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<document id="80AFDEC5FE8E698F5521B4FEBEA08EBD" ID-CLB-Dataset="305799" ID-DOI="10.11646/zootaxa.5539.1.1" ID-GBIF-Dataset="fa176ed2-307a-4b37-a666-f1d25247f53b" ID-ISSN="1175-5334" ID-Zenodo-Dep="14240426" ID-ZooBank="7BDB9666-CA90-40BF-8F65-B9897CDA1F2B" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="valdenar" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1732831664062" checkinUser="plazi" docAuthor="Runyon, Justin B." docDate="2024" docId="FD1A87BFA51DFFD650D3FB79FEB7FB16" docLanguage="en" docName="zootaxa.5539.1.1.pdf" docOrigin="Zootaxa 5539 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5539.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Calyxochaetus binodatus" docType="treatment" docVersion="11" lastPageNumber="52" masterDocId="0123FFC7A52CFFE55044FFD3FFFBFFD3" masterDocTitle="Revision of the genus Calyxochaetus Bigot (Diptera: Dolichopodidae: Sympycninae)" masterLastPageNumber="74" masterPageNumber="1" pageNumber="50" updateTime="1738760000286" updateUser="valdenar">
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<mods:title id="40A756B1420EF661220E308F76716126">Revision of the genus Calyxochaetus Bigot (Diptera: Dolichopodidae: Sympycninae)</mods:title>
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<mods:namePart id="26E80B30B395AA021DE9372CFB191C94">Runyon, Justin B.</mods:namePart>
<mods:nameIdentifier id="A4AD141CF815663ACB487B20C548DAB9" type="ORCID">0000-0002-0271-0511</mods:nameIdentifier>
<mods:affiliation id="E4EBFF4F2C22A1AB5B977F85A8437187">Rocky Mountain Research Station, USDA Forest Service, 1648 S. 7 th Avenue, Bozeman, Montana 59717, USA</mods:affiliation>
<mods:nameIdentifier id="BD0F6E2F5869CE4F8685E5732D6CD52A" type="email">justin.runyon@usda.gov</mods:nameIdentifier>
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<mods:date id="911439988980DBF4D347FA68E13BED80">2024</mods:date>
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<mods:identifier id="55BBC3B73DDE380B0EA8C5CE23D56EDA" type="DOI">10.11646/zootaxa.5539.1.1</mods:identifier>
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<treatment id="FD1A87BFA51DFFD650D3FB79FEB7FB16" ID-DOI="http://doi.org/10.5281/zenodo.14278193" ID-GBIF-Taxon="248203384" ID-Zenodo-Dep="14278193" LSID="urn:lsid:plazi:treatment:FD1A87BFA51DFFD650D3FB79FEB7FB16" httpUri="http://treatment.plazi.org/id/FD1A87BFA51DFFD650D3FB79FEB7FB16" lastPageId="51" lastPageNumber="52" pageId="49" pageNumber="50" scope_family="Dolichopodidae" scope_order="Diptera" scope_subFamily="Sympycninae">
<subSubSection id="3DA96522A51DFFD450D3FB79FCFFFB17" box="[151,772,1194,1221]" pageId="49" pageNumber="50" type="nomenclature">
<paragraph id="750C36A9A51DFFD450D3FB79FCFFFB17" blockId="49.[151,772,1194,1257]" box="[151,772,1194,1221]" pageId="49" pageNumber="50">
<heading id="2E4481C5A51DFFD450D3FB79FCFFFB17" bold="true" box="[151,772,1194,1221]" fontSize="11" level="2" pageId="49" pageNumber="50" reason="4">
<taxonomicName id="B2B34D2AA51DFFD450D3FB79FCFFFB17" authority="(Harmston &amp; Knowlton)" baseAuthorityName="Harmston &amp; Knowlton" baseAuthorityYear="1940" box="[151,772,1194,1221]" class="Insecta" family="Dolichopodidae" genus="Calyxochaetus" kingdom="Animalia" order="Diptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="binodatus">
<emphasis id="47C7EABBA51DFFD450D3FB79FCFFFB17" bold="true" box="[151,772,1194,1221]" pageId="49" pageNumber="50">
<emphasis id="47C7EABBA51DFFD450D3FB79FE3CFB16" bold="true" box="[151,455,1194,1221]" italics="true" pageId="49" pageNumber="50">Calyxochaetus binodatus</emphasis>
(Harmston &amp; Knowlton)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3DA96522A51DFFD450D3FB1CFE11FB3A" box="[151,490,1231,1257]" pageId="49" pageNumber="50" type="description">
<paragraph id="750C36A9A51DFFD450D3FB1CFE11FB3A" blockId="49.[151,772,1194,1257]" box="[151,490,1231,1257]" pageId="49" pageNumber="50">
(
<figureCitation id="ED882A2CA51DFFD450DBFB1CFEC7FB3A" box="[159,316,1231,1257]" captionStart="FIGURES 104–119" captionStartId="50.[151,264,1472,1496]" captionTargetBox="[151,1437,181,1448]" captionTargetId="figure-27@50.[151,1437,181,1448]" captionTargetPageId="50" captionText="FIGURES 104–119. Male antennae (lateral view) and forelegs of the Calyxochaetus frontalis species group. 104. C. binodatus (Harmston &amp; Knowlton). 105. C. nodatus (Loew). 106. C. desertus sp. nov. 107. C. pictipes (Harmston &amp; Knowlton). 108. C. sobrinus (Wheeler). 109. C. ornatus Parent. 110. C. remifer sp. nov. 111. C. tohono sp. nov. 112. C. frontalis (Loew). 113. C. furcatus (Van Duzee). 114. C. lamellicornis Parent (photo by Emmanuel Delfosse, MNHN). 115. C. lacrima sp. nov. 116. C. binodatus, fore tarsus, anterior view. 117. C. sobrinus, fore tarsus, anterior view. 118. C. remifer sp. nov., fore tarsus, dorsal view. 119. C. lacrima sp. nov., fore tibia and tarsus, posterior view. Scale bars = 0.5 mm; arista-like styli approximately 1 mm long." figureDoi="http://doi.org/10.5281/zenodo.14240476" httpUri="https://zenodo.org/record/14240476/files/figure.png" pageId="49" pageNumber="50">Figs 104, 116</figureCitation>
,
<figureCitation id="ED882A2CA51DFFD45103FB1CFE88FB3A" box="[327,371,1231,1257]" captionStart="FIGURES 120–131" captionStartId="52.[151,264,1711,1735]" captionTargetBox="[196,1392,181,1688]" captionTargetId="figure-27@52.[196,1392,181,1688]" captionTargetPageId="52" captionText="FIGURES 120–131. Male mid tarsi of the Calyxochaetus frontalis species group (anterior on left side). 120. C. binodatus (Harmston &amp; Knowlton), dorsal view. 121. C. nodatus (Loew), ventral view. 122. C. bovanodatus sp. nov., dorsal view. 123. C. desertus sp. nov., mid tarsomeres 2–5, dorsal view. 124. C. sobrinus (Wheeler), mid tarsomeres 2–5, dorsal view. 125. C. pictipes (Harmston &amp; Knowlton), dorsal view. 126. C. ornatus Parent, mid tarsomeres 2–5, ventral view. 127. C. tohono sp. nov., mid tarsomeres 2–5, dorsal view. 128. C. remifer sp. nov., dorsal view. 129. C. frontalis (Loew), dorsal view. 130. C. furcatus (Van Duzee), dorsal view. 131. C. lacrima sp. nov., dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14240480" httpUri="https://zenodo.org/record/14240480/files/figure.png" pageId="49" pageNumber="50">120</figureCitation>
,
<figureCitation id="ED882A2CA51DFFD4513BFB1CFE50FB3A" box="[383,427,1231,1257]" captionStart="FIGURES 132–138" captionStartId="54.[151,264,1423,1447]" captionTargetBox="[152,1436,381,1400]" captionTargetId="figure-114@54.[152,1436,381,1400]" captionTargetPageId="54" captionText="FIGURES 132–138. Male hind legs and wings of the Calyxochaetus frontalis species group. 132. C. binodatus (Harmston &amp; Knowlton), hind tibia, anterior view. 133. C. nodatus (Loew), hind tibia, anterior view. 134. C. ornatus Parent, hind tibia, posterior view. 135. C. frontalis (Loew), hind tibia, posterior view. 136. C. lacrima sp. nov., apical third of hind femur and basal half of hind tibia, posterior view. 137. C. nodatus, wing. 138. C. ornatus, wing. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14240484" httpUri="https://zenodo.org/record/14240484/files/figure.png" pageId="49" pageNumber="50">132</figureCitation>
,
<figureCitation id="ED882A2CA51DFFD451F3FB1CFE19FB3A" box="[439,482,1231,1257]" captionStart="FIGURES 144–146" captionStartId="64.[151,264,1684,1708]" captionTargetBox="[151,1436,181,1660]" captionTargetId="figure-27@64.[151,1436,181,1660]" captionTargetPageId="64" captionText="FIGURES 144–146. Known geographical distributions of species in the Calyxochaetus frontalis group. 144. C. nodatus (Loew), C. binodatus (Harmston &amp; Knowlton) and C. bovanodatus sp. nov. 145. C. sobrinus (Wheeler) and C. pictipes (Harmston &amp; Knowlton). 146. C. frontalis (Loew) and C. desertus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.14240490" httpUri="https://zenodo.org/record/14240490/files/figure.png" pageId="49" pageNumber="50">144</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="3DA96522A51DFFD450D3FAC0FD6BFA99" pageId="49" pageNumber="50" type="reference_group">
<paragraph id="750C36A9A51DFFD450D3FAC0FD27FAF8" blockId="49.[151,732,1299,1354]" box="[151,732,1299,1323]" pageId="49" pageNumber="50">
<treatmentCitationGroup id="55A31187A51DFFD450D3FAC0FD27FAF8" box="[151,732,1299,1323]" pageId="49" pageNumber="50">
<treatmentCitation id="F41210B8A51DFFD450D3FAC0FD23FAF8" author="Harmston, F. C. &amp; Knowlton, G. F." box="[151,728,1299,1323]" page="397" pageId="49" pageNumber="50" year="1940">
<taxonomicName id="B2B34D2AA51DFFD450D3FAC0FD23FAF8" ID-CoL="53SF8" authority="Harmston &amp; Knowlton, 1940: 397" authorityName="Harmston &amp; Knowlton" authorityPageNumber="397" authorityYear="1940" box="[151,728,1299,1323]" class="Insecta" family="Dolichopodidae" genus="Sympycnus" kingdom="Animalia" order="Diptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="binodatus">
<emphasis id="47C7EABBA51DFFD450D3FAC0FE89FAF8" box="[151,370,1299,1323]" italics="true" pageId="49" pageNumber="50">Sympycnus binodatus</emphasis>
<bibRefCitation id="11224B58A51DFFD4513CFAC7FD23FAF8" author="Harmston, F. C. &amp; Knowlton, G. F." box="[376,728,1299,1323]" pageId="49" pageNumber="50" pagination="395 - 403" refId="ref72695" refString="Harmston, F. C. &amp; Knowlton, G. F. (1940) The genus Sympycnus in Utah (Dolichopodidae: Diptera). Annals of the Entomological Society of America, 33, 395 - 403. https: // doi. org / 10.1093 / aesa / 33.2.395" type="journal article" year="1940">Harmston &amp; Knowlton, 1940: 397</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="750C36A9A51DFFD450D3FAE1FD6BFA99" blockId="49.[151,732,1299,1354]" box="[151,656,1330,1354]" pageId="49" pageNumber="50">
<treatmentCitationGroup id="55A31187A51DFFD450D3FAE1FD6BFA99" box="[151,656,1330,1354]" pageId="49" pageNumber="50">
<taxonomicName id="B2B34D2AA51DFFD450D3FAE1FE6CFA99" baseAuthorityName="Harmston &amp; Knowlton" baseAuthorityYear="1940" box="[151,407,1330,1354]" class="Insecta" family="Dolichopodidae" genus="Calyxochaetus" kingdom="Animalia" order="Diptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="binodatus">
<emphasis id="47C7EABBA51DFFD450D3FAE1FE6CFA99" box="[151,407,1330,1354]" italics="true" pageId="49" pageNumber="50">Calyxochaetus binodatus</emphasis>
</taxonomicName>
:
<treatmentCitation id="F41210B8A51DFFD451E0FAE1FD77FA99" author="Pollet, M. A. A. &amp; Brooks, S. E. &amp; Cumming, J. M." box="[420,652,1330,1354]" page="74" pageId="49" pageNumber="50" year="2004">
<bibRefCitation id="11224B58A51DFFD451E0FAE1FD77FA99" author="Pollet, M. A. A. &amp; Brooks, S. E. &amp; Cumming, J. M." box="[420,652,1330,1354]" pageId="49" pageNumber="50" pagination="1 - 114" refId="ref73053" refString="Pollet, M. A. A., Brooks, S. E. &amp; Cumming, J. M. (2004) Catalog of the Dolichopodidae (Diptera) of America North of Mexico. Bulletin of the American Museum of Natural History, 283, 1 - 114. https: // doi. org / 10.1206 / 0003 - 0090 (2004) 283 % 3 C 0001: COTDDO % 3 E 2.0. CO; 2" type="journal article" year="2004">
Pollet
<emphasis id="47C7EABBA51DFFD451A1FAE0FDE9FA99" box="[485,530,1330,1354]" italics="true" pageId="49" pageNumber="50">et al</emphasis>
. (2004: 74)
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="3DA96522A51DFFD450D3FAABFA79F929" pageId="49" pageNumber="50" type="materials_examined">
<paragraph id="750C36A9A51DFFD450D3FAABFDD4FA09" blockId="49.[151,1437,1400,1859]" pageId="49" pageNumber="50">
<emphasis id="47C7EABBA51DFFD450D3FAABFE3FFA41" bold="true" box="[151,452,1400,1426]" pageId="49" pageNumber="50">Type material examined.</emphasis>
<materialsCitation id="C5DB3CF4A51DFFD45189FAABFDD4FA09" ID-GBIF-Occurrence="4987889399" collectingDate="1938-09-13" collectionCode="USNM" collectorName="G. F. Knowlton &amp; F. C. Harmston" country="United States of America" location="Leeds" pageId="49" pageNumber="50" specimenCount="1" specimenCount-male="1" stateProvince="Utah" typeStatus="holotype">
<emphasis id="47C7EABBA51DFFD45189FAABFD97FA41" bold="true" box="[461,620,1400,1426]" pageId="49" pageNumber="50">
<typeStatus id="AA08880BA51DFFD45189FAABFD97FA41" box="[461,620,1400,1426]" pageId="49" pageNumber="50" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
<specimenCount id="63B5FD20A51DFFD45231FAAAFD77FA40" box="[629,652,1401,1427]" count="1" pageId="49" pageNumber="50" type="male">♂</specimenCount>
, labeled: “
<collectorName id="D846537FA51DFFD45343FAABFC4BFA41" box="[775,944,1400,1426]" pageId="49" pageNumber="50">G.F. Knowlton</collectorName>
/
<collectorName id="D846537FA51DFFD45386FAABFB9CFA41" box="[962,1127,1400,1426]" pageId="49" pageNumber="50">F.C. Harmston</collectorName>
/
<location id="706C6072A51DFFD4543DFAABFB44FA41" LSID="urn:lsid:plazi:treatment:FD1A87BFA51DFFD650D3FB79FEB7FB16:706C6072A51DFFD4543DFAABFB44FA41" box="[1145,1215,1400,1426]" country="United States of America" name="Leeds" pageId="49" pageNumber="50" stateProvince="Utah">Leeds</location>
,
<collectingRegion id="B777F84BA51DFFD45488FAABFAF9FA41" box="[1228,1282,1400,1426]" country="United States of America" name="Utah" pageId="49" pageNumber="50">Utah</collectingRegion>
/
<date id="010D1069A51DFFD45557FAABFA77FA41" box="[1299,1420,1400,1426]" pageId="49" pageNumber="50" value="1938-09-13">
<collectingDate id="1149E981A51DFFD45557FAABFA77FA41" box="[1299,1420,1400,1426]" pageId="49" pageNumber="50" value="1938-09-13">9-13-1938</collectingDate>
</date>
”; “Type No/ 54297/ U.S.N.M.”; “Holo Type/
<taxonomicName id="B2B34D2AA51DFFD452C2FA4FFCF9FA65" baseAuthorityName="Sinclair &amp; Cumming" baseAuthorityYear="2006" box="[646,770,1436,1462]" class="Insecta" family="Dolichopodidae" genus="Sympycnus" kingdom="Animalia" order="Diptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="47C7EABBA51DFFD452C2FA4FFCF9FA65" box="[646,770,1436,1462]" italics="true" pageId="49" pageNumber="50">Sympycnus</emphasis>
</taxonomicName>
/
<taxonomicName id="B2B34D2AA51DFFD45356FA4FFC7BFA65" baseAuthorityName="Harmston &amp; Knowlton" baseAuthorityYear="1940" box="[786,896,1436,1462]" class="Insecta" family="Dolichopodidae" genus="Calyxochaetus" kingdom="Animalia" order="Diptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="binodatus">
<emphasis id="47C7EABBA51DFFD45356FA4FFC7BFA65" box="[786,896,1436,1462]" italics="true" pageId="49" pageNumber="50">binodatus</emphasis>
</taxonomicName>
/ n. sp. F.C.H.-G.F.K.”; “
<taxonomicName id="B2B34D2AA51DFFD454DEFA4FFAEDFA65" baseAuthorityName="Sinclair &amp; Cumming" baseAuthorityYear="2006" box="[1178,1302,1436,1462]" class="Insecta" family="Dolichopodidae" genus="Sympycnus" kingdom="Animalia" order="Diptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="47C7EABBA51DFFD454DEFA4FFAEDFA65" box="[1178,1302,1436,1462]" italics="true" pageId="49" pageNumber="50">Sympycnus</emphasis>
</taxonomicName>
/
<taxonomicName id="B2B34D2AA51DFFD45562FA4FFA6FFA65" baseAuthorityName="Harmston &amp; Knowlton" baseAuthorityYear="1940" box="[1318,1428,1436,1462]" class="Insecta" family="Dolichopodidae" genus="Calyxochaetus" kingdom="Animalia" order="Diptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="binodatus">
<emphasis id="47C7EABBA51DFFD45562FA4FFA6FFA65" box="[1318,1428,1436,1462]" italics="true" pageId="49" pageNumber="50">binodatus</emphasis>
</taxonomicName>
/ Det. 1940 F.C. Harmston” (
<collectionCode id="13A2AE6CA51DFFD4518FFA13FDD9FA09" box="[459,546,1472,1498]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" pageId="49" pageNumber="50">USNM</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="750C36A9A51DFFD45083FA37FA79F929" blockId="49.[151,1437,1400,1859]" pageId="49" pageNumber="50">
<emphasis id="47C7EABBA51DFFD45083FA37FE04FA2D" bold="true" box="[199,511,1508,1534]" pageId="49" pageNumber="50">Other material examined.</emphasis>
<materialsCitation id="C5DB3CF4A51DFFD45243FA37FDBCF9F1" ID-GBIF-Occurrence="4987889595" collectingDate="2002-04-23" collectingDateMax="2003-04-15" collectingDateMin="2002-04-23" collectionCode="MTEC" collectorName="RLH &amp; JBR" country="United States of America" county="Cochise County" elevation="1227" location="San Pedro River" municipality="San Pedro River" pageId="49" pageNumber="50" specimenCount="4" specimenCount-female="4" specimenCount-male="3" stateProvince="Arizona">
<emphasis id="47C7EABBA51DFFD45243FA37FD41FA2D" bold="true" box="[519,698,1508,1534]" pageId="49" pageNumber="50">
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:
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:
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:
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E Sierra Vista
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<collectingDate id="1149E981A51DFFD450D3F9DBFEEBF9F1" box="[151,272,1544,1570]" pageId="49" pageNumber="50" value="2002-04-23">23.iv.2002</collectingDate>
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&amp;
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(
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<materialsCitation id="C5DB3CF4A51DFFD4520AF9DBFDC5F995" ID-GBIF-Occurrence="4987889372" collectingDate="2003-04-13" collectingDateMax="2003-04-15" collectingDateMin="2002-04-23" collectionCode="CNC, MTEC" collectorName="JBR &amp; RLH" country="United States of America" county="Cochise County" elevation="594" location="Mohave County" municipality="San Pedro River" pageId="49" pageNumber="50" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Arizona">
<location id="706C6072A51DFFD4520AF9DBFCFCF9F1" LSID="urn:lsid:plazi:treatment:FD1A87BFA51DFFD650D3FB79FEB7FB16:706C6072A51DFFD4520AF9DBFCFCF9F1" box="[590,775,1544,1570]" country="United States of America" county="Cochise County" municipality="San Pedro River" name="Mohave County" pageId="49" pageNumber="50" stateProvince="Arizona">Mohave County</location>
:
<location id="706C6072A51DFFD45355F9DAFC1FF9F1" LSID="urn:lsid:plazi:treatment:FD1A87BFA51DFFD650D3FB79FEB7FB16:706C6072A51DFFD45355F9DAFC1FF9F1" box="[785,996,1544,1571]" country="United States of America" county="Cochise County" municipality="San Pedro River" name="Beaver Dam Wash" pageId="49" pageNumber="50" stateProvince="Arizona">Beaver Dam Wash</location>
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<location id="706C6072A51DFFD453B4F9DAFB84F9F0" LSID="urn:lsid:plazi:treatment:FD1A87BFA51DFFD650D3FB79FEB7FB16:706C6072A51DFFD453B4F9DAFB84F9F0" box="[1008,1151,1545,1571]" country="United States of America" county="Cochise County" municipality="San Pedro River" name="Beaver Dam" pageId="49" pageNumber="50" stateProvince="Arizona">Beaver Dam</location>
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<date id="010D1069A51DFFD454ACF9DBFA9BF9F1" box="[1256,1376,1544,1570]" pageId="49" pageNumber="50" value="2003-04-13">
<collectingDate id="1149E981A51DFFD454ACF9DBFA9BF9F1" box="[1256,1376,1544,1570]" pageId="49" pageNumber="50" value="2003-04-13">13.iv.2003</collectingDate>
</date>
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<collectorName id="D846537FA51DFFD45528F9DAFA67F9F0" box="[1388,1436,1545,1571]" pageId="49" pageNumber="50">JBR</collectorName>
&amp;
<collectorName id="D846537FA51DFFD450F1F9FEFF16F994" box="[181,237,1581,1607]" pageId="49" pageNumber="50">RLH</collectorName>
(
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<collectionCode id="13A2AE6CA51DFFD45175F9FFFE95F995" box="[305,366,1580,1606]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" pageId="49" pageNumber="50">CNC</collectionCode>
;
<specimenCount id="63B5FD20A51DFFD4513FF9FFFE64F995" box="[379,415,1580,1606]" count="1" pageId="49" pageNumber="50" type="male">1♂</specimenCount>
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<specimenCount id="63B5FD20A51DFFD451E9F9FFFE2AF995" box="[429,465,1580,1606]" count="2" pageId="49" pageNumber="50" type="female">2♀</specimenCount>
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<collectionCode id="13A2AE6CA51DFFD4519BF9FFFDC9F995" box="[479,562,1580,1606]" collectionName="USA, Montana, Bozeman, Montana State University" pageId="49" pageNumber="50">MTEC</collectionCode>
);
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<materialsCitation id="C5DB3CF4A51DFFD45202F9FFFEDFF9B9" ID-GBIF-Occurrence="4987889505" collectingDate="2003-04-15" collectingDateMax="2003-04-15" collectingDateMin="2002-04-23" collectionCode="MTEC" collectorName="JBR &amp; RLH" country="United States of America" county="Cochise County" elevation="1143" location="Santa Cruz County" municipality="San Pedro River" pageId="49" pageNumber="50" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Arizona">
<location id="706C6072A51DFFD45202F9FFFCD8F995" LSID="urn:lsid:plazi:treatment:FD1A87BFA51DFFD650D3FB79FEB7FB16:706C6072A51DFFD45202F9FFFCD8F995" box="[582,803,1580,1606]" country="United States of America" county="Cochise County" municipality="San Pedro River" name="Santa Cruz County" pageId="49" pageNumber="50" stateProvince="Arizona">Santa Cruz County</location>
:
<location id="706C6072A51DFFD4536AF9FFFC1BF995" LSID="urn:lsid:plazi:treatment:FD1A87BFA51DFFD650D3FB79FEB7FB16:706C6072A51DFFD4536AF9FFFC1BF995" box="[814,992,1580,1606]" country="United States of America" county="Cochise County" municipality="San Pedro River" name="Patagonia Lake" pageId="49" pageNumber="50" stateProvince="Arizona">Patagonia Lake</location>
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<quantity id="B24B9B4CA51DFFD453A8F9FFFBBBF995" box="[1004,1088,1580,1607]" metricMagnitude="3" metricUnit="m" metricValue="1.143" pageId="49" pageNumber="50" unit="ft" value="3750.0">
<elevation id="FE9ED19AA51DFFD453A8F9FFFBBBF995" box="[1004,1088,1580,1607]" metricMagnitude="3" metricUnit="m" metricValue="1.143" pageId="49" pageNumber="50" unit="ft" value="3750.0">3750 ft</elevation>
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<date id="010D1069A51DFFD45409F9FFFB3EF994" box="[1101,1221,1580,1607]" pageId="49" pageNumber="50" value="2003-04-15">
<collectingDate id="1149E981A51DFFD45409F9FFFB3EF994" box="[1101,1221,1580,1607]" pageId="49" pageNumber="50" value="2003-04-15">15.iv.2003</collectingDate>
</date>
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<collectorName id="D846537FA51DFFD45496F9FEFAF9F994" box="[1234,1282,1581,1607]" pageId="49" pageNumber="50">JBR</collectorName>
&amp;
<collectorName id="D846537FA51DFFD4556DF9FEFA9AF994" box="[1321,1377,1581,1607]" pageId="49" pageNumber="50">RLH</collectorName>
(
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<specimenCount id="63B5FD20A51DFFD450D3F983FF41F9B9" box="[151,186,1616,1642]" count="1" pageId="49" pageNumber="50" type="female">1♀</specimenCount>
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<collectionCode id="13A2AE6CA51DFFD4508EF983FEE6F9B9" box="[202,285,1616,1642]" collectionName="USA, Montana, Bozeman, Montana State University" pageId="49" pageNumber="50">MTEC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="C5DB3CF4A51DFFD45174F983FAE1F9B9" ID-GBIF-Occurrence="4987889553" box="[304,1306,1616,1643]" collectingDate="1972-06-01" collectingMethod="Malaise" collectionCode="USNM" collectorName="W. W. Wirth" country="United States of America" county="Catron County" location="Glenwood" pageId="49" pageNumber="50" specimenCount="2" specimenCount-male="2" stateProvince="New Mexico">
<emphasis id="47C7EABBA51DFFD45174F983FE35F9B9" bold="true" box="[304,462,1616,1642]" pageId="49" pageNumber="50">
<collectingRegion id="B777F84BA51DFFD45174F983FE33F9B9" box="[304,456,1616,1642]" country="United States of America" name="New Mexico" pageId="49" pageNumber="50">New Mexico</collectingRegion>
:
</emphasis>
<collectingCounty id="9C6D4E25A51DFFD45192F983FD79F9B9" box="[470,642,1616,1642]" pageId="49" pageNumber="50">Catron County</collectingCounty>
:
<location id="706C6072A51DFFD452C9F983FCFEF9B9" LSID="urn:lsid:plazi:treatment:FD1A87BFA51DFFD650D3FB79FEB7FB16:706C6072A51DFFD452C9F983FCFEF9B9" box="[653,773,1616,1642]" country="United States of America" county="Catron County" name="Glenwood" pageId="49" pageNumber="50" stateProvince="New Mexico">Glenwood</location>
,
<date id="010D1069A51DFFD45355F983FC86F9B9" box="[785,893,1616,1642]" pageId="49" pageNumber="50" value="1972-06-01">
<collectingDate id="1149E981A51DFFD45355F983FC86F9B9" box="[785,893,1616,1642]" pageId="49" pageNumber="50" value="1972-06-01">1.vi.1972</collectingDate>
</date>
,
<collectingMethod id="ACF24EBEA51DFFD453CEF983FC1EF9B9" box="[906,997,1616,1642]" pageId="49" pageNumber="50">Malaise</collectingMethod>
,
<collectorName id="D846537FA51DFFD453B6F982FB83F9B9" box="[1010,1144,1616,1643]" pageId="49" pageNumber="50">W.W. Wirth</collectorName>
(
<specimenCount id="63B5FD20A51DFFD454CDF983FB54F9B9" box="[1161,1199,1616,1642]" count="2" pageId="49" pageNumber="50" type="male">2♂</specimenCount>
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<collectionCode id="13A2AE6CA51DFFD454F8F983FAE9F9B9" box="[1212,1298,1616,1642]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" pageId="49" pageNumber="50">USNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="C5DB3CF4A51DFFD45561F983FCD2F95D" ID-GBIF-Occurrence="4987889603" collectingDate="1950-08-30" collectingDateMax="1987-09-10" collectingDateMin="1940-07-27" collectionCode="FSCA" collectorName="FCH" country="United States of America" county="San Juan County" location="Monticello" municipality="Monticello" pageId="49" pageNumber="50" specimenCount="1" specimenCount-male="1" stateProvince="Utah">
<emphasis id="47C7EABBA51DFFD45561F983FA91F9B9" bold="true" box="[1317,1386,1616,1642]" pageId="49" pageNumber="50">
<collectingRegion id="B777F84BA51DFFD45561F983FA9FF9B9" box="[1317,1380,1616,1642]" country="United States of America" name="Utah" pageId="49" pageNumber="50">Utah</collectingRegion>
:
</emphasis>
<collectingCounty id="9C6D4E25A51DFFD45536F983FED1F95D" pageId="49" pageNumber="50">San Juan County</collectingCounty>
:
<collectingMunicipality id="9568ACD3A51DFFD45172F9A7FE4FF95D" box="[310,436,1652,1678]" pageId="49" pageNumber="50">Monticello</collectingMunicipality>
,
<date id="010D1069A51DFFD45184F9A7FDB1F95C" box="[448,586,1652,1679]" pageId="49" pageNumber="50" value="1950-08-30">
<collectingDate id="1149E981A51DFFD45184F9A7FDB1F95C" box="[448,586,1652,1679]" pageId="49" pageNumber="50" value="1950-08-30">30.viii.1950</collectingDate>
</date>
,
<collectorName id="D846537FA51DFFD45213F9A7FD76F95D" box="[599,653,1652,1678]" pageId="49" pageNumber="50">FCH</collectorName>
(
<specimenCount id="63B5FD20A51DFFD452E5F9A7FD3EF95D" box="[673,709,1652,1678]" count="1" pageId="49" pageNumber="50" type="male">1♂</specimenCount>
,
<collectionCode id="13A2AE6CA51DFFD45296F9A7FCE6F95D" box="[722,797,1652,1678]" collectionName="USA, Florida, Gainesville, Division of Plant Industry, Florida State Collection of Arthropods" pageId="49" pageNumber="50">FSCA</collectionCode>
);
</materialsCitation>
<materialsCitation id="C5DB3CF4A51DFFD45376F9A7FD35F961" ID-GBIF-Occurrence="4987889350" collectingDate="1965-07-27" collectingDateMax="1987-09-10" collectingDateMin="1940-07-27" collectingMethod="Malaise" collectionCode="LACM" collectorName="W. J. Hanson" country="United States of America" county="San Juan County" location="Zion National Park" municipality="Monticello" pageId="49" pageNumber="50" specimenCount="1" specimenCount-male="1" stateProvince="Washington County">
<collectingRegion id="B777F84BA51DFFD45376F9A7FBEDF95D" box="[818,1046,1652,1678]" pageId="49" pageNumber="50">Washington County</collectingRegion>
:
<location id="706C6072A51DFFD45466F9A7FB04F95D" LSID="urn:lsid:plazi:treatment:FD1A87BFA51DFFD650D3FB79FEB7FB16:706C6072A51DFFD45466F9A7FB04F95D" box="[1058,1279,1652,1678]" country="United States of America" county="San Juan County" municipality="Monticello" name="Zion National Park" pageId="49" pageNumber="50" stateProvince="Washington County">Zion National Park</location>
,
<location id="706C6072A51DFFD45548F9A7FA63F95D" LSID="urn:lsid:plazi:treatment:FD1A87BFA51DFFD650D3FB79FEB7FB16:706C6072A51DFFD45548F9A7FA63F95D" box="[1292,1432,1652,1678]" country="United States of America" county="San Juan County" municipality="Monticello" name="Birch Creek" pageId="49" pageNumber="50" stateProvince="Washington County">Birch Creek</location>
,
<date id="010D1069A51DFFD450D3F94BFEE2F960" box="[151,281,1688,1715]" pageId="49" pageNumber="50" value="1965-07-27">
<collectingDate id="1149E981A51DFFD450D3F94BFEE2F960" box="[151,281,1688,1715]" pageId="49" pageNumber="50" value="1965-07-27">27.vii.1965</collectingDate>
</date>
,
<collectingMethod id="ACF24EBEA51DFFD4516DF94BFE7FF961" box="[297,388,1688,1714]" pageId="49" pageNumber="50">Malaise</collectingMethod>
,
<collectorName id="D846537FA51DFFD451D7F94AFDDFF960" box="[403,548,1689,1715]" pageId="49" pageNumber="50">W.J. Hanson</collectorName>
(
<specimenCount id="63B5FD20A51DFFD4527DF94BFDA6F961" box="[569,605,1688,1714]" count="1" pageId="49" pageNumber="50" type="male">1♂</specimenCount>
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<collectionCode id="13A2AE6CA51DFFD45229F94BFD38F961" box="[621,707,1688,1714]" collectionName="USA, California, Los Angeles, Los Angeles County Museum of Natural History" pageId="49" pageNumber="50">LACM</collectionCode>
);
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<materialsCitation id="C5DB3CF4A51DFFD4529DF94AFE6FF905" ID-GBIF-Occurrence="4987889376" collectingDate="1987-07-01" collectingDateMax="1987-07-02" collectingDateMin="1987-07-01" collectionCode="LACM" collectorName="W. J. Hanson" country="United States of America" county="San Juan County" location="Beaver Dam Wash" municipality="Monticello" pageId="49" pageNumber="50" specimenCount="1" specimenCount-male="1" stateProvince="Utah">
<location id="706C6072A51DFFD4529DF94AFC4FF961" LSID="urn:lsid:plazi:treatment:FD1A87BFA51DFFD650D3FB79FEB7FB16:706C6072A51DFFD4529DF94AFC4FF961" box="[729,948,1688,1715]" country="United States of America" county="San Juan County" municipality="Monticello" name="Beaver Dam Wash" pageId="49" pageNumber="50" stateProvince="Utah">Beaver Dam Wash</location>
,
<location id="706C6072A51DFFD45380F94BFB46F960" LSID="urn:lsid:plazi:treatment:FD1A87BFA51DFFD650D3FB79FEB7FB16:706C6072A51DFFD45380F94BFB46F960" box="[964,1213,1688,1715]" country="United States of America" county="San Juan County" municipality="Monticello" name="Lytle Ranch Preserve" pageId="49" pageNumber="50" stateProvince="Utah">Lytle Ranch Preserve</location>
,
<date id="010D1069A51DFFD45489F94BFAA6F961" box="[1229,1373,1688,1714]" pageId="49" pageNumber="50" value="1987-07-01" valueMax="1987-07-02" valueMin="1987-07-01">
<collectingDate id="1149E981A51DFFD45489F94BFAA6F961" box="[1229,1373,1688,1714]" pageId="49" pageNumber="50" value="1987-07-01" valueMax="1987-07-02" valueMin="1987-07-01">1–2.vii.1987</collectingDate>
</date>
,
<collectorName id="D846537FA51DFFD4552FF94AFF16F904" pageId="49" pageNumber="50">W.J. Hanson</collectorName>
(
<specimenCount id="63B5FD20A51DFFD45144F96FFEDFF905" box="[256,292,1724,1750]" count="1" pageId="49" pageNumber="50" type="male">1♂</specimenCount>
,
<collectionCode id="13A2AE6CA51DFFD45177F96FFE72F905" box="[307,393,1724,1750]" collectionName="USA, California, Los Angeles, Los Angeles County Museum of Natural History" pageId="49" pageNumber="50">LACM</collectionCode>
);
</materialsCitation>
<materialsCitation id="C5DB3CF4A51DFFD451DAF96EFC13F905" ID-GBIF-Occurrence="4987889427" box="[414,1000,1724,1751]" collectingDate="1987-08-19" collectingDateMax="1987-09-10" collectingDateMin="1987-08-19" collectionCode="LACM" collectorName=" W. J. Hanson " country="United States of America" county="San Juan County" location="Zion National Park" municipality="Monticello" pageId="49" pageNumber="50" specimenCount="1" specimenCount-male="1" stateProvince="Utah">
same data except,
<date id="010D1069A51DFFD45236F96FFCBBF905" box="[626,832,1724,1750]" pageId="49" pageNumber="50" value="1987-08-19" valueMax="1987-09-10" valueMin="1987-08-19">
<collectingDate id="1149E981A51DFFD45236F96FFCBBF905" box="[626,832,1724,1750]" pageId="49" pageNumber="50" value="1987-08-19" valueMax="1987-09-10" valueMin="1987-08-19">19.viii–10.ix.1987</collectingDate>
</date>
(
<specimenCount id="63B5FD20A51DFFD45310F96FFC83F905" box="[852,888,1724,1750]" count="1" pageId="49" pageNumber="50" type="male">1♂</specimenCount>
,
<collectionCode id="13A2AE6CA51DFFD453C2F96FFC27F905" box="[902,988,1724,1750]" collectionName="USA, California, Los Angeles, Los Angeles County Museum of Natural History" pageId="49" pageNumber="50">LACM</collectionCode>
);
</materialsCitation>
<materialsCitation id="C5DB3CF4A51DFFD453B5F96FFE95F929" ID-GBIF-Occurrence="4987889309" collectingDate="1940-07-27" collectingDateMax="1987-09-10" collectingDateMin="1940-07-27" collectionCode="MTEC" collectorName="G. F. Knowlton &amp; FCH" country="United States of America" county="San Juan County" location="Leeds" municipality="Monticello" pageId="49" pageNumber="50" specimenCount="2" specimenCount-male="2" stateProvince="Utah">
<location id="706C6072A51DFFD453B5F96FFBCCF905" LSID="urn:lsid:plazi:treatment:FD1A87BFA51DFFD650D3FB79FEB7FB16:706C6072A51DFFD453B5F96FFBCCF905" box="[1009,1079,1724,1750]" country="United States of America" county="San Juan County" municipality="Monticello" name="Leeds" pageId="49" pageNumber="50" stateProvince="Utah">Leeds</location>
,
<date id="010D1069A51DFFD45400F96FFB3CF905" box="[1092,1223,1724,1750]" pageId="49" pageNumber="50" value="1940-07-27">
<collectingDate id="1149E981A51DFFD45400F96FFB3CF905" box="[1092,1223,1724,1750]" pageId="49" pageNumber="50" value="1940-07-27">27.vii.1940</collectingDate>
</date>
,
<collectorName id="D846537FA51DFFD45490F96FFA86F905" box="[1236,1405,1724,1750]" pageId="49" pageNumber="50">G.F. Knowlton</collectorName>
&amp;
<collectorName id="D846537FA51DFFD450D3F933FF36F929" box="[151,205,1760,1786]" pageId="49" pageNumber="50">FCH</collectorName>
(
<specimenCount id="63B5FD20A51DFFD4509AF933FEF8F929" box="[222,259,1760,1786]" count="2" pageId="49" pageNumber="50" type="male">2♂</specimenCount>
,
<collectionCode id="13A2AE6CA51DFFD4514BF933FE99F929" box="[271,354,1760,1786]" collectionName="USA, Montana, Bozeman, Montana State University" pageId="49" pageNumber="50">MTEC</collectionCode>
);
</materialsCitation>
<materialsCitation id="C5DB3CF4A51DFFD45131F932FCA0F929" ID-GBIF-Occurrence="4987889639" box="[373,859,1760,1787]" collectingDate="1941-04-25" collectingDateMax="1987-09-10" collectingDateMin="1940-07-27" collectionCode="USNM" collectorName=" G. F. Knowlton &amp; FCH" country="United States of America" county="San Juan County" location="Zion National Park" municipality="Monticello" pageId="49" pageNumber="50" specimenCount="1" specimenCount-male="1" stateProvince="Utah">
same data except,
<date id="010D1069A51DFFD45206F933FD43F928" box="[578,696,1760,1787]" pageId="49" pageNumber="50" value="1941-04-25">
<collectingDate id="1149E981A51DFFD45206F933FD43F928" box="[578,696,1760,1787]" pageId="49" pageNumber="50" value="1941-04-25">25.iv.1941</collectingDate>
</date>
(
<specimenCount id="63B5FD20A51DFFD4528DF933FD16F929" box="[713,749,1760,1786]" count="1" pageId="49" pageNumber="50" type="male">1♂</specimenCount>
,
<collectionCode id="13A2AE6CA51DFFD452BDF933FCB4F929" box="[761,847,1760,1786]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" pageId="49" pageNumber="50">USNM</collectionCode>
);
</materialsCitation>
<materialsCitation id="C5DB3CF4A51DFFD45326F932FA85F929" ID-GBIF-Occurrence="4987889456" box="[866,1406,1760,1787]" collectingDate="1951-04-20" collectingDateMax="1987-09-10" collectingDateMin="1940-07-27" collectionCode="FSCA" collectorName="FCH" country="United States of America" county="San Juan County" location="Zion National Park" municipality="Monticello" pageId="49" pageNumber="50" specimenCount="2" specimenCount-male="2" stateProvince="Utah">
same data except,
<date id="010D1069A51DFFD45474F933FB53F928" box="[1072,1192,1760,1787]" pageId="49" pageNumber="50" value="1951-04-20">
<collectingDate id="1149E981A51DFFD45474F933FB53F928" box="[1072,1192,1760,1787]" pageId="49" pageNumber="50" value="1951-04-20">20.iv.1951</collectingDate>
</date>
,
<collectorName id="D846537FA51DFFD454F0F933FB11F929" box="[1204,1258,1760,1786]" pageId="49" pageNumber="50">FCH</collectorName>
(
<specimenCount id="63B5FD20A51DFFD454BEF933FADBF929" box="[1274,1312,1760,1786]" count="2" pageId="49" pageNumber="50" type="male">2♂</specimenCount>
,
<collectionCode id="13A2AE6CA51DFFD4556FF933FA8DF929" box="[1323,1398,1760,1786]" collectionName="USA, Florida, Gainesville, Division of Plant Industry, Florida State Collection of Arthropods" pageId="49" pageNumber="50">FSCA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3DA96522A51DFFD45083F8D7FCD1F891" pageId="49" pageNumber="50" type="diagnosis">
<paragraph id="750C36A9A51DFFD45083F8D7FCD1F891" blockId="49.[151,1437,1400,1859]" pageId="49" pageNumber="50">
<emphasis id="47C7EABBA51DFFD45083F8D7FEB9F8CD" bold="true" box="[199,322,1796,1822]" pageId="49" pageNumber="50">Diagnosis.</emphasis>
This is the only known species of
<taxonomicName id="B2B34D2AA51DFFD4528CF8D7FC96F8CD" box="[712,877,1796,1822]" class="Insecta" family="Dolichopodidae" genus="Calyxochaetus" kingdom="Animalia" order="Diptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="47C7EABBA51DFFD4528CF8D7FC96F8CD" box="[712,877,1796,1822]" italics="true" pageId="49" pageNumber="50">Calyxochaetus</emphasis>
</taxonomicName>
whose males have two lamellae on the arista-like stylus (
<figureCitation id="ED882A2CA51DFFD450ACF8FBFEB1F891" box="[232,330,1832,1858]" captionStart="FIGURES 104–119" captionStartId="50.[151,264,1472,1496]" captionTargetBox="[151,1437,181,1448]" captionTargetId="figure-27@50.[151,1437,181,1448]" captionTargetPageId="50" captionText="FIGURES 104–119. Male antennae (lateral view) and forelegs of the Calyxochaetus frontalis species group. 104. C. binodatus (Harmston &amp; Knowlton). 105. C. nodatus (Loew). 106. C. desertus sp. nov. 107. C. pictipes (Harmston &amp; Knowlton). 108. C. sobrinus (Wheeler). 109. C. ornatus Parent. 110. C. remifer sp. nov. 111. C. tohono sp. nov. 112. C. frontalis (Loew). 113. C. furcatus (Van Duzee). 114. C. lamellicornis Parent (photo by Emmanuel Delfosse, MNHN). 115. C. lacrima sp. nov. 116. C. binodatus, fore tarsus, anterior view. 117. C. sobrinus, fore tarsus, anterior view. 118. C. remifer sp. nov., fore tarsus, dorsal view. 119. C. lacrima sp. nov., fore tibia and tarsus, posterior view. Scale bars = 0.5 mm; arista-like styli approximately 1 mm long." figureDoi="http://doi.org/10.5281/zenodo.14240476" httpUri="https://zenodo.org/record/14240476/files/figure.png" pageId="49" pageNumber="50">Fig. 104</figureCitation>
). Females have a semicircular postpedicel.
</paragraph>
</subSubSection>
<caption id="21CC6621A51EFFD750D3FA13FF30F963" ID-DOI="http://doi.org/10.5281/zenodo.14240476" ID-Zenodo-Dep="14240476" httpUri="https://zenodo.org/record/14240476/files/figure.png" pageId="50" pageNumber="51" startId="50.[151,264,1472,1496]" targetBox="[151,1437,181,1448]" targetPageId="50" targetType="figure">
<paragraph id="750C36A9A51EFFD750D3FA13FF30F963" blockId="50.[151,1437,1472,1712]" pageId="50" pageNumber="51">
<emphasis id="47C7EABBA51EFFD750D3FA13FE97FA0B" bold="true" box="[151,364,1472,1496]" pageId="50" pageNumber="51">FIGURES 104–119.</emphasis>
Male antennae (lateral view) and forelegs of the
<taxonomicName id="B2B34D2AA51EFFD75318FA13FBB7FA0B" baseAuthorityName="Loew" baseAuthorityYear="1861" box="[860,1100,1472,1496]" class="Insecta" family="Dolichopodidae" genus="Calyxochaetus" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="47C7EABBA51EFFD75318FA13FBB7FA0B" box="[860,1100,1472,1496]" italics="true" pageId="50" pageNumber="51">Calyxochaetus frontalis</emphasis>
</taxonomicName>
species group.
<emphasis id="47C7EABBA51EFFD754ADFA13FAEEFA0B" bold="true" box="[1257,1301,1472,1496]" pageId="50" pageNumber="51">104.</emphasis>
<taxonomicName id="B2B34D2AA51EFFD7555FFA12FE6DFA2F" authority="(Harmston &amp; Knowlton)" baseAuthorityName="Harmston &amp; Knowlton" baseAuthorityYear="1940" class="Insecta" family="Dolichopodidae" genus="Calyxochaetus" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="binodatus">
<emphasis id="47C7EABBA51EFFD7555FFA12FA67FA0B" box="[1307,1436,1472,1496]" italics="true" pageId="50" pageNumber="51">C. binodatus</emphasis>
(Harmston &amp; Knowlton)
</taxonomicName>
.
<emphasis id="47C7EABBA51EFFD751DBFA37FE37FA2F" bold="true" box="[415,460,1508,1532]" pageId="50" pageNumber="51">105.</emphasis>
<taxonomicName id="B2B34D2AA51EFFD75197FA36FD6EFA2F" authority="(Loew)" baseAuthorityName="Loew" box="[467,661,1508,1532]" class="Insecta" family="Dolichopodidae" genus="Sympycnus" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="nodatus">
<emphasis id="47C7EABBA51EFFD75197FA36FDBBFA2F" box="[467,576,1508,1532]" italics="true" pageId="50" pageNumber="51">C. nodatus</emphasis>
(Loew)
</taxonomicName>
.
<emphasis id="47C7EABBA51EFFD752DBFA37FD37FA2F" bold="true" box="[671,716,1508,1532]" pageId="50" pageNumber="51">106.</emphasis>
<taxonomicName id="B2B34D2AA51EFFD75296FA36FCBFFA2F" authorityName="Runyon" authorityYear="2024" box="[722,836,1508,1532]" class="Insecta" family="Dolichopodidae" genus="Calyxochaetus" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="desertus" status="sp. nov.">
<emphasis id="47C7EABBA51EFFD75296FA36FCBFFA2F" box="[722,836,1508,1532]" italics="true" pageId="50" pageNumber="51">C. desertus</emphasis>
</taxonomicName>
<emphasis id="47C7EABBA51EFFD7530FFA37FC2BFA2F" bold="true" box="[843,976,1508,1532]" pageId="50" pageNumber="51">
<taxonomicNameLabel id="5CF457C0A51EFFD7530FFA37FC67FA2F" box="[843,924,1508,1532]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
107.
</emphasis>
<taxonomicName id="B2B34D2AA51EFFD75392FA36FAB3FA2F" authority="(Harmston &amp; Knowlton)" baseAuthorityName="Harmston &amp; Knowlton" baseAuthorityYear="1940" box="[982,1352,1508,1532]" class="Insecta" family="Dolichopodidae" genus="Calyxochaetus" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="pictipes">
<emphasis id="47C7EABBA51EFFD75392FA36FBB8FA2F" box="[982,1091,1509,1532]" italics="true" pageId="50" pageNumber="51">C. pictipes</emphasis>
(Harmston &amp; Knowlton)
</taxonomicName>
.
<emphasis id="47C7EABBA51EFFD75516FA37FA84FA2F" bold="true" box="[1362,1407,1508,1532]" pageId="50" pageNumber="51">108.</emphasis>
<taxonomicName id="B2B34D2AA51EFFD755C1FA36FE9BF9F3" authority="(Wheeler)" baseAuthorityName="Wheeler" baseAuthorityYear="1899" class="Insecta" family="Dolichopodidae" genus="Calyxochaetus" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="sobrinus">
<emphasis id="47C7EABBA51EFFD755C1FA36FF15F9F3" italics="true" pageId="50" pageNumber="51">C. sobrinus</emphasis>
(Wheeler)
</taxonomicName>
.
<emphasis id="47C7EABBA51EFFD7512EF9DBFE6DF9F3" bold="true" box="[362,406,1544,1568]" pageId="50" pageNumber="51">109.</emphasis>
<taxonomicName id="B2B34D2AA51EFFD751D9F9DAFDA3F9F3" authority="Parent." authorityName="Parent." box="[413,600,1545,1568]" class="Insecta" family="Dolichopodidae" genus="Calyxochaetus" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="ornatus">
<emphasis id="47C7EABBA51EFFD751D9F9DAFDF2F9F3" box="[413,521,1545,1568]" italics="true" pageId="50" pageNumber="51">C. ornatus</emphasis>
Parent.
</taxonomicName>
<emphasis id="47C7EABBA51EFFD7521AF9DBFD71F9F3" bold="true" box="[606,650,1544,1568]" pageId="50" pageNumber="51">110.</emphasis>
<taxonomicName id="B2B34D2AA51EFFD752D5F9DAFD02F9F3" authorityName="Runyon" authorityYear="2024" box="[657,761,1544,1568]" class="Insecta" family="Dolichopodidae" genus="Calyxochaetus" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="remifer" status="sp. nov.">
<emphasis id="47C7EABBA51EFFD752D5F9DAFD02F9F3" box="[657,761,1544,1568]" italics="true" pageId="50" pageNumber="51">C. remifer</emphasis>
</taxonomicName>
<emphasis id="47C7EABBA51EFFD75344F9DBFC78F9F3" bold="true" box="[768,899,1544,1568]" pageId="50" pageNumber="51">
<taxonomicNameLabel id="5CF457C0A51EFFD75344F9DBFCAAF9F3" box="[768,849,1544,1568]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
111.
</emphasis>
<taxonomicName id="B2B34D2AA51EFFD753CDF9DAFC15F9F3" authorityName="Runyon" authorityYear="2024" box="[905,1006,1544,1568]" class="Insecta" family="Dolichopodidae" genus="Calyxochaetus" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="tohono" status="sp. nov.">
<emphasis id="47C7EABBA51EFFD753CDF9DAFC15F9F3" box="[905,1006,1544,1568]" italics="true" pageId="50" pageNumber="51">C. tohono</emphasis>
</taxonomicName>
<emphasis id="47C7EABBA51EFFD753B1F9DBFB81F9F3" bold="true" box="[1013,1146,1544,1568]" pageId="50" pageNumber="51">
<taxonomicNameLabel id="5CF457C0A51EFFD753B1F9DBFBBCF9F3" box="[1013,1095,1544,1568]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
112.
</emphasis>
<taxonomicName id="B2B34D2AA51EFFD754C4F9DAFAB3F9F3" authority="(Loew)" baseAuthorityName="Loew" baseAuthorityYear="1861" box="[1152,1352,1544,1568]" class="Insecta" family="Dolichopodidae" genus="Calyxochaetus" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="47C7EABBA51EFFD754C4F9DAFB0FF9F3" box="[1152,1268,1544,1568]" italics="true" pageId="50" pageNumber="51">C. frontalis</emphasis>
(Loew)
</taxonomicName>
.
<emphasis id="47C7EABBA51EFFD75517F9DBFA85F9F3" bold="true" box="[1363,1406,1544,1568]" pageId="50" pageNumber="51">113.</emphasis>
<taxonomicName id="B2B34D2AA51EFFD755C1F9DAFE8EF997" authority="(Van Duzee)" baseAuthorityName="Van Duzee" baseAuthorityYear="1929" class="Insecta" family="Dolichopodidae" genus="Calyxochaetus" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="furcatus">
<emphasis id="47C7EABBA51EFFD755C1F9DAFF12F997" italics="true" pageId="50" pageNumber="51">C. furcatus</emphasis>
(Van Duzee)
</taxonomicName>
.
<emphasis id="47C7EABBA51EFFD7513BF9FFFE50F997" bold="true" box="[383,427,1580,1604]" pageId="50" pageNumber="51">114.</emphasis>
<taxonomicName id="B2B34D2AA51EFFD751F6F9FEFD64F997" authority="Parent" authorityName="Parent" box="[434,671,1580,1604]" class="Insecta" family="Dolichopodidae" genus="Calyxochaetus" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="lamellicornis">
<emphasis id="47C7EABBA51EFFD751F6F9FEFDACF997" box="[434,599,1580,1604]" italics="true" pageId="50" pageNumber="51">C. lamellicornis</emphasis>
Parent
</taxonomicName>
(photo by Emmanuel Delfosse, MNHN).
<emphasis id="47C7EABBA51EFFD75415F9FFFB87F997" bold="true" box="[1105,1148,1580,1604]" pageId="50" pageNumber="51">115.</emphasis>
<taxonomicName id="B2B34D2AA51EFFD754C7F9FEFB0AF997" authorityName="Runyon" authorityYear="2024" box="[1155,1265,1580,1604]" class="Insecta" family="Dolichopodidae" genus="Calyxochaetus" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="lacrima" status="sp. nov.">
<emphasis id="47C7EABBA51EFFD754C7F9FEFB0AF997" box="[1155,1265,1580,1604]" italics="true" pageId="50" pageNumber="51">C. lacrima</emphasis>
</taxonomicName>
<emphasis id="47C7EABBA51EFFD754BDF9FFFA85F997" bold="true" box="[1273,1406,1580,1604]" pageId="50" pageNumber="51">
<taxonomicNameLabel id="5CF457C0A51EFFD754BDF9FFFAB1F997" box="[1273,1354,1580,1604]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
116.
</emphasis>
<taxonomicName id="B2B34D2AA51EFFD755C1F9FEFF00F9BB" baseAuthorityName="Harmston &amp; Knowlton" baseAuthorityYear="1940" class="Insecta" family="Dolichopodidae" genus="Calyxochaetus" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="binodatus">
<emphasis id="47C7EABBA51EFFD755C1F9FEFF00F9BB" italics="true" pageId="50" pageNumber="51">C. binodatus</emphasis>
</taxonomicName>
, fore tarsus, anterior view.
<emphasis id="47C7EABBA51EFFD7525CF983FDBFF9BB" bold="true" box="[536,580,1616,1640]" pageId="50" pageNumber="51">117.</emphasis>
<taxonomicName id="B2B34D2AA51EFFD7520FF982FD39F9BB" baseAuthorityName="Wheeler" baseAuthorityYear="1899" box="[587,706,1616,1640]" class="Insecta" family="Dolichopodidae" genus="Calyxochaetus" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="sobrinus">
<emphasis id="47C7EABBA51EFFD7520FF982FD39F9BB" box="[587,706,1616,1640]" italics="true" pageId="50" pageNumber="51">C. sobrinus</emphasis>
</taxonomicName>
, fore tarsus, anterior view.
<emphasis id="47C7EABBA51EFFD7539BF983FBF0F9BB" bold="true" box="[991,1035,1616,1640]" pageId="50" pageNumber="51">118.</emphasis>
<taxonomicName id="B2B34D2AA51EFFD75456F982FB80F9BB" authorityName="Runyon" authorityYear="2024" box="[1042,1147,1616,1640]" class="Insecta" family="Dolichopodidae" genus="Calyxochaetus" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="remifer" status="sp. nov.">
<emphasis id="47C7EABBA51EFFD75456F982FB80F9BB" box="[1042,1147,1616,1640]" italics="true" pageId="50" pageNumber="51">C. remifer</emphasis>
</taxonomicName>
<emphasis id="47C7EABBA51EFFD754C7F983FB2EF9BB" bold="true" box="[1155,1237,1616,1640]" pageId="50" pageNumber="51">
<taxonomicNameLabel id="5CF457C0A51EFFD754C7F983FB2EF9BB" box="[1155,1237,1616,1640]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, fore tarsus, dorsal view.
<emphasis id="47C7EABBA51EFFD75090F9A7FF04F95F" bold="true" box="[212,255,1652,1676]" pageId="50" pageNumber="51">119.</emphasis>
<taxonomicName id="B2B34D2AA51EFFD75141F9A6FE89F95F" authorityName="Runyon" authorityYear="2024" box="[261,370,1652,1676]" class="Insecta" family="Dolichopodidae" genus="Calyxochaetus" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="lacrima" status="sp. nov.">
<emphasis id="47C7EABBA51EFFD75141F9A6FE89F95F" box="[261,370,1652,1676]" italics="true" pageId="50" pageNumber="51">C. lacrima</emphasis>
</taxonomicName>
<emphasis id="47C7EABBA51EFFD7513DF9A7FE31F95F" bold="true" box="[377,458,1652,1676]" pageId="50" pageNumber="51">
<taxonomicNameLabel id="5CF457C0A51EFFD7513DF9A7FE31F95F" box="[377,458,1652,1676]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, fore tibia and tarsus, posterior view. Scale bars = 0.5 mm; arista-like styli approximately 1 mm long.
</paragraph>
</caption>
<subSubSection id="3DA96522A51EFFD65083F8D1FD1DFBC2" lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51" type="description">
<paragraph id="750C36A9A51EFFD65083F8D1FB35FCC6" blockId="50.[151,1436,1794,2037]" lastBlockId="51.[151,1437,150,1221]" lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51">
<emphasis id="47C7EABBA51EFFD75083F8D1FE3EF8CF" bold="true" box="[199,453,1794,1820]" pageId="50" pageNumber="51">Redescription. Male.</emphasis>
Wing length 2.7–3.0 mm.
<emphasis id="47C7EABBA51EFFD7534BF8D1FCA3F8CF" bold="true" box="[783,856,1794,1820]" pageId="50" pageNumber="51">Head:</emphasis>
Frons brown pruinose, sparser on either side of ocellar tubercle revealing some blue-violet ground color, not glabrous. Face silver-white, very narrow, about one ommatidium wide on ventral half. Antenna (
<figureCitation id="ED882A2CA51EFFD752CCF899FD1CF8B7" box="[648,743,1866,1892]" captionStart="FIGURES 104–119" captionStartId="50.[151,264,1472,1496]" captionTargetBox="[151,1437,181,1448]" captionTargetId="figure-27@50.[151,1437,181,1448]" captionTargetPageId="50" captionText="FIGURES 104–119. Male antennae (lateral view) and forelegs of the Calyxochaetus frontalis species group. 104. C. binodatus (Harmston &amp; Knowlton). 105. C. nodatus (Loew). 106. C. desertus sp. nov. 107. C. pictipes (Harmston &amp; Knowlton). 108. C. sobrinus (Wheeler). 109. C. ornatus Parent. 110. C. remifer sp. nov. 111. C. tohono sp. nov. 112. C. frontalis (Loew). 113. C. furcatus (Van Duzee). 114. C. lamellicornis Parent (photo by Emmanuel Delfosse, MNHN). 115. C. lacrima sp. nov. 116. C. binodatus, fore tarsus, anterior view. 117. C. sobrinus, fore tarsus, anterior view. 118. C. remifer sp. nov., fore tarsus, dorsal view. 119. C. lacrima sp. nov., fore tibia and tarsus, posterior view. Scale bars = 0.5 mm; arista-like styli approximately 1 mm long." figureDoi="http://doi.org/10.5281/zenodo.14240476" httpUri="https://zenodo.org/record/14240476/files/figure.png" pageId="50" pageNumber="51">Fig. 104</figureCitation>
) dark brown; scape stout; pedicel slightly longer and rounded medially; postpedicel semicircular; arista-like stylus with small oblong black lamella near 1/2 and larger rounded black lamella at apex. Palpus small, elongate oval, brown with apex yellow.
<emphasis id="47C7EABBA51EFFD753A9F841FBB4F87F" bold="true" box="[1005,1103,1938,1964]" pageId="50" pageNumber="51">Thorax:</emphasis>
Pleura brown with thick gray pruinosity; metepimeron yellow on ventral half or less. Scutellum without distinct blue-violet reflections.
<emphasis id="47C7EABBA51EFFD7551FF865FA67F803" bold="true" box="[1371,1436,1974,2000]" pageId="50" pageNumber="51">Legs:</emphasis>
<emphasis id="47C7EABBA51EFFD750D3F809FF14F827" box="[151,239,2010,2036]" italics="true" pageId="50" pageNumber="51">Foreleg</emphasis>
: Yellow, except coxa sometimes infuscated at very base, tarsomere 1 brown apically, tarsomere 2 sometimes yellow-brown, tarsomeres 3–5 dark brown. Femur with small yellow ventral setae on basal half. Tibia with small posterodorsal seta near 2/5. Tarsus (
<figureCitation id="ED882A2CA51FFFD6527CFF68FD61FF06" box="[568,666,187,213]" captionStart="FIGURES 104–119" captionStartId="50.[151,264,1472,1496]" captionTargetBox="[151,1437,181,1448]" captionTargetId="figure-27@50.[151,1437,181,1448]" captionTargetPageId="50" captionText="FIGURES 104–119. Male antennae (lateral view) and forelegs of the Calyxochaetus frontalis species group. 104. C. binodatus (Harmston &amp; Knowlton). 105. C. nodatus (Loew). 106. C. desertus sp. nov. 107. C. pictipes (Harmston &amp; Knowlton). 108. C. sobrinus (Wheeler). 109. C. ornatus Parent. 110. C. remifer sp. nov. 111. C. tohono sp. nov. 112. C. frontalis (Loew). 113. C. furcatus (Van Duzee). 114. C. lamellicornis Parent (photo by Emmanuel Delfosse, MNHN). 115. C. lacrima sp. nov. 116. C. binodatus, fore tarsus, anterior view. 117. C. sobrinus, fore tarsus, anterior view. 118. C. remifer sp. nov., fore tarsus, dorsal view. 119. C. lacrima sp. nov., fore tibia and tarsus, posterior view. Scale bars = 0.5 mm; arista-like styli approximately 1 mm long." figureDoi="http://doi.org/10.5281/zenodo.14240476" httpUri="https://zenodo.org/record/14240476/files/figure.png" pageId="51" pageNumber="52">Fig. 116</figureCitation>
) with tarsomere 1 very short, about as wide as long. Tarsomere 3 with small (less than length tarsomere 4) curved anterodorsal apical seta. Tarsomere 4 with dorsal surface clothed in dense white microsetulae. Tarsomere 5 dorsally with sparser white microsetulae.
<emphasis id="47C7EABBA51FFFD6541CFED0FB5CFECE" box="[1112,1191,259,285]" italics="true" pageId="51" pageNumber="52">Midleg</emphasis>
: Yellow, except coxa infuscated with yellow ground color visible, tarsus brown from apex of tarsomere 1. Femur with anteroventral row of slender yellow to yellow-brown setae on basal three-fifths, increasing in size distally (largest subequal to femur width). Tibia with anteroventral full-length row of long (2X tibia width) slender yellow to yellow-brown setae, bent downward at tips, with conspicuous black preapical anteroventral seta. Tarsus (
<figureCitation id="ED882A2CA51FFFD6544DFE40FB93FE7E" box="[1033,1128,403,429]" captionStart="FIGURES 120–131" captionStartId="52.[151,264,1711,1735]" captionTargetBox="[196,1392,181,1688]" captionTargetId="figure-27@52.[196,1392,181,1688]" captionTargetPageId="52" captionText="FIGURES 120–131. Male mid tarsi of the Calyxochaetus frontalis species group (anterior on left side). 120. C. binodatus (Harmston &amp; Knowlton), dorsal view. 121. C. nodatus (Loew), ventral view. 122. C. bovanodatus sp. nov., dorsal view. 123. C. desertus sp. nov., mid tarsomeres 2–5, dorsal view. 124. C. sobrinus (Wheeler), mid tarsomeres 2–5, dorsal view. 125. C. pictipes (Harmston &amp; Knowlton), dorsal view. 126. C. ornatus Parent, mid tarsomeres 2–5, ventral view. 127. C. tohono sp. nov., mid tarsomeres 2–5, dorsal view. 128. C. remifer sp. nov., dorsal view. 129. C. frontalis (Loew), dorsal view. 130. C. furcatus (Van Duzee), dorsal view. 131. C. lacrima sp. nov., dorsal view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14240480" httpUri="https://zenodo.org/record/14240480/files/figure.png" pageId="51" pageNumber="52">Fig. 120</figureCitation>
) short, half as long as tibia. Tarsomere 1 bowed slightly, longer than combined length of tarsomeres 2–5. Tarsomeres 2–4 widened and pointed anteriorly, with white microsetulae at anterior tips. Tarsomere 2 broadening from base anteriorly, longer and pointed anteroapically (overlapping tarsomere 3). Tarsomeres 3–4 small, subequal in size, wider than long.
<emphasis id="47C7EABBA51FFFD654A5FE2CFAC0FDCA" box="[1249,1339,511,537]" italics="true" pageId="51" pageNumber="52">Hindleg</emphasis>
: Yellow, except coxa infuscated at base, femur light brownish dorsoapically, tarsus brown from near middle of tarsomere 1. Femur with long (subequal to femur width) anteroventral brown seta near 3/4 (
<specimenCount id="63B5FD20A51FFFD65468FD94FB3CFDB2" box="[1068,1223,583,609]" count="1" pageId="51" pageNumber="52" type="generic">one specimen</specimenCount>
has two setae near 3/4). Tibia (
<figureCitation id="ED882A2CA51FFFD6515AFDB8FE85FD56" box="[286,382,619,645]" captionStart="FIGURES 132–138" captionStartId="54.[151,264,1423,1447]" captionTargetBox="[152,1436,381,1400]" captionTargetId="figure-114@54.[152,1436,381,1400]" captionTargetPageId="54" captionText="FIGURES 132–138. Male hind legs and wings of the Calyxochaetus frontalis species group. 132. C. binodatus (Harmston &amp; Knowlton), hind tibia, anterior view. 133. C. nodatus (Loew), hind tibia, anterior view. 134. C. ornatus Parent, hind tibia, posterior view. 135. C. frontalis (Loew), hind tibia, posterior view. 136. C. lacrima sp. nov., apical third of hind femur and basal half of hind tibia, posterior view. 137. C. nodatus, wing. 138. C. ornatus, wing. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.14240484" httpUri="https://zenodo.org/record/14240484/files/figure.png" pageId="51" pageNumber="52">Fig. 132</figureCitation>
) with clump of 3–5 very long (3X tibia width or longer) yellow-brown ventral setae at base, tibia slightly bowed just beyond clump.
<emphasis id="47C7EABBA51FFFD65221FD5DFD54FD7B" bold="true" box="[613,687,654,680]" pageId="51" pageNumber="52">Wing:</emphasis>
Hyaline. Anal lobe constricted at very base (basal to fork of R
<subScript id="E93734ECA51FFFD655C7FD72FA67FD7C" attach="left" box="[1411,1436,673,687]" fontSize="6" pageId="51" pageNumber="52">2+3</subScript>
and R
<subScript id="E93734ECA51FFFD6509DFD16FF08FD00" attach="left" box="[217,243,709,723]" fontSize="6" pageId="51" pageNumber="52">4+5</subScript>
), otherwise well-developed. Distal section of M
<subScript id="E93734ECA51FFFD6534DFD16FCEAFD00" attach="left" box="[777,785,709,723]" fontSize="6" pageId="51" pageNumber="52">4</subScript>
about 4X longer than crossvein dm-m.
<emphasis id="47C7EABBA51FFFD6548EFD61FAB3FD1F" bold="true" box="[1226,1352,690,716]" pageId="51" pageNumber="52">Abdomen:</emphasis>
Tergite 1 brown, pruinose, concolorous to thorax. Tergites 2–3 mostly yellow, brown dorsally. Tergite 4 broadly yellow at base.
<emphasis id="47C7EABBA51FFFD65093FD28FE9DFCC7" box="[215,358,763,788]" italics="true" pageId="51" pageNumber="52">Hypopygium</emphasis>
: Postgonites with apical lobes narrow, apex rounded, translucent light brown.
</paragraph>
<paragraph id="750C36A9A51FFFD65083FCCCFD1DFBC2" blockId="51.[151,1437,150,1221]" pageId="51" pageNumber="52">
<emphasis id="47C7EABBA51FFFD65083FCCCFEDEFCEA" bold="true" box="[199,293,799,825]" pageId="51" pageNumber="52">Female.</emphasis>
Wing length 3.0–
<quantity id="B24B9B4CA51FFFD651B4FCCCFDB1FCEA" box="[496,586,799,825]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="51" pageNumber="52" unit="mm" value="3.4">3.4 mm</quantity>
. Similar to male except as follows:
<emphasis id="47C7EABBA51FFFD65399FCCCFBDDFCEA" bold="true" box="[989,1062,799,825]" pageId="51" pageNumber="52">Head:</emphasis>
Face wide, nearly parallel-sided, gray. Antenna with postpedicel short, semicircular; arista-like stylus unmodified, blunted at apex. Palpus much larger, triangular, rounded apically, brown, with white setulae and brown setae.
<emphasis id="47C7EABBA51FFFD65453FCB4FB82FC52" bold="true" box="[1047,1145,871,897]" pageId="51" pageNumber="52">Thorax:</emphasis>
Proepimeron with cluster of 5–6 small white hairs.
<emphasis id="47C7EABBA51FFFD651EAFC58FE14FC76" bold="true" box="[430,495,907,933]" pageId="51" pageNumber="52">Legs:</emphasis>
Unmodified, without specialized setae. Hind femur with indistinct anteroventral seta near 3/4 (corresponding to long seta in male). Hind tarsomere 1 yellow with brown apex.
<emphasis id="47C7EABBA51FFFD65438FC7DFB3DFC1B" bold="true" box="[1148,1222,942,968]" pageId="51" pageNumber="52">Wing:</emphasis>
Anal lobe broader, not constricted at base. Distal section of M
<subScript id="E93734ECA51FFFD652C5FC36FD72FC20" attach="left" box="[641,649,997,1011]" fontSize="6" pageId="51" pageNumber="52">4</subScript>
about 3X longer than crossvein dm-m.
<emphasis id="47C7EABBA51FFFD65414FC01FB35FC3F" bold="true" box="[1104,1230,978,1004]" pageId="51" pageNumber="52">Abdomen:</emphasis>
Tergites 2–3 with broad yellow lateral spots, remaining tergites brown.
</paragraph>
</subSubSection>
<subSubSection id="3DA96522A51FFFD65083FBC9FD1EFB8A" pageId="51" pageNumber="52" type="distribution">
<paragraph id="750C36A9A51FFFD65083FBC9FD1EFB8A" blockId="51.[151,1437,150,1221]" pageId="51" pageNumber="52">
<emphasis id="47C7EABBA51FFFD65083FBC9FD6FFBE6" bold="true" box="[199,660,1050,1077]" pageId="51" pageNumber="52">Distribution and seasonal occurrence.</emphasis>
Known from
<collectingRegion id="B777F84BA51FFFD6537EFBC8FC6CFBE6" box="[826,919,1051,1077]" country="United States of America" name="Arizona" pageId="51" pageNumber="52">Arizona</collectingRegion>
,
<collectingRegion id="B777F84BA51FFFD653E1FBC8FBC1FBE6" box="[933,1082,1051,1077]" country="United States of America" name="New Mexico" pageId="51" pageNumber="52">New Mexico</collectingRegion>
and southern
<collectingRegion id="B777F84BA51FFFD654A5FBC8FAECFBE6" box="[1249,1303,1051,1077]" country="United States of America" name="Utah" pageId="51" pageNumber="52">Utah</collectingRegion>
(
<figureCitation id="ED882A2CA51FFFD6556EFBC8FA6BFBE6" box="[1322,1424,1051,1077]" captionStart="FIGURES 144–146" captionStartId="64.[151,264,1684,1708]" captionTargetBox="[151,1436,181,1660]" captionTargetId="figure-27@64.[151,1436,181,1660]" captionTargetPageId="64" captionText="FIGURES 144–146. Known geographical distributions of species in the Calyxochaetus frontalis group. 144. C. nodatus (Loew), C. binodatus (Harmston &amp; Knowlton) and C. bovanodatus sp. nov. 145. C. sobrinus (Wheeler) and C. pictipes (Harmston &amp; Knowlton). 146. C. frontalis (Loew) and C. desertus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.14240490" httpUri="https://zenodo.org/record/14240490/files/figure.png" pageId="51" pageNumber="52">Fig. 144</figureCitation>
). Adults have been collected from April to September.
</paragraph>
</subSubSection>
<subSubSection id="3DA96522A51FFFD65083FBB0FEB7FB16" pageId="51" pageNumber="52" type="discussion">
<paragraph id="750C36A9A51FFFD65083FBB0FEB7FB16" blockId="51.[151,1437,150,1221]" pageId="51" pageNumber="52">
<emphasis id="47C7EABBA51FFFD65083FBB0FEC0FBAE" bold="true" box="[199,315,1123,1149]" pageId="51" pageNumber="52">Remarks.</emphasis>
Adults were collected in Mojave County
<collectingRegion id="B777F84BA51FFFD65355FBB0FC90FBAE" box="[785,875,1123,1149]" country="United States of America" name="Arizona" pageId="51" pageNumber="52">Arizona</collectingRegion>
from low vegetation in the shade of the Highway 91 bridge at Beaver Dam, and in Santa Cruz County on shaded vegetation along Sonoita Creek near the inlet to Patagonia Lake.
</paragraph>
</subSubSection>
</treatment>
</document>
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