247 lines
33 KiB
XML
247 lines
33 KiB
XML
<document id="02B680E77AD3F4EBC8AD000D570D6F10" ID-DOI="10.4202/app.00061.2014" ID-ISSN="1732-2421" ID-Zenodo-Dep="10626402" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.taxonomicNames_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1707249081010" checkinUser="felipe" docAuthor="Denayer, Julien" docDate="2016" docId="03B687FCFFEC194DFF48F8FC34D685E5" docLanguage="en" docName="ActaPalaeontolPol.61.1.51-70.pdf" docOrigin="Acta Palaeontologica Polonica 61 (1)" docSource="http://dx.doi.org/10.4202/app.00061.2014" docStyle="DocumentStyle:E4780F6D34569985EDF9CACFB9BC780E.4:ActaPalaeontolPol.2015-.journal_article" docStyleId="E4780F6D34569985EDF9CACFB9BC780E" docStyleName="ActaPalaeontolPol.2015-.journal_article" docStyleVersion="4" docTitle="Caninophyllum charli Denayer 2016, sp. nov." docType="treatment" docVersion="3" lastPageNumber="64" masterDocId="FF8FFF84FFE01940FFDFFFE0367F8762" masterDocTitle="Rugose corals across the Devonian-Carboniferous boundary in NW Turkey" masterLastPageNumber="70" masterPageNumber="51" pageNumber="63" updateTime="1713405178588" updateUser="karina" zenodo-license-document="CC-BY-4.0">
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<mods:title id="361C52654011675B0FD1AEB53BFC095D">Rugose corals across the Devonian-Carboniferous boundary in NW Turkey</mods:title>
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<mods:namePart id="2F30639D6A1B5D6CBACD931FA76772AC">Denayer, Julien</mods:namePart>
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<mods:title id="41BDA582C52F83D7447596E525CD22BA">Acta Palaeontologica Polonica</mods:title>
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<mods:part id="14660875F6FD4EFC6397ED9EE016DB42">
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<mods:date id="D3071C5722393D295E8B860BA7C3FBBB">2016</mods:date>
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<mods:number id="99238C33903D19C2C7F76C1C8D50CC90">2014-10-07</mods:number>
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<mods:number id="237F4960B0EA43B98863F849DB45C06D">61</mods:number>
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<mods:identifier id="63740B86064AA43559F39ADB7ECE18C8" type="DOI">10.4202/app.00061.2014</mods:identifier>
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<mods:identifier id="741B5C66C36E338E358D9E199960734C" type="ISSN">1732-2421</mods:identifier>
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<treatment id="03B687FCFFEC194DFF48F8FC34D685E5" LSID="urn:lsid:plazi:treatment:03B687FCFFEC194DFF48F8FC34D685E5" httpUri="http://treatment.plazi.org/id/03B687FCFFEC194DFF48F8FC34D685E5" lastPageId="13" lastPageNumber="64" pageId="12" pageNumber="63">
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<subSubSection id="C3056561FFEC194CFF48F8FC3464805A" box="[151,539,1820,1848]" pageId="12" pageNumber="63" type="nomenclature">
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<paragraph id="8BA036EAFFEC194CFF48F8FC3464805A" blockId="12.[151,539,1820,1848]" box="[151,539,1820,1848]" pageId="12" pageNumber="63">
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<heading id="D0E88186FFEC194CFF48F8FC3464805A" box="[151,539,1820,1848]" fontSize="12" level="2" pageId="12" pageNumber="63" reason="2">
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<taxonomicName id="4C1F4D69FFEC194CFF48F8FC37CE805A" authority="Denayer, 2016" authorityName="Denayer" authorityYear="2016" box="[151,433,1820,1848]" class="Anthozoa" family="Bothrophyllidae" genus="Caninophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stauriida" pageId="12" pageNumber="63" phylum="Cnidaria" rank="species" species="charli" status="sp. nov.">
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<emphasis id="B96BEAF8FFEC194CFF48F8FC37CE805A" box="[151,433,1820,1848]" italics="true" pageId="12" pageNumber="63">Caninophyllum charli</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2585783FFEC194CFE66F8FC3464805A" box="[441,539,1820,1848]" pageId="12" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3056561FFEC194CFF48F8A9379380C4" pageId="12" pageNumber="63" type="description">
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||
<paragraph id="8BA036EAFFEC194CFF48F8A9377D803C" blockId="12.[151,258,1865,1886]" box="[151,258,1865,1886]" pageId="12" pageNumber="63">
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<figureCitation id="13242A6FFFEC194CFF48F8A93681803C" box="[151,254,1865,1886]" captionStart="Fig" captionStartId="11.[96,129,1614,1634]" captionTargetBox="[98,1478,161,1587]" captionTargetId="figure-79@11.[98,1479,161,1588]" captionTargetPageId="11" captionText="Fig. 8. Devonian–Carboniferous rugose corals from Turkey. A–E. Caninophyllum charli sp. nov. from the lower Tournaisian (upper Hastarian) of Zonguldak, Gökgöl section. A. Holotype, G.8.1.1, successive TS (A1, A2); close-up view of the dissepimentarium, TS (A3). B. G.8.6.1, successive TS (B1, B2). C. G.8.4.3, TS. D. G.8.4.2, TS. E. G.8.3.2, LS. F, G. Uralinia simplex (Yü, 1933) from the lower Tournaisian (lower Hastarian) of Bartın, Topluca section. F. ET.9c.7, TS. G. ET.9c.8, TS. H. Amplexocarinia rozkowskae (Fedorowski, 2003) from the uppermost Famennian (Strunian) of Bartın, Dallıca section, D.2.4.II’, TS. I.?Metriophyllum sp. from the uppermost Famennian (Strunian) of Bartın, Topluca section, ET.11.X, TS. J–L. Bounophyllum praecursor (Frech, 1895) form the uppermost Famennian (Strunian) of Bartın, Topluca section. J. ET.12a.1.III, successive TS (J1–J3). K. ET.12a.1.II, TS. L. ET.12a.1.V, LS. Scale bar A–G, 5 mm; H, 3 mm; I, 1.9 mm; J–L, 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10626415" httpUri="https://zenodo.org/record/10626415/files/figure.png" pageId="12" pageNumber="63">Fig. 8A–E</figureCitation>
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.
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</paragraph>
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<paragraph id="8BA036EAFFEC194CFF48F895379380C4" blockId="12.[151,824,1909,1958]" pageId="12" pageNumber="63">
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<emphasis id="B96BEAF8FFEC194CFF48F895368180E8" box="[151,254,1909,1930]" italics="true" pageId="12" pageNumber="63">Etymology</emphasis>
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: In honour of François Charles, who first described corals in the Carboniferous of
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<collectingRegion id="49DBF808FFEC194CFEA3F871379880C4" box="[380,487,1937,1958]" country="Turkey" name="Zonguldak" pageId="12" pageNumber="63">Zonguldak</collectingRegion>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3056561FFEC194DFC81FF4034D685E5" lastPageId="13" lastPageNumber="64" pageId="12" pageNumber="63" type="materials_examined">
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<paragraph id="8BA036EAFFEC194CFC81FF40323A87B3" blockId="12.[862,1535,160,209]" pageId="12" pageNumber="63">
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<materialsCitation id="3B773CB7FFEC194CFC81FF40323A87B3" collectionCode="TS" location="Zonguldak" pageId="12" pageNumber="63" specimenCode="G.8.1.1" specimenCount="1" stateProvince="Zonguldak" typeStatus="holotype">
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<emphasis id="B96BEAF8FFEC194CFC81FF4035C987D7" box="[862,950,160,181]" italics="true" pageId="12" pageNumber="63">
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<typeStatus id="54A48848FFEC194CFC81FF4035C987D7" box="[862,950,160,181]" pageId="12" pageNumber="63">Holotype</typeStatus>
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</emphasis>
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: Corallum included in a rock matrix, specimen
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<specimenCode id="DBB99E91FFEC194CFA6EFF40338587D7" box="[1457,1530,160,181]" pageId="12" pageNumber="63">G.8.1.1</specimenCode>
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,
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<collectingRegion id="49DBF808FFEC194CFC81FF5C35B987B3" box="[862,966,188,209]" country="Turkey" name="Zonguldak" pageId="12" pageNumber="63">Zonguldak</collectingRegion>
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2011 (5 TS).
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</materialsCitation>
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</paragraph>
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<paragraph id="8BA036EAFFEC194CFC81FF3D33BC8790" blockId="12.[862,1475,221,242]" box="[862,1475,221,242]" pageId="12" pageNumber="63">
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<emphasis id="B96BEAF8FFEC194CFC81FF3D35A68790" box="[862,985,221,242]" italics="true" pageId="12" pageNumber="63">Type locality</emphasis>
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: Gökgöl section, south of
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<collectingRegion id="49DBF808FFEC194CFB08FF3D333E8790" box="[1239,1345,221,242]" country="Turkey" name="Zonguldak" pageId="12" pageNumber="63">Zonguldak</collectingRegion>
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, NW
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<collectingCountry id="F308767AFFEC194CFAA7FF3D33C18790" box="[1400,1470,221,242]" name="Turkey" pageId="12" pageNumber="63">Turkey</collectingCountry>
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.
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</paragraph>
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<paragraph id="8BA036EAFFEC194CFC81FF1F32AA8652" blockId="12.[862,1535,255,304]" pageId="12" pageNumber="63">
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<emphasis id="B96BEAF8FFEC194CFC81FF1F35A58676" box="[862,986,255,276]" italics="true" pageId="12" pageNumber="63">Type horizon</emphasis>
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: Yılanlı Formation, unit G8, lower Tournaisian (Hastarian, MFZ3 biozone of
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<bibRefCitation id="EF8E4B1BFFEC194CFBEFFEFB32B58652" author="Poty, E. & Devuyst, F. - X. & Hance, L." box="[1072,1226,283,304]" pageId="12" pageNumber="63" pagination="829 - 857" refId="ref16230" refString="Poty, E., Devuyst, F. - X., and Hance, L. 2006. Upper Devonian and Mississippian foraminiferal and rugose coral zonations of Belgium and northern France: a tool for Eurasian correlations. Geological Magazine 143: 829 - 857." type="journal article" year="2006">Poty et al. 2006</bibRefCitation>
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).
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</paragraph>
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<paragraph id="8BA036EAFFEC194CFC81FEA3328E86FF" blockId="12.[862,1535,323,413]" pageId="12" pageNumber="63">
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<emphasis id="B96BEAF8FFEC194CFC81FEA335BF863E" box="[862,960,323,348]" italics="true" pageId="12" pageNumber="63">Material</emphasis>
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.—
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<materialsCitation id="3B773CB7FFEC194CFC3EFEA333D4861F" collectionCode="TS, LS" county="Gokgol" location="Dallica" municipality="Dallica" pageId="12" pageNumber="63" specimenCount="18" stateProvince="Zonguldak" typeStatus="holotype">
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Eighteen specimens (27 TS, 2 LS):
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<specimenCount id="9D19FD63FFEC194CFAB2FEA333F6863E" box="[1389,1417,323,348]" pageId="12" pageNumber="63">15</specimenCount>
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, including the holotype, coming from
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<collectingCounty id="62C14E66FFEC194CFB4BFE843298861F" box="[1172,1255,356,381]" pageId="12" pageNumber="63">Gökgöl</collectingCounty>
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||
(
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<collectingRegion id="49DBF808FFEC194CFB29FE84330C861F" box="[1270,1395,356,381]" country="Turkey" name="Zonguldak" pageId="12" pageNumber="63">Zonguldak</collectingRegion>
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) and
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</materialsCitation>
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<materialsCitation id="3B773CB7FFEC194CFA6DFE84329286FF" collectionCode="TS, LS" county="Gokgol" location="Dallica" municipality="Dallica" pageId="12" pageNumber="63" specimenCount="18" stateProvince="Zonguldak" typeStatus="holotype">
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<specimenCount id="9D19FD63FFEC194CFA6DFE8433BF861F" box="[1458,1472,356,381]" pageId="12" pageNumber="63">3</specimenCount>
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from
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<collectingMunicipality id="6BC4AC90FFEC194CFC81FE6435D086FF" box="[862,943,388,413]" pageId="12" pageNumber="63">Dallıca</collectingMunicipality>
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(
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<collectingRegion id="49DBF808FFEC194CFC61FE64327486FF" box="[958,1035,388,413]" country="Turkey" name="Bartin" pageId="12" pageNumber="63">Bartın</collectingRegion>
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),
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<geologicalTimeScale id="F93FB641FFEC194CFBC1FE64329286FF" box="[1054,1261,388,413]" pageId="12" pageNumber="63">lower Tournaisian</geologicalTimeScale>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BA036EAFFEC194CFC81FE4F334785CB" blockId="12.[862,1535,431,681]" pageId="12" pageNumber="63">
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<emphasis id="B96BEAF8FFEC194CFC81FE4F35AF86AA" box="[862,976,431,456]" italics="true" pageId="12" pageNumber="63">Diagnosis</emphasis>
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.—Small
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<taxonomicName id="4C1F4D69FFEC194CFBE1FE4F329486AA" authorityName="Lewis" authorityYear="1929" box="[1086,1259,431,456]" class="Anthozoa" family="Bothrophyllidae" genus="Caninophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="63" phylum="Cnidaria" rank="genus">
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<emphasis id="B96BEAF8FFEC194CFBE1FE4F329486AA" box="[1086,1259,431,456]" italics="true" pageId="12" pageNumber="63">Caninophyllum</emphasis>
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</taxonomicName>
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,
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<quantity id="4CE79B0FFFEC194CFB25FE4F330B86AA" box="[1274,1396,431,456]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" metricValueMax="3.2" metricValueMin="3.0" pageId="12" pageNumber="63" unit="mm" value="31.0" valueMax="32.0" valueMin="30.0">30–32 mm</quantity>
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in diameter (
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<quantity id="4CE79B0FFFEC194CFCB9FE2F35A1868A" box="[870,990,463,488]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="12" pageNumber="63" unit="mm" value="17.5" valueMax="20.0" valueMin="15.0">15–20 mm</quantity>
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for the tabularium), having 42–46 septa of each order. Major septa extending to the axis in the juvenile stages but withdrawn in the mature stages, thickened in the cardinal parts of the tabularium. Minor septa rudimentary. Dissepimentarium narrow but complex, constituted of concentric, herringbone and arched interseptal dissepiments. Cardinal fossula conspicuous and opened.
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</paragraph>
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<paragraph id="8BA036EAFFEC194CFC81FD5B32B88159" blockId="12.[862,1535,699,1595]" pageId="12" pageNumber="63">
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<emphasis id="B96BEAF8FFEC194CFC81FD5B359A85B6" box="[862,997,699,724]" italics="true" pageId="12" pageNumber="63">Description</emphasis>
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.—The corallum is cylindrical,
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<quantity id="4CE79B0FFFEC194CFA81FD5B33C885B6" box="[1374,1463,699,724]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="12" pageNumber="63" unit="cm" value="4.0" valueMax="5.0" valueMin="3.0">3–5 cm</quantity>
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high. The mean diameter is
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<quantity id="4CE79B0FFFEC194CFBBAFD3B32C98596" box="[1125,1206,731,756]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="12" pageNumber="63" unit="mm" value="32.0">32 mm</quantity>
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(maximum
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<quantity id="4CE79B0FFFEC194CFA99FD3B33E78596" box="[1350,1432,731,756]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="12" pageNumber="63" unit="mm" value="38.0">38 mm</quantity>
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) and the tabularium is in average
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<quantity id="4CE79B0FFFEC194CFB5FFD1C32B18477" box="[1152,1230,764,789]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="12" pageNumber="63" unit="mm" value="17.0">17 mm</quantity>
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||
wide (maximum
|
||
<quantity id="4CE79B0FFFEC194CFA7FFD1C338E8477" box="[1440,1521,764,789]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="12" pageNumber="63" unit="mm" value="21.0">21 mm</quantity>
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). There are, on average, 46 septa of each order (maximum 48). The major septa are long, reaching the axis in the juvenile stages but withdraw toward the periphery during the growth, leaving a central zone of 5–6 mm-wide in the centre of the tabularium (
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<figureCitation id="13242A6FFFEC194CFB56FC7D329E84D4" box="[1161,1249,925,950]" captionStart="Fig" captionStartId="11.[96,129,1614,1634]" captionTargetBox="[98,1478,161,1587]" captionTargetId="figure-79@11.[98,1479,161,1588]" captionTargetPageId="11" captionText="Fig. 8. Devonian–Carboniferous rugose corals from Turkey. A–E. Caninophyllum charli sp. nov. from the lower Tournaisian (upper Hastarian) of Zonguldak, Gökgöl section. A. Holotype, G.8.1.1, successive TS (A1, A2); close-up view of the dissepimentarium, TS (A3). B. G.8.6.1, successive TS (B1, B2). C. G.8.4.3, TS. D. G.8.4.2, TS. E. G.8.3.2, LS. F, G. Uralinia simplex (Yü, 1933) from the lower Tournaisian (lower Hastarian) of Bartın, Topluca section. F. ET.9c.7, TS. G. ET.9c.8, TS. H. Amplexocarinia rozkowskae (Fedorowski, 2003) from the uppermost Famennian (Strunian) of Bartın, Dallıca section, D.2.4.II’, TS. I.?Metriophyllum sp. from the uppermost Famennian (Strunian) of Bartın, Topluca section, ET.11.X, TS. J–L. Bounophyllum praecursor (Frech, 1895) form the uppermost Famennian (Strunian) of Bartın, Topluca section. J. ET.12a.1.III, successive TS (J1–J3). K. ET.12a.1.II, TS. L. ET.12a.1.V, LS. Scale bar A–G, 5 mm; H, 3 mm; I, 1.9 mm; J–L, 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10626415" httpUri="https://zenodo.org/record/10626415/files/figure.png" pageId="12" pageNumber="63">Fig. 8A</figureCitation>
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<figureCitation id="13242A6FFFEC194CFB3DFC48329384DB" box="[1250,1260,936,953]" captionStart="Fig" captionStartId="2.[150,183,1190,1210]" captionTargetBox="[242,1443,169,1165]" captionTargetId="graphics-377@2.[242,1394,197,1165]" captionTargetPageId="2" captionText="Fig. 2. Simplified lithological columns of the main sampled sections with the stratigraphic range of the rugose corals in the Yılanlı Formation and the position of samples with foraminifers (DFZ7, MFZ1, MFZ2, MFZ3 referring to the biostratigraphic zones of Poty et al. 2006). The position of the D–C boundary, based on foraminiferal assemblage, is indicated by the double arrow." figureDoi="http://doi.org/10.5281/zenodo.10626406" httpUri="https://zenodo.org/record/10626406/files/figure.png" pageId="12" pageNumber="63">
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<subScript id="179B34AFFFEC194CFB3DFC48329384DB" attach="right" box="[1250,1260,936,953]" fontSize="7" pageId="12" pageNumber="63">2</subScript>
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</figureCitation>
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, C, D). They are thickened but their axial ends are sharp. They bend toward the cardinal fossula in the cardinal quadrants but are straighter in the counter quadrants. The initial thickening occurring in the whole tabularium rapidly decreases during the growth but maintains in the cardinal quadrants in the mature stages. The cardinal septum is shorter and thicker than its neighbouring septa. The minor septa are rudimentary or restricted to the peripheral part of the dissepimentarium. The cardinal fossula is well marked. The dissepimentarium counts 5–12 rows of concentric and herringbone dissepiments and several arched dissepiments in the peripheral part. Second order lonsdaleoid dissepiments occur in some specimens. The inner row of dissepiments is thickened in continuity with the septa (
|
||
<figureCitation id="13242A6FFFEC194CFB5FFA8032AB821B" box="[1152,1236,1376,1401]" captionStart="Fig" captionStartId="11.[96,129,1614,1634]" captionTargetBox="[98,1478,161,1587]" captionTargetId="figure-79@11.[98,1479,161,1588]" captionTargetPageId="11" captionText="Fig. 8. Devonian–Carboniferous rugose corals from Turkey. A–E. Caninophyllum charli sp. nov. from the lower Tournaisian (upper Hastarian) of Zonguldak, Gökgöl section. A. Holotype, G.8.1.1, successive TS (A1, A2); close-up view of the dissepimentarium, TS (A3). B. G.8.6.1, successive TS (B1, B2). C. G.8.4.3, TS. D. G.8.4.2, TS. E. G.8.3.2, LS. F, G. Uralinia simplex (Yü, 1933) from the lower Tournaisian (lower Hastarian) of Bartın, Topluca section. F. ET.9c.7, TS. G. ET.9c.8, TS. H. Amplexocarinia rozkowskae (Fedorowski, 2003) from the uppermost Famennian (Strunian) of Bartın, Dallıca section, D.2.4.II’, TS. I.?Metriophyllum sp. from the uppermost Famennian (Strunian) of Bartın, Topluca section, ET.11.X, TS. J–L. Bounophyllum praecursor (Frech, 1895) form the uppermost Famennian (Strunian) of Bartın, Topluca section. J. ET.12a.1.III, successive TS (J1–J3). K. ET.12a.1.II, TS. L. ET.12a.1.V, LS. Scale bar A–G, 5 mm; H, 3 mm; I, 1.9 mm; J–L, 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10626415" httpUri="https://zenodo.org/record/10626415/files/figure.png" pageId="12" pageNumber="63">Fig. 8A</figureCitation>
|
||
<figureCitation id="13242A6FFFEC194CFB0BFA8B32A2821E" box="[1236,1245,1387,1404]" captionStart="Fig" captionStartId="3.[98,131,662,682]" captionTargetBox="[99,1480,160,636]" captionTargetId="figure-697@3.[99,1481,160,636]" captionTargetPageId="3" captionText="Fig. 3. Uppermost Famennian (Strunian) facies in the Topluca section, Turkey. A. Lower stromatoporoid biostrome (unit ET12a in Fig. 2). B. Bioclastic facies of the unit ET11 crowded with large campophyllid solitary rugose corals in a packstone matrix." figureDoi="http://doi.org/10.5281/zenodo.10626408" httpUri="https://zenodo.org/record/10626408/files/figure.png" pageId="12" pageNumber="63">
|
||
<subScript id="179B34AFFFEC194CFB0BFA8B32A2821E" attach="left" box="[1236,1245,1387,1404]" fontSize="7" pageId="12" pageNumber="63">3</subScript>
|
||
</figureCitation>
|
||
). The wall is thin, regular or slightly undulating. In longitudinal section, the tabulae are incomplete, horizontal or domed in the axial part of the tabularium, domed or mesa-shaped in the peripheral part (
|
||
<figureCitation id="13242A6FFFEC194CFCB9FA0135C58298" box="[870,954,1505,1530]" captionStart="Fig" captionStartId="11.[96,129,1614,1634]" captionTargetBox="[98,1478,161,1587]" captionTargetId="figure-79@11.[98,1479,161,1588]" captionTargetPageId="11" captionText="Fig. 8. Devonian–Carboniferous rugose corals from Turkey. A–E. Caninophyllum charli sp. nov. from the lower Tournaisian (upper Hastarian) of Zonguldak, Gökgöl section. A. Holotype, G.8.1.1, successive TS (A1, A2); close-up view of the dissepimentarium, TS (A3). B. G.8.6.1, successive TS (B1, B2). C. G.8.4.3, TS. D. G.8.4.2, TS. E. G.8.3.2, LS. F, G. Uralinia simplex (Yü, 1933) from the lower Tournaisian (lower Hastarian) of Bartın, Topluca section. F. ET.9c.7, TS. G. ET.9c.8, TS. H. Amplexocarinia rozkowskae (Fedorowski, 2003) from the uppermost Famennian (Strunian) of Bartın, Dallıca section, D.2.4.II’, TS. I.?Metriophyllum sp. from the uppermost Famennian (Strunian) of Bartın, Topluca section, ET.11.X, TS. J–L. Bounophyllum praecursor (Frech, 1895) form the uppermost Famennian (Strunian) of Bartın, Topluca section. J. ET.12a.1.III, successive TS (J1–J3). K. ET.12a.1.II, TS. L. ET.12a.1.V, LS. Scale bar A–G, 5 mm; H, 3 mm; I, 1.9 mm; J–L, 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10626415" httpUri="https://zenodo.org/record/10626415/files/figure.png" pageId="12" pageNumber="63">Fig. 8E</figureCitation>
|
||
). A peripheral gutter develops in the mature stages. The dissepiments are 1.5–
|
||
<quantity id="4CE79B0FFFEC194CFB5EF9E132C08178" box="[1153,1215,1537,1562]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="63" unit="mm" value="3.0">3 mm</quantity>
|
||
long,
|
||
<quantity id="4CE79B0FFFEC194CFADCF9E1333C8178" box="[1283,1347,1537,1562]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="63" unit="mm" value="1.0">1 mm</quantity>
|
||
high and steeply declined toward the tabularium.
|
||
</paragraph>
|
||
<paragraph id="8BA036EAFFEC194DFC81F9AC369A86B9" blockId="12.[862,1535,1612,1960]" lastBlockId="13.[97,771,160,475]" lastPageId="13" lastPageNumber="64" pageId="12" pageNumber="63">
|
||
<emphasis id="B96BEAF8FFEC194CFC81F9AC35BF8107" box="[862,960,1612,1637]" italics="true" pageId="12" pageNumber="63">Remarks</emphasis>
|
||
.—The small size and number of septa are diagnostic of
|
||
<taxonomicName id="4C1F4D69FFEC194CFC13F98D32B881E4" authority="Denayer, 2016" authorityName="Denayer" authorityYear="2016" box="[972,1223,1645,1670]" class="Anthozoa" family="Bothrophyllidae" genus="Caninophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="63" phylum="Cnidaria" rank="species" species="charli" status="sp. nov.">
|
||
<emphasis id="B96BEAF8FFEC194CFC13F98D32B881E4" box="[972,1223,1645,1670]" italics="true" pageId="12" pageNumber="63">Caninophyllum charli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2585783FFEC194CFB0EF98D335581E4" box="[1233,1322,1645,1670]" pageId="12" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and allow its discrimination from
|
||
<taxonomicName id="4C1F4D69FFEC194CFBF4F96D35E581A4" authority="(Milne-Edwards and Haime, 1852)" baseAuthorityName="Milne-Edwards and Haime" baseAuthorityYear="1852" class="Anthozoa" family="Cyathophyllidae" genus="Cyathophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="63" phylum="Cnidaria" rank="species" species="archiaci">
|
||
<emphasis id="B96BEAF8FFEC194CFBF4F96D32D281C4" box="[1067,1197,1677,1702]" italics="true" pageId="12" pageNumber="63">C. archiaci</emphasis>
|
||
(Milne-Edwards and Haime, 1852)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1F4D69FFEC194CFC0BF94D334981A4" authority="(Tolmachev, 1931)" baseAuthorityName="Tolmachev" baseAuthorityYear="1931" box="[980,1334,1709,1734]" class="Anthozoa" family="Bothrophyllidae" genus="Caninophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="63" phylum="Cnidaria" rank="species" species="tomiense">
|
||
<emphasis id="B96BEAF8FFEC194CFC0BF94D322481A4" box="[980,1115,1709,1734]" italics="true" pageId="12" pageNumber="63">C. tomiense</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8E4B1BFFEC194CFBB4F94D335181A4" author="Tolmachev, I. P." box="[1131,1326,1709,1734]" pageId="12" pageNumber="63" pagination="1 - 663" refId="ref17050" refString="Tolmachev, I. P. 1931. Faune du Calcaire du Bassin du Kuznetsk. Comite geologique de Leningrad, materiaux pour la geologie generale et appliquee 25: 1 - 663." type="journal article" year="1931">Tolmachev, 1931</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. The new species shares its dimensions with
|
||
<taxonomicName id="4C1F4D69FFEC194CFB4FF92D33AB8184" authority="(Michelin, 1846)" baseAuthorityName="Michelin" baseAuthorityYear="1846" box="[1168,1492,1741,1766]" class="Anthozoa" family="Bothrophyllidae" genus="Caninophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="63" phylum="Cnidaria" rank="species" species="patulum">
|
||
<emphasis id="B96BEAF8FFEC194CFB4FF92D33718184" box="[1168,1294,1741,1766]" italics="true" pageId="12" pageNumber="63">C. patulum</emphasis>
|
||
(Michelin, 1846)
|
||
</taxonomicName>
|
||
but the latter show a larger dissepimentarium usually composed only of concentric and herringbone dissepiments. The septa are less thickened and more withdrawn toward the periphery in
|
||
<taxonomicName id="4C1F4D69FFEC194CFC77F8AE32588005" baseAuthorityName="Michelin" baseAuthorityYear="1846" box="[936,1063,1870,1895]" class="Anthozoa" family="Bothrophyllidae" genus="Caninophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="63" phylum="Cnidaria" rank="species" species="patulum">
|
||
<emphasis id="B96BEAF8FFEC194CFC77F8AE32588005" box="[936,1063,1870,1895]" italics="true" pageId="12" pageNumber="63">C. patulum</emphasis>
|
||
</taxonomicName>
|
||
while its cardinal septum is longer in the juvenile stage. The tabularium is also different: the tabulae are flat or slightly domed in the central part in
|
||
<taxonomicName id="4C1F4D69FFEC194CFA49F86F338180CA" authorityName="Denayer" authorityYear="2016" box="[1430,1534,1935,1960]" class="Anthozoa" family="Bothrophyllidae" genus="Caninophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="12" pageNumber="63" phylum="Cnidaria" rank="species" species="charli">
|
||
<emphasis id="B96BEAF8FFEC194CFA49F86F338180CA" box="[1430,1534,1935,1960]" italics="true" pageId="12" pageNumber="63">C. charli</emphasis>
|
||
</taxonomicName>
|
||
but are clearly domed in
|
||
<taxonomicName id="4C1F4D69FFED194DFE55FF40347387DB" baseAuthorityName="Michelin" baseAuthorityYear="1846" box="[394,524,160,185]" class="Anthozoa" family="Bothrophyllidae" genus="Caninophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="64" phylum="Cnidaria" rank="species" species="patulum">
|
||
<emphasis id="B96BEAF8FFED194DFE55FF40347387DB" box="[394,524,160,185]" italics="true" pageId="13" pageNumber="64">C. patulum</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="EF8E4B1BFFED194DFDC2FF40357D87DB" author="Poty, E. & Xu, S." box="[541,770,160,185]" pageId="13" pageNumber="64" pagination="89 - 139" refId="ref16110" refString="Poty, E. and Xu, S. 1996. Rugosa from the Devonian - Carboniferous transition in Hunan, China. Memoires de l'Institut de Geologie de l'Universite de Louvain 36: 89 - 139." type="journal article" year="1996">Poty and Xu (1996)</bibRefCitation>
|
||
figured two species of small
|
||
<taxonomicName id="4C1F4D69FFED194DFE78FF20342B87BB" authorityName="Lewis" authorityYear="1929" box="[423,596,192,217]" class="Anthozoa" family="Bothrophyllidae" genus="Caninophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="64" phylum="Cnidaria" rank="genus">
|
||
<emphasis id="B96BEAF8FFED194DFE78FF20342B87BB" box="[423,596,192,217]" italics="true" pageId="13" pageNumber="64">Caninophyllum</emphasis>
|
||
</taxonomicName>
|
||
from the lower Tournaisian
|
||
<taxonomicName id="4C1F4D69FFED194DFF2EFF003432879B" authority="Zone" authorityName="Zone" box="[241,589,224,249]" class="Anthozoa" family="Uraliniidae" genus="Uralinia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="64" phylum="Cnidaria" rank="species" species="tangpakouensis">
|
||
<emphasis id="B96BEAF8FFED194DFF2EFF003473879B" box="[241,524,224,249]" italics="true" pageId="13" pageNumber="64">Uralinia tangpakouensis</emphasis>
|
||
Zone
|
||
</taxonomicName>
|
||
of S
|
||
<collectingCountry id="F308767AFFED194DFD4CFF0034A6879B" box="[659,729,224,249]" name="China" pageId="13" pageNumber="64">China</collectingCountry>
|
||
.
|
||
<taxonomicName id="4C1F4D69FFED194DFD37FF00372E867B" authority="Jiang, 1982" authorityName="Jiang" authorityYear="1982" class="Anthozoa" family="Bothrophyllidae" genus="Caninophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="64" phylum="Cnidaria" rank="species" species="cystosum">
|
||
<emphasis id="B96BEAF8FFED194DFD37FF0036B6867B" italics="true" pageId="13" pageNumber="64">C. cystosum</emphasis>
|
||
<bibRefCitation id="EF8E4B1BFFED194DFF0EFEE0372E867B" author="Jiang, S." box="[209,337,256,281]" pageId="13" pageNumber="64" refId="ref14407" refString="Jiang, S. 1982. Class Anthozoa. The Palaeontological Atlas of Hunan. 189 pp. Geological Publishing House, Beijing." type="book" year="1982">Jiang, 1982</bibRefCitation>
|
||
</taxonomicName>
|
||
has 41–46 septa for a diameter of
|
||
<quantity id="4CE79B0FFFED194DFD07FEE036CE8658" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="13" pageNumber="64" unit="mm" value="30.0" valueMax="35.0" valueMin="25.0">25– 35 mm</quantity>
|
||
but differs from
|
||
<taxonomicName id="4C1F4D69FFED194DFEA7FEC137A18658" authorityName="Denayer" authorityYear="2016" box="[376,478,289,314]" class="Anthozoa" family="Bothrophyllidae" genus="Caninophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="64" phylum="Cnidaria" rank="species" species="charli">
|
||
<emphasis id="B96BEAF8FFED194DFEA7FEC137A18658" box="[376,478,289,314]" italics="true" pageId="13" pageNumber="64">C. charli</emphasis>
|
||
</taxonomicName>
|
||
by an inconspicuous cardinal fossula and a long counter septum often connected to the cardinal septum.
|
||
<taxonomicName id="4C1F4D69FFED194DFE92FE8134EE8618" authority="Jiang, 1982" authorityName="Jiang" authorityYear="1982" box="[333,657,353,378]" class="Anthozoa" family="Bothrophyllidae" genus="Caninophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="64" phylum="Cnidaria" rank="species" species="shaoyangense">
|
||
<emphasis id="B96BEAF8FFED194DFE92FE8134748618" box="[333,523,353,378]" italics="true" pageId="13" pageNumber="64">C. shaoyangense</emphasis>
|
||
<bibRefCitation id="EF8E4B1BFFED194DFDCDFE8134EE8618" author="Jiang, S." box="[530,657,353,378]" pageId="13" pageNumber="64" refId="ref14407" refString="Jiang, S. 1982. Class Anthozoa. The Palaeontological Atlas of Hunan. 189 pp. Geological Publishing House, Beijing." type="book" year="1982">Jiang, 1982</bibRefCitation>
|
||
</taxonomicName>
|
||
is smaller
|
||
<quantity id="4CE79B0FFFED194DFFB6FE6136C686F8" box="[105,185,385,410]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="13" pageNumber="64" unit="mm" value="17.0">17 mm</quantity>
|
||
and 38 septa of both orders) and has a very large dissepimentarium and septa almost not thickened in the tabularium.
|
||
</paragraph>
|
||
<paragraph id="8BA036EAFFED194DFFBDFE0D34D685E5" blockId="13.[98,771,493,647]" pageId="13" pageNumber="64">
|
||
<emphasis id="B96BEAF8FFED194DFFBDFE0D34528564" box="[98,557,493,518]" italics="true" pageId="13" pageNumber="64">Stratigraphic and geographic range</emphasis>
|
||
.—
|
||
<taxonomicName id="4C1F4D69FFED194DFD8EFE0D36DA8544" authority="Denayer, 2016" authorityName="Denayer" authorityYear="2016" class="Anthozoa" family="Bothrophyllidae" genus="Caninophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="64" phylum="Cnidaria" rank="species" species="charli" status="sp. nov.">
|
||
<emphasis id="B96BEAF8FFED194DFD8EFE0D36DA8544" italics="true" pageId="13" pageNumber="64">Caninophyllum charli</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2585783FFED194DFF74FDED377F8544" box="[171,256,525,550]" pageId="13" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
|
||
was collected in a bioclastic level dated of the
|
||
<geologicalTimeScale id="F93FB641FFED194DFFBDFDCD37508524" box="[98,303,557,582]" pageId="13" pageNumber="64">lower Tournaisian</geologicalTimeScale>
|
||
(Hastarian) by foraminifers (MFZ3 biozone) in the
|
||
<collectingMunicipality id="6BC4AC90FFED194DFF2FFDAD373C8504" box="[240,323,589,614]" pageId="13" pageNumber="64">Gökgöl</collectingMunicipality>
|
||
(
|
||
<collectingRegion id="49DBF808FFED194DFE8CFDAD37AC8504" box="[339,467,589,614]" country="Turkey" name="Zonguldak" pageId="13" pageNumber="64">Zonguldak</collectingRegion>
|
||
). Crushed specimens were collected in equivalent levels in the
|
||
<location id="8EC06031FFED194DFE29FD8E343885E5" LSID="urn:lsid:plazi:treatment:03B687FCFFEC194DFF48F8FC34D685E5:8EC06031FFED194DFE29FD8E343885E5" box="[502,583,622,647]" county="lower Tournaisian" municipality="Gokgol" name="Dallica" pageId="13" pageNumber="64" stateProvince="Zonguldak">Dallıca</location>
|
||
(
|
||
<collectingRegion id="49DBF808FFED194DFD89FD8E34DF85E5" box="[598,672,622,647]" country="Turkey" name="Bartin" pageId="13" pageNumber="64">Bartın</collectingRegion>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |