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<document id="F6AFC9E33406337C4204576958C9BA48" ID-CLB-Dataset="4741" ID-DOI="10.11646/zootaxa.4500.1.8" ID-GBIF-Dataset="f5739d78-7cb5-4373-82dd-09eb248f7c30" ID-ISSN="1175-5326" ID-Zenodo-Dep="5297416" ID-ZooBank="B2F30EBD-60D2-4413-B269-5863F5FD1D5B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630082613700" checkinUser="felipe" docAuthor="Ng, Heok Hee &amp; Lalronunga, Samuel" docDate="2018" docId="03D987DFAB4AA31810CAFF5CFE87F824" docLanguage="en" docName="zootaxa.4500.1.8.pdf" docOrigin="Zootaxa 4500 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pterocryptis subrisa Ng &amp; Lalronunga 2018, new species" docType="treatment" docUuid="946302FD-5BEF-4533-9D7F-384B8804E507" docUuidSource="ZooBank" docVersion="8" lastPageNumber="128" masterDocId="FFE0FFA7AB4BA31A105DFFCBFFADFFFF" masterDocTitle="Pterocryptis subrisa, a new silurid catfish (Teleostei: Siluridae) from northeastern India" masterLastPageNumber="134" masterPageNumber="126" pageNumber="127" updateTime="1699146240577" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="2E9FC0EC0FE5DEBF09242F7E0ECA1615" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="7BE936323995FA2712C50796504852E1">Pterocryptis subrisa, a new silurid catfish (Teleostei: Siluridae) from northeastern India</mods:title>
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<mods:name id="94553BD616E3F3D25FFA8BC933785C48" type="personal">
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<mods:namePart id="E46444D9FA061C57CD493FDB84E68296">Ng, Heok Hee</mods:namePart>
<mods:affiliation id="8DC64CC86ECC5AE65F617DADAB26EE8F">c / o Lee Kong Chian Natural History Museum, 2 Conservatory Drive, Singapore 117377. &amp; Department of Zoology, Pachhunga University College, Aizawl, Mizoram, India 796001. E-mail: lrl _ zoo @ yahoo. co. in</mods:affiliation>
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<mods:role id="55217459B4E9FB36B46132F04E178D05">
<mods:roleTerm id="C272B34D2164EA8A8F35C0AAF67CEECA">Author</mods:roleTerm>
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<mods:namePart id="3C9EECEC059F33A78A3FA4E7260F8797">Lalronunga, Samuel</mods:namePart>
<mods:affiliation id="67D76EC27C59A9D92AA4F57913AF990F">Department of Zoology, Pachhunga University College, Aizawl, Mizoram 796001, India. E-mail: samuellrna @ gmail. com</mods:affiliation>
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<mods:title id="B8468E2E1D46833703FB415155150D3B">Zootaxa</mods:title>
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<mods:part id="472FF53B0BBC3C8D55E6761AC375B1F9">
<mods:date id="307FA0BAA8247B13ABD4BA01DCDD6B3A">2018</mods:date>
<mods:detail id="9ADE5B89B78FCF30125BA8EFE9E856B1" type="pubDate">
<mods:number id="DBAC3F23E6110A057507B677E6E84CC7">2018-10-15</mods:number>
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<mods:detail id="2187A068A6F813974EA344C094C39BE5" type="volume">
<mods:number id="ADB38D31DF77B43BBA49356609F24C49">4500</mods:number>
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<mods:classification id="D0D01D2CF708DBACFE580B7033B35015">journal article</mods:classification>
<mods:identifier id="38CFA7881B6A9C49B23D240EDAFD5E59" type="CLB-Dataset">4741</mods:identifier>
<mods:identifier id="AB69219817B066AB82154DFBD9F5169F" type="DOI">10.11646/zootaxa.4500.1.8</mods:identifier>
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<mods:identifier id="ED71F59AC7DF28E8FF1D57692FD96FB7" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03D987DFAB4AA31810CAFF5CFE87F824" ID-DOI="http://doi.org/10.5281/zenodo.5303561" ID-GBIF-Taxon="185118362" ID-Zenodo-Dep="5303561" LSID="urn:lsid:zoobank.org:act:946302FD-5BEF-4533-9D7F-384B8804E507" httpUri="http://treatment.plazi.org/id/03D987DFAB4AA31810CAFF5CFE87F824" lastPageId="2" lastPageNumber="128" pageId="1" pageNumber="127">
<subSubSection id="C36A6542AB4AA31B10CAFF5CFC18FF2C" pageId="1" pageNumber="127" type="nomenclature">
<paragraph id="8BCF36C9AB4AA31B10CAFF5CFDB3FF4D" blockId="1.[151,949,151,247]" box="[151,542,151,178]" pageId="1" pageNumber="127">
<heading id="D08781A5AB4AA31B10CAFF5CFDB3FF4D" bold="true" box="[151,542,151,178]" fontSize="11" level="1" pageId="1" pageNumber="127" reason="1">
<emphasis id="B904EADBAB4AA31B10CAFF5CFDB3FF4D" bold="true" box="[151,542,151,178]" pageId="1" pageNumber="127">
<taxonomicName id="4C704D4AAB4AA31B10CAFF5CFE2AFF4E" authority="Ng &amp; Lalronunga, 2018" authorityName="Ng &amp; Lalronunga" authorityYear="2018" box="[151,391,151,177]" class="Actinopterygii" family="Siluridae" genus="Pterocryptis" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="subrisa" status="sp. nov.">
<emphasis id="B904EADBAB4AA31B10CAFF5CFE2AFF4E" bold="true" box="[151,391,151,177]" italics="true" pageId="1" pageNumber="127">Pterocryptis subrisa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23757A0AB4AA31B11D3FF53FDB3FF4D" box="[398,542,152,178]" pageId="1" pageNumber="127" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF36C9AB4AA31B10CAFF71FC18FF2C" blockId="1.[151,949,151,247]" box="[151,949,186,211]" pageId="1" pageNumber="127">
<uri id="FFE13ACBAB4AA31B10CAFF71FC18FF2C" box="[151,949,186,211]" pageId="1" pageNumber="127">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="C36A6542AB4AA31B10CAFF15FF75FF08" box="[151,216,222,247]" pageId="1" pageNumber="127" type="description">
<paragraph id="8BCF36C9AB4AA31B10CAFF15FF75FF08" blockId="1.[151,949,151,247]" box="[151,216,222,247]" pageId="1" pageNumber="127">
<figureCitation id="134B2A4CAB4AA31B10CAFF15FF75FF08" box="[151,216,222,247]" captionStart="FIGURE 1" captionStartId="1.[151,250,1908,1930]" captionTargetBox="[151,1436,734,1883]" captionTargetId="figure-238@1.[150,1436,734,1883]" captionTargetPageId="1" captionText="FIGURE 1. Pterocryptis subrisa, holotype, ZSI FF 7870, 91.8 mm SL; India: Maisa River." figureDoi="http://doi.org/10.5281/zenodo.5297418" httpUri="https://zenodo.org/record/5297418/files/figure.png" pageId="1" pageNumber="127">Fig. 1</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36A6542AB4AA31B10CAFEECFE91FD58" pageId="1" pageNumber="127" type="materials_examined">
<paragraph id="8BCF36C9AB4AA31B10CAFEECFCABFE9C" blockId="1.[151,1437,294,679]" pageId="1" pageNumber="127">
<materialsCitation id="3B183C94AB4AA31B10CAFEECFCABFE9C" ID-GBIF-Occurrence="3339716302" collectingDate="2011-12-09" country="India" county="Saiha District" latitude="22.297222" location="Maisa" longLatPrecision="21" longitude="92.917496" municipality="Maisa River" pageId="1" pageNumber="127" specimenCount="1" stateProvince="Mizoram" typeStatus="holotype">
<emphasis id="B904EADBAB4AA31B10CAFEECFEA6FEC0" bold="true" box="[151,267,295,319]" inLineHeading="true" pageId="1" pageNumber="127" reason="1">
<typeStatus id="54CB886BAB4AA31B10CAFEECFEABFEC0" box="[151,262,295,319]" pageId="1" pageNumber="127" type="holotype">Holotype</typeStatus>
.
</emphasis>
ZSI FF 7870,
<quantity id="4C889B2CAB4AA31B119FFEEDFD86FEC1" box="[450,555,294,319]" metricMagnitude="-2" metricUnit="m" metricValue="9.18" pageId="1" pageNumber="127" unit="mm" value="91.8">91.8 mm</quantity>
SL;
<collectingCountry id="F3677659AB4AA31B1236FEEDFD04FEC0" box="[619,681,294,319]" name="India" pageId="1" pageNumber="127">India</collectingCountry>
:
<collectingRegion id="49B4F82BAB4AA31B12E5FEEDFC8FFEC0" box="[696,802,294,319]" country="India" name="Mizoram" pageId="1" pageNumber="127">Mizoram</collectingRegion>
,
<collectingCounty id="62AE4E45AB4AA31B136CFEEDFC7FFEC0" box="[817,978,294,319]" pageId="1" pageNumber="127">Saiha District</collectingCounty>
,
<collectingMunicipality id="6BABACB3AB4AA31B13BFFEEDFBDDFEC0" box="[994,1136,294,319]" pageId="1" pageNumber="127">Maisa River</collectingMunicipality>
in the vicinity of
<location id="8EAF6012AB4AA31B150CFEEDFA35FEC0" LSID="urn:lsid:plazi:treatment:03D987DFAB4AA31810CAFF5CFE87F824:8EAF6012AB4AA31B150CFEEDFA35FEC0" box="[1361,1432,294,319]" country="India" county="Saiha District" latitude="22.297222" longLatPrecision="21" longitude="92.917496" municipality="Maisa River" name="Maisa" pageId="1" pageNumber="127" stateProvince="Mizoram">Maisa</location>
,
<geoCoordinate id="EE44500EAB4AA31B10CAFE81FEB7FE9C" box="[151,282,330,355]" degrees="22" direction="north" minutes="17" orientation="latitude" pageId="1" pageNumber="127" precision="15" seconds="50" value="22.297222">22°17'50&quot;N</geoCoordinate>
<geoCoordinate id="EE44500EAB4AA31B117FFE81FE35FE9C" box="[290,408,330,355]" degrees="92" direction="east" minutes="55" orientation="longitude" pageId="1" pageNumber="127" precision="15" seconds="3" value="92.917496">92°55'3&quot;E</geoCoordinate>
;
<location id="8EAF6012AB4AA31B11FEFE81FD80FE9C" LSID="urn:lsid:plazi:treatment:03D987DFAB4AA31810CAFF5CFE87F824:8EAF6012AB4AA31B11FEFE81FD80FE9C" box="[419,557,330,355]" country="India" county="Saiha District" latitude="22.297222" longLatPrecision="21" longitude="92.917496" municipality="Maisa River" name="Lalramliana" pageId="1" pageNumber="127" stateProvince="Mizoram">Lalramliana</location>
,
<date id="FFCE1009AB4AA31B1265FE81FCAEFE9C" box="[568,771,330,355]" pageId="1" pageNumber="127" value="2011-12-09">
<collectingDate id="EF8AE9E1AB4AA31B1265FE81FCAEFE9C" box="[568,771,330,355]" pageId="1" pageNumber="127" value="2011-12-09">9 December 2011</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8BCF36C9AB4AA31B109AFEA4FE91FD58" blockId="1.[151,1437,294,679]" pageId="1" pageNumber="127">
<materialsCitation id="3B183C94AB4AA31B109AFEA4FBECFE54" ID-GBIF-Occurrence="3339716305" collectingDate="2012-11-15" collectorName="S Lalronunga &amp; Lalramliana." country="India" county="Saiha District" latitude="22.194166" location="Mipu" longLatPrecision="21" longitude="92.89611" municipality="Palak River" pageId="1" pageNumber="127" specimenCount="1" stateProvince="Mizoram" typeStatus="paratype">
<emphasis id="B904EADBAB4AA31B109AFEA4FEEBFE78" bold="true" box="[199,326,367,391]" inLineHeading="true" pageId="1" pageNumber="127" reason="1">
<typeStatus id="54CB886BAB4AA31B109AFEA4FEECFE78" box="[199,321,367,391]" pageId="1" pageNumber="127" type="paratype">Paratypes</typeStatus>
.
</emphasis>
ZSI FF 7871 (2),
<quantity id="4C889B2CAB4AA31B1248FEA5FD79FE79" box="[533,724,366,391]" metricMagnitude="-1" metricUnit="m" metricValue="1.4164999999999996" metricValueMax="1.677" metricValueMin="1.156" pageId="1" pageNumber="127" unit="mm" value="141.64999999999998" valueMax="167.7" valueMin="115.6">115.6–167.7 mm</quantity>
SL;
<collectingCountry id="F3677659AB4AA31B1351FEA5FCE7FE78" box="[780,842,366,391]" name="India" pageId="1" pageNumber="127">India</collectingCountry>
:
<collectingRegion id="49B4F82BAB4AA31B1308FEA5FC12FE78" box="[853,959,366,391]" country="India" name="Mizoram" pageId="1" pageNumber="127">Mizoram</collectingRegion>
,
<collectingCounty id="62AE4E45AB4AA31B1397FEA5FBCAFE78" box="[970,1127,366,391]" pageId="1" pageNumber="127">Saiha District</collectingCounty>
,
<collectingMunicipality id="6BABACB3AB4AA31B142EFEA5FB5AFE78" box="[1139,1271,366,391]" pageId="1" pageNumber="127">Palak River</collectingMunicipality>
in the vicinity of
<location id="8EAF6012AB4AA31B10E9FE59FF59FE54" LSID="urn:lsid:plazi:treatment:03D987DFAB4AA31810CAFF5CFE87F824:8EAF6012AB4AA31B10E9FE59FF59FE54" box="[180,244,402,427]" country="India" county="Saiha District" latitude="22.194166" longLatPrecision="21" longitude="92.89611" municipality="Palak River" name="Mipu" pageId="1" pageNumber="127" stateProvince="Mizoram">Mipu</location>
,
<geoCoordinate id="EE44500EAB4AA31B10A2FE59FE2FFE54" box="[255,386,402,427]" degrees="22" direction="north" minutes="11" orientation="latitude" pageId="1" pageNumber="127" precision="15" seconds="39" value="22.194166">22°11'39&quot;N</geoCoordinate>
<geoCoordinate id="EE44500EAB4AA31B11D5FE59FDA1FE54" box="[392,524,402,427]" degrees="92" direction="east" minutes="53" orientation="longitude" pageId="1" pageNumber="127" precision="15" seconds="46" value="92.89611">92°53'46&quot;E</geoCoordinate>
;
<collectorName id="2685531FAB4AA31B124AFE59FD03FE54" box="[535,686,402,427]" pageId="1" pageNumber="127">S Lalronunga</collectorName>
&amp;
<collectorName id="2685531FAB4AA31B128CFE59FCCDFE54" box="[721,864,402,427]" pageId="1" pageNumber="127">Lalramliana.</collectorName>
<date id="FFCE1009AB4AA31B133BFE59FBECFE54" box="[870,1089,402,427]" pageId="1" pageNumber="127" value="2012-11-15">
<collectingDate id="EF8AE9E1AB4AA31B133BFE59FBECFE54" box="[870,1089,402,427]" pageId="1" pageNumber="127" value="2012-11-15">15 November 2012</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B183C94AB4AA31B1416FE59FDA5FE0C" ID-GBIF-Occurrence="3339716303" collectingDate="2012-11-16" collectionCode="PUCMF" collectorName="S Lalronunga &amp; Lalramliana" country="India" county="Saiha District" latitude="22.195555" location="Khopai" longLatPrecision="21" longitude="93.024445" municipality="Tuisi River" pageId="1" pageNumber="127" specimenCode="PUCMF 18010" specimenCount="1" stateProvince="Mizoram" typeStatus="paratype">
<specimenCode id="DBD69EB2AB4AA31B1416FE59FB5BFE54" box="[1099,1270,402,427]" collectionCode="PUCMF" pageId="1" pageNumber="127">PUCMF 18010</specimenCode>
(1),
<quantity id="4C889B2CAB4AA31B1576FE59FA30FE55" box="[1323,1437,402,427]" metricMagnitude="-1" metricUnit="m" metricValue="1.594" pageId="1" pageNumber="127" unit="mm" value="159.4">159.4 mm</quantity>
SL;
<collectingCountry id="F3677659AB4AA31B109BFE7DFEA9FE30" box="[198,260,438,463]" name="India" pageId="1" pageNumber="127">India</collectingCountry>
:
<collectingRegion id="49B4F82BAB4AA31B1153FE7DFEDAFE30" box="[270,375,438,463]" country="India" name="Mizoram" pageId="1" pageNumber="127">Mizoram</collectingRegion>
,
<collectingCounty id="62AE4E45AB4AA31B11DEFE7DFDB2FE30" box="[387,543,438,463]" pageId="1" pageNumber="127">Saiha District</collectingCounty>
,
<collectingMunicipality id="6BABACB3AB4AA31B1277FE7DFD0AFE30" box="[554,679,438,463]" pageId="1" pageNumber="127">Tuisi River</collectingMunicipality>
in the vicinity of
<location id="8EAF6012AB4AA31B1333FE7DFC6EFE30" LSID="urn:lsid:plazi:treatment:03D987DFAB4AA31810CAFF5CFE87F824:8EAF6012AB4AA31B1333FE7DFC6EFE30" box="[878,963,438,463]" country="India" county="Saiha District" latitude="22.195555" longLatPrecision="21" longitude="93.024445" municipality="Tuisi River" name="Khopai" pageId="1" pageNumber="127" stateProvince="Mizoram">Khopai</location>
,
<geoCoordinate id="EE44500EAB4AA31B1392FE7DFBFCFE30" box="[975,1105,438,463]" degrees="22" direction="north" minutes="11" orientation="latitude" pageId="1" pageNumber="127" precision="15" seconds="44" value="22.195555">22°11'44&quot;N</geoCoordinate>
<geoCoordinate id="EE44500EAB4AA31B1405FE7DFB70FE30" box="[1112,1245,438,463]" degrees="93" direction="east" minutes="01" orientation="longitude" pageId="1" pageNumber="127" precision="15" seconds="28" value="93.024445">93°01'28&quot;E</geoCoordinate>
;
<collectorName id="2685531FAB4AA31B14B5FE7DFAD2FE30" box="[1256,1407,438,463]" pageId="1" pageNumber="127">S Lalronunga</collectorName>
&amp;
<collectorName id="2685531FAB4AA31B10CAFE11FE8CFE0C" box="[151,289,474,499]" pageId="1" pageNumber="127">Lalramliana</collectorName>
,
<date id="FFCE1009AB4AA31B1171FE11FDA5FE0C" box="[300,520,474,499]" pageId="1" pageNumber="127" value="2012-11-16">
<collectingDate id="EF8AE9E1AB4AA31B1171FE11FDA5FE0C" box="[300,520,474,499]" pageId="1" pageNumber="127" value="2012-11-16">16 November 2012</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B183C94AB4AA31B124FFE11FB15FE0C" ID-GBIF-Occurrence="3339716301" box="[530,1208,474,499]" collectingDate="2011-12-09" collectionCode="PUCMF" country="India" county="Saiha District" latitude="22.297222" location="Maisa" longLatPrecision="21" longitude="92.917496" municipality="Maisa River" pageId="1" pageNumber="127" specimenCode="PUCMF 18011" specimenCount="1" stateProvince="Mizoram" typeStatus="paratype">
<specimenCode id="DBD69EB2AB4AA31B124FFE11FD10FE0C" box="[530,701,474,499]" collectionCode="PUCMF" pageId="1" pageNumber="127">PUCMF 18011</specimenCode>
(5),
<quantity id="4C889B2CAB4AA31B12ACFE11FC0CFE0D" box="[753,929,474,499]" metricMagnitude="-1" metricUnit="m" metricValue="1.015" metricValueMax="1.149" metricValueMin="0.8809999999999999" pageId="1" pageNumber="127" unit="mm" value="101.5" valueMax="114.9" valueMin="88.1">88.1–114.9 mm</quantity>
SL; data as for holotype
</materialsCitation>
.
<materialsCitation id="3B183C94AB4AA31B149FFE11FDEEFDC4" ID-GBIF-Occurrence="3339716306" collectingDate="2012-11-15" collectionCode="PUCMF" collectorName="Lalramliana &amp; S Lalronunga" country="India" latitude="22.397223" location="Kawlchaw" longLatPrecision="21" longitude="92.96972" municipality="Kawlchaw River" pageId="1" pageNumber="127" specimenCode="PUCMF 12071" specimenCount="1" stateProvince="Mizoram" typeStatus="paratype">
<specimenCode id="DBD69EB2AB4AA31B149FFE11FAC3FE0C" box="[1218,1390,474,499]" collectionCode="PUCMF" pageId="1" pageNumber="127">PUCMF 12071</specimenCode>
(1),
<quantity id="4C889B2CAB4AA31B10CAFE35FEA5FDE9" box="[151,264,510,535]" metricMagnitude="-1" metricUnit="m" metricValue="1.204" pageId="1" pageNumber="127" unit="mm" value="120.4">120.4 mm</quantity>
SL;
<collectingCountry id="F3677659AB4AA31B1160FE35FED6FDE8" box="[317,379,510,535]" name="India" pageId="1" pageNumber="127">India</collectingCountry>
:
<collectingRegion id="49B4F82BAB4AA31B11D8FE35FE42FDE8" box="[389,495,510,535]" country="India" name="Mizoram" pageId="1" pageNumber="127">Mizoram</collectingRegion>
,
<collectingMunicipality id="6BABACB3AB4AA31B11A5FE35FD18FDE8" box="[504,693,510,535]" pageId="1" pageNumber="127">Kawlchaw River</collectingMunicipality>
in the vicinity of
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,
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<geoCoordinate id="EE44500EAB4AA31B14DAFE35FAA7FDE8" box="[1159,1290,510,535]" degrees="92" direction="east" minutes="58" orientation="longitude" pageId="1" pageNumber="127" precision="15" seconds="11" value="92.96972">92°58'11&quot;E</geoCoordinate>
;
<collectorName id="2685531FAB4AA31B1548FE35FA31FDE8" box="[1301,1436,510,535]" pageId="1" pageNumber="127">Lalramliana</collectorName>
&amp;
<collectorName id="2685531FAB4AA31B10EBFDE9FEF9FDC4" box="[182,340,546,571]" pageId="1" pageNumber="127">S Lalronunga</collectorName>
,
<date id="FFCE1009AB4AA31B113CFDE9FDEEFDC4" box="[353,579,546,571]" pageId="1" pageNumber="127" value="2012-11-15">
<collectingDate id="EF8AE9E1AB4AA31B113CFDE9FDEEFDC4" box="[353,579,546,571]" pageId="1" pageNumber="127" value="2012-11-15">15 November 2012</collectingDate>
</date>
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.
<materialsCitation id="3B183C94AB4AA31B120DFDE9FBA4FDA0" ID-GBIF-Occurrence="3339716304" collectingDate="2015-11-24" collectionCode="PUCMF" collectorName="Lalramliana" country="India" latitude="22.397223" location="Kawlchaw River" longLatPrecision="21" longitude="92.96972" municipality="Saiha District" pageId="1" pageNumber="127" specimenCode="PUCMF 15042" specimenCount="1" stateProvince="Mizoram" typeStatus="paratype">
<specimenCode id="DBD69EB2AB4AA31B120DFDE9FD53FDC4" box="[592,766,546,571]" collectionCode="PUCMF" pageId="1" pageNumber="127">PUCMF 15042</specimenCode>
(2),
<quantity id="4C889B2CAB4AA31B1364FDE9FC56FDC5" box="[825,1019,546,571]" metricMagnitude="-1" metricUnit="m" metricValue="1.3735" metricValueMax="1.399" metricValueMin="1.348" pageId="1" pageNumber="127" unit="mm" value="137.35" valueMax="139.9" valueMin="134.8">134.8–139.9 mm</quantity>
SL;
<collectingCountry id="F3677659AB4AA31B146AFDE9FBD8FDC4" box="[1079,1141,546,571]" name="India" pageId="1" pageNumber="127">India</collectingCountry>
:
<collectingRegion id="49B4F82BAB4AA31B14DFFDE9FB41FDC4" box="[1154,1260,546,571]" country="India" name="Mizoram" pageId="1" pageNumber="127">Mizoram</collectingRegion>
,
<collectingMunicipality id="6BABACB3AB4AA31B14A4FDE9FA35FDC4" box="[1273,1432,546,571]" pageId="1" pageNumber="127">Saiha District</collectingMunicipality>
,
<location id="8EAF6012AB4AA31B10CAFD8DFEF9FDA0" LSID="urn:lsid:plazi:treatment:03D987DFAB4AA31810CAFF5CFE87F824:8EAF6012AB4AA31B10CAFD8DFEF9FDA0" box="[151,340,582,607]" country="India" latitude="22.397223" longLatPrecision="21" longitude="92.96972" municipality="Saiha District" name="Kawlchaw River" pageId="1" pageNumber="127" stateProvince="Mizoram">Kawlchaw River</location>
at
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<geoCoordinate id="EE44500EAB4AA31B1258FD8DFD25FDA0" box="[517,648,582,607]" degrees="92" direction="east" minutes="58" orientation="longitude" pageId="1" pageNumber="127" precision="15" seconds="11" value="92.96972">92°58'11&quot;E</geoCoordinate>
;
<collectorName id="2685531FAB4AA31B12C8FD8DFCB2FDA0" box="[661,799,582,607]" pageId="1" pageNumber="127">Lalramliana</collectorName>
,
<date id="FFCE1009AB4AA31B1376FD8DFBA4FDA0" box="[811,1033,582,607]" pageId="1" pageNumber="127" value="2015-11-24">
<collectingDate id="EF8AE9E1AB4AA31B1376FD8DFBA4FDA0" box="[811,1033,582,607]" pageId="1" pageNumber="127" value="2015-11-24">24 November 2015</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B183C94AB4AA31B1449FD8DFE95FD58" ID-GBIF-Occurrence="3339716307" collectingDate="2016-03-02" collectionCode="PUCMF" collectorName="Lalramliana" country="India" county="Saiha District" latitude="22.195555" location="Khopai village" longLatPrecision="21" longitude="93.024445" municipality="Tuisi River" pageId="1" pageNumber="127" specimenCode="PUCMF 16020" specimenCount="1" stateProvince="Mizoram" typeStatus="paratype">
<specimenCode id="DBD69EB2AB4AA31B1449FD8DFB6FFDA0" box="[1044,1218,582,607]" collectionCode="PUCMF" pageId="1" pageNumber="127">PUCMF 16020</specimenCode>
(1),
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SL;
<collectingCountry id="F3677659AB4AA31B10CAFDA1FF78FD7C" box="[151,213,618,643]" name="India" pageId="1" pageNumber="127">India</collectingCountry>
:
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,
<collectingCounty id="62AE4E45AB4AA31B1109FDA1FE5DFD7C" box="[340,496,618,643]" pageId="1" pageNumber="127">Saiha District</collectingCounty>
,
<collectingMunicipality id="6BABACB3AB4AA31B11A6FDA1FDD4FD7C" box="[507,633,618,643]" pageId="1" pageNumber="127">Tuisi River</collectingMunicipality>
in the vicinity of
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,
<geoCoordinate id="EE44500EAB4AA31B13A8FDA1FBDAFD7C" box="[1013,1143,618,643]" degrees="22" direction="north" minutes="11" orientation="latitude" pageId="1" pageNumber="127" precision="15" seconds="44" value="22.195555">22°11'44&quot;N</geoCoordinate>
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,
<collectorName id="2685531FAB4AA31B1553FDA1FA35FD7C" box="[1294,1432,618,643]" pageId="1" pageNumber="127">Lalramliana</collectorName>
,
<date id="FFCE1009AB4AA31B10CAFD45FE95FD58" box="[151,312,654,679]" pageId="1" pageNumber="127" value="2016-03-02">
<collectingDate id="EF8AE9E1AB4AA31B10CAFD45FE95FD58" box="[151,312,654,679]" pageId="1" pageNumber="127" value="2016-03-02">2 March 2016</collectingDate>
</date>
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.
</paragraph>
</subSubSection>
<caption id="DF0F6641AB4AA31B10CAF8BFFB99F875" ID-DOI="http://doi.org/10.5281/zenodo.5297418" ID-Zenodo-Dep="5297418" box="[151,1076,1908,1930]" httpUri="https://zenodo.org/record/5297418/files/figure.png" pageId="1" pageNumber="127" startId="1.[151,250,1908,1930]" targetBox="[151,1436,734,1883]" targetPageId="1">
<paragraph id="8BCF36C9AB4AA31B10CAF8BFFB99F875" blockId="1.[151,1076,1908,1930]" box="[151,1076,1908,1930]" pageId="1" pageNumber="127">
<emphasis id="B904EADBAB4AA31B10CAF8BFFEBEF876" bold="true" box="[151,275,1908,1930]" pageId="1" pageNumber="127">FIGURE 1.</emphasis>
<taxonomicName id="4C704D4AAB4AA31B1144F8BFFE4BF875" authorityName="Ng &amp; Lalronunga" authorityYear="2018" box="[281,486,1908,1930]" class="Actinopterygii" family="Siluridae" genus="Pterocryptis" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="subrisa">
<emphasis id="B904EADBAB4AA31B1144F8BFFE4BF875" box="[281,486,1908,1930]" italics="true" pageId="1" pageNumber="127">Pterocryptis subrisa</emphasis>
</taxonomicName>
, holotype, ZSI FF 7870, 91.8 mm SL; India: Maisa River.
</paragraph>
</caption>
<subSubSection id="C36A6542AB4AA318109AF87DFC14FE77" lastPageId="2" lastPageNumber="128" pageId="1" pageNumber="127" type="diagnosis">
<paragraph id="8BCF36C9AB4AA318109AF87DFC14FE77" blockId="1.[151,1440,1974,2035]" lastBlockId="2.[151,1437,151,2011]" lastPageId="2" lastPageNumber="128" pageId="1" pageNumber="127">
<emphasis id="B904EADBAB4AA31B109AF87DFEECF830" bold="true" box="[199,321,1974,1999]" inLineHeading="true" pageId="1" pageNumber="127" reason="1">Diagnosis.</emphasis>
<taxonomicName id="4C704D4AAB4AA31B1115F87CFD81F831" authorityName="Ng &amp; Lalronunga" authorityYear="2018" box="[328,556,1974,1998]" class="Actinopterygii" family="Siluridae" genus="Pterocryptis" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="subrisa">
<emphasis id="B904EADBAB4AA31B1115F87CFD81F831" box="[328,556,1974,1998]" italics="true" pageId="1" pageNumber="127">Pterocryptis subrisa</emphasis>
</taxonomicName>
can be distinguished from all congeners except
<taxonomicName id="4C704D4AAB4AA31B1415F87CFB70F831" box="[1096,1245,1974,1998]" class="Actinopterygii" family="Siluridae" genus="Pterocryptis" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="barakensis">
<emphasis id="B904EADBAB4AA31B1415F87CFB70F831" box="[1096,1245,1974,1998]" italics="true" pageId="1" pageNumber="127">P. barakensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C704D4AAB4AA31B14B6F87CFAD7F831" box="[1259,1402,1974,1998]" class="Actinopterygii" family="Siluridae" genus="Pterocryptis" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="berdmorei">
<emphasis id="B904EADBAB4AA31B14B6F87CFAD7F831" box="[1259,1402,1974,1998]" italics="true" pageId="1" pageNumber="127">P. berdmorei</emphasis>
</taxonomicName>
,
<taxonomicName id="4C704D4AAB4AA31B15DAF87CFEABF80D" class="Actinopterygii" family="Siluridae" genus="Pterocryptis" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="gangelica">
<emphasis id="B904EADBAB4AA31B15DAF87CFEABF80D" italics="true" pageId="1" pageNumber="127">P. gangelica</emphasis>
</taxonomicName>
and
<taxonomicName id="4C704D4AAB4AA31B1160F810FE32F80D" authorityName=": ZSI FF" authorityYear="1699" box="[317,415,2010,2034]" class="Actinopterygii" family="Siluridae" genus="Pterocryptis" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="indica">
<emphasis id="B904EADBAB4AA31B1160F810FE32F80D" box="[317,415,2010,2034]" italics="true" pageId="1" pageNumber="127">P. indica</emphasis>
</taxonomicName>
in having fewer dorsal-fin rays (2 vs. 4–5). It differs from
<taxonomicName id="4C704D4AAB4AA31B1464F810FB63F80D" box="[1081,1230,2010,2034]" class="Actinopterygii" family="Siluridae" genus="Pterocryptis" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="barakensis">
<emphasis id="B904EADBAB4AA31B1464F810FB63F80D" box="[1081,1230,2010,2034]" italics="true" pageId="1" pageNumber="127">P. barakensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C704D4AAB4AA31B1480F810FAC1F80D" box="[1245,1388,2010,2034]" class="Actinopterygii" family="Siluridae" genus="Pterocryptis" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="127" phylum="Chordata" rank="species" species="berdmorei">
<emphasis id="B904EADBAB4AA31B1480F810FAC1F80D" box="[1245,1388,2010,2034]" italics="true" pageId="1" pageNumber="127">P. berdmorei</emphasis>
</taxonomicName>
and
<taxonomicName id="4C704D4AAB49A31810CAFF52FE8CFF4F" box="[151,289,152,176]" class="Actinopterygii" family="Siluridae" genus="Pterocryptis" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="128" phylum="Chordata" rank="species" species="gangelica">
<emphasis id="B904EADBAB49A31810CAFF52FE8CFF4F" box="[151,289,152,176]" italics="true" pageId="2" pageNumber="128">P. gangelica</emphasis>
</taxonomicName>
in having a deeper head (10.6–11.9% SL vs. 8.3–10.8), from
<taxonomicName id="4C704D4AAB49A318138BFF52FBC6FF4F" box="[982,1131,152,176]" class="Actinopterygii" family="Siluridae" genus="Pterocryptis" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="128" phylum="Chordata" rank="species" species="barakensis">
<emphasis id="B904EADBAB49A318138BFF52FBC6FF4F" box="[982,1131,152,176]" italics="true" pageId="2" pageNumber="128">P. barakensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C704D4AAB49A31814FFFF52FA81FF4F" box="[1186,1324,152,176]" class="Actinopterygii" family="Siluridae" genus="Pterocryptis" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="128" phylum="Chordata" rank="species" species="gangelica">
<emphasis id="B904EADBAB49A31814FFFF52FA81FF4F" box="[1186,1324,152,176]" italics="true" pageId="2" pageNumber="128">P. gangelica</emphasis>
</taxonomicName>
in having a longer supralabial fold (extending posteriorly beyond vertical through posterior orbital margin vs. not or just reaching vertical through posterior orbital margin;
<figureCitation id="134B2A4CAB49A318128CFF14FCB8FF07" box="[721,789,223,248]" captionStart="FIGURE 2" captionStartId="3.[151,250,1944,1966]" captionTargetBox="[193,1398,195,1927]" captionTargetId="figure-75@3.[193,1398,195,1927]" captionTargetPageId="3" captionText="FIGURE 2. Lateral views of heads of: a. P.subrisa, PUCMF 18011, paratype, 93.6 mm SL; b. P. gangelica, ZRC 60007, 92.8 mm SL showing posterior extent of supralabial fold (indicated with black arrows) in relation to the posterior orbital margin (indicated with white dashed line). Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.5297420" httpUri="https://zenodo.org/record/5297420/files/figure.png" pageId="2" pageNumber="128">Fig. 2</figureCitation>
), and from
<taxonomicName id="4C704D4AAB49A31813C5FF2AFC57FF07" authorityName=": ZSI FF" authorityYear="1699" box="[920,1018,224,248]" class="Actinopterygii" family="Siluridae" genus="Pterocryptis" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="128" phylum="Chordata" rank="species" species="indica">
<emphasis id="B904EADBAB49A31813C5FF2AFC57FF07" box="[920,1018,224,248]" italics="true" pageId="2" pageNumber="128">P. indica</emphasis>
</taxonomicName>
in having fewer anal-fin rays (66–75 vs. 85). It is additionally diagnosed from congeners in having the following unique combination of characters: nearly circular eye; head length 17.6–19.6% SL; dorsal-fin height 2.6–4.7% SL; pectoral-fin length 11.8–14.0% SL; body depth at anus 14.4–16.7% SL; caudal peduncle depth 6.8–8.5% SL; confluent anal and caudal fins separated by deep notch; 17 principal caudal-fin rays; and 57 vertebrae.
</paragraph>
</subSubSection>
<subSubSection id="C36A6542AB49A318109AFE58FB08F91F" pageId="2" pageNumber="128" type="description">
<paragraph id="8BCF36C9AB49A318109AFE58FEADFD9F" blockId="2.[151,1437,151,2011]" pageId="2" pageNumber="128">
<emphasis id="B904EADBAB49A318109AFE58FEF5FE53" bold="true" box="[199,344,403,428]" inLineHeading="true" pageId="2" pageNumber="128" reason="1">Description.</emphasis>
Morphometric data is given in
<tableCitation id="C6F20372AB49A31812B3FE5FFCE7FE52" box="[750,842,404,429]" captionStart="TABLE 1" captionStartId="5.[151,239,476,499]" captionText="TABLE 1. Morphometric data for Pterocryptis subrisa (n=9)" pageId="2" pageNumber="128">Table 1</tableCitation>
. Body subcylindrical in preanal region, laterally compressed posteriorly. Head moderately depressed. Dorsal profile straight, descending gently from dorsal-fin origin to snout tip. Anterior profile of snout rounded. Anterior pair of nostrils tubular and anteromedial to maxillary barbel base. Posterior pair of nostrils bordered by fleshy dorsal and ventral membranes and situated posteromedial to maxillary barbel base. Eyes small, subcutaneous; located in anterior half of head; visible dorsally, but not ventrally.
</paragraph>
<paragraph id="8BCF36C9AB49A318109AFDA7FD45FD0F" blockId="2.[151,1437,151,2011]" pageId="2" pageNumber="128">Mouth subterminal; gape horizontal. Well-developed rictal fold present, consisting of large and fleshy upper lobe joined at corner of mouth with lower lobe; lower lobe subtended by short submandibular groove. Poorlydeveloped supralabial fold present, extending from posterodorsal corner of upper rictal lobe to immediately posterior to vertical through posterior orbital margin.</paragraph>
<paragraph id="8BCF36C9AB49A318109AFD37FDEBFCA2" blockId="2.[151,1437,151,2011]" pageId="2" pageNumber="128">Teeth villiform. Dentary teeth in slightly curved, elongate bands narrowing posteriorly, reaching from symphysis almost to mouth corners; premaxillary teeth in broader, slightly curved rectangular bands; vomerine teeth in a single crescent-shaped band.</paragraph>
<paragraph id="8BCF36C9AB49A318109AFCACFEF7FC37" blockId="2.[151,1437,151,2011]" pageId="2" pageNumber="128">Maxillary barbels slightly flattened, reaching to anterior third of anal fin. One pair of mandibular barbels present; located slightly anterolateral to gular fold; barbels flattened for most of length, reaching to middle of pectoral-fin base.</paragraph>
<paragraph id="8BCF36C9AB49A318109AFC1FFF13FBCA" blockId="2.[151,1437,151,2011]" pageId="2" pageNumber="128">Gill membranes separate and overlapping, free from isthmus. Branchiostegal rays 11* (4) or 12 (5). Gill rakers short, anteriormost rakers on lower first arch small and widely spaced; 2+3 (1), 2+4* (3), 2+5 (2), 2+6 (1) or 3+5 (2).</paragraph>
<paragraph id="8BCF36C9AB49A318109AFBF4FD00FA3F" blockId="2.[151,1437,151,2011]" pageId="2" pageNumber="128">Distal margin of dorsal fin pointed, with i, 1 (8) rays; segments of first ray not co-ossified to form spine. Distal margin of pectoral fin broadly convex, with 12* (7), 13 (1) or 14 (1) rays. Segments of the proximal two-thirds of first pectoral-fin element co-ossified, forming spine. Pectoral spine and articulated segments sexually dimorphic in mature individuals. Spine in males with broad and somewhat flattened dorsoventrally, with 0–8 serrations on posterior edge, increasing in size distally; proximal articulated segments with 0–1 serrations on posterior edge. Spine in females or juveniles slender, without serrations on posterior edges of spine proper and proximal articulated segments. Distal margin of pelvic fin convex, with i,7,i (9) rays. Distal margin of anal fin straight, with 66 (1), 68* (3), 69 (1), 72 (1), 73 (1) 74 (1) or 75 (1) rays; joined to caudal fin for length of last anal-fin ray. Integument over anal fin thickened proximally for slightly more than half of ray lengths; fin-ray erector muscles extending along anterior edges of anal-fin rays, ventral most extent of muscles that of thickened integument. Caudal fin emarginate; principal rays i,7,8,i (9).</paragraph>
<paragraph id="8BCF36C9AB49A318109AFA07FC84F9F7" blockId="2.[151,1437,151,2011]" pageId="2" pageNumber="128">Urogenital papillae of both sexes located immediately posterior to anus. Males with a conical papilla; females with a similar shaped, but smaller, slightly broader papilla.</paragraph>
<paragraph id="8BCF36C9AB49A318109AF9DFFD4FF9D2" blockId="2.[151,1437,151,2011]" box="[199,738,1556,1581]" pageId="2" pageNumber="128">Vertebrae 15 + 42 = 57* (5) or 16 + 41 = 57 (1).</paragraph>
<paragraph id="8BCF36C9AB49A318109AF9FCFB08F91F" blockId="2.[151,1437,151,2011]" pageId="2" pageNumber="128">
<emphasis id="B904EADBAB49A318109AF9FCFEE2F9AF" bold="true" box="[199,335,1591,1616]" inLineHeading="true" pageId="2" pageNumber="128" reason="1">Coloration.</emphasis>
In 70% ethanol: Dorsal surface and sides of head uniformly brown flanks and thickened integument over anal fin pale brown, with faint mottled pattern. Ventral surfaces of head, breast and belly yellow with scattered melanophores; melanophores less dense ventral to level of eye. Maxillary and mandibular barbels brown, fading to yellow distally. Anal fin with hyaline ventral margin. Caudal, pectoral and pelvic fins with dusky fin rays and hyaline fin membrane. Color in life similar, with a more yellowish tinge overall.
</paragraph>
</subSubSection>
<subSubSection id="C36A6542AB49A318109AF920FB34F890" pageId="2" pageNumber="128" type="distribution">
<paragraph id="8BCF36C9AB49A318109AF920FA84F8FA" blockId="2.[151,1437,151,2011]" box="[199,1321,1771,1797]" pageId="2" pageNumber="128">
<emphasis id="B904EADBAB49A318109AF920FECDF8FB" bold="true" box="[199,352,1771,1796]" inLineHeading="true" pageId="2" pageNumber="128" reason="1">Distribution.</emphasis>
This species is known from the Kaladan River drainage in southern
<collectingRegion id="49B4F82BAB49A318143DF927FB6BF8FA" box="[1120,1222,1772,1797]" country="India" name="Mizoram" pageId="2" pageNumber="128">Mizoram</collectingRegion>
(
<figureCitation id="134B2A4CAB49A318148BF927FAB6F8FA" box="[1238,1307,1772,1797]" captionStart="FIGURE 3" captionStartId="4.[151,250,1117,1139]" captionTargetBox="[152,1435,193,1084]" captionTargetId="figure-415@4.[151,1436,193,1096]" captionTargetPageId="4" captionText="FIGURE 3. Map showing collection localities of Pterocryptis subrisa. Open symbol represents type locality." figureDoi="http://doi.org/10.5281/zenodo.5297422" httpUri="https://zenodo.org/record/5297422/files/figure.png" pageId="2" pageNumber="128">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8BCF36C9AB49A318109AF8C4FB34F890" blockId="2.[151,1437,151,2011]" pageId="2" pageNumber="128">
<emphasis id="B904EADBAB49A318109AF8C4FE87F8D7" bold="true" box="[199,298,1807,1832]" inLineHeading="true" pageId="2" pageNumber="128" reason="1">Habitat.</emphasis>
<taxonomicName id="4C704D4AAB49A3181168F8DBFDB1F8D7" authorityName="Ng &amp; Lalronunga" authorityYear="2018" box="[309,540,1808,1832]" class="Actinopterygii" family="Siluridae" genus="Pterocryptis" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="128" phylum="Chordata" rank="species" species="subrisa">
<emphasis id="B904EADBAB49A3181168F8DBFDB1F8D7" box="[309,540,1808,1832]" italics="true" pageId="2" pageNumber="128">Pterocryptis subrisa</emphasis>
</taxonomicName>
has been collected from moderately flowing streams with vegetation on the bank, and a substrate of pebble, sand, gravel and small boulders. The water temperature ranging from 20 to 25 °C and the species is found syntopically with
<taxonomicName id="4C704D4AAB49A318122FF893FD40F890" box="[626,749,1880,1903]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="128" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B904EADBAB49A318122FF893FD15F890" box="[626,696,1880,1903]" italics="true" pageId="2" pageNumber="128">Garra</emphasis>
spp.
</taxonomicName>
<taxonomicName id="4C704D4AAB49A31812A9F893FCF6F890" box="[756,859,1880,1904]" class="Actinopterygii" family="Badidae" genus="Badis" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="128" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B904EADBAB49A31812A9F893FC99F88F" box="[756,820,1880,1904]" italics="true" pageId="2" pageNumber="128">Badis</emphasis>
sp.
</taxonomicName>
and
<taxonomicName id="4C704D4AAB49A31813CCF893FB3EF890" baseAuthorityName="McClelland" baseAuthorityYear="1839" box="[913,1171,1880,1903]" class="Actinopterygii" family="Cyprinidae" genus="Devario" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="128" phylum="Chordata" rank="species" species="aequipinnatus">
<emphasis id="B904EADBAB49A31813CCF893FB3EF890" box="[913,1171,1880,1903]" italics="true" pageId="2" pageNumber="128">Devario aequipinnatus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C36A6542AB49A318109AF8B7FE87F824" pageId="2" pageNumber="128" type="etymology">
<paragraph id="8BCF36C9AB49A318109AF8B7FE87F824" blockId="2.[151,1437,151,2011]" pageId="2" pageNumber="128">
<emphasis id="B904EADBAB49A318109AF8B7FEE3F86B" bold="true" box="[199,334,1916,1940]" inLineHeading="true" pageId="2" pageNumber="128" reason="1">Etymology.</emphasis>
The specific epithet is the perfect passive participle of the Latin verb subrideo, meaning to smile. The name is used in allusion to the longer supralabial fold in this species, which vaguely resembles a smiling countenance.
</paragraph>
</subSubSection>
</treatment>
</document>
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