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<document id="031844E6BF956029998521D865DDD78A" ID-DOI="10.5281/zenodo.35388" ID-GBIF-Dataset="b184dfcc-9b5f-4abb-b401-0f45f76006f6" checkinTime="1450029366780" checkinUser="donat" docAuthor="Dijkstra, Klaas-Douwe B., Kipping, Jens &amp; Mézière, Nicolas" docDate="2015" docId="03A25264CA67FF6AEF2AFC5A4333FD00" docLanguage="en" docName="Odonatologica_44-4-low_res.pdf.imf" docOrigin="Odonatologica 44 (4)" docTitle="Notogomphus gorilla Dijkstra, sp. nov." docType="treatment" docVersion="20" lastPageNumber="573" masterDocId="FF9B2A1CCA19FFEAEE22FFED4105FFB2" masterDocTitle="Sixty new dragonfly and damselfly species from Africa (Odonata)" masterLastPageNumber="678" masterPageNumber="447" pageNumber="571" updateTime="1695001031402" updateUser="admin">
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<mods:title id="656DC3E1DA85571C5E3DBE76663BE2B9">Sixty new dragonfly and damselfly species from Africa (Odonata)</mods:title>
</mods:titleInfo>
<mods:name id="D31C92E7781B987712ED73DC9E97C406" type="personal">
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<mods:namePart id="5AA8FDA4BE0A6196FC6175A2A37AC99B">Dijkstra, Klaas-Douwe B.</mods:namePart>
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<mods:namePart id="F67B77AF8DDDE9835D33CBAEDE802791">Kipping, Jens</mods:namePart>
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<mods:name id="30A2B4D04C635B6DDD5D66FD2B688B4A" type="personal">
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<mods:namePart id="9EA7A54751A6FE40312B8E9C0C0E086D">Mézière, Nicolas</mods:namePart>
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<mods:title id="C0DF182648C518ECBB2E102014E4F603">Odonatologica</mods:title>
</mods:titleInfo>
<mods:part id="4340655CBAF9A82B8BD475387FCD2280">
<mods:date id="592054F1A5A8E70EC10C2EF69C309C69">2015</mods:date>
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<mods:number id="0AA957ADAA08A7376496F67D9E942599">44</mods:number>
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<mods:identifier id="E6472B9E98BC400597A50770CC89B406" type="DOI">10.5281/zenodo.35388</mods:identifier>
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<treatment id="03A25264CA67FF6AEF2AFC5A4333FD00" ID-DOI="http://doi.org/10.5281/zenodo.5640256" ID-GBIF-Taxon="114923440" ID-Zenodo-Dep="5640256" LSID="urn:lsid:zoobank.org:act:03A25264-CA67-FF6A-EF2A-FC5A4333FD00" httpUri="http://treatment.plazi.org/id/03A25264CA67FF6AEF2AFC5A4333FD00" lastPageId="128" lastPageNumber="573" pageId="126" pageNumber="571">
<subSubSection id="C311B0F9CA67FF94EF2AFC5A43FCFC4F" pageId="126" pageNumber="571" type="nomenclature">
<paragraph id="8BB4E372CA67FF94EF2AFC5A43FCFC4F" pageId="126" pageNumber="571">
<heading id="D0FC541ECA67FF94EF2AFC5A43FCFC4F" bold="true" centered="true" fontSize="11" level="2" pageId="126" pageNumber="571" reason="4">
<emphasis id="B97F3F60CA67FF94EF2AFC5A42A7FC60" bold="true" box="[264,930,951,978]" pageId="126" pageNumber="571">
<taxonomicName id="4C0B98F1CA67FF94EF2AFC5A4368FC60" LSID-ZBK="urn:lsid:zoobank.org:act:03A25264-CA67-FF6A-EF2A-FC5A4333FD00" authority="Dijkstra" authorityName="Dijkstra" box="[264,621,951,978]" class="Insecta" family="Gomphidae" genus="Notogomphus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="126" pageNumber="571" phylum="Arthropoda" rank="species" species="gorilla" status="sp. nov.">
<emphasis id="B97F3F60CA67FF94EF2AFC5A430FFC60" bold="true" box="[264,522,951,978]" italics="true" pageId="126" pageNumber="571">Notogomphus gorilla</emphasis>
Dijkstra
</taxonomicName>
<taxonomicNameLabel id="A24C821BCA67FF94EC56FC5A43D4FC60" box="[628,721,951,978]" pageId="126" pageNumber="571" rank="species">sp. nov.</taxonomicNameLabel>
Gorilla Longleg
</emphasis>
(
<typeStatus id="54B05DD0CA67FF94EF9AFC0C40F7FC4F" box="[440,498,993,1021]" pageId="126" pageNumber="571">Type</typeStatus>
<figureCitation id="1330FFF7CA67FF94EFDBFC0C436DFC4E" box="[505,616,993,1020]" captionStart="Photo 33" captionStartId="88.[159,225,1497,1522]" captionTargetBox="[159,1035,875,1455]" captionTargetId="figure@88.[159,1036,874,1458]" captionTargetPageId="88" captionText="Photo 33. Pseudagrion sarepi, male perching on vegetation low over the water; Longa River, Angola. Photo: JK (20 - v- 2012)" pageId="126" pageNumber="571">Photo 33</figureCitation>
,
<figureCitation id="1330FFF7CA67FF94EC50FC0C43CDFC4E" box="[626,712,993,1021]" captionStart="Figure 18" captionStartId="121.[159,227,1466,1491]" captionTargetBox="[249,933,252,1430]" captionTargetId="figure@121.[246,946,249,1429]" captionTargetPageId="121" captionText="Figure 18. Head in rostral (left) and thorax in dorsal (middle) and lateral view (right) of dark Notogomphus species of central Africa; N. bosumbuli sp. nov. is not shown but the head is like N. leroyi and thorax like N. gorilla sp. nov." pageId="126" pageNumber="571">Figs 18</figureCitation>
,
<figureCitation id="1330FFF7CA67FF94ECF0FC0C43F4FC4F" box="[722,753,993,1021]" captionStart="Figure 19" captionStartId="122.[159,227,1496,1521]" captionTargetBox="[158,1034,686,1452]" captionTargetId="figure@122.[156,1038,674,1455]" captionTargetPageId="122" captionText="Figure 19. Male appendages in dorsal (left) and lateral view (right) of selection of Notogomphus species." pageId="126" pageNumber="571">19</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CA67FF94EEBDFBFD401AFB98" box="[159,287,1040,1066]" pageId="126" pageNumber="571" type="description">
<paragraph id="8BB4E372CA67FF94EEBDFBFD401AFB98" box="[159,287,1040,1066]" pageId="126" pageNumber="571">
<heading id="D0FC541ECA67FF94EEBDFBFD401AFB98" bold="true" box="[159,287,1040,1066]" fontSize="11" level="3" pageId="126" pageNumber="571" reason="6">
<emphasis id="B97F3F60CA67FF94EEBDFBFD401AFB98" bold="true" box="[159,287,1040,1066]" pageId="126" pageNumber="571">Taxonomy</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CA67FF95EEBDFBD94374FC0A" lastPageId="127" lastPageNumber="572" pageId="126" pageNumber="571" type="materials_examined">
<paragraph id="8BB4E372CA67FF95EEBDFBD940FBFE4A" lastPageId="127" lastPageNumber="572" pageId="126" pageNumber="571">
<bibRefCitation id="A93350BEF6C40A5B4BB8D365314FA0BA" author="Pinhey E. C. G." journalOrPublisher="British Museum (Natural History), London" refId="ref68651" refString="Pinhey E. C. G. 1961 b. A survey of the dragonflies (order Odonata) of eastern Africa. British Museum (Natural History), London" title="A survey of the dragonflies (order Odonata) of eastern Africa" type="book" year="1961">Pinhey (1961 b)</bibRefCitation>
described a pair of »large
<taxonomicName id="4C0B98F1CA67FF94EC81FBD94244FBFD" box="[675,833,1076,1103]" class="Insecta" family="Gomphidae" genus="Notogomphus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="126" pageNumber="571" phylum="Arthropoda" rank="genus">
<emphasis id="B97F3F60CA67FF94EC81FBD94244FBFD" box="[675,833,1076,1103]" italics="true" pageId="126" pageNumber="571">Notogomphus</emphasis>
</taxonomicName>
, closely allied to
<taxonomicName id="4C0B98F1CA67FF94EEBDFBB7401BFBC6" box="[159,286,1114,1140]" class="Insecta" family="Gomphidae" genus="Notogomphus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="126" pageNumber="571" phylum="Arthropoda" rank="species" species="butoloensis">
<emphasis id="B97F3F60CA67FF94EEBDFBB7401BFBC6" box="[159,286,1114,1140]" italics="true" pageId="126" pageNumber="571">butoloensis</emphasis>
</taxonomicName>
« from Bwamba Forest (= Semliki National Park) in western Uganda under the name
<taxonomicName id="4C0B98F1CA67FF94EFE0FB6D4320FB28" box="[450,549,1151,1178]" class="Insecta" family="Gomphidae" genus="Notogomphus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="126" pageNumber="571" phylum="Arthropoda" rank="species" species="leroyi">
<emphasis id="B97F3F60CA67FF94EFE0FB6D4320FB28" box="[450,549,1151,1178]" italics="true" pageId="126" pageNumber="571">N. leroyi</emphasis>
</taxonomicName>
.
<collectionCode id="ED1A7BB7CA67FF94EC10FB6D4380FB28" box="[562,645,1152,1178]" collectionName="Zimbabwe, Bulawayo, Natural History Museum of Zimbabwe" pageId="126" pageNumber="571">NMBZ</collectionCode>
has the male of this pair and two from sites nearby, all identified as
<taxonomicName id="4C0B98F1CA67FF94EC06FB484383FB0D" box="[548,646,1188,1215]" class="Insecta" family="Gomphidae" genus="Notogomphus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="126" pageNumber="571" phylum="Arthropoda" rank="species" species="leroyi">
<emphasis id="B97F3F60CA67FF94EC06FB484383FB0D" box="[548,646,1188,1215]" italics="true" pageId="126" pageNumber="571">N. leroyi</emphasis>
</taxonomicName>
. These agree with two large males collected recently in south-western Uganda. The
<typeStatus id="54B05DD0CA67FF94ECF7FB274242FB56" box="[725,839,1226,1252]" pageId="126" pageNumber="571">holotypes</typeStatus>
of
<taxonomicName id="4C0B98F1CA67FF94ED49FB26402AFAB8" authority="Fraser, 1952" authorityName="Fraser" authorityYear="1952" class="Insecta" family="Gomphidae" genus="Notogomphus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="126" pageNumber="571" phylum="Arthropoda" rank="species" species="butoloensis">
<emphasis id="B97F3F60CA67FF94ED49FB26450EFB56" box="[875,1035,1226,1252]" italics="true" pageId="126" pageNumber="571">N. butoloensis</emphasis>
Fraser, 1952
</taxonomicName>
(
<collectionCode id="ED1A7BB7CA67FF94EF63FB024098FAB8" box="[321,413,1263,1290]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="126" pageNumber="571">BMNH</collectionCode>
) and
<taxonomicName id="4C0B98F1CA67FF94EFC7FB1D434EFAB8" box="[485,587,1263,1290]" class="Insecta" family="Gomphidae" genus="Notogomphus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="126" pageNumber="571" phylum="Arthropoda" rank="species" species="leroyi">
<emphasis id="B97F3F60CA67FF94EFC7FB1D434EFAB8" box="[485,587,1263,1290]" italics="true" pageId="126" pageNumber="571">N. leroyi</emphasis>
</taxonomicName>
(
<collectionCode id="ED1A7BB7CA67FF94EC7FFB0243B1FAB8" box="[605,692,1263,1290]" collectionName="Belgium, Tervuren, Musee Royal de l'Afrique Centrale" pageId="126" pageNumber="571">MRAC</collectionCode>
) are similar in their moderate size, markings (
<figureCitation id="1330FFF7CA67FF94EFA1FAF940D2FA9D" box="[387,471,1300,1327]" captionStart="Figure 18" captionStartId="121.[159,227,1466,1491]" captionTargetBox="[249,933,252,1430]" captionTargetId="figure@121.[246,946,249,1429]" captionTargetPageId="121" captionText="Figure 18. Head in rostral (left) and thorax in dorsal (middle) and lateral view (right) of dark Notogomphus species of central Africa; N. bosumbuli sp. nov. is not shown but the head is like N. leroyi and thorax like N. gorilla sp. nov." pageId="126" pageNumber="571">Fig. 18</figureCitation>
), clubbed abdomen and distinctive appendages (
<figureCitation id="1330FFF7CA67FF94EE8AFAD741F8FAE6" box="[168,253,1338,1364]" captionStart="Figure 19" captionStartId="122.[159,227,1496,1521]" captionTargetBox="[158,1034,686,1452]" captionTargetId="figure@122.[156,1038,674,1455]" captionTargetPageId="122" captionText="Figure 19. Male appendages in dorsal (left) and lateral view (right) of selection of Notogomphus species." pageId="126" pageNumber="571">Fig. 19</figureCitation>
). While apparently conspecific with each other [new synonymy], they differ from the large males. The female
<typeStatus id="54B05DD0CA67FF94EC9FFAB24221FAC8" box="[701,804,1375,1402]" pageId="126" pageNumber="571">holotype</typeStatus>
of
<taxonomicName id="4C0B98F1CA67FF94ED6DFA8D41D2FA2D" authority="Fraser, 1955" authorityName="Fraser" authorityYear="1955" class="Insecta" family="Gomphidae" genus="Notogomphus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="126" pageNumber="571" phylum="Arthropoda" rank="species" species="anaci">
<emphasis id="B97F3F60CA67FF94ED6DFA8D42B1FAC8" box="[847,948,1376,1402]" italics="true" pageId="126" pageNumber="571">N. anaci</emphasis>
Fraser, 1955
</taxonomicName>
(
<collectionCode id="ED1A7BB7CA67FF94EECAFA694045FA2D" box="[232,320,1412,1439]" collectionName="Belgium, Tervuren, Musee Royal de l'Afrique Centrale" pageId="126" pageNumber="571">MRAC</collectionCode>
) is large and from the same
<typeStatus id="54B05DD0CA67FF94ECBBFA6843C9FA2D" box="[665,716,1413,1439]" pageId="126" pageNumber="571">type</typeStatus>
locality as
<taxonomicName id="4C0B98F1CA67FF94ED76FA6842BCFA2D" box="[852,953,1412,1439]" class="Insecta" family="Gomphidae" genus="Notogomphus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="126" pageNumber="571" phylum="Arthropoda" rank="species" species="leroyi">
<emphasis id="B97F3F60CA67FF94ED76FA6842BCFA2D" box="[852,953,1412,1439]" italics="true" pageId="126" pageNumber="571">N. leroyi</emphasis>
</taxonomicName>
. Three matching topotypical males agree with the
<typeStatus id="54B05DD0CA67FF94ECA9FA4743F8FA76" box="[651,765,1450,1476]" pageId="126" pageNumber="571">holotypes</typeStatus>
of
<taxonomicName id="4C0B98F1CA67FF94ED3CFA46429BFA76" box="[798,926,1451,1476]" class="Insecta" family="Gomphidae" genus="Notogomphus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="126" pageNumber="571" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B97F3F60CA67FF94ED3CFA46429BFA76" box="[798,926,1451,1476]" italics="true" pageId="126" pageNumber="571">N. spinosus</emphasis>
</taxonomicName>
(
<collectionCode id="ED1A7BB7CA67FF94ED8EFA474506FA76" box="[940,1027,1450,1476]" collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" pageId="126" pageNumber="571">ZMHB</collectionCode>
) and
<taxonomicName id="4C0B98F1CA67FF94EEF6FA3D4373FA58" authority="Schouteden, 1934" authorityName="Schouteden" authorityYear="1934" box="[212,630,1487,1514]" class="Insecta" family="Gomphidae" genus="Notogomphus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="126" pageNumber="571" phylum="Arthropoda" rank="species" species="verschuereni">
<emphasis id="B97F3F60CA67FF94EEF6FA3D4082FA58" box="[212,391,1487,1514]" italics="true" pageId="126" pageNumber="571">N. verschuereni</emphasis>
(Schouteden, 1934)
</taxonomicName>
(
<collectionCode id="ED1A7BB7CA67FF94ECA5FA2243DBFA58" box="[647,734,1487,1514]" collectionName="Belgium, Tervuren, Musee Royal de l'Afrique Centrale" pageId="126" pageNumber="571">MRAC</collectionCode>
) in their large size, facial markings (
<figureCitation id="1330FFF7CA67FF94EF3CFA194076F9BD" box="[286,371,1524,1551]" captionStart="Figure 18" captionStartId="121.[159,227,1466,1491]" captionTargetBox="[249,933,252,1430]" captionTargetId="figure@121.[246,946,249,1429]" captionTargetPageId="121" captionText="Figure 18. Head in rostral (left) and thorax in dorsal (middle) and lateral view (right) of dark Notogomphus species of central Africa; N. bosumbuli sp. nov. is not shown but the head is like N. leroyi and thorax like N. gorilla sp. nov." pageId="126" pageNumber="571">Fig. 18</figureCitation>
), appendages (
<figureCitation id="1330FFF7CA67FF94EC07FA19437CF9BD" box="[549,633,1524,1551]" captionStart="Figure 19" captionStartId="122.[159,227,1496,1521]" captionTargetBox="[158,1034,686,1452]" captionTargetId="figure@122.[156,1038,674,1455]" captionTargetPageId="122" captionText="Figure 19. Male appendages in dorsal (left) and lateral view (right) of selection of Notogomphus species." pageId="126" pageNumber="571">Fig. 19</figureCitation>
), and unique green dorsum of S 2 with only two apical black marks. The head structure and vulvar scale, as well as size and coloration, of the
<taxonomicName id="4C0B98F1CA66FF95EC01FECE4381FE8F" box="[547,644,291,317]" class="Insecta" family="Gomphidae" genus="Notogomphus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="127" pageNumber="572" phylum="Arthropoda" rank="species" species="anaci">
<emphasis id="B97F3F60CA66FF95EC01FECE4381FE8F" box="[547,644,291,317]" italics="true" pageId="127" pageNumber="572">N. anaci</emphasis>
</taxonomicName>
<typeStatus id="54B05DD0CA66FF95ECA8FECF43F4FE8F" box="[650,753,290,317]" pageId="127" pageNumber="572">holotype</typeStatus>
and topotypical females of
<taxonomicName id="4C0B98F1CA66FF95EE9FFEA4403AFED0" box="[189,319,329,354]" class="Insecta" family="Gomphidae" genus="Notogomphus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="127" pageNumber="572" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B97F3F60CA66FF95EE9FFEA4403AFED0" box="[189,319,329,354]" italics="true" pageId="127" pageNumber="572">N. spinosus</emphasis>
</taxonomicName>
in
<collectionCode id="ED1A7BB7CA66FF95EF44FEA440BCFED0" box="[358,441,329,354]" collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" pageId="127" pageNumber="572">ZMHB</collectionCode>
and
<collectionCode id="ED1A7BB7CA66FF95EFD1FEA54337FED0" box="[499,562,328,354]" collectionName="Belgium, Brussels, Institut Royal des Sciences Naturelles de Belgique" pageId="127" pageNumber="572">ISNB</collectionCode>
also agree. However, while
<taxonomicName id="4C0B98F1CA66FF95ED55FEA442DDFED0" box="[887,984,329,354]" class="Insecta" family="Gomphidae" genus="Notogomphus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="127" pageNumber="572" phylum="Arthropoda" rank="species" species="anaci">
<emphasis id="B97F3F60CA66FF95ED55FEA442DDFED0" box="[887,984,329,354]" italics="true" pageId="127" pageNumber="572">N. anaci</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0B98F1CA66FF95EEBDFE834055FE3A" box="[159,336,365,392]" class="Insecta" family="Gomphidae" genus="Notogomphus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="127" pageNumber="572" phylum="Arthropoda" rank="species" species="verschuereni">
<emphasis id="B97F3F60CA66FF95EEBDFE834055FE3A" box="[159,336,365,392]" italics="true" pageId="127" pageNumber="572">N. verschuereni</emphasis>
</taxonomicName>
thus appear conspecific with
<taxonomicName id="4C0B98F1CA66FF95EC85FE83422DFE3A" box="[679,808,366,392]" class="Insecta" family="Gomphidae" genus="Notogomphus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="127" pageNumber="572" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B97F3F60CA66FF95EC85FE83422DFE3A" box="[679,808,366,392]" italics="true" pageId="127" pageNumber="572">N. spinosus</emphasis>
</taxonomicName>
[new synonymies], Pinheys species differs and remains unnamed. No genetic data is available, but the species was treated as distinct by
<bibRefCitation id="D840F83D86515CB811407DCCCDB9DE88" author="Dijkstra K. - D. B. &amp; Clausnitzer V." journalOrPublisher="Royal Museum for Central Africa, Tervuren" refId="ref66897" refString="Dijkstra K. - D. B. &amp; Clausnitzer V. 2014. The dragonflies and damselflies of eastern Africa: handbook for all Odonata from Sudan to Zimbabwe. Royal Museum for Central Africa, Tervuren" title="The dragonflies and damselflies of eastern Africa: handbook for all Odonata from Sudan to Zimbabwe" type="book" year="2014">Dijkstra &amp; Clausnitzer (2014)</bibRefCitation>
on account of its morphology.
</paragraph>
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<emphasis id="B97F3F60CA66FF95EEBDFDC5406DFDF1" bold="true" box="[159,360,552,579]" pageId="127" pageNumber="572">Material studied</emphasis>
</heading>
</paragraph>
<paragraph id="8BB4E372CA66FF95EEBDFDA34374FC0A" pageId="127" pageNumber="572">
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,
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,
<location id="152AE9C35093936063FD0970D9FDA2B3" LSID="urn:lsid:plazi:treatment:03A25264CA67FF6AEF2AFC5A4333FD00:152AE9C35093936063FD0970D9FDA2B3" country="Uganda" latitude="-0.9916" longLatPrecision="7" longitude="29.6083" municipality="Kanungu District" name="Buhoma" stateProvince="Western Region">Buhoma</location>
,
<location id="2803CE1A5EE0D1656BA84093C2CC0E1D" LSID="urn:lsid:plazi:treatment:03A25264CA67FF6AEF2AFC5A4333FD00:2803CE1A5EE0D1656BA84093C2CC0E1D" country="Uganda" latitude="-0.9916" longLatPrecision="7" longitude="29.6083" municipality="Kanungu District" name="Munyaga Valley" stateProvince="Western Region">Munyaga Valley</location>
,
<location id="62C88ED12B282609DB3FE50168FC2BF9" LSID="urn:lsid:plazi:treatment:03A25264CA67FF6AEF2AFC5A4333FD00:62C88ED12B282609DB3FE50168FC2BF9" country="Uganda" latitude="-0.9916" longLatPrecision="7" longitude="29.6083" municipality="Kanungu District" name="Munyaga Stream" stateProvince="Western Region">Munyaga Stream</location>
,
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a.s.l. (
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),
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</date>
, leg.
<collectorName id="5296C663C503EA7819761BE4E888A07C">K.-D.B. Dijkstra</collectorName>
,
<collectionCode id="ED1A7BB7CA66FF95ED7BFD7442B3FD00" box="[857,950,665,690]" collectionName="ZMMU" pageId="127" pageNumber="572">ZMMU</collectionCode>
</materialsCitation>
.
<emphasis id="B97F3F60CA66FF95EEBDFD53407DFD6A" bold="true" box="[159,376,702,728]" pageId="127" pageNumber="572">Further material.</emphasis>
<materialsCitation id="3B63E92FCA66FF95EFA6FD53450DFD4F" ID-GBIF-Occurrence="1224547857" collectingDate="1957-10-05" collectionCode="NMBZ" collectorName="E. S. Ross &amp; &amp; R. E. Leech" country="Democratic Republic of the Congo" elevation="1250" location="Ituri, Mont Hoyo" pageId="127" pageNumber="572" specimenCount="1" specimenCount-male="1" stateProvince="Province Orientale">
<collectingCountry id="F31CA3E2CA66FF95EFA6FD534380FD6A" box="[388,645,702,728]" name="Democratic Republic of the Congo" pageId="127" pageNumber="572" value="Democratic Republic of the Congo">CONGO-KINSHASA</collectingCountry>
(
<collectingRegion id="49CF2D90CA66FF95ECB8FD504286FD6A" box="[666,899,701,728]" pageId="127" pageNumber="572">Province Orientale</collectingRegion>
):
<specimenCount id="9D0D28FBCA66FF95EDBEFD5342BBFD65" box="[924,958,701,728]" pageId="127" pageNumber="572" type="generic">1 ♂</specimenCount>
,
<location id="0CFAADA2CE01182F7442D5F624829E12" LSID="urn:lsid:plazi:treatment:03A25264CA67FF6AEF2AFC5A4333FD00:0CFAADA2CE01182F7442D5F624829E12" country="Democratic Republic of the Congo" name="Ituri, Mont Hoyo" stateProvince="Province Orientale">Ituri, Mont Hoyo</location>
,
<elevation id="47016335237ED1038B17D0B8B9B16343" box="[319,406,739,765]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="127" pageNumber="572" unit="m" value="1250.0">
<quantity id="D82F1D93EB700CE44F5361F7C8D3A333" box="[319,406,739,765]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="127" pageNumber="572" unit="m" value="1250.0">1250 m</quantity>
</elevation>
a.s.l.,
<date id="FFB5C5B2CA66FF95EFC2FD0E435CFD4F" box="[480,601,739,765]" pageId="127" pageNumber="572" value="1957-10-05">
<collectingDate id="0C15AC8FB517C6756E9B16DEDA5B453D" box="[480,601,739,765]" pageId="127" pageNumber="572" value="1957-10-05">05-x- 1957</collectingDate>
</date>
, leg.
<collectorName id="34B42D0969A0064A039F804EBD9655F7">E.S. Ross &amp;</collectorName>
<collectorName id="5EB56C5B893F5CCC16434003B7D51A82">R.E. Leech</collectorName>
,
<collectionCode id="ED1A7BB7CA66FF95ED90FD0E450DFD4F" box="[946,1032,739,765]" collectionName="Zimbabwe, Bulawayo, Natural History Museum of Zimbabwe" pageId="127" pageNumber="572">NMBZ</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B63E92FCA66FF95EEBDFCE5450DFCDF" ID-GBIF-Occurrence="1224547863" collectingDate="2002-01-01" collectingDateMax="2003-12-31" collectingDateMin="2002-01-01" collectionCode="ZMMU" collectorName="J. J. Kisakye &amp; &amp; S. G. Kyobe" country="Uganda" elevation="1500" latitude="-0.5912" location="Bwindi Impenetrable National Park" longLatPrecision="7" longitude="29.3642" municipality="Kanungu District" pageId="127" pageNumber="572" specimenCount="1" specimenCount-male="1" stateProvince="Western Region">
<collectingCountry id="F31CA3E2CA66FF95EEBDFCE54019FC90" box="[159,284,776,802]" name="Uganda" pageId="127" pageNumber="572" value="Uganda">UGANDA</collectingCountry>
(
<collectingRegion id="49CF2D90CA66FF95EF0FFCE540F5FC90" box="[301,496,776,802]" pageId="127" pageNumber="572">Western Region</collectingRegion>
):
<specimenCount id="9D0D28FBCA66FF95EC26FCE54323FC90" box="[516,550,776,802]" pageId="127" pageNumber="572" type="generic">1 ♂</specimenCount>
,
<collectingMunicipality id="DC98AF2B19B21667CE9EF283D82EB72E">Kanungu District</collectingMunicipality>
,
<location id="40B94EAF21D19D99CBC979ED9F53AE84" LSID="urn:lsid:plazi:treatment:03A25264CA67FF6AEF2AFC5A4333FD00:40B94EAF21D19D99CBC979ED9F53AE84" country="Uganda" latitude="-0.5912" longLatPrecision="7" longitude="29.3642" municipality="Kanungu District" name="Bwindi Impenetrable National Park" stateProvince="Western Region">Bwindi Impenetrable National Park</location>
,
<location id="69807C62093EB2356A6E3495A53B9010" LSID="urn:lsid:plazi:treatment:03A25264CA67FF6AEF2AFC5A4333FD00:69807C62093EB2356A6E3495A53B9010" country="Uganda" latitude="-0.5912" longLatPrecision="7" longitude="29.3642" municipality="Kanungu District" name="Buhoma" stateProvince="Western Region">Buhoma</location>
,
<location id="33809ECC894B36F7FBBE5B3E42863405" LSID="urn:lsid:plazi:treatment:03A25264CA67FF6AEF2AFC5A4333FD00:33809ECC894B36F7FBBE5B3E42863405" country="Uganda" latitude="-0.5912" longLatPrecision="7" longitude="29.3642" municipality="Kanungu District" name="Munyaga Valley" stateProvince="Western Region">Munyaga Valley</location>
,
<location id="368FB466E416D270F1E093DC929060B0" LSID="urn:lsid:plazi:treatment:03A25264CA67FF6AEF2AFC5A4333FD00:368FB466E416D270F1E093DC929060B0" country="Uganda" latitude="-0.5912" longLatPrecision="7" longitude="29.3642" municipality="Kanungu District" name="Bizenga Stream" stateProvince="Western Region">Bizenga Stream</location>
,
<elevation id="32CF698356D8E04646DA05F26339F6C0" box="[896,974,814,840]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="127" pageNumber="572" unit="m" value="1500.0">
<quantity id="85E7C076C082CD7CCCD177CCB7C14A90" box="[896,974,814,840]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="127" pageNumber="572" unit="m" value="1500.0">1500 m</quantity>
</elevation>
a.s.l. (
<geoCoordinate id="EE3F85B5CA66FF95EE85FCBF400BFCDF" box="[167,270,850,877]" direction="south" orientation="latitude" pageId="127" pageNumber="572" precision="5" value="-0.5912">0.5912 ° S</geoCoordinate>
<geoCoordinate id="EE3F85B5CA66FF95EF3AFCBF4097FCDF" box="[280,402,850,877]" direction="east" orientation="longitude" pageId="127" pageNumber="572" precision="5" value="29.3642">29.3642 ° E</geoCoordinate>
),
<collectingDate id="12A8CE5FFC626FC7A2E27D83E7620165">20022003</collectingDate>
, leg.
<collectorName id="A6BEDA3194DC4CCC072E6994411B4740">J.J. Kisakye &amp;</collectorName>
<collectorName id="80D3285DE7A84FFB022DC345388C3928">S.G. Kyobe</collectorName>
,
<collectionCode id="ED1A7BB7CA66FF95ED8EFCBE450DFCDF" box="[940,1032,851,877]" collectionName="ZMMU" pageId="127" pageNumber="572">ZMMU</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B63E92FCA66FF95EEBDFC954285FC20" ID-GBIF-Occurrence="1224547852" box="[159,896,888,914]" collectingDate="1954-05" collectionCode="NMBZ" collectorName="T. H. E. Jackson" country="Uganda" location="Bwamba" municipality="Bundibugyo" pageId="127" pageNumber="572" specimenCount="1" specimenCount-male="1" stateProvince="Western Region">
<specimenCount id="9D0D28FBCA66FF95EEBDFC9541C4FC20" box="[159,193,888,914]" pageId="127" pageNumber="572" type="generic">1 ♂</specimenCount>
,
<collectingMunicipality id="92A19CB6B481734EB2452A7CB4895BC1">Bundibugyo</collectingMunicipality>
,
<location id="75A8DBD168D367FB1BEDC525C9DF88EB" LSID="urn:lsid:plazi:treatment:03A25264CA67FF6AEF2AFC5A4333FD00:75A8DBD168D367FB1BEDC525C9DF88EB" country="Uganda" municipality="Bundibugyo" name="Bwamba" stateProvince="Western Region">Bwamba</location>
,
<date id="FFB5C5B2CA66FF95EFC3FC954336FC20" box="[481,563,888,914]" pageId="127" pageNumber="572" value="1954-05">
<collectingDate id="80EFC4720E3AAAA89854AE077656A603" box="[481,563,888,914]" pageId="127" pageNumber="572" value="1954-05">v- 1954</collectingDate>
</date>
, leg.
<collectorName id="11BA5708274352F7616900285C64ABF3">T.H.E. Jackson</collectorName>
,
<collectionCode id="ED1A7BB7CA66FF95ED08FC944285FC20" box="[810,896,889,914]" collectionName="Zimbabwe, Bulawayo, Natural History Museum of Zimbabwe" pageId="127" pageNumber="572">NMBZ</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B63E92FCA66FF95EDA9FC954368FC0A" ID-GBIF-Occurrence="1224547860" collectingDate="1957-05" collectionCode="NMBZ" collectorName="unknown" country="Uganda" location="Kayonzi" municipality="Kigezi" pageId="127" pageNumber="572" specimenCount="1" specimenCount-male="1" stateProvince="Western Region">
<specimenCount id="9D0D28FBCA66FF95EDA9FC9542A8FC20" box="[907,941,888,914]" pageId="127" pageNumber="572" type="generic">1 ♂</specimenCount>
,
<collectingMunicipality id="AC32B83F6F541B678063CBCF14961E25">Kigezi</collectingMunicipality>
,
<location id="D9023BA6B2D35E81A3B3820FE9AD4D33" LSID="urn:lsid:plazi:treatment:03A25264CA67FF6AEF2AFC5A4333FD00:D9023BA6B2D35E81A3B3820FE9AD4D33" country="Uganda" municipality="Kigezi" name="Kayonzi" stateProvince="Western Region">Kayonzi</location>
,
<date id="FFB5C5B2CA66FF95EF2EFC73405BFC0A" box="[268,350,926,952]" pageId="127" pageNumber="572" value="1957-05">
<collectingDate id="E4ECA368347B0665A174A3EC146C0283" box="[268,350,926,952]" pageId="127" pageNumber="572" value="1957-05">v- 1957</collectingDate>
</date>
, leg.
<collectorName id="E747021C54E4ABA3AB4AC213CC1D71A5">unknown</collectorName>
,
<collectionCode id="ED1A7BB7CA66FF95EC35FC734368FC0A" box="[535,621,926,952]" collectionName="Zimbabwe, Bulawayo, Natural History Museum of Zimbabwe" pageId="127" pageNumber="572">NMBZ</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CA66FF95EEBDFC05406AFB9A" box="[159,367,1000,1064]" pageId="127" pageNumber="572" type="description">
<paragraph id="8BB4E372CA66FF95EEBDFC05400CFBB1" box="[159,265,1000,1027]" pageId="127" pageNumber="572">
<heading id="D0FC541ECA66FF95EEBDFC05400CFBB1" bold="true" box="[159,265,1000,1027]" fontSize="11" level="3" pageId="127" pageNumber="572" reason="6">
<emphasis id="B97F3F60CA66FF95EEBDFC05400CFBB1" bold="true" box="[159,265,1000,1027]" pageId="127" pageNumber="572">Genetics</emphasis>
</heading>
</paragraph>
<paragraph id="8BB4E372CA66FF95EEBDFBE3406AFB9A" box="[159,367,1037,1064]" pageId="127" pageNumber="572">No data available.</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CA66FF6AEEBDFBB440F1FE3A" lastPageId="128" lastPageNumber="573" pageId="127" pageNumber="572" type="diagnosis">
<paragraph id="8BB4E372CA66FF95EEBDFBB44317FBC1" box="[159,530,1112,1139]" pageId="127" pageNumber="572">
<heading id="D0FC541ECA66FF95EEBDFBB44317FBC1" bold="true" box="[159,530,1112,1139]" fontSize="11" level="3" pageId="127" pageNumber="572" reason="6">
<emphasis id="B97F3F60CA66FF95EEBDFBB44317FBC1" bold="true" box="[159,530,1112,1139]" pageId="127" pageNumber="572">Male morphological diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BB4E372CA66FF6AEEBDFB9340F1FE3A" lastPageId="128" lastPageNumber="573" pageId="127" pageNumber="572">
Nearest to
<taxonomicName id="4C0B98F1CA66FF95EF3FFB93407AFB2A" box="[285,383,1149,1176]" class="Insecta" family="Gomphidae" genus="Notogomphus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="127" pageNumber="572" phylum="Arthropoda" rank="species" species="leroyi">
<emphasis id="B97F3F60CA66FF95EF3FFB93407AFB2A" box="[285,383,1149,1176]" italics="true" pageId="127" pageNumber="572">N. leroyi</emphasis>
</taxonomicName>
by its dark coloration with (a) largely black lower frons, clypeus and labrum; (b) distinct black interpleural and metapleural stripes on the thorax as well as well-defined pale postdorsal and ante-humeral stripes separated from each other and the pale area of the middorsal carina (
<figureCitation id="1330FFF7CA66FF95EE85FAFF41F9FA9F" box="[167,252,1298,1325]" captionStart="Figure 18" captionStartId="121.[159,227,1466,1491]" captionTargetBox="[249,933,252,1430]" captionTargetId="figure@121.[246,946,249,1429]" captionTargetPageId="121" captionText="Figure 18. Head in rostral (left) and thorax in dorsal (middle) and lateral view (right) of dark Notogomphus species of central Africa; N. bosumbuli sp. nov. is not shown but the head is like N. leroyi and thorax like N. gorilla sp. nov." pageId="127" pageNumber="572">Fig. 18</figureCitation>
); (c) widely clubbed S 79, with S 8 about as long as high or wide; (d)quite uniformly dark rufous S 810; and (e) strongly diverging branches of epiproct (
<figureCitation id="1330FFF7CA66FF95EF0EFAB0407BFACA" box="[300,382,1373,1400]" captionStart="Figure 19" captionStartId="122.[159,227,1496,1521]" captionTargetBox="[158,1034,686,1452]" captionTargetId="figure@122.[156,1038,674,1455]" captionTargetPageId="122" captionText="Figure 19. Male appendages in dorsal (left) and lateral view (right) of selection of Notogomphus species." pageId="127" pageNumber="572">Fig. 19</figureCitation>
). Differs, and nearer
<taxonomicName id="4C0B98F1CA66FF95EC53FAB343F7FACA" box="[625,754,1374,1400]" class="Insecta" family="Gomphidae" genus="Notogomphus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="127" pageNumber="572" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B97F3F60CA66FF95EC53FAB343F7FACA" box="[625,754,1374,1400]" italics="true" pageId="127" pageNumber="572">N. spinosus</emphasis>
</taxonomicName>
, by its (1) great size, Hw 36.038.0 mm (n = 5) rather than 31.034.0 mm (n = 4); (2) very small pair of basal green spots on the black labrum; (3) green rather than black occiput, although with a broad black border; (4) humped rather than smoothly curved anterior border of the posterior hamule; and (5) cerci parallel and of rather even width throughout in dorsal view, rather than converging and distally widened (
<figureCitation id="1330FFF7CA99FF6AEF4EFECF40BBFE8F" box="[364,446,290,317]" captionStart="Figure 19" captionStartId="122.[159,227,1496,1521]" captionTargetBox="[158,1034,686,1452]" captionTargetId="figure@122.[156,1038,674,1455]" captionTargetPageId="122" captionText="Figure 19. Male appendages in dorsal (left) and lateral view (right) of selection of Notogomphus species." pageId="128" pageNumber="573">Fig. 19</figureCitation>
). Unlike both
<taxonomicName id="4C0B98F1CA99FF6AEC46FECE43C3FE8F" box="[612,710,290,317]" class="Insecta" family="Gomphidae" genus="Notogomphus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="128" pageNumber="573" phylum="Arthropoda" rank="species" species="leroyi">
<emphasis id="B97F3F60CA99FF6AEC46FECE43C3FE8F" box="[612,710,290,317]" italics="true" pageId="128" pageNumber="573">N. leroyi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0B98F1CA99FF6AECDCFECE427AFE8F" box="[766,895,291,317]" class="Insecta" family="Gomphidae" genus="Notogomphus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="128" pageNumber="573" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B97F3F60CA99FF6AECDCFECE427AFE8F" box="[766,895,291,317]" italics="true" pageId="128" pageNumber="573">N. spinosus</emphasis>
</taxonomicName>
(6) the dorsum of S 7 is black with only yellow smears laterally; and (7) the cerci have a relatively small ventral tooth.
</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CA99FF6AEEBDFE5442E7FDAF" pageId="128" pageNumber="573" type="etymology">
<paragraph id="8BB4E372CA99FF6AEEBDFE544021FE61" box="[159,292,441,467]" pageId="128" pageNumber="573">
<heading id="D0FC541ECA99FF6AEEBDFE544021FE61" bold="true" box="[159,292,441,467]" fontSize="11" level="3" pageId="128" pageNumber="573" reason="6">
<emphasis id="B97F3F60CA99FF6AEEBDFE544021FE61" bold="true" box="[159,292,441,467]" pageId="128" pageNumber="573">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BB4E372CA99FF6AEEBDFE3042E7FDAF" pageId="128" pageNumber="573">
This large dark species is named after the Mountain
<taxonomicName id="4C0B98F1CA99FF6AECD4FE30424CFE4A" box="[758,841,477,504]" class="Mammalia" family="Hominidae" genus="Gorilla" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Primates" pageId="128" pageNumber="573" phylum="Chordata" rank="genus">Gorilla</taxonomicName>
(
<emphasis id="B97F3F60CA99FF6AED7BFE3041FEFDAF" italics="true" pageId="128" pageNumber="573">
<taxonomicName id="4C0B98F1CA99FF6AED7BFE304509FE4A" box="[857,1036,477,504]" class="Mammalia" family="Hominidae" genus="Gorilla" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Primates" pageId="128" pageNumber="573" phylum="Chordata" rank="species" species="beringei">Gorilla beringei</taxonomicName>
<taxonomicName id="4C0B98F1CA99FF6AEEBDFDEF41FEFDAF" box="[159,251,514,541]" class="Mammalia" family="Hominidae" genus="Gorilla" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Primates" pageId="128" pageNumber="573" phylum="Chordata" rank="species" species="beringei">beringei</taxonomicName>
</emphasis>
), for which its
<typeStatus id="54B05DD0CA99FF6AEF89FDEE40D8FDAF" box="[427,477,515,541]" pageId="128" pageNumber="573">type</typeStatus>
locality is best known (noun in apposition).
</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CA99FF6AEEBDFDA34333FD00" pageId="128" pageNumber="573" type="distribution">
<paragraph id="8BB4E372CA99FF6AEEBDFDA34081FDDA" box="[159,388,590,616]" pageId="128" pageNumber="573">
<heading id="D0FC541ECA99FF6AEEBDFDA34081FDDA" bold="true" box="[159,388,590,616]" fontSize="11" level="3" pageId="128" pageNumber="573" reason="6">
<emphasis id="B97F3F60CA99FF6AEEBDFDA34081FDDA" bold="true" box="[159,388,590,616]" pageId="128" pageNumber="573">Range and ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BB4E372CA99FF6AEEBDFD9E4333FD00" pageId="128" pageNumber="573">
Forest streams between about 700 and 1
<quantity id="4CF34E97CA99FF6AEC49FD9E43BAFD3F" box="[619,703,627,653]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="128" pageNumber="573" unit="m" value="600.0">600 m</quantity>
a.s.l. in western Uganda and adjacent Congo-Kinshasa (
<figureCitation id="1330FFF7CA99FF6AEFF8FD754322FD00" box="[474,551,664,690]" captionStart="Map 8" captionStartId="120.[159,210,1463,1488]" captionTargetBox="[159,1035,506,1423]" captionTargetId="figure@120.[156,1038,492,1425]" captionTargetPageId="120" captionText="Map 8. Distribution of N. bosumbuli sp. nov., N. cobyae sp. nov., N. gorilla sp. nov., N. intermedius sp. nov., N. kimpavita sp. nov. and all other species of Notogomphus." pageId="128" pageNumber="573">Map 8</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>