381 lines
49 KiB
XML
381 lines
49 KiB
XML
<document id="7981D7C1F717DE888F96EF44F029BFF7" ID-CLB-Dataset="5606" ID-DOI="10.11646/zootaxa.3795.5.1" ID-GBIF-Dataset="27b8f170-95d1-4132-a493-c0b78c109163" ID-ISSN="1175-5326" ID-Zenodo-Dep="4915064" ID-ZooBank="3034E613-829A-4E56-A860-CA2A7C23B8FA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623197981020" checkinUser="felipe" docAuthor="Fritz, Uwe, Petzold, Alice, Kehlmaier, Christian, Kindler, Carolin, Campbell, Patrick, Hofmeyr, Margaretha D. & Branch, William R." docDate="2014" docId="03B68780CF59102018C3EDFCFBBFFB03" docLanguage="en" docName="zt03795p522.pdf" docOrigin="Zootaxa 3795 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pentonyx gehafie Ruppell 1835" docType="treatment" docVersion="8" lastPageNumber="513" masterDocId="FF8FFFF8CF50102C1854EB39FF7EFFA3" masterDocTitle="Disentangling the Pelomedusa complex using type specimens and historical DNA (Testudines: Pelomedusidae)" masterLastPageNumber="522" masterPageNumber="501" pageNumber="510" updateTime="1699006212259" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="0B204DEE55B7CAF31803770EFA0259A6">Disentangling the Pelomedusa complex using type specimens and historical DNA (Testudines: Pelomedusidae)</mods:title>
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<mods:namePart id="25FDE5805B1B80AF20AB9A900D8C231A">Fritz, Uwe</mods:namePart>
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<mods:affiliation id="8D76453B4C5E83D8676A8B8A9D1939DE">Museum of Zoology, Senckenberg Dresden, A. B. Meyer Building, 01109 Dresden, Germany</mods:affiliation>
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<mods:nameIdentifier id="313BBE1DBF04F3C6981DDEFC30AD134D" type="email">uwe.fritz@senckenberg.de</mods:nameIdentifier>
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<mods:namePart id="F831EB7414014F7BC839303937D33ED1">Petzold, Alice</mods:namePart>
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<mods:affiliation id="52BCB6CA59EB5A9D5AB94DE495A54B7A">Museum of Zoology, Senckenberg Dresden, A. B. Meyer Building, 01109 Dresden, Germany</mods:affiliation>
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<mods:namePart id="ACC7CB556CF68679A8D8C6F2F814437A">Kehlmaier, Christian</mods:namePart>
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<mods:affiliation id="340523C7EB39E0C4D1DDBDEC44E0D9E2">Museum of Zoology, Senckenberg Dresden, A. B. Meyer Building, 01109 Dresden, Germany</mods:affiliation>
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<mods:namePart id="2CA39368383D384CBDD026C63CF3015F">Kindler, Carolin</mods:namePart>
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<mods:affiliation id="4F76594C11BB23A31E1C1590AAD02CA6">Museum of Zoology, Senckenberg Dresden, A. B. Meyer Building, 01109 Dresden, Germany</mods:affiliation>
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<mods:namePart id="C7AF70C316E224358EA5EC7E9CA908BB">Campbell, Patrick</mods:namePart>
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<mods:affiliation id="35EB03E48336A0537AE984A3E3C4E210">Department of Zoology, Darwin Centre (DC 1), Natural History Museum, Cromwell Road, South Kensington, London SW 7 5 BD, England</mods:affiliation>
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<mods:namePart id="E4941B9F53846359BCAF384822FA0EBB">Hofmeyr, Margaretha D.</mods:namePart>
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<mods:affiliation id="41560E2623E7E861207175AB64534746">Chelonian Biodiversity and Conservation, Department of Biodiversity and Conservation Biology, University of the Western Cape, Private Bag X 17, Bellville 7535, South Africa</mods:affiliation>
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<mods:namePart id="1E7853432C958C60F9D895BE75B87292">Branch, William R.</mods:namePart>
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<mods:affiliation id="F2F67AD95366986BB30B2DD3BD293A5C">Department of Herpetology, Port Elizabeth Museum, P. O. Box 13147, Humewood 6013, South Africa & Department of Zoology, P. O. Box 77000, Nelson Mandela Metropolitan University, Port Elizabeth 6031, South Africa Corresponding author. E-mail: uwe. fritz @ senckenberg. de</mods:affiliation>
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<treatment id="03B68780CF59102018C3EDFCFBBFFB03" ID-DOI="http://doi.org/10.5281/zenodo.5082885" ID-GBIF-Taxon="183403832" ID-Zenodo-Dep="5082885" LSID="urn:lsid:plazi:treatment:03B68780CF59102018C3EDFCFBBFFB03" httpUri="http://treatment.plazi.org/id/03B68780CF59102018C3EDFCFBBFFB03" lastPageId="12" lastPageNumber="513" pageId="9" pageNumber="510">
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<taxonomicName id="4C1F4D15CF59102518C3EDFCFD60F97C" ID-CoL="8SJZW" authority="Ruppell, 1835" authorityName="Ruppell" authorityYear="1835" box="[151,542,1733,1759]" class="Testudines" family="Pelomedusidae" genus="Pentonyx" higherTaxonomySource="CoL" kingdom="Animalia" order="Testudines" pageId="9" pageNumber="510" phylum="Chordata" rank="species" species="gehafie">
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<emphasis id="B96BEA84CF59102518C3EDFCFD60F97C" bold="true" box="[151,542,1733,1759]" pageId="9" pageNumber="510">
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<emphasis id="B96BEA84CF59102518C3EDFCFE17F97C" bold="true" box="[151,361,1733,1759]" italics="true" pageId="9" pageNumber="510">Pentonyx gehafie</emphasis>
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<bibRefCitation id="EF8E4B67CF5910251925EDFCFD60F97C" author="Ruppell, E." box="[369,542,1733,1759]" pageId="9" pageNumber="510" refId="ref13026" refString="Ruppell, E. (1835) Neue Wirbelthiere zu der Fauna von Abyssinien gehorig. Amphibien. Siegmund Schmerber, Frankfurt am Main, 18 pp., 6 plates. http: // dx. doi. org / 10.5962 / bhl. title. 53778" type="book" year="1835">Rüppell, 1835</bibRefCitation>
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</emphasis>
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</taxonomicName>
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<paragraph id="8BA03696CF59102618C3EC35FE5AFE56" blockId="9.[151,1437,1804,2009]" lastBlockId="10.[151,1436,151,501]" lastPageId="10" lastPageNumber="511" pageId="9" pageNumber="510">
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<bibRefCitation id="EF8E4B67CF59102518C3EC35FE42F886" author="Ruppell, E." box="[151,316,1804,1829]" pageId="9" pageNumber="510" refId="ref13026" refString="Ruppell, E. (1835) Neue Wirbelthiere zu der Fauna von Abyssinien gehorig. Amphibien. Siegmund Schmerber, Frankfurt am Main, 18 pp., 6 plates. http: // dx. doi. org / 10.5962 / bhl. title. 53778" type="book" year="1835">Rüppell (1835</bibRefCitation>
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: pp. 2–3) described
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<taxonomicName id="4C1F4D15CF5910251A48EC34FDA0F886" authorityName="Ruppell" authorityYear="1835" box="[540,734,1804,1829]" class="Testudines" family="Pelomedusidae" genus="Pentonyx" higherTaxonomySource="CoL" kingdom="Animalia" order="Testudines" pageId="9" pageNumber="510" phylum="Chordata" rank="species" species="gehafie">
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<emphasis id="B96BEA84CF5910251A48EC34FDA0F886" box="[540,734,1804,1829]" italics="true" pageId="9" pageNumber="510">Pentonyx gehafie</emphasis>
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</taxonomicName>
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based on terrapins which he found “häufig in allen fliessenden oder stagnirenden Gewässern auf dem östlichen Abhange der abyssinischen Küstengebirge” [frequently in all flowing and stagnant waters on the eastern slope of the Abyssinian coastal mountains]. Rüppell diagnosed
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<taxonomicName id="4C1F4D15CF59102518C3EC40FE25F833" authorityName="Ruppell" authorityYear="1835" box="[151,347,1911,1936]" class="Testudines" family="Pelomedusidae" genus="Pentonyx" higherTaxonomySource="CoL" kingdom="Animalia" order="Testudines" pageId="9" pageNumber="510" phylum="Chordata" rank="species" species="gehafie">
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<emphasis id="B96BEA84CF59102518C3EC40FE25F833" box="[151,347,1911,1936]" italics="true" pageId="9" pageNumber="510">Pentonyx gehafie</emphasis>
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</taxonomicName>
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from the
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<taxonomicName id="4C1F4D15CF5910251984EC40FDEDF833" box="[464,659,1911,1936]" class="Testudines" family="Pelomedusidae" genus="Pentonyx" higherTaxonomySource="CoL" kingdom="Animalia" order="Testudines" pageId="9" pageNumber="510" phylum="Chordata" rank="species" species="undetermined">
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<emphasis id="B96BEA84CF5910251984EC40FD46F833" box="[464,568,1913,1936]" italics="true" pageId="9" pageNumber="510">Pentonyx</emphasis>
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species
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</taxonomicName>
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occurring at the
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<collectingRegion id="49DBF874CF5910251B0FEC4EFCEBF833" box="[859,917,1911,1936]" country="South Africa" name="Western Cape" pageId="9" pageNumber="510">Cape</collectingRegion>
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of Good Hope by the plastral scutation: The pectoral scutes of his new species are triangular and do not reach the median plastral seam, whereas in the
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<collectingRegion id="49DBF874CF5910251D37ECA5FAE3F816" box="[1379,1437,1948,1973]" country="South Africa" name="Western Cape" pageId="9" pageNumber="510">Cape</collectingRegion>
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species the pectoral scutes are in contact. The character state of the Abyssinian specimens is nicely depicted in Rüppell’s plate 1. Obviously Rüppell collected quite a number of helmeted terrapins, which originally had all
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<typeStatus id="54A48834CF5A102618C3EB85FF90FF77" box="[151,238,188,212]" pageId="10" pageNumber="511" type="syntype">syntype</typeStatus>
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status.
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<bibRefCitation id="EF8E4B67CF5A10261913EB85FE8FFF76" author="Mertens, R." box="[327,497,188,213]" pageId="10" pageNumber="511" pagination="139 - 142" refId="ref12650" refString="Mertens, R. (1937) Bemerkungen uber die Rassen von Pelomedusa subrufa (La Cepede). Zoologischer Anzeiger, 117, 139 - 142." type="journal article" year="1937">Mertens (1937</bibRefCitation>
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: p. 140) mentions that seven of these specimens are in the Senckenberg Museum, Frankfurt am Main. However, there are eight, and not seven, terrapins collected by Rüppell with identical data (SMF 7947–7949, SMF 7953, SMF 7960–7962, SMF 33054). Whilst SMF 7947–7949 and SMF 7960–7962 are alcohol-preserved, SMF 7953 and SMF 33054 are dry specimens (
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<tableCitation id="C69D032DCF5A10261BD5EA1EFCA6FEE3" box="[897,984,295,320]" captionStart="TABLE 2" captionStartId="4.[147,233,594,616]" captionTargetPageId="4" captionText="TABLE 2. Pelomedusa type specimens studied genetically. Lengths of successfully sequenced mtDNA fragments indicated. For GenBank accession numbers, see Table S1. The cyt b sequence of ZFMK 17076 was generated by Wong et al. (2010). For museum acronyms, see Table 1." httpUri="http://table.plazi.org/id/DF60661ECF54102818C7E96BFBB0FD26" pageId="10" pageNumber="511" tableUuid="DF60661ECF54102818C7E96BFBB0FD26">Table 2</tableCitation>
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). SMF 7947 was designated by
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<bibRefCitation id="EF8E4B67CF5A10261D14EA11FFA2FEC6" author="Mertens, R." pageId="10" pageNumber="511" pagination="139 - 142" refId="ref12650" refString="Mertens, R. (1937) Bemerkungen uber die Rassen von Pelomedusa subrufa (La Cepede). Zoologischer Anzeiger, 117, 139 - 142." type="journal article" year="1937">Mertens (1937</bibRefCitation>
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: p. 140) as
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<typeStatus id="54A48834CF5A10261935EA75FEB4FEC6" box="[353,458,332,357]" pageId="10" pageNumber="511" type="lectotype">lectotype</typeStatus>
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, which is why the remaining terrapins bear today the status of
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<typeStatus id="54A48834CF5A10261CCDEA75FA40FEC6" box="[1177,1342,332,357]" pageId="10" pageNumber="511" type="paralectotype">paralectotypes</typeStatus>
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. Further
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<typeStatus id="54A48834CF5A102618C3EA56FE47FE2B" box="[151,313,367,392]" pageId="10" pageNumber="511" type="paralectotype">paralectotypes</typeStatus>
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collected by Rüppell are in the Natural History Museum, London (
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<bibRefCitation id="EF8E4B67CF5A10261C18EA56FBB2FE2B" author="Gray, J. E." box="[1100,1228,367,392]" pageId="10" pageNumber="511" refId="ref12142" refString="Gray, J. E. (1856 [" 1855 "]) Catalogue of Shield Reptiles in the Collection of the British Museum. Part I. Testudinata (Tortoises). British Museum, London, (1) + 79 + (2) pp., 42 plates. http: // dx. doi. org / 10.5962 / bhl. title. 5491" type="book" year="1856">Gray 1856</bibRefCitation>
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: p. 53;
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<bibRefCitation id="EF8E4B67CF5A10261D7CEA56FFADFE0E" author="Boulenger, G. A." pageId="10" pageNumber="511" refId="ref11241" refString="Boulenger, G. A. (1889) Catalogue of the Chelonians, Rhynchocephalians, and Crocodiles in the British Museum (Natural History). New Edition. British Museum (Natural History), London, x + 311 pp., 6 plates. http: // dx. doi. org / 10.5962 / bhl. title. 13541" type="book" year="1889">Boulenger 1889</bibRefCitation>
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: p. 199) and the Muséum National d’Histoire naturelle, Paris (see below under
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<taxonomicName id="4C1F4D15CF5A10261CD4EAADFD36FE73" authority="Vaillant & Grandidier, 1910" authorityName="Vaillant & Grandidier" authorityYear="1910" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="511" phylum="Chordata" rank="variety" species="galeata" variety="disjuncta">
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<emphasis id="B96BEA84CF5A10261CD4EAADFA16FE0E" box="[1152,1384,404,429]" italics="true" pageId="10" pageNumber="511">Pelomedusa galeata</emphasis>
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var.
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<emphasis id="B96BEA84CF5A102618C3EA8EFF80FE73" box="[151,254,439,464]" italics="true" pageId="10" pageNumber="511">disjuncta</emphasis>
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<bibRefCitation id="EF8E4B67CF5A10261952EA8EFD36FE73" author="Vaillant, L. & Grandidier, G." box="[262,584,439,464]" pageId="10" pageNumber="511" pagination="1 - 86" refId="ref13530" refString="Vaillant, L. & Grandidier, G. (1910) Volume XVII. Histoire naturelle des reptiles. Premiere partie: Crocodiles et tortues. In: Grandidier, A. & Grandidier, G. (Eds.), Histoire physique, naturelle et politique de Madagascar. Imprimerie Nationale, Paris, pp. 1 - 86, plates 1 - 27." type="book chapter" year="1910">Vaillant & Grandidier, 1910</bibRefCitation>
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</taxonomicName>
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). The
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<typeStatus id="54A48834CF5A10261AC5EA8EFD89FE73" box="[657,759,439,464]" pageId="10" pageNumber="511" type="lectotype">lectotype</typeStatus>
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is a subadult female of
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<quantity id="4CE79B73CF5A10261C5AEA8EFB08FE6C" box="[1038,1142,439,464]" metricMagnitude="-1" metricUnit="m" metricValue="1.142" pageId="10" pageNumber="511" unit="cm" value="11.42">11.42 cm</quantity>
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straight carapacial length (
|
||
<figureCitation id="13242A13CF5A102618CBEAE5FF9DFE56" box="[159,227,476,501]" captionStart="FIGURE 5" captionStartId="11.[151,250,1953,1975]" captionTargetBox="[236,1375,210,1925]" captionTargetId="figure-77@11.[188,1399,193,1932]" captionTargetPageId="11" captionText="FIGURE 5. Dorsal and ventral views of the lectotype of Pentonyx gehafie Rüppell, 1835 (SMF 7947, subadult female, straight carapacial length 11.42 cm; top) and of one of the paralectotypes in the Natural History Museum, London (BMNH 1947.3.5.78, adult male, straight carapacial length 16.70 cm; bottom). Scale bars, 3 cm. Photos: E. Morawa, P. Campbell." figureDoi="http://doi.org/10.5281/zenodo.4915074" httpUri="https://zenodo.org/record/4915074/files/figure.png" pageId="10" pageNumber="511">Fig. 5</figureCitation>
|
||
, top).
|
||
</paragraph>
|
||
<caption id="DF60661ECF5A102618C3ED88FCD8F945" ID-DOI="http://doi.org/10.5281/zenodo.4915072" ID-Zenodo-Dep="4915072" httpUri="https://zenodo.org/record/4915072/files/figure.png" pageId="10" pageNumber="511" startId="10.[151,250,1713,1735]" targetBox="[235,1368,561,1646]" targetPageId="10">
|
||
<paragraph id="8BA03696CF5A102618C3ED88FCD8F945" blockId="10.[151,1436,1713,1766]" pageId="10" pageNumber="511">
|
||
<emphasis id="B96BEA84CF5A102618C3ED88FE6BF965" bold="true" box="[151,277,1713,1735]" pageId="10" pageNumber="511">FIGURE 4.</emphasis>
|
||
Dorsal and ventral view of the lectotype of
|
||
<taxonomicName id="4C1F4D15CF5A10261ABAED8BFBC4F964" authority="Dumeril & Bibron, 1835" authorityName="Dumeril & Bibron" authorityYear="1835" box="[750,1210,1713,1735]" class="Testudines" family="Pelomedusidae" genus="Pentonyx" higherTaxonomySource="CoL" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="511" phylum="Chordata" rank="species" species="capensis">
|
||
<emphasis id="B96BEA84CF5A10261ABAED8BFCD3F964" box="[750,941,1714,1735]" italics="true" pageId="10" pageNumber="511">Pentonyx capensis</emphasis>
|
||
<bibRefCitation id="EF8E4B67CF5A10261BE2ED88FBC4F964" author="Dumeril, A. M. C. & Bibron, G." box="[950,1210,1713,1735]" pageId="10" pageNumber="511" refId="ref11700" refString="Dumeril, A. M. C. & Bibron, G. (1835) Erpetologie generale ou histoire naturelle complete des reptiles. Tome second. Librairie encyclopedique de Roret, Paris, 680 pp." type="book" year="1835">Duméril & Bibron, 1835</bibRefCitation>
|
||
</taxonomicName>
|
||
(MNHN 9506, male, straight carapacial length 23.45 cm). Scale bar, 3 cm. Photos: M.D. Hofmeyr.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA03696CF5A10261893EC2AFBD4F818" blockId="10.[151,1436,1810,1979]" pageId="10" pageNumber="511">
|
||
<materialsCitation id="3B773CCBCF5A10261893EC2AFBD8F818" ID-GBIF-Occurrence="3319793302" collectionCode="ICZN" country="Eritrea" county="International Code of Zoological Nomenclature" location="However" municipality="Article" pageId="10" pageNumber="511" specimenCode="ICZN 1999" specimenCount="5">
|
||
<bibRefCitation id="EF8E4B67CF5A10261893EC2AFE11F888" author="Mertens, R." box="[199,367,1810,1835]" pageId="10" pageNumber="511" pagination="139 - 142" refId="ref12650" refString="Mertens, R. (1937) Bemerkungen uber die Rassen von Pelomedusa subrufa (La Cepede). Zoologischer Anzeiger, 117, 139 - 142." type="journal article" year="1937">Mertens (1937</bibRefCitation>
|
||
: p. 140) restricted the
|
||
<typeStatus id="54A48834CF5A10261A33EC2AFDE9F888" box="[615,663,1811,1835]" pageId="10" pageNumber="511">type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="4C1F4D15CF5A10261B43EC2AFCA6F888" authorityName="Ruppell" authorityYear="1835" box="[791,984,1810,1835]" class="Testudines" family="Pelomedusidae" genus="Pentonyx" higherTaxonomySource="CoL" kingdom="Animalia" order="Testudines" pageId="10" pageNumber="511" phylum="Chordata" rank="species" species="gehafie">
|
||
<emphasis id="B96BEA84CF5A10261B43EC2AFCA6F888" box="[791,984,1810,1835]" italics="true" pageId="10" pageNumber="511">Pentonyx gehafie</emphasis>
|
||
</taxonomicName>
|
||
to “Massaua” [also known as Massawa, Missiwa or Mitsiwa,
|
||
<collectingCountry id="F3087606CF5A102619D7EC0FFEABF8EC" box="[387,469,1846,1871]" name="Eritrea" pageId="10" pageNumber="511">Eritrea</collectingCountry>
|
||
], a town within the source region of the
|
||
<typeStatus id="54A48834CF5A10261BC6EC0EFCBCF8EC" box="[914,962,1847,1871]" pageId="10" pageNumber="511">type</typeStatus>
|
||
specimens.
|
||
<location id="8EC0604DCF5A10261C1EEC0EFBCCF8ED" LSID="urn:lsid:plazi:treatment:03B68780CF59102018C3EDFCFBBFFB03:8EC0604DCF5A10261C1EEC0EFBCCF8ED" box="[1098,1202,1847,1870]" country="Eritrea" county="International Code of Zoological Nomenclature" municipality="Article" name="However" pageId="10" pageNumber="511">However</location>
|
||
, according to
|
||
<collectingMunicipality id="6BC4ACECCF5A10261D1AEC0FFAE2F8EC" box="[1358,1436,1846,1871]" pageId="10" pageNumber="511">Article</collectingMunicipality>
|
||
76.1 of the
|
||
<collectingCounty id="62C14E1ACF5A10261941EC63FC56F8D0" box="[277,808,1882,1907]" pageId="10" pageNumber="511">International Code of Zoological Nomenclature</collectingCounty>
|
||
(
|
||
<specimenCode id="DBB99EEDCF5A10261B62EC63FCC5F8D0" box="[822,955,1882,1907]" collectionCode="ICZN" pageId="10" pageNumber="511">ICZN 1999</specimenCode>
|
||
), the
|
||
<typeStatus id="54A48834CF5A10261BA3EC62FB59F8D0" box="[1015,1063,1883,1907]" pageId="10" pageNumber="511">type</typeStatus>
|
||
locality “is the geographical […] place of capture, collection or observation of the name-bearing
|
||
<typeStatus id="54A48834CF5A10261B37EC47FCE6F834" box="[867,920,1918,1943]" pageId="10" pageNumber="511">type</typeStatus>
|
||
.” Thus, Rüppell’s original
|
||
<typeStatus id="54A48834CF5A10261C92EC46FB88F834" box="[1222,1270,1919,1943]" pageId="10" pageNumber="511">type</typeStatus>
|
||
locality has to remain unchanged and is to be identified with the eastern slope of coastal mountains,
|
||
<collectingCountry id="F3087606CF5A10261C02EC9BFBD8F818" box="[1110,1190,1954,1979]" name="Eritrea" pageId="10" pageNumber="511">Eritrea</collectingCountry>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF60661ECF5B102718C3EC98FBA0F857" ID-DOI="http://doi.org/10.5281/zenodo.4915074" ID-Zenodo-Dep="4915074" httpUri="https://zenodo.org/record/4915074/files/figure.png" pageId="11" pageNumber="512" startId="11.[151,250,1953,1975]" targetBox="[236,1375,210,1925]" targetPageId="11">
|
||
<paragraph id="8BA03696CF5B102718C3EC98FBA0F857" blockId="11.[151,1436,1953,2036]" pageId="11" pageNumber="512">
|
||
<emphasis id="B96BEA84CF5B102718C3EC98FE6CF814" bold="true" box="[151,274,1953,1975]" pageId="11" pageNumber="512">FIGURE 5.</emphasis>
|
||
Dorsal and ventral views of the lectotype of
|
||
<taxonomicName id="4C1F4D15CF5B10271A8CEC9BFB63F814" authority="Ruppell, 1835" authorityName="Ruppell" authorityYear="1835" box="[728,1053,1953,1975]" class="Testudines" family="Pelomedusidae" genus="Pentonyx" higherTaxonomySource="CoL" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="512" phylum="Chordata" rank="species" species="gehafie">
|
||
<emphasis id="B96BEA84CF5B10271A8CEC9BFCF9F814" box="[728,903,1953,1975]" italics="true" pageId="11" pageNumber="512">Pentonyx gehafie</emphasis>
|
||
<bibRefCitation id="EF8E4B67CF5B10271BD9EC98FB63F814" author="Ruppell, E." box="[909,1053,1953,1975]" pageId="11" pageNumber="512" refId="ref13026" refString="Ruppell, E. (1835) Neue Wirbelthiere zu der Fauna von Abyssinien gehorig. Amphibien. Siegmund Schmerber, Frankfurt am Main, 18 pp., 6 plates. http: // dx. doi. org / 10.5962 / bhl. title. 53778" type="book" year="1835">Rüppell, 1835</bibRefCitation>
|
||
</taxonomicName>
|
||
(SMF 7947, subadult female, straight carapacial length 11.42 cm; top) and of one of the paralectotypes in the Natural History Museum, London (BMNH 1947.3.5.78, adult male, straight carapacial length 16.70 cm; bottom). Scale bars, 3 cm. Photos: E. Morawa, P. Campbell.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA03696CF5C10201893EBAEFA24FCB6" blockId="12.[151,1436,151,1184]" pageId="12" pageNumber="513">
|
||
<bibRefCitation id="EF8E4B67CF5C10201893EBAEFEF9FF13" author="Boulenger, G. A." box="[199,391,151,176]" pageId="12" pageNumber="513" pagination="146 - 151" refId="ref11208" refString="Boulenger, G. A. (1880) Sur l'existence d'une seule espece du genre Pelomedusa Wagler. Bulletin de la Societe zoologique de France, 5, 146 - 151." type="journal article" year="1880">Boulenger (1880</bibRefCitation>
|
||
: p. 151) synonymized
|
||
<taxonomicName id="4C1F4D15CF5C10201AD7EBAEFC1BFF13" baseAuthorityName="Rueppell" baseAuthorityYear="1835" box="[643,869,151,176]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="513" phylum="Chordata" rank="species" species="gehafie">
|
||
<emphasis id="B96BEA84CF5C10201AD7EBAEFC1BFF13" box="[643,869,151,176]" italics="true" pageId="12" pageNumber="513">Pelomedusa gehafie</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="4C1F4D15CF5C10201BF0EBAEFBF9FF13" baseAuthorityName="Schoepff" baseAuthorityYear="1792" box="[932,1159,151,176]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="513" phylum="Chordata" rank="species" species="galeata">
|
||
<emphasis id="B96BEA84CF5C10201BF0EBAEFBF9FF13" box="[932,1159,151,176]" italics="true" pageId="12" pageNumber="513">Pelomedusa galeata</emphasis>
|
||
</taxonomicName>
|
||
(=
|
||
<taxonomicName id="4C1F4D15CF5C10201CF9EBAEFAECFF13" baseAuthorityName="Bonnaterre" baseAuthorityYear="1789" box="[1197,1426,151,176]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="513" phylum="Chordata" rank="species" species="subrufa">
|
||
<emphasis id="B96BEA84CF5C10201CF9EBAEFAECFF13" box="[1197,1426,151,176]" italics="true" pageId="12" pageNumber="513">Pelomedusa subrufa</emphasis>
|
||
</taxonomicName>
|
||
) because he studied specimens with intermediate plastral characters from
|
||
<collectingCountry id="F3087606CF5C10201B9BEB85FB26FF76" box="[975,1112,188,213]" name="Madagascar" pageId="12" pageNumber="513">Madagascar</collectingCountry>
|
||
, the Upper Nile Region, and
|
||
<collectingCountry id="F3087606CF5C102018C3EBE6FF83FF5B" box="[151,253,223,248]" name="Tanzania" pageId="12" pageNumber="513">Zanzibar</collectingCountry>
|
||
, even though he acknowledged that separated triangular pectorals never occur in South African terrapins and that in
|
||
<collectingCountry id="F3087606CF5C10201971EA3DFEEBFEBE" box="[293,405,260,285]" name="Ethiopia" pageId="12" pageNumber="513">Abyssinia</collectingCountry>
|
||
the terrapins have consistently widely separated triangular pectorals. Based on similar observations, but drawing a completely different conclusion,
|
||
<bibRefCitation id="EF8E4B67CF5C10201B3AEA1EFB73FEE3" author="Hewitt, J." box="[878,1037,295,320]" pageId="12" pageNumber="513" pagination="283 - 357" refId="ref12349" refString="Hewitt, J. (1935) Some new forms of batrachians and reptiles from South Africa. Records of the Albany Museum, 4, 283 - 357, plates XXVII - XXXVI." type="journal article" year="1935">Hewitt (1935</bibRefCitation>
|
||
: p. 325) recognized “
|
||
<taxonomicName id="4C1F4D15CF5C10201D47EA1EFF89FEC6" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="513" phylum="Chordata" rank="species" species="gehafiae">
|
||
<emphasis id="B96BEA84CF5C10201D47EA1EFF89FEC6" italics="true" pageId="12" pageNumber="513">Pelomedusa gehafiae</emphasis>
|
||
</taxonomicName>
|
||
” [sic] as a valid species and opposed it to his polytypic species
|
||
<taxonomicName id="4C1F4D15CF5C10201B9EEA75FBD0FEC6" baseAuthorityName="Schoepff" baseAuthorityYear="1792" box="[970,1198,332,357]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="513" phylum="Chordata" rank="species" species="galeata">
|
||
<emphasis id="B96BEA84CF5C10201B9EEA75FBD0FEC6" box="[970,1198,332,357]" italics="true" pageId="12" pageNumber="513">Pelomedusa galeata</emphasis>
|
||
</taxonomicName>
|
||
. Hewitt noted that at Mt. Elgon (border region of
|
||
<collectingCountry id="F3087606CF5C102019B6EA56FD47FE2B" box="[482,569,367,392]" name="Uganda" pageId="12" pageNumber="513">Uganda</collectingCountry>
|
||
and
|
||
<collectingCountry id="F3087606CF5C10201A21EA56FDBEFE2B" box="[629,704,367,392]" name="Kenya" pageId="12" pageNumber="513">Kenya</collectingCountry>
|
||
) helmeted terrapins with separated pectorals are found together with terrapins having the pectorals in contact. Finally, based on the occurrence of morphologically intermediate terrapins in some regions,
|
||
<bibRefCitation id="EF8E4B67CF5C102019EAEA8EFD2DFE73" author="Parker, H. W." box="[446,595,439,464]" pageId="12" pageNumber="513" pagination="594 - 609" refId="ref12724" refString="Parker, H. W. (1936) Reptiles and amphibians collected by the Lake Rudolf Rift Valley Expedition, 1934. Annals and Magazine of Natural History, 18, 594 - 609. http: // dx. doi. org / 10.1080 / 00222933608655232" type="journal article" year="1936">Parker (1936</bibRefCitation>
|
||
: p. 609) and
|
||
<bibRefCitation id="EF8E4B67CF5C10201AB6EA81FCF4FE73" author="Mertens, R." box="[738,906,439,464]" pageId="12" pageNumber="513" pagination="139 - 142" refId="ref12650" refString="Mertens, R. (1937) Bemerkungen uber die Rassen von Pelomedusa subrufa (La Cepede). Zoologischer Anzeiger, 117, 139 - 142." type="journal article" year="1937">Mertens (1937</bibRefCitation>
|
||
: p. 140) recognized
|
||
<taxonomicName id="4C1F4D15CF5C10201C3FEA80FA52FE73" authorityName="Ruppell" authorityYear="1835" box="[1131,1324,439,464]" class="Testudines" family="Pelomedusidae" genus="Pentonyx" higherTaxonomySource="CoL" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="513" phylum="Chordata" rank="species" species="gehafie">
|
||
<emphasis id="B96BEA84CF5C10201C3FEA80FA52FE73" box="[1131,1324,439,464]" italics="true" pageId="12" pageNumber="513">Pentonyx gehafie</emphasis>
|
||
</taxonomicName>
|
||
as a valid subspecies of the helmeted terrapin and used the names
|
||
<taxonomicName id="4C1F4D15CF5C10201B7AEAE5FB0DFE56" box="[814,1139,476,501]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="513" phylum="Chordata" rank="subSpecies" species="galeata" subSpecies="gehafie">
|
||
<emphasis id="B96BEA84CF5C10201B7AEAE5FB0DFE56" box="[814,1139,476,501]" italics="true" pageId="12" pageNumber="513">Pelomedusa galeata gehafie</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1F4D15CF5C10201CE6EAE5FF94FDBB" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="513" phylum="Chordata" rank="subSpecies" species="subrufa" subSpecies="gehafie">
|
||
<emphasis id="B96BEA84CF5C10201CE6EAE5FF94FDBB" italics="true" pageId="12" pageNumber="513">Pelomedusa subrufa gehafie</emphasis>
|
||
</taxonomicName>
|
||
, respectively. In a similar vein,
|
||
<bibRefCitation id="EF8E4B67CF5C10201A18EAC6FC72FDBB" author="Loveridge, A." box="[588,780,511,536]" pageId="12" pageNumber="513" pagination="467 - 524" refId="ref12616" refString="Loveridge, A. (1941) Revision of the African terrapin [sic] of the family Pelomedusidae. Bulletin of the Museum of Comparative Zoology, 88, 467 - 524." type="journal article" year="1941">Loveridge (1941</bibRefCitation>
|
||
: p. 480) accepted the validity of a subspecies characterized by separated triangular pectoral scutes, but placed
|
||
<taxonomicName id="4C1F4D15CF5C10201AB9E91CFCCDFD9E" authorityName="Ruppell" authorityYear="1835" box="[749,947,548,573]" class="Testudines" family="Pelomedusidae" genus="Pentonyx" higherTaxonomySource="CoL" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="513" phylum="Chordata" rank="species" species="gehafie">
|
||
<emphasis id="B96BEA84CF5C10201AB9E91CFCCDFD9E" box="[749,947,548,573]" italics="true" pageId="12" pageNumber="513">Pentonyx gehafie</emphasis>
|
||
</taxonomicName>
|
||
in the synonymy of
|
||
<taxonomicName id="4C1F4D15CF5C10201CE6E91DFE94FDC3" authority="(Schweigger, 1812)" baseAuthorityName="Schweigger" baseAuthorityYear="1812" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="513" phylum="Chordata" rank="subSpecies" species="subrufa" subSpecies="olivacea">
|
||
<emphasis id="B96BEA84CF5C10201CE6E91DFF88FDC3" italics="true" pageId="12" pageNumber="513">Pelomedusa subrufa olivacea</emphasis>
|
||
(Schweigger, 1812)
|
||
</taxonomicName>
|
||
. Summarizing his own observations and those of previous authors, Loveridge believed that this taxon occurs in “the drier regions of a belt extending from
|
||
<collectingCountry id="F3087606CF5C10201C54E955FB27FD26" box="[1024,1113,620,645]" name="Senegal" pageId="12" pageNumber="513">Senegal</collectingCountry>
|
||
to
|
||
<collectingCountry id="F3087606CF5C10201C2BE955FBB0FD26" box="[1151,1230,620,645]" name="Eritrea" pageId="12" pageNumber="513">Eritrea</collectingCountry>
|
||
, intergrading with the typical form in the
|
||
<collectingCountry id="F3087606CF5C102019CEE9B6FDE1FD0B" box="[410,671,655,680]" name="Sudan" pageId="12" pageNumber="513">Anglo-Egyptian Sudan</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3087606CF5C10201AFFE9B6FC73FD0B" box="[683,781,655,680]" name="Ethiopia" pageId="12" pageNumber="513">Ethiopia</collectingCountry>
|
||
, Somaliland, northern
|
||
<collectingCountry id="F3087606CF5C10201C59E9A9FB28FD0B" box="[1037,1110,656,680]" name="Kenya" pageId="12" pageNumber="513">Kenya</collectingCountry>
|
||
and
|
||
<collectingCountry id="F3087606CF5C10201CDAE9B6FB96FD0B" box="[1166,1256,655,680]" name="Uganda" pageId="12" pageNumber="513">Uganda</collectingCountry>
|
||
” (p. 481). Later authors identified
|
||
<taxonomicName id="4C1F4D15CF5C1020193FE98DFDC1FD6E" baseAuthorityName="Schweigger" baseAuthorityYear="1812" box="[363,703,692,717]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="513" phylum="Chordata" rank="subSpecies" species="subrufa" subSpecies="olivacea">
|
||
<emphasis id="B96BEA84CF5C1020193FE98DFDC1FD6E" box="[363,703,692,717]" italics="true" pageId="12" pageNumber="513">Pelomedusa subrufa olivacea</emphasis>
|
||
</taxonomicName>
|
||
generally with the more northern populations of the helmeted terrapin, including the populations of the Arabian Peninsula (
|
||
<emphasis id="B96BEA84CF5C10201B19E9EEFC1FFD53" box="[845,865,727,752]" italics="true" pageId="12" pageNumber="513">cf</emphasis>
|
||
. Ernst &
|
||
<bibRefCitation id="EF8E4B67CF5C10201B85E9EEFB0EFD53" author="Barbour, R. W." box="[977,1136,727,752]" pageId="12" pageNumber="513" refId="ref11784" refString="Ernst, C. H. & Barbour, R. W. (1989) Turtles of the World. Smithsonian Institution Press, Washington, D. C., xii + 313 pp., 16 plates." type="book" year="1989">Barbour 1989</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8E4B67CF5C10201C28E9E1FA6DFD53" author="Iverson, J. B." box="[1148,1299,727,752]" pageId="12" pageNumber="513" refId="ref12440" refString="Iverson, J. B. (1992) A Revised Checklist with Distribution Maps of the Turtles of the World. Privately Printed, Richmond, IN, xiii + 363 pp." type="book" year="1992">Iverson 1992</bibRefCitation>
|
||
; Ernst
|
||
<emphasis id="B96BEA84CF5C10201D37E9E0FAE2FD53" box="[1379,1436,727,752]" italics="true" pageId="12" pageNumber="513">et al.</emphasis>
|
||
2000;
|
||
<bibRefCitation id="EF8E4B67CF5C10201889E9C5FD79FCB6" author="Boycott, R. C. & Bourquin, O." box="[221,519,764,789]" pageId="12" pageNumber="513" pagination="1788" refId="ref11440" refString="Boycott, R. C. & Bourquin, O. (2008) Pelomedusa subrufa (Lacepede 1788), helmeted turtle, helmeted terrapin. In: Rhodin, A. G. J., Pritchard, P. C. H., van Dijk, P. P., Saumure, R. A., Buhlmann, K. A. & Iverson, J. B. (Eds.), Conservation Biology of Freshwater Turtles and Tortoises: A Compilation Project of the IUCN / SSC Tortoise and Freshwater Turtle Specialist Group. Chelonian Research Monographs, No. 5. Chelonian Research Foundation, Lunenburg, MA, pp. 007.1 - 007.6. http: // dx. doi. org / 10.3854 / crm. 5.007. subrufa. v 1.2008" type="book chapter" year="2008">Boycott & Bourquin 2008</bibRefCitation>
|
||
), until the usage of subspecies was abandoned after
|
||
<bibRefCitation id="EF8E4B67CF5C10201C1BE9C5FA28FCB6" author="Gasperetti, J. & Stimson, A. F. & Miller, J. D. & Ross, J. P. & Gasperetti, P. R." box="[1103,1366,764,789]" pageId="12" pageNumber="513" pagination="170 - 367" refId="ref12043" refString="Gasperetti, J., Stimson, A. F., Miller, J. D., Ross, J. P. & Gasperetti, P. R. (1993) Turtles of Arabia. Fauna of Saudi Arabia, 13, 170 - 367." type="journal article" year="1993">
|
||
Gasperetti
|
||
<emphasis id="B96BEA84CF5C10201C9DE9C4FA7FFCB6" box="[1225,1281,764,789]" italics="true" pageId="12" pageNumber="513">et al.</emphasis>
|
||
(1993)
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C305651DCF5C10201893E819FAFEFC06" pageId="12" pageNumber="513" type="materials_examined">
|
||
<paragraph id="8BA03696CF5C10201893E819FAFEFC06" blockId="12.[151,1436,151,1184]" pageId="12" pageNumber="513">
|
||
<materialsCitation id="3B773CCBCF5C10201893E819FA02FC06" ID-GBIF-Occurrence="3319793303" county="Frankfurt" location="Frankfurt am Main" municipality="Tissue" pageId="12" pageNumber="513" specimenCount="1" typeStatus="lectotype">
|
||
For the present investigation, we studied the
|
||
<typeStatus id="54A48834CF5C10201A8DE826FC3EFC9B" box="[729,832,799,824]" pageId="12" pageNumber="513" type="lectotype">lectotype</typeStatus>
|
||
and the
|
||
<specimenCount id="9D19FD1FCF5C10201BFFE826FBF7FC9B" box="[939,1161,799,824]" pageId="12" pageNumber="513" type="generic" typeStatus="paralectotypes">nine paralectotypes</specimenCount>
|
||
of
|
||
<taxonomicName id="4C1F4D15CF5C10201CE1E818FA02FC9B" authorityName="Ruppell" authorityYear="1835" box="[1205,1404,799,824]" class="Testudines" family="Pelomedusidae" genus="Pentonyx" higherTaxonomySource="CoL" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="513" phylum="Chordata" rank="species" species="gehafie">
|
||
<emphasis id="B96BEA84CF5C10201CE1E818FA02FC9B" box="[1205,1404,799,824]" italics="true" pageId="12" pageNumber="513">Pentonyx gehafie</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<location id="8EC0604DCF5C102018C3E87DFE12FCFE" LSID="urn:lsid:plazi:treatment:03B68780CF59102018C3EDFCFBBFFB03:8EC0604DCF5C102018C3E87DFE12FCFE" box="[151,364,836,861]" county="Frankfurt" municipality="Tissue" name="Frankfurt am Main" pageId="12" pageNumber="513">Frankfurt am Main</location>
|
||
and
|
||
<location id="8EC0604DCF5C102019F6E87DFE83FCFE" LSID="urn:lsid:plazi:treatment:03B68780CF59102018C3EDFCFBBFFB03:8EC0604DCF5C102019F6E87DFE83FCFE" box="[418,509,836,861]" county="Frankfurt" municipality="Tissue" name="London" pageId="12" pageNumber="513">London</location>
|
||
; all specimens conform well to the description by
|
||
<bibRefCitation id="EF8E4B67CF5C10201C7BE87DFBA7FCFE" author="Ruppell, E." box="[1071,1241,836,861]" pageId="12" pageNumber="513" refId="ref13026" refString="Ruppell, E. (1835) Neue Wirbelthiere zu der Fauna von Abyssinien gehorig. Amphibien. Siegmund Schmerber, Frankfurt am Main, 18 pp., 6 plates. http: // dx. doi. org / 10.5962 / bhl. title. 53778" type="book" year="1835">Rüppell (1835)</bibRefCitation>
|
||
in having widely separated triangular pectoral scutes (
|
||
<figureCitation id="13242A13CF5C10201A65E85EFD08FC23" box="[561,630,871,896]" captionStart="FIGURE 5" captionStartId="11.[151,250,1953,1975]" captionTargetBox="[236,1375,210,1925]" captionTargetId="figure-77@11.[188,1399,193,1932]" captionTargetPageId="11" captionText="FIGURE 5. Dorsal and ventral views of the lectotype of Pentonyx gehafie Rüppell, 1835 (SMF 7947, subadult female, straight carapacial length 11.42 cm; top) and of one of the paralectotypes in the Natural History Museum, London (BMNH 1947.3.5.78, adult male, straight carapacial length 16.70 cm; bottom). Scale bars, 3 cm. Photos: E. Morawa, P. Campbell." figureDoi="http://doi.org/10.5281/zenodo.4915074" httpUri="https://zenodo.org/record/4915074/files/figure.png" pageId="12" pageNumber="513">Fig. 5</figureCitation>
|
||
).
|
||
<collectingMunicipality id="6BC4ACECCF5C10201ADFE85EFDADFC23" box="[651,723,871,896]" pageId="12" pageNumber="513">Tissue</collectingMunicipality>
|
||
for genetic examination was extracted from the eight
|
||
<collectingCounty id="62C14E1ACF5C10201D66E85EFAE2FC23" box="[1330,1436,871,896]" pageId="12" pageNumber="513">Frankfurt</collectingCounty>
|
||
specimens, but DNA sequences could be generated only for three terrapins, among them the
|
||
<typeStatus id="54A48834CF5C10201CF2E8B5FA73FC06" box="[1190,1293,908,933]" pageId="12" pageNumber="513" type="lectotype">lectotype</typeStatus>
|
||
(
|
||
<tableCitation id="C69D032DCF5C10201D4FE8B5FA0DFC06" box="[1307,1395,908,933]" captionStart="TABLE 2" captionStartId="4.[147,233,594,616]" captionTargetPageId="4" captionText="TABLE 2. Pelomedusa type specimens studied genetically. Lengths of successfully sequenced mtDNA fragments indicated. For GenBank accession numbers, see Table S1. The cyt b sequence of ZFMK 17076 was generated by Wong et al. (2010). For museum acronyms, see Table 1." httpUri="http://table.plazi.org/id/DF60661ECF54102818C7E96BFBB0FD26" pageId="12" pageNumber="513" tableUuid="DF60661ECF54102818C7E96BFBB0FD26">Table 2</tableCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C305651DCF5C10201893E889FBBFFB03" pageId="12" pageNumber="513" type="discussion">
|
||
<paragraph id="8BA03696CF5C10201893E889FBBFFB03" blockId="12.[151,1436,151,1184]" pageId="12" pageNumber="513">
|
||
In phylogenetic analyses (
|
||
<figureCitation id="13242A13CF5C102019A2E896FD45FC6B" box="[502,571,943,968]" captionStart="FIGURE 1" captionStartId="5.[151,250,1100,1122]" captionTargetBox="[157,1429,201,1075]" captionTargetId="figure-427@5.[151,1436,195,1080]" captionTargetPageId="5" captionText="FIGURE 1. Maximum Likelihood tree for Pelomedusa, based on 1848 bp of mitochondrial DNA (12S, cyt b, ND4+tRNAs), rooted with Pelusios sinuatus. The tree topology completely matches the one inferred by Bayesian analyses. Numbers along nodes indicate bootstrap support and Bayesian posterior probabilities greater than 50 or 0.95, respectively (not shown for some terminal clades with short branches). Sample codes are voucher numbers; see Table S1 for explanation. Name-bearing types (red) and paralectotypes or topotypic samples (blue) highlighted. On the right, mitochondrial lineages of Vargas-Ramírez et al. (2010) indicated. No topotypic material of Pelomedusa galeata orangensis Hewitt, 1935 could be sequenced. However, the lost holotype is thought to come from the vicinity of Kimberley (South Africa) and all South African samples belong to lineage IX, so it can be concluded that Pelomedusa galeata orangensis represents the same lineage. Inset: Helmeted terrapin from the Addo Elephant National Park, South Africa. Photo: M. Gregorič." figureDoi="http://doi.org/10.5281/zenodo.4915066" httpUri="https://zenodo.org/record/4915066/files/figure.png" pageId="12" pageNumber="513">Fig. 1</figureCitation>
|
||
), the sequences of the
|
||
<typeStatus id="54A48834CF5C10201B1BE889FC01FC6B" box="[847,895,944,968]" pageId="12" pageNumber="513">type</typeStatus>
|
||
specimens constitute a deeply divergent clade being sister to Kenyan terrapins representing lineage V
|
||
<emphasis id="B96BEA84CF5C10201B7EE8ECFC16FC4F" box="[810,872,981,1004]" italics="true" pageId="12" pageNumber="513">sensu</emphasis>
|
||
<bibRefCitation id="EF8E4B67CF5C10201B20E8EDFBB6FC4E" author="Vargas-Ramirez, M. & Vences, M. & Branch, W. R. & Daniels, S. R. & Glaw, F. & Hofmeyr, M. D. & Kuchling, G. & Maran, J. & Papenfuss, T. J. & Siroky, P. & Vieites, D. R. & Fritz, U." box="[884,1224,980,1005]" pageId="12" pageNumber="513" pagination="428 - 440" refId="ref13781" refString="Vargas-Ramirez, M., Vences, M., Branch, W. R., Daniels, S. R., Glaw, F., Hofmeyr, M. D., Kuchling, G., Maran, J., Papenfuss, T. J., Siroky, P., Vieites, D. R. & Fritz, U. (2010) Deep genealogical lineages in the widely distributed African helmeted terrapin: evidence from mitochondrial and nuclear DNA (Testudines: Pelomedusidae: Pelomedusa subrufa). Molecular Phylogenetics and Evolution, 56, 428 - 440. http: // dx. doi. org / 10.1016 / j. ympev. 2010.03.019" type="journal article" year="2010">
|
||
Vargas-Ramírez
|
||
<emphasis id="B96BEA84CF5C10201C67E8ECFB11FC4E" box="[1075,1135,980,1005]" italics="true" pageId="12" pageNumber="513">et al.</emphasis>
|
||
(2010)
|
||
</bibRefCitation>
|
||
. We examined 18
|
||
<taxonomicName id="4C1F4D15CF5C102018C3E8CEFE5EFBB3" box="[151,288,1015,1040]" class="Reptilia" family="Pelomedusidae" genus="Pelomedusa" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="513" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96BEA84CF5C102018C3E8CEFE5EFBB3" box="[151,288,1015,1040]" italics="true" pageId="12" pageNumber="513">Pelomedusa</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F3087606CF5C10201939E8C1FEC8FBB3" box="[365,438,1016,1040]" name="Kenya" pageId="12" pageNumber="513">Kenya</collectingCountry>
|
||
morphologically (7 live terrapins and
|
||
<specimenCount id="9D19FD1FCF5C10201BD6E8CEFB63FBB3" box="[898,1053,1015,1040]" pageId="12" pageNumber="513" type="generic">11 specimens</specimenCount>
|
||
in the collections of the natural history museums of Berlin, Bonn, London, and Vienna) and these differ consistently from the
|
||
<typeStatus id="54A48834CF5C10201CA9EF25FA53FB97" box="[1277,1325,1052,1076]" pageId="12" pageNumber="513">type</typeStatus>
|
||
series of
|
||
<taxonomicName id="4C1F4D15CF5C102018C3EF78FE24FBFB" authorityName="Ruppell" authorityYear="1835" box="[151,346,1087,1112]" class="Testudines" family="Pelomedusidae" genus="Pentonyx" higherTaxonomySource="CoL" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="513" phylum="Chordata" rank="species" species="gehafie">
|
||
<emphasis id="B96BEA84CF5C102018C3EF78FE24FBFB" box="[151,346,1087,1112]" italics="true" pageId="12" pageNumber="513">Pentonyx gehafie</emphasis>
|
||
</taxonomicName>
|
||
in that their pectoral scutes are in wide contact in the midline, suggesting that
|
||
<taxonomicName id="4C1F4D15CF5C10201C8DEF78FAE2FBFB" authorityName="Ruppell" authorityYear="1835" box="[1241,1436,1087,1112]" class="Testudines" family="Pelomedusidae" genus="Pentonyx" higherTaxonomySource="CoL" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="513" phylum="Chordata" rank="species" species="gehafie">
|
||
<emphasis id="B96BEA84CF5C10201C8DEF78FAE2FBFB" box="[1241,1436,1087,1112]" italics="true" pageId="12" pageNumber="513">Pentonyx gehafie</emphasis>
|
||
</taxonomicName>
|
||
could be a distinct taxon. At present, we refer to the genetic lineage of the
|
||
<typeStatus id="54A48834CF5C10201BB3EF5DFB69FBDF" box="[999,1047,1124,1148]" pageId="12" pageNumber="513">type</typeStatus>
|
||
specimens of
|
||
<taxonomicName id="4C1F4D15CF5C10201CEEEF5CFA03FBDE" authorityName="Ruppell" authorityYear="1835" box="[1210,1405,1124,1149]" class="Testudines" family="Pelomedusidae" genus="Pentonyx" higherTaxonomySource="CoL" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="513" phylum="Chordata" rank="species" species="gehafie">
|
||
<emphasis id="B96BEA84CF5C10201CEEEF5CFA03FBDE" box="[1210,1405,1124,1149]" italics="true" pageId="12" pageNumber="513">Pentonyx gehafie</emphasis>
|
||
</taxonomicName>
|
||
as lineage X, and do not relate it to any of the lineages identified by
|
||
<bibRefCitation id="EF8E4B67CF5C10201B22EFBEFBC3FB03" author="Vargas-Ramirez, M. & Vences, M. & Branch, W. R. & Daniels, S. R. & Glaw, F. & Hofmeyr, M. D. & Kuchling, G. & Maran, J. & Papenfuss, T. J. & Siroky, P. & Vieites, D. R. & Fritz, U." box="[886,1213,1159,1184]" pageId="12" pageNumber="513" pagination="428 - 440" refId="ref13781" refString="Vargas-Ramirez, M., Vences, M., Branch, W. R., Daniels, S. R., Glaw, F., Hofmeyr, M. D., Kuchling, G., Maran, J., Papenfuss, T. J., Siroky, P., Vieites, D. R. & Fritz, U. (2010) Deep genealogical lineages in the widely distributed African helmeted terrapin: evidence from mitochondrial and nuclear DNA (Testudines: Pelomedusidae: Pelomedusa subrufa). Molecular Phylogenetics and Evolution, 56, 428 - 440. http: // dx. doi. org / 10.1016 / j. ympev. 2010.03.019" type="journal article" year="2010">
|
||
Vargas-Ramírez
|
||
<emphasis id="B96BEA84CF5C10201C64EFB0FB17FB03" box="[1072,1129,1159,1184]" italics="true" pageId="12" pageNumber="513">et al.</emphasis>
|
||
(2010)
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |