treatments-xml/data/03/BF/87/03BF87A3C853FFA3FF3DF9F4FE29A20A.xml
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<mods:title id="1D1A8EC179DA92ED65219894EB845AC8">A revision of the Pontocaspian gastropods of the subfamily Caspiinae (Caenogastropoda: Hydrobiidae)</mods:title>
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<mods:affiliation id="3CAECF3C0E0B9961BC60DE32F4C3D73C">Department of Invertebrate Fauna and Systematics, Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, B. Khmelnytsky Str. 15, 01030 Kiev, Ukraine.</mods:affiliation>
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<mods:namePart id="6485F6194F9A163CF53F047FFC148C4A">Neubauer, Thomas A.</mods:namePart>
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<mods:affiliation id="C94DFB6A6E4C9CC993CC652B7BC4F258">Department of Animal Ecology and Systematics, Justus Liebig University, Heinrich-Buff-Ring 26 - 32 IFZ, 35392 Giessen, Germany. tneub @ zo. jlug. de; https: // orcid. org / 0000 - 0002 - 1398 - 9941 &amp; Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, The Netherlands.</mods:affiliation>
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<mods:namePart id="9BB1881514CA1E7B0705723F45866B09">Anistratenko, Olga Yu.</mods:namePart>
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<mods:affiliation id="696F03AB75BB91D30DD6E267294E6A6C">Department of Invertebrate Fauna and Systematics, Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, B. Khmelnytsky Str. 15, 01030 Kiev, Ukraine. &amp; Department of Cainozoic Deposits, Institute of Geological Sciences, National Academy of Sciences of Ukraine, O. Gontchar Str. 55 b, 01054 Kiev, Ukraine.</mods:affiliation>
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<mods:namePart id="05EC9399A782C6EA3224846723A18C46">Kijashko, Pavel V.</mods:namePart>
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<mods:affiliation id="F6AF78F247549F90A6F26AB1224E0C26">Zoological Institute, Russian Academy of Sciences, Universitetskaya Emb. 1, 199034 Saint Petersburg, Russia.</mods:affiliation>
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<mods:affiliation id="641E222C393702662870B73487400C2E">Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, The Netherlands. &amp; Department of Earth Sciences, Utrecht University, Budapestlaan 17, 3584 CD Utrecht, The Netherlands</mods:affiliation>
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<paragraph id="8BA936B5C853FFA5FF3DF9F4FD1DA333" blockId="18.[151,696,1578,1641]" box="[151,696,1578,1605]" pageId="18" pageNumber="169">
<heading id="D0E181D9C853FFA5FF3DF9F4FD1DA333" bold="true" box="[151,696,1578,1605]" fontSize="11" level="1" pageId="18" pageNumber="169" reason="1">
<taxonomicName id="4C164D36C853FFA5FF3DF9F4FD1DA333" ID-CoL="873PD" authority="(Tadjalli-Pour, 1977)" baseAuthorityName="Tadjalli-Pour" baseAuthorityYear="1977" box="[151,696,1578,1605]" class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="18" pageNumber="169" phylum="Mollusca" rank="species" species="gaillardi">
<emphasis id="B962EAA7C853FFA5FF3DF9F4FD1DA333" bold="true" box="[151,696,1578,1605]" pageId="18" pageNumber="169">
<emphasis id="B962EAA7C853FFA5FF3DF9F4FE0AA332" bold="true" box="[151,431,1578,1605]" italics="true" pageId="18" pageNumber="169">Clathrocaspia gaillardi</emphasis>
(
<bibRefCitation id="EF874B44C853FFA5FE14F9F4FD15A333" author="Tadjalli-Pour, M." box="[446,688,1578,1604]" pageId="18" pageNumber="169" pagination="87 - 117" refId="ref33728" refString="Tadjalli-Pour, M. (1977) Les Mollusques marins des cotes Iraniennes de la Mer Caspienne (Astara-Hachtpar). Journal de Conchyliologie, 114 (3 - 4), 87 - 117." type="journal article" year="1977">Tadjalli-Pour, 1977</bibRefCitation>
)
</emphasis>
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<figureCitation id="132D2A30C853FFA5FF3DF990FF7BA31E" box="[151,222,1614,1641]" captionStart="FIGURE 9" captionStartId="19.[151,250,1789,1813]" captionTargetBox="[151,1436,273,1766]" captionTargetId="figure-68@19.[151,1436,273,1766]" captionTargetPageId="19" captionText="FIGURE 9. Clathrocaspia gaillardi (Tadjalli-Pour, 1977) from the Middle and South Caspian basins. (ae) Five specimens from locality 13 (ZIN, no number). (fi) Three specimens labelled as “Caspia subgmelini” from locality 18 (ZIN, no number). (jn) Five specimens from Holocene deposits of the Kura delta (locality 25d, RGM 13102991310303). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4550172" httpUri="https://zenodo.org/record/4550172/files/figure.png" pageId="18" pageNumber="169">Figs 9</figureCitation>
,
<figureCitation id="132D2A30C853FFA5FF40F991FEA3A31E" box="[234,262,1615,1641]" captionStart="FIGURE 10" captionStartId="20.[151,250,1460,1484]" captionTargetBox="[151,1436,525,1437]" captionTargetId="figure-190@20.[151,1436,525,1437]" captionTargetPageId="20" captionText="FIGURE 10. Protoconch views of Clathrocaspia gaillardi (Tadjalli-Pour, 1977) from Holocene deposits of the Kura delta. (a, g) RGM 1310299 (locality 25d). (b, h) RGM 1310300 (locality 25d). (c, f) RGM 1310304 (locality 25f). (d) RGM 1310302 (locality 25d). (e) RGM 1310301 (locality 25d). (i) RGM 1310303 (locality 25d). Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.4550174" httpUri="https://zenodo.org/record/4550174/files/figure.png" pageId="18" pageNumber="169">10</figureCitation>
</paragraph>
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<taxonomicName id="4C164D36C853FFA5FF3DF94DFE00A3DC" baseAuthorityName="Tadjalli-Pour" baseAuthorityYear="1977" box="[151,421,1683,1707]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgula" kingdom="Animalia" order="Littorinimorpha" pageId="18" pageNumber="169" phylum="Mollusca" rank="species" species="gaillardi" subGenus="Caspia">
<emphasis id="B962EAA7C853FFA5FF3DF94DFF4DA3DC" box="[151,232,1683,1707]" italics="true" pageId="18" pageNumber="169">Pyrgula</emphasis>
(
<emphasis id="B962EAA7C853FFA5FF5DF94DFE9AA3DD" box="[247,319,1683,1706]" italics="true" pageId="18" pageNumber="169">Caspia</emphasis>
)
<emphasis id="B962EAA7C853FFA5FEE7F94DFE00A3DC" box="[333,421,1683,1707]" italics="true" pageId="18" pageNumber="169">gaillardi</emphasis>
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, N. Espece—Tadjalli-Pour 1977: 107, fig. 8.
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<emphasis id="B962EAA7C853FFA5FF3DF96CFF0AA3BE" box="[151,175,1714,1737]" italics="true" pageId="18" pageNumber="169">C.</emphasis>
[
<emphasis id="B962EAA7C853FFA5FF1DF96CFF4BA3BE" box="[183,238,1714,1737]" italics="true" pageId="18" pageNumber="169">aspia</emphasis>
] (
<emphasis id="B962EAA7C853FFA5FEAFF96CFE81A3BD" box="[261,292,1714,1738]" italics="true" pageId="18" pageNumber="169">Cl.</emphasis>
[
<emphasis id="B962EAA7C853FFA5FE86F96CFE01A3BD" box="[300,420,1714,1738]" italics="true" pageId="18" pageNumber="169">athrocaspia</emphasis>
])
<taxonomicName id="4C164D36C853FFA5FE11F96CFD4FA3BD" authority="(Tadjalli-Pour, 1977)" baseAuthorityName="Tadjalli-Pour" baseAuthorityYear="1977" box="[443,746,1714,1738]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgula" kingdom="Animalia" order="Littorinimorpha" pageId="18" pageNumber="169" phylum="Mollusca" rank="species" species="gaillardi">
<emphasis id="B962EAA7C853FFA5FE11F96CFDB6A3BD" box="[443,531,1714,1738]" italics="true" pageId="18" pageNumber="169">gaillardi</emphasis>
(
<bibRefCitation id="EF874B44C853FFA5FD8AF96CFD46A3BD" author="Tadjalli-Pour, M." box="[544,739,1714,1738]" pageId="18" pageNumber="169" pagination="87 - 117" refId="ref33728" refString="Tadjalli-Pour, M. (1977) Les Mollusques marins des cotes Iraniennes de la Mer Caspienne (Astara-Hachtpar). Journal de Conchyliologie, 114 (3 - 4), 87 - 117." type="journal article" year="1977">Tadjalli-Pour, 1977</bibRefCitation>
)
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<bibRefCitation id="EF874B44C853FFA5FCAFF96CFB6FA3BE" author="Sitnikova, T. Ya. &amp; Starobogatov, Ya. I." box="[773,1226,1713,1738]" pageId="18" pageNumber="169" pagination="1357 - 1367" refId="ref33455" refString="Sitnikova, T. Ya. &amp; Starobogatov, Ya. I. (1998) Polovaya sistema i radula kaspiyskikh Pyrgulidae (podsemeystva Turricaspiinae i Caspiinae, Gastropoda, Pectinibranchia). Zoologicheskiy Zhurnal, 77 (12), 1357 - 1367." type="journal article" year="1998">Sitnikova &amp; Starobogatov 1998: 13631364</bibRefCitation>
, fig. 1.13.
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<taxonomicName id="4C164D36C853FFA5FF3DF90FFDB0A39F" ID-CoL="84W5P" authority="(Tadjalli-Pour, 1977)" baseAuthorityName="Tadjalli-Pour" baseAuthorityYear="1977" box="[151,533,1744,1768]" class="Gastropoda" family="Hydrobiidae" genus="Caspia" kingdom="Animalia" order="Littorinimorpha" pageId="18" pageNumber="169" phylum="Mollusca" rank="species" species="gaillardi">
<emphasis id="B962EAA7C853FFA5FF3DF90FFE98A39F" box="[151,317,1744,1768]" italics="true" pageId="18" pageNumber="169">Caspia gaillardi</emphasis>
(
<bibRefCitation id="EF874B44C853FFA5FEE0F90EFDABA39F" author="Tadjalli-Pour, M." box="[330,526,1744,1768]" pageId="18" pageNumber="169" pagination="87 - 117" refId="ref33728" refString="Tadjalli-Pour, M. (1977) Les Mollusques marins des cotes Iraniennes de la Mer Caspienne (Astara-Hachtpar). Journal de Conchyliologie, 114 (3 - 4), 87 - 117." type="journal article" year="1977">Tadjalli-Pour, 1977</bibRefCitation>
)
</taxonomicName>
<treatmentCitation id="0AB710A4C853FFA5FD85F90EFCE1A39F" author="Kantor, Yu. I. &amp; Sysoev, A. V." box="[559,836,1744,1768]" page="87" pageId="18" pageNumber="169" year="2006">
<bibRefCitation id="EF874B44C853FFA5FD85F90EFCE1A39F" author="Kantor, Yu. I. &amp; Sysoev, A. V." box="[559,836,1744,1768]" pageId="18" pageNumber="169" refId="ref31309" refString="Kantor, Yu. I. &amp; Sysoev, A. V. (2006) Morskiye i solonovatovodnyye bryukhonogiye mollyuski Rossii i sopredel'nykh stran: illyustrirovannyy katalog. KMK Scientific Press, Moscow, 372 pp., 140 pls pp." type="book" year="2006">Kantor &amp; Sysoev 2006: 87</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
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<emphasis id="B962EAA7C853FFA5FF3DF8C9FEE3A247" bold="true" box="[151,326,1814,1841]" pageId="18" pageNumber="169">Type material.</emphasis>
<materialsCitation id="3B7E3CE8C853FFA5FEE4F8C8FE51A20F" ID-GBIF-Occurrence="3042095305" collectionCode="ZIN" location="One" municipality="Faculte de Medecine Veterinaire de l'Universite" pageId="18" pageNumber="169" specimenCount="3" stateProvince="Tehran" typeStatus="holotype">
<typeStatus id="54AD8817C853FFA5FEE4F8C8FE1CA247" box="[334,441,1814,1840]" pageId="18" pageNumber="169" type="holotype">Holotype</typeStatus>
,
<collectingMunicipality id="6BCDACCFC853FFA5FE6FF8C8FC40A247" box="[453,997,1814,1840]" pageId="18" pageNumber="169">Faculté de Médecine Vétérinaire de lUniversité</collectingMunicipality>
de
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; not studied.
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<typeStatus id="54AD8817C853FFA5FA96F8C9FA39A246" box="[1340,1436,1815,1841]" pageId="18" pageNumber="169" type="paratype">paratype</typeStatus>
should be in
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(“Un spécimen
<typeStatus id="54AD8817C853FFA5FDAFF8E5FDC0A222" box="[517,613,1851,1877]" pageId="18" pageNumber="169" type="paratype">paratype</typeStatus>
a été déposé au Musée Malacologique de
<collectingRegion id="49D2F857C853FFA5FB9DF8E4FB15A223" box="[1079,1200,1850,1876]" country="Russia" name="Leningrad" pageId="18" pageNumber="169">Leningrad</collectingRegion>
”;
<bibRefCitation id="EF874B44C853FFA5FB69F8E4FF61A20E" author="Tadjalli-Pour, M." pageId="18" pageNumber="169" pagination="87 - 117" refId="ref33728" refString="Tadjalli-Pour, M. (1977) Les Mollusques marins des cotes Iraniennes de la Mer Caspienne (Astara-Hachtpar). Journal de Conchyliologie, 114 (3 - 4), 87 - 117." type="journal article" year="1977">Tadjalli-Pour 1977: 107</bibRefCitation>
), but it could not be found.
</materialsCitation>
</paragraph>
<paragraph id="8BA936B5C853FFA5FF6DF85DFCC9A2EA" blockId="18.[151,1437,1814,2021]" box="[199,876,1922,1949]" pageId="18" pageNumber="169">
<materialsCitation id="3B7E3CE8C853FFA5FF6DF85DFCC9A2EA" ID-GBIF-Occurrence="3042095336" box="[199,876,1922,1949]" country="Iran" elevation="15" location="Caspian Sea" municipality="Astara-Hachtpar" pageId="18" pageNumber="169" specimenCount="1" typeStatus="holotype">
<emphasis id="B962EAA7C853FFA5FF6DF85DFEC3A2EB" bold="true" box="[199,358,1922,1949]" pageId="18" pageNumber="169">
<typeStatus id="54AD8817C853FFA5FF6DF85DFEA7A2EA" box="[199,258,1923,1949]" pageId="18" pageNumber="169">Type</typeStatus>
locality.
</emphasis>
<location id="8EC9606EC853FFA5FEC6F85CFE5CA2EA" LSID="urn:lsid:plazi:treatment:03BF87A3C853FFA3FF3DF9F4FE29A20A:8EC9606EC853FFA5FEC6F85CFE5CA2EA" box="[364,505,1922,1949]" country="Iran" municipality="Astara-Hachtpar" name="Caspian Sea" pageId="18" pageNumber="169">Caspian Sea</location>
,
<collectingMunicipality id="6BCDACCFC853FFA5FDA8F85CFD1EA2EB" box="[514,699,1922,1948]" pageId="18" pageNumber="169">Astara-Hachtpar</collectingMunicipality>
(
<collectingCountry id="F3017625C853FFA5FD61F85DFD58A2EA" box="[715,765,1923,1949]" name="Iran" pageId="18" pageNumber="169">Iran</collectingCountry>
),
<quantity id="4CEE9B50C853FFA5FCBAF85DFCCDA2EA" box="[784,872,1923,1949]" metricMagnitude="1" metricUnit="m" metricValue="1.5" metricValueMax="3.0" metricValueMin="0.0" pageId="18" pageNumber="169" unit="m" value="15.0" valueMax="30.0" valueMin="0.0">
<elevation id="003BD186C853FFA5FCBAF85DFCCDA2EA" box="[784,872,1923,1949]" metricMagnitude="1" metricUnit="m" metricValue="1.5" metricValueMax="3.0" metricValueMin="0.0" pageId="18" pageNumber="169" unit="m" value="15.0" valueMax="30.0" valueMin="0.0">030 m</elevation>
</quantity>
.
</materialsCitation>
</paragraph>
<paragraph id="8BA936B5C853FFA4FF6DF878FA32A58E" blockId="18.[151,1437,1814,2021]" lastBlockId="19.[151,1437,151,249]" lastPageId="19" lastPageNumber="170" pageId="18" pageNumber="169">
<emphasis id="B962EAA7C853FFA5FF6DF878FE27A2B7" bold="true" box="[199,386,1958,1984]" pageId="18" pageNumber="169">Other material.</emphasis>
<bibRefCitation id="EF874B44C853FFA5FE23F879FD6FA2B6" author="Kantor, Yu. I. &amp; Sysoev, A. V." box="[393,714,1958,1985]" pageId="18" pageNumber="169" refId="ref31309" refString="Kantor, Yu. I. &amp; Sysoev, A. V. (2006) Morskiye i solonovatovodnyye bryukhonogiye mollyuski Rossii i sopredel'nykh stran: illyustrirovannyy katalog. KMK Scientific Press, Moscow, 372 pp., 140 pls pp." type="book" year="2006">Kantor &amp; Sysoev (2006: 87)</bibRefCitation>
were unable to locate identified samples in the
<collectionCode id="ED07AE70C853FFA5FB4CF879FAB0A2B6" box="[1254,1301,1959,1985]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="18" pageNumber="169" type="Museum">ZIN</collectionCode>
collections, but we retrieved a lot in the Starobogatov collection containing
<specimenCount id="9D10FD3CC853FFA5FCD2F815FBB7A293" box="[888,1042,1994,2021]" pageId="18" pageNumber="169" type="generic">12 specimens</specimenCount>
determined as “
<taxonomicName id="4C164D36C853FFA5FB65F815FA2CA293" baseAuthorityName="Tadjalli-Pour" baseAuthorityYear="1977" box="[1231,1417,1994,2021]" class="Gastropoda" family="Hydrobiidae" genus="Caspia" kingdom="Animalia" order="Littorinimorpha" pageId="18" pageNumber="169" phylum="Mollusca" rank="species" species="gaillardi">
<emphasis id="B962EAA7C853FFA5FB65F815FA2CA293" box="[1231,1417,1994,2021]" italics="true" pageId="18" pageNumber="169">Caspia gaillardi</emphasis>
</taxonomicName>
”, collected in the Middle Caspian Basin off Apsheron Peninsula (
<tableCitation id="C694030EC852FFA4FCDCFF49FC6BA5C6" box="[886,974,151,177]" captionStart="TABLE 1" captionStartId="3.[151,239,473,499]" captionTargetPageId="3" captionText="TABLE 1. Locality information of the studied material. Coordinates of collecting sites are given based on data in museum catalogue cards, labels and corresponding publications. Asterisks mark localities with approximate geographic coordinates based on indirect data provided in the ZIN catalogue. Abbreviations: q.—quadrant, st.—station." httpUri="http://table.plazi.org/id/DF69663DC842FFB4FF3DFE07F8C6A760" pageId="19" pageNumber="170" tableUuid="DF69663DC842FFB4FF3DFE07F8C6A760">Table 1</tableCitation>
, locality 13). Additional five specimens come from the South Caspian Basin off Cheleken Peninsula (locality 18, labelled as “
<taxonomicName id="4C164D36C852FFA4FBF7FF65FA8AA5A2" box="[1117,1327,187,213]" class="Gastropoda" family="Hydrobiidae" genus="Caspia" kingdom="Animalia" order="Littorinimorpha" pageId="19" pageNumber="170" phylum="Mollusca" rank="species" species="subgmelini">
<emphasis id="B962EAA7C852FFA4FBF7FF65FA8AA5A2" box="[1117,1327,187,213]" italics="true" pageId="19" pageNumber="170">Caspia subgmelini</emphasis>
</taxonomicName>
”) and six specimens derive from late Holocene deposits of the Kura delta (localities 25a and 25f,
<collectionCode id="ED07AE70C852FFA4FBC7FF01FB08A58E" box="[1133,1197,223,249]" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="19" pageNumber="170" type="Museum">RGM</collectionCode>
13102991310304).
</paragraph>
</subSubSection>
<caption id="DF69663DC852FFA4FF3DF923FAD3A22A" ID-DOI="http://doi.org/10.5281/zenodo.4550172" ID-Zenodo-Dep="4550172" httpUri="https://zenodo.org/record/4550172/files/figure.png" pageId="19" pageNumber="170" startId="19.[151,250,1789,1813]" targetBox="[151,1436,273,1766]" targetPageId="19">
<paragraph id="8BA936B5C852FFA4FF3DF923FAD3A22A" blockId="19.[151,1437,1789,1885]" pageId="19" pageNumber="170">
<emphasis id="B962EAA7C852FFA4FF3DF923FEB0A262" bold="true" box="[151,277,1789,1813]" pageId="19" pageNumber="170">FIGURE 9.</emphasis>
<taxonomicName id="4C164D36C852FFA4FEB4F923FD48A262" authority="(Tadjalli-Pour, 1977)" baseAuthorityName="Tadjalli-Pour" baseAuthorityYear="1977" box="[286,749,1789,1813]" class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="19" pageNumber="170" phylum="Mollusca" rank="species" species="gaillardi">
<emphasis id="B962EAA7C852FFA4FEB4F923FDABA262" box="[286,526,1789,1813]" italics="true" pageId="19" pageNumber="170">Clathrocaspia gaillardi</emphasis>
(
<bibRefCitation id="EF874B44C852FFA4FDB7F923FD43A262" author="Tadjalli-Pour, M." box="[541,742,1789,1813]" pageId="19" pageNumber="170" pagination="87 - 117" refId="ref33728" refString="Tadjalli-Pour, M. (1977) Les Mollusques marins des cotes Iraniennes de la Mer Caspienne (Astara-Hachtpar). Journal de Conchyliologie, 114 (3 - 4), 87 - 117." type="journal article" year="1977">Tadjalli-Pour, 1977</bibRefCitation>
)
</taxonomicName>
from the Middle and South Caspian basins. (ae) Five specimens from locality 13 (ZIN, no number). (fi) Three specimens labelled as “
<taxonomicName id="4C164D36C852FFA4FCCDF8FCFB81A24E" box="[871,1060,1825,1849]" class="Gastropoda" family="Hydrobiidae" genus="Caspia" kingdom="Animalia" order="Littorinimorpha" pageId="19" pageNumber="170" phylum="Mollusca" rank="species" species="subgmelini">
<emphasis id="B962EAA7C852FFA4FCCDF8FCFB81A24E" box="[871,1060,1825,1849]" italics="true" pageId="19" pageNumber="170">Caspia subgmelini</emphasis>
</taxonomicName>
” from locality 18 (ZIN, no number). (jn) Five specimens from Holocene deposits of the Kura delta (locality 25d, RGM 13102991310303). Scale bar = 1 mm.
</paragraph>
</caption>
<subSubSection id="C30C653EC852FFA3FF6DF855FBF8A46A" lastPageId="20" lastPageNumber="171" pageId="19" pageNumber="170" type="description">
<paragraph id="8BA936B5C852FFA3FF6DF855FBF8A46A" blockId="19.[151,1437,1931,2030]" lastBlockId="20.[151,1437,151,501]" lastPageId="20" lastPageNumber="171" pageId="19" pageNumber="170">
<emphasis id="B962EAA7C852FFA4FF6DF855FEFCA2D2" bold="true" box="[199,345,1931,1957]" pageId="19" pageNumber="170">Description.</emphasis>
Shell comparably large, broadly conical, consisting of up to 5 low to moderately convex, slightly stepped whorls. Protoconch bulbous, ranges around
<quantity id="4CEE9B50C852FFA4FD45F871FC24A2BD" box="[751,897,1967,1994]" metricMagnitude="-1" metricUnit="m" metricValue="3.275" metricValueMax="3.35" metricValueMin="3.2" pageId="19" pageNumber="170" unit="mm" value="327.5" valueMax="335.0" valueMin="320.0">320335 µm</quantity>
and consists of c. 1.2 whorls; surface strongly sculptured, with distinct wrinkles that increase in size and become more irregular towards teleoconch; final triangu-lar part of protoconch always devoid of wrinkles (
<figureCitation id="132D2A30C855FFA3FD67FF49FCBBA5C5" box="[717,798,151,178]" captionStart="FIGURE 10" captionStartId="20.[151,250,1460,1484]" captionTargetBox="[151,1436,525,1437]" captionTargetId="figure-190@20.[151,1436,525,1437]" captionTargetPageId="20" captionText="FIGURE 10. Protoconch views of Clathrocaspia gaillardi (Tadjalli-Pour, 1977) from Holocene deposits of the Kura delta. (a, g) RGM 1310299 (locality 25d). (b, h) RGM 1310300 (locality 25d). (c, f) RGM 1310304 (locality 25f). (d) RGM 1310302 (locality 25d). (e) RGM 1310301 (locality 25d). (i) RGM 1310303 (locality 25d). Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.4550174" httpUri="https://zenodo.org/record/4550174/files/figure.png" pageId="20" pageNumber="171">Fig. 10</figureCitation>
ai); additionally, spiral threads occur on first half whorl (
<figureCitation id="132D2A30C855FFA3FF0AFF65FE9AA5A1" box="[160,319,187,214]" captionStart="FIGURE 10" captionStartId="20.[151,250,1460,1484]" captionTargetBox="[151,1436,525,1437]" captionTargetId="figure-190@20.[151,1436,525,1437]" captionTargetPageId="20" captionText="FIGURE 10. Protoconch views of Clathrocaspia gaillardi (Tadjalli-Pour, 1977) from Holocene deposits of the Kura delta. (a, g) RGM 1310299 (locality 25d). (b, h) RGM 1310300 (locality 25d). (c, f) RGM 1310304 (locality 25f). (d) RGM 1310302 (locality 25d). (e) RGM 1310301 (locality 25d). (i) RGM 1310303 (locality 25d). Scale bars = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.4550174" httpUri="https://zenodo.org/record/4550174/files/figure.png" pageId="20" pageNumber="171">Fig. 10e, g, h</figureCitation>
). Teleoconch covered with strong reticulate sculpture, with sharp ribs and subquadrangular interspaces; spirals dominate, axials faintly sigmoid. Aperture ovoid, sometimes detached, leaving slit-like umbilicus; in other cases, basal columellar margin expands beyond columella and covers umbilicus.
</paragraph>
</subSubSection>
<subSubSection id="C30C653EC855FFA3FF6DFEF9FE93A242" pageId="20" pageNumber="171" type="discussion">
<paragraph id="8BA936B5C855FFA3FF6DFEF9FA83A482" blockId="20.[151,1437,151,501]" pageId="20" pageNumber="171">
<emphasis id="B962EAA7C855FFA3FF6DFEF9FE9EA436" bold="true" box="[199,315,295,321]" pageId="20" pageNumber="171">Remarks.</emphasis>
The material in the Starobogatovs collection fits well to
<taxonomicName id="4C164D36C855FFA3FC6AFEF9FBE7A436" baseAuthorityName="Tadjalli-Pour" baseAuthorityYear="1977" box="[960,1090,295,321]" class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="20" pageNumber="171" phylum="Mollusca" rank="species" species="gaillardi">
<emphasis id="B962EAA7C855FFA3FC6AFEF9FBE7A436" box="[960,1090,295,321]" italics="true" pageId="20" pageNumber="171">C. gaillardi</emphasis>
</taxonomicName>
as described and illustrated by
<bibRefCitation id="EF874B44C855FFA3FF3DFE95FEDAA412" author="Tadjalli-Pour, M." box="[151,383,331,357]" pageId="20" pageNumber="171" pagination="87 - 117" refId="ref33728" refString="Tadjalli-Pour, M. (1977) Les Mollusques marins des cotes Iraniennes de la Mer Caspienne (Astara-Hachtpar). Journal de Conchyliologie, 114 (3 - 4), 87 - 117." type="journal article" year="1977">Tadjalli-Pour (1977)</bibRefCitation>
, regarding the broad conical shape with five whorls, the marked sutures, the strong sculpture and the comparably large size. Similarly, five specimens labelled in Starobogatovs collection as “
<taxonomicName id="4C164D36C855FFA3FAAFFEB1FF78A4DA" class="Gastropoda" family="Hydrobiidae" genus="Caspia" kingdom="Animalia" order="Littorinimorpha" pageId="20" pageNumber="171" phylum="Mollusca" rank="species" species="subgmelini">
<emphasis id="B962EAA7C855FFA3FAAFFEB1FF78A4DA" italics="true" pageId="20" pageNumber="171">Caspia subgmelini</emphasis>
</taxonomicName>
” (an unavailable collection name;
<figureCitation id="132D2A30C855FFA3FDCFFE4DFD03A4D9" box="[613,678,403,430]" captionStart="FIGURE 9" captionStartId="19.[151,250,1789,1813]" captionTargetBox="[151,1436,273,1766]" captionTargetId="figure-68@19.[151,1436,273,1766]" captionTargetPageId="19" captionText="FIGURE 9. Clathrocaspia gaillardi (Tadjalli-Pour, 1977) from the Middle and South Caspian basins. (ae) Five specimens from locality 13 (ZIN, no number). (fi) Three specimens labelled as “Caspia subgmelini” from locality 18 (ZIN, no number). (jn) Five specimens from Holocene deposits of the Kura delta (locality 25d, RGM 13102991310303). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4550172" httpUri="https://zenodo.org/record/4550172/files/figure.png" pageId="20" pageNumber="171">Fig. 9</figureCitation>
fi) collected off Cheleken Peninsula fit well to
<taxonomicName id="4C164D36C855FFA3FB6EFE4DFAE0A4DA" baseAuthorityName="Tadjalli-Pour" baseAuthorityYear="1977" box="[1220,1349,403,429]" class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="20" pageNumber="171" phylum="Mollusca" rank="species" species="gaillardi">
<emphasis id="B962EAA7C855FFA3FB6EFE4DFAE0A4DA" box="[1220,1349,403,429]" italics="true" pageId="20" pageNumber="171">C. gaillardi</emphasis>
</taxonomicName>
, and so do specimens from the Holocene deposits off the Kura delta (
<figureCitation id="132D2A30C855FFA3FCFFFE69FC3DA4A5" box="[853,920,439,466]" captionStart="FIGURE 9" captionStartId="19.[151,250,1789,1813]" captionTargetBox="[151,1436,273,1766]" captionTargetId="figure-68@19.[151,1436,273,1766]" captionTargetPageId="19" captionText="FIGURE 9. Clathrocaspia gaillardi (Tadjalli-Pour, 1977) from the Middle and South Caspian basins. (ae) Five specimens from locality 13 (ZIN, no number). (fi) Three specimens labelled as “Caspia subgmelini” from locality 18 (ZIN, no number). (jn) Five specimens from Holocene deposits of the Kura delta (locality 25d, RGM 13102991310303). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4550172" httpUri="https://zenodo.org/record/4550172/files/figure.png" pageId="20" pageNumber="171">Fig. 9</figureCitation>
jn). Some of the specimens only have a less stepped spire than is typical of
<taxonomicName id="4C164D36C855FFA3FE5EFE05FDD0A482" baseAuthorityName="Tadjalli-Pour" baseAuthorityYear="1977" box="[500,629,475,501]" class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="20" pageNumber="171" phylum="Mollusca" rank="species" species="gaillardi">
<emphasis id="B962EAA7C855FFA3FE5EFE05FDD0A482" box="[500,629,475,501]" italics="true" pageId="20" pageNumber="171">C. gaillardi</emphasis>
</taxonomicName>
, but we consider this to range within intraspecific variability.
</paragraph>
<caption id="DF69663DC855FFA3FF3DFA6AFB0AA362" ID-DOI="http://doi.org/10.5281/zenodo.4550174" ID-Zenodo-Dep="4550174" httpUri="https://zenodo.org/record/4550174/files/figure.png" pageId="20" pageNumber="171" startId="20.[151,250,1460,1484]" targetBox="[151,1436,525,1437]" targetPageId="20">
<paragraph id="8BA936B5C855FFA3FF3DFA6AFB0AA362" blockId="20.[151,1437,1460,1557]" pageId="20" pageNumber="171">
<emphasis id="B962EAA7C855FFA3FF3DFA6AFE85A0BB" bold="true" box="[151,288,1460,1484]" pageId="20" pageNumber="171">FIGURE 10.</emphasis>
Protoconch views of
<taxonomicName id="4C164D36C855FFA3FE55FA6AFC6CA0BB" authority="(Tadjalli-Pour, 1977)" baseAuthorityName="Tadjalli-Pour" baseAuthorityYear="1977" box="[511,969,1460,1484]" class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="20" pageNumber="171" phylum="Mollusca" rank="species" species="gaillardi">
<emphasis id="B962EAA7C855FFA3FE55FA6AFD48A0BB" box="[511,749,1460,1484]" italics="true" pageId="20" pageNumber="171">Clathrocaspia gaillardi</emphasis>
(
<bibRefCitation id="EF874B44C855FFA3FD50FA6AFC67A0BB" author="Tadjalli-Pour, M." box="[762,962,1460,1484]" pageId="20" pageNumber="171" pagination="87 - 117" refId="ref33728" refString="Tadjalli-Pour, M. (1977) Les Mollusques marins des cotes Iraniennes de la Mer Caspienne (Astara-Hachtpar). Journal de Conchyliologie, 114 (3 - 4), 87 - 117." type="journal article" year="1977">Tadjalli-Pour, 1977</bibRefCitation>
)
</taxonomicName>
from Holocene deposits of the Kura delta. (a, g) RGM 1310299 (locality 25d). (b, h) RGM 1310300 (locality 25d). (c, f) RGM 1310304 (locality 25f). (d) RGM 1310302 (locality 25d). (e) RGM 1310301 (locality 25d). (i) RGM 1310303 (locality 25d). Scale bars = 100 μm.
</paragraph>
</caption>
<paragraph id="8BA936B5C855FFA3FF6DF99CFE93A242" blockId="20.[151,1437,1602,1917]" pageId="20" pageNumber="171">
<bibRefCitation id="EF874B44C855FFA3FF6DF99CFDE2A32A" author="Sitnikova, T. Ya. &amp; Starobogatov, Ya. I." box="[199,583,1602,1629]" pageId="20" pageNumber="171" pagination="1357 - 1367" refId="ref33455" refString="Sitnikova, T. Ya. &amp; Starobogatov, Ya. I. (1998) Polovaya sistema i radula kaspiyskikh Pyrgulidae (podsemeystva Turricaspiinae i Caspiinae, Gastropoda, Pectinibranchia). Zoologicheskiy Zhurnal, 77 (12), 1357 - 1367." type="journal article" year="1998">Sitnikova &amp; Starobogatov (1998)</bibRefCitation>
provided data on the female reproductive anatomy and the radula of this species. They noted that the studied morphological characters of
<taxonomicName id="4C164D36C855FFA3FCDDF9B9FC5FA3F7" baseAuthorityName="Tadjalli-Pour" baseAuthorityYear="1977" box="[887,1018,1638,1665]" class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="20" pageNumber="171" phylum="Mollusca" rank="species" species="gaillardi">
<emphasis id="B962EAA7C855FFA3FCDDF9B9FC5FA3F7" box="[887,1018,1638,1665]" italics="true" pageId="20" pageNumber="171">C. gaillardi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C164D36C855FFA3FB99F9B9FB15A3F7" box="[1075,1200,1638,1665]" class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="20" pageNumber="171" phylum="Mollusca" rank="species" species="gmelinii">
<emphasis id="B962EAA7C855FFA3FB99F9B9FB15A3F7" box="[1075,1200,1638,1665]" italics="true" pageId="20" pageNumber="171">C. gmelinii</emphasis>
</taxonomicName>
are almost identical. Also
<bibRefCitation id="EF874B44C855FFA3FF78F954FE45A3D2" author="Wesselingh, F. P. &amp; Neubauer, T. A. &amp; Anistratenko, V. V. &amp; Vinarski, M. V. &amp; Yanina, T. &amp; ter Poorten, J. J. &amp; Kijashko, P. V. &amp; Albrecht, C. &amp; Anistratenko, O. Yu. &amp; D'Hont, A. &amp; Frolov, P. &amp; Martinez Gandara, A. &amp; Gittenberger, A. &amp; Gogaladze, A. &amp; Karpinsky, M. &amp; Lattuada, M. &amp; Popa, L. &amp; Sands, A. F. &amp; van de Velde, S. &amp; Vandendorpe, J. &amp; Wilke, T." box="[210,480,1674,1701]" pageId="20" pageNumber="171" pagination="31 - 124" refId="ref34307" refString="Wesselingh, F. P., Neubauer, T. A., Anistratenko, V. V., Vinarski, M. V., Yanina, T., ter Poorten, J. J., Kijashko, P. V., Albrecht, C., Anistratenko, O. Yu., D'Hont, A., Frolov, P., Martinez Gandara, A., Gittenberger, A., Gogaladze, A., Karpinsky, M., Lattuada, M., Popa, L., Sands, A. F., van de Velde, S., Vandendorpe, J. &amp; Wilke, T. (2019) Mollusc species from the Pontocaspian region - an expert opinion list. ZooKeys, 827, 31 - 124. https: // doi. org / 10.3897 / zookeys. 827.31365" type="journal article" year="2019">
Wesselingh
<emphasis id="B962EAA7C855FFA3FEF2F955FE2DA3D3" box="[344,392,1674,1700]" italics="true" pageId="20" pageNumber="171">et al</emphasis>
. (2019)
</bibRefCitation>
listed the species as a tentative synonym of
<taxonomicName id="4C164D36C855FFA3FC62F955FBE1A3D3" box="[968,1092,1674,1701]" class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="20" pageNumber="171" phylum="Mollusca" rank="species" species="gmelinii">
<emphasis id="B962EAA7C855FFA3FC62F955FBE1A3D3" box="[968,1092,1674,1701]" italics="true" pageId="20" pageNumber="171">C. gmelinii</emphasis>
</taxonomicName>
. However,
<taxonomicName id="4C164D36C855FFA3FB6EF955FA9AA3D3" box="[1220,1343,1674,1701]" class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="20" pageNumber="171" phylum="Mollusca" rank="species" species="gmelinii">
<emphasis id="B962EAA7C855FFA3FB6EF955FA9AA3D3" box="[1220,1343,1674,1701]" italics="true" pageId="20" pageNumber="171">C. gmelinii</emphasis>
</taxonomicName>
is much smaller and has a less stepped outline and a weaker reticulate sculpture. Specimens of
<taxonomicName id="4C164D36C855FFA3FB2CF971FAAFA3BF" baseAuthorityName="Tadjalli-Pour" baseAuthorityYear="1977" box="[1158,1290,1710,1737]" class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="20" pageNumber="171" phylum="Mollusca" rank="species" species="gaillardi">
<emphasis id="B962EAA7C855FFA3FB2CF971FAAFA3BF" box="[1158,1290,1710,1737]" italics="true" pageId="20" pageNumber="171">C. gaillardi</emphasis>
</taxonomicName>
also show a certain similarity to
<taxonomicName id="4C164D36C855FFA3FED3F90DFE4BA39B" authorityName="Clessin &amp; W. Dybowski" authorityYear="1887" box="[377,494,1746,1773]" class="Gastropoda" family="Hydrobiidae" genus="Caspia" kingdom="Animalia" order="Littorinimorpha" pageId="20" pageNumber="171" phylum="Mollusca" rank="species" species="pallasii">
<emphasis id="B962EAA7C855FFA3FED3F90DFE4BA39B" box="[377,494,1746,1773]" italics="true" pageId="20" pageNumber="171">C. pallasii</emphasis>
</taxonomicName>
, but the latter species can be distinguished by its larger, bulbous protoconch and the more slender shell. The stepped teleoconch of
<taxonomicName id="4C164D36C855FFA3FD0BF929FC86A267" baseAuthorityName="Tadjalli-Pour" baseAuthorityYear="1977" box="[673,803,1782,1809]" class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="20" pageNumber="171" phylum="Mollusca" rank="species" species="gaillardi">
<emphasis id="B962EAA7C855FFA3FD0BF929FC86A267" box="[673,803,1782,1809]" italics="true" pageId="20" pageNumber="171">C. gaillardi</emphasis>
</taxonomicName>
and the comparably high last whorl also differentiates it from
<taxonomicName id="4C164D36C855FFA3FF7FF8C5FE8AA243" baseAuthorityName="Logvinenko &amp; Starobogatov" baseAuthorityYear="1969" box="[213,303,1818,1845]" class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="20" pageNumber="171" phylum="Mollusca" rank="species" species="isseli">
<emphasis id="B962EAA7C855FFA3FF7FF8C5FE8AA243" box="[213,303,1818,1845]" italics="true" pageId="20" pageNumber="171">C. isseli</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C30C653EC855FFA3FF6DF8E0FE29A20A" pageId="20" pageNumber="171" type="distribution">
<paragraph id="8BA936B5C855FFA3FF6DF8E0FE29A20A" blockId="20.[151,1437,1602,1917]" pageId="20" pageNumber="171">
<emphasis id="B962EAA7C855FFA3FF6DF8E0FEC4A22F" bold="true" box="[199,353,1854,1880]" pageId="20" pageNumber="171">Distribution.</emphasis>
Caspian Sea, South Basin to southern part of Middle Basin, known only from shallow waters (max. depth of
<quantity id="4CEE9B50C855FFA3FEE8F8BDFEDAA20A" box="[322,383,1891,1917]" metricMagnitude="1" metricUnit="m" metricValue="4.4" pageId="20" pageNumber="171" unit="m" value="44.0">44 m</quantity>
).
</paragraph>
</subSubSection>
</treatment>
</document>