC4C" blockId="10.[151,730,906,966]" box="[151,730,906,933]" pageId="10" pageNumber="178">
<heading id="D0E181D11B21FFC39AC49FC034B5FC4C" bold="true" box="[151,730,906,933]" fontSize="11" level="1" pageId="10" pageNumber="178" reason="1">
<emphasis id="B962EAAF1B21FFC39AC49FC034B5FC4C" bold="true" box="[151,730,906,933]" pageId="10" pageNumber="178">
<taxonomicName id="4C164D3E1B21FFC39AC49FC03422FC4C" authority="(McDonald, 1982)" baseAuthorityName="McDonald" baseAuthorityYear="1982" box="[151,589,906,933]" class="Insecta" family="Pentatomidae" genus="Braunus" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="178" phylum="Arthropoda" rank="species" species="asperus">
<emphasis id="B962EAAF1B21FFC39AC49FC0370AFC4D" bold="true" box="[151,357,906,932]" italics="true" pageId="10" pageNumber="178">Braunus asperus</emphasis>
(
<bibRefCitation id="EF874B4C1B21FFC39B279FC1342BFC4C" author="McDonald" box="[372,580,907,933]" pageId="10" pageNumber="178" refString="McDonald, F. J. D. (1982) Lojus asperus, a new genus and species (Hemiptera: Pentatomidae). Annals of the Entomological Society of America, 75, 327 - 329. http: // dx. doi. org / 10.1093 / aesa / 75.3.327" type="journal article" year="1982">McDonald, 1982</bibRefCitation>
)
</taxonomicName>
comb. nov.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30C65361B21FFC39AC49FE73753FC2F" box="[151,316,941,966]" pageId="10" pageNumber="178" type="description">
<paragraph id="8BA936BD1B21FFC39AC49FE73753FC2F" blockId="10.[151,730,906,966]" box="[151,316,941,966]" pageId="10" pageNumber="178">
(
<figureCitation id="132D2A381B21FFC39ACD9FE73764FC2C" box="[158,267,941,966]" captionStart="FIGURES 1 – 11" captionStartId="3.[151,264,1812,1835]" captionTargetBox="[151,1436,949,1788]" captionTargetId="figure@3.[151,1436,945,1789]" captionTargetPageId="3" captionText="FIGURES 1 – 11. Schematic drawings of pronotum of Braunus Distant, 1899 in dorsal view. 1 – 3. B. sciocorinus (Walker, 1867); 4. B. machadoi Barão &amp; Garbelotto sp. nov.; 5. B. gibbus Garbelotto &amp; Barão sp. nov.; 6. B. prionotus Barão &amp; Garbelotto sp. nov.; 7. B. asperus (McDonald, 1982) comb. nov.; 8. B. ateuchus (Brailovski &amp; Rolston, 1986) comb. nov.; 9. B. bordoni (Thomas, 1997) comb. nov.; 10. B. brettlizorum (Thomas, 1997) comb. nov.; 11. B. ocellatus (Thomas, 2001) comb. nov. Scale bars = 1 mm." httpUri="https://zenodo.org/record/261495/files/figure.png" pageId="10" pageNumber="178">Figures 7</figureCitation>
,
<figureCitation id="132D2A381B21FFC39B449FE4375BFC2F" box="[279,308,942,966]" captionStart="FIGURES 78 – 82" captionStartId="13.[151,264,1110,1133]" captionTargetBox="[151,1436,591,1086]" captionTargetId="figure@13.[151,1436,591,1086]" captionTargetPageId="13" captionText="FIGURES 78 – 82. Distribution of species of Braunus Distant, 1899 and localities of some collection events. 78. map of localities; 79. a collection locality of B. sciocorinus (Walker, 1867) and B. ocellatus (Thomas, 2001) comb. nov. (ECUADOR, Napo, Yanayacu Biological Station; photo by D. Forero); 80. type locality of B. machadoi Barão &amp; Garbelotto sp. nov. (PERU, Cuzco, Wayqecha Research Center; photo by C. Weirauch); 81 – 82. a collection locality of B. ocellatus (Thomas, 2001) comb. nov. (ECUADOR, Napo, 1.3 km SW of Baeza; photos by J. Skuhrovec)." httpUri="https://zenodo.org/record/261505/files/figure.png" pageId="10" pageNumber="178">78</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30C65361B21FFC39AC49FBC351BFBE2" box="[151,884,1013,1035]" pageId="10" pageNumber="178" type="reference_group">
<paragraph id="8BA936BD1B21FFC39AC49FBC351BFBE2" blockId="10.[151,884,1013,1035]" box="[151,884,1013,1035]" pageId="10" pageNumber="178">
<treatmentCitationGroup id="AB0611931B21FFC39AC49FBC351BFBE2" box="[151,884,1013,1035]" pageId="10" pageNumber="178">
<taxonomicName id="4C164D3E1B21FFC39AC49FBC3465FBE2" authority="McDonald, 1982: 327" authorityName="McDonald" authorityPageNumber="327" authorityYear="1982" box="[151,522,1013,1035]" class="Insecta" family="Pentatomidae" genus="Lojus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="178" phylum="Arthropoda" rank="species" species="asperus">
<emphasis id="B962EAAF1B21FFC39AC49FBC374BFBE2" box="[151,292,1014,1035]" italics="true" pageId="10" pageNumber="178">Lojus asperus</emphasis>
<treatmentCitation id="0AB710AC1B21FFC39B789FBF3465FBE2" author="McDonald" box="[299,522,1013,1035]" page="327" pageId="10" pageNumber="178" year="1982">
<bibRefCitation id="EF874B4C1B21FFC39B789FBF37B5FBE2" author="McDonald" box="[299,474,1013,1035]" pageId="10" pageNumber="178" refString="McDonald, F. J. D. (1982) Lojus asperus, a new genus and species (Hemiptera: Pentatomidae). Annals of the Entomological Society of America, 75, 327 - 329. http: // dx. doi. org / 10.1093 / aesa / 75.3.327" type="journal article" year="1982">McDonald, 1982</bibRefCitation>
: 327
</treatmentCitation>
</taxonomicName>
–329, figs 1–9;
<treatmentCitation id="0AB710AC1B21FFC398FB9FBF3500FBE2" author="Thomas" box="[680,879,1013,1035]" page="572" pageId="10" pageNumber="178" year="1997">
<bibRefCitation id="EF874B4C1B21FFC398FB9FBF3553FBE2" author="Thomas" box="[680,828,1013,1035]" pageId="10" pageNumber="178" refString="Thomas, D. B. (1997) The anocellate, flightless genus Lojus McDonald (Heteroptera: Pentatomidae). Annals of the Entomological Society of America, 90, 569 - 574. http: // dx. doi. org / 10.1093 / aesa / 90.5.569" type="journal article" year="1997">Thomas, 1997</bibRefCitation>
: 572
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30C65361B21FFC39AC4987233E3FBB9" box="[151,1420,1080,1105]" pageId="10" pageNumber="178" type="materials_examined">
<paragraph id="8BA936BD1B21FFC39AC4987233E3FBB9" blockId="10.[151,1437,1080,1501]" box="[151,1420,1080,1105]" pageId="10" pageNumber="178">
<materialsCitation id="3B7E3CE01B21FFC39AC4987233E7FBB9" ID-GBIF-Occurrence="2636403309" box="[151,1416,1080,1105]" collectingDate="1970-11-26" collectingDateMax="1970-11-27" collectingDateMin="1970-11-26" collectionCode="USNM" collectorName="L. E. Pena" country="Ecuador" elevation="1800" location="Loja a Zamora" pageId="10" pageNumber="178" specimenCount="1" specimenCount-male="1" stateProvince="Loja" typeStatus="holotype">
<emphasis id="B962EAAF1B21FFC39AC4987237EFFBB8" bold="true" box="[151,384,1080,1105]" pageId="10" pageNumber="178">Material examined.</emphasis>
<typeStatus id="54AD881F1B21FFC39BD498723780FBB8" box="[391,495,1080,1105]" pageId="10" pageNumber="178" type="holotype">Holotype</typeStatus>
<specimenCount id="9D10FD341B21FFC39BA598733462FBB9" box="[502,525,1081,1104]" pageId="10" pageNumber="178" type="male">♂</specimenCount>
,
<collectingCountry id="F301762D1B21FFC3984A987234CCFBB9" box="[537,675,1080,1104]" name="Ecuador" pageId="10" pageNumber="178" value="Ecuador">ECUADOR</collectingCountry>
,
<location id="8EC960661B21FFC398FD98723539FBB9" LSID="urn:lsid:plazi:treatment:03BF87AB1B21FFC39AC49FC03464FA34:8EC960661B21FFC398FD98723539FBB9" box="[686,854,1080,1105]" country="Ecuador" name="Loja a Zamora" pageId="10" pageNumber="178" stateProvince="Loja">Loja a Zamora</location>
,
<quantity id="4CEE9B581B21FFC39932987235D7FBB9" box="[865,952,1080,1104]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="10" pageNumber="178" unit="m" value="1800.0">
<elevation id="003BD18E1B21FFC39932987235D7FBB9" box="[865,952,1080,1104]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="10" pageNumber="178" unit="m" value="1800.0">1800 m</elevation>
</quantity>
,
<date id="FFA8107D1B21FFC399979872321FFBB8" box="[964,1136,1080,1105]" pageId="10" pageNumber="178" value="1970-11-26" valueMax="1970-11-27" valueMin="1970-11-26">
<collectingDate id="EFECE9951B21FFC399979872321FFBB8" box="[964,1136,1080,1105]" pageId="10" pageNumber="178" value="1970-11-26" valueMax="1970-11-27" valueMin="1970-11-26">26–27.XI.1970</collectingDate>
</date>
,
<collectorName id="26E3536B1B21FFC39E2898733286FBB9" box="[1147,1257,1081,1104]" pageId="10" pageNumber="178">L.E. Pena</collectorName>
leg. (
<collectionCode id="ED07AE781B21FFC39F7B987233EFFBB9" LSID="urn:lsid:biocol.org:col:34871" box="[1320,1408,1080,1104]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="178">USNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30C65361B21FFC39A9498163565FA7C" pageId="10" pageNumber="178" type="diagnosis">
<paragraph id="8BA936BD1B21FFC39A9498163565FA7C" blockId="10.[151,1437,1080,1501]" pageId="10" pageNumber="178">
<emphasis id="B962EAAF1B21FFC39A949816372EFB9C" bold="true" box="[199,321,1116,1141]" pageId="10" pageNumber="178">Diagnosis.</emphasis>
Body color mottled yellowish and brown; predominantly yellowish dorsally and brown ventrally. Each anterior angle of pronotum with a slender spine-like projection, surpassing the lateral margin of each compound eye; anterolateral margins strongly concave, each bearing 3+3 spine-like processes; humeral angles truncate apically, each with a finger-like process directed posteriorly; posterolateral margins sinuate, each with a tubercle medially; posterior margin straight (
<figureCitation id="132D2A381B21FFC398DC98A634BBFAEC" box="[655,724,1260,1285]" captionStart="FIGURES 1 – 11" captionStartId="3.[151,264,1812,1835]" captionTargetBox="[151,1436,949,1788]" captionTargetId="figure@3.[151,1436,945,1789]" captionTargetPageId="3" captionText="FIGURES 1 – 11. Schematic drawings of pronotum of Braunus Distant, 1899 in dorsal view. 1 – 3. B. sciocorinus (Walker, 1867); 4. B. machadoi Barão &amp; Garbelotto sp. nov.; 5. B. gibbus Garbelotto &amp; Barão sp. nov.; 6. B. prionotus Barão &amp; Garbelotto sp. nov.; 7. B. asperus (McDonald, 1982) comb. nov.; 8. B. ateuchus (Brailovski &amp; Rolston, 1986) comb. nov.; 9. B. bordoni (Thomas, 1997) comb. nov.; 10. B. brettlizorum (Thomas, 1997) comb. nov.; 11. B. ocellatus (Thomas, 2001) comb. nov. Scale bars = 1 mm." httpUri="https://zenodo.org/record/261495/files/figure.png" pageId="10" pageNumber="178">Fig. 7</figureCitation>
). Apex of scutellum reaching abdominal segment VII. Apex of hemelytral radial vein calloused; corium reaching posterior margin of abdominal segment VII; membrane surpassing apex of abdomen. Male genitalia: each posterolateral angle of pygophore rounded; ventral rim sinuous, inferior layer of ventral rim emarginate with one projection medially. Phallus: lateroventral conjunctival processes each digitiform; ductus seminis distalis dorsally curved.
</paragraph>
</subSubSection>
<subSubSection id="C30C65361B21FFC39A9499EA3488FA51" box="[199,743,1440,1465]" pageId="10" pageNumber="178" type="distribution">
<paragraph id="8BA936BD1B21FFC39A9499EA3488FA51" blockId="10.[151,1437,1080,1501]" box="[199,743,1440,1465]" pageId="10" pageNumber="178">
<emphasis id="B962EAAF1B21FFC39A9499EA370FFA50" bold="true" box="[199,352,1440,1465]" pageId="10" pageNumber="178">Distribution.</emphasis>
Ecuador (Loja, Zamora) (
<figureCitation id="132D2A381B21FFC398D599EA34B5FA51" box="[646,730,1440,1465]" captionStart="FIGURES 78 – 82" captionStartId="13.[151,264,1110,1133]" captionTargetBox="[151,1436,591,1086]" captionTargetId="figure@13.[151,1436,591,1086]" captionTargetPageId="13" captionText="FIGURES 78 – 82. Distribution of species of Braunus Distant, 1899 and localities of some collection events. 78. map of localities; 79. a collection locality of B. sciocorinus (Walker, 1867) and B. ocellatus (Thomas, 2001) comb. nov. (ECUADOR, Napo, Yanayacu Biological Station; photo by D. Forero); 80. type locality of B. machadoi Barão &amp; Garbelotto sp. nov. (PERU, Cuzco, Wayqecha Research Center; photo by C. Weirauch); 81 – 82. a collection locality of B. ocellatus (Thomas, 2001) comb. nov. (ECUADOR, Napo, 1.3 km SW of Baeza; photos by J. Skuhrovec)." httpUri="https://zenodo.org/record/261505/files/figure.png" pageId="10" pageNumber="178">Fig. 78</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30C65361B21FFC39A94998E3464FA34" box="[199,523,1476,1501]" pageId="10" pageNumber="178" type="discussion">
<paragraph id="8BA936BD1B21FFC39A94998E3464FA34" blockId="10.[151,1437,1080,1501]" box="[199,523,1476,1501]" pageId="10" pageNumber="178">
<emphasis id="B962EAAF1B21FFC39A94998E3754FA34" bold="true" box="[199,315,1476,1501]" pageId="10" pageNumber="178">Remarks.</emphasis>
Female unknown.
</paragraph>
</subSubSection>
</treatment>
</document>
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