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<document id="770A89AE1A71C205BD01991BCE5AB412" ID-DOI="10.5281/zenodo.35388" ID-GBIF-Dataset="b184dfcc-9b5f-4abb-b401-0f45f76006f6" checkinTime="1450029366780" checkinUser="donat" docAuthor="Dijkstra, Klaas-Douwe B., Kipping, Jens &amp; Mézière, Nicolas" docDate="2015" docId="03A25264CAA6FF28EF0CFCAB407DFB9A" docLanguage="en" docName="Odonatologica_44-4-low_res.pdf.imf" docOrigin="Odonatologica 44 (4)" docTitle="Orthetrum kafwi Dijkstra, sp. nov." docType="treatment" docVersion="19" lastPageNumber="639" masterDocId="FF9B2A1CCA19FFEAEE22FFED4105FFB2" masterDocTitle="Sixty new dragonfly and damselfly species from Africa (Odonata)" masterLastPageNumber="678" masterPageNumber="447" pageNumber="636" updateTime="1695001031402" updateUser="admin">
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<mods:title id="454E3170593DBBDD711CB4DEC3824934">Sixty new dragonfly and damselfly species from Africa (Odonata)</mods:title>
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<mods:namePart id="F00AFF555E512ADAD991FF9B29F8ABC5">Dijkstra, Klaas-Douwe B.</mods:namePart>
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<mods:namePart id="CF2A1AEAD60C18B53AFDAB485D7662FA">Kipping, Jens</mods:namePart>
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<mods:namePart id="C49AE96F7D1C8AAAA16B20BBAEFF4054">Mézière, Nicolas</mods:namePart>
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<mods:title id="0DB348AC341AD392BE4F3014DB4BDCB4">Odonatologica</mods:title>
</mods:titleInfo>
<mods:part id="CE502622C0B60296CFDC771C01251141">
<mods:date id="991B6EEEF5A24A3DF825B1A3D2603612">2015</mods:date>
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<mods:number id="971A522E3B45EAAC58868D990FA151C4">44</mods:number>
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<mods:classification id="9AA881B3A1689D90E1D07D23D4F9F2A9">journal article</mods:classification>
<mods:identifier id="70304FD5D38C25D3EDAE2049F811EEFF" type="DOI">10.5281/zenodo.35388</mods:identifier>
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<treatment id="03A25264CAA6FF28EF0CFCAB407DFB9A" ID-DOI="http://doi.org/10.5281/zenodo.5640294" ID-GBIF-Taxon="114923461" ID-Zenodo-Dep="5640294" LSID="urn:lsid:zoobank.org:act:A5E8E1BA-D518-45D8-AB70-7368B16DCC2E" httpUri="http://treatment.plazi.org/id/03A25264CAA6FF28EF0CFCAB407DFB9A" lastPageId="194" lastPageNumber="639" pageId="191" pageNumber="636">
<subSubSection id="C311B0F9CAA6FF55EF0CFCAB421BFC3F" pageId="191" pageNumber="636" type="nomenclature">
<paragraph id="8BB4E372CAA6FF55EF0CFCAB421BFC3F" pageId="191" pageNumber="636">
<heading id="D0FC541ECAA6FF55EF0CFCAB421BFC3F" bold="true" centered="true" fontSize="11" level="2" pageId="191" pageNumber="636" reason="4">
<emphasis id="B97F3F60CAA6FF55EF0CFCAB4278FCD0" bold="true" box="[302,893,838,866]" pageId="191" pageNumber="636">
<taxonomicName id="4C0B98F1CAA6FF55EF0CFCAB4359FCD0" LSID-ZBK="urn:lsid:zoobank.org:act:A5E8E1BA-D518-45D8-AB70-7368B16DCC2E" authority="Dijkstra" authorityName="Dijkstra" box="[302,604,838,866]" class="Insecta" family="Libellulidae" genus="Orthetrum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="191" pageNumber="636" phylum="Arthropoda" rank="species" species="kafwi" status="sp. nov.">
<emphasis id="B97F3F60CAA6FF55EF0CFCAB40FCFCD0" bold="true" box="[302,505,838,866]" italics="true" pageId="191" pageNumber="636">Orthetrum kafwi</emphasis>
Dijkstra
</taxonomicName>
<taxonomicNameLabel id="A24C821BCAA6FF55EC41FCAA43C5FCD0" box="[611,704,839,866]" pageId="191" pageNumber="636" rank="species">sp. nov.</taxonomicNameLabel>
Bog Skimmer
</emphasis>
(
<typeStatus id="07A6111C5E6446635096FB8E0F8AEC98">Type</typeStatus>
<figureCitation id="1330FFF7CAA6FF55EFF6FC9C4346FC3E" box="[468,579,881,908]" captionStart="Photo 51" captionStartId="149.[159,225,1498,1522]" captionTargetBox="[159,1035,801,1457]" captionTargetId="figure@149.[159,1038,801,1459]" captionTargetPageId="149" captionText="Photo 51. Paragomphus darwalli, male; Boubou stream near Bongoville, Gabon. Photo: NM (20 - xi- 2011)" pageId="191" pageNumber="636">Photo 51</figureCitation>
,
<figureCitation id="1330FFF7CAA6FF55EC6CFC9C43B8FC3E" box="[590,701,881,908]" captionStart="Photo 67" captionStartId="193.[159,225,1263,1287]" captionText="Photo 67. Bog at the source of the Kafwi River on the Kibara Plateau in Upemba National Park, Katanga, Congo-Kinshasa. Both Orthetrum kafwi and O. lusinga were found here. Photo: KDBD (15 - - ii- 2011)" pageId="191" pageNumber="636">Photo 67</figureCitation>
,
<figureCitation id="1330FFF7CAA6FF55ECE5FC9C4213FC3F" box="[711,790,881,909]" captionStart="Figure 31" captionStartId="189.[784,852,1410,1435]" captionTargetBox="[159,745,910,1550]" captionTargetId="figure@189.[156,749,904,1552]" captionTargetPageId="189" captionText="Figure 31. Secondary genitalia in lateral view of the saegeri - group of Orthetrum and three similar species." pageId="191" pageNumber="636">Fig. 31</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CAA6FF55EEBDFC4D401AFC08" box="[159,287,928,954]" pageId="191" pageNumber="636" type="description">
<paragraph id="8BB4E372CAA6FF55EEBDFC4D401AFC08" box="[159,287,928,954]" pageId="191" pageNumber="636">
<heading id="D0FC541ECAA6FF55EEBDFC4D401AFC08" bold="true" box="[159,287,928,954]" fontSize="11" level="3" pageId="191" pageNumber="636" reason="6">
<emphasis id="B97F3F60CAA6FF55EEBDFC4D401AFC08" bold="true" box="[159,287,928,954]" pageId="191" pageNumber="636">Taxonomy</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CAA6FF55EEBDFC29423FFBFD" pageId="191" pageNumber="636" type="discussion">
<paragraph id="8BB4E372CAA6FF55EEBDFC29423FFBFD" pageId="191" pageNumber="636">
Morphologically similar to
<taxonomicName id="4C0B98F1CAA6FF55EFCEFC28420EFC6D" authority="Schmidt, 1951" authorityName="Schmidt" authorityYear="1951" box="[492,779,964,991]" class="Insecta" family="Libellulidae" genus="Orthetrum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="191" pageNumber="636" phylum="Arthropoda" rank="species" species="hintzi">
<emphasis id="B97F3F60CAA6FF55EFCEFC284351FC6D" box="[492,596,964,991]" italics="true" pageId="191" pageNumber="636">O. hintzi</emphasis>
Schmidt, 1951
</taxonomicName>
among which it was discovered, but genetically closer to the
<taxonomicName id="4C0B98F1CAA6FF55EC5AFC0643C3FBB6" box="[632,710,1003,1028]" class="Insecta" family="Libellulidae" genus="Orthetrum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="191" pageNumber="636" phylum="Arthropoda" rank="species" species="saegeri">
<emphasis id="B97F3F60CAA6FF55EC5AFC0643C3FBB6" box="[632,710,1003,1028]" italics="true" pageId="191" pageNumber="636">saegeri</emphasis>
</taxonomicName>
-group (see
<taxonomicName id="4C0B98F1CAA6FF55ED76FC0742E1FBB6" box="[852,996,1002,1028]" class="Insecta" family="Libellulidae" genus="Orthetrum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="191" pageNumber="636" phylum="Arthropoda" rank="species" species="agaricum" status="sp. nov.">
<emphasis id="B97F3F60CAA6FF55ED76FC0742E1FBB6" box="[852,996,1002,1028]" italics="true" pageId="191" pageNumber="636">O. agaricum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C821BCAA6FF55EDCEFC0641D6FB98" pageId="191" pageNumber="636" rank="species">sp. nov.</taxonomicNameLabel>
), which includes the also syntopic
<taxonomicName id="4C0B98F1CAA6FF55EC4FFBE243E4FB98" box="[621,737,1039,1066]" class="Insecta" family="Libellulidae" genus="Orthetrum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="191" pageNumber="636" phylum="Arthropoda" rank="species" species="lusinga" status="sp. nov.">
<emphasis id="B97F3F60CAA6FF55EC4FFBE243E4FB98" box="[621,737,1039,1066]" italics="true" pageId="191" pageNumber="636">O. lusinga</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C821BCAA6FF55ECCBFBFD4244FB98" box="[745,833,1040,1066]" pageId="191" pageNumber="636" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="1330FFF7CAA6FF55ED72FBE24298FB98" box="[848,925,1039,1066]" captionStart="Tree 8" captionStartId="188.[159,207,1438,1463]" captionTargetBox="[159,1034,577,1399]" captionTargetId="figure@188.[156,1038,571,1402]" captionTargetPageId="188" captionText="Tree 8. COI gene tree from Bayesian inference analysis for the saegeri - group of Orthetrum and two outgroup taxa. For relevant branches posterior probabilities are shown first, as is bootstrap support from a 50 % majority rule consensus of 1 0 0 0 trees from Maximum Likelihood analysis (both as percentages)." pageId="191" pageNumber="636">Tree 8</figureCitation>
). Treated as a distinct species by
<bibRefCitation id="6F32837D215C617E0E6B10D270136F8F" author="Dijkstra K. - D. B. &amp; Clausnitzer V." journalOrPublisher="Royal Museum for Central Africa, Tervuren" refId="ref66897" refString="Dijkstra K. - D. B. &amp; Clausnitzer V. 2014. The dragonflies and damselflies of eastern Africa: handbook for all Odonata from Sudan to Zimbabwe. Royal Museum for Central Africa, Tervuren" title="The dragonflies and damselflies of eastern Africa: handbook for all Odonata from Sudan to Zimbabwe" type="book" year="2014">Dijkstra &amp; Clausnitzer (2014)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CAA6FF55EEBDFB6D40A1F9BD" pageId="191" pageNumber="636" type="materials_examined">
<paragraph id="8BB4E372CAA6FF55EEBDFB6D406DFB28" box="[159,360,1152,1178]" pageId="191" pageNumber="636">
<heading id="D0FC541ECAA6FF55EEBDFB6D406DFB28" bold="true" box="[159,360,1152,1178]" fontSize="11" level="3" pageId="191" pageNumber="636" reason="6">
<emphasis id="B97F3F60CAA6FF55EEBDFB6D406DFB28" bold="true" box="[159,360,1152,1178]" pageId="191" pageNumber="636">Material studied</emphasis>
</heading>
</paragraph>
<paragraph id="8BB4E372CAA6FF55EEBDFB484206FA9D" pageId="191" pageNumber="636">
<materialsCitation id="7530E266090B8CE33C51DCD795BC24E4" ID-GBIF-Occurrence="3396361302" collectingDate="2011-11-15" collectionCode="RMNH" collectorName="K. - D. B. Dijkstra" country="Democratic Republic of the Congo" elevation="1795" httpUri="http://www.boldsystems.org/index.php/Public_RecordView?processid=NSPAF331-15" latitude="-8.937" location="Upemba National Park, Kabwekanono, source area of Kafwi" longLatPrecision="1" longitude="27.166" specimenCode="RMNH.INS.505544" specimenCount="1" specimenCount-male="1" stateProvince="Katanga" typeStatus="holotype">
<emphasis id="B97F3F60CAA6FF55EEBDFB484015FB0D" bold="true" box="[159,272,1189,1215]" pageId="191" pageNumber="636">
<typeStatus id="F2AADB2A24490B95B207F4A8AFA22934">Holotype</typeStatus>
</emphasis>
<emphasis id="B97F3F60CAA6FF55EF3AFB494029FB0C" bold="true" box="[280,300,1188,1214]" pageId="191" pageNumber="636">
<specimenCount id="72081FE2AC2BC159995925BB665BDE59" type="male"></specimenCount>
</emphasis>
<emphasis id="B97F3F60CAA6FF55EF0EFB484031FB0D" bold="true" box="[300,308,1189,1215]" pageId="191" pageNumber="636">.</emphasis>
<specimenCode id="EB8C671EF89E68663D287C581F911619">RMNH.INS.505544</specimenCode>
,
<collectingCountry id="4C8FCC643CFDAF447BC8F9B6CB1B8E45" name="Democratic Republic of the Congo" value="Democratic Republic of the Congo">Congo-Kinshasa</collectingCountry>
,
<taxonomicName id="4C0B98F1CAA6FF55ED26FB484262FB0D" box="[772,871,1189,1215]" class="Insecta" family="Libellulidae" genus="Neodythemis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="191" pageNumber="631" phylum="Arthropoda" rank="species" species="katanga">
<collectingRegion id="EC5E694D5B05E739ABCA3813FA721FBF">Katanga</collectingRegion>
</taxonomicName>
,
<location id="342EA9D7860B6D2F82537C75FEFC027A" LSID="urn:lsid:plazi:treatment:03A25264CAA6FF28EF0CFCAB407DFB9A:342EA9D7860B6D2F82537C75FEFC027A" country="Democratic Republic of the Congo" latitude="-8.937" longLatPrecision="1" longitude="27.166" name="Upemba National Park, Kabwekanono, source area of Kafwi" stateProvince="Katanga">
Upemba National Park, Kabwekanono, source area of
<taxonomicName id="4C0B98F1CAA6FF55ECA3FB2743CDFB56" box="[641,712,1226,1252]" class="Insecta" family="Libellulidae" genus="Orthetrum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="191" pageNumber="639" phylum="Arthropoda" rank="species" species="kafwi" status="sp. nov.">Kafwi</taxonomicName>
</location>
, stream through gallery forest and bogs in open grassy plains (
<figureCitation id="1330FFF7CAA6FF55EC1AFB0243A3FAB8" box="[568,678,1263,1290]" captionStart="Photo 67" captionStartId="193.[159,225,1263,1287]" captionText="Photo 67. Bog at the source of the Kafwi River on the Kibara Plateau in Upemba National Park, Katanga, Congo-Kinshasa. Both Orthetrum kafwi and O. lusinga were found here. Photo: KDBD (15 - - ii- 2011)" pageId="191" pageNumber="636">Photo 67</figureCitation>
),
<elevation id="672190E07397520C88120B2E6C274B6D" metricMagnitude="3" metricUnit="m" metricValue="1.795" metricValueMax="1.8199999999999998" metricValueMin="1.77" unit="m" value="1795.0" valueMax="1820.0" valueMin="1770.0">
<quantity id="D744C67EA68A0B3104904C1903B18BAF" metricMagnitude="3" metricUnit="m" metricValue="1.795" metricValueMax="1.8199999999999998" metricValueMin="1.77" unit="m" value="1795.0" valueMax="1820.0" valueMin="1770.0">17701820 m</quantity>
</elevation>
. a.s.l. (
<geoCoordinate id="153D64DB2001162AB135CB2607B0A627" direction="south" orientation="latitude" precision="1" value="-8.937">8.937°S</geoCoordinate>
<geoCoordinate id="6593B96DC8972283B41C83A8B3AD6FB6" direction="east" orientation="longitude" precision="1" value="27.166">27.166°E</geoCoordinate>
),
<date id="0BFC6E2E9596C4CEC52CE0ECD2897099" value="2011-11-15">
<collectingDate id="F64B512F2087F9CEF4FFF38F7587FA4B" value="2011-11-15">15-xi-2011</collectingDate>
</date>
, leg.
<collectorName id="96052EB8187330D2EF77572A050087FA">K.-D.B. Dijkstra</collectorName>
,
<collectionCode id="74F67334FBCB3C35C2BEA5B1A950DE1A" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]">RMNH</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB4E372CAA6FF55EEBDFAD640A1F9BD" pageId="191" pageNumber="636">
<materialsCitation id="51D465554791A9589CD0D594FF908BFF" ID-GBIF-Occurrence="1224547945" collectingDate="2011-11-15" collectionCode="RMNH" country="Democratic Republic of the Congo" httpUri-0="http://www.boldsystems.org/index.php/Public_RecordView?processid=NSPAF335-15" httpUri-1="http://www.boldsystems.org/index.php/Public_RecordView?processid=NSPAF342-15" httpUri-2="http://www.boldsystems.org/index.php/Public_RecordView?processid=NSPAF334-15" location="Katanga" specimenCode="RMNH.INS.505554, RMNH.INS.505579" specimenCount="10" specimenCount-female="4" specimenCount-male="6" stateProvince="Katanga">
<emphasis id="B97F3F60CAA6FF55EEBDFAD6408CFAE6" bold="true" box="[159,393,1338,1364]" pageId="191" pageNumber="636">Further material.</emphasis>
<collectingCountry id="F1F96D6A18E7450C4CA6978FB72B345B" name="Democratic Republic of the Congo" value="Democratic Republic of the Congo">CONGO-KINSHASA</collectingCountry>
(
<taxonomicName id="4C0B98F1CAA6FF55ECF3FAD7423DFAE6" box="[721,824,1338,1364]" class="Insecta" family="Libellulidae" genus="Neodythemis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="191" pageNumber="631" phylum="Arthropoda" rank="species" species="katanga">
<collectingRegion id="15331EE87DE3D17193E8092DF53D41BE">Katanga</collectingRegion>
</taxonomicName>
):
<specimenCount id="38A4F929F2C4324C963A51019AD1EBFE" type="generic">4♂</specimenCount>
(
<specimenCode id="15B91461C87094CBF31FF992CAECB39C">RMNH.INS.505554</specimenCode>
,
<specimenCode id="0A312FBF714C57C157691FFC4D6BB817">RMNH.INS.505579</specimenCode>
),
<specimenCount id="D63FD5CDF16AB9589342CB45ABBD8E36" type="female">2♀</specimenCount>
(
<specimenCode id="5DB98BF9EF038F3081C9016124677B46">RMNH.INS.505553</specimenCode>
),
<specimenCount id="595D425C204C8DFA4F406EF6D2663A4A" type="generic">2♂</specimenCount>
<specimenCount id="252E78AC139FE582EAD1B6C3A559B021" type="female"></specimenCount>
, as holotype,
<collectionCode id="18E1CD6B8EF1362CB0E7BDCE078D33A1" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]">RMNH</collectionCode>
.
</materialsCitation>
<materialsCitation id="175BEC4103A624ED9B1589334ED04EA0" ID-GBIF-Occurrence="1224547958" collectingDate="2011-11-11" collectionCode="RMNH" collectorName="K. - D. B. Dijkstra" country="Democratic Republic of the Congo" elevation="1780" latitude="-8.933" location="Upemba National Park, source area of Lusinga near park headquarters" longLatPrecision="1" longitude="27.199" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Katanga">
<specimenCount id="424C1091D3B012AD829D82B855221CFB" type="generic">1♂</specimenCount>
<specimenCount id="FED6913D6506BC22427D3909D632AE85" type="female">1♀</specimenCount>
,
<location id="B84B2AA44073C726C61BF0AC6EB710F7" LSID="urn:lsid:plazi:treatment:03A25264CAA6FF28EF0CFCAB407DFB9A:B84B2AA44073C726C61BF0AC6EB710F7" country="Democratic Republic of the Congo" latitude="-8.933" longLatPrecision="1" longitude="27.199" name="Upemba National Park, source area of Lusinga near park headquarters" stateProvince="Katanga">
Upemba National Park, source area of
<taxonomicName id="4C0B98F1CAA6FF55ED51FA6942CAFA2D" box="[883,975,1412,1439]" class="Insecta" family="Libellulidae" genus="Orthetrum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="191" pageNumber="642" phylum="Arthropoda" rank="species" species="lusinga" status="sp. nov.">Lusinga</taxonomicName>
near park headquarters
</location>
, spring streams in gallery forest and adjacent bog, dam and channel,
<elevation id="0DFFFBCCB8B0827351809AAE5A95E6C7" metricMagnitude="3" metricUnit="m" metricValue="1.78" metricValueMax="1.8" metricValueMin="1.76" unit="m" value="1780.0" valueMax="1800.0" valueMin="1760.0">
<quantity id="CA3E8E50F34B417EE62C3676165C02D8" metricMagnitude="3" metricUnit="m" metricValue="1.78" metricValueMax="1.8" metricValueMin="1.76" unit="m" value="1780.0" valueMax="1800.0" valueMin="1760.0">17601800 m</quantity>
</elevation>
a.s.l. (
<geoCoordinate id="B7FB6C4C892ED8FC375F60680C9A29FA" direction="south" orientation="latitude" precision="1" value="-8.933">8.933°S</geoCoordinate>
<geoCoordinate id="0A6ECE77513E0251494A5C8C897B6B2C" direction="east" orientation="longitude" precision="1" value="27.199">27.199°E</geoCoordinate>
),
<date id="8EA99ED4EE42A8D2455F9121CB469FBF" value="2011-11-11">
<collectingDate id="0912BD8BC0EEE7F5D392E73FD32B8C3D" value="2011-11-11">11-xi-2011</collectingDate>
</date>
, leg.
<collectorName id="A5A027C4E580BE2AB7022C3435DE3B25">K.- D.B. Dijkstra</collectorName>
,
<collectionCode id="FF3AA746412DB9CC15441C1F3D937A71" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]">RMNH</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CAD9FF2AEEBDFF13404FFED0" pageId="192" pageNumber="637" type="description">
<paragraph id="8BB4E372CAD9FF2AEEBDFF13400CFEAA" box="[159,265,254,280]" pageId="192" pageNumber="637">
<emphasis id="B97F3F60CAD9FF2AEEBDFF13400CFEAA" bold="true" box="[159,265,254,280]" pageId="192" pageNumber="637">Genetics</emphasis>
</paragraph>
<paragraph id="8BB4E372CAD9FF2AEEBDFECE404FFED0" pageId="192" pageNumber="637">
One unique haplotype (n =4) is perhaps the most distinct within
<taxonomicName id="4C0B98F1CAD9FF2AED90FECE4504FE8F" box="[946,1025,291,317]" class="Insecta" family="Libellulidae" genus="Orthetrum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="192" pageNumber="637" phylum="Arthropoda" rank="species" species="saegeri">
<emphasis id="B97F3F60CAD9FF2AED90FECE4504FE8F" box="[946,1025,291,317]" italics="true" pageId="192" pageNumber="637">saegeri</emphasis>
</taxonomicName>
- group (
<figureCitation id="1330FFF7CAD9FF2AEED1FEA5403AFED0" box="[243,319,328,354]" captionStart="Tree 8" captionStartId="188.[159,207,1438,1463]" captionTargetBox="[159,1034,577,1399]" captionTargetId="figure@188.[156,1038,571,1402]" captionTargetPageId="188" captionText="Tree 8. COI gene tree from Bayesian inference analysis for the saegeri - group of Orthetrum and two outgroup taxa. For relevant branches posterior probabilities are shown first, as is bootstrap support from a 50 % majority rule consensus of 1 0 0 0 trees from Maximum Likelihood analysis (both as percentages)." pageId="192" pageNumber="637">Tree 8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CAD9FF28EF68FEA540F9FD00" lastPageId="194" lastPageNumber="639" pageId="192" pageNumber="637" type="diagnosis">
<paragraph id="8BB4E372CAD9FF2BEF68FEA54317FEAA" lastPageId="193" lastPageNumber="638" pageId="192" pageNumber="637">
.
<emphasis id="B97F3F60CAD8FF2BEEBDFF134317FEAA" bold="true" box="[159,530,254,280]" pageId="193" pageNumber="638">Male morphological diagnosis</emphasis>
</paragraph>
<caption id="DF74B3FACAD9FF2AEEBDFA354269F9BD" pageId="192" pageNumber="637" targetBox="[158,1035,505,1454]" targetPageId="192">
<paragraph id="8BB4E372CAD9FF2AEEBDFA354269F9BD" pageId="192" pageNumber="637">
<emphasis id="B97F3F60CAD9FF2AEEBDFA3541F8FA43" bold="true" box="[159,253,1496,1521]" pageId="192" pageNumber="637">Map 11.</emphasis>
Distribution of the
<taxonomicName id="4C0B98F1CAD9FF2AEFFEFA35432EFA42" box="[476,555,1496,1520]" class="Insecta" family="Libellulidae" genus="Orthetrum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="192" pageNumber="637" phylum="Arthropoda" rank="species" species="saegeri">
<emphasis id="B97F3F60CAD9FF2AEFFEFA35432EFA42" box="[476,555,1496,1520]" italics="true" pageId="192" pageNumber="637">saegeri</emphasis>
</taxonomicName>
-group of
<taxonomicName id="4C0B98F1CAD9FF2AECB9FA35420BFA42" box="[667,782,1496,1520]" class="Insecta" family="Libellulidae" genus="Orthetrum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="192" pageNumber="637" phylum="Arthropoda" rank="genus">
<emphasis id="B97F3F60CAD9FF2AECB9FA35420BFA42" box="[667,782,1496,1520]" italics="true" pageId="192" pageNumber="637">Orthetrum</emphasis>
</taxonomicName>
(
<taxonomicName id="4C0B98F1CAD9FF2AED3DFA3542A2FA42" box="[799,935,1496,1520]" class="Insecta" family="Libellulidae" genus="Orthetrum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="192" pageNumber="637" phylum="Arthropoda" rank="species" species="agaricum" status="sp. nov.">
<emphasis id="B97F3F60CAD9FF2AED3DFA3542A2FA42" box="[799,935,1496,1520]" italics="true" pageId="192" pageNumber="637">O. agaricum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C821BCAD9FF2AED92FA35450FFA43" box="[944,1034,1496,1521]" pageId="192" pageNumber="637" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C0B98F1CAD9FF2AEEBDFA1B41F2F9BC" box="[159,247,1526,1550]" class="Insecta" family="Libellulidae" genus="Orthetrum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="192" pageNumber="637" phylum="Arthropoda" rank="species" species="kafwi" status="sp. nov.">
<emphasis id="B97F3F60CAD9FF2AEEBDFA1B41F2F9BC" box="[159,247,1526,1550]" italics="true" pageId="192" pageNumber="637">O. kafwi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C821BCAD9FF2AEEDFFA184056F9BC" box="[253,339,1525,1550]" pageId="192" pageNumber="637" rank="species">sp. nov.</taxonomicNameLabel>
,
<emphasis id="B97F3F60CAD9FF2AEF79FA1B4361F9BC" box="[347,612,1526,1550]" italics="true" pageId="192" pageNumber="637">
<taxonomicName id="4C0B98F1CAD9FF2AEF79FA1B40CCF9BC" box="[347,457,1526,1550]" class="Insecta" family="Libellulidae" genus="Orthetrum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="192" pageNumber="637" phylum="Arthropoda" rank="species" species="lusinga">O. lusinga</taxonomicName>
,
<taxonomicName id="4C0B98F1CAD9FF2AEFF1FA1B4361F9BC" box="[467,612,1526,1550]" class="Insecta" family="Libellulidae" genus="Orthetrum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="192" pageNumber="637" phylum="Arthropoda" rank="species" species="umbratum" status="sp. nov.">O. umbratum</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A24C821BCAD9FF2AEC48FA1843B8F9BC" box="[618,701,1525,1550]" pageId="192" pageNumber="637" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C0B98F1CAD9FF2AECD3FA1B4258F9BC" box="[753,861,1526,1550]" class="Insecta" family="Libellulidae" genus="Orthetrum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="192" pageNumber="637" phylum="Arthropoda" rank="species" species="saegeri">
<emphasis id="B97F3F60CAD9FF2AECD3FA1B4258F9BC" box="[753,861,1526,1550]" italics="true" pageId="192" pageNumber="637">O. saegeri</emphasis>
</taxonomicName>
).
</paragraph>
</caption>
<paragraph id="8BB4E372CAD8FF2BEEBDFECF4509FED0" pageId="193" pageNumber="638">
Recalls the widespread and syntopic
<taxonomicName id="4C0B98F1CAD8FF2BEC7AFECE43BBFE8F" box="[600,702,290,317]" class="Insecta" family="Libellulidae" genus="Orthetrum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="193" pageNumber="638" phylum="Arthropoda" rank="species" species="hintzi">
<emphasis id="B97F3F60CAD8FF2BEC7AFECE43BBFE8F" box="[600,702,290,317]" italics="true" pageId="193" pageNumber="638">O. hintzi</emphasis>
</taxonomicName>
by (a) small size, Hw 23.7 26.0 mm (n = 6); (b) the absence of cell-doublings in the radial planate and
</paragraph>
<caption id="DF74B3FACAD8FF2BEEBDFBC843E5FBE9" pageId="193" pageNumber="638" targetBox="[158,1035,385,967]" targetPageId="193">
<paragraph id="8BB4E372CAD8FF2BEEBDFBC843E5FBE9" pageId="193" pageNumber="638">
<emphasis id="B97F3F60CAD8FF2BEEBDFBC84049FB8C" bold="true" box="[159,332,1061,1086]" pageId="193" pageNumber="638">
<typeStatus id="FA27DD45FFE200825F407A34B2BF0E19">Type</typeStatus>
<figureCitation id="1330FFF7CAD8FF2BEEFFFBC84042FB8C" box="[221,327,1061,1086]" captionStart="Photo 51" captionStartId="149.[159,225,1498,1522]" captionTargetBox="[159,1035,801,1457]" captionTargetId="figure@149.[159,1038,801,1459]" captionTargetPageId="149" captionText="Photo 51. Paragomphus darwalli, male; Boubou stream near Bongoville, Gabon. Photo: NM (20 - xi- 2011)" pageId="193" pageNumber="638">Photo 51</figureCitation>
.
</emphasis>
<typeStatus id="5E95DD613F553619689B0BB9A8BCBC7E">Holotype</typeStatus>
of
<taxonomicName id="4C0B98F1CAD8FF2BEFC1FBC8439CFB8C" box="[483,665,1061,1086]" class="Insecta" family="Libellulidae" genus="Orthetrum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="193" pageNumber="638" phylum="Arthropoda" rank="species" species="kafwi" status="sp. nov.">
<emphasis id="B97F3F60CAD8FF2BEFC1FBC8439CFB8C" box="[483,665,1061,1086]" italics="true" pageId="193" pageNumber="638">Orthetrum kafwi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C821BCAD8FF2BEC81FBC843FBFB8C" box="[675,766,1061,1086]" pageId="193" pageNumber="638" rank="species">sp. nov.</taxonomicNameLabel>
,
<collectionCode id="6031CDDEE46A07CD7AC38AB7EB3799D1" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]">RMNH</collectionCode>
. Pale abdominal pruinosity has been partly discolored in preservation.
</paragraph>
</caption>
<caption id="DF74B3FACAD8FF2BEEBDFB0241FCF9A2" pageId="193" pageNumber="638">
<paragraph id="8BB4E372CAD8FF2BEEBDFB0241FCF9A2" pageId="193" pageNumber="638">
<emphasis id="B97F3F60CAD8FF2BEEBDFB024014FAB5" bold="true" box="[159,273,1263,1287]" pageId="193" pageNumber="638">Photo 67.</emphasis>
Bog at the source of the
<taxonomicName id="4C0B98F1CAD8FF2BEF43FAE14098FA97" box="[353,413,1292,1317]" class="Insecta" family="Libellulidae" genus="Orthetrum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="193" pageNumber="639" phylum="Arthropoda" rank="species" species="kafwi" status="sp. nov.">Kafwi</taxonomicName>
River on the Kibara Plateau in Upemba National Park,
<taxonomicName id="4C0B98F1CAD8FF2BEF62FA89409FFACF" box="[320,410,1380,1405]" class="Insecta" family="Libellulidae" genus="Neodythemis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="193" pageNumber="631" phylum="Arthropoda" rank="species" species="katanga">Katanga</taxonomicName>
, Congo-Kinshasa. Both
<taxonomicName id="4C0B98F1CAD8FF2BEEBDFA724065FA05" box="[159,352,1439,1463]" class="Insecta" family="Libellulidae" genus="Orthetrum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="193" pageNumber="638" phylum="Arthropoda" rank="species" species="kafwi">
<emphasis id="B97F3F60CAD8FF2BEEBDFA724065FA05" box="[159,352,1439,1463]" italics="true" pageId="193" pageNumber="638">Orthetrum kafwi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0B98F1CAD8FF2BEEBDFA514015FA67" box="[159,272,1468,1493]" class="Insecta" family="Libellulidae" genus="Orthetrum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="193" pageNumber="638" phylum="Arthropoda" rank="species" species="lusinga">
<emphasis id="B97F3F60CAD8FF2BEEBDFA514015FA67" box="[159,272,1468,1493]" italics="true" pageId="193" pageNumber="638">O. lusinga</emphasis>
</taxonomicName>
were found here. Photo: KDBD (15- -
<date id="7FD84210D7F161097080E560E9723771" value="2011-02">ii-2011</date>
)
</paragraph>
</caption>
<paragraph id="8BB4E372CADBFF28EEBDFF1040F9FD00" pageId="194" pageNumber="639">
thus a single row of cells there; (c) the pale subcostal Ax; (d) the large pale Pt that are about 1314 % of Hw length, 3.2
<quantity id="745643F727B48E9B27225F233C639C32" metricMagnitude="-3" metricUnit="m" metricValue="3.5" unit="mm" value="3.5">3.5 mm</quantity>
; and (e) the narrow space between the triangular and out-turned hook and prominent lobe of the hamule (
<figureCitation id="1330FFF7CADBFF28EF11FE804082FE3A" box="[307,391,365,392]" captionStart="Figure 31" captionStartId="189.[784,852,1410,1435]" captionTargetBox="[159,745,910,1550]" captionTargetId="figure@189.[156,749,904,1552]" captionTargetPageId="189" captionText="Figure 31. Secondary genitalia in lateral view of the saegeri - group of Orthetrum and three similar species." pageId="194" pageNumber="639">Fig. 31</figureCitation>
). When not pruinose also similar by (f) the absence of a black line on the metepimeral carina; (g) the dorsa of S410 yellow with black lateral halves; and (h) the cerci always dark. However, (1) the frons anterior to the vertex, as well as the labium, is always pale and never marked with black; and (2) the hamule has a straight rather than convex anterior border, ending in a slender hook that is distinctly longer rather than shorter than it is wide at the point where it turns outward when seen in ventral view (
<figureCitation id="1330FFF7CADBFF28EE85FD9F41FFFD3F" box="[167,250,626,653]" captionStart="Figure 31" captionStartId="189.[784,852,1410,1435]" captionTargetBox="[159,745,910,1550]" captionTargetId="figure@189.[156,749,904,1552]" captionTargetPageId="189" captionText="Figure 31. Secondary genitalia in lateral view of the saegeri - group of Orthetrum and three similar species." pageId="194" pageNumber="639">Fig. 31</figureCitation>
). Also, when not yet pruinose (3) the dorsal carina of S39 is always marked with a thin black line.
</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CADBFF28EEBDFD0941FDFCDF" pageId="194" pageNumber="639" type="etymology">
<paragraph id="8BB4E372CADBFF28EEBDFD094021FD4F" box="[159,292,740,765]" pageId="194" pageNumber="639">
<heading id="D0FC541ECADBFF28EEBDFD094021FD4F" bold="true" box="[159,292,740,765]" fontSize="11" level="3" pageId="194" pageNumber="639" reason="6">
<emphasis id="B97F3F60CADBFF28EEBDFD094021FD4F" bold="true" box="[159,292,740,765]" pageId="194" pageNumber="639">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BB4E372CADBFF28EEBDFCE541FDFCDF" pageId="194" pageNumber="639">
Named after the
<taxonomicName id="4C0B98F1CADBFF28EF46FCE540ACFC90" box="[356,425,776,802]" class="Insecta" family="Libellulidae" genus="Orthetrum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="194" pageNumber="639" phylum="Arthropoda" rank="species" species="kafwi" status="sp. nov.">Kafwi</taxonomicName>
River, of which the
<typeStatus id="073C4211C8D9D55BD242687C24992AA1">type</typeStatus>
locality is the source (noun in apposition). The name also mimics the genitive suffix of most related species.
</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CADBFF28EEBDFC73407DFB9A" pageId="194" pageNumber="639" type="distribution">
<paragraph id="8BB4E372CADBFF28EEBDFC734081FC0A" box="[159,388,926,952]" pageId="194" pageNumber="639">
<heading id="D0FC541ECADBFF28EEBDFC734081FC0A" bold="true" box="[159,388,926,952]" fontSize="11" level="3" pageId="194" pageNumber="639" reason="6">
<emphasis id="B97F3F60CADBFF28EEBDFC734081FC0A" bold="true" box="[159,388,926,952]" pageId="194" pageNumber="639">Range and ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BB4E372CADBFF28EEBDFC2F407DFB9A" pageId="194" pageNumber="639">
Found between 1 760 and 1
<quantity id="12CB5664F8984146FF1642729D04DA3A" metricMagnitude="2" metricUnit="m" metricValue="8.2" unit="m" value="820.0">820 m</quantity>
a.s.l. at boggy areas bordering forest patches on the otherwise grassy Kibara Plateau in Upemba National Park,
<taxonomicName id="4C0B98F1CADBFF28EEBDFBE341FAFB9A" box="[159,255,1038,1064]" class="Insecta" family="Libellulidae" genus="Neodythemis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="194" pageNumber="631" phylum="Arthropoda" rank="species" species="katanga">Katanga</taxonomicName>
(
<figureCitation id="1330FFF7CADBFF28EF2CFBE0406CFB9A" box="[270,361,1037,1064]" captionStart="Map 11" captionStartId="192.[159,210,1496,1521]" captionTargetBox="[158,1035,505,1454]" captionTargetId="figure@192.[156,1038,505,1458]" captionTargetPageId="192" captionText="Map 11. Distribution of the saegeri - group of Orthetrum (O. agaricum sp. nov., O. kafwi sp. nov., O. lusinga, O. umbratum sp. nov. and O. saegeri)." pageId="194" pageNumber="639">Map 11</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>