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<document id="3EC3C1E83DE35579BCD8A530ED969762" ID-CLB-Dataset="32021" ID-DOI="10.11646/zootaxa.4323.3.1" ID-GBIF-Dataset="f1be4566-adeb-4995-83c7-2aebab3c7458" ID-ISSN="1175-5326" ID-Zenodo-Dep="919606" ID-ZooBank="3E58F99B-2A30-437A-95Ae-D68A1D3Ecc45" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1506057765384" checkinUser="plazi" docAuthor="Schmidt, Joachim" docDate="2017" docId="03956A03A3737F62FF67FF368E6CD8E0" docLanguage="en" docName="zootaxa.4323.3.1.pdf" docOrigin="Zootaxa 4323 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trichotichnus (Bottchrus) minutus Schmidt 2017, sp. n." docType="treatment" docVersion="7" lastPageNumber="332" masterDocId="FFAC127BA36D7F7DFFF0FFAE8B5CD930" masterDocTitle="Brachypterous ground beetles of the Trichotichnus subgenus Bottchrus Jedlička (Coleoptera, Carabidae) from the Himalaya, with description of fifteen new species" masterLastPageNumber="358" masterPageNumber="301" pageNumber="331" updateTime="1698468866275" updateUser="ExternalLinkService">
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<mods:title id="E648FC33E048E9D869AC4BC947FD9734">Brachypterous ground beetles of the Trichotichnus subgenus Bottchrus Jedlička (Coleoptera, Carabidae) from the Himalaya, with description of fifteen new species</mods:title>
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<paragraph id="8B83DB15A3737F63FF67FF3689DBD982" blockId="30.[151,647,152,211]" box="[151,647,152,178]" pageId="30" pageNumber="331">
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<emphasis id="B9480707A3737F63FF67FF3689DBD982" bold="true" box="[151,647,152,178]" pageId="30" pageNumber="331">
<taxonomicName id="4C3CA096A3737F63FF67FF368961D982" ID-CoL="58LHB" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[151,573,152,178]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="331" phylum="Arthropoda" rank="species" species="minutus" status="sp. nov." subGenus="Bottchrus">
<emphasis id="B9480707A3737F63FF67FF368A1DD982" bold="true" box="[151,321,152,178]" italics="true" pageId="30" pageNumber="331">Trichotichnus</emphasis>
(
<emphasis id="B9480707A3737F63FEA2FF368A95D982" bold="true" box="[338,457,152,178]" italics="true" pageId="30" pageNumber="331">Bottchrus</emphasis>
)
<emphasis id="B9480707A3737F63FE2AFF368961D982" bold="true" box="[474,573,152,178]" italics="true" pageId="30" pageNumber="331">minutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BBA7CA3737F63FDB4FF3689DBD982" box="[580,647,152,178]" pageId="30" pageNumber="331" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B83DB15A3737F63FF67FF148AECD9E3" blockId="30.[151,647,152,211]" box="[151,432,186,211]" pageId="30" pageNumber="331">
(
<figureCitation id="1307C790A3737F63FF6FFF148BA9D9E3" box="[159,245,186,211]" captionStart="FIGURES 67 70" captionStartId="25.[151,264,1947,1970]" captionTargetBox="[311,1277,200,1923]" captionTargetId="figure@25.[305,1282,193,1927]" captionTargetPageId="25" captionText="FIGURES 67 70. Himalayan species of Trichotichnus subgenus Bottchrus, photographs of habitus. 67, T. pusillus sp. n., paratype, male, from the Bara Pokhari Lekh mountain range; 68, T. pusillus sp. n., paratype, male, from the Dudh Pokhari Lekh mountain range; 69, T. ganeshensis sp. n., paratype, male; 70, T. minutus sp. n., paratype, male." httpUri="https://zenodo.org/record/919638/files/figure.png" pageId="30" pageNumber="331">Figs 70</figureCitation>
,
<figureCitation id="1307C790A3737F63FEF1FF158A16D9E3" box="[257,330,187,211]" captionStart="FIGURES 87 90" captionStartId="30.[151,264,1780,1803]" captionTargetBox="[188,1402,964,1744]" captionTargetId="figure@30.[177,1410,948,1759]" captionTargetPageId="30" captionText="FIGURES 87 90. Trichotichnus (Bottchrus) minutus sp. n. (87, paratype; 88 90, holotype). 87 88, 90, median lobe of aedeagus; 89, terminal lamella and apical capitulum. 87 88, lateral view; 89, caudal view; 90, dorsal view. Scale bars = 0.25 mm (A: Figs 87 88, 90; B: Fig. 89)." httpUri="https://zenodo.org/record/919650/files/figure.png" pageId="30" pageNumber="331">8790</figureCitation>
,
<figureCitation id="1307C790A3737F63FEA5FF148ADDD9E3" box="[341,385,186,211]" captionStart="FIGURE 151" captionStartId="53.[151,250,1080,1102]" captionTargetBox="[151,1436,339,1059]" captionTargetId="figure@53.[151,1436,339,1059]" captionTargetPageId="53" captionText="FIGURE 151. Distribution of the Himalayan species of Trichotichnus subgenus Bottchrus. 1, T. holzschuhi Kirschenhofer, 1992; 2, T. baglungensis sp. n.; 3, T. parvulus sp. n.; 4, T. bubsaensis sp. n.; 5, T. schawalleri sp. n.; 6, T. obliquebasalis sp. n.; 7, T. panchhaseensis sp. n.; 8, T. siklesensis sp. n.; 9, T. pusillus sp. n.; 10, T. ganeshensis sp. n.; 11, T. minutus sp. n.; 12, T. cyanescens Ito, 1998; 13, T. martensi sp. n.; 14, T. opacus Ito, 1998." httpUri="https://zenodo.org/record/919694/files/figure.png" pageId="30" pageNumber="331">151</figureCitation>
,
<figureCitation id="1307C790A3737F63FE7DFF158AF9D9E3" box="[397,421,187,211]" captionStart="FIGURES 5 13" captionStartId="6.[151,264,1196,1219]" captionTargetBox="[152,1425,180,1152]" captionTargetId="figure@6.[151,1436,180,1193]" captionTargetPageId="6" captionText="FIGURES 5 13. Trichotichnus (Bottchrus) holzschuhi Kirschenhofer, 1992 (5 7, paratype; 8 13, specimens from Sete). 5, terminal lamella of aedeagus; 6 8, 11, median lobe of aedeagus; 9, apical capitulum; 10, left metepisternum; 12, apex of last (VII) abdominal sternite, male; 13, same, female. 5, 7, 8, dorsal view; 6, 11, lateral view; 9, caudal view; 10, 12, 13, ventral view. Scale bars = 0.25 mm (A: Figs 5 8, 10 13; B: Fig. 9)." httpUri="https://zenodo.org/record/919610/files/figure.png" pageId="30" pageNumber="331">
<emphasis id="B9480707A3737F63FE7DFF158AF9D9E3" box="[397,421,187,211]" italics="true" pageId="30" pageNumber="331">11</emphasis>
</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C326889EA3737F63FF67FEAC8A14D8B7" pageId="30" pageNumber="331" type="materials_examined">
<paragraph id="8B83DB15A3737F63FF67FEAC8AB7D80F" blockId="30.[151,1437,258,895]" pageId="30" pageNumber="331">
<materialsCitation id="3B54D148A3737F63FF67FEAC8AB7D80F" ID-GBIF-Occurrence="1655986837" collectingDate="2001-05-06" collectorName="Godawari, Mt. &amp; O. Jager" country="Nepal" elevation="1750" location="Kathmandu-Tal Godawari" municipality="Mt. Phulchoki" pageId="30" pageNumber="331" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B9480707A3737F63FF67FEAC8A63D82B" bold="true" box="[151,319,258,283]" pageId="30" pageNumber="331">
<typeStatus id="548765B7A3737F63FF67FEAC8B8ED82B" box="[151,210,258,283]" pageId="30" pageNumber="331">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="548765B7A3737F63FEBEFEAC8AE7D82B" box="[334,443,258,283]" pageId="30" pageNumber="331" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D3A109CA3737F63FE36FEAD8A81D82A" box="[454,477,259,282]" pageId="30" pageNumber="331" type="male"></specimenCount>
, “
<collectingCountry id="F32B9B85A3737F63FE05FEAD890ED82B" box="[501,594,259,283]" name="Nepal" pageId="30" pageNumber="331">NEPAL</collectingCountry>
,
<location id="8EE38DCEA3737F63FDAFFEAC88D4D82B" LSID="urn:lsid:plazi:treatment:03956A03A3737F62FF67FF368E6CD8E0:8EE38DCEA3737F63FDAFFEAC88D4D82B" box="[607,904,258,283]" country="Nepal" municipality="Mt. Phulchoki" name="Kathmandu-Tal Godawari" pageId="30" pageNumber="331">
Kathmandu-Tal
<collectorName id="26C9BEC3A3737F63FCE7FEAC88D4D82B" box="[791,904,258,283]" pageId="30" pageNumber="331">Godawari</collectorName>
</location>
,
<collectingMunicipality id="6BE7416FA3737F63FC64FEAD8F6BD82B" box="[916,1079,258,283]" pageId="30" pageNumber="331">Mt. Phulchoki</collectingMunicipality>
N-Hang,
<elevation id="00113C26A3737F63FB58FEAC8E5DD82A" box="[1192,1281,258,283]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="30" pageNumber="331" unit="m" value="1750.0">
<quantity id="4CC476F0A3737F63FB58FEAC8E5DD82A" box="[1192,1281,258,283]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="30" pageNumber="331" unit="m" value="1750.0">1750 m</quantity>
</elevation>
, Ufergesiebe,
<collectingDate id="EFC6043DA3737F63FF67FE888BA2D80F" box="[151,254,294,319]" pageId="30" pageNumber="331" value="2001-05-06">
<date id="FF82FDD5A3737F63FF67FE888BA2D80F" box="[151,254,294,319]" pageId="30" pageNumber="331" value="2001-05-06">6.V.2001</date>
</collectingDate>
, leg.
<collectorName id="26C9BEC3A3737F63FECAFE898AC5D80F" box="[314,409,295,319]" pageId="30" pageNumber="331">O. Jäger</collectorName>
” (cJS).
</materialsCitation>
</paragraph>
<paragraph id="8B83DB15A3737F63FF37FEE58A14D8B7" blockId="30.[151,1437,258,895]" pageId="30" pageNumber="331">
<materialsCitation id="3B54D148A3737F63FF37FEE58A14D8B7" ID-GBIF-Occurrence="1655986830" collectingDate="05-05-23" collectorName="N Kathmandu Shivapuri Lekh &amp; Schmidt" country="Nepal" elevation="1151" location="Bagmati" pageId="30" pageNumber="331" specimenCount="1" specimenCount-male="1" stateProvince="Bagmati" typeStatus="paratype">
<typeStatus id="548765B7A3737F63FF37FEE58A71D853" box="[199,301,331,355]" pageId="30" pageNumber="331" type="paratype">Paratype</typeStatus>
:
<specimenCount id="9D3A109CA3737F63FECBFEE58A36D852" box="[315,362,331,355]" pageId="30" pageNumber="331" type="male">1 ♂</specimenCount>
, “
<collectingCountry id="F32B9B85A3737F63FE72FEE58A81D853" box="[386,477,331,355]" name="Nepal" pageId="30" pageNumber="331">NEPAL</collectingCountry>
<collectorName id="26C9BEC3A3737F63FE16FEE5881DD853" box="[486,833,330,355]" pageId="30" pageNumber="331">N Kathmandu Shivapuri Lekh</collectorName>
<elevation id="00113C26A3737F63FCBCFEE588E3D852" box="[844,959,331,355]" metricMagnitude="3" metricUnit="m" metricValue="1.151" metricValueMax="2.3" metricValueMin="0.002" pageId="30" pageNumber="331" unit="m" value="1151.0" valueMax="2300.0" valueMin="2.0">
<quantity id="4CC476F0A3737F63FCBCFEE588E3D852" box="[844,959,331,355]" metricMagnitude="3" metricUnit="m" metricValue="1.151" metricValueMax="2.3" metricValueMin="0.002" pageId="30" pageNumber="331" unit="m" value="1151.0" valueMax="2300.0" valueMin="2.0">22300 m</quantity>
</elevation>
slope W of
<collectingRegion id="49F815F7A3737F63FBA4FEE48FEFD853" box="[1108,1203,330,355]" country="Nepal" name="Bagmati" pageId="30" pageNumber="331">Bagmati</collectingRegion>
riv. 22
<collectingDate id="EFC6043DA3737F63FAE5FEE48E35D853" box="[1301,1385,330,355]" pageId="30" pageNumber="331" value="05-05-23">
<date id="FF82FDD5A3737F63FAE5FEE48E35D853" box="[1301,1385,330,355]" pageId="30" pageNumber="331" value="05-05-23">23.5.05</date>
</collectingDate>
leg.
<collectorName id="26C9BEC3A3737F63FF67FEC08BABD8B7" box="[151,247,366,391]" pageId="30" pageNumber="331">Schmidt</collectorName>
” (cJS).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C326889EA3737F63FF37FE3C8A52DA4F" pageId="30" pageNumber="331" type="description">
<paragraph id="8B83DB15A3737F63FF37FE3C8E4EDA23" blockId="30.[151,1437,258,895]" pageId="30" pageNumber="331">
<emphasis id="B9480707A3737F63FF37FE3C8A0DD89B" bold="true" box="[199,337,402,427]" pageId="30" pageNumber="331">Description</emphasis>
. Habitus as in
<figureCitation id="1307C790A3737F63FE07FE3C8916D89B" box="[503,586,402,427]" captionStart="FIGURES 67 70" captionStartId="25.[151,264,1947,1970]" captionTargetBox="[311,1277,200,1923]" captionTargetId="figure@25.[305,1282,193,1927]" captionTargetPageId="25" captionText="FIGURES 67 70. Himalayan species of Trichotichnus subgenus Bottchrus, photographs of habitus. 67, T. pusillus sp. n., paratype, male, from the Bara Pokhari Lekh mountain range; 68, T. pusillus sp. n., paratype, male, from the Dudh Pokhari Lekh mountain range; 69, T. ganeshensis sp. n., paratype, male; 70, T. minutus sp. n., paratype, male." httpUri="https://zenodo.org/record/919638/files/figure.png" pageId="30" pageNumber="331">Fig. 70</figureCitation>
. Body length 5.56.0 mm, width
<quantity id="4CC476F0A3737F63FC4EFE3C8F14D89A" box="[958,1096,402,427]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" metricValueMax="2.7" metricValueMin="2.5" pageId="30" pageNumber="331" unit="mm" value="2.6" valueMax="2.7" valueMin="2.5">2.52.7 mm</quantity>
. Characters as described for
<taxonomicName id="4C3CA096A3737F63FA77FE3A8A0DD8FF" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="331" phylum="Arthropoda" rank="species" species="ganeshensis" status="sp. nov." subGenus="Bottchrus">
<emphasis id="B9480707A3737F63FA77FE3A8EC1D89B" box="[1415,1437,404,427]" italics="true" pageId="30" pageNumber="331">T.</emphasis>
(
<emphasis id="B9480707A3737F63FF50FE198BEDD8FE" box="[160,177,439,462]" italics="true" pageId="30" pageNumber="331">B</emphasis>
.)
<emphasis id="B9480707A3737F63FF3AFE188A0DD8FF" box="[202,337,438,463]" italics="true" pageId="30" pageNumber="331">ganeshensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BBA7CA3737F63FEA9FE188AC6D8FF" box="[345,410,438,463]" pageId="30" pageNumber="331" rank="species">
<emphasis id="B9480707A3737F63FEA9FE188AC6D8FF" bold="true" box="[345,410,438,463]" pageId="30" pageNumber="331">sp. n.</emphasis>
</taxonomicNameLabel>
except as follows: Eyes in lateral view slightly more elongate. Pronotum relatively wider, with basal angles subdenticulate; basal margin vaguely bordered laterally (in
<typeStatus id="548765B7A3737F63FBE0FE748F28D8C3" box="[1040,1140,474,499]" pageId="30" pageNumber="331" type="holotype">holotype</typeStatus>
) or almost throughout (in
<typeStatus id="548765B7A3737F63FF67FE518BA1DB27" box="[151,253,511,535]" pageId="30" pageNumber="331" type="paratype">paratype</typeStatus>
); pronotal lateral gutter very narrow almost equal throughout (in
<typeStatus id="548765B7A3737F63FC14FE508F14DB27" box="[996,1096,510,535]" pageId="30" pageNumber="331" type="holotype">holotype</typeStatus>
) or slightly widened in apical third (in
<typeStatus id="548765B7A3737F63FEF3FD8D8A35DB0B" box="[259,361,547,571]" pageId="30" pageNumber="331" type="paratype">paratype</typeStatus>
). Elytral striae smooth, not crenulate, slightly impressed along entire length; intervals with slightly more distinct microsculpture. Legs yellowish brown, with femora infuscate. Median lobe of aedeagus (
<figureCitation id="1307C790A3737F63FA9AFDE88B82DBB3" captionStart="FIGURES 87 90" captionStartId="30.[151,264,1780,1803]" captionTargetBox="[188,1402,964,1744]" captionTargetId="figure@30.[177,1410,948,1759]" captionTargetPageId="30" captionText="FIGURES 87 90. Trichotichnus (Bottchrus) minutus sp. n. (87, paratype; 88 90, holotype). 87 88, 90, median lobe of aedeagus; 89, terminal lamella and apical capitulum. 87 88, lateral view; 89, caudal view; 90, dorsal view. Scale bars = 0.25 mm (A: Figs 87 88, 90; B: Fig. 89)." httpUri="https://zenodo.org/record/919650/files/figure.png" pageId="30" pageNumber="331">Figs 8790</figureCitation>
) with ventral margin slightly convex in middle portion, terminal lamella angularly bent ventrad and sinuate at sides; apical capitulum oblique in lateral view, comparatively large, markedly larger than in
<taxonomicName id="4C3CA096A3737F63FB36FD218EC7DB97" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[1222,1435,654,679]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="331" phylum="Arthropoda" rank="species" species="ganeshensis" status="sp. nov." subGenus="Bottchrus">
<emphasis id="B9480707A3737F63FB36FD218F87DB96" box="[1222,1243,655,678]" italics="true" pageId="30" pageNumber="331">T.</emphasis>
(
<emphasis id="B9480707A3737F63FB1BFD218FA0DB96" box="[1259,1276,655,678]" italics="true" pageId="30" pageNumber="331">B</emphasis>
.)
<emphasis id="B9480707A3737F63FAE4FD208EC7DB97" box="[1300,1435,654,679]" italics="true" pageId="30" pageNumber="331">ganeshensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BBA7CA3737F63FF67FD1C8B85DBFB" box="[151,217,690,715]" pageId="30" pageNumber="331" rank="species">
<emphasis id="B9480707A3737F63FF67FD1C8B85DBFB" bold="true" box="[151,217,690,715]" pageId="30" pageNumber="331">sp. n.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="4C3CA096A3737F63FEE4FD1A8AE2DBFB" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[276,446,690,715]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="331" phylum="Arthropoda" rank="species" species="pusillus" status="sp. nov." subGenus="Bottchrus">
<emphasis id="B9480707A3737F63FEE4FD1A8A74DBFB" box="[276,296,692,715]" italics="true" pageId="30" pageNumber="331">T.</emphasis>
(
<emphasis id="B9480707A3737F63FECCFD1A8A11DBFB" box="[316,333,692,715]" italics="true" pageId="30" pageNumber="331">B</emphasis>
.)
<emphasis id="B9480707A3737F63FE97FD1C8AE2DBFB" box="[359,446,690,715]" italics="true" pageId="30" pageNumber="331">pusillus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BBA7CA3737F63FE38FD1C8956DBFB" box="[456,522,690,715]" pageId="30" pageNumber="331" rank="species">
<emphasis id="B9480707A3737F63FE38FD1C8956DBFB" bold="true" box="[456,522,690,715]" pageId="30" pageNumber="331">sp. n.</emphasis>
</taxonomicNameLabel>
, approximately equally prominent ventrally and dorsally, in caudal view apical capitulum, as in
<taxonomicName id="4C3CA096A3737F63FEA0FD798978DBDF" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[336,548,726,751]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="331" phylum="Arthropoda" rank="species" species="ganeshensis" status="sp. nov." subGenus="Bottchrus">
<emphasis id="B9480707A3737F63FEA0FD798A39DBDE" box="[336,357,727,750]" italics="true" pageId="30" pageNumber="331">T.</emphasis>
(
<emphasis id="B9480707A3737F63FE84FD798AD9DBDE" box="[372,389,727,750]" italics="true" pageId="30" pageNumber="331">B</emphasis>
.)
<emphasis id="B9480707A3737F63FE6DFD788978DBDF" box="[413,548,726,751]" italics="true" pageId="30" pageNumber="331">ganeshensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BBA7CA3737F63FDDBFD788936DBDF" box="[555,618,726,751]" pageId="30" pageNumber="331" rank="species">
<emphasis id="B9480707A3737F63FDDBFD788936DBDF" bold="true" box="[555,618,726,751]" pageId="30" pageNumber="331">sp. n.</emphasis>
</taxonomicNameLabel>
, somewhat semi-oval, but lightly more elongate, narrowly rounded at tip (
<figureCitation id="1307C790A3737F63FF6FFD548BAFDA23" box="[159,243,762,787]" captionStart="FIGURES 87 90" captionStartId="30.[151,264,1780,1803]" captionTargetBox="[188,1402,964,1744]" captionTargetId="figure@30.[177,1410,948,1759]" captionTargetPageId="30" captionText="FIGURES 87 90. Trichotichnus (Bottchrus) minutus sp. n. (87, paratype; 88 90, holotype). 87 88, 90, median lobe of aedeagus; 89, terminal lamella and apical capitulum. 87 88, lateral view; 89, caudal view; 90, dorsal view. Scale bars = 0.25 mm (A: Figs 87 88, 90; B: Fig. 89)." httpUri="https://zenodo.org/record/919650/files/figure.png" pageId="30" pageNumber="331">Fig. 89</figureCitation>
). Internal sac with characteristic pattern of spiny folders in the middle portion of median lobe.
</paragraph>
<paragraph id="8B83DB15A3737F63FF37FCB08A52DA4F" blockId="30.[151,1437,258,895]" pageId="30" pageNumber="331">Proportions (males): PWmax/PL = 1.421.43; EL/EW = 1.321.36; EL/PL = 2.362.42; EW/PWmax = 1.24 1.25; HWmax/PWmax = 0.67; HWmin/PWmax = 0.540.55; PWmax/PWmin = 1.231.25; HWmax/HWmin = 1.221.23.</paragraph>
</subSubSection>
<caption id="DF438B9DA3737F63FF67F95A895ADE78" httpUri="https://zenodo.org/record/919650/files/figure.png" pageId="30" pageNumber="331" targetBox="[188,1402,964,1744]" targetPageId="30">
<paragraph id="8B83DB15A3737F63FF67F95A895ADE78" blockId="30.[151,1436,1780,1864]" pageId="30" pageNumber="331">
<emphasis id="B9480707A3737F63FF67F95A8A05DE3A" bold="true" box="[151,345,1780,1803]" pageId="30" pageNumber="331">FIGURES 8790.</emphasis>
<taxonomicName id="4C3CA096A3737F63FE94F95A8995DE3B" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[356,713,1780,1803]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="331" phylum="Arthropoda" rank="species" species="minutus" status="sp. nov." subGenus="Bottchrus">
<emphasis id="B9480707A3737F63FE94F95A8AB3DE3A" box="[356,495,1780,1802]" italics="true" pageId="30" pageNumber="331">Trichotichnus</emphasis>
(
<emphasis id="B9480707A3737F63FDF3F95A8939DE3A" box="[515,613,1780,1802]" italics="true" pageId="30" pageNumber="331">Bottchrus</emphasis>
)
<emphasis id="B9480707A3737F63FD89F9588995DE3B" box="[633,713,1782,1803]" italics="true" pageId="30" pageNumber="331">minutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BBA7CA3737F63FD24F95B884EDE3A" box="[724,786,1781,1802]" pageId="30" pageNumber="331" rank="species">
<emphasis id="B9480707A3737F63FD24F95B884EDE3A" bold="true" box="[724,786,1781,1802]" pageId="30" pageNumber="331">sp. n.</emphasis>
</taxonomicNameLabel>
(87, paratype; 8890, holotype). 8788, 90, median lobe of aedeagus; 89, terminal lamella and apical capitulum. 8788, lateral view; 89, caudal view; 90, dorsal view. Scale bars = 0.25 mm (A: Figs 8788, 90; B: Fig. 89).
</paragraph>
</caption>
<subSubSection id="C326889EA3737F62FF37F8DA8FE8D854" lastPageId="31" lastPageNumber="332" pageId="30" pageNumber="331" type="etymology">
<paragraph id="8B83DB15A3737F63FF37F8DA8894DEBD" blockId="30.[151,1436,1908,2005]" box="[199,968,1908,1933]" pageId="30" pageNumber="331">
<emphasis id="B9480707A3737F63FF37F8DA8A1ADEBD" bold="true" box="[199,326,1908,1933]" pageId="30" pageNumber="331">Etymology</emphasis>
. The specific name refers to the small size of the beetles.
</paragraph>
<paragraph id="8B83DB15A3737F62FF37F8368FE8D854" blockId="30.[151,1436,1908,2005]" lastBlockId="31.[151,1437,151,464]" lastPageId="31" lastPageNumber="332" pageId="30" pageNumber="331">
<emphasis id="B9480707A3737F63FF37F8368A06DE81" bold="true" box="[199,346,1944,1969]" pageId="30" pageNumber="331">Comparison</emphasis>
. This new species is very similar in size and external morphology to
<taxonomicName id="4C3CA096A3737F63FB85F8348E10DE81" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[1141,1356,1944,1969]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="331" phylum="Arthropoda" rank="species" species="ganeshensis" status="sp. nov." subGenus="Bottchrus">
<emphasis id="B9480707A3737F63FB85F8348FD6DE81" box="[1141,1162,1946,1969]" italics="true" pageId="30" pageNumber="331">T.</emphasis>
(
<emphasis id="B9480707A3737F63FB6BF8348FF0DE81" box="[1179,1196,1946,1969]" italics="true" pageId="30" pageNumber="331">B</emphasis>
.)
<emphasis id="B9480707A3737F63FB35F8368E10DE81" box="[1221,1356,1944,1969]" italics="true" pageId="30" pageNumber="331">ganeshensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BBA7CA3737F63FAA4F8368EC9DE81" box="[1364,1429,1944,1969]" pageId="30" pageNumber="331" rank="species">
<emphasis id="B9480707A3737F63FAA4F8368EC9DE81" bold="true" box="[1364,1429,1944,1969]" pageId="30" pageNumber="331">sp. n.</emphasis>
</taxonomicNameLabel>
, but distinctly differs from it in the male genitalia as follows: aedeagal median lobe (
<figureCitation id="1307C790A3737F63FBB4F8128F98DEE5" box="[1092,1220,1980,2005]" captionStart="FIGURES 87 90" captionStartId="30.[151,264,1780,1803]" captionTargetBox="[188,1402,964,1744]" captionTargetId="figure@30.[177,1410,948,1759]" captionTargetPageId="30" captionText="FIGURES 87 90. Trichotichnus (Bottchrus) minutus sp. n. (87, paratype; 88 90, holotype). 87 88, 90, median lobe of aedeagus; 89, terminal lamella and apical capitulum. 87 88, lateral view; 89, caudal view; 90, dorsal view. Scale bars = 0.25 mm (A: Figs 87 88, 90; B: Fig. 89)." httpUri="https://zenodo.org/record/919650/files/figure.png" pageId="30" pageNumber="331">Figs 8790</figureCitation>
) relatively smaller, with ventral margin in lateral view slightly convex in middle portion, apical capitulum much larger, approximately equally prominent ventrally and dorsally [dorsal flange of the apical capitulum in
<taxonomicName id="4C3CA096A3727F62FBCBFF138E4DD9E5" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[1083,1297,188,213]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="332" phylum="Arthropoda" rank="species" species="ganeshensis" status="sp. nov." subGenus="Bottchrus">
<emphasis id="B9480707A3727F62FBCBFF138F0CD9E4" box="[1083,1104,189,212]" italics="true" pageId="31" pageNumber="332">T.</emphasis>
(
<emphasis id="B9480707A3727F62FB91FF138F2ED9E4" box="[1121,1138,189,212]" italics="true" pageId="31" pageNumber="332">B</emphasis>
.)
<emphasis id="B9480707A3727F62FB7AFF128E4DD9E5" box="[1162,1297,188,213]" italics="true" pageId="31" pageNumber="332">ganeshensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BBA7CA3727F62FAEAFF128E07D9E5" box="[1306,1371,188,213]" pageId="31" pageNumber="332" rank="species">
<emphasis id="B9480707A3727F62FAEAFF128E07D9E5" bold="true" box="[1306,1371,188,213]" pageId="31" pageNumber="332">sp. n.</emphasis>
</taxonomicNameLabel>
more prominent than ventral flange, figs. 8182], in caudal view apical capitulum, as in
<taxonomicName id="4C3CA096A3727F62FB9AFF4F8E1AD9C8" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[1130,1350,223,248]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="332" phylum="Arthropoda" rank="species" species="ganeshensis" status="sp. nov." subGenus="Bottchrus">
<emphasis id="B9480707A3727F62FB9AFF4F8F23D9C8" box="[1130,1151,225,248]" italics="true" pageId="31" pageNumber="332">T.</emphasis>
(
<emphasis id="B9480707A3727F62FB63FF4F8FF8D9C8" box="[1171,1188,225,248]" italics="true" pageId="31" pageNumber="332">B</emphasis>
.)
<emphasis id="B9480707A3727F62FB4FFF718E1AD9C8" box="[1215,1350,223,248]" italics="true" pageId="31" pageNumber="332">ganeshensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BBA7CA3727F62FAA2FF718EC9D9C8" box="[1362,1429,223,248]" pageId="31" pageNumber="332" rank="species">
<emphasis id="B9480707A3727F62FAA2FF718EC9D9C8" bold="true" box="[1362,1429,223,248]" pageId="31" pageNumber="332">sp. n.</emphasis>
</taxonomicNameLabel>
, somewhat semi-oval, but slightly more elongate, narrowly rounded at tip (
<figureCitation id="1307C790A3727F62FC0FFEAA8F0BD82C" box="[1023,1111,260,285]" captionStart="FIGURES 87 90" captionStartId="30.[151,264,1780,1803]" captionTargetBox="[188,1402,964,1744]" captionTargetId="figure@30.[177,1410,948,1759]" captionTargetPageId="30" captionText="FIGURES 87 90. Trichotichnus (Bottchrus) minutus sp. n. (87, paratype; 88 90, holotype). 87 88, 90, median lobe of aedeagus; 89, terminal lamella and apical capitulum. 87 88, lateral view; 89, caudal view; 90, dorsal view. Scale bars = 0.25 mm (A: Figs 87 88, 90; B: Fig. 89)." httpUri="https://zenodo.org/record/919650/files/figure.png" pageId="31" pageNumber="332">Fig. 89</figureCitation>
); internal sac with different pattern of spiny folders in the middle portion of the median lobe. Additionally, pronotum in
<taxonomicName id="4C3CA096A3727F62FB5CFE878E08D870" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[1196,1364,296,320]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="332" phylum="Arthropoda" rank="species" species="minutus" status="sp. nov." subGenus="Bottchrus">
<emphasis id="B9480707A3727F62FB5CFE878F9DD870" box="[1196,1217,297,320]" italics="true" pageId="31" pageNumber="332">T.</emphasis>
(
<emphasis id="B9480707A3727F62FB22FE878FBFD870" box="[1234,1251,297,320]" italics="true" pageId="31" pageNumber="332">B</emphasis>
.)
<emphasis id="B9480707A3727F62FB0BFE878E08D870" box="[1275,1364,297,320]" italics="true" pageId="31" pageNumber="332">minutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BBA7CA3727F62FAACFE898EC0D870" box="[1372,1436,295,320]" pageId="31" pageNumber="332" rank="species">
<emphasis id="B9480707A3727F62FAACFE898EC0D870" bold="true" box="[1372,1436,295,320]" pageId="31" pageNumber="332">sp. n.</emphasis>
</taxonomicNameLabel>
relatively wider than in
<taxonomicName id="4C3CA096A3727F62FE52FEE3892AD855" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[418,630,332,357]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="332" phylum="Arthropoda" rank="species" species="ganeshensis" status="sp. nov." subGenus="Bottchrus">
<emphasis id="B9480707A3727F62FE52FEE38AEBD854" box="[418,439,333,356]" italics="true" pageId="31" pageNumber="332">T.</emphasis>
(
<emphasis id="B9480707A3727F62FE37FEE38A84D854" box="[455,472,333,356]" italics="true" pageId="31" pageNumber="332">B</emphasis>
.)
<emphasis id="B9480707A3727F62FE1FFEE2892AD855" box="[495,630,332,357]" italics="true" pageId="31" pageNumber="332">ganeshensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BBA7CA3727F62FD8EFEE289E1D855" box="[638,701,332,357]" pageId="31" pageNumber="332" rank="species">
<emphasis id="B9480707A3727F62FD8EFEE289E1D855" bold="true" box="[638,701,332,357]" pageId="31" pageNumber="332">sp. n.</emphasis>
</taxonomicNameLabel>
(PWmax/PL = 1.421.43 versus 1.321.39).
</paragraph>
</subSubSection>
<subSubSection id="C326889EA3727F62FF37FEC18E6CD8E0" pageId="31" pageNumber="332" type="distribution">
<paragraph id="8B83DB15A3727F62FF37FEC18E6CD8E0" blockId="31.[151,1437,151,464]" pageId="31" pageNumber="332">
<emphasis id="B9480707A3727F62FF37FEC18A05D8B8" bold="true" box="[199,345,367,392]" pageId="31" pageNumber="332">Distribution</emphasis>
(
<figureCitation id="1307C790A3727F62FE9AFEC18A97D8B8" box="[362,459,367,392]" captionStart="FIGURE 151" captionStartId="53.[151,250,1080,1102]" captionTargetBox="[151,1436,339,1059]" captionTargetId="figure@53.[151,1436,339,1059]" captionTargetPageId="53" captionText="FIGURE 151. Distribution of the Himalayan species of Trichotichnus subgenus Bottchrus. 1, T. holzschuhi Kirschenhofer, 1992; 2, T. baglungensis sp. n.; 3, T. parvulus sp. n.; 4, T. bubsaensis sp. n.; 5, T. schawalleri sp. n.; 6, T. obliquebasalis sp. n.; 7, T. panchhaseensis sp. n.; 8, T. siklesensis sp. n.; 9, T. pusillus sp. n.; 10, T. ganeshensis sp. n.; 11, T. minutus sp. n.; 12, T. cyanescens Ito, 1998; 13, T. martensi sp. n.; 14, T. opacus Ito, 1998." httpUri="https://zenodo.org/record/919694/files/figure.png" pageId="31" pageNumber="332">Fig. 151</figureCitation>
,
<figureCitation id="1307C790A3727F62FE29FEDE8AADD8B8" box="[473,497,368,392]" captionStart="FIGURES 5 13" captionStartId="6.[151,264,1196,1219]" captionTargetBox="[152,1425,180,1152]" captionTargetId="figure@6.[151,1436,180,1193]" captionTargetPageId="6" captionText="FIGURES 5 13. Trichotichnus (Bottchrus) holzschuhi Kirschenhofer, 1992 (5 7, paratype; 8 13, specimens from Sete). 5, terminal lamella of aedeagus; 6 8, 11, median lobe of aedeagus; 9, apical capitulum; 10, left metepisternum; 12, apex of last (VII) abdominal sternite, male; 13, same, female. 5, 7, 8, dorsal view; 6, 11, lateral view; 9, caudal view; 10, 12, 13, ventral view. Scale bars = 0.25 mm (A: Figs 5 8, 10 13; B: Fig. 9)." httpUri="https://zenodo.org/record/919610/files/figure.png" pageId="31" pageNumber="332">
<emphasis id="B9480707A3727F62FE29FEDE8AADD8B8" box="[473,497,368,392]" italics="true" pageId="31" pageNumber="332">11</emphasis>
</figureCitation>
). Up to today only known from the mountains surrounding the Kathmandu Valley (Shivapuri Lekh, Mt. Phulchoki), at altitudes of
<quantity id="4CC476F0A3727F62FD40FE3A8812D89C" box="[688,846,404,429]" metricMagnitude="3" metricUnit="m" metricValue="2.025" metricValueMax="2.3" metricValueMin="1.75" pageId="31" pageNumber="332" unit="m" value="2025.0" valueMax="2300.0" valueMin="1750.0">17502300 m</quantity>
. This species seems to be a geographical vicariant of
<taxonomicName id="4C3CA096A3727F62FF67FE178A37D8E0" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[151,363,439,464]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="332" phylum="Arthropoda" rank="species" species="ganeshensis" status="sp. nov." subGenus="Bottchrus">
<emphasis id="B9480707A3727F62FF67FE178BF0D8E0" box="[151,172,441,464]" italics="true" pageId="31" pageNumber="332">T.</emphasis>
(
<emphasis id="B9480707A3727F62FF4CFE178B91D8E0" box="[188,205,441,464]" italics="true" pageId="31" pageNumber="332">B</emphasis>
.)
<emphasis id="B9480707A3727F62FF14FE198A37D8E0" box="[228,363,439,464]" italics="true" pageId="31" pageNumber="332">ganeshensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27BBA7CA3727F62FE82FE198AEED8E0" box="[370,434,439,464]" pageId="31" pageNumber="332" rank="species">
<emphasis id="B9480707A3727F62FE82FE198AEED8E0" bold="true" box="[370,434,439,464]" pageId="31" pageNumber="332">sp. n.</emphasis>
</taxonomicNameLabel>
which is occurs on the south-western slope of the Ganesh Himal (
<figureCitation id="1307C790A3727F62FB68FE198FA4D8E0" box="[1176,1272,439,464]" captionStart="FIGURE 151" captionStartId="53.[151,250,1080,1102]" captionTargetBox="[151,1436,339,1059]" captionTargetId="figure@53.[151,1436,339,1059]" captionTargetPageId="53" captionText="FIGURE 151. Distribution of the Himalayan species of Trichotichnus subgenus Bottchrus. 1, T. holzschuhi Kirschenhofer, 1992; 2, T. baglungensis sp. n.; 3, T. parvulus sp. n.; 4, T. bubsaensis sp. n.; 5, T. schawalleri sp. n.; 6, T. obliquebasalis sp. n.; 7, T. panchhaseensis sp. n.; 8, T. siklesensis sp. n.; 9, T. pusillus sp. n.; 10, T. ganeshensis sp. n.; 11, T. minutus sp. n.; 12, T. cyanescens Ito, 1998; 13, T. martensi sp. n.; 14, T. opacus Ito, 1998." httpUri="https://zenodo.org/record/919694/files/figure.png" pageId="31" pageNumber="332">Fig. 151</figureCitation>
,
<figureCitation id="1307C790A3727F62FAF4FE168E7CD8E0" box="[1284,1312,440,464]" captionStart="FIGURES 5 13" captionStartId="6.[151,264,1196,1219]" captionTargetBox="[152,1425,180,1152]" captionTargetId="figure@6.[151,1436,180,1193]" captionTargetPageId="6" captionText="FIGURES 5 13. Trichotichnus (Bottchrus) holzschuhi Kirschenhofer, 1992 (5 7, paratype; 8 13, specimens from Sete). 5, terminal lamella of aedeagus; 6 8, 11, median lobe of aedeagus; 9, apical capitulum; 10, left metepisternum; 12, apex of last (VII) abdominal sternite, male; 13, same, female. 5, 7, 8, dorsal view; 6, 11, lateral view; 9, caudal view; 10, 12, 13, ventral view. Scale bars = 0.25 mm (A: Figs 5 8, 10 13; B: Fig. 9)." httpUri="https://zenodo.org/record/919610/files/figure.png" pageId="31" pageNumber="332">
<emphasis id="B9480707A3727F62FAF4FE168E7CD8E0" box="[1284,1312,440,464]" italics="true" pageId="31" pageNumber="332">10</emphasis>
</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>