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<mods:title id="7C4394BFDE5264C119783468D0BCA5FB">A revision of the Pontocaspian gastropods of the subfamily Caspiinae (Caenogastropoda: Hydrobiidae)</mods:title>
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<mods:affiliation id="A6142754C0A750F6CA2E3A280CC56A7D">Department of Animal Ecology and Systematics, Justus Liebig University, Heinrich-Buff-Ring 26 - 32 IFZ, 35392 Giessen, Germany. tneub @ zo. jlug. de; https: // orcid. org / 0000 - 0002 - 1398 - 9941 &amp; Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, The Netherlands.</mods:affiliation>
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<mods:affiliation id="6E58EFD92BDF1F417D50947D4D04ED5A">Department of Invertebrate Fauna and Systematics, Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, B. Khmelnytsky Str. 15, 01030 Kiev, Ukraine. &amp; Department of Cainozoic Deposits, Institute of Geological Sciences, National Academy of Sciences of Ukraine, O. Gontchar Str. 55 b, 01054 Kiev, Ukraine.</mods:affiliation>
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<mods:affiliation id="CC814EFF95F87F96A7F7B5A72E22AFE5">Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, The Netherlands. &amp; Department of Earth Sciences, Utrecht University, Budapestlaan 17, 3584 CD Utrecht, The Netherlands</mods:affiliation>
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<heading id="D0E181D9C85FFFA9FF3DFC28FC20A167" bold="true" box="[151,901,1014,1041]" fontSize="11" level="1" pageId="30" pageNumber="181" reason="1">
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<taxonomicName id="4C164D36C85FFFA9FF3DFC28FE95A166" ID-CoL="7TJ8Y" authority="Clessin &amp; W. Dybowski" box="[151,304,1014,1041]" class="Gastropoda" family="Hydrobiidae" genus="Ulskia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="181" phylum="Mollusca" rank="species" species="ulskii">
<emphasis id="B962EAA7C85FFFA9FF3DFC28FE95A166" bold="true" box="[151,304,1014,1041]" italics="true" pageId="30" pageNumber="181">Ulskia ulskii</emphasis>
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(Clessin &amp; W. Dybowski in W.
<bibRefCitation id="EF874B44C85FFFA9FD1CFC28FCD8A167" author="Dybowski, W." box="[694,893,1014,1040]" pageId="30" pageNumber="181" pagination="1 - 64" refId="ref30278" refString="Dybowski, W. (1887 - 1888) Die Gasteropoden-Fauna des Kaspischen Meeres. Nach der Sammlung des Akademikers Dr. K. E. v. Baer. Malakozoologische Blatter, Neue Folge, 10 (1 - 3), 1 - 64 (issue 1, 1887), 65 - 79 (issue 2, 1888), pls. 1 - 3 (issue 3, 1888)." type="journal article" year="1887">Dybowski, 1887</bibRefCitation>
)
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<paragraph id="8BA936B5C85FFFA9FF3DFBC4FEB1A143" blockId="30.[151,901,1014,1076]" box="[151,276,1050,1076]" pageId="30" pageNumber="181">
<figureCitation id="132D2A30C85FFFA9FF3DFBC4FF49A143" box="[151,236,1050,1076]" captionStart="FIGURE 16" captionStartId="31.[151,250,1760,1784]" captionTargetBox="[301,1286,181,1736]" captionTargetId="figure-17@31.[301,1286,181,1736]" captionTargetPageId="31" captionText="FIGURE 16. Ulskia ulskii (Clessin &amp; Dybowski in Dybowski, 1887) from the Caspian Sea Basin. (a) Lectotype, locality 26 (ZB-M W.Dyb. 76). (b) Paralectotype, locality 26 (ZB-M W.Dyb. 78). (c) Paralectotype, locality 26 (ZB-M W.Dyb. 78). (d) Paralectotype, locality 26 (ZB-M W.Dyb. 76). (e) Paralectotype, locality 26 (ZB-M W.Dyb. 76). (f, k) Specimen from locality 17 (ZIN, no number). (g, h) Holotype of Pyrgula nana, locality 15 (ZIN, no number). (i, j) Specimen labelled as “Caspia clessiniolaeformis, holotype”, locality 17 (ZIN, no number). (l) Holotype of Pyrgula behningi, locality 23 (ZIN #1). (m) Paratype of P. behningi, locality 14 (ZIN #2). (n, o) Paratype of P. behningi, locality 14 (ZIN #2). (ps, u) Five specimens from the Holocene of the Kura delta (localities 25d and 25f, RGM 13103061310310). (t, vz) Three specimens from Selitrennoye (locality 1, RGM 1309810, 1309809, 1309856). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4550190" httpUri="https://zenodo.org/record/4550190/files/figure.png" pageId="30" pageNumber="181">Figs 16</figureCitation>
,
<figureCitation id="132D2A30C85FFFA9FF52FBC4FEB1A143" box="[248,276,1050,1076]" captionStart="FIGURE 17" captionStartId="32.[151,250,1836,1860]" captionTargetBox="[151,1436,309,1812]" captionTargetId="figure-93@32.[151,1436,309,1812]" captionTargetPageId="32" captionText="FIGURE 17. Protoconch views of Ulskia ulskii (Clessin &amp; Dybowski in Dybowski, 1887). (a) Paralectotype, locality 26 (ZB- M W.Dyb. 76). (b, j, m) Paralectotype, locality 26 (ZB-M W.Dyb. 116). (c, i) Specimen from the Holocene of the Kura delta (locality 25f, RGM 1310308). (d, k) Specimen from the same area (locality 25d, RGM 1310310). (e) Specimen from the same area (locality 25f, RGM 1310306). (f, l) Specimen from the same area (locality 25d, RGM 1310309). (g, h, n, o) Specimen from the same area (locality 25d, RGM 1310307). Scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.4550192" httpUri="https://zenodo.org/record/4550192/files/figure.png" pageId="30" pageNumber="181">17</figureCitation>
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<taxonomicName id="4C164D36C85FFFA9FF3DFB81FE87A100" ID-CoL="7NWCX" authority="Ulskii" authorityName="Ulskii" box="[151,290,1119,1143]" class="Gastropoda" family="Hydrobiidae" genus="Caspia" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="181" phylum="Mollusca" rank="genus">
<emphasis id="B962EAA7C85FFFA9FF3DFB81FE87A100" box="[151,290,1119,1143]" italics="true" pageId="30" pageNumber="181">Caspia Ulskii</emphasis>
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nob.—Clessin &amp; W. Dybowski in W.
<treatmentCitation id="0AB710A4C85FFFA9FD0FFB81FC33A100" author="Dybowski, W." box="[677,918,1119,1143]" page="38" pageId="30" pageNumber="181" year="1887">
<bibRefCitation id="EF874B44C85FFFA9FD0FFB81FC33A100" author="Dybowski, W." box="[677,918,1119,1143]" pageId="30" pageNumber="181" pagination="1 - 64" refId="ref30278" refString="Dybowski, W. (1887 - 1888) Die Gasteropoden-Fauna des Kaspischen Meeres. Nach der Sammlung des Akademikers Dr. K. E. v. Baer. Malakozoologische Blatter, Neue Folge, 10 (1 - 3), 1 - 64 (issue 1, 1887), 65 - 79 (issue 2, 1888), pls. 1 - 3 (issue 3, 1888)." type="journal article" year="1887">Dybowski 1887: 3839</bibRefCitation>
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.
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[
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<emphasis id="B962EAA7C85FFFA9FF0AFBA0FF4DA1E2" box="[160,232,1150,1173]" italics="true" pageId="30" pageNumber="181">Caspia</emphasis>
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<taxonomicName id="4C164D36C85FFFA9FF5CFBA3FE96A1E2" authority="Anistratenko &amp; Neubauer &amp; Anistratenko &amp; Kijashko &amp; Wesselingh, 2021" authorityName="Anistratenko &amp; Neubauer &amp; Anistratenko &amp; Kijashko &amp; Wesselingh" authorityYear="2021" box="[246,307,1149,1173]" pageId="30" pageNumber="181" rank="species" species="ulskii" status="sp. nov.">
<emphasis id="B962EAA7C85FFFA9FF5CFBA3FE96A1E2" box="[246,307,1149,1173]" italics="true" pageId="30" pageNumber="181">Ulskii</emphasis>
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—W. Dybowski 1888: 79, pl. 3, fig. 8a, b.
</paragraph>
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<emphasis id="B962EAA7C85FFFA9FF3DFB42FF4DA1C3" box="[151,232,1180,1204]" italics="true" pageId="30" pageNumber="181">Pyrgula</emphasis>
[(
<emphasis id="B962EAA7C85FFFA9FF55FB42FEE4A1C3" box="[255,321,1180,1204]" italics="true" pageId="30" pageNumber="181">Ulskia</emphasis>
)]
<emphasis id="B962EAA7C85FFFA9FEF2FB42FE14A1C3" box="[344,433,1180,1204]" italics="true" pageId="30" pageNumber="181">behningi</emphasis>
Logv. et Star.
</taxonomicName>
<taxonomicNameLabel id="A25157DCC85FFFA9FDEFFB42FDDDA1C3" box="[581,632,1180,1204]" pageId="30" pageNumber="181" rank="species">sp. n.</taxonomicNameLabel>
<treatmentCitation id="0AB710A4C85FFFA9FD38FB42FB81A1C3" author="Logvinenko, B. M. &amp; Starobogatov, Ya. I." box="[658,1060,1180,1204]" page="380" pageId="30" pageNumber="181" year="1969">
<bibRefCitation id="EF874B44C85FFFA9FD38FB42FB81A1C3" author="Logvinenko, B. M. &amp; Starobogatov, Ya. I." box="[658,1060,1180,1204]" pageId="30" pageNumber="181" pagination="308 - 385" refId="ref32049" refString="Logvinenko, B. M. &amp; Starobogatov, Ya. I. (1969 [&quot; 1968 &quot;]) Mollusca. In: Birshtein, Y. A., Vinogradov, L. G., Kondakov, N. N., Kuhn, M. S., Astakhova, T. V. &amp; Romanova, N. N. (Eds.), Atlas bespozvonochnykh Kaspiyskogo morya. Pishchevaya Promyshlennost (Vsesoyuznyi Nauchno-issledovatel'skii Institut Morskogo Rybnogo Khozyaistva i Okeanografii), Moskva, pp. 308 - 385." type="book chapter" year="1969">Logvinenko &amp; Starobogatov 1969: 380</bibRefCitation>
</treatmentCitation>
, fig. 36713).
</treatmentCitationGroup>
</paragraph>
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<treatmentCitationGroup id="AB06119BC85FFFA9FF3DFB65FB11A1A4" box="[151,1204,1211,1235]" pageId="30" pageNumber="181">
<taxonomicName id="4C164D36C85FFFA9FF3DFB65FDBCA1A4" ID-CoL="7PFDQ" authority="Logv. et Star." authorityName="De Cristofori &amp; Jan" authorityYear="1832" box="[151,537,1211,1235]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="181" phylum="Mollusca" rank="genus" status="sp. nov.">
<emphasis id="B962EAA7C85FFFA9FF3DFB65FF4DA1A4" box="[151,232,1211,1235]" italics="true" pageId="30" pageNumber="181">Pyrgula</emphasis>
[(
<emphasis id="B962EAA7C85FFFA9FF55FB65FEE4A1A4" box="[255,321,1211,1235]" italics="true" pageId="30" pageNumber="181">Ulskia</emphasis>
)]
<emphasis id="B962EAA7C85FFFA9FEF2FB65FE2EA1A5" box="[344,395,1211,1234]" italics="true" pageId="30" pageNumber="181">nana</emphasis>
Logv. et Star.
</taxonomicName>
<taxonomicNameLabel id="A25157DCC85FFFA9FDB5FB65FDF7A1A4" box="[543,594,1211,1235]" pageId="30" pageNumber="181" rank="species">sp. n.</taxonomicNameLabel>
<treatmentCitation id="0AB710A4C85FFFA9FDC6FB65FB97A1A4" author="Logvinenko, B. M. &amp; Starobogatov, Ya. I." box="[620,1074,1211,1235]" page="379" pageId="30" pageNumber="181" year="1969">
<bibRefCitation id="EF874B44C85FFFA9FDC6FB65FB97A1A4" author="Logvinenko, B. M. &amp; Starobogatov, Ya. I." box="[620,1074,1211,1235]" pageId="30" pageNumber="181" pagination="308 - 385" refId="ref32049" refString="Logvinenko, B. M. &amp; Starobogatov, Ya. I. (1969 [&quot; 1968 &quot;]) Mollusca. In: Birshtein, Y. A., Vinogradov, L. G., Kondakov, N. N., Kuhn, M. S., Astakhova, T. V. &amp; Romanova, N. N. (Eds.), Atlas bespozvonochnykh Kaspiyskogo morya. Pishchevaya Promyshlennost (Vsesoyuznyi Nauchno-issledovatel'skii Institut Morskogo Rybnogo Khozyaistva i Okeanografii), Moskva, pp. 308 - 385." type="book chapter" year="1969">Logvinenko &amp; Starobogatov 1969: 379380</bibRefCitation>
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, fig. 36712).
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</paragraph>
<paragraph id="8BA936B5C85FFFA9FF3DFB07FB26A186" blockId="30.[151,1246,1119,1449]" box="[151,1155,1241,1265]" pageId="30" pageNumber="181">
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<emphasis id="B962EAA7C85FFFA9FF3DFB07FF4DA186" box="[151,232,1241,1265]" italics="true" pageId="30" pageNumber="181">Pyrgula</emphasis>
[(
<emphasis id="B962EAA7C85FFFA9FF55FB07FEE4A186" box="[255,321,1241,1265]" italics="true" pageId="30" pageNumber="181">Ulskia</emphasis>
)]
<emphasis id="B962EAA7C85FFFA9FEF2FB07FE2AA186" box="[344,399,1241,1265]" italics="true" pageId="30" pageNumber="181">ulskii</emphasis>
(Cless. et W. Dyb.)
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<bibRefCitation id="EF874B44C85FFFA9FDC7FB07FBA5A186" author="Logvinenko, B. M. &amp; Starobogatov, Ya. I." box="[621,1024,1241,1265]" pageId="30" pageNumber="181" pagination="308 - 385" refId="ref32049" refString="Logvinenko, B. M. &amp; Starobogatov, Ya. I. (1969 [&quot; 1968 &quot;]) Mollusca. In: Birshtein, Y. A., Vinogradov, L. G., Kondakov, N. N., Kuhn, M. S., Astakhova, T. V. &amp; Romanova, N. N. (Eds.), Atlas bespozvonochnykh Kaspiyskogo morya. Pishchevaya Promyshlennost (Vsesoyuznyi Nauchno-issledovatel'skii Institut Morskogo Rybnogo Khozyaistva i Okeanografii), Moskva, pp. 308 - 385." type="book chapter" year="1969">Logvinenko &amp; Starobogatov 1969: 379</bibRefCitation>
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, fig. 36710).
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</paragraph>
<paragraph id="8BA936B5C85FFFA9FF3DFB26FB31A078" blockId="30.[151,1246,1119,1449]" box="[151,1172,1272,1296]" pageId="30" pageNumber="181">
<treatmentCitationGroup id="AB06119BC85FFFA9FF3DFB26FB31A078" box="[151,1172,1272,1296]" pageId="30" pageNumber="181">
<taxonomicName id="4C164D36C85FFFA9FF3DFB26FD0CA067" ID-CoL="7PFDQ" authority="Logvinenko et Starobogatov, 1968" authorityName="Logvinenko et Starobogatov" authorityYear="1968" box="[151,681,1272,1296]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgula" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="181" phylum="Mollusca" rank="species" species="behningi">
<emphasis id="B962EAA7C85FFFA9FF3DFB26FEE2A067" box="[151,327,1272,1296]" italics="true" pageId="30" pageNumber="181">Pyrgula behningi</emphasis>
Logvinenko et Starobogatov, 1968
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<treatmentCitation id="0AB710A4C85FFFA9FD69FB26FC58A067" author="Kantor, Yu. I. &amp; Sysoev, A. V." box="[707,1021,1272,1296]" page="97" pageId="30" pageNumber="181" year="2006">
<bibRefCitation id="EF874B44C85FFFA9FD69FB26FC58A067" author="Kantor, Yu. I. &amp; Sysoev, A. V." box="[707,1021,1272,1296]" pageId="30" pageNumber="181" refId="ref31309" refString="Kantor, Yu. I. &amp; Sysoev, A. V. (2006) Morskiye i solonovatovodnyye bryukhonogiye mollyuski Rossii i sopredel'nykh stran: illyustrirovannyy katalog. KMK Scientific Press, Moscow, 372 pp., 140 pls pp." type="book" year="2006">Kantor &amp; Sysoev 2006: 9798</bibRefCitation>
</treatmentCitation>
, pl. 45, fig. M.
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</paragraph>
<paragraph id="8BA936B5C85FFFA9FF3DFAC9FBF5A059" blockId="30.[151,1246,1119,1449]" box="[151,1104,1303,1327]" pageId="30" pageNumber="181">
<treatmentCitationGroup id="AB06119BC85FFFA9FF3DFAC9FBF5A059" box="[151,1104,1303,1327]" pageId="30" pageNumber="181">
<taxonomicName id="4C164D36C85FFFA9FF3DFAC9FD26A058" ID-CoL="7PFDQ" authority="Logvinenko et Starobogatov, 1968" authorityName="Logvinenko et Starobogatov" authorityYear="1968" box="[151,643,1303,1327]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgula" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="181" phylum="Mollusca" rank="species" species="nana">
<emphasis id="B962EAA7C85FFFA9FF3DFAC9FE84A059" box="[151,289,1303,1327]" italics="true" pageId="30" pageNumber="181">Pyrgula nana</emphasis>
Logvinenko et Starobogatov, 1968
</taxonomicName>
<treatmentCitation id="0AB710A4C85FFFA9FD37FAC9FC18A058" author="Kantor, Yu. I. &amp; Sysoev, A. V." box="[669,957,1303,1327]" page="101" pageId="30" pageNumber="181" year="2006">
<bibRefCitation id="EF874B44C85FFFA9FD37FAC9FC18A058" author="Kantor, Yu. I. &amp; Sysoev, A. V." box="[669,957,1303,1327]" pageId="30" pageNumber="181" refId="ref31309" refString="Kantor, Yu. I. &amp; Sysoev, A. V. (2006) Morskiye i solonovatovodnyye bryukhonogiye mollyuski Rossii i sopredel'nykh stran: illyustrirovannyy katalog. KMK Scientific Press, Moscow, 372 pp., 140 pls pp." type="book" year="2006">Kantor &amp; Sysoev 2006: 101</bibRefCitation>
</treatmentCitation>
, pl. 47, fig. D.
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</paragraph>
<paragraph id="8BA936B5C85FFFA9FF3DFAEBFB7BA03A" blockId="30.[151,1246,1119,1449]" box="[151,1246,1333,1357]" pageId="30" pageNumber="181">
<treatmentCitationGroup id="AB06119BC85FFFA9FF3DFAEBFB7BA03A" box="[151,1246,1333,1357]" pageId="30" pageNumber="181">
<taxonomicName id="4C164D36C85FFFA9FF3DFAEBFE80A03A" ID-CoL="7PFDQ" baseAuthorityName="Clessin &amp; W.Dybowski" baseAuthorityYear="1888" box="[151,293,1333,1357]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgula" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="181" phylum="Mollusca" rank="species" species="ulskii">
<emphasis id="B962EAA7C85FFFA9FF3DFAEBFE80A03A" box="[151,293,1333,1357]" italics="true" pageId="30" pageNumber="181">Pyrgula ulskii</emphasis>
</taxonomicName>
(Clessin et W. Dybowski in W. Dybowski, 1888)—
<treatmentCitation id="0AB710A4C85FFFA9FC9AFAEBFBF4A03A" author="Kantor, Yu. I. &amp; Sysoev, A. V." box="[816,1105,1333,1357]" page="104" pageId="30" pageNumber="181" year="2006">
<bibRefCitation id="EF874B44C85FFFA9FC9AFAEBFBF4A03A" author="Kantor, Yu. I. &amp; Sysoev, A. V." box="[816,1105,1333,1357]" pageId="30" pageNumber="181" refId="ref31309" refString="Kantor, Yu. I. &amp; Sysoev, A. V. (2006) Morskiye i solonovatovodnyye bryukhonogiye mollyuski Rossii i sopredel'nykh stran: illyustrirovannyy katalog. KMK Scientific Press, Moscow, 372 pp., 140 pls pp." type="book" year="2006">Kantor &amp; Sysoev 2006: 104</bibRefCitation>
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, pl. 45, fig. F.
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</paragraph>
<paragraph id="8BA936B5C85FFFA9FF3DFA8AFB64A01B" blockId="30.[151,1246,1119,1449]" box="[151,1217,1364,1388]" pageId="30" pageNumber="181">
<treatmentCitationGroup id="AB06119BC85FFFA9FF3DFA8AFB64A01B" box="[151,1217,1364,1388]" pageId="30" pageNumber="181">
<taxonomicName id="4C164D36C85FFFA9FF3DFA8AFEB2A01B" ID-CoL="7TJ8Y" authority="(Clessin &amp; W. Dybowski in W. Dybowski, 1887)" box="[151,279,1364,1388]" class="Gastropoda" family="Hydrobiidae" genus="Ulskia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="181" phylum="Mollusca" rank="species" species="ulskii">
<emphasis id="B962EAA7C85FFFA9FF3DFA8AFEB2A01B" box="[151,279,1364,1388]" italics="true" pageId="30" pageNumber="181">Ulskia ulskii</emphasis>
</taxonomicName>
(Clessin &amp; W. Dybowski in W.
<bibRefCitation id="EF874B44C85FFFA9FDF6FA8AFCA4A01B" author="Dybowski, W." box="[604,769,1364,1388]" pageId="30" pageNumber="181" pagination="1 - 64" refId="ref30278" refString="Dybowski, W. (1887 - 1888) Die Gasteropoden-Fauna des Kaspischen Meeres. Nach der Sammlung des Akademikers Dr. K. E. v. Baer. Malakozoologische Blatter, Neue Folge, 10 (1 - 3), 1 - 64 (issue 1, 1887), 65 - 79 (issue 2, 1888), pls. 1 - 3 (issue 3, 1888)." type="journal article" year="1887">Dybowski, 1887</bibRefCitation>
)—
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<bibRefCitation id="EF874B44C85FFFA9FC89FA8AFBECA01B" author="Neubauer, T. A. &amp; van de Velde, S. &amp; Yanina, T. A. &amp; Wesselingh, F. P." box="[803,1097,1364,1388]" pageId="30" pageNumber="181" pagination="43 - 103" refId="ref32549" refString="Neubauer, T. A., van de Velde, S., Yanina, T. A. &amp; Wesselingh, F. P. (2018) A late Pleistocene gastropod fauna from the northern Caspian Sea with implications for Pontocaspian gastropod taxonomy. ZooKeys, 770, 43 - 103. https: // doi. org / 10.3897 / zookeys. 770.25365" type="journal article" year="2018">
Neubauer
<emphasis id="B962EAA7C85FFFA9FC24FA8BFC1EA01B" box="[910,955,1364,1388]" italics="true" pageId="30" pageNumber="181">et al</emphasis>
. 2018: 5254
</bibRefCitation>
</treatmentCitation>
, fig. 5AK.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA936B5C85FFFA9FF3DFAADFBA7A0FC" blockId="30.[151,1246,1119,1449]" box="[151,1026,1395,1419]" pageId="30" pageNumber="181">
<treatmentCitationGroup id="AB06119BC85FFFA9FF3DFAADFBA7A0FC" box="[151,1026,1395,1419]" pageId="30" pageNumber="181">
<taxonomicName id="4C164D36C85FFFA9FF3DFAADFD0EA0FC" ID-CoL="84ZDZ" authority="(Logvinenko &amp; Starobogatov, 1969)" baseAuthorityName="Logvinenko &amp; Starobogatov" baseAuthorityYear="1969" box="[151,683,1395,1419]" class="Gastropoda" family="Hydrobiidae" genus="Ulskia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="181" phylum="Mollusca" rank="species" species="behningi">
<emphasis id="B962EAA7C85FFFA9FF3DFAADFE9CA0FC" box="[151,313,1395,1419]" italics="true" pageId="30" pageNumber="181">Ulskia behningi</emphasis>
(
<bibRefCitation id="EF874B44C85FFFA9FEEFFAADFD00A0FC" author="Logvinenko, B. M. &amp; Starobogatov, Ya. I." box="[325,677,1395,1419]" pageId="30" pageNumber="181" pagination="308 - 385" refId="ref32049" refString="Logvinenko, B. M. &amp; Starobogatov, Ya. I. (1969 [&quot; 1968 &quot;]) Mollusca. In: Birshtein, Y. A., Vinogradov, L. G., Kondakov, N. N., Kuhn, M. S., Astakhova, T. V. &amp; Romanova, N. N. (Eds.), Atlas bespozvonochnykh Kaspiyskogo morya. Pishchevaya Promyshlennost (Vsesoyuznyi Nauchno-issledovatel'skii Institut Morskogo Rybnogo Khozyaistva i Okeanografii), Moskva, pp. 308 - 385." type="book chapter" year="1969">Logvinenko &amp; Starobogatov, 1969</bibRefCitation>
)
</taxonomicName>
<treatmentCitation id="0AB710A4C85FFFA9FD6FFAADFC5AA0FC" author="Wesselingh, F. P. &amp; Neubauer, T. A. &amp; Anistratenko, V. V. &amp; Vinarski, M. V. &amp; Yanina, T. &amp; ter Poorten, J. J. &amp; Kijashko, P. V. &amp; Albrecht, C. &amp; Anistratenko, O. Yu. &amp; D'Hont, A. &amp; Frolov, P. &amp; Martinez Gandara, A. &amp; Gittenberger, A. &amp; Gogaladze, A. &amp; Karpinsky, M. &amp; Lattuada, M. &amp; Popa, L. &amp; Sands, A. F. &amp; van de Velde, S. &amp; Vandendorpe, J. &amp; Wilke, T." box="[709,1023,1395,1419]" httpUri="http://treatment.plazi.org/id/48C7D9004E54C5585471B4B822C0F65B" page="72" pageId="30" pageNumber="181" year="2019">
<bibRefCitation id="EF874B44C85FFFA9FD6FFAADFC5AA0FC" author="Wesselingh, F. P. &amp; Neubauer, T. A. &amp; Anistratenko, V. V. &amp; Vinarski, M. V. &amp; Yanina, T. &amp; ter Poorten, J. J. &amp; Kijashko, P. V. &amp; Albrecht, C. &amp; Anistratenko, O. Yu. &amp; D'Hont, A. &amp; Frolov, P. &amp; Martinez Gandara, A. &amp; Gittenberger, A. &amp; Gogaladze, A. &amp; Karpinsky, M. &amp; Lattuada, M. &amp; Popa, L. &amp; Sands, A. F. &amp; van de Velde, S. &amp; Vandendorpe, J. &amp; Wilke, T." box="[709,1023,1395,1419]" pageId="30" pageNumber="181" pagination="31 - 124" refId="ref34307" refString="Wesselingh, F. P., Neubauer, T. A., Anistratenko, V. V., Vinarski, M. V., Yanina, T., ter Poorten, J. J., Kijashko, P. V., Albrecht, C., Anistratenko, O. Yu., D'Hont, A., Frolov, P., Martinez Gandara, A., Gittenberger, A., Gogaladze, A., Karpinsky, M., Lattuada, M., Popa, L., Sands, A. F., van de Velde, S., Vandendorpe, J. &amp; Wilke, T. (2019) Mollusc species from the Pontocaspian region - an expert opinion list. ZooKeys, 827, 31 - 124. https: // doi. org / 10.3897 / zookeys. 827.31365" type="journal article" year="2019">
Wesselingh
<emphasis id="B962EAA7C85FFFA9FCE9FAADFCD5A0FC" box="[835,880,1395,1419]" italics="true" pageId="30" pageNumber="181">et al</emphasis>
. 2019: 7273
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA936B5C85FFFA9FF3DFA4FFC53A0DE" blockId="30.[151,1246,1119,1449]" box="[151,1014,1425,1449]" pageId="30" pageNumber="181">
<treatmentCitationGroup id="AB06119BC85FFFA9FF3DFA4FFC53A0DE" box="[151,1014,1425,1449]" pageId="30" pageNumber="181">
<taxonomicName id="4C164D36C85FFFA9FF3DFA4FFEB2A0DE" ID-CoL="7TJ8Y" authority="(Clessin &amp; Dybowski in Dybowski, 1887)" box="[151,279,1425,1449]" class="Gastropoda" family="Hydrobiidae" genus="Ulskia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="181" phylum="Mollusca" rank="species" species="ulskii">
<emphasis id="B962EAA7C85FFFA9FF3DFA4FFEB2A0DE" box="[151,279,1425,1449]" italics="true" pageId="30" pageNumber="181">Ulskia ulskii</emphasis>
</taxonomicName>
(Clessin &amp; Dybowski in
<bibRefCitation id="EF874B44C85FFFA9FDB3FA4FFD1BA0DE" author="Dybowski, W." box="[537,702,1425,1449]" pageId="30" pageNumber="181" pagination="1 - 64" refId="ref30278" refString="Dybowski, W. (1887 - 1888) Die Gasteropoden-Fauna des Kaspischen Meeres. Nach der Sammlung des Akademikers Dr. K. E. v. Baer. Malakozoologische Blatter, Neue Folge, 10 (1 - 3), 1 - 64 (issue 1, 1887), 65 - 79 (issue 2, 1888), pls. 1 - 3 (issue 3, 1888)." type="journal article" year="1887">Dybowski, 1887</bibRefCitation>
)—
<treatmentCitation id="0AB710A4C85FFFA9FD75FA4FFC57A0DE" author="Wesselingh, F. P. &amp; Neubauer, T. A. &amp; Anistratenko, V. V. &amp; Vinarski, M. V. &amp; Yanina, T. &amp; ter Poorten, J. J. &amp; Kijashko, P. V. &amp; Albrecht, C. &amp; Anistratenko, O. Yu. &amp; D'Hont, A. &amp; Frolov, P. &amp; Martinez Gandara, A. &amp; Gittenberger, A. &amp; Gogaladze, A. &amp; Karpinsky, M. &amp; Lattuada, M. &amp; Popa, L. &amp; Sands, A. F. &amp; van de Velde, S. &amp; Vandendorpe, J. &amp; Wilke, T." box="[735,1010,1425,1449]" httpUri="http://treatment.plazi.org/id/9AFCF7F38B5BADFE72D5129962CAB2F1" page="73" pageId="30" pageNumber="181" year="2019">
<bibRefCitation id="EF874B44C85FFFA9FD75FA4FFC57A0DE" author="Wesselingh, F. P. &amp; Neubauer, T. A. &amp; Anistratenko, V. V. &amp; Vinarski, M. V. &amp; Yanina, T. &amp; ter Poorten, J. J. &amp; Kijashko, P. V. &amp; Albrecht, C. &amp; Anistratenko, O. Yu. &amp; D'Hont, A. &amp; Frolov, P. &amp; Martinez Gandara, A. &amp; Gittenberger, A. &amp; Gogaladze, A. &amp; Karpinsky, M. &amp; Lattuada, M. &amp; Popa, L. &amp; Sands, A. F. &amp; van de Velde, S. &amp; Vandendorpe, J. &amp; Wilke, T." box="[735,1010,1425,1449]" pageId="30" pageNumber="181" pagination="31 - 124" refId="ref34307" refString="Wesselingh, F. P., Neubauer, T. A., Anistratenko, V. V., Vinarski, M. V., Yanina, T., ter Poorten, J. J., Kijashko, P. V., Albrecht, C., Anistratenko, O. Yu., D'Hont, A., Frolov, P., Martinez Gandara, A., Gittenberger, A., Gogaladze, A., Karpinsky, M., Lattuada, M., Popa, L., Sands, A. F., van de Velde, S., Vandendorpe, J. &amp; Wilke, T. (2019) Mollusc species from the Pontocaspian region - an expert opinion list. ZooKeys, 827, 31 - 124. https: // doi. org / 10.3897 / zookeys. 827.31365" type="journal article" year="2019">
Wesselingh
<emphasis id="B962EAA7C85FFFA9FCF7FA4CFC2CA0DE" box="[861,905,1425,1449]" italics="true" pageId="30" pageNumber="181">et al</emphasis>
. 2019: 73
</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="C30C653EC85FFFA9FF3DFA06FB2EA2B1" pageId="30" pageNumber="181" type="materials_examined">
<paragraph id="8BA936B5C85FFFA9FF3DFA06FB4DA329" blockId="30.[151,1437,1495,1990]" pageId="30" pageNumber="181">
<emphasis id="B962EAA7C85FFFA9FF3DFA06FEEDA086" bold="true" box="[151,328,1495,1522]" pageId="30" pageNumber="181">Type material.</emphasis>
<materialsCitation id="3B7E3CE8C85FFFA9FEFBFA06FB4DA329" ID-GBIF-Occurrence="3042095330" collectingDate="1956-06-26" collectingDateMax="1957-08-04" collectingDateMin="1956-06-26" collectionCode="ZB-M, ZIN" collectorName=" B. M. Logvinenko" pageId="30" pageNumber="181" specimenCode="ZIN #1" specimenCount="115" typeStatus="lectotype">
<typeStatus id="54AD8817C85FFFA9FEFBFA06FE64A085" box="[337,449,1496,1522]" pageId="30" pageNumber="181" type="lectotype">Lectotype</typeStatus>
(
<collectionCode id="ED07AE70C85FFFA9FE79FA06FDBEA085" box="[467,539,1496,1522]" pageId="30" pageNumber="181">ZB-M</collectionCode>
<collectorName id="26E35363C85FFFA9FD8EFA09FDDFA086" box="[548,634,1495,1521]" pageId="30" pageNumber="181">W.Dyb.</collectorName>
76) and
<specimenCount id="9D10FD3CC85FFFA9FD4FFA06FC09A086" box="[741,940,1495,1522]" pageId="30" pageNumber="181" type="generic" typeStatus="paralectotypes">33 paralectotypes</specimenCount>
(
<collectionCode id="ED07AE70C85FFFA9FC17FA06FBA0A085" box="[957,1029,1496,1522]" pageId="30" pageNumber="181">ZB-M</collectionCode>
<collectorName id="26E35363C85FFFA9FBA4FA09FBC0A086" box="[1038,1125,1495,1521]" pageId="30" pageNumber="181">W.Dyb.</collectorName>
76 and 99) (designated by
<bibRefCitation id="EF874B44C85FFFA9FF3DFA25FE0BA361" author="Anistratenko, V. V. &amp; Anistratenko, O. Yu. &amp; Kadolsky, D." box="[151,430,1531,1558]" pageId="30" pageNumber="181" pagination="35 - 62" refId="ref29151" refString="Anistratenko, V. V., Anistratenko, O. Yu. &amp; Kadolsky, D. (2019) Karl E. von Baer's collection of Caspian Sea molluscs stored in the Zoological Museum of Lviv University, Ukraine. Part 2. Type materials of gastropod species described by Stephan Clessin and Wladyslaw Dybowski in 1887 - 1888. Archiv fur Molluskenkunde, 148 (1), 35 - 62. https: // doi. org / 10.1127 / arch. moll / 148 / 035 - 062" type="journal article" year="2019">
Anistratenko
<emphasis id="B962EAA7C85FFFA9FE9AFA22FEC7A362" box="[304,354,1531,1557]" italics="true" pageId="30" pageNumber="181">et al</emphasis>
. 2019
</bibRefCitation>
).
<collectorName id="26E35363C85FFFA9FE6BFA25FE48A362" box="[449,493,1531,1557]" pageId="30" pageNumber="181">The</collectorName>
<typeStatus id="54AD8817C85FFFA9FE5FFA25FDF2A362" box="[501,599,1531,1557]" pageId="30" pageNumber="181" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C164D36C85FFFA9FDD5FA22FD5BA362" baseAuthorityName="Logvinenko &amp; Starobogatov" baseAuthorityYear="1969" box="[639,766,1531,1557]" class="Gastropoda" family="Hydrobiidae" genus="Ulskia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="181" phylum="Mollusca" rank="species" species="behningi">
<emphasis id="B962EAA7C85FFFA9FDD5FA22FD5BA362" box="[639,766,1531,1557]" italics="true" pageId="30" pageNumber="181">P. behningi</emphasis>
</taxonomicName>
(
<specimenCode id="DBB09ECEC85FFFA9FCA7FA22FCC7A361" box="[781,866,1532,1558]" collectionCode="ZIN" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="30" pageNumber="181" type="Museum">ZIN #1</specimenCode>
) and
<specimenCount id="9D10FD3CC85FFFA9FC0EFA22FB96A361" box="[932,1075,1532,1558]" pageId="30" pageNumber="181" type="generic" typeStatus="paratypes">82 paratypes</specimenCount>
(
<collectionCode id="ED07AE70C85FFFA9FBE9FA22FBD6A361" box="[1091,1139,1532,1558]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="30" pageNumber="181" type="Museum">ZIN</collectionCode>
##211) are listed in
<collectionCode id="ED07AE70C85FFFA9FAC4FA22FA38A361" box="[1390,1437,1532,1558]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="30" pageNumber="181" type="Museum">ZIN</collectionCode>
systematic catalogue, collected by
<collectorName id="26E35363C85FFFA9FDB4F9FEFD43A34E" box="[542,742,1567,1594]" pageId="30" pageNumber="181">B.M. Logvinenko</collectorName>
in
<date id="FFA81075C85FFFA9FCA1F9C1FC2EA34D" box="[779,907,1567,1594]" pageId="30" pageNumber="181" value="1956-06-26">
<collectingDate id="EFECE99DC85FFFA9FCA1F9C1FC2EA34D" box="[779,907,1567,1594]" pageId="30" pageNumber="181" value="1956-06-26">26/06/1956</collectingDate>
</date>
<date id="FFA81075C85FFFA9FC31F9C1FBBEA34D" box="[923,1051,1567,1594]" pageId="30" pageNumber="181" value="1957-08-04">
<collectingDate id="EFECE99DC85FFFA9FC31F9C1FBBEA34D" box="[923,1051,1567,1594]" pageId="30" pageNumber="181" value="1957-08-04">04/08/1957</collectingDate>
</date>
. As for
<taxonomicName id="4C164D36C85FFFA9FBD9F9FEFB62A34E" authorityName="Logvinenko et Starobogatov" authorityYear="1968" box="[1139,1223,1568,1593]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgula" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="181" phylum="Mollusca" rank="species" species="nana">
<emphasis id="B962EAA7C85FFFA9FBD9F9FEFB62A34E" box="[1139,1223,1568,1593]" italics="true" pageId="30" pageNumber="181">P. nana</emphasis>
</taxonomicName>
, we found a specimen labelled as “
<typeStatus id="54AD8817C85FFFA9FEFCF99DFE18A32A" box="[342,445,1603,1629]" pageId="30" pageNumber="181" type="holotype">holotype</typeStatus>
” in the Starobogatov collection (
<collectionCode id="ED07AE70C85FFFA9FC84F99AFCC7A329" box="[814,866,1604,1630]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="30" pageNumber="181" type="Museum">ZIN</collectionCode>
), as well as 13 probable
<typeStatus id="54AD8817C85FFFA9FBDFF99AFB41A329" box="[1141,1252,1604,1630]" pageId="30" pageNumber="181" type="paratype">paratypes</typeStatus>
.
</materialsCitation>
</paragraph>
<paragraph id="8BA936B5C85FFFA9FF6DF9B6FB8AA265" blockId="30.[151,1437,1495,1990]" pageId="30" pageNumber="181">
<emphasis id="B962EAA7C85FFFA9FF6DF9B6FECCA3F6" bold="true" box="[199,361,1639,1666]" pageId="30" pageNumber="181">Type locality.</emphasis>
“Kaspi-See” (Caspian Sea, without further details;
<tableCitation id="C694030EC85FFFA9FC6CF9B9FBB8A3F5" box="[966,1053,1639,1666]" captionStart="TABLE 1" captionStartId="3.[151,239,473,499]" captionTargetPageId="3" captionText="TABLE 1. Locality information of the studied material. Coordinates of collecting sites are given based on data in museum catalogue cards, labels and corresponding publications. Asterisks mark localities with approximate geographic coordinates based on indirect data provided in the ZIN catalogue. Abbreviations: q.—quadrant, st.—station." httpUri="http://table.plazi.org/id/DF69663DC842FFB4FF3DFE07F8C6A760" pageId="30" pageNumber="181" tableUuid="DF69663DC842FFB4FF3DFE07F8C6A760">Table 1</tableCitation>
, locality 26). Type locality of
<taxonomicName id="4C164D36C85FFFA9FA22F9B6FF5CA3D2" baseAuthorityName="Logvinenko &amp; Starobogatov" baseAuthorityYear="1969" class="Gastropoda" family="Hydrobiidae" genus="Ulskia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="181" phylum="Mollusca" rank="species" species="behningi">
<emphasis id="B962EAA7C85FFFA9FA22F9B6FF5CA3D2" italics="true" pageId="30" pageNumber="181">P. behningi</emphasis>
</taxonomicName>
: Western part of the South Caspian Sea in the vicinity of the Kura River mouth,
<quantity id="4CEE9B50C85FFFA9FB23F952FB74A3D1" box="[1161,1233,1676,1702]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="30" pageNumber="181" unit="m" value="120.0">120 m</quantity>
(locality 23). The
<typeStatus id="54AD8817C85FFFA9FF3DF96EFEA7A3BD" box="[151,258,1712,1738]" pageId="30" pageNumber="181" type="paratype">paratypes</typeStatus>
were collected off Apsheron Peninsula (
<collectingCountry id="F3017625C85FFFA9FD6BF971FCE5A3BD" box="[705,832,1711,1738]" name="Azerbaijan" pageId="30" pageNumber="181">Azerbaijan</collectingCountry>
) (locality 14). Type locality of
<taxonomicName id="4C164D36C85FFFA9FB3CF96EFB4CA3BE" authorityName="Logvinenko et Starobogatov" authorityYear="1968" box="[1174,1257,1712,1737]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgula" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="181" phylum="Mollusca" rank="species" species="nana">
<emphasis id="B962EAA7C85FFFA9FB3CF96EFB4CA3BE" box="[1174,1257,1712,1737]" italics="true" pageId="30" pageNumber="181">P. nana</emphasis>
</taxonomicName>
: western part of the Caspian Sea near Apsheron peninsula,
<quantity id="4CEE9B50C85FFFA9FDC7F90AFD00A399" box="[621,677,1748,1774]" metricMagnitude="1" metricUnit="m" metricValue="8.8" pageId="30" pageNumber="181" unit="m" value="88.0">88 m</quantity>
(locality 15); the 13 probable
<typeStatus id="54AD8817C85FFFA9FC58F90AFBF8A399" box="[1010,1117,1748,1774]" pageId="30" pageNumber="181" type="paratype">paratypes</typeStatus>
were retrieved in the eastern part of the South Caspian Sea off Cheleken Peninsula,
<collectingCountry id="F3017625C85FFFA9FD56F929FC32A266" box="[764,919,1783,1809]" name="Turkmenistan" pageId="30" pageNumber="181">Turkmenistan</collectingCountry>
(locality 20).
</paragraph>
<paragraph id="8BA936B5C85FFFA9FF6DF8C5FB2EA2B1" blockId="30.[151,1437,1495,1990]" pageId="30" pageNumber="181">
<materialsCitation id="3B7E3CE8C85FFFA9FF6DF8C5FA38A2D5" ID-GBIF-Occurrence="3042095316" collectionCode="ZIN" elevation="64" pageId="30" pageNumber="181" specimenCode="RGM 1309810, 1309809, 1309856" specimenCount="3" typeStatus="holotype">
<emphasis id="B962EAA7C85FFFA9FF6DF8C5FE26A242" bold="true" box="[199,387,1819,1845]" pageId="30" pageNumber="181">Other material.</emphasis>
The “
<typeStatus id="54AD8817C85FFFA9FE62F8C5FD8AA242" box="[456,559,1819,1845]" pageId="30" pageNumber="181" type="holotype">holotype</typeStatus>
” and 6 “
<typeStatus id="54AD8817C85FFFA9FD39F8C2FCA1A241" box="[659,772,1820,1846]" pageId="30" pageNumber="181" type="paratype">paratypes</typeStatus>
” of “
<taxonomicName id="4C164D36C85FFFA9FCE8F8C2FBC3A242" box="[834,1126,1819,1846]" class="Gastropoda" family="Hydrobiidae" genus="Caspia" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="181" phylum="Mollusca" rank="species" species="clessiniolaeformis">
<emphasis id="B962EAA7C85FFFA9FCE8F8C2FBC3A242" box="[834,1126,1819,1846]" italics="true" pageId="30" pageNumber="181">Caspia clessiniolaeformis</emphasis>
</taxonomicName>
”, an unavailable collection name coined by Starobogatov (
<collectionCode id="ED07AE70C85FFFA9FE5AF89EFD86A22D" box="[496,547,1856,1882]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="30" pageNumber="181" type="Museum">ZIN</collectionCode>
, no number), collected in 1957 and
<quantity id="4CEE9B50C85FFFA9FC04F8E1FBA7A22E" box="[942,1026,1855,1881]" metricMagnitude="1" metricUnit="m" metricValue="4.98348" pageId="30" pageNumber="181" unit="in" value="1962.0">1962 in</quantity>
the eastern part of the Middle Caspi-an Sea at a depth of
<quantity id="4CEE9B50C85FFFA9FED9F8BDFE71A209" box="[371,468,1891,1918]" metricMagnitude="1" metricUnit="m" metricValue="6.35" metricValueMax="7.1" metricValueMin="5.6" pageId="30" pageNumber="181" unit="m" value="63.5" valueMax="71.0" valueMin="56.0">
<elevation id="003BD186C85FFFA9FED9F8BDFE71A209" box="[371,468,1891,1918]" metricMagnitude="1" metricUnit="m" metricValue="6.35" metricValueMax="7.1" metricValueMin="5.6" pageId="30" pageNumber="181" unit="m" value="63.5" valueMax="71.0" valueMin="56.0">5671 m</elevation>
</quantity>
(localities 5 and 17). Further three specimens labelled by Starobogatov as
<taxonomicName id="4C164D36C85FFFA9FAA0F8BAFA38A20A" authorityName="Clessin &amp; W. Dybowski" authorityYear="1887" box="[1290,1437,1891,1918]" class="Gastropoda" family="Hydrobiidae" genus="Caspia" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="181" phylum="Mollusca" rank="species" species="ulskii">
<emphasis id="B962EAA7C85FFFA9FAA0F8BAFA38A20A" box="[1290,1437,1891,1918]" italics="true" pageId="30" pageNumber="181">Caspia ulskii</emphasis>
</taxonomicName>
come from locality 17 (
<collectionCode id="ED07AE70C85FFFA9FE08F856FE70A2D5" box="[418,469,1928,1954]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="30" pageNumber="181" type="Museum">ZIN</collectionCode>
, no number), three from Selitrennoye (locality 1,
<specimenCode id="DBB09ECEC85FFFA9FBA2F856FB11A2D5" box="[1032,1204,1928,1954]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="30" pageNumber="181" type="Museum">RGM 1309810</specimenCode>
,
<specimenCode id="DBB09ECEC85FFFA9FB6AF856FA80A2D5" box="[1216,1317,1928,1954]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="30" pageNumber="181" type="Museum">1309809</specimenCode>
,
<specimenCode id="DBB09ECEC85FFFA9FA9BF859FA30A2D5" box="[1329,1429,1927,1954]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="30" pageNumber="181" type="Museum">1309856</specimenCode>
)
</materialsCitation>
<materialsCitation id="3B7E3CE8C85FFFA9FF3DF875FB22A2B1" ID-GBIF-Occurrence="3042095310" box="[151,1159,1963,1990]" collectionCode="RGM" pageId="30" pageNumber="181" specimenCode="RGM 1310306-1310310" specimenCount="1">
and five specimens from the Kura delta (localities 25d and 25f,
<specimenCode id="DBB09ECEC85FFFA9FCCBF872FBDAA2B1" box="[865,1151,1963,1990]" collectionCode="RGM" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="30" pageNumber="181" type="Museum">RGM 13103061310310</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF69663DC85EFFA8FF3DF93EFD2FA284" ID-DOI="http://doi.org/10.5281/zenodo.4550190" ID-Zenodo-Dep="4550190" httpUri="https://zenodo.org/record/4550190/files/figure.png" pageId="31" pageNumber="182" startId="31.[151,250,1760,1784]" targetBox="[301,1286,181,1736]" targetPageId="31">
<paragraph id="8BA936B5C85EFFA8FF3DF93EFD2FA284" blockId="31.[151,1437,1760,2036]" pageId="31" pageNumber="182">
<emphasis id="B962EAA7C85EFFA8FF3DF93EFE84A38F" bold="true" box="[151,289,1760,1784]" pageId="31" pageNumber="182">FIGURE 16.</emphasis>
<taxonomicName id="4C164D36C85EFFA8FE83F93EFE0FA38F" box="[297,426,1760,1784]" class="Gastropoda" family="Hydrobiidae" genus="Ulskia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="182" phylum="Mollusca" rank="species" species="ulskii">
<emphasis id="B962EAA7C85EFFA8FE83F93EFE0FA38F" box="[297,426,1760,1784]" italics="true" pageId="31" pageNumber="182">Ulskia ulskii</emphasis>
</taxonomicName>
(Clessin &amp; Dybowski in
<bibRefCitation id="EF874B44C85EFFA8FD18F93EFCF8A38F" author="Dybowski, W." box="[690,861,1760,1784]" pageId="31" pageNumber="182" pagination="1 - 64" refId="ref30278" refString="Dybowski, W. (1887 - 1888) Die Gasteropoden-Fauna des Kaspischen Meeres. Nach der Sammlung des Akademikers Dr. K. E. v. Baer. Malakozoologische Blatter, Neue Folge, 10 (1 - 3), 1 - 64 (issue 1, 1887), 65 - 79 (issue 2, 1888), pls. 1 - 3 (issue 3, 1888)." type="journal article" year="1887">Dybowski, 1887</bibRefCitation>
) from the Caspian Sea Basin. (a) Lectotype, locality 26 (ZB-M W.Dyb. 76). (b) Paralectotype, locality 26 (ZB-M W.Dyb. 78). (c) Paralectotype, locality 26 (ZB-M W.Dyb. 78). (d) Paralectotype, locality 26 (ZB-M W.Dyb. 76). (e) Paralectotype, locality 26 (ZB-M W.Dyb. 76). (f, k) Specimen from locality 17 (ZIN, no number). (g, h) Holotype of
<taxonomicName id="4C164D36C85EFFA8FDBCF892FD05A213" authorityName="Logvinenko et Starobogatov" authorityYear="1968" box="[534,672,1868,1892]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgula" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="182" phylum="Mollusca" rank="species" species="nana">
<emphasis id="B962EAA7C85EFFA8FDBCF892FD05A213" box="[534,672,1868,1892]" italics="true" pageId="31" pageNumber="182">Pyrgula nana</emphasis>
</taxonomicName>
, locality 15 (ZIN, no number). (i, j) Specimen labelled as “
<taxonomicName id="4C164D36C85EFFA8FB50F892FEA6A2FF" class="Gastropoda" family="Hydrobiidae" genus="Caspia" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="182" phylum="Mollusca" rank="species" species="clessiniolaeformis">
<emphasis id="B962EAA7C85EFFA8FB50F892FEA6A2FF" italics="true" pageId="31" pageNumber="182">Caspia clessiniolaeformis</emphasis>
</taxonomicName>
, holotype”, locality 17 (ZIN, no number). (l) Holotype of
<taxonomicName id="4C164D36C85EFFA8FCFAF8AEFBA5A2FF" authorityName="Logvinenko et Starobogatov" authorityYear="1968" box="[848,1024,1904,1928]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgula" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="182" phylum="Mollusca" rank="species" species="behningi">
<emphasis id="B962EAA7C85EFFA8FCFAF8AEFBA5A2FF" box="[848,1024,1904,1928]" italics="true" pageId="31" pageNumber="182">Pyrgula behningi</emphasis>
</taxonomicName>
, locality 23 (ZIN #1). (m) Paratype of
<taxonomicName id="4C164D36C85EFFA8FA20F8AFFF55A2DB" authorityName="Logvinenko et Starobogatov" authorityYear="1968" class="Gastropoda" family="Hydrobiidae" genus="Pyrgula" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="182" phylum="Mollusca" rank="species" species="behningi">
<emphasis id="B962EAA7C85EFFA8FA20F8AFFF55A2DB" italics="true" pageId="31" pageNumber="182">P. behningi</emphasis>
</taxonomicName>
, locality 14 (ZIN #2). (n, o) Paratype of
<taxonomicName id="4C164D36C85EFFA8FD29F84BFD51A2DB" authorityName="Logvinenko et Starobogatov" authorityYear="1968" box="[643,756,1940,1964]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgula" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="182" phylum="Mollusca" rank="species" species="behningi">
<emphasis id="B962EAA7C85EFFA8FD29F84BFD51A2DB" box="[643,756,1940,1964]" italics="true" pageId="31" pageNumber="182">P. behningi</emphasis>
</taxonomicName>
, locality 14 (ZIN #2). (ps, u) Five specimens from the Holocene of the Kura delta (localities 25d and 25f, RGM 13103061310310). (t, vz) Three specimens from Selitrennoye (locality 1, RGM 1309810, 1309809, 1309856). Scale bar = 1 mm.
</paragraph>
</caption>
<subSubSection id="C30C653EC861FF96FF6DFF49FB43A715" lastPageId="33" lastPageNumber="184" pageId="32" pageNumber="183" type="discussion">
<paragraph id="8BA936B5C861FF97FF6DFF49FB71A46A" blockId="32.[151,1437,150,285]" pageId="32" pageNumber="183">
<emphasis id="B962EAA7C861FF97FF6DFF49FE91A5C6" bold="true" box="[199,308,151,177]" pageId="32" pageNumber="183">Remarks</emphasis>
. The characteristic features of
<taxonomicName id="4C164D36C861FF97FD0AFF49FC95A5C6" box="[672,816,151,177]" class="Gastropoda" family="Hydrobiidae" genus="Ulskia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="32" pageNumber="183" phylum="Mollusca" rank="species" species="ulskii">
<emphasis id="B962EAA7C861FF97FD0AFF49FC95A5C6" box="[672,816,151,177]" italics="true" pageId="32" pageNumber="183">Ulskia ulskii</emphasis>
</taxonomicName>
are the small (up to
<quantity id="4CEE9B50C861FF97FB81FF48FB36A5C6" box="[1067,1171,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" pageId="32" pageNumber="183" unit="mm" value="2.15">2.15 mm</quantity>
in height,
<quantity id="4CEE9B50C861FF97FABFFF49FAD9A5C6" box="[1301,1404,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="32" pageNumber="183" unit="mm" value="1.2">1.20 mm</quantity>
in width), slender and usually stepped shell with a tightly coiled aperture that rarely leaves an umbilicus (
<figureCitation id="132D2A30C861FF97FABFFF65FAC0A5A1" box="[1301,1381,187,214]" captionStart="FIGURE 16" captionStartId="31.[151,250,1760,1784]" captionTargetBox="[301,1286,181,1736]" captionTargetId="figure-17@31.[301,1286,181,1736]" captionTargetPageId="31" captionText="FIGURE 16. Ulskia ulskii (Clessin &amp; Dybowski in Dybowski, 1887) from the Caspian Sea Basin. (a) Lectotype, locality 26 (ZB-M W.Dyb. 76). (b) Paralectotype, locality 26 (ZB-M W.Dyb. 78). (c) Paralectotype, locality 26 (ZB-M W.Dyb. 78). (d) Paralectotype, locality 26 (ZB-M W.Dyb. 76). (e) Paralectotype, locality 26 (ZB-M W.Dyb. 76). (f, k) Specimen from locality 17 (ZIN, no number). (g, h) Holotype of Pyrgula nana, locality 15 (ZIN, no number). (i, j) Specimen labelled as “Caspia clessiniolaeformis, holotype”, locality 17 (ZIN, no number). (l) Holotype of Pyrgula behningi, locality 23 (ZIN #1). (m) Paratype of P. behningi, locality 14 (ZIN #2). (n, o) Paratype of P. behningi, locality 14 (ZIN #2). (ps, u) Five specimens from the Holocene of the Kura delta (localities 25d and 25f, RGM 13103061310310). (t, vz) Three specimens from Selitrennoye (locality 1, RGM 1309810, 1309809, 1309856). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4550190" httpUri="https://zenodo.org/record/4550190/files/figure.png" pageId="32" pageNumber="183">Fig. 16</figureCitation>
az), a striatedwrinkled protoconch demarcated by a distinct but not particularly bulgy growth rim (
<figureCitation id="132D2A30C861FF97FB79FF01FA86A58E" box="[1235,1315,223,249]" captionStart="FIGURE 17" captionStartId="32.[151,250,1836,1860]" captionTargetBox="[151,1436,309,1812]" captionTargetId="figure-93@32.[151,1436,309,1812]" captionTargetPageId="32" captionText="FIGURE 17. Protoconch views of Ulskia ulskii (Clessin &amp; Dybowski in Dybowski, 1887). (a) Paralectotype, locality 26 (ZB- M W.Dyb. 76). (b, j, m) Paralectotype, locality 26 (ZB-M W.Dyb. 116). (c, i) Specimen from the Holocene of the Kura delta (locality 25f, RGM 1310308). (d, k) Specimen from the same area (locality 25d, RGM 1310310). (e) Specimen from the same area (locality 25f, RGM 1310306). (f, l) Specimen from the same area (locality 25d, RGM 1310309). (g, h, n, o) Specimen from the same area (locality 25d, RGM 1310307). Scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.4550192" httpUri="https://zenodo.org/record/4550192/files/figure.png" pageId="32" pageNumber="183">Fig. 17</figureCitation>
ao) and a teleoconch microsculpture that consists of tiny, irregular, elongated beads (
<figureCitation id="132D2A30C861FF97FC77FEDDFB62A46A" box="[989,1223,259,285]" captionStart="FIGURE 17" captionStartId="32.[151,250,1836,1860]" captionTargetBox="[151,1436,309,1812]" captionTargetId="figure-93@32.[151,1436,309,1812]" captionTargetPageId="32" captionText="FIGURE 17. Protoconch views of Ulskia ulskii (Clessin &amp; Dybowski in Dybowski, 1887). (a) Paralectotype, locality 26 (ZB- M W.Dyb. 76). (b, j, m) Paralectotype, locality 26 (ZB-M W.Dyb. 116). (c, i) Specimen from the Holocene of the Kura delta (locality 25f, RGM 1310308). (d, k) Specimen from the same area (locality 25d, RGM 1310310). (e) Specimen from the same area (locality 25f, RGM 1310306). (f, l) Specimen from the same area (locality 25d, RGM 1310309). (g, h, n, o) Specimen from the same area (locality 25d, RGM 1310307). Scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.4550192" httpUri="https://zenodo.org/record/4550192/files/figure.png" pageId="32" pageNumber="183">Fig. 17b, c, f, i, k, m</figureCitation>
).
</paragraph>
<caption id="DF69663DC861FF97FF3DF8F2FC95A2A2" ID-DOI="http://doi.org/10.5281/zenodo.4550192" ID-Zenodo-Dep="4550192" httpUri="https://zenodo.org/record/4550192/files/figure.png" pageId="32" pageNumber="183" startId="32.[151,250,1836,1860]" targetBox="[151,1436,309,1812]" targetPageId="32">
<paragraph id="8BA936B5C861FF97FF3DF8F2FC95A2A2" blockId="32.[151,1436,1836,2005]" pageId="32" pageNumber="183">
<emphasis id="B962EAA7C861FF97FF3DF8F2FE85A233" bold="true" box="[151,288,1836,1860]" pageId="32" pageNumber="183">FIGURE 17.</emphasis>
Protoconch views of
<taxonomicName id="4C164D36C861FF97FE54F8F2FDDBA233" box="[510,638,1836,1860]" class="Gastropoda" family="Hydrobiidae" genus="Ulskia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="32" pageNumber="183" phylum="Mollusca" rank="species" species="ulskii">
<emphasis id="B962EAA7C861FF97FE54F8F2FDDBA233" box="[510,638,1836,1860]" italics="true" pageId="32" pageNumber="183">Ulskia ulskii</emphasis>
</taxonomicName>
(Clessin &amp; Dybowski in
<bibRefCitation id="EF874B44C861FF97FC28F8F2FB8BA233" author="Dybowski, W." box="[898,1070,1836,1860]" pageId="32" pageNumber="183" pagination="1 - 64" refId="ref30278" refString="Dybowski, W. (1887 - 1888) Die Gasteropoden-Fauna des Kaspischen Meeres. Nach der Sammlung des Akademikers Dr. K. E. v. Baer. Malakozoologische Blatter, Neue Folge, 10 (1 - 3), 1 - 64 (issue 1, 1887), 65 - 79 (issue 2, 1888), pls. 1 - 3 (issue 3, 1888)." type="journal article" year="1887">Dybowski, 1887</bibRefCitation>
). (a) Paralectotype, locality 26 (ZB- M W.Dyb. 76). (b, j, m) Paralectotype, locality 26 (ZB-M W.Dyb. 116). (c, i) Specimen from the Holocene of the Kura delta (locality 25f, RGM 1310308). (d, k) Specimen from the same area (locality 25d, RGM 1310310). (e) Specimen from the same area (locality 25f, RGM 1310306). (f, l) Specimen from the same area (locality 25d, RGM 1310309). (g, h, n, o) Specimen from the same area (locality 25d, RGM 1310307). Scale bar = 100 μm.
</paragraph>
</caption>
<paragraph id="8BA936B5C860FF96FF6DFF49FE31A412" blockId="33.[151,1437,151,717]" pageId="33" pageNumber="184">
<taxonomicName id="4C164D36C860FF96FF6DFF49FEF1A5C6" box="[199,340,151,177]" class="Gastropoda" family="Hydrobiidae" genus="Ulskia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="33" pageNumber="184" phylum="Mollusca" rank="species" species="ulskii">
<emphasis id="B962EAA7C860FF96FF6DFF49FEF1A5C6" box="[199,340,151,177]" italics="true" pageId="33" pageNumber="184">Ulskia ulskii</emphasis>
</taxonomicName>
<materialsCitation id="3B7E3CE8C860FF96FEF1FF49FE35A412" ID-GBIF-Occurrence="3042095343" country="Russia" location="Pleistocene" municipality="These" pageId="33" pageNumber="184" specimenCount="1" typeStatus="lectotype">
is a highly variable species with respect to shell shape.
<bibRefCitation id="EF874B44C860FF96FC68FF49FB1AA5C6" author="Neubauer, T. A. &amp; van de Velde, S. &amp; Yanina, T. A. &amp; Wesselingh, F. P." box="[962,1215,151,177]" pageId="33" pageNumber="184" pagination="43 - 103" refId="ref32549" refString="Neubauer, T. A., van de Velde, S., Yanina, T. A. &amp; Wesselingh, F. P. (2018) A late Pleistocene gastropod fauna from the northern Caspian Sea with implications for Pontocaspian gastropod taxonomy. ZooKeys, 770, 43 - 103. https: // doi. org / 10.3897 / zookeys. 770.25365" type="journal article" year="2018">
Neubauer
<emphasis id="B962EAA7C860FF96FB9CFF49FBC2A5C6" box="[1078,1127,151,177]" italics="true" pageId="33" pageNumber="184">et al</emphasis>
. (2018)
</bibRefCitation>
provided a detailed description for material from the
<location id="8EC9606EC860FF96FDB0FF65FD3FA5A2" LSID="urn:lsid:plazi:treatment:03BF87A3C85FFF96FF3DFC28FEB3A7BA:8EC9606EC860FF96FDB0FF65FD3FA5A2" box="[538,666,187,213]" country="Russia" municipality="These" name="Pleistocene" pageId="33" pageNumber="184">Pleistocene</location>
of
<collectingCountry id="F3017625C860FF96FD6FFF65FCB5A5A2" box="[709,784,187,213]" name="Russia" pageId="33" pageNumber="184">Russia</collectingCountry>
(
<figureCitation id="132D2A30C860FF96FC88FF65FCDAA5A2" box="[802,895,187,213]" captionStart="FIGURE 16" captionStartId="31.[151,250,1760,1784]" captionTargetBox="[301,1286,181,1736]" captionTargetId="figure-17@31.[301,1286,181,1736]" captionTargetPageId="31" captionText="FIGURE 16. Ulskia ulskii (Clessin &amp; Dybowski in Dybowski, 1887) from the Caspian Sea Basin. (a) Lectotype, locality 26 (ZB-M W.Dyb. 76). (b) Paralectotype, locality 26 (ZB-M W.Dyb. 78). (c) Paralectotype, locality 26 (ZB-M W.Dyb. 78). (d) Paralectotype, locality 26 (ZB-M W.Dyb. 76). (e) Paralectotype, locality 26 (ZB-M W.Dyb. 76). (f, k) Specimen from locality 17 (ZIN, no number). (g, h) Holotype of Pyrgula nana, locality 15 (ZIN, no number). (i, j) Specimen labelled as “Caspia clessiniolaeformis, holotype”, locality 17 (ZIN, no number). (l) Holotype of Pyrgula behningi, locality 23 (ZIN #1). (m) Paratype of P. behningi, locality 14 (ZIN #2). (n, o) Paratype of P. behningi, locality 14 (ZIN #2). (ps, u) Five specimens from the Holocene of the Kura delta (localities 25d and 25f, RGM 13103061310310). (t, vz) Three specimens from Selitrennoye (locality 1, RGM 1309810, 1309809, 1309856). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4550190" httpUri="https://zenodo.org/record/4550190/files/figure.png" pageId="33" pageNumber="184">
Fig.
<quantity id="4CEE9B50C860FF96FCF3FF65FCDAA5A2" box="[857,895,187,213]" metricMagnitude="4" metricUnit="kg" metricValue="1.6" pageId="33" pageNumber="184" unit="t" value="16.0">16t</quantity>
</figureCitation>
, vz).
<collectingMunicipality id="6BCDACCFC860FF96FC7AFF65FBB6A5A2" box="[976,1043,187,213]" pageId="33" pageNumber="184">These</collectingMunicipality>
specimens are more elongate and slightly larger but otherwise fit well to the
<typeStatus id="54AD8817C860FF96FD2DFF01FD4BA58E" box="[647,750,223,249]" pageId="33" pageNumber="184" type="lectotype">lectotype</typeStatus>
series of
<taxonomicName id="4C164D36C860FF96FCCAFF01FC67A58E" box="[864,962,223,249]" class="Gastropoda" family="Hydrobiidae" genus="Ulskia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="33" pageNumber="184" phylum="Mollusca" rank="species" species="ulskii">
<emphasis id="B962EAA7C860FF96FCCAFF01FC67A58E" box="[864,962,223,249]" italics="true" pageId="33" pageNumber="184">U. ulskii</emphasis>
</taxonomicName>
. Similarly, Holocene specimens from the Kura delta tend to be more slender and have a less stepped outline (
<figureCitation id="132D2A30C860FF96FC24FEDDFC7BA46A" box="[910,990,259,285]" captionStart="FIGURE 16" captionStartId="31.[151,250,1760,1784]" captionTargetBox="[301,1286,181,1736]" captionTargetId="figure-17@31.[301,1286,181,1736]" captionTargetPageId="31" captionText="FIGURE 16. Ulskia ulskii (Clessin &amp; Dybowski in Dybowski, 1887) from the Caspian Sea Basin. (a) Lectotype, locality 26 (ZB-M W.Dyb. 76). (b) Paralectotype, locality 26 (ZB-M W.Dyb. 78). (c) Paralectotype, locality 26 (ZB-M W.Dyb. 78). (d) Paralectotype, locality 26 (ZB-M W.Dyb. 76). (e) Paralectotype, locality 26 (ZB-M W.Dyb. 76). (f, k) Specimen from locality 17 (ZIN, no number). (g, h) Holotype of Pyrgula nana, locality 15 (ZIN, no number). (i, j) Specimen labelled as “Caspia clessiniolaeformis, holotype”, locality 17 (ZIN, no number). (l) Holotype of Pyrgula behningi, locality 23 (ZIN #1). (m) Paratype of P. behningi, locality 14 (ZIN #2). (n, o) Paratype of P. behningi, locality 14 (ZIN #2). (ps, u) Five specimens from the Holocene of the Kura delta (localities 25d and 25f, RGM 13103061310310). (t, vz) Three specimens from Selitrennoye (locality 1, RGM 1309810, 1309809, 1309856). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4550190" httpUri="https://zenodo.org/record/4550190/files/figure.png" pageId="33" pageNumber="184">Fig. 16</figureCitation>
ps, u), but the morphological variation is fluent and does not allow establishing clear species boundaries. Rather we consider all these differences to reflect intraspecific variation
</materialsCitation>
.
</paragraph>
<paragraph id="8BA936B5C860FF96FF6DFEB1FCE4A482" blockId="33.[151,1437,151,717]" pageId="33" pageNumber="184">
The range of variation also covers the
<typeStatus id="54AD8817C860FF96FDC6FEB1FD6BA4FE" box="[620,718,367,393]" pageId="33" pageNumber="184" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C164D36C860FF96FD5AFEB1FAA8A4FE" authority="Logvinenko &amp; Starobogatov, 1969" authorityName="Logvinenko &amp; Starobogatov" authorityYear="1969" box="[752,1293,367,393]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgula" kingdom="Animalia" order="Littorinimorpha" pageId="33" pageNumber="184" phylum="Mollusca" rank="species" species="nana">
<emphasis id="B962EAA7C860FF96FD5AFEB1FC22A4FF" box="[752,903,367,393]" italics="true" pageId="33" pageNumber="184">Pyrgula nana</emphasis>
<bibRefCitation id="EF874B44C860FF96FC27FEB1FAA8A4FE" author="Logvinenko, B. M. &amp; Starobogatov, Ya. I." box="[909,1293,367,393]" pageId="33" pageNumber="184" pagination="308 - 385" refId="ref32049" refString="Logvinenko, B. M. &amp; Starobogatov, Ya. I. (1969 [&quot; 1968 &quot;]) Mollusca. In: Birshtein, Y. A., Vinogradov, L. G., Kondakov, N. N., Kuhn, M. S., Astakhova, T. V. &amp; Romanova, N. N. (Eds.), Atlas bespozvonochnykh Kaspiyskogo morya. Pishchevaya Promyshlennost (Vsesoyuznyi Nauchno-issledovatel'skii Institut Morskogo Rybnogo Khozyaistva i Okeanografii), Moskva, pp. 308 - 385." type="book chapter" year="1969">Logvinenko &amp; Starobogatov, 1969</bibRefCitation>
</taxonomicName>
(
<figureCitation id="132D2A30C860FF96FAB1FEB1FA31A4FD" box="[1307,1428,367,394]" captionStart="FIGURE 16" captionStartId="31.[151,250,1760,1784]" captionTargetBox="[301,1286,181,1736]" captionTargetId="figure-17@31.[301,1286,181,1736]" captionTargetPageId="31" captionText="FIGURE 16. Ulskia ulskii (Clessin &amp; Dybowski in Dybowski, 1887) from the Caspian Sea Basin. (a) Lectotype, locality 26 (ZB-M W.Dyb. 76). (b) Paralectotype, locality 26 (ZB-M W.Dyb. 78). (c) Paralectotype, locality 26 (ZB-M W.Dyb. 78). (d) Paralectotype, locality 26 (ZB-M W.Dyb. 76). (e) Paralectotype, locality 26 (ZB-M W.Dyb. 76). (f, k) Specimen from locality 17 (ZIN, no number). (g, h) Holotype of Pyrgula nana, locality 15 (ZIN, no number). (i, j) Specimen labelled as “Caspia clessiniolaeformis, holotype”, locality 17 (ZIN, no number). (l) Holotype of Pyrgula behningi, locality 23 (ZIN #1). (m) Paratype of P. behningi, locality 14 (ZIN #2). (n, o) Paratype of P. behningi, locality 14 (ZIN #2). (ps, u) Five specimens from the Holocene of the Kura delta (localities 25d and 25f, RGM 13103061310310). (t, vz) Three specimens from Selitrennoye (locality 1, RGM 1309810, 1309809, 1309856). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4550190" httpUri="https://zenodo.org/record/4550190/files/figure.png" pageId="33" pageNumber="184">
Fig.
<quantity id="4CEE9B50C860FF96FAE4FEB1FADFA4FE" box="[1358,1402,367,393]" metricMagnitude="-2" metricUnit="kg" metricValue="1.6" pageId="33" pageNumber="184" unit="g" value="16.0">16g</quantity>
, h
</figureCitation>
) and
<taxonomicName id="4C164D36C860FF96FF6CFE4DFD68A4DA" authority="Logvinenko &amp; Starobogatov, 1969" authorityName="Logvinenko &amp; Starobogatov" authorityYear="1969" box="[198,717,403,429]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgula" kingdom="Animalia" order="Littorinimorpha" pageId="33" pageNumber="184" phylum="Mollusca" rank="species" species="behningi">
<emphasis id="B962EAA7C860FF96FF6CFE4DFEE6A4DA" box="[198,323,403,429]" italics="true" pageId="33" pageNumber="184">P. behningi</emphasis>
<bibRefCitation id="EF874B44C860FF96FEE0FE4DFD68A4DA" author="Logvinenko, B. M. &amp; Starobogatov, Ya. I." box="[330,717,403,429]" pageId="33" pageNumber="184" pagination="308 - 385" refId="ref32049" refString="Logvinenko, B. M. &amp; Starobogatov, Ya. I. (1969 [&quot; 1968 &quot;]) Mollusca. In: Birshtein, Y. A., Vinogradov, L. G., Kondakov, N. N., Kuhn, M. S., Astakhova, T. V. &amp; Romanova, N. N. (Eds.), Atlas bespozvonochnykh Kaspiyskogo morya. Pishchevaya Promyshlennost (Vsesoyuznyi Nauchno-issledovatel'skii Institut Morskogo Rybnogo Khozyaistva i Okeanografii), Moskva, pp. 308 - 385." type="book chapter" year="1969">Logvinenko &amp; Starobogatov, 1969</bibRefCitation>
</taxonomicName>
(
<figureCitation id="132D2A30C860FF96FD71FE4DFC8EA4D9" box="[731,811,403,430]" captionStart="FIGURE 16" captionStartId="31.[151,250,1760,1784]" captionTargetBox="[301,1286,181,1736]" captionTargetId="figure-17@31.[301,1286,181,1736]" captionTargetPageId="31" captionText="FIGURE 16. Ulskia ulskii (Clessin &amp; Dybowski in Dybowski, 1887) from the Caspian Sea Basin. (a) Lectotype, locality 26 (ZB-M W.Dyb. 76). (b) Paralectotype, locality 26 (ZB-M W.Dyb. 78). (c) Paralectotype, locality 26 (ZB-M W.Dyb. 78). (d) Paralectotype, locality 26 (ZB-M W.Dyb. 76). (e) Paralectotype, locality 26 (ZB-M W.Dyb. 76). (f, k) Specimen from locality 17 (ZIN, no number). (g, h) Holotype of Pyrgula nana, locality 15 (ZIN, no number). (i, j) Specimen labelled as “Caspia clessiniolaeformis, holotype”, locality 17 (ZIN, no number). (l) Holotype of Pyrgula behningi, locality 23 (ZIN #1). (m) Paratype of P. behningi, locality 14 (ZIN #2). (n, o) Paratype of P. behningi, locality 14 (ZIN #2). (ps, u) Five specimens from the Holocene of the Kura delta (localities 25d and 25f, RGM 13103061310310). (t, vz) Three specimens from Selitrennoye (locality 1, RGM 1309810, 1309809, 1309856). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4550190" httpUri="https://zenodo.org/record/4550190/files/figure.png" pageId="33" pageNumber="184">Fig. 16</figureCitation>
lo), which closely resemble some of the
<typeStatus id="54AD8817C860FF96FB50FE4DFA39A4DA" box="[1274,1436,403,429]" pageId="33" pageNumber="184" type="paralectotype">paralectotypes</typeStatus>
of
<taxonomicName id="4C164D36C860FF96FF1CFE69FEB0A4A6" box="[182,277,439,465]" class="Gastropoda" family="Hydrobiidae" genus="Ulskia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="33" pageNumber="184" phylum="Mollusca" rank="species" species="ulskii">
<emphasis id="B962EAA7C860FF96FF1CFE69FEB0A4A6" box="[182,277,439,465]" italics="true" pageId="33" pageNumber="184">U. ulskii</emphasis>
</taxonomicName>
. We thus follow the previous opinion of
<bibRefCitation id="EF874B44C860FF96FD4BFE69FC45A4A6" author="Neubauer, T. A. &amp; van de Velde, S. &amp; Yanina, T. A. &amp; Wesselingh, F. P." box="[737,992,439,465]" pageId="33" pageNumber="184" pagination="43 - 103" refId="ref32549" refString="Neubauer, T. A., van de Velde, S., Yanina, T. A. &amp; Wesselingh, F. P. (2018) A late Pleistocene gastropod fauna from the northern Caspian Sea with implications for Pontocaspian gastropod taxonomy. ZooKeys, 770, 43 - 103. https: // doi. org / 10.3897 / zookeys. 770.25365" type="journal article" year="2018">
Neubauer
<emphasis id="B962EAA7C860FF96FCFFFE69FC22A4A6" box="[853,903,439,465]" italics="true" pageId="33" pageNumber="184">et al</emphasis>
. (2018)
</bibRefCitation>
and consider
<taxonomicName id="4C164D36C860FF96FBD7FE69FB75A4A7" authorityName="Logvinenko &amp; Starobogatov" authorityYear="1969" box="[1149,1232,439,464]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgula" kingdom="Animalia" order="Littorinimorpha" pageId="33" pageNumber="184" phylum="Mollusca" rank="species" species="nana">
<emphasis id="B962EAA7C860FF96FBD7FE69FB75A4A7" box="[1149,1232,439,464]" italics="true" pageId="33" pageNumber="184">P. nana</emphasis>
</taxonomicName>
a junior synonym of
<taxonomicName id="4C164D36C860FF96FF1CFE05FEB0A482" box="[182,277,475,501]" class="Gastropoda" family="Hydrobiidae" genus="Ulskia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="33" pageNumber="184" phylum="Mollusca" rank="species" species="ulskii">
<emphasis id="B962EAA7C860FF96FF1CFE05FEB0A482" box="[182,277,475,501]" italics="true" pageId="33" pageNumber="184">U. ulskii</emphasis>
</taxonomicName>
;
<taxonomicName id="4C164D36C860FF96FE89FE05FE40A482" authorityName="Logvinenko &amp; Starobogatov" authorityYear="1969" box="[291,485,475,501]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgula" kingdom="Animalia" order="Littorinimorpha" pageId="33" pageNumber="184" phylum="Mollusca" rank="species" species="behningi">
<emphasis id="B962EAA7C860FF96FE89FE05FE40A482" box="[291,485,475,501]" italics="true" pageId="33" pageNumber="184">Pyrgula behningi</emphasis>
</taxonomicName>
is a new synonym of
<taxonomicName id="4C164D36C860FF96FD71FE05FC9FA482" box="[731,826,475,501]" class="Gastropoda" family="Hydrobiidae" genus="Ulskia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="33" pageNumber="184" phylum="Mollusca" rank="species" species="ulskii">
<emphasis id="B962EAA7C860FF96FD71FE05FC9FA482" box="[731,826,475,501]" italics="true" pageId="33" pageNumber="184">U. ulskii</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA936B5C860FF96FF6DFE21FB43A715" blockId="33.[151,1437,151,717]" pageId="33" pageNumber="184">
<materialsCitation id="3B7E3CE8C860FF96FF6DFE21FB47A715" ID-GBIF-Occurrence="3042095325" collectionCode="ZIN" pageId="33" pageNumber="184" specimenCount="2" typeStatus="holotype">
Furthermore, we found in the
<collectionCode id="ED07AE70C860FF96FDB3FE21FDEDA76E" box="[537,584,511,537]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="33" pageNumber="184" type="Museum">ZIN</collectionCode>
collection of Starobogatov material labelled with the unavailable collection name “
<taxonomicName id="4C164D36C860FF96FF43FDFDFDA8A74A" box="[233,525,547,573]" class="Gastropoda" family="Hydrobiidae" genus="Caspia" kingdom="Animalia" order="Littorinimorpha" pageId="33" pageNumber="184" phylum="Mollusca" rank="species" species="clessiniolaeformis">
<emphasis id="B962EAA7C860FF96FF43FDFDFDA8A74A" box="[233,525,547,573]" italics="true" pageId="33" pageNumber="184">Caspia clessiniolaeformis</emphasis>
</taxonomicName>
”. The “
<typeStatus id="54AD8817C860FF96FDCCFDFDFD68A74A" box="[614,717,547,573]" pageId="33" pageNumber="184" type="holotype">holotype</typeStatus>
” and 6 “
<typeStatus id="54AD8817C860FF96FC9BFDFDFC07A74A" box="[817,930,547,573]" pageId="33" pageNumber="184" type="paratype">paratypes</typeStatus>
” of “
<taxonomicName id="4C164D36C860FF96FC4AFDFDFAA1A74A" box="[992,1284,547,573]" class="Gastropoda" family="Hydrobiidae" genus="Caspia" kingdom="Animalia" order="Littorinimorpha" pageId="33" pageNumber="184" phylum="Mollusca" rank="species" species="clessiniolaeformis">
<emphasis id="B962EAA7C860FF96FC4AFDFDFAA1A74A" box="[992,1284,547,573]" italics="true" pageId="33" pageNumber="184">Caspia clessiniolaeformis</emphasis>
</taxonomicName>
” collected in the eastern part of the Middle Caspian Sea fall in the morphological range of
<taxonomicName id="4C164D36C860FF96FC53FD99FBFCA716" box="[1017,1113,583,609]" class="Gastropoda" family="Hydrobiidae" genus="Ulskia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="33" pageNumber="184" phylum="Mollusca" rank="species" species="ulskii">
<emphasis id="B962EAA7C860FF96FC53FD99FBFCA716" box="[1017,1113,583,609]" italics="true" pageId="33" pageNumber="184">U. ulskii</emphasis>
</taxonomicName>
(
<figureCitation id="132D2A30C860FF96FBC2FD99FB7DA715" box="[1128,1240,583,610]" captionStart="FIGURE 16" captionStartId="31.[151,250,1760,1784]" captionTargetBox="[301,1286,181,1736]" captionTargetId="figure-17@31.[301,1286,181,1736]" captionTargetPageId="31" captionText="FIGURE 16. Ulskia ulskii (Clessin &amp; Dybowski in Dybowski, 1887) from the Caspian Sea Basin. (a) Lectotype, locality 26 (ZB-M W.Dyb. 76). (b) Paralectotype, locality 26 (ZB-M W.Dyb. 78). (c) Paralectotype, locality 26 (ZB-M W.Dyb. 78). (d) Paralectotype, locality 26 (ZB-M W.Dyb. 76). (e) Paralectotype, locality 26 (ZB-M W.Dyb. 76). (f, k) Specimen from locality 17 (ZIN, no number). (g, h) Holotype of Pyrgula nana, locality 15 (ZIN, no number). (i, j) Specimen labelled as “Caspia clessiniolaeformis, holotype”, locality 17 (ZIN, no number). (l) Holotype of Pyrgula behningi, locality 23 (ZIN #1). (m) Paratype of P. behningi, locality 14 (ZIN #2). (n, o) Paratype of P. behningi, locality 14 (ZIN #2). (ps, u) Five specimens from the Holocene of the Kura delta (localities 25d and 25f, RGM 13103061310310). (t, vz) Three specimens from Selitrennoye (locality 1, RGM 1309810, 1309809, 1309856). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4550190" httpUri="https://zenodo.org/record/4550190/files/figure.png" pageId="33" pageNumber="184">Fig. 16i, j</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30C653EC860FF96FF6DFDB4FEB3A7BA" pageId="33" pageNumber="184" type="distribution">
<paragraph id="8BA936B5C860FF96FF6DFDB4FEB3A7BA" blockId="33.[151,1437,151,717]" pageId="33" pageNumber="184">
<emphasis id="B962EAA7C860FF96FF6DFDB4FEC4A7F3" bold="true" box="[199,353,618,644]" pageId="33" pageNumber="184">Distribution.</emphasis>
Endemic to the Caspian Sea, reported from water depths between 45 and
<quantity id="4CEE9B50C860FF96FB0AFDB5FB4DA7F2" box="[1184,1256,619,645]" metricMagnitude="2" metricUnit="m" metricValue="1.7" pageId="33" pageNumber="184" unit="m" value="170.0">170 m</quantity>
(
<bibRefCitation id="EF874B44C860FF96FB5CFDB5FECAA7DE" author="Logvinenko, B. M. &amp; Starobogatov, Ya. I." pageId="33" pageNumber="184" pagination="308 - 385" refId="ref32049" refString="Logvinenko, B. M. &amp; Starobogatov, Ya. I. (1969 [&quot; 1968 &quot;]) Mollusca. In: Birshtein, Y. A., Vinogradov, L. G., Kondakov, N. N., Kuhn, M. S., Astakhova, T. V. &amp; Romanova, N. N. (Eds.), Atlas bespozvonochnykh Kaspiyskogo morya. Pishchevaya Promyshlennost (Vsesoyuznyi Nauchno-issledovatel'skii Institut Morskogo Rybnogo Khozyaistva i Okeanografii), Moskva, pp. 308 - 385." type="book chapter" year="1969">Logvinenko &amp; Starobogatov 1969</bibRefCitation>
; data of ZIN catalogue).
<bibRefCitation id="EF874B44C860FF96FD20FD51FCE6A7DD" author="Tarasov, A. G." box="[650,835,655,682]" pageId="33" pageNumber="184" pagination="117 - 126" refId="ref33795" refString="Tarasov, A. G. (1996 b) Novyye dannyye o kaspiyskikh glubokovodnykh Gastropoda. Ruthenica, 6 (2), 117 - 126." type="journal article" year="1996">Tarasov (1996b)</bibRefCitation>
mentioned the species from
<quantity id="4CEE9B50C860FF96FB23FD51FB74A7DE" box="[1161,1233,655,681]" metricMagnitude="2" metricUnit="m" metricValue="4.779999999999999" pageId="33" pageNumber="184" unit="m" value="478.0">478 m</quantity>
in the South Caspian Basin.
</paragraph>
</subSubSection>
</treatment>
</document>