1026 lines
217 KiB
XML
1026 lines
217 KiB
XML
<document id="4EA3D2F270147D298759DB9C4EF61339" ID-CLB-Dataset="22296" ID-DOI="10.5852/ejt.2018.456" ID-GBIF-Dataset="0566a11e-d6a3-4283-89c7-c7feea96fdbf" ID-Zenodo-Dep="3817823" ID-ZooBank="DE1B1DEE-9871-4803-9F67-025F2B439560" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588965964057" checkinUser="valdenar" docAuthor="Sørensen, Martin V., Rohal, Melissa & Thistle, David" docDate="2018" docId="F64287A25045FF851442FA79FE8309D0" docLanguage="en" docName="ejt-456_sorensen_rohal_thistle.pdf.imf" docOrigin="European Journal of Taxonomy 456" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Echinoderes hamiltonorum Sørensen & Rohal & Thistle 2018, sp. nov." docType="treatment" docUuid="B3EE641D-72E3-4BCA-A4A9-3AD3A9BD8C05" docUuidSource="ZooBank" docVersion="8" lastPageNumber="28" masterDocId="0A7BFFDA5056FF9E1600FFDEFFDE0C64" masterDocTitle="Deep-sea Echinoderidae (Kinorhyncha: Cyclorhagida) from the Northwest Pacific" masterLastPageNumber="75" masterPageNumber="1" pageNumber="20" updateTime="1698839057356" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="301961F7EAF9AAA38E0F1296C00C88FE">
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<mods:title id="7C1B163E4A3240C2926006CE1A2A12B3">Deep-sea Echinoderidae (Kinorhyncha: Cyclorhagida) from the Northwest Pacific</mods:title>
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<mods:name id="442A2EF9F3A211D843B99FC4023BB1EE" type="personal">
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<mods:namePart id="39B2DA43A43BC9CF6A0EF05E8F5FECFF">Sørensen, Martin V.</mods:namePart>
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<mods:nameIdentifier id="8003CDDE19F25B06362423DEEEEC27F1" type="LSID">4143D650-12FC-4914-93F5-2C39339A7156</mods:nameIdentifier>
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<mods:affiliation id="65A09B6C503584D13A670EF356076964">Section for GeoGenetics, Natural History Museum of Denmark, University of Copenhagen, DK- 1350 Copenhagen, Denmark. Harte Research Institute, Texas A & M University - Corpus Christi, Corpus Christi, TX 78412, USA. Department of Earth, Ocean and Atmospheric Sciences, Florida State University, Tallahassee, FL 32306 - 4520, USA.</mods:affiliation>
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<mods:nameIdentifier id="1664069A856C02E5E916FC4D89599892" type="email">mvsorensen@snm.ku.dk</mods:nameIdentifier>
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<mods:namePart id="604A43C06B4FA2475F1AE6FDEF05ADFB">Rohal, Melissa</mods:namePart>
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<mods:nameIdentifier id="C8C171A14E9AF8F67EEE8ADFC0B16A74" type="email">mrohal@islander.tamucc.edu</mods:nameIdentifier>
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<mods:namePart id="098926F0B63EB98D5B967C28A8BCE17B">Thistle, David</mods:namePart>
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<mods:nameIdentifier id="34AB5287E6FD5096A1107E61705EA45D" type="email">dthistle@fsu.edu</mods:nameIdentifier>
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<mods:title id="53CFFFB4C086F3161132719248FBDE0D">European Journal of Taxonomy</mods:title>
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<mods:part id="1252A2E4CDDDA6FEC3E9E3B8183BB3FA">
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<mods:date id="FABE188F7729826B93EA1B7B1F58C7ED">2018</mods:date>
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<mods:number id="7D0084E3C84161E61932BC08128C0C73">2018-08-17</mods:number>
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<mods:number id="EE8FAAD38A1DF83CC9EF9B83CB1D3B0A">456</mods:number>
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<treatment id="F64287A25045FF851442FA79FE8309D0" ID-DOI="http://doi.org/10.5281/zenodo.3818838" ID-GBIF-Taxon="163993852" ID-Zenodo-Dep="3818838" LSID="urn:lsid:zoobank.org:act:B3EE641D-72E3-4BCA-A4A9-3AD3A9BD8C05" httpUri="http://treatment.plazi.org/id/F64287A25045FF851442FA79FE8309D0" lastPageId="27" lastPageNumber="28" pageId="19" pageNumber="20">
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<subSubSection id="36F1653F5045FF8D1442FA79FB08098D" pageId="19" pageNumber="20" type="nomenclature">
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<paragraph id="7E5436B45045FF8D1442FA79FC2C09A6" blockId="19.[350,1238,1447,1552]" box="[578,1010,1447,1474]" pageId="19" pageNumber="20">
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<heading id="251C81D85045FF8D1442FA79FC2C09A6" box="[578,1010,1447,1474]" centered="true" fontSize="11" level="2" pageId="19" pageNumber="20" reason="2">
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<taxonomicName id="B9EB4D375045FF8D1442FA79FC5109A6" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[578,911,1447,1474]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="19" pageNumber="20" phylum="Cephalorhyncha" rank="species" species="hamiltonorum" status="sp. nov.">
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<emphasis id="4C9FEAA65045FF8D1442FA79FC5109A6" bold="true" box="[578,911,1447,1474]" italics="true" pageId="19" pageNumber="20">Echinoderes hamiltonorum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="57AC57DD5045FF8D1597FA76FC2C09A6" box="[919,1010,1448,1474]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<paragraph id="7E5436B45045FF8D175EFA10FB08098D" blockId="19.[350,1238,1447,1552]" box="[350,1238,1486,1513]" pageId="19" pageNumber="20">
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<uri id="0A7A3AB65045FF8D175EFA10FB08098D" box="[350,1238,1486,1513]" pageId="19" pageNumber="20">
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urn:lsid:zoobank.org:act:
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<uuid id="0A4D0C615045FF8D147FFA10FB08098D" box="[639,1238,1486,1513]" pageId="19" pageNumber="20">B3EE641D-72E3-4BCA-A4A9-3AD3A9BD8C05</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="36F1653F5045FF8D14A0FA28FC4A0A74" box="[672,916,1526,1552]" pageId="19" pageNumber="20" type="description">
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<paragraph id="7E5436B45045FF8D14A0FA28FC4A0A74" blockId="19.[350,1238,1447,1552]" box="[672,916,1526,1552]" pageId="19" pageNumber="20">
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<figureCitation id="E6D02A315045FF8D14A0FA28FCD70A74" box="[672,777,1526,1552]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="21.[189,232,1753,1779]" captionStartId-1="22.[189,232,1611,1637]" captionStartId-2="23.[189,232,1508,1534]" captionTargetBox-0="[315,1235,269,1735]" captionTargetBox-1="[256,1332,265,1599]" captionTargetId-0="figure@21.[315,1273,265,1735]" captionTargetId-1="figure@22.[256,1332,265,1599]" captionTargetId-2="figure@24.[189,1398,260,2026]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionTargetPageId-2="24" captionText-0="Fig. 7. Line art illustrations of Echinoderes hamiltonorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♂, segments 10 to 11, dorsal view. D. ♂, segments 10 to 11, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." captionText-1="Fig. 8. Light micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov., holotype, ♀ (A–D, H) (NHMD-223916), paratype, ♂ (E–F) (NHMD-223918) and paratype, ♀ (G) (NHMD-223920). A. Ventral overview. B. Segments 1 to 5, dorsal view. C. Segments 1 to 5, ventral view. D. Segments 6 to 8, dorsal view. E. Segments 6 to 9, ventral view. F. Detail showing sublateral glandular cell outlets type 2 on segments 1 and 2. G. Segments 8 to 9, lateral view. H. Segments 10 to 11 in female, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; pdgco1 = paradorsal glandular cell outlet type 1; pdss = paradorsal sensory spot; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." captionText-2="Fig. 9 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov. A. Lateral overview. B. Ventral overview. C. Mouth cone. D. Introvert sector 8. E. Segments 1 to 3, dorsolateral view; note that laterodorsal sensory spots are missing in this particular specimen. F. Segments 1 to 3, ventral view. G. Detail showing lateral side of segment 8 without any indication of glandular cell outlets type 2. H. Segments 8 to 9, lateral view. I. Detail showing midlateral to lateroventral parts of segments 5 and 6. J. ♂, segments 10 to 11, dorsolateral view. K. ♂, segments 10 to 11, ventral view. L. ♀, segments 10 to 11, laterodorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdss = middorsal sensory spot; mlss = midlateral sensory spot; oos = outer oral style; pdss = paradorsal sensory spot; pe = penile spines; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlgco1 = ventrolateral glandular cell outlet type 1; vlss = ventrolateral sensory spot; vlt = ventrolateral tube; vmss = ventromedial sensory spot." figureDoi-0="http://doi.org/10.5281/zenodo.3817837" figureDoi-1="http://doi.org/10.5281/zenodo.3817839" figureDoi-2="http://doi.org/10.5281/zenodo.3817841" httpUri-0="https://zenodo.org/record/3817837/files/figure.png" httpUri-1="https://zenodo.org/record/3817839/files/figure.png" httpUri-2="https://zenodo.org/record/3817841/files/figure.png" pageId="19" pageNumber="20">Figs 7–9</figureCitation>
|
||
,
|
||
<tableCitation id="3369030F5045FF8D1514FA28FC4A0A74" box="[788,916,1526,1552]" captionStart-0="Table 6" captionStart-1="Table 7" captionStartId-0="23.[189,258,268,294]" captionStartId-1="25.[189,258,268,294]" captionTargetPageId-0="23" captionTargetPageId-1="25" captionText-0="Table 6. Measurements from light microscopy of Echinoderes hamiltonorum sp. nov. (in µm), including number of measured specimens (n) and standard deviation (SD). Abbreviations: (ac) = acicular spine; LTAS = lateral terminal accessory spine; LTS = lateral terminal spine; LV = lateroventral; MD = middorsal; MSW-8 = maximum sternal width, measured on segment 8 in this species; S = segment lengths; SW-10 = standard width, always measured on segment 10; TL = trunk length." captionText-1="Table 7. Summary of nature and location of sensory spots, glandular cell outlets, tubes and spines arranged by series in Echinoderes hamiltonorum sp. nov. Abbreviations: LA = lateral accessory; LD = laterodorsal; LV = lateroventral; MD = middorsal; ML = midlateral; PD = paradorsal; PV = paraventral; SD = subdorsal; SL = sublateral; VL = ventrolateral; VM = ventromedial; ac = acicular spine; gco1/2 = glandular cell outlet type 1/2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; pe = penile spines; si = sieve plate; ss = sensory spot; tu = tube; (♀) = female and (♂) = male conditions of sexually dimorphic characters; * = character not present in all specimens." httpUri-1="http://table.plazi.org/id/2A94663C504FFF8716BDFED2FBC50D9D" pageId="19" pageNumber="20" tableUuid-1="2A94663C504FFF8716BDFED2FBC50D9D">Tables 6–7</tableCitation>
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</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="36F1653F5045FF8D16BDF9E5FB6A0B2F" pageId="19" pageNumber="20" type="diagnosis">
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||
<paragraph id="7E5436B45045FF8D16BDF9E5FEE80A31" blockId="19.[189,310,1595,1621]" box="[189,310,1595,1621]" pageId="19" pageNumber="20">
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<heading id="251C81D85045FF8D16BDF9E5FEE80A31" bold="true" box="[189,310,1595,1621]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
|
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<emphasis id="4C9FEAA65045FF8D16BDF9E5FEE80A31" bold="true" box="[189,310,1595,1621]" pageId="19" pageNumber="20">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="7E5436B45045FF8D16BDF9B0FB6A0B2F" blockId="19.[189,1399,1646,1868]" pageId="19" pageNumber="20">
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<taxonomicName id="B9EB4D375045FF8D16BDF9B0FE910AEC" authorityName="Claparede" authorityYear="1863" box="[189,335,1646,1672]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="19" pageNumber="20" phylum="Cephalorhyncha" rank="genus">
|
||
<emphasis id="4C9FEAA65045FF8D16BDF9B0FE910AEC" box="[189,335,1646,1672]" italics="true" pageId="19" pageNumber="20">Echinoderes</emphasis>
|
||
</taxonomicName>
|
||
with middorsal spines on segments 4, 6 and 8, and spines in lateroventral positions on segments 6 to 9. Tubes present in ventrolateral positions on segment 2, in lateroventral positions on segment 5, and in laterodorsal positions on segment 10. Minute glandular cell outlets
|
||
<typeStatus id="A15088165045FF8D12B2F963FB3B0AB3" box="[1202,1253,1725,1751]" pageId="19" pageNumber="20">type</typeStatus>
|
||
2 present in sublateral positions on segment 1, and in subdorsal, laterodorsal and sublateral positions on segment 2. Larger glandular cell outlets
|
||
<typeStatus id="A15088165045FF8D1431F8D5FDBA0B41" box="[561,612,1803,1829]" pageId="19" pageNumber="20">type</typeStatus>
|
||
2 may or may not be present in sublateral positions on segment 8. Males with three pairs of penile spines; females with lateral terminal accessory spines.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="36F1653F5045FF8D16BDF8A9FB110B8F" pageId="19" pageNumber="20" type="etymology">
|
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<paragraph id="7E5436B45045FF8D16BDF8A9FE9A0BF5" blockId="19.[189,324,1911,1937]" box="[189,324,1911,1937]" pageId="19" pageNumber="20">
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<heading id="251C81D85045FF8D16BDF8A9FE9A0BF5" bold="true" box="[189,324,1911,1937]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
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<emphasis id="4C9FEAA65045FF8D16BDF8A9FE9A0BF5" bold="true" box="[189,324,1911,1937]" pageId="19" pageNumber="20">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="7E5436B45045FF8D16BDF877FB110B8F" blockId="19.[189,1398,1961,2027]" pageId="19" pageNumber="20">The second author (MR) dedicates this species to her mother and the Hamiltons, i.e., her mother’s side of the family, for always being there, supporting her and helping her achieve her dreams.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="36F1653F5042FF8A16BDFED5FB040E99" pageId="20" pageNumber="21" type="materials_examined">
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<paragraph id="7E5436B45042FF8A16BDFED5FE740D41" blockId="20.[189,426,267,293]" box="[189,426,267,293]" pageId="20" pageNumber="21">
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<heading id="251C81D85042FF8A16BDFED5FE740D41" bold="true" box="[189,426,267,293]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
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||
<emphasis id="4C9FEAA65042FF8A16BDFED5FE740D41" bold="true" box="[189,426,267,293]" pageId="20" pageNumber="21">Material examined</emphasis>
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</heading>
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</paragraph>
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<materialsCitation id="CE833CE95042FF8A16EAFEE5FB010DA6" ID-GBIF-Occurrence="2611490303" collectingDate="2008-09-23" collectionCode="US" collectorName="St. & Fluoromount G & See" country="United States of America" latitude="36.796112" location="Monterey" longLatPrecision="19" longitude="-123.69833" municipality="West Coast" pageId="20" pageNumber="21" specimenCount="1" specimenCount-female="1" stateProvince="California" typeStatus="holotype">
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<paragraph id="7E5436B45042FF8A16EAFEE5FE820D31" blockId="20.[189,1399,315,451]" box="[234,348,315,341]" pageId="20" pageNumber="21">
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<emphasis id="4C9FEAA65042FF8A16EAFEE5FE820D31" bold="true" box="[234,348,315,341]" pageId="20" pageNumber="21">
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<typeStatus id="A15088165042FF8A16EAFEE5FE820D31" box="[234,348,315,341]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
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</emphasis>
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</paragraph>
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<paragraph id="7E5436B45042FF8A16BDFEBEFB010DA6" blockId="20.[189,1399,315,451]" pageId="20" pageNumber="21">
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<collectingCountry id="06FC76245042FF8A16BDFEBEFD920D1D" box="[189,588,351,378]" name="United States of America" pageId="20" pageNumber="21">UNITED STATES OF AMERICA</collectingCountry>
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:adult
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<specimenCount id="68EDFD3D5042FF8A1492FEBEFD760D1E" box="[658,680,352,378]" pageId="20" pageNumber="21" type="female">♀</specimenCount>
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,
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<collectionCode id="18FAAE715042FF8A14B3FE81FD060D1D" box="[691,728,351,377]" country="South Africa" name="University of Stellenbosch" pageId="20" pageNumber="21">US</collectionCode>
|
||
<collectingMunicipality id="9E30ACCE5042FF8A14DCFEBEFCBE0D1D" box="[732,864,351,378]" pageId="20" pageNumber="21">West Coast</collectingMunicipality>
|
||
, central
|
||
<collectingRegion id="BC2FF8565042FF8A15BCFE81FBE80D1D" box="[956,1078,351,377]" country="United States of America" name="California" pageId="20" pageNumber="21">California</collectingRegion>
|
||
, off
|
||
<location id="7B34606F5042FF8A1264FEBEFB090D1E" LSID="urn:lsid:plazi:treatment:F64287A25045FF851442FA79FE8309D0:7B34606F5042FF8A1264FEBEFB090D1E" box="[1124,1239,352,378]" country="United States of America" latitude="36.796112" longLatPrecision="19" longitude="-123.69833" municipality="West Coast" name="Monterey" pageId="20" pageNumber="21" stateProvince="California">Monterey</location>
|
||
,
|
||
<geoCoordinate id="1BDF50735042FF8A12DFFE83FAAC0D1E" box="[1247,1394,349,378]" degrees="36" direction="north" minutes="47" orientation="latitude" pageId="20" pageNumber="21" precision="15" seconds="46" value="36.796112">36°47′46″ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="1BDF50735042FF8A16BDFE5CFEBB0DFA" box="[189,357,386,414]" degrees="123" direction="west" minutes="41" orientation="longitude" pageId="20" pageNumber="21" precision="15" seconds="54" value="-123.69833">123°41′54″ W</geoCoordinate>
|
||
,
|
||
<collectorName id="D31E53625042FF8A1771FE5AFE4F0DFA" box="[369,401,388,414]" pageId="20" pageNumber="21">St.</collectorName>
|
||
5, 3679 m deep, collected from mud,
|
||
<date id="0A5510745042FF8A1551FE5AFC310DFA" box="[849,1007,388,414]" pageId="20" pageNumber="21" value="2008-09-23">
|
||
<collectingDate id="1A11E99C5042FF8A1551FE5AFC310DFA" box="[849,1007,388,414]" pageId="20" pageNumber="21" value="2008-09-23">23 Sep. 2008</collectingDate>
|
||
</date>
|
||
, mounted in
|
||
<collectorName id="D31E53625042FF8A1287FE5DFAE30DFA" box="[1159,1341,387,414]" pageId="20" pageNumber="21">Fluoromount G</collectorName>
|
||
on a glass slide (NHMD-223916).
|
||
<collectorName id="D31E53625042FF8A141BFE76FD9B0DA6" box="[539,581,424,450]" pageId="20" pageNumber="21">See</collectorName>
|
||
<figureCitation id="E6D02A315042FF8A144DFE76FD4C0DA6" box="[589,658,424,450]" captionStart="Fig" captionStartId="2.[189,232,2000,2026]" captionTargetBox="[241,1347,1019,1957]" captionTargetId="figure@2.[241,1347,1019,1958]" captionTargetPageId="2" captionText="Fig. 1. Map showing the eight sampling stations off the US West Coast. See Table 1 for further details." figureDoi="http://doi.org/10.5281/zenodo.3817825" httpUri="https://zenodo.org/record/3817825/files/figure.png" pageId="20" pageNumber="21">Fig. 1</figureCitation>
|
||
for localities and
|
||
<tableCitation id="3369030F5042FF8A1565FE76FC620DA6" box="[869,956,424,450]" captionStart="Table 1" captionStartId="3.[189,258,268,294]" captionTargetBox="[202,1399,283,441]" captionTargetPageId="4" captionText="Table 1 (continued on next page). Summary of data on stations, species identities, and catalogue numbers." httpUri="http://table.plazi.org/id/2A94663C5055FF9D16BDFED2FAA90D41" pageId="20" pageNumber="21" tableUuid="2A94663C5055FF9D16BDFED2FAA90D41">Table 1</tableCitation>
|
||
for detailed station data.
|
||
</paragraph>
|
||
</materialsCitation>
|
||
<paragraph id="7E5436B45042FF8A16EAFE35FEB90E61" blockId="20.[189,1406,491,626]" box="[234,359,491,517]" pageId="20" pageNumber="21">
|
||
<materialsCitation id="CE833CE95042FF8A16EAFE35FEB90E61" ID-GBIF-Occurrence="2611490304" box="[234,359,491,517]" pageId="20" pageNumber="21" typeStatus="paratype">
|
||
<emphasis id="4C9FEAA65042FF8A16EAFE35FEB90E61" bold="true" box="[234,359,491,517]" pageId="20" pageNumber="21">
|
||
<typeStatus id="A15088165042FF8A16EAFE35FEB90E61" box="[234,359,491,517]" pageId="20" pageNumber="21" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="7E5436B45042FF8A16BDFDD1FED60E16" blockId="20.[189,1406,491,626]" pageId="20" pageNumber="21">
|
||
<collectingCountry id="06FC76245042FF8A16BDFDD1FDBE0E4D" box="[189,608,527,553]" name="United States of America" pageId="20" pageNumber="21">UNITED STATES OF AMERICA</collectingCountry>
|
||
: 2 ♂♂, same collecting data as for
|
||
<typeStatus id="A15088165042FF8A1211FDD1FBA30E4D" box="[1041,1149,527,553]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
|
||
;
|
||
<specimenCount id="68EDFD3D5042FF8A128BFDD1FB0E0E4D" box="[1163,1232,527,553]" pageId="20" pageNumber="21" type="female">2 ♀♀</specimenCount>
|
||
, St. 2;
|
||
<specimenCount id="68EDFD3D5042FF8A132BFDD1FAA80E4D" box="[1323,1398,527,553]" pageId="20" pageNumber="21" type="female">2 ♀♀</specimenCount>
|
||
, St. 3;
|
||
<specimenCount id="68EDFD3D5042FF8A170AFDEDFEE40E2A" box="[266,314,563,590]" pageId="20" pageNumber="21" type="female">1 ♀</specimenCount>
|
||
, St. 6 (NHMD-223917–223923). All
|
||
<typeStatus id="A15088165042FF8A150CFDEAFCA00E2A" box="[780,894,564,590]" pageId="20" pageNumber="21" type="paratype">paratypes</typeStatus>
|
||
are mounted in Fluoromount G on glass slides.
|
||
</paragraph>
|
||
<paragraph id="7E5436B45042FF8A16EAFD44FD890ED0" blockId="20.[189,1398,666,765]" box="[234,599,666,693]" pageId="20" pageNumber="21">
|
||
<emphasis id="4C9FEAA65042FF8A16EAFD44FD890ED0" bold="true" box="[234,599,666,693]" pageId="20" pageNumber="21">Additional non-type material</emphasis>
|
||
</paragraph>
|
||
<paragraph id="7E5436B45042FF8A16BDFD61FB040E99" blockId="20.[189,1398,666,765]" pageId="20" pageNumber="21">
|
||
<collectingCountry id="06FC76245042FF8A16BDFD61FD870EBD" box="[189,601,703,729]" name="United States of America" pageId="20" pageNumber="21">UNITED STATES OF AMERICA</collectingCountry>
|
||
:
|
||
<specimenCount id="68EDFD3D5042FF8A1464FD61FD4F0EBD" box="[612,657,703,729]" pageId="20" pageNumber="21" type="female">1 ♀</specimenCount>
|
||
, St. 3;
|
||
<specimenCount id="68EDFD3D5042FF8A14E8FD61FCCB0EBD" box="[744,789,703,729]" pageId="20" pageNumber="21" type="female">1 ♀</specimenCount>
|
||
, 1 ♂, St. 5;
|
||
<specimenCount id="68EDFD3D5042FF8A15A8FD61FC350EBD" box="[936,1003,703,729]" pageId="20" pageNumber="21" type="female">2 ♀♀</specimenCount>
|
||
, 2 ♂♂, St. 6;
|
||
<specimenCount id="68EDFD3D5042FF8A1293FD61FB1E0EBD" box="[1171,1216,703,729]" pageId="20" pageNumber="21" type="female">1 ♀</specimenCount>
|
||
, 1 ♂, St. 8. All specimens mounted for SEM and stored in the first author’s personal reference collection.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="36F1653F5042FF8416BDFCF8FC4F0A38" lastPageId="26" lastPageNumber="27" pageId="20" pageNumber="21" type="description">
|
||
<paragraph id="7E5436B45042FF8A16BDFCF8FE900F24" blockId="20.[189,334,806,832]" box="[189,334,806,832]" pageId="20" pageNumber="21">
|
||
<heading id="251C81D85042FF8A16BDFCF8FE900F24" bold="true" box="[189,334,806,832]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
|
||
<emphasis id="4C9FEAA65042FF8A16BDFCF8FE900F24" bold="true" box="[189,334,806,832]" pageId="20" pageNumber="21">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="7E5436B45042FF8A16BDFC88FD64082F" blockId="20.[189,1399,854,1099]" pageId="20" pageNumber="21">
|
||
Adults with head, neck and eleven trunk segments (
|
||
<figureCitation id="E6D02A315042FF8A151EFC88FCB60F14" box="[798,872,854,880]" captionStart="Fig" captionStartId="21.[189,232,1753,1779]" captionTargetBox="[315,1235,269,1735]" captionTargetId="figure@21.[315,1273,265,1735]" captionTargetPageId="21" captionText="Fig. 7. Line art illustrations of Echinoderes hamiltonorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♂, segments 10 to 11, dorsal view. D. ♂, segments 10 to 11, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817837" httpUri="https://zenodo.org/record/3817837/files/figure.png" pageId="20" pageNumber="21">Figs 7</figureCitation>
|
||
A–B, 8A, 9A–B). The species is quite small, and trunk and sternal plates appear nearly parallel-sided from segments 2 to 8 (
|
||
<figureCitation id="E6D02A315042FF8A126FFCA4FB110FF0" box="[1135,1231,890,916]" captionStart="Fig" captionStartId="22.[189,232,1611,1637]" captionTargetBox="[256,1332,265,1599]" captionTargetId="figure@22.[256,1332,265,1599]" captionTargetPageId="22" captionText="Fig. 8. Light micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov., holotype, ♀ (A–D, H) (NHMD-223916), paratype, ♂ (E–F) (NHMD-223918) and paratype, ♀ (G) (NHMD-223920). A. Ventral overview. B. Segments 1 to 5, dorsal view. C. Segments 1 to 5, ventral view. D. Segments 6 to 8, dorsal view. E. Segments 6 to 9, ventral view. F. Detail showing sublateral glandular cell outlets type 2 on segments 1 and 2. G. Segments 8 to 9, lateral view. H. Segments 10 to 11 in female, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; pdgco1 = paradorsal glandular cell outlet type 1; pdss = paradorsal sensory spot; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817839" httpUri="https://zenodo.org/record/3817839/files/figure.png" pageId="20" pageNumber="21">Fig. 8A</figureCitation>
|
||
). Intraspecific variation is present on segment 8 where some specimens, but not all, independent of gender and population may have a pair of sublateral glandular cell outlets
|
||
<typeStatus id="A15088165042FF8A1524FC1AFC890FBA" box="[804,855,964,990]" pageId="20" pageNumber="21">type</typeStatus>
|
||
2. Lateral terminal spines are relatively short and slender, 40% to 65% of trunk length (
|
||
<figureCitation id="E6D02A315042FF8A14A0FC39FD370866" box="[672,745,999,1026]" captionStart="Fig" captionStartId="21.[189,232,1753,1779]" captionTargetBox="[315,1235,269,1735]" captionTargetId="figure@21.[315,1273,265,1735]" captionTargetPageId="21" captionText="Fig. 7. Line art illustrations of Echinoderes hamiltonorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♂, segments 10 to 11, dorsal view. D. ♂, segments 10 to 11, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817837" httpUri="https://zenodo.org/record/3817837/files/figure.png" pageId="20" pageNumber="21">Figs 7</figureCitation>
|
||
A–B, 8A). For complete overview of measurements and dimensions, see
|
||
<tableCitation id="3369030F5042FF8A1781FBD2FE020842" box="[385,476,1036,1062]" captionStart="Table 6" captionStartId="23.[189,258,268,294]" captionTargetPageId="23" captionText="Table 6. Measurements from light microscopy of Echinoderes hamiltonorum sp. nov. (in µm), including number of measured specimens (n) and standard deviation (SD). Abbreviations: (ac) = acicular spine; LTAS = lateral terminal accessory spine; LTS = lateral terminal spine; LV = lateroventral; MD = middorsal; MSW-8 = maximum sternal width, measured on segment 8 in this species; S = segment lengths; SW-10 = standard width, always measured on segment 10; TL = trunk length." pageId="20" pageNumber="21">Table 6</tableCitation>
|
||
. Distribution of cuticular structures, i.e., sensory spots, glandular cell outlets, spines and tubes, is summarized in
|
||
<tableCitation id="3369030F5042FF8A145BFBEEFD6A082F" box="[603,692,1072,1099]" captionStart="Table 7" captionStartId="25.[189,258,268,294]" captionTargetPageId="25" captionText="Table 7. Summary of nature and location of sensory spots, glandular cell outlets, tubes and spines arranged by series in Echinoderes hamiltonorum sp. nov. Abbreviations: LA = lateral accessory; LD = laterodorsal; LV = lateroventral; MD = middorsal; ML = midlateral; PD = paradorsal; PV = paraventral; SD = subdorsal; SL = sublateral; VL = ventrolateral; VM = ventromedial; ac = acicular spine; gco1/2 = glandular cell outlet type 1/2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; pe = penile spines; si = sieve plate; ss = sensory spot; tu = tube; (♀) = female and (♂) = male conditions of sexually dimorphic characters; * = character not present in all specimens." httpUri="http://table.plazi.org/id/2A94663C504FFF8716BDFED2FBC50D9D" pageId="20" pageNumber="21" tableUuid="2A94663C504FFF8716BDFED2FBC50D9D">Table 7</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="7E5436B45042FF8A16BDFBADFE47099D" blockId="20.[189,1399,1139,1529]" pageId="20" pageNumber="21">
|
||
The head consists of a retractable mouth cone and an introvert (
|
||
<figureCitation id="E6D02A315042FF8A159CFBADFC0108E9" box="[924,991,1139,1165]" captionStart="Fig" captionStartId="23.[189,232,1508,1534]" captionTargetId="figure@24.[189,1398,260,2026]" captionTargetPageId="24" captionText="Fig. 9 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov. A. Lateral overview. B. Ventral overview. C. Mouth cone. D. Introvert sector 8. E. Segments 1 to 3, dorsolateral view; note that laterodorsal sensory spots are missing in this particular specimen. F. Segments 1 to 3, ventral view. G. Detail showing lateral side of segment 8 without any indication of glandular cell outlets type 2. H. Segments 8 to 9, lateral view. I. Detail showing midlateral to lateroventral parts of segments 5 and 6. J. ♂, segments 10 to 11, dorsolateral view. K. ♂, segments 10 to 11, ventral view. L. ♀, segments 10 to 11, laterodorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdss = middorsal sensory spot; mlss = midlateral sensory spot; oos = outer oral style; pdss = paradorsal sensory spot; pe = penile spines; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlgco1 = ventrolateral glandular cell outlet type 1; vlss = ventrolateral sensory spot; vlt = ventrolateral tube; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817841" httpUri="https://zenodo.org/record/3817841/files/figure.png" pageId="20" pageNumber="21">Fig. 9</figureCitation>
|
||
C–D). Inner oral styles are present, but their exact number and arrangement could not be determined. The external mouth cone armature consists of nine outer oral styles, each with five basal fringe tips (
|
||
<figureCitation id="E6D02A315042FF8A15DBFB62FBE408B2" box="[987,1082,1212,1238]" captionStart="Fig" captionStartId="23.[189,232,1508,1534]" captionTargetId="figure@24.[189,1398,260,2026]" captionTargetPageId="24" captionText="Fig. 9 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov. A. Lateral overview. B. Ventral overview. C. Mouth cone. D. Introvert sector 8. E. Segments 1 to 3, dorsolateral view; note that laterodorsal sensory spots are missing in this particular specimen. F. Segments 1 to 3, ventral view. G. Detail showing lateral side of segment 8 without any indication of glandular cell outlets type 2. H. Segments 8 to 9, lateral view. I. Detail showing midlateral to lateroventral parts of segments 5 and 6. J. ♂, segments 10 to 11, dorsolateral view. K. ♂, segments 10 to 11, ventral view. L. ♀, segments 10 to 11, laterodorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdss = middorsal sensory spot; mlss = midlateral sensory spot; oos = outer oral style; pdss = paradorsal sensory spot; pe = penile spines; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlgco1 = ventrolateral glandular cell outlet type 1; vlss = ventrolateral sensory spot; vlt = ventrolateral tube; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817841" httpUri="https://zenodo.org/record/3817841/files/figure.png" pageId="20" pageNumber="21">Fig. 9C</figureCitation>
|
||
). The introvert sectors are defined by ten primary spinoscalids in Ring 01. Each primary spinoscalid consists of a basal sheath and a distal end piece with a blunt tip. The basal sheaths have marginal extensions forming two layers of transverse fringes. End pieces are smooth. The arrangement of scalids follows the same pattern as in
|
||
<taxonomicName id="B9EB4D375042FF8A16DEFA93FE2D0903" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[222,499,1357,1383]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="20" pageNumber="21" phylum="Cephalorhyncha" rank="species" species="hviidarum" status="sp. nov.">
|
||
<emphasis id="4C9FEAA65042FF8A16DEFA93FE2D0903" box="[222,499,1357,1383]" italics="true" pageId="20" pageNumber="21">Echinoderes hviidarum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD5042FF8A17FCFA90FD84090C" box="[508,602,1358,1384]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
|
||
described below; hence, see
|
||
<figureCitation id="E6D02A315042FF8A15BEFA93FBCC090C" box="[958,1042,1357,1384]" captionStart="Fig" captionStartId="30.[189,232,1930,1956]" captionTargetBox="[201,1388,269,1911]" captionTargetId="figure@30.[200,1389,265,1912]" captionTargetPageId="30" captionText="Fig. 11. Diagram of mouth cone (grey area), introvert and placids in Echinoderes hviidarum sp. nov., showing distribution of outer oral styles, scalids and trichoscalids. Table shows the scalid arrangement by sector; single-lined boxes mark quincunxes, double-lined boxes mark “double diamonds”." figureDoi="http://doi.org/10.5281/zenodo.3817845" httpUri="https://zenodo.org/record/3817845/files/figure.png" pageId="20" pageNumber="21">Fig. 11</figureCitation>
|
||
for a complete overview. All spinoscalids in these rings consist of a basal sheath and a pointed end piece (
|
||
<figureCitation id="E6D02A315042FF8A1237FAACFB4B09E8" box="[1079,1173,1394,1420]" captionStart="Fig" captionStartId="23.[189,232,1508,1534]" captionTargetId="figure@24.[189,1398,260,2026]" captionTargetPageId="24" captionText="Fig. 9 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov. A. Lateral overview. B. Ventral overview. C. Mouth cone. D. Introvert sector 8. E. Segments 1 to 3, dorsolateral view; note that laterodorsal sensory spots are missing in this particular specimen. F. Segments 1 to 3, ventral view. G. Detail showing lateral side of segment 8 without any indication of glandular cell outlets type 2. H. Segments 8 to 9, lateral view. I. Detail showing midlateral to lateroventral parts of segments 5 and 6. J. ♂, segments 10 to 11, dorsolateral view. K. ♂, segments 10 to 11, ventral view. L. ♀, segments 10 to 11, laterodorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdss = middorsal sensory spot; mlss = midlateral sensory spot; oos = outer oral style; pdss = paradorsal sensory spot; pe = penile spines; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlgco1 = ventrolateral glandular cell outlet type 1; vlss = ventrolateral sensory spot; vlt = ventrolateral tube; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817841" httpUri="https://zenodo.org/record/3817841/files/figure.png" pageId="20" pageNumber="21">Fig. 9D</figureCitation>
|
||
). The basal sheaths terminate in fringed margins on spinoscalids of Rings 02 and 04, whereas the sheaths of Ring 03 have a median spike only. Spinoscalids of Rings 05 to 07 are composed as those in the preceding ring, but with shorter end pieces.
|
||
</paragraph>
|
||
<paragraph id="7E5436B45042FF8A16BDF9FCFE8A0ACD" blockId="20.[189,1398,1570,1705]" pageId="20" pageNumber="21">
|
||
The neck has 16 placids, measuring
|
||
<quantity id="B9139B515042FF8A147CF9FCFD120A58" box="[636,716,1570,1596]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="21" unit="mm" value="10.0">10 µm</quantity>
|
||
in length. The midventral placid is broadest, measuring
|
||
<quantity id="B9139B515042FF8A16BDF998FED60A05" box="[189,264,1606,1633]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="20" pageNumber="21" unit="mm" value="11.0">11 µm</quantity>
|
||
in width at its base, whereas all other are narrower, measuring
|
||
<quantity id="B9139B515042FF8A15EBF999FBF70A05" box="[1003,1065,1607,1633]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="20" pageNumber="21" unit="mm" value="7.0">7 µm</quantity>
|
||
in width at their bases. Four dorsal and two ventral trichoscalid plates are present; the plates are well-developed and hat-shaped (
|
||
<figureCitation id="E6D02A315042FF8A16C6F951FED50ACD" box="[198,267,1679,1705]" captionStart="Fig" captionStartId="21.[189,232,1753,1779]" captionTargetBox="[315,1235,269,1735]" captionTargetId="figure@21.[315,1273,265,1735]" captionTargetPageId="21" captionText="Fig. 7. Line art illustrations of Echinoderes hamiltonorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♂, segments 10 to 11, dorsal view. D. ♂, segments 10 to 11, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817837" httpUri="https://zenodo.org/record/3817837/files/figure.png" pageId="20" pageNumber="21">Fig. 7</figureCitation>
|
||
A–B).
|
||
</paragraph>
|
||
<paragraph id="7E5436B45042FF8A16BDF90CFD4E0B8F" blockId="20.[189,1399,1746,2027]" pageId="20" pageNumber="21">
|
||
Segment 1 consists of a complete cuticular ring. A pair of minute glandular cell outlets
|
||
<typeStatus id="A15088165042FF8A12B8F90CFB350A88" box="[1208,1259,1746,1772]" pageId="20" pageNumber="21">type</typeStatus>
|
||
2 is present in sublateral positions (
|
||
<figureCitation id="E6D02A315042FF8A17D5F928FDE60B75" box="[469,568,1782,1809]" captionStart="Fig" captionStartId="21.[189,232,1753,1779]" captionTargetBox="[315,1235,269,1735]" captionTargetId="figure@21.[315,1273,265,1735]" captionTargetPageId="21" captionText="Fig. 7. Line art illustrations of Echinoderes hamiltonorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♂, segments 10 to 11, dorsal view. D. ♂, segments 10 to 11, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817837" httpUri="https://zenodo.org/record/3817837/files/figure.png" pageId="20" pageNumber="21">Figs 7B</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A315042FF8A1447F928FD550B75" box="[583,651,1782,1809]" captionStart="Fig" captionStartId="22.[189,232,1611,1637]" captionTargetBox="[256,1332,265,1599]" captionTargetId="figure@22.[256,1332,265,1599]" captionTargetPageId="22" captionText="Fig. 8. Light micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov., holotype, ♀ (A–D, H) (NHMD-223916), paratype, ♂ (E–F) (NHMD-223918) and paratype, ♀ (G) (NHMD-223920). A. Ventral overview. B. Segments 1 to 5, dorsal view. C. Segments 1 to 5, ventral view. D. Segments 6 to 8, dorsal view. E. Segments 6 to 9, ventral view. F. Detail showing sublateral glandular cell outlets type 2 on segments 1 and 2. G. Segments 8 to 9, lateral view. H. Segments 10 to 11 in female, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; pdgco1 = paradorsal glandular cell outlet type 1; pdss = paradorsal sensory spot; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817839" httpUri="https://zenodo.org/record/3817839/files/figure.png" pageId="20" pageNumber="21">8C, F</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A315042FF8A1499F928FD790B74" box="[665,679,1782,1808]" captionStart="Fig" captionStartId="23.[189,232,1508,1534]" captionTargetId="figure@24.[189,1398,260,2026]" captionTargetPageId="24" captionText="Fig. 9 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov. A. Lateral overview. B. Ventral overview. C. Mouth cone. D. Introvert sector 8. E. Segments 1 to 3, dorsolateral view; note that laterodorsal sensory spots are missing in this particular specimen. F. Segments 1 to 3, ventral view. G. Detail showing lateral side of segment 8 without any indication of glandular cell outlets type 2. H. Segments 8 to 9, lateral view. I. Detail showing midlateral to lateroventral parts of segments 5 and 6. J. ♂, segments 10 to 11, dorsolateral view. K. ♂, segments 10 to 11, ventral view. L. ♀, segments 10 to 11, laterodorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdss = middorsal sensory spot; mlss = midlateral sensory spot; oos = outer oral style; pdss = paradorsal sensory spot; pe = penile spines; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlgco1 = ventrolateral glandular cell outlet type 1; vlss = ventrolateral sensory spot; vlt = ventrolateral tube; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817841" httpUri="https://zenodo.org/record/3817841/files/figure.png" pageId="20" pageNumber="21">9</figureCitation>
|
||
E–F). Sensory spots are located medially on the segment in subdorsal and laterodorsal positions (
|
||
<figureCitation id="E6D02A315042FF8A1476F8C4FD040B51" box="[630,730,1818,1845]" captionStart="Fig" captionStartId="21.[189,232,1753,1779]" captionTargetBox="[315,1235,269,1735]" captionTargetId="figure@21.[315,1273,265,1735]" captionTargetPageId="21" captionText="Fig. 7. Line art illustrations of Echinoderes hamiltonorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♂, segments 10 to 11, dorsal view. D. ♂, segments 10 to 11, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817837" httpUri="https://zenodo.org/record/3817837/files/figure.png" pageId="20" pageNumber="21">Figs 7A</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A315042FF8A14E6F8C5FCD40B51" box="[742,778,1819,1845]" captionStart="Fig" captionStartId="22.[189,232,1611,1637]" captionTargetBox="[256,1332,265,1599]" captionTargetId="figure@22.[256,1332,265,1599]" captionTargetPageId="22" captionText="Fig. 8. Light micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov., holotype, ♀ (A–D, H) (NHMD-223916), paratype, ♂ (E–F) (NHMD-223918) and paratype, ♀ (G) (NHMD-223920). A. Ventral overview. B. Segments 1 to 5, dorsal view. C. Segments 1 to 5, ventral view. D. Segments 6 to 8, dorsal view. E. Segments 6 to 9, ventral view. F. Detail showing sublateral glandular cell outlets type 2 on segments 1 and 2. G. Segments 8 to 9, lateral view. H. Segments 10 to 11 in female, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; pdgco1 = paradorsal glandular cell outlet type 1; pdss = paradorsal sensory spot; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817839" httpUri="https://zenodo.org/record/3817839/files/figure.png" pageId="20" pageNumber="21">8B</figureCitation>
|
||
) (laterodorsal sensory spots are missing in one nontype specimen,
|
||
<figureCitation id="E6D02A315042FF8A1777F8E1FE0D0B3D" box="[375,467,1855,1881]" captionStart="Fig" captionStartId="23.[189,232,1508,1534]" captionTargetId="figure@24.[189,1398,260,2026]" captionTargetPageId="24" captionText="Fig. 9 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov. A. Lateral overview. B. Ventral overview. C. Mouth cone. D. Introvert sector 8. E. Segments 1 to 3, dorsolateral view; note that laterodorsal sensory spots are missing in this particular specimen. F. Segments 1 to 3, ventral view. G. Detail showing lateral side of segment 8 without any indication of glandular cell outlets type 2. H. Segments 8 to 9, lateral view. I. Detail showing midlateral to lateroventral parts of segments 5 and 6. J. ♂, segments 10 to 11, dorsolateral view. K. ♂, segments 10 to 11, ventral view. L. ♀, segments 10 to 11, laterodorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdss = middorsal sensory spot; mlss = midlateral sensory spot; oos = outer oral style; pdss = paradorsal sensory spot; pe = penile spines; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlgco1 = ventrolateral glandular cell outlet type 1; vlss = ventrolateral sensory spot; vlt = ventrolateral tube; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817841" httpUri="https://zenodo.org/record/3817841/files/figure.png" pageId="20" pageNumber="21">Fig. 9E</figureCitation>
|
||
); sensory spots on this and following nine segments are medium-sized, with a tuft of micropapillae and long marginal cuticular hairs. Glandular cell outlets
|
||
<typeStatus id="A15088165042FF8A123CF8BAFBB10B1A" box="[1084,1135,1892,1918]" pageId="20" pageNumber="21">type</typeStatus>
|
||
1 present in middorsal and ventrolateral positions (
|
||
<figureCitation id="E6D02A315042FF8A140BF856FD890BC6" box="[523,599,1928,1954]" captionStart="Fig" captionStartId="21.[189,232,1753,1779]" captionTargetBox="[315,1235,269,1735]" captionTargetId="figure@21.[315,1273,265,1735]" captionTargetPageId="21" captionText="Fig. 7. Line art illustrations of Echinoderes hamiltonorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♂, segments 10 to 11, dorsal view. D. ♂, segments 10 to 11, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817837" httpUri="https://zenodo.org/record/3817837/files/figure.png" pageId="20" pageNumber="21">Figs 7</figureCitation>
|
||
A–B, 9F). Cuticular hairs arranged in a single transverse row near posterior segment margin. The posterior segment margin is straight, terminating in a pectinate fringe with rather long and flexible fringe tips.
|
||
</paragraph>
|
||
<caption id="2A94663C5043FF8B16BDF907FA8C0B8E" ID-DOI="http://doi.org/10.5281/zenodo.3817837" ID-Zenodo-Dep="3817837" httpUri="https://zenodo.org/record/3817837/files/figure.png" pageId="21" pageNumber="22" startId="21.[189,232,1753,1779]" targetBox="[315,1235,269,1735]" targetPageId="21">
|
||
<paragraph id="7E5436B45043FF8B16BDF907FA8C0B8E" blockId="21.[189,1399,1753,2027]" pageId="21" pageNumber="22">
|
||
<emphasis id="4C9FEAA65043FF8B16BDF907FED00A90" bold="true" box="[189,270,1753,1780]" pageId="21" pageNumber="22">Fig. 7.</emphasis>
|
||
Line art illustrations of
|
||
<taxonomicName id="B9EB4D375043FF8B1439F904FCA50A90" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[569,891,1754,1780]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="21" pageNumber="22" phylum="Cephalorhyncha" rank="species" species="hamiltonorum" status="sp. nov.">
|
||
<emphasis id="4C9FEAA65043FF8B1439F904FCA50A90" box="[569,891,1754,1780]" italics="true" pageId="21" pageNumber="22">Echinoderes hamiltonorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD5043FF8B1586F904FC3A0A90" box="[902,996,1754,1780]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="4C9FEAA65043FF8B15EEF907FBDD0A97" bold="true" box="[1006,1027,1753,1779]" pageId="21" pageNumber="22">A</emphasis>
|
||
. ♀, dorsal view.
|
||
<emphasis id="4C9FEAA65043FF8B12D8F904FB320A90" bold="true" box="[1240,1260,1754,1780]" pageId="21" pageNumber="22">B</emphasis>
|
||
. ♀, ventral view.
|
||
<emphasis id="4C9FEAA65043FF8B1706F923FEC50B73" bold="true" box="[262,283,1789,1815]" pageId="21" pageNumber="22">C</emphasis>
|
||
. ♂, segments 10 to 11, dorsal view.
|
||
<emphasis id="4C9FEAA65043FF8B14E3F923FD260B73" bold="true" box="[739,760,1789,1815]" pageId="21" pageNumber="22">D</emphasis>
|
||
. ♂, segments 10 to 11, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="2A94663C5040FF8816BDF995FE4D0B8D" ID-DOI="http://doi.org/10.5281/zenodo.3817839" ID-Zenodo-Dep="3817839" httpUri="https://zenodo.org/record/3817839/files/figure.png" pageId="22" pageNumber="23" startId="22.[189,232,1611,1637]" targetBox="[256,1332,265,1599]" targetPageId="22">
|
||
<paragraph id="7E5436B45040FF8816BDF995FE4D0B8D" blockId="22.[189,1399,1611,2025]" pageId="22" pageNumber="23">
|
||
<emphasis id="4C9FEAA65040FF8816BDF995FECF0A01" bold="true" box="[189,273,1611,1637]" pageId="22" pageNumber="23">Fig. 8.</emphasis>
|
||
Light micrographs showing overviews and details of
|
||
<taxonomicName id="B9EB4D375040FF8815BBF992FADF0A02" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[955,1281,1612,1638]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="22" pageNumber="23" phylum="Cephalorhyncha" rank="species" species="hamiltonorum" status="sp. nov.">
|
||
<emphasis id="4C9FEAA65040FF8815BBF992FADF0A02" box="[955,1281,1612,1638]" italics="true" pageId="22" pageNumber="23">Echinoderes hamiltonorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD5040FF88130FF992FAAC0A02" box="[1295,1394,1612,1638]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, ♀ (A–D, H) (NHMD-223916), paratype, ♂ (E–F) (NHMD-223918) and paratype, ♀ (G) (NHMD-223920).
|
||
<emphasis id="4C9FEAA65040FF88179DF94CFE6C0AC8" bold="true" box="[413,434,1682,1708]" pageId="22" pageNumber="23">A</emphasis>
|
||
. Ventral overview.
|
||
<emphasis id="4C9FEAA65040FF88149CF94CFD6E0AC8" bold="true" box="[668,688,1682,1708]" pageId="22" pageNumber="23">B</emphasis>
|
||
. Segments 1 to 5, dorsal view.
|
||
<emphasis id="4C9FEAA65040FF88122CF94CFB9F0AC8" bold="true" box="[1068,1089,1682,1708]" pageId="22" pageNumber="23">C</emphasis>
|
||
. Segments 1 to 5, ventral view.
|
||
<emphasis id="4C9FEAA65040FF881704F96BFEC70AAB" bold="true" box="[260,281,1717,1743]" pageId="22" pageNumber="23">D</emphasis>
|
||
. Segments 6 to 8, dorsal view.
|
||
<emphasis id="4C9FEAA65040FF881491F96BFD7B0AAB" bold="true" box="[657,677,1717,1743]" pageId="22" pageNumber="23">E</emphasis>
|
||
. Segments 6 to 9, ventral view.
|
||
<emphasis id="4C9FEAA65040FF881226F96BFBE60AAB" bold="true" box="[1062,1080,1717,1743]" pageId="22" pageNumber="23">F</emphasis>
|
||
. Detail showing sublateral glandular cell outlets type 2 on segments 1 and 2.
|
||
<emphasis id="4C9FEAA65040FF88150FF906FCF80A96" bold="true" box="[783,806,1752,1778]" pageId="22" pageNumber="23">G</emphasis>
|
||
. Segments 8 to 9, lateral view.
|
||
<emphasis id="4C9FEAA65040FF88129AF907FB6F0A97" bold="true" box="[1178,1201,1753,1779]" pageId="22" pageNumber="23">H</emphasis>
|
||
. Segments 10 to 11 in female, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; pdgco1 = paradorsal glandular cell outlet type 1; pdss = paradorsal sensory spot; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="2A94663C5041FF8916BDFED2FB6F0DD6" pageId="23" pageNumber="24" targetBox="[357,1199,470,1400]" targetIsTable="true" targetPageId="23">
|
||
<paragraph id="7E5436B45041FF8916BDFED2FB6F0DD6" blockId="23.[189,1399,267,435]" pageId="23" pageNumber="24">
|
||
<emphasis id="4C9FEAA65041FF8916BDFED2FEC00D41" bold="true" box="[189,286,267,294]" pageId="23" pageNumber="24">Table 6.</emphasis>
|
||
Measurements from light microscopy of
|
||
<taxonomicName id="B9EB4D375041FF891501FED5FBE00D41" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[769,1086,267,293]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="23" pageNumber="24" phylum="Cephalorhyncha" rank="species" species="hamiltonorum" status="sp. nov.">
|
||
<emphasis id="4C9FEAA65041FF891501FED5FBE00D41" box="[769,1086,267,293]" italics="true" pageId="23" pageNumber="24">Echinoderes hamiltonorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD5041FF891244FED2FB430D42" box="[1092,1181,268,294]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(in µm), including number of measured specimens (n) and standard deviation (SD). Abbreviations: (ac) = acicular spine; LTAS = lateral terminal accessory spine; LTS = lateral terminal spine; LV = lateroventral; MD = middorsal; MSW-8 = maximum sternal width, measured on segment 8 in this species; S = segment lengths; SW-10 = standard width, always measured on segment 10; TL = trunk length.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="7E5436B45041FF891769FE08FB7D091C" pageId="23" pageNumber="24">
|
||
<table id="0CEBC414504100611765FE08FB71091C" box="[357,1199,470,1400]" colsContinueFrom="15.[355,1204,464,1477]" colsContinueIn="32.[357,1199,455,1351]" gridcols="5" gridrows="27" pageId="23" pageNumber="24">
|
||
<tr id="C0DB34F6504100611765FE08FB710D95" box="[357,1199,470,497]" gridrow="0" pageId="23" pageNumber="24">
|
||
<th id="830A5D8A504100611765FE08FE360D95" box="[357,488,470,497]" gridcol="0" gridrow="0" pageId="23" pageNumber="24">
|
||
<emphasis id="4C9FEAA65041FF891769FE08FE3A0D95" bold="true" box="[361,484,470,497]" pageId="23" pageNumber="24">Character</emphasis>
|
||
</th>
|
||
<th id="830A5D8A50410061143DFE08FD930D95" box="[573,589,470,497]" gridcol="1" gridrow="0" pageId="23" pageNumber="24">
|
||
<emphasis id="4C9FEAA65041FF89143DFE09FD930D95" bold="true" box="[573,589,471,497]" pageId="23" pageNumber="24">n</emphasis>
|
||
</th>
|
||
<th id="830A5D8A5041006114A0FE08FCF90D95" box="[672,807,470,497]" gridcol="2" gridrow="0" pageId="23" pageNumber="24">
|
||
<emphasis id="4C9FEAA65041FF8914BDFE09FCD70D95" bold="true" box="[701,777,471,497]" pageId="23" pageNumber="24">Range</emphasis>
|
||
</th>
|
||
<th id="830A5D8A504100611593FE08FC020D95" box="[915,988,470,497]" gridcol="3" gridrow="0" pageId="23" pageNumber="24">
|
||
<emphasis id="4C9FEAA65041FF891595FE09FC070D95" bold="true" box="[917,985,471,497]" pageId="23" pageNumber="24">Mean</emphasis>
|
||
</th>
|
||
<th id="830A5D8A504100611267FE08FB710D95" box="[1127,1199,470,497]" gridcol="4" gridrow="0" pageId="23" pageNumber="24">
|
||
<emphasis id="4C9FEAA65041FF891279FE08FB430D95" bold="true" box="[1145,1181,470,497]" pageId="23" pageNumber="24">SD</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="C0DB34F6504100611765FE27FB710E70" box="[357,1199,505,532]" gridrow="1" pageId="23" pageNumber="24">
|
||
<th id="830A5D8A504100611765FE27FE360E70" box="[357,488,505,532]" gridcol="0" gridrow="1" pageId="23" pageNumber="24">TL</th>
|
||
<td id="830A5D8A50410061143DFE27FD930E70" box="[573,589,505,532]" gridcol="1" gridrow="1" pageId="23" pageNumber="24">7</td>
|
||
<td id="830A5D8A5041006114A0FE27FCF90E70" box="[672,807,505,532]" gridcol="2" gridrow="1" pageId="23" pageNumber="24">197–263</td>
|
||
<td id="830A5D8A504100611593FE27FC020E70" box="[915,988,505,532]" gridcol="3" gridrow="1" pageId="23" pageNumber="24">233</td>
|
||
<td id="830A5D8A504100611267FE27FB710E70" box="[1127,1199,505,532]" gridcol="4" gridrow="1" pageId="23" pageNumber="24">21.20</td>
|
||
</tr>
|
||
<tr id="C0DB34F6504100611765FDC5FB710E51" box="[357,1199,539,565]" gridrow="2" pageId="23" pageNumber="24">
|
||
<th id="830A5D8A504100611765FDC5FE360E51" box="[357,488,539,565]" gridcol="0" gridrow="2" pageId="23" pageNumber="24">MSW-8</th>
|
||
<td id="830A5D8A50410061143DFDC5FD930E51" box="[573,589,539,565]" gridcol="1" gridrow="2" pageId="23" pageNumber="24">4</td>
|
||
<td id="830A5D8A5041006114A0FDC5FCF90E51" box="[672,807,539,565]" gridcol="2" gridrow="2" pageId="23" pageNumber="24">42–46</td>
|
||
<td id="830A5D8A504100611593FDC5FC020E51" box="[915,988,539,565]" gridcol="3" gridrow="2" pageId="23" pageNumber="24">44</td>
|
||
<td id="830A5D8A504100611267FDC5FB710E51" box="[1127,1199,539,565]" gridcol="4" gridrow="2" pageId="23" pageNumber="24">2.06</td>
|
||
</tr>
|
||
<tr id="C0DB34F6504100611765FDE5FB710E32" box="[357,1199,571,598]" gridrow="3" pageId="23" pageNumber="24">
|
||
<th id="830A5D8A504100611765FDE5FE360E32" box="[357,488,571,598]" gridcol="0" gridrow="3" pageId="23" pageNumber="24">MSW-8/TL</th>
|
||
<td id="830A5D8A50410061143DFDE5FD930E32" box="[573,589,571,598]" gridcol="1" gridrow="3" pageId="23" pageNumber="24">4</td>
|
||
<td id="830A5D8A5041006114A0FDE5FCF90E32" box="[672,807,571,598]" gridcol="2" gridrow="3" pageId="23" pageNumber="24">17.7–19.3%</td>
|
||
<td id="830A5D8A504100611593FDE5FC020E32" box="[915,988,571,598]" gridcol="3" gridrow="3" pageId="23" pageNumber="24">18.5%</td>
|
||
<td id="830A5D8A504100611267FDE5FB710E32" box="[1127,1199,571,598]" gridcol="4" gridrow="3" pageId="23" pageNumber="24">0.83%</td>
|
||
</tr>
|
||
<tr id="C0DB34F6504100611765FD80FB710E1C" box="[357,1199,606,632]" gridrow="4" pageId="23" pageNumber="24">
|
||
<th id="830A5D8A504100611765FD80FE360E1C" box="[357,488,606,632]" gridcol="0" gridrow="4" pageId="23" pageNumber="24">SW-10</th>
|
||
<td id="830A5D8A50410061143DFD80FD930E1C" box="[573,589,606,632]" gridcol="1" gridrow="4" pageId="23" pageNumber="24">4</td>
|
||
<td id="830A5D8A5041006114A0FD80FCF90E1C" box="[672,807,606,632]" gridcol="2" gridrow="4" pageId="23" pageNumber="24">36–41</td>
|
||
<td id="830A5D8A504100611593FD80FC020E1C" box="[915,988,606,632]" gridcol="3" gridrow="4" pageId="23" pageNumber="24">39</td>
|
||
<td id="830A5D8A504100611267FD80FB710E1C" box="[1127,1199,606,632]" gridcol="4" gridrow="4" pageId="23" pageNumber="24">2.36%</td>
|
||
</tr>
|
||
<tr id="C0DB34F6504100611765FDA1FB710EFE" box="[357,1199,639,666]" gridrow="5" pageId="23" pageNumber="24">
|
||
<th id="830A5D8A504100611765FDA1FE360EFE" box="[357,488,639,666]" gridcol="0" gridrow="5" pageId="23" pageNumber="24">SW-10/TL</th>
|
||
<td id="830A5D8A50410061143DFDA1FD930EFE" box="[573,589,639,666]" gridcol="1" gridrow="5" pageId="23" pageNumber="24">4</td>
|
||
<td id="830A5D8A5041006114A0FDA1FCF90EFE" box="[672,807,639,666]" gridcol="2" gridrow="5" pageId="23" pageNumber="24">15.1–17.3%</td>
|
||
<td id="830A5D8A504100611593FDA1FC020EFE" box="[915,988,639,666]" gridcol="3" gridrow="5" pageId="23" pageNumber="24">16.4%</td>
|
||
<td id="830A5D8A504100611267FDA1FB710EFE" box="[1127,1199,639,666]" gridcol="4" gridrow="5" pageId="23" pageNumber="24">0.99%</td>
|
||
</tr>
|
||
<tr id="C0DB34F6504100611765FD7FFB710ED8" box="[357,1199,673,700]" gridrow="6" pageId="23" pageNumber="24">
|
||
<th id="830A5D8A504100611765FD7FFE360ED8" box="[357,488,673,700]" gridcol="0" gridrow="6" pageId="23" pageNumber="24">S1</th>
|
||
<td id="830A5D8A50410061143DFD7FFD930ED8" box="[573,589,673,700]" gridcol="1" gridrow="6" pageId="23" pageNumber="24">7</td>
|
||
<td id="830A5D8A5041006114A0FD7FFCF90ED8" box="[672,807,673,700]" gridcol="2" gridrow="6" pageId="23" pageNumber="24">24–28</td>
|
||
<td id="830A5D8A504100611593FD7FFC020ED8" box="[915,988,673,700]" gridcol="3" gridrow="6" pageId="23" pageNumber="24">27</td>
|
||
<td id="830A5D8A504100611267FD7FFB710ED8" box="[1127,1199,673,700]" gridcol="4" gridrow="6" pageId="23" pageNumber="24">1.62</td>
|
||
</tr>
|
||
<tr id="C0DB34F6504100611765FD1DFB710EB9" box="[357,1199,707,733]" gridrow="7" pageId="23" pageNumber="24">
|
||
<th id="830A5D8A504100611765FD1DFE360EB9" box="[357,488,707,733]" gridcol="0" gridrow="7" pageId="23" pageNumber="24">S2</th>
|
||
<td id="830A5D8A50410061143DFD1DFD930EB9" box="[573,589,707,733]" gridcol="1" gridrow="7" pageId="23" pageNumber="24">7</td>
|
||
<td id="830A5D8A5041006114A0FD1DFCF90EB9" box="[672,807,707,733]" gridcol="2" gridrow="7" pageId="23" pageNumber="24">21–27</td>
|
||
<td id="830A5D8A504100611593FD1DFC020EB9" box="[915,988,707,733]" gridcol="3" gridrow="7" pageId="23" pageNumber="24">24</td>
|
||
<td id="830A5D8A504100611267FD1DFB710EB9" box="[1127,1199,707,733]" gridcol="4" gridrow="7" pageId="23" pageNumber="24">1.77</td>
|
||
</tr>
|
||
<tr id="C0DB34F6504100611765FD3AFB710E9B" box="[357,1199,740,767]" gridrow="8" pageId="23" pageNumber="24">
|
||
<th id="830A5D8A504100611765FD3AFE360E9B" box="[357,488,740,767]" gridcol="0" gridrow="8" pageId="23" pageNumber="24">S3</th>
|
||
<td id="830A5D8A50410061143DFD3AFD930E9B" box="[573,589,740,767]" gridcol="1" gridrow="8" pageId="23" pageNumber="24">7</td>
|
||
<td id="830A5D8A5041006114A0FD3AFCF90E9B" box="[672,807,740,767]" gridcol="2" gridrow="8" pageId="23" pageNumber="24">24–28</td>
|
||
<td id="830A5D8A504100611593FD3AFC020E9B" box="[915,988,740,767]" gridcol="3" gridrow="8" pageId="23" pageNumber="24">26</td>
|
||
<td id="830A5D8A504100611267FD3AFB710E9B" box="[1127,1199,740,767]" gridcol="4" gridrow="8" pageId="23" pageNumber="24">1.40</td>
|
||
</tr>
|
||
<tr id="C0DB34F6504100611765FCD8FB710F44" box="[357,1199,774,800]" gridrow="9" pageId="23" pageNumber="24">
|
||
<th id="830A5D8A504100611765FCD8FE360F44" box="[357,488,774,800]" gridcol="0" gridrow="9" pageId="23" pageNumber="24">S4</th>
|
||
<td id="830A5D8A50410061143DFCD8FD930F44" box="[573,589,774,800]" gridcol="1" gridrow="9" pageId="23" pageNumber="24">7</td>
|
||
<td id="830A5D8A5041006114A0FCD8FCF90F44" box="[672,807,774,800]" gridcol="2" gridrow="9" pageId="23" pageNumber="24">26–32</td>
|
||
<td id="830A5D8A504100611593FCD8FC020F44" box="[915,988,774,800]" gridcol="3" gridrow="9" pageId="23" pageNumber="24">29</td>
|
||
<td id="830A5D8A504100611267FCD8FB710F44" box="[1127,1199,774,800]" gridcol="4" gridrow="9" pageId="23" pageNumber="24">2.44</td>
|
||
</tr>
|
||
<tr id="C0DB34F6504100611765FCF9FB710F26" box="[357,1199,807,834]" gridrow="10" pageId="23" pageNumber="24">
|
||
<th id="830A5D8A504100611765FCF9FE360F26" box="[357,488,807,834]" gridcol="0" gridrow="10" pageId="23" pageNumber="24">S5</th>
|
||
<td id="830A5D8A50410061143DFCF9FD930F26" box="[573,589,807,834]" gridcol="1" gridrow="10" pageId="23" pageNumber="24">7</td>
|
||
<td id="830A5D8A5041006114A0FCF9FCF90F26" box="[672,807,807,834]" gridcol="2" gridrow="10" pageId="23" pageNumber="24">28–34</td>
|
||
<td id="830A5D8A504100611593FCF9FC020F26" box="[915,988,807,834]" gridcol="3" gridrow="10" pageId="23" pageNumber="24">32</td>
|
||
<td id="830A5D8A504100611267FCF9FB710F26" box="[1127,1199,807,834]" gridcol="4" gridrow="10" pageId="23" pageNumber="24">2.57</td>
|
||
</tr>
|
||
<tr id="C0DB34F6504100611765FC97FB710F00" box="[357,1199,841,868]" gridrow="11" pageId="23" pageNumber="24">
|
||
<th id="830A5D8A504100611765FC97FE360F00" box="[357,488,841,868]" gridcol="0" gridrow="11" pageId="23" pageNumber="24">S6</th>
|
||
<td id="830A5D8A50410061143DFC97FD930F00" box="[573,589,841,868]" gridcol="1" gridrow="11" pageId="23" pageNumber="24">7</td>
|
||
<td id="830A5D8A5041006114A0FC97FCF90F00" box="[672,807,841,868]" gridcol="2" gridrow="11" pageId="23" pageNumber="24">31–37</td>
|
||
<td id="830A5D8A504100611593FC97FC020F00" box="[915,988,841,868]" gridcol="3" gridrow="11" pageId="23" pageNumber="24">34</td>
|
||
<td id="830A5D8A504100611267FC97FB710F00" box="[1127,1199,841,868]" gridcol="4" gridrow="11" pageId="23" pageNumber="24">2.37</td>
|
||
</tr>
|
||
<tr id="C0DB34F6504100611765FCB5FB710FE1" box="[357,1199,875,901]" gridrow="12" pageId="23" pageNumber="24">
|
||
<th id="830A5D8A504100611765FCB5FE360FE1" box="[357,488,875,901]" gridcol="0" gridrow="12" pageId="23" pageNumber="24">S7</th>
|
||
<td id="830A5D8A50410061143DFCB5FD930FE1" box="[573,589,875,901]" gridcol="1" gridrow="12" pageId="23" pageNumber="24">7</td>
|
||
<td id="830A5D8A5041006114A0FCB5FCF90FE1" box="[672,807,875,901]" gridcol="2" gridrow="12" pageId="23" pageNumber="24">36–42</td>
|
||
<td id="830A5D8A504100611593FCB5FC020FE1" box="[915,988,875,901]" gridcol="3" gridrow="12" pageId="23" pageNumber="24">38</td>
|
||
<td id="830A5D8A504100611267FCB5FB710FE1" box="[1127,1199,875,901]" gridcol="4" gridrow="12" pageId="23" pageNumber="24">2.24</td>
|
||
</tr>
|
||
<tr id="C0DB34F6504100611765FC52FB710FC3" box="[357,1199,908,935]" gridrow="13" pageId="23" pageNumber="24">
|
||
<th id="830A5D8A504100611765FC52FE360FC3" box="[357,488,908,935]" gridcol="0" gridrow="13" pageId="23" pageNumber="24">S8</th>
|
||
<td id="830A5D8A50410061143DFC52FD930FC3" box="[573,589,908,935]" gridcol="1" gridrow="13" pageId="23" pageNumber="24">7</td>
|
||
<td id="830A5D8A5041006114A0FC52FCF90FC3" box="[672,807,908,935]" gridcol="2" gridrow="13" pageId="23" pageNumber="24">36–42</td>
|
||
<td id="830A5D8A504100611593FC52FC020FC3" box="[915,988,908,935]" gridcol="3" gridrow="13" pageId="23" pageNumber="24">40</td>
|
||
<td id="830A5D8A504100611267FC52FB710FC3" box="[1127,1199,908,935]" gridcol="4" gridrow="13" pageId="23" pageNumber="24">2.08</td>
|
||
</tr>
|
||
<tr id="C0DB34F6504100611765FC73FB710FAC" box="[357,1199,941,968]" gridrow="14" pageId="23" pageNumber="24">
|
||
<th id="830A5D8A504100611765FC73FE360FAC" box="[357,488,941,968]" gridcol="0" gridrow="14" pageId="23" pageNumber="24">S9</th>
|
||
<td id="830A5D8A50410061143DFC73FD930FAC" box="[573,589,941,968]" gridcol="1" gridrow="14" pageId="23" pageNumber="24">7</td>
|
||
<td id="830A5D8A5041006114A0FC73FCF90FAC" box="[672,807,941,968]" gridcol="2" gridrow="14" pageId="23" pageNumber="24">39–47</td>
|
||
<td id="830A5D8A504100611593FC73FC020FAC" box="[915,988,941,968]" gridcol="3" gridrow="14" pageId="23" pageNumber="24">42</td>
|
||
<td id="830A5D8A504100611267FC73FB710FAC" box="[1127,1199,941,968]" gridcol="4" gridrow="14" pageId="23" pageNumber="24">2.85</td>
|
||
</tr>
|
||
<tr id="C0DB34F6504100611765FC11FB710F8E" box="[357,1199,975,1002]" gridrow="15" pageId="23" pageNumber="24">
|
||
<th id="830A5D8A504100611765FC11FE360F8E" box="[357,488,975,1002]" gridcol="0" gridrow="15" pageId="23" pageNumber="24">S10</th>
|
||
<td id="830A5D8A50410061143DFC11FD930F8E" box="[573,589,975,1002]" gridcol="1" gridrow="15" pageId="23" pageNumber="24">7</td>
|
||
<td id="830A5D8A5041006114A0FC11FCF90F8E" box="[672,807,975,1002]" gridcol="2" gridrow="15" pageId="23" pageNumber="24">29–35</td>
|
||
<td id="830A5D8A504100611593FC11FC020F8E" box="[915,988,975,1002]" gridcol="3" gridrow="15" pageId="23" pageNumber="24">32</td>
|
||
<td id="830A5D8A504100611267FC11FB710F8E" box="[1127,1199,975,1002]" gridcol="4" gridrow="15" pageId="23" pageNumber="24">2.04</td>
|
||
</tr>
|
||
<tr id="C0DB34F6504100611765FC2EFB710868" box="[357,1199,1008,1036]" gridrow="16" pageId="23" pageNumber="24">
|
||
<th id="830A5D8A504100611765FC2EFE360868" box="[357,488,1008,1036]" gridcol="0" gridrow="16" pageId="23" pageNumber="24">S11</th>
|
||
<td id="830A5D8A50410061143DFC2EFD930868" box="[573,589,1008,1036]" gridcol="1" gridrow="16" pageId="23" pageNumber="24">7</td>
|
||
<td id="830A5D8A5041006114A0FC2EFCF90868" box="[672,807,1008,1036]" gridcol="2" gridrow="16" pageId="23" pageNumber="24">22–37</td>
|
||
<td id="830A5D8A504100611593FC2EFC020868" box="[915,988,1008,1036]" gridcol="3" gridrow="16" pageId="23" pageNumber="24">24</td>
|
||
<td id="830A5D8A504100611267FC2EFB710868" box="[1127,1199,1008,1036]" gridcol="4" gridrow="16" pageId="23" pageNumber="24">1.95</td>
|
||
</tr>
|
||
<tr id="C0DB34F6504100611765FBC5FB710853" box="[357,1199,1051,1079]" gridrow="17" pageId="23" pageNumber="24">
|
||
<th id="830A5D8A504100611765FBC5FE360853" box="[357,488,1051,1079]" gridcol="0" gridrow="17" pageId="23" pageNumber="24">MD4 (ac)</th>
|
||
<td id="830A5D8A50410061143DFBC5FD930853" box="[573,589,1051,1079]" gridcol="1" gridrow="17" pageId="23" pageNumber="24">7</td>
|
||
<td id="830A5D8A5041006114A0FBC5FCF90853" box="[672,807,1051,1079]" gridcol="2" gridrow="17" pageId="23" pageNumber="24">41–63</td>
|
||
<td id="830A5D8A504100611593FBC5FC020853" box="[915,988,1051,1079]" gridcol="3" gridrow="17" pageId="23" pageNumber="24">54</td>
|
||
<td id="830A5D8A504100611267FBC5FB710853" box="[1127,1199,1051,1079]" gridcol="4" gridrow="17" pageId="23" pageNumber="24">7.72</td>
|
||
</tr>
|
||
<tr id="C0DB34F6504100611765FBE3FB71083C" box="[357,1199,1085,1112]" gridrow="18" pageId="23" pageNumber="24">
|
||
<th id="830A5D8A504100611765FBE3FE36083C" box="[357,488,1085,1112]" gridcol="0" gridrow="18" pageId="23" pageNumber="24">MD6 (ac)</th>
|
||
<td id="830A5D8A50410061143DFBE3FD93083C" box="[573,589,1085,1112]" gridcol="1" gridrow="18" pageId="23" pageNumber="24">7</td>
|
||
<td id="830A5D8A5041006114A0FBE3FCF9083C" box="[672,807,1085,1112]" gridcol="2" gridrow="18" pageId="23" pageNumber="24">59–88</td>
|
||
<td id="830A5D8A504100611593FBE3FC02083C" box="[915,988,1085,1112]" gridcol="3" gridrow="18" pageId="23" pageNumber="24">74</td>
|
||
<td id="830A5D8A504100611267FBE3FB71083C" box="[1127,1199,1085,1112]" gridcol="4" gridrow="18" pageId="23" pageNumber="24">10.80</td>
|
||
</tr>
|
||
<tr id="C0DB34F6504100611765FB81FB71081E" box="[357,1199,1119,1146]" gridrow="19" pageId="23" pageNumber="24">
|
||
<th id="830A5D8A504100611765FB81FE36081E" box="[357,488,1119,1146]" gridcol="0" gridrow="19" pageId="23" pageNumber="24">MD8 (ac)</th>
|
||
<td id="830A5D8A50410061143DFB81FD93081E" box="[573,589,1119,1146]" gridcol="1" gridrow="19" pageId="23" pageNumber="24">7</td>
|
||
<td id="830A5D8A5041006114A0FB81FCF9081E" box="[672,807,1119,1146]" gridcol="2" gridrow="19" pageId="23" pageNumber="24">64–87</td>
|
||
<td id="830A5D8A504100611593FB81FC02081E" box="[915,988,1119,1146]" gridcol="3" gridrow="19" pageId="23" pageNumber="24">75</td>
|
||
<td id="830A5D8A504100611267FB81FB71081E" box="[1127,1199,1119,1146]" gridcol="4" gridrow="19" pageId="23" pageNumber="24">10.68</td>
|
||
</tr>
|
||
<tr id="C0DB34F6504100611765FB55FB7108C1" box="[357,1199,1163,1189]" gridrow="20" pageId="23" pageNumber="24">
|
||
<th id="830A5D8A504100611765FB55FE3608C1" box="[357,488,1163,1189]" gridcol="0" gridrow="20" pageId="23" pageNumber="24">LV6 (ac)</th>
|
||
<td id="830A5D8A50410061143DFB55FD9308C1" box="[573,589,1163,1189]" gridcol="1" gridrow="20" pageId="23" pageNumber="24">6</td>
|
||
<td id="830A5D8A5041006114A0FB55FCF908C1" box="[672,807,1163,1189]" gridcol="2" gridrow="20" pageId="23" pageNumber="24">20–28</td>
|
||
<td id="830A5D8A504100611593FB55FC0208C1" box="[915,988,1163,1189]" gridcol="3" gridrow="20" pageId="23" pageNumber="24">24</td>
|
||
<td id="830A5D8A504100611267FB55FB7108C1" box="[1127,1199,1163,1189]" gridcol="4" gridrow="20" pageId="23" pageNumber="24">3.39</td>
|
||
</tr>
|
||
<tr id="C0DB34F6504100611765FB75FB7108A3" box="[357,1199,1195,1223]" gridrow="21" pageId="23" pageNumber="24">
|
||
<th id="830A5D8A504100611765FB75FE3608A3" box="[357,488,1195,1223]" gridcol="0" gridrow="21" pageId="23" pageNumber="24">LV7 (ac)</th>
|
||
<td id="830A5D8A50410061143DFB75FD9308A3" box="[573,589,1195,1223]" gridcol="1" gridrow="21" pageId="23" pageNumber="24">7</td>
|
||
<td id="830A5D8A5041006114A0FB75FCF908A3" box="[672,807,1195,1223]" gridcol="2" gridrow="21" pageId="23" pageNumber="24">21–28</td>
|
||
<td id="830A5D8A504100611593FB75FC0208A3" box="[915,988,1195,1223]" gridcol="3" gridrow="21" pageId="23" pageNumber="24">25</td>
|
||
<td id="830A5D8A504100611267FB75FB7108A3" box="[1127,1199,1195,1223]" gridcol="4" gridrow="21" pageId="23" pageNumber="24">2.93</td>
|
||
</tr>
|
||
<tr id="C0DB34F6504100611765FB10FB71088C" box="[357,1199,1230,1256]" gridrow="22" pageId="23" pageNumber="24">
|
||
<th id="830A5D8A504100611765FB10FE36088C" box="[357,488,1230,1256]" gridcol="0" gridrow="22" pageId="23" pageNumber="24">LV8 (ac)</th>
|
||
<td id="830A5D8A50410061143DFB10FD93088C" box="[573,589,1230,1256]" gridcol="1" gridrow="22" pageId="23" pageNumber="24">7</td>
|
||
<td id="830A5D8A5041006114A0FB10FCF9088C" box="[672,807,1230,1256]" gridcol="2" gridrow="22" pageId="23" pageNumber="24">24–30</td>
|
||
<td id="830A5D8A504100611593FB10FC02088C" box="[915,988,1230,1256]" gridcol="3" gridrow="22" pageId="23" pageNumber="24">27</td>
|
||
<td id="830A5D8A504100611267FB10FB71088C" box="[1127,1199,1230,1256]" gridcol="4" gridrow="22" pageId="23" pageNumber="24">2.29</td>
|
||
</tr>
|
||
<tr id="C0DB34F6504100611765FB2EFB71096E" box="[357,1199,1264,1290]" gridrow="23" pageId="23" pageNumber="24">
|
||
<th id="830A5D8A504100611765FB2EFE36096E" box="[357,488,1264,1290]" gridcol="0" gridrow="23" pageId="23" pageNumber="24">LV9 (ac)</th>
|
||
<td id="830A5D8A50410061143DFB2EFD93096E" box="[573,589,1264,1290]" gridcol="1" gridrow="23" pageId="23" pageNumber="24">7</td>
|
||
<td id="830A5D8A5041006114A0FB2EFCF9096E" box="[672,807,1264,1290]" gridcol="2" gridrow="23" pageId="23" pageNumber="24">23–33</td>
|
||
<td id="830A5D8A504100611593FB2EFC02096E" box="[915,988,1264,1290]" gridcol="3" gridrow="23" pageId="23" pageNumber="24">28</td>
|
||
<td id="830A5D8A504100611267FB2EFB71096E" box="[1127,1199,1264,1290]" gridcol="4" gridrow="23" pageId="23" pageNumber="24">3.99</td>
|
||
</tr>
|
||
<tr id="C0DB34F6504100611765FAC4FB710951" box="[357,1199,1306,1333]" gridrow="24" pageId="23" pageNumber="24">
|
||
<th id="830A5D8A504100611765FAC4FE360951" box="[357,488,1306,1333]" gridcol="0" gridrow="24" pageId="23" pageNumber="24">LTS</th>
|
||
<td id="830A5D8A50410061143DFAC4FD930951" box="[573,589,1306,1333]" gridcol="1" gridrow="24" pageId="23" pageNumber="24">7</td>
|
||
<td id="830A5D8A5041006114A0FAC4FCF90951" box="[672,807,1306,1333]" gridcol="2" gridrow="24" pageId="23" pageNumber="24">102–149</td>
|
||
<td id="830A5D8A504100611593FAC4FC020951" box="[915,988,1306,1333]" gridcol="3" gridrow="24" pageId="23" pageNumber="24">125</td>
|
||
<td id="830A5D8A504100611267FAC4FB710951" box="[1127,1199,1306,1333]" gridcol="4" gridrow="24" pageId="23" pageNumber="24">20.10</td>
|
||
</tr>
|
||
<tr id="C0DB34F6504100611765FAE5FB710933" box="[357,1199,1339,1367]" gridrow="25" pageId="23" pageNumber="24">
|
||
<th id="830A5D8A504100611765FAE5FE360933" box="[357,488,1339,1367]" gridcol="0" gridrow="25" pageId="23" pageNumber="24">LTS/TL</th>
|
||
<td id="830A5D8A50410061143DFAE5FD930933" box="[573,589,1339,1367]" gridcol="1" gridrow="25" pageId="23" pageNumber="24">7</td>
|
||
<td id="830A5D8A5041006114A0FAE5FCF90933" box="[672,807,1339,1367]" gridcol="2" gridrow="25" pageId="23" pageNumber="24">42.5–65.4%</td>
|
||
<td id="830A5D8A504100611593FAE5FC020933" box="[915,988,1339,1367]" gridcol="3" gridrow="25" pageId="23" pageNumber="24">53.8%</td>
|
||
<td id="830A5D8A504100611267FAE5FB710933" box="[1127,1199,1339,1367]" gridcol="4" gridrow="25" pageId="23" pageNumber="24">8.81%</td>
|
||
</tr>
|
||
<tr id="C0DB34F6504100611765FA83FB71091C" box="[357,1199,1373,1400]" gridrow="26" pageId="23" pageNumber="24">
|
||
<th id="830A5D8A504100611765FA83FE36091C" box="[357,488,1373,1400]" gridcol="0" gridrow="26" pageId="23" pageNumber="24">LTAS</th>
|
||
<td id="830A5D8A50410061143DFA83FD93091C" box="[573,589,1373,1400]" gridcol="1" gridrow="26" pageId="23" pageNumber="24">5</td>
|
||
<td id="830A5D8A5041006114A0FA83FCF9091C" box="[672,807,1373,1400]" gridcol="2" gridrow="26" pageId="23" pageNumber="24">34–43</td>
|
||
<td id="830A5D8A504100611593FA83FC02091C" box="[915,988,1373,1400]" gridcol="3" gridrow="26" pageId="23" pageNumber="24">39</td>
|
||
<td id="830A5D8A504100611267FA83FB71091C" box="[1127,1199,1373,1400]" gridcol="4" gridrow="26" pageId="23" pageNumber="24">4.04</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="2A94663C5041FF8916BDFA3AFD8A0B88" ID-DOI="http://doi.org/10.5281/zenodo.3817841" ID-Zenodo-Dep="3817841" httpUri="https://zenodo.org/record/3817841/files/figure.png" pageId="23" pageNumber="24" subCaptionStartIDs="23.[967,1138,1719,1745]" subCaptionStarts="Abbr" targetBox="[193,1393,264,2020]" targetPageId="24">
|
||
<paragraph id="7E5436B45041FF8916BDFA3AFD8A0B88" blockId="23.[189,1399,1508,2028]" pageId="23" pageNumber="24">
|
||
<emphasis id="4C9FEAA65041FF8916BDFA3AFEDB099A" bold="true" box="[189,261,1508,1534]" pageId="23" pageNumber="24">Fig. 9</emphasis>
|
||
(opposite page). Scanning electron micrographs showing overviews and details of
|
||
<taxonomicName id="B9EB4D375041FF8912E5FA3AFEBD0A45" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="23" pageNumber="24" phylum="Cephalorhyncha" rank="species" species="hamiltonorum" status="sp. nov.">
|
||
<emphasis id="4C9FEAA65041FF8912E5FA3AFEBD0A45" italics="true" pageId="23" pageNumber="24">Echinoderes hamiltonorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD5041FF89176BF9D6FE180A46" box="[363,454,1544,1570]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="4C9FEAA65041FF8917CDF9D9FE3C0A45" bold="true" box="[461,482,1543,1569]" pageId="23" pageNumber="24">A</emphasis>
|
||
. Lateral overview.
|
||
<emphasis id="4C9FEAA65041FF8914C7F9D6FD050A46" bold="true" box="[711,731,1544,1570]" pageId="23" pageNumber="24">B</emphasis>
|
||
. Ventral overview.
|
||
<emphasis id="4C9FEAA65041FF8915BFF9D9FC0A0A45" bold="true" box="[959,980,1543,1569]" pageId="23" pageNumber="24">C</emphasis>
|
||
. Mouth cone.
|
||
<emphasis id="4C9FEAA65041FF891280F9D6FB4B0A46" bold="true" box="[1152,1173,1544,1570]" pageId="23" pageNumber="24">D</emphasis>
|
||
. Introvert sector 8.
|
||
<emphasis id="4C9FEAA65041FF8916BDF9F5FF0F0A21" bold="true" box="[189,209,1579,1605]" pageId="23" pageNumber="24">E</emphasis>
|
||
. Segments 1 to 3, dorsolateral view; note that laterodorsal sensory spots are missing in this particular specimen.
|
||
<emphasis id="4C9FEAA65041FF89173CF990FE900A0C" bold="true" box="[316,334,1614,1640]" pageId="23" pageNumber="24">F</emphasis>
|
||
. Segments 1 to 3, ventral view.
|
||
<emphasis id="4C9FEAA65041FF8914D4F990FD350A0C" bold="true" box="[724,747,1614,1640]" pageId="23" pageNumber="24">G</emphasis>
|
||
. Detail showing lateral side of segment 8 without any indication of glandular cell outlets
|
||
<typeStatus id="A15088165041FF891454F9ACFD590AE8" box="[596,647,1650,1676]" pageId="23" pageNumber="24">type</typeStatus>
|
||
2.
|
||
<emphasis id="4C9FEAA65041FF8914AAF9AFFD1F0AEF" bold="true" box="[682,705,1649,1675]" pageId="23" pageNumber="24">H</emphasis>
|
||
. Segments 8 to 9, lateral view.
|
||
<emphasis id="4C9FEAA65041FF89122DF9AFFBE60AEF" bold="true" box="[1069,1080,1649,1675]" pageId="23" pageNumber="24">I</emphasis>
|
||
. Detail showing midlateral to lateroventral parts of segments 5 and 6.
|
||
<emphasis id="4C9FEAA65041FF8914B9F94BFD160ACB" bold="true" box="[697,712,1685,1711]" pageId="23" pageNumber="24">J</emphasis>
|
||
. ♂, segments 10 to 11, dorsolateral view.
|
||
<emphasis id="4C9FEAA65041FF8912BDF94BFB0A0ACB" bold="true" box="[1213,1236,1685,1711]" pageId="23" pageNumber="24">K</emphasis>
|
||
. ♂, segments 10 to 11, ventral view.
|
||
<emphasis id="4C9FEAA65041FF8917C9F966FE030AB6" bold="true" box="[457,477,1720,1746]" pageId="23" pageNumber="24">L</emphasis>
|
||
.
|
||
<specimenCount id="68EDFD3D5041FF8917EBF966FDDF0AB6" box="[491,513,1720,1746]" pageId="23" pageNumber="24" type="female">♀</specimenCount>
|
||
, segments 10 to 11, laterodorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet
|
||
<typeStatus id="A15088165041FF8917BDF905FE2E0A91" box="[445,496,1755,1781]" pageId="23" pageNumber="24">type</typeStatus>
|
||
2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdss = middorsal sensory spot; mlss = midlateral sensory spot; oos = outer oral style; pdss = paradorsal sensory spot; pe = penile spines; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet
|
||
<typeStatus id="A15088165041FF8916BDF8B6FF2E0BE6" box="[189,240,1896,1922]" pageId="23" pageNumber="24">type</typeStatus>
|
||
2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet
|
||
<typeStatus id="A15088165041FF891325F8B6FA860BE6" box="[1317,1368,1896,1922]" pageId="23" pageNumber="24">type</typeStatus>
|
||
2; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlgco1 = ventrolateral glandular cell outlet
|
||
<typeStatus id="A15088165041FF89145AF870FD530BAC" box="[602,653,1966,1992]" pageId="23" pageNumber="24">type</typeStatus>
|
||
1; vlss = ventrolateral sensory spot; vlt = ventrolateral tube; vmss = ventromedial sensory spot.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="2A94663C504FFF8716BDFED2FBC50D9D" ID-Table-UUID="2A94663C504FFF8716BDFED2FBC50D9D" httpUri="http://table.plazi.org/id/2A94663C504FFF8716BDFED2FBC50D9D" pageId="25" pageNumber="26" targetBox="[207,1376,558,995]" targetIsTable="true" targetPageId="25">
|
||
<paragraph id="7E5436B4504FFF8716BDFED2FBC50D9D" blockId="25.[189,1399,268,506]" pageId="25" pageNumber="26">
|
||
<emphasis id="4C9FEAA6504FFF8716BDFED2FEFD0D42" bold="true" box="[189,291,268,294]" pageId="25" pageNumber="26">Table 7.</emphasis>
|
||
Summary of nature and location of sensory spots, glandular cell outlets, tubes and spines arranged by series in
|
||
<taxonomicName id="B9EB4D37504FFF8717BCFEF1FD250D2D" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[444,763,303,329]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="25" pageNumber="26" phylum="Cephalorhyncha" rank="species" species="hamiltonorum" status="sp. nov.">
|
||
<emphasis id="4C9FEAA6504FFF8717BCFEF1FD250D2D" box="[444,763,303,329]" italics="true" pageId="25" pageNumber="26">Echinoderes hamiltonorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD504FFF871504FEEEFC810D2E" box="[772,863,304,330]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Abbreviations: LA = lateral accessory; LD = laterodorsal; LV = lateroventral; MD = middorsal; ML = midlateral; PD = paradorsal; PV = paraventral; SD = subdorsal; SL = sublateral; VL = ventrolateral; VM = ventromedial; ac = acicular spine; gco1/2 = glandular cell outlet type 1/2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; pe = penile spines; si = sieve plate; ss = sensory spot; tu = tube; (♀) = female and (♂) = male conditions of sexually dimorphic characters; * = character not present in all specimens.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="7E5436B4504FFF8714BCFDF0FBE50F87" pageId="25" pageNumber="26">
|
||
<table id="0CEBC414504F006116CFFDF0FABE0F87" box="[207,1376,558,995]" gridcols="12" gridrows="13" pageId="25" pageNumber="26">
|
||
<tr id="C0DB34F6504F006116CFFDF0FABE0E22" box="[207,1376,558,582]" gridrow="0" pageId="25" pageNumber="26" rowspan-0="1" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-3="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
||
<th id="830A5D8A504F00611483FDF0FCE10E22" box="[643,831,558,582]" colspan="2" colspanRight="1" gridcol="4" gridrow="0" pageId="25" pageNumber="26">
|
||
<emphasis id="4C9FEAA6504FFF8714BCFDF0FCC60E22" bold="true" box="[700,792,558,582]" pageId="25" pageNumber="26">Position</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="C0DB34F6504F006116CFFD85FABE0E10" box="[207,1376,603,628]" gridrow="1" pageId="25" pageNumber="26">
|
||
<th id="830A5D8A504F006116CFFD85FEF20E10" box="[207,300,603,628]" gridcol="0" gridrow="1" pageId="25" pageNumber="26">
|
||
<emphasis id="4C9FEAA6504FFF8716CFFD85FEF20E10" bold="true" box="[207,300,603,628]" pageId="25" pageNumber="26">Segment</emphasis>
|
||
</th>
|
||
<td id="830A5D8A504F00611742FD85FE7D0E10" box="[322,419,603,628]" gridcol="1" gridrow="1" pageId="25" pageNumber="26">
|
||
<emphasis id="4C9FEAA6504FFF871765FD85FE510E17" bold="true" box="[357,399,603,627]" pageId="25" pageNumber="26">MD</emphasis>
|
||
</td>
|
||
<td id="830A5D8A504F006117BFFD85FDCE0E10" box="[447,528,603,628]" gridcol="2" gridrow="1" pageId="25" pageNumber="26">
|
||
<emphasis id="4C9FEAA6504FFF8717D7FD85FE270E17" bold="true" box="[471,505,603,627]" pageId="25" pageNumber="26">PD</emphasis>
|
||
</td>
|
||
<td id="830A5D8A504F00611431FD85FDBC0E10" box="[561,610,603,628]" gridcol="3" gridrow="1" pageId="25" pageNumber="26">
|
||
<emphasis id="4C9FEAA6504FFF87143AFD85FD840E10" bold="true" box="[570,602,603,628]" pageId="25" pageNumber="26">SD</emphasis>
|
||
</td>
|
||
<td id="830A5D8A504F00611483FD85FD630E10" box="[643,701,603,628]" gridcol="4" gridrow="1" pageId="25" pageNumber="26">
|
||
<emphasis id="4C9FEAA6504FFF87149AFD85FD630E17" bold="true" box="[666,701,603,627]" pageId="25" pageNumber="26">LD</emphasis>
|
||
</td>
|
||
<td id="830A5D8A504F006114FBFD85FCE10E10" box="[763,831,603,628]" gridcol="5" gridrow="1" pageId="25" pageNumber="26">
|
||
<emphasis id="4C9FEAA6504FFF8714FBFD85FCFA0E17" bold="true" box="[763,804,603,627]" pageId="25" pageNumber="26">ML</emphasis>
|
||
</td>
|
||
<td id="830A5D8A504F00611552FD85FC510E10" box="[850,911,603,628]" gridcol="6" gridrow="1" pageId="25" pageNumber="26">
|
||
<emphasis id="4C9FEAA6504FFF871562FD85FC5F0E10" bold="true" box="[866,897,603,628]" pageId="25" pageNumber="26">SL</emphasis>
|
||
</td>
|
||
<td id="830A5D8A504F006115AFFD85FC280E10" box="[943,1014,603,628]" gridcol="7" gridrow="1" pageId="25" pageNumber="26">
|
||
<emphasis id="4C9FEAA6504FFF8715C2FD85FC3B0E10" bold="true" box="[962,997,603,628]" pageId="25" pageNumber="26">LA</emphasis>
|
||
</td>
|
||
<td id="830A5D8A504F0061121FFD85FBE00E10" box="[1055,1086,603,628]" gridcol="8" gridrow="1" pageId="25" pageNumber="26">
|
||
<emphasis id="4C9FEAA6504FFF87121FFD85FBE00E17" bold="true" box="[1055,1086,603,627]" pageId="25" pageNumber="26">LV</emphasis>
|
||
</td>
|
||
<td id="830A5D8A504F00611272FD85FB7D0E10" box="[1138,1187,603,628]" gridcol="9" gridrow="1" pageId="25" pageNumber="26">
|
||
<emphasis id="4C9FEAA6504FFF87127AFD85FB430E17" bold="true" box="[1146,1181,603,627]" pageId="25" pageNumber="26">VL</emphasis>
|
||
</td>
|
||
<td id="830A5D8A504F006112D4FD85FB200E10" box="[1236,1278,603,628]" gridcol="10" gridrow="1" pageId="25" pageNumber="26">
|
||
<emphasis id="4C9FEAA6504FFF8712D4FD85FB200E17" bold="true" box="[1236,1278,603,627]" pageId="25" pageNumber="26">VM</emphasis>
|
||
</td>
|
||
<td id="830A5D8A504F0061132FFD85FABE0E10" box="[1327,1376,603,628]" gridcol="11" gridrow="1" pageId="25" pageNumber="26">
|
||
<emphasis id="4C9FEAA6504FFF871337FD85FA870E17" bold="true" box="[1335,1369,603,627]" pageId="25" pageNumber="26">PV</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr id="C0DB34F6504F006116CFFD5DFABE0EFF" box="[207,1376,643,667]" gridrow="2" pageId="25" pageNumber="26" rowspan-10="1" rowspan-11="1" rowspan-2="1" rowspan-5="1" rowspan-7="1" rowspan-8="1">
|
||
<th id="830A5D8A504F006116CFFD5DFEF20EFF" box="[207,300,643,667]" gridcol="0" gridrow="2" pageId="25" pageNumber="26">1</th>
|
||
<td id="830A5D8A504F00611742FD5DFE7D0EFF" box="[322,419,643,667]" gridcol="1" gridrow="2" pageId="25" pageNumber="26">gco1</td>
|
||
<td id="830A5D8A504F00611431FD5DFDBC0EFF" box="[561,610,643,667]" gridcol="3" gridrow="2" pageId="25" pageNumber="26">ss</td>
|
||
<td id="830A5D8A504F00611483FD5DFD630EFF" box="[643,701,643,667]" gridcol="4" gridrow="2" pageId="25" pageNumber="26">ss*</td>
|
||
<td id="830A5D8A504F00611552FD5DFC510EFF" box="[850,911,643,667]" gridcol="6" gridrow="2" pageId="25" pageNumber="26">gco2</td>
|
||
<td id="830A5D8A504F00611272FD5DFB7D0EFF" box="[1138,1187,643,667]" gridcol="9" gridrow="2" pageId="25" pageNumber="26">gco1</td>
|
||
</tr>
|
||
<tr id="C0DB34F6504F006116CFFD7DFABE0EDF" box="[207,1376,675,699]" gridrow="3" pageId="25" pageNumber="26" rowspan-11="1" rowspan-2="1" rowspan-7="1" rowspan-8="1">
|
||
<th id="830A5D8A504F006116CFFD7DFEF20EDF" box="[207,300,675,699]" gridcol="0" gridrow="3" pageId="25" pageNumber="26">2</th>
|
||
<td id="830A5D8A504F00611742FD7DFE7D0EDF" box="[322,419,675,699]" gridcol="1" gridrow="3" pageId="25" pageNumber="26">gco1,ss</td>
|
||
<td id="830A5D8A504F00611431FD7DFDBC0EDF" box="[561,610,675,699]" gridcol="3" gridrow="3" pageId="25" pageNumber="26">gco2</td>
|
||
<td id="830A5D8A504F00611483FD7DFCE10EDF" box="[643,831,675,699]" colspan="2" colspanRight="1" gridcol="4" gridrow="3" pageId="25" pageNumber="26">ss, gco2</td>
|
||
<td id="830A5D8A504F00611552FD7DFC510EDF" box="[850,911,675,699]" gridcol="6" gridrow="3" pageId="25" pageNumber="26">gco2</td>
|
||
<td id="830A5D8A504F00611272FD7DFB7D0EDF" box="[1138,1187,675,699]" gridcol="9" gridrow="3" pageId="25" pageNumber="26">tu</td>
|
||
<td id="830A5D8A504F006112D4FD7DFB200EDF" box="[1236,1278,675,699]" gridcol="10" gridrow="3" pageId="25" pageNumber="26">ss</td>
|
||
</tr>
|
||
<tr id="C0DB34F6504F006116CFFD1AFABE0EB8" box="[207,1376,708,732]" gridrow="4" pageId="25" pageNumber="26" rowspan-10="1" rowspan-2="1" rowspan-4="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
||
<th id="830A5D8A504F006116CFFD1AFEF20EB8" box="[207,300,708,732]" gridcol="0" gridrow="4" pageId="25" pageNumber="26">3</th>
|
||
<td id="830A5D8A504F00611742FD1AFE7D0EB8" box="[322,419,708,732]" gridcol="1" gridrow="4" pageId="25" pageNumber="26">gco1</td>
|
||
<td id="830A5D8A504F00611431FD1AFDBC0EB8" box="[561,610,708,732]" gridcol="3" gridrow="4" pageId="25" pageNumber="26">ss</td>
|
||
<td id="830A5D8A504F006114FBFD1AFCE10EB8" box="[763,831,708,732]" gridcol="5" gridrow="4" pageId="25" pageNumber="26">ss</td>
|
||
<td id="830A5D8A504F0061132FFD1AFABE0EB8" box="[1327,1376,708,732]" gridcol="11" gridrow="4" pageId="25" pageNumber="26">gco1</td>
|
||
</tr>
|
||
<tr id="C0DB34F6504F006116CFFD3BFABE0E99" box="[207,1376,741,765]" gridrow="5" pageId="25" pageNumber="26" rowspan-10="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
|
||
<th id="830A5D8A504F006116CFFD3BFEF20E99" box="[207,300,741,765]" gridcol="0" gridrow="5" pageId="25" pageNumber="26">4</th>
|
||
<td id="830A5D8A504F00611742FD3BFE7D0E99" box="[322,419,741,765]" gridcol="1" gridrow="5" pageId="25" pageNumber="26">ac</td>
|
||
<td id="830A5D8A504F006117BFFD3BFDCE0E99" box="[447,528,741,765]" gridcol="2" gridrow="5" pageId="25" pageNumber="26">gco1</td>
|
||
<td id="830A5D8A504F0061132FFD3BFABE0E99" box="[1327,1376,741,765]" gridcol="11" gridrow="5" pageId="25" pageNumber="26">gco1</td>
|
||
</tr>
|
||
<tr id="C0DB34F6504F006116CFFCDBFABE0F7A" box="[207,1376,773,798]" gridrow="6" pageId="25" pageNumber="26" rowspan-10="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-6="1" rowspan-7="1" rowspan-9="1">
|
||
<th id="830A5D8A504F006116CFFCDBFEF20F7A" box="[207,300,773,798]" gridcol="0" gridrow="6" pageId="25" pageNumber="26">5</th>
|
||
<td id="830A5D8A504F00611742FCDBFE7D0F7A" box="[322,419,773,798]" gridcol="1" gridrow="6" pageId="25" pageNumber="26">gco1</td>
|
||
<td id="830A5D8A504F006114FBFCDBFCE10F7A" box="[763,831,773,798]" gridcol="5" gridrow="6" pageId="25" pageNumber="26">ss</td>
|
||
<td id="830A5D8A504F0061121FFCDBFBE00F7A" box="[1055,1086,773,798]" gridcol="8" gridrow="6" pageId="25" pageNumber="26">tu</td>
|
||
<td id="830A5D8A504F0061132FFCDBFABE0F7A" box="[1327,1376,773,798]" gridcol="11" gridrow="6" pageId="25" pageNumber="26">gco1</td>
|
||
</tr>
|
||
<tr id="C0DB34F6504F006116CFFCFBFABE0F5A" box="[207,1376,805,830]" gridrow="7" pageId="25" pageNumber="26" rowspan-3="1" rowspan-4="1" rowspan-6="1" rowspan-7="1" rowspan-9="1">
|
||
<th id="830A5D8A504F006116CFFCFBFEF20F5A" box="[207,300,805,830]" gridcol="0" gridrow="7" pageId="25" pageNumber="26">6</th>
|
||
<td id="830A5D8A504F00611742FCFBFE7D0F5A" box="[322,419,805,830]" gridcol="1" gridrow="7" pageId="25" pageNumber="26">ac</td>
|
||
<td id="830A5D8A504F006117BFFCFBFDCE0F5A" box="[447,528,805,830]" gridcol="2" gridrow="7" pageId="25" pageNumber="26">ss, gco1</td>
|
||
<td id="830A5D8A504F006114FBFCFBFCE10F5A" box="[763,831,805,830]" gridcol="5" gridrow="7" pageId="25" pageNumber="26">ss</td>
|
||
<td id="830A5D8A504F0061121FFCFBFBE00F5A" box="[1055,1086,805,830]" gridcol="8" gridrow="7" pageId="25" pageNumber="26">ac</td>
|
||
<td id="830A5D8A504F006112D4FCFBFB200F5A" box="[1236,1278,805,830]" gridcol="10" gridrow="7" pageId="25" pageNumber="26">ss</td>
|
||
<td id="830A5D8A504F0061132FFCFBFABE0F5A" box="[1327,1376,805,830]" gridcol="11" gridrow="7" pageId="25" pageNumber="26">gco1</td>
|
||
</tr>
|
||
<tr id="C0DB34F6504F006116CFFC99FABE0F3B" box="[207,1376,839,863]" gridrow="8" pageId="25" pageNumber="26" rowspan-3="1" rowspan-4="1" rowspan-6="1" rowspan-7="1" rowspan-9="1">
|
||
<th id="830A5D8A504F006116CFFC99FEF20F3B" box="[207,300,839,863]" gridcol="0" gridrow="8" pageId="25" pageNumber="26">7</th>
|
||
<td id="830A5D8A504F00611742FC99FE7D0F3B" box="[322,419,839,863]" gridcol="1" gridrow="8" pageId="25" pageNumber="26">gco1</td>
|
||
<td id="830A5D8A504F006117BFFC99FDCE0F3B" box="[447,528,839,863]" gridcol="2" gridrow="8" pageId="25" pageNumber="26">ss</td>
|
||
<td id="830A5D8A504F006114FBFC99FCE10F3B" box="[763,831,839,863]" gridcol="5" gridrow="8" pageId="25" pageNumber="26">ss</td>
|
||
<td id="830A5D8A504F0061121FFC99FBE00F3B" box="[1055,1086,839,863]" gridcol="8" gridrow="8" pageId="25" pageNumber="26">ac</td>
|
||
<td id="830A5D8A504F006112D4FC99FB200F3B" box="[1236,1278,839,863]" gridcol="10" gridrow="8" pageId="25" pageNumber="26">ss</td>
|
||
<td id="830A5D8A504F0061132FFC99FABE0F3B" box="[1327,1376,839,863]" gridcol="11" gridrow="8" pageId="25" pageNumber="26">gco1</td>
|
||
</tr>
|
||
<tr id="C0DB34F6504F006116CFFCB9FABE0F1B" box="[207,1376,871,895]" gridrow="9" pageId="25" pageNumber="26" rowspan-10="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-7="1" rowspan-9="1">
|
||
<th id="830A5D8A504F006116CFFCB9FEF20F1B" box="[207,300,871,895]" gridcol="0" gridrow="9" pageId="25" pageNumber="26">8</th>
|
||
<td id="830A5D8A504F00611742FCB9FE7D0F1B" box="[322,419,871,895]" gridcol="1" gridrow="9" pageId="25" pageNumber="26">ac</td>
|
||
<td id="830A5D8A504F006117BFFCB9FDCE0F1B" box="[447,528,871,895]" gridcol="2" gridrow="9" pageId="25" pageNumber="26">ss, gco1</td>
|
||
<td id="830A5D8A504F00611552FCB9FC510F1B" box="[850,911,871,895]" gridcol="6" gridrow="9" pageId="25" pageNumber="26">gco2*</td>
|
||
<td id="830A5D8A504F0061121FFCB9FBE00F1B" box="[1055,1086,871,895]" gridcol="8" gridrow="9" pageId="25" pageNumber="26">ac</td>
|
||
<td id="830A5D8A504F0061132FFCB9FABE0F1B" box="[1327,1376,871,895]" gridcol="11" gridrow="9" pageId="25" pageNumber="26">gco1</td>
|
||
</tr>
|
||
<tr id="C0DB34F6504F006116CFFC56FABE0FC5" box="[207,1376,904,929]" gridrow="10" pageId="25" pageNumber="26" rowspan-1="1" rowspan-10="1" rowspan-3="1" rowspan-5="1" rowspan-6="1">
|
||
<th id="830A5D8A504F006116CFFC56FEF20FC5" box="[207,300,904,929]" gridcol="0" gridrow="10" pageId="25" pageNumber="26">9</th>
|
||
<td id="830A5D8A504F006117BFFC56FDCE0FC5" box="[447,528,904,929]" gridcol="2" gridrow="10" pageId="25" pageNumber="26">ss, gco1</td>
|
||
<td id="830A5D8A504F00611483FC56FD630FC5" box="[643,701,904,929]" gridcol="4" gridrow="10" pageId="25" pageNumber="26">ss</td>
|
||
<td id="830A5D8A504F006115AFFC56FC280FC5" box="[943,1014,904,929]" gridcol="7" gridrow="10" pageId="25" pageNumber="26">si</td>
|
||
<td id="830A5D8A504F0061121FFC56FBE00FC5" box="[1055,1086,904,929]" gridcol="8" gridrow="10" pageId="25" pageNumber="26">ac</td>
|
||
<td id="830A5D8A504F00611272FC56FB7D0FC5" box="[1138,1187,904,929]" gridcol="9" gridrow="10" pageId="25" pageNumber="26">ss</td>
|
||
<td id="830A5D8A504F0061132FFC56FABE0FC5" box="[1327,1376,904,929]" gridcol="11" gridrow="10" pageId="25" pageNumber="26">gco1</td>
|
||
</tr>
|
||
<tr id="C0DB34F6504F006116CFFC74FABE0FA6" box="[207,1376,938,962]" gridrow="11" pageId="25" pageNumber="26" rowspan-10="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
|
||
<th id="830A5D8A504F006116CFFC74FEF20FA6" box="[207,300,938,962]" gridcol="0" gridrow="11" pageId="25" pageNumber="26">10</th>
|
||
<td id="830A5D8A504F00611742FC74FDCE0FA6" box="[322,528,938,962]" colspan="2" colspanRight="1" gridcol="1" gridrow="11" pageId="25" pageNumber="26">gco1, gco1 ss</td>
|
||
<td id="830A5D8A504F00611483FC74FD630FA6" box="[643,701,938,962]" gridcol="4" gridrow="11" pageId="25" pageNumber="26">tu</td>
|
||
<td id="830A5D8A504F00611272FC74FB7D0FA6" box="[1138,1187,938,962]" gridcol="9" gridrow="11" pageId="25" pageNumber="26">ss</td>
|
||
<td id="830A5D8A504F0061132FFC74FABE0FA6" box="[1327,1376,938,962]" gridcol="11" gridrow="11" pageId="25" pageNumber="26">gco1</td>
|
||
</tr>
|
||
<tr id="C0DB34F6504F006116CFFC14FABE0F87" box="[207,1376,970,995]" gridrow="12" pageId="25" pageNumber="26" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-3="1" rowspan-6="1" rowspan-9="1">
|
||
<th id="830A5D8A504F006116CFFC14FEF20F87" box="[207,300,970,995]" gridcol="0" gridrow="12" pageId="25" pageNumber="26">11</th>
|
||
<td id="830A5D8A504F006117BFFC14FDCE0F87" box="[447,528,970,995]" gridcol="2" gridrow="12" pageId="25" pageNumber="26">ss</td>
|
||
<td id="830A5D8A504F00611483FC14FCE10F87" box="[643,831,970,995]" colspan="2" colspanRight="1" gridcol="4" gridrow="12" pageId="25" pageNumber="26">3 x pe(♂)</td>
|
||
<td id="830A5D8A504F006115AFFC14FC280F87" box="[943,1014,970,995]" gridcol="7" gridrow="12" pageId="25" pageNumber="26">ltas(♀)</td>
|
||
<td id="830A5D8A504F0061121FFC14FBE00F87" box="[1055,1086,970,995]" gridcol="8" gridrow="12" pageId="25" pageNumber="26">lts</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="7E5436B4504FFF8716BDFBD6FBF5095F" blockId="25.[189,1399,1031,1340]" pageId="25" pageNumber="26">
|
||
Segment 2 consists of a complete cuticular ring. Pachycyclus of the anterior segment margin is relatively thin and not interrupted. Tubes present in ventrolateral positions (
|
||
<figureCitation id="E6D02A31504FFF8715D4FBF5FBE80821" box="[980,1078,1067,1093]" captionStart="Fig" captionStartId="21.[189,232,1753,1779]" captionTargetBox="[315,1235,269,1735]" captionTargetId="figure@21.[315,1273,265,1735]" captionTargetPageId="21" captionText="Fig. 7. Line art illustrations of Echinoderes hamiltonorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♂, segments 10 to 11, dorsal view. D. ♂, segments 10 to 11, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817837" httpUri="https://zenodo.org/record/3817837/files/figure.png" pageId="25" pageNumber="26">Figs 7B</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A31504FFF871245FBF5FBB40821" box="[1093,1130,1067,1093]" captionStart="Fig" captionStartId="22.[189,232,1611,1637]" captionTargetBox="[256,1332,265,1599]" captionTargetId="figure@22.[256,1332,265,1599]" captionTargetPageId="22" captionText="Fig. 8. Light micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov., holotype, ♀ (A–D, H) (NHMD-223916), paratype, ♂ (E–F) (NHMD-223918) and paratype, ♀ (G) (NHMD-223920). A. Ventral overview. B. Segments 1 to 5, dorsal view. C. Segments 1 to 5, ventral view. D. Segments 6 to 8, dorsal view. E. Segments 6 to 9, ventral view. F. Detail showing sublateral glandular cell outlets type 2 on segments 1 and 2. G. Segments 8 to 9, lateral view. H. Segments 10 to 11 in female, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; pdgco1 = paradorsal glandular cell outlet type 1; pdss = paradorsal sensory spot; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817839" httpUri="https://zenodo.org/record/3817839/files/figure.png" pageId="25" pageNumber="26">8C</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A31504FFF871279FBF5FB450821" box="[1145,1179,1067,1093]" captionStart="Fig" captionStartId="23.[189,232,1508,1534]" captionTargetId="figure@24.[189,1398,260,2026]" captionTargetPageId="24" captionText="Fig. 9 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov. A. Lateral overview. B. Ventral overview. C. Mouth cone. D. Introvert sector 8. E. Segments 1 to 3, dorsolateral view; note that laterodorsal sensory spots are missing in this particular specimen. F. Segments 1 to 3, ventral view. G. Detail showing lateral side of segment 8 without any indication of glandular cell outlets type 2. H. Segments 8 to 9, lateral view. I. Detail showing midlateral to lateroventral parts of segments 5 and 6. J. ♂, segments 10 to 11, dorsolateral view. K. ♂, segments 10 to 11, ventral view. L. ♀, segments 10 to 11, laterodorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdss = middorsal sensory spot; mlss = midlateral sensory spot; oos = outer oral style; pdss = paradorsal sensory spot; pe = penile spines; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlgco1 = ventrolateral glandular cell outlet type 1; vlss = ventrolateral sensory spot; vlt = ventrolateral tube; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817841" httpUri="https://zenodo.org/record/3817841/files/figure.png" pageId="25" pageNumber="26">9F</figureCitation>
|
||
), and very minute glandular cell outlets
|
||
<typeStatus id="A1508816504FFF8717C1FB90FE2A080C" box="[449,500,1102,1128]" pageId="25" pageNumber="26">type</typeStatus>
|
||
<quantity id="B9139B51504FFF8717FDFB90FDF3080C" box="[509,557,1102,1128]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="25" pageNumber="26" unit="in" value="2.0">2 in</quantity>
|
||
subdorsal, laterodorsal and sublateral (
|
||
<figureCitation id="E6D02A31504FFF871204FB90FB83080C" box="[1028,1117,1102,1128]" captionStart="Fig" captionStartId="22.[189,232,1611,1637]" captionTargetBox="[256,1332,265,1599]" captionTargetId="figure@22.[256,1332,265,1599]" captionTargetPageId="22" captionText="Fig. 8. Light micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov., holotype, ♀ (A–D, H) (NHMD-223916), paratype, ♂ (E–F) (NHMD-223918) and paratype, ♀ (G) (NHMD-223920). A. Ventral overview. B. Segments 1 to 5, dorsal view. C. Segments 1 to 5, ventral view. D. Segments 6 to 8, dorsal view. E. Segments 6 to 9, ventral view. F. Detail showing sublateral glandular cell outlets type 2 on segments 1 and 2. G. Segments 8 to 9, lateral view. H. Segments 10 to 11 in female, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; pdgco1 = paradorsal glandular cell outlet type 1; pdss = paradorsal sensory spot; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817839" httpUri="https://zenodo.org/record/3817839/files/figure.png" pageId="25" pageNumber="26">Fig. 8F</figureCitation>
|
||
) positions (
|
||
<figureCitation id="E6D02A31504FFF8712EBFB90FAE9080C" box="[1259,1335,1102,1128]" captionStart="Fig" captionStartId="21.[189,232,1753,1779]" captionTargetBox="[315,1235,269,1735]" captionTargetId="figure@21.[315,1273,265,1735]" captionTargetPageId="21" captionText="Fig. 7. Line art illustrations of Echinoderes hamiltonorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♂, segments 10 to 11, dorsal view. D. ♂, segments 10 to 11, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817837" httpUri="https://zenodo.org/record/3817837/files/figure.png" pageId="25" pageNumber="26">Figs 7</figureCitation>
|
||
A–B, 8B–C, 9E–F). Sensory spots are located in middorsal, laterodorsal and ventromedial positions (
|
||
<figureCitation id="E6D02A31504FFF87130AFBAFFA8F08EF" box="[1290,1361,1137,1163]" captionStart="Fig" captionStartId="21.[189,232,1753,1779]" captionTargetBox="[315,1235,269,1735]" captionTargetId="figure@21.[315,1273,265,1735]" captionTargetPageId="21" captionText="Fig. 7. Line art illustrations of Echinoderes hamiltonorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♂, segments 10 to 11, dorsal view. D. ♂, segments 10 to 11, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817837" httpUri="https://zenodo.org/record/3817837/files/figure.png" pageId="25" pageNumber="26">Figs 7</figureCitation>
|
||
A– B, 9E–F). Glandular cell outlet
|
||
<typeStatus id="A1508816504FFF87142EFB4BFDBF08CB" box="[558,609,1173,1199]" pageId="25" pageNumber="26">type</typeStatus>
|
||
1 present in middorsal position only, near anterior segment margin. Secondary pectinate fringe present near anterior segment margin of this and the following segments, but it is usually covered by the preceding segment. On this and the following eight segments, the cuticular hairs are bracteate. Fairly long hairs densely cover the segment. The posterior segment margin is nearly straight, terminating in a pectinate fringe with long and flexible fringe tips.
|
||
</paragraph>
|
||
<paragraph id="7E5436B4504FFF8716BDFABDFD1E0A48" blockId="25.[189,1399,1378,1580]" pageId="25" pageNumber="26">
|
||
Segment 3, and remaining segments, consisting of one tergal and two sternal plates (
|
||
<figureCitation id="E6D02A31504FFF87129CFABCFB220919" box="[1180,1276,1378,1405]" captionStart="Fig" captionStartId="21.[189,232,1753,1779]" captionTargetBox="[315,1235,269,1735]" captionTargetId="figure@21.[315,1273,265,1735]" captionTargetPageId="21" captionText="Fig. 7. Line art illustrations of Echinoderes hamiltonorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♂, segments 10 to 11, dorsal view. D. ♂, segments 10 to 11, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817837" httpUri="https://zenodo.org/record/3817837/files/figure.png" pageId="25" pageNumber="26">Figs 7B</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A31504FFF871308FABDFAAF0919" box="[1288,1393,1379,1405]" captionStart="Fig" captionStartId="22.[189,232,1611,1637]" captionTargetBox="[256,1332,265,1599]" captionTargetId="figure@22.[256,1332,265,1599]" captionTargetPageId="22" captionText="Fig. 8. Light micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov., holotype, ♀ (A–D, H) (NHMD-223916), paratype, ♂ (E–F) (NHMD-223918) and paratype, ♀ (G) (NHMD-223920). A. Ventral overview. B. Segments 1 to 5, dorsal view. C. Segments 1 to 5, ventral view. D. Segments 6 to 8, dorsal view. E. Segments 6 to 9, ventral view. F. Detail showing sublateral glandular cell outlets type 2 on segments 1 and 2. G. Segments 8 to 9, lateral view. H. Segments 10 to 11 in female, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; pdgco1 = paradorsal glandular cell outlet type 1; pdss = paradorsal sensory spot; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817839" httpUri="https://zenodo.org/record/3817839/files/figure.png" pageId="25" pageNumber="26">8A, C, E</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A31504FFF8716BDFA58FF3D09C4" box="[189,227,1414,1440]" captionStart="Fig" captionStartId="23.[189,232,1508,1534]" captionTargetId="figure@24.[189,1398,260,2026]" captionTargetPageId="24" captionText="Fig. 9 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov. A. Lateral overview. B. Ventral overview. C. Mouth cone. D. Introvert sector 8. E. Segments 1 to 3, dorsolateral view; note that laterodorsal sensory spots are missing in this particular specimen. F. Segments 1 to 3, ventral view. G. Detail showing lateral side of segment 8 without any indication of glandular cell outlets type 2. H. Segments 8 to 9, lateral view. I. Detail showing midlateral to lateroventral parts of segments 5 and 6. J. ♂, segments 10 to 11, dorsolateral view. K. ♂, segments 10 to 11, ventral view. L. ♀, segments 10 to 11, laterodorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdss = middorsal sensory spot; mlss = midlateral sensory spot; oos = outer oral style; pdss = paradorsal sensory spot; pe = penile spines; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlgco1 = ventrolateral glandular cell outlet type 1; vlss = ventrolateral sensory spot; vlt = ventrolateral tube; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817841" httpUri="https://zenodo.org/record/3817841/files/figure.png" pageId="25" pageNumber="26">9B</figureCitation>
|
||
). Pachycyclus of the anterior segment margin is thin and interrupted at the tergosternal junctions and middorsally. Sensory spots present in subdorsal and midlateral positions (
|
||
<figureCitation id="E6D02A31504FFF871208FA77FBB609A7" box="[1032,1128,1449,1475]" captionStart="Fig" captionStartId="21.[189,232,1753,1779]" captionTargetBox="[315,1235,269,1735]" captionTargetId="figure@21.[315,1273,265,1735]" captionTargetPageId="21" captionText="Fig. 7. Line art illustrations of Echinoderes hamiltonorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♂, segments 10 to 11, dorsal view. D. ♂, segments 10 to 11, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817837" httpUri="https://zenodo.org/record/3817837/files/figure.png" pageId="25" pageNumber="26">Figs 7A</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A31504FFF871271FA77FB4809A7" box="[1137,1174,1449,1475]" captionStart="Fig" captionStartId="23.[189,232,1508,1534]" captionTargetId="figure@24.[189,1398,260,2026]" captionTargetPageId="24" captionText="Fig. 9 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov. A. Lateral overview. B. Ventral overview. C. Mouth cone. D. Introvert sector 8. E. Segments 1 to 3, dorsolateral view; note that laterodorsal sensory spots are missing in this particular specimen. F. Segments 1 to 3, ventral view. G. Detail showing lateral side of segment 8 without any indication of glandular cell outlets type 2. H. Segments 8 to 9, lateral view. I. Detail showing midlateral to lateroventral parts of segments 5 and 6. J. ♂, segments 10 to 11, dorsolateral view. K. ♂, segments 10 to 11, ventral view. L. ♀, segments 10 to 11, laterodorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdss = middorsal sensory spot; mlss = midlateral sensory spot; oos = outer oral style; pdss = paradorsal sensory spot; pe = penile spines; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlgco1 = ventrolateral glandular cell outlet type 1; vlss = ventrolateral sensory spot; vlt = ventrolateral tube; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817841" httpUri="https://zenodo.org/record/3817841/files/figure.png" pageId="25" pageNumber="26">9E</figureCitation>
|
||
), and glandular cell outlets
|
||
<typeStatus id="A1508816504FFF871713FA13FE980983" box="[275,326,1485,1511]" pageId="25" pageNumber="26">type</typeStatus>
|
||
<quantity id="B9139B51504FFF87174DFA12FEA40982" box="[333,378,1484,1510]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="25" pageNumber="26" unit="in" value="1.0">1 in</quantity>
|
||
middorsal and paraventral positions. Posterior segment margin straight, terminating in pectinate fringe with long and flexible fringe tips. Cuticular hairs cover tergal plate, except in middorsal position, and lateral halves of sternal plates.
|
||
</paragraph>
|
||
<paragraph id="7E5436B4504FFF8716BDF98AFBB00AD1" blockId="25.[189,1398,1620,1717]" pageId="25" pageNumber="26">
|
||
Segment 4 with acicular spine in middorsal position (
|
||
<figureCitation id="E6D02A31504FFF871551F98AFC690A0A" box="[849,951,1620,1646]" captionStart="Fig" captionStartId="21.[189,232,1753,1779]" captionTargetBox="[315,1235,269,1735]" captionTargetId="figure@21.[315,1273,265,1735]" captionTargetPageId="21" captionText="Fig. 7. Line art illustrations of Echinoderes hamiltonorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♂, segments 10 to 11, dorsal view. D. ♂, segments 10 to 11, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817837" httpUri="https://zenodo.org/record/3817837/files/figure.png" pageId="25" pageNumber="26">Figs 7A</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A31504FFF8715C7F98AFC320A0A" box="[967,1004,1620,1646]" captionStart="Fig" captionStartId="22.[189,232,1611,1637]" captionTargetBox="[256,1332,265,1599]" captionTargetId="figure@22.[256,1332,265,1599]" captionTargetPageId="22" captionText="Fig. 8. Light micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov., holotype, ♀ (A–D, H) (NHMD-223916), paratype, ♂ (E–F) (NHMD-223918) and paratype, ♀ (G) (NHMD-223920). A. Ventral overview. B. Segments 1 to 5, dorsal view. C. Segments 1 to 5, ventral view. D. Segments 6 to 8, dorsal view. E. Segments 6 to 9, ventral view. F. Detail showing sublateral glandular cell outlets type 2 on segments 1 and 2. G. Segments 8 to 9, lateral view. H. Segments 10 to 11 in female, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; pdgco1 = paradorsal glandular cell outlet type 1; pdss = paradorsal sensory spot; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817839" httpUri="https://zenodo.org/record/3817839/files/figure.png" pageId="25" pageNumber="26">8B</figureCitation>
|
||
). Sensory spots not present, but glandular cell outlets
|
||
<typeStatus id="A1508816504FFF8717C9F9A6FE220AF6" box="[457,508,1656,1682]" pageId="25" pageNumber="26">type</typeStatus>
|
||
1 present in paradorsal (
|
||
<figureCitation id="E6D02A31504FFF871536F9A9FC4B0AF5" box="[822,917,1655,1681]" captionStart="Fig" captionStartId="22.[189,232,1611,1637]" captionTargetBox="[256,1332,265,1599]" captionTargetId="figure@22.[256,1332,265,1599]" captionTargetPageId="22" captionText="Fig. 8. Light micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov., holotype, ♀ (A–D, H) (NHMD-223916), paratype, ♂ (E–F) (NHMD-223918) and paratype, ♀ (G) (NHMD-223920). A. Ventral overview. B. Segments 1 to 5, dorsal view. C. Segments 1 to 5, ventral view. D. Segments 6 to 8, dorsal view. E. Segments 6 to 9, ventral view. F. Detail showing sublateral glandular cell outlets type 2 on segments 1 and 2. G. Segments 8 to 9, lateral view. H. Segments 10 to 11 in female, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; pdgco1 = paradorsal glandular cell outlet type 1; pdss = paradorsal sensory spot; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817839" httpUri="https://zenodo.org/record/3817839/files/figure.png" pageId="25" pageNumber="26">Fig. 8B</figureCitation>
|
||
) and paraventral positions. Pachycycli, pectinate fringe of posterior margin and cuticular hairs as on preceding segment.
|
||
</paragraph>
|
||
<paragraph id="7E5436B4504FFF8716BDF902FB230B59" blockId="25.[189,1399,1756,1853]" pageId="25" pageNumber="26">
|
||
Segment 5 with a pair of minute tubes in lateroventral positions (
|
||
<figureCitation id="E6D02A31504FFF8715AFF902FBD00A92" box="[943,1038,1756,1782]" captionStart="Fig" captionStartId="21.[189,232,1753,1779]" captionTargetBox="[315,1235,269,1735]" captionTargetId="figure@21.[315,1273,265,1735]" captionTargetPageId="21" captionText="Fig. 7. Line art illustrations of Echinoderes hamiltonorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♂, segments 10 to 11, dorsal view. D. ♂, segments 10 to 11, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817837" httpUri="https://zenodo.org/record/3817837/files/figure.png" pageId="25" pageNumber="26">Figs 7B</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A31504FFF871219F902FBE00A92" box="[1049,1086,1756,1782]" captionStart="Fig" captionStartId="22.[189,232,1611,1637]" captionTargetBox="[256,1332,265,1599]" captionTargetId="figure@22.[256,1332,265,1599]" captionTargetPageId="22" captionText="Fig. 8. Light micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov., holotype, ♀ (A–D, H) (NHMD-223916), paratype, ♂ (E–F) (NHMD-223918) and paratype, ♀ (G) (NHMD-223920). A. Ventral overview. B. Segments 1 to 5, dorsal view. C. Segments 1 to 5, ventral view. D. Segments 6 to 8, dorsal view. E. Segments 6 to 9, ventral view. F. Detail showing sublateral glandular cell outlets type 2 on segments 1 and 2. G. Segments 8 to 9, lateral view. H. Segments 10 to 11 in female, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; pdgco1 = paradorsal glandular cell outlet type 1; pdss = paradorsal sensory spot; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817839" httpUri="https://zenodo.org/record/3817839/files/figure.png" pageId="25" pageNumber="26">8B</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A31504FFF87124AF902FBBB0A92" box="[1098,1125,1756,1782]" captionStart="Fig" captionStartId="23.[189,232,1508,1534]" captionTargetId="figure@24.[189,1398,260,2026]" captionTargetPageId="24" captionText="Fig. 9 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov. A. Lateral overview. B. Ventral overview. C. Mouth cone. D. Introvert sector 8. E. Segments 1 to 3, dorsolateral view; note that laterodorsal sensory spots are missing in this particular specimen. F. Segments 1 to 3, ventral view. G. Detail showing lateral side of segment 8 without any indication of glandular cell outlets type 2. H. Segments 8 to 9, lateral view. I. Detail showing midlateral to lateroventral parts of segments 5 and 6. J. ♂, segments 10 to 11, dorsolateral view. K. ♂, segments 10 to 11, ventral view. L. ♀, segments 10 to 11, laterodorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdss = middorsal sensory spot; mlss = midlateral sensory spot; oos = outer oral style; pdss = paradorsal sensory spot; pe = penile spines; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlgco1 = ventrolateral glandular cell outlet type 1; vlss = ventrolateral sensory spot; vlt = ventrolateral tube; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817841" httpUri="https://zenodo.org/record/3817841/files/figure.png" pageId="25" pageNumber="26">9I</figureCitation>
|
||
). Sensory spots present in midlateral positions (
|
||
<figureCitation id="E6D02A31504FFF8717D1F921FDFC0B7D" box="[465,546,1791,1817]" captionStart="Fig" captionStartId="23.[189,232,1508,1534]" captionTargetId="figure@24.[189,1398,260,2026]" captionTargetPageId="24" captionText="Fig. 9 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov. A. Lateral overview. B. Ventral overview. C. Mouth cone. D. Introvert sector 8. E. Segments 1 to 3, dorsolateral view; note that laterodorsal sensory spots are missing in this particular specimen. F. Segments 1 to 3, ventral view. G. Detail showing lateral side of segment 8 without any indication of glandular cell outlets type 2. H. Segments 8 to 9, lateral view. I. Detail showing midlateral to lateroventral parts of segments 5 and 6. J. ♂, segments 10 to 11, dorsolateral view. K. ♂, segments 10 to 11, ventral view. L. ♀, segments 10 to 11, laterodorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdss = middorsal sensory spot; mlss = midlateral sensory spot; oos = outer oral style; pdss = paradorsal sensory spot; pe = penile spines; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlgco1 = ventrolateral glandular cell outlet type 1; vlss = ventrolateral sensory spot; vlt = ventrolateral tube; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817841" httpUri="https://zenodo.org/record/3817841/files/figure.png" pageId="25" pageNumber="26">Fig. 9I</figureCitation>
|
||
), and glandular cell outlets
|
||
<typeStatus id="A1508816504FFF871563F921FC480B7D" box="[867,918,1791,1817]" pageId="25" pageNumber="26">type</typeStatus>
|
||
<quantity id="B9139B51504FFF87159CF921FC160B7D" box="[924,968,1791,1817]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="25" pageNumber="26" unit="in" value="1.0">1 in</quantity>
|
||
middorsal and paraventral positions. Pachycycli, pectinate fringe of posterior margin and cuticular hairs as on preceding segment.
|
||
</paragraph>
|
||
<paragraph id="7E5436B4504FFF8716BDF8BAFCFE0B8C" blockId="25.[189,1399,1891,2024]" pageId="25" pageNumber="26">
|
||
Segment 6 with acicular spines in middorsal and lateroventral positions (
|
||
<figureCitation id="E6D02A31504FFF871208F8BDFB8E0B1A" box="[1032,1104,1891,1918]" captionStart="Fig" captionStartId="21.[189,232,1753,1779]" captionTargetBox="[315,1235,269,1735]" captionTargetId="figure@21.[315,1273,265,1735]" captionTargetPageId="21" captionText="Fig. 7. Line art illustrations of Echinoderes hamiltonorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♂, segments 10 to 11, dorsal view. D. ♂, segments 10 to 11, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817837" httpUri="https://zenodo.org/record/3817837/files/figure.png" pageId="25" pageNumber="26">Figs 7</figureCitation>
|
||
A–B, 8D–E, 9I). Sensory spots present in paradorsal (
|
||
<figureCitation id="E6D02A31504FFF871403F859FD810BC5" box="[515,607,1927,1953]" captionStart="Fig" captionStartId="22.[189,232,1611,1637]" captionTargetBox="[256,1332,265,1599]" captionTargetId="figure@22.[256,1332,265,1599]" captionTargetPageId="22" captionText="Fig. 8. Light micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov., holotype, ♀ (A–D, H) (NHMD-223916), paratype, ♂ (E–F) (NHMD-223918) and paratype, ♀ (G) (NHMD-223920). A. Ventral overview. B. Segments 1 to 5, dorsal view. C. Segments 1 to 5, ventral view. D. Segments 6 to 8, dorsal view. E. Segments 6 to 9, ventral view. F. Detail showing sublateral glandular cell outlets type 2 on segments 1 and 2. G. Segments 8 to 9, lateral view. H. Segments 10 to 11 in female, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; pdgco1 = paradorsal glandular cell outlet type 1; pdss = paradorsal sensory spot; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817839" httpUri="https://zenodo.org/record/3817839/files/figure.png" pageId="25" pageNumber="26">Fig. 8D</figureCitation>
|
||
), midlateral (
|
||
<figureCitation id="E6D02A31504FFF8714FBF859FC950BC5" box="[763,843,1927,1953]" captionStart="Fig" captionStartId="23.[189,232,1508,1534]" captionTargetId="figure@24.[189,1398,260,2026]" captionTargetPageId="24" captionText="Fig. 9 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov. A. Lateral overview. B. Ventral overview. C. Mouth cone. D. Introvert sector 8. E. Segments 1 to 3, dorsolateral view; note that laterodorsal sensory spots are missing in this particular specimen. F. Segments 1 to 3, ventral view. G. Detail showing lateral side of segment 8 without any indication of glandular cell outlets type 2. H. Segments 8 to 9, lateral view. I. Detail showing midlateral to lateroventral parts of segments 5 and 6. J. ♂, segments 10 to 11, dorsolateral view. K. ♂, segments 10 to 11, ventral view. L. ♀, segments 10 to 11, laterodorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdss = middorsal sensory spot; mlss = midlateral sensory spot; oos = outer oral style; pdss = paradorsal sensory spot; pe = penile spines; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlgco1 = ventrolateral glandular cell outlet type 1; vlss = ventrolateral sensory spot; vlt = ventrolateral tube; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817841" httpUri="https://zenodo.org/record/3817841/files/figure.png" pageId="25" pageNumber="26">Fig. 9I</figureCitation>
|
||
) and ventromedial positions, and glandular cell outlets
|
||
<typeStatus id="A1508816504FFF871712F874FE9B0BA0" box="[274,325,1962,1988]" pageId="25" pageNumber="26">type</typeStatus>
|
||
<quantity id="B9139B51504FFF87174BF874FEA90BA0" box="[331,375,1962,1988]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="25" pageNumber="26" unit="in" value="1.0">1 in</quantity>
|
||
paradorsal (
|
||
<figureCitation id="E6D02A31504FFF871407F874FDBF0BA0" box="[519,609,1962,1988]" captionStart="Fig" captionStartId="22.[189,232,1611,1637]" captionTargetBox="[256,1332,265,1599]" captionTargetId="figure@22.[256,1332,265,1599]" captionTargetPageId="22" captionText="Fig. 8. Light micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov., holotype, ♀ (A–D, H) (NHMD-223916), paratype, ♂ (E–F) (NHMD-223918) and paratype, ♀ (G) (NHMD-223920). A. Ventral overview. B. Segments 1 to 5, dorsal view. C. Segments 1 to 5, ventral view. D. Segments 6 to 8, dorsal view. E. Segments 6 to 9, ventral view. F. Detail showing sublateral glandular cell outlets type 2 on segments 1 and 2. G. Segments 8 to 9, lateral view. H. Segments 10 to 11 in female, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; pdgco1 = paradorsal glandular cell outlet type 1; pdss = paradorsal sensory spot; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817839" httpUri="https://zenodo.org/record/3817839/files/figure.png" pageId="25" pageNumber="26">Fig. 8D</figureCitation>
|
||
) and paraventral positions. Pachycycli, pectinate fringe of posterior margin and cuticular hairs as on preceding segment.
|
||
</paragraph>
|
||
<paragraph id="7E5436B4504CFF8416BDFED5FBF70DD0" blockId="26.[189,1399,267,437]" pageId="26" pageNumber="27">
|
||
Segment 7 with acicular spines in lateroventral positions (
|
||
<figureCitation id="E6D02A31504CFF84157EFED5FC3F0D42" box="[894,993,267,294]" captionStart="Fig" captionStartId="21.[189,232,1753,1779]" captionTargetBox="[315,1235,269,1735]" captionTargetId="figure@21.[315,1273,265,1735]" captionTargetPageId="21" captionText="Fig. 7. Line art illustrations of Echinoderes hamiltonorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♂, segments 10 to 11, dorsal view. D. ♂, segments 10 to 11, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817837" httpUri="https://zenodo.org/record/3817837/files/figure.png" pageId="26" pageNumber="27">Figs 7B</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A31504CFF8415F0FED5FC230D41" box="[1008,1021,267,293]" captionStart="Fig" captionStartId="22.[189,232,1611,1637]" captionTargetBox="[256,1332,265,1599]" captionTargetId="figure@22.[256,1332,265,1599]" captionTargetPageId="22" captionText="Fig. 8. Light micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov., holotype, ♀ (A–D, H) (NHMD-223916), paratype, ♂ (E–F) (NHMD-223918) and paratype, ♀ (G) (NHMD-223920). A. Ventral overview. B. Segments 1 to 5, dorsal view. C. Segments 1 to 5, ventral view. D. Segments 6 to 8, dorsal view. E. Segments 6 to 9, ventral view. F. Detail showing sublateral glandular cell outlets type 2 on segments 1 and 2. G. Segments 8 to 9, lateral view. H. Segments 10 to 11 in female, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; pdgco1 = paradorsal glandular cell outlet type 1; pdss = paradorsal sensory spot; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817839" httpUri="https://zenodo.org/record/3817839/files/figure.png" pageId="26" pageNumber="27">8</figureCitation>
|
||
D–E). Sensory spots present in paradorsal (
|
||
<figureCitation id="E6D02A31504CFF841745FEF1FE7E0D2D" box="[325,416,303,329]" captionStart="Fig" captionStartId="22.[189,232,1611,1637]" captionTargetBox="[256,1332,265,1599]" captionTargetId="figure@22.[256,1332,265,1599]" captionTargetPageId="22" captionText="Fig. 8. Light micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov., holotype, ♀ (A–D, H) (NHMD-223916), paratype, ♂ (E–F) (NHMD-223918) and paratype, ♀ (G) (NHMD-223920). A. Ventral overview. B. Segments 1 to 5, dorsal view. C. Segments 1 to 5, ventral view. D. Segments 6 to 8, dorsal view. E. Segments 6 to 9, ventral view. F. Detail showing sublateral glandular cell outlets type 2 on segments 1 and 2. G. Segments 8 to 9, lateral view. H. Segments 10 to 11 in female, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; pdgco1 = paradorsal glandular cell outlet type 1; pdss = paradorsal sensory spot; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817839" httpUri="https://zenodo.org/record/3817839/files/figure.png" pageId="26" pageNumber="27">Fig. 8D</figureCitation>
|
||
), midlateral and ventromedial positions, and glandular cell outlets
|
||
<typeStatus id="A1508816504CFF84129AFEEEFB130D2E" box="[1178,1229,304,330]" pageId="26" pageNumber="27">type</typeStatus>
|
||
<quantity id="B9139B51504CFF8412D2FEF1FB220D2D" box="[1234,1276,303,329]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="26" pageNumber="27" unit="in" value="1.0">1 in</quantity>
|
||
middorsal (
|
||
<figureCitation id="E6D02A31504CFF8416C6FE8DFEFE0D09" box="[198,288,339,365]" captionStart="Fig" captionStartId="22.[189,232,1611,1637]" captionTargetBox="[256,1332,265,1599]" captionTargetId="figure@22.[256,1332,265,1599]" captionTargetPageId="22" captionText="Fig. 8. Light micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov., holotype, ♀ (A–D, H) (NHMD-223916), paratype, ♂ (E–F) (NHMD-223918) and paratype, ♀ (G) (NHMD-223920). A. Ventral overview. B. Segments 1 to 5, dorsal view. C. Segments 1 to 5, ventral view. D. Segments 6 to 8, dorsal view. E. Segments 6 to 9, ventral view. F. Detail showing sublateral glandular cell outlets type 2 on segments 1 and 2. G. Segments 8 to 9, lateral view. H. Segments 10 to 11 in female, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; pdgco1 = paradorsal glandular cell outlet type 1; pdss = paradorsal sensory spot; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817839" httpUri="https://zenodo.org/record/3817839/files/figure.png" pageId="26" pageNumber="27">Fig. 8D</figureCitation>
|
||
) and paraventral positions. Pachycycli and pectinate fringe of posterior margin as on preceding segment; cuticular hair covering also as on preceding segment, but with middorsal hairless area being broader, posteriorly expanding into paradorsal and subdorsal areas as well.
|
||
</paragraph>
|
||
<paragraph id="7E5436B4504CFF8416BDFE02FC230EA9" blockId="26.[189,1399,476,717]" pageId="26" pageNumber="27">
|
||
Segment 8 with acicular spines in middorsal and lateroventral positions (
|
||
<figureCitation id="E6D02A31504CFF841231FE02FBA00D92" box="[1073,1150,476,502]" captionStart="Fig" captionStartId="21.[189,232,1753,1779]" captionTargetBox="[315,1235,269,1735]" captionTargetId="figure@21.[315,1273,265,1735]" captionTargetPageId="21" captionText="Fig. 7. Line art illustrations of Echinoderes hamiltonorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♂, segments 10 to 11, dorsal view. D. ♂, segments 10 to 11, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817837" httpUri="https://zenodo.org/record/3817837/files/figure.png" pageId="26" pageNumber="27">Figs 7</figureCitation>
|
||
A–B, 8D–E, 9G–H). Some
|
||
<typeStatus id="A1508816504CFF841708FDDEFEA60E7E" box="[264,376,512,538]" pageId="26" pageNumber="27" type="paratype">paratypes</typeStatus>
|
||
with fairly large sublateral glandular cell outlets type 2 (
|
||
<figureCitation id="E6D02A31504CFF841217FDDEFBA60E7E" box="[1047,1144,512,538]" captionStart="Fig" captionStartId="21.[189,232,1753,1779]" captionTargetBox="[315,1235,269,1735]" captionTargetId="figure@21.[315,1273,265,1735]" captionTargetPageId="21" captionText="Fig. 7. Line art illustrations of Echinoderes hamiltonorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♂, segments 10 to 11, dorsal view. D. ♂, segments 10 to 11, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817837" httpUri="https://zenodo.org/record/3817837/files/figure.png" pageId="26" pageNumber="27">Figs 7B</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A31504CFF841285FDDEFB740E7E" box="[1157,1194,512,538]" captionStart="Fig" captionStartId="22.[189,232,1611,1637]" captionTargetBox="[256,1332,265,1599]" captionTargetId="figure@22.[256,1332,265,1599]" captionTargetPageId="22" captionText="Fig. 8. Light micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov., holotype, ♀ (A–D, H) (NHMD-223916), paratype, ♂ (E–F) (NHMD-223918) and paratype, ♀ (G) (NHMD-223920). A. Ventral overview. B. Segments 1 to 5, dorsal view. C. Segments 1 to 5, ventral view. D. Segments 6 to 8, dorsal view. E. Segments 6 to 9, ventral view. F. Detail showing sublateral glandular cell outlets type 2 on segments 1 and 2. G. Segments 8 to 9, lateral view. H. Segments 10 to 11 in female, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; pdgco1 = paradorsal glandular cell outlet type 1; pdss = paradorsal sensory spot; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817839" httpUri="https://zenodo.org/record/3817839/files/figure.png" pageId="26" pageNumber="27">8G</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A31504CFF8412B8FDDEFB030E7E" box="[1208,1245,512,538]" captionStart="Fig" captionStartId="23.[189,232,1508,1534]" captionTargetId="figure@24.[189,1398,260,2026]" captionTargetPageId="24" captionText="Fig. 9 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov. A. Lateral overview. B. Ventral overview. C. Mouth cone. D. Introvert sector 8. E. Segments 1 to 3, dorsolateral view; note that laterodorsal sensory spots are missing in this particular specimen. F. Segments 1 to 3, ventral view. G. Detail showing lateral side of segment 8 without any indication of glandular cell outlets type 2. H. Segments 8 to 9, lateral view. I. Detail showing midlateral to lateroventral parts of segments 5 and 6. J. ♂, segments 10 to 11, dorsolateral view. K. ♂, segments 10 to 11, ventral view. L. ♀, segments 10 to 11, laterodorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdss = middorsal sensory spot; mlss = midlateral sensory spot; oos = outer oral style; pdss = paradorsal sensory spot; pe = penile spines; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlgco1 = ventrolateral glandular cell outlet type 1; vlss = ventrolateral sensory spot; vlt = ventrolateral tube; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817841" httpUri="https://zenodo.org/record/3817841/files/figure.png" pageId="26" pageNumber="27">9H</figureCitation>
|
||
) present, but others (including the
|
||
<typeStatus id="A1508816504CFF8417BCFDFDFDFB0E5A" box="[444,549,547,574]" pageId="26" pageNumber="27" type="holotype">holotype</typeStatus>
|
||
) without such outlets (compare
|
||
<figureCitation id="E6D02A31504CFF8415A7FDFDFBDE0E5A" box="[935,1024,547,574]" captionStart="Fig" captionStartId="22.[189,232,1611,1637]" captionTargetBox="[256,1332,265,1599]" captionTargetId="figure@22.[256,1332,265,1599]" captionTargetPageId="22" captionText="Fig. 8. Light micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov., holotype, ♀ (A–D, H) (NHMD-223916), paratype, ♂ (E–F) (NHMD-223918) and paratype, ♀ (G) (NHMD-223920). A. Ventral overview. B. Segments 1 to 5, dorsal view. C. Segments 1 to 5, ventral view. D. Segments 6 to 8, dorsal view. E. Segments 6 to 9, ventral view. F. Detail showing sublateral glandular cell outlets type 2 on segments 1 and 2. G. Segments 8 to 9, lateral view. H. Segments 10 to 11 in female, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; pdgco1 = paradorsal glandular cell outlet type 1; pdss = paradorsal sensory spot; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817839" httpUri="https://zenodo.org/record/3817839/files/figure.png" pageId="26" pageNumber="27">Fig. 8E</figureCitation>
|
||
vs 8G and
|
||
<figureCitation id="E6D02A31504CFF84128CFDFDFB390E5A" box="[1164,1255,547,574]" captionStart="Fig" captionStartId="23.[189,232,1508,1534]" captionTargetId="figure@24.[189,1398,260,2026]" captionTargetPageId="24" captionText="Fig. 9 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov. A. Lateral overview. B. Ventral overview. C. Mouth cone. D. Introvert sector 8. E. Segments 1 to 3, dorsolateral view; note that laterodorsal sensory spots are missing in this particular specimen. F. Segments 1 to 3, ventral view. G. Detail showing lateral side of segment 8 without any indication of glandular cell outlets type 2. H. Segments 8 to 9, lateral view. I. Detail showing midlateral to lateroventral parts of segments 5 and 6. J. ♂, segments 10 to 11, dorsolateral view. K. ♂, segments 10 to 11, ventral view. L. ♀, segments 10 to 11, laterodorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdss = middorsal sensory spot; mlss = midlateral sensory spot; oos = outer oral style; pdss = paradorsal sensory spot; pe = penile spines; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlgco1 = ventrolateral glandular cell outlet type 1; vlss = ventrolateral sensory spot; vlt = ventrolateral tube; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817841" httpUri="https://zenodo.org/record/3817841/files/figure.png" pageId="26" pageNumber="27">Fig. 9H</figureCitation>
|
||
vs 9G); the outlets were present in eight females and one male, from stations 2, 3, 6 and 8, and were missing in three females and four males, from stations 3, 5 and 6. Sensory spots present in paradorsal positions (
|
||
<figureCitation id="E6D02A31504CFF84130CFDB5FAB60EE1" box="[1292,1384,619,645]" captionStart="Fig" captionStartId="22.[189,232,1611,1637]" captionTargetBox="[256,1332,265,1599]" captionTargetId="figure@22.[256,1332,265,1599]" captionTargetPageId="22" captionText="Fig. 8. Light micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov., holotype, ♀ (A–D, H) (NHMD-223916), paratype, ♂ (E–F) (NHMD-223918) and paratype, ♀ (G) (NHMD-223920). A. Ventral overview. B. Segments 1 to 5, dorsal view. C. Segments 1 to 5, ventral view. D. Segments 6 to 8, dorsal view. E. Segments 6 to 9, ventral view. F. Detail showing sublateral glandular cell outlets type 2 on segments 1 and 2. G. Segments 8 to 9, lateral view. H. Segments 10 to 11 in female, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; pdgco1 = paradorsal glandular cell outlet type 1; pdss = paradorsal sensory spot; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817839" httpUri="https://zenodo.org/record/3817839/files/figure.png" pageId="26" pageNumber="27">Fig. 8D</figureCitation>
|
||
), and glandular cell outlets type
|
||
<quantity id="B9139B51504CFF841423FD51FD910ECC" box="[547,591,654,681]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="26" pageNumber="27" unit="in" value="1.0">1 in</quantity>
|
||
paradorsal (
|
||
<figureCitation id="E6D02A31504CFF8414DFFD50FCE40ECD" box="[735,826,654,681]" captionStart="Fig" captionStartId="22.[189,232,1611,1637]" captionTargetBox="[256,1332,265,1599]" captionTargetId="figure@22.[256,1332,265,1599]" captionTargetPageId="22" captionText="Fig. 8. Light micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov., holotype, ♀ (A–D, H) (NHMD-223916), paratype, ♂ (E–F) (NHMD-223918) and paratype, ♀ (G) (NHMD-223920). A. Ventral overview. B. Segments 1 to 5, dorsal view. C. Segments 1 to 5, ventral view. D. Segments 6 to 8, dorsal view. E. Segments 6 to 9, ventral view. F. Detail showing sublateral glandular cell outlets type 2 on segments 1 and 2. G. Segments 8 to 9, lateral view. H. Segments 10 to 11 in female, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; pdgco1 = paradorsal glandular cell outlet type 1; pdss = paradorsal sensory spot; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817839" httpUri="https://zenodo.org/record/3817839/files/figure.png" pageId="26" pageNumber="27">Fig. 8D</figureCitation>
|
||
) and paraventral positions. Pachycycli, pectinate fringe of posterior margin and cuticular hairs as on preceding segment.
|
||
</paragraph>
|
||
<paragraph id="7E5436B4504CFF8416BDFD2AFC620FA5" blockId="26.[189,1399,756,961]" pageId="26" pageNumber="27">
|
||
Segment 9 with acicular spines in lateroventral positions (
|
||
<figureCitation id="E6D02A31504CFF841585FD2AFC370F6B" box="[901,1001,756,783]" captionStart="Fig" captionStartId="21.[189,232,1753,1779]" captionTargetBox="[315,1235,269,1735]" captionTargetId="figure@21.[315,1273,265,1735]" captionTargetPageId="21" captionText="Fig. 7. Line art illustrations of Echinoderes hamiltonorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♂, segments 10 to 11, dorsal view. D. ♂, segments 10 to 11, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817837" httpUri="https://zenodo.org/record/3817837/files/figure.png" pageId="26" pageNumber="27">Figs 7B</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A31504CFF8415F9FD2AFBC10F6A" box="[1017,1055,756,782]" captionStart="Fig" captionStartId="22.[189,232,1611,1637]" captionTargetBox="[256,1332,265,1599]" captionTargetId="figure@22.[256,1332,265,1599]" captionTargetPageId="22" captionText="Fig. 8. Light micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov., holotype, ♀ (A–D, H) (NHMD-223916), paratype, ♂ (E–F) (NHMD-223918) and paratype, ♀ (G) (NHMD-223920). A. Ventral overview. B. Segments 1 to 5, dorsal view. C. Segments 1 to 5, ventral view. D. Segments 6 to 8, dorsal view. E. Segments 6 to 9, ventral view. F. Detail showing sublateral glandular cell outlets type 2 on segments 1 and 2. G. Segments 8 to 9, lateral view. H. Segments 10 to 11 in female, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; pdgco1 = paradorsal glandular cell outlet type 1; pdss = paradorsal sensory spot; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817839" httpUri="https://zenodo.org/record/3817839/files/figure.png" pageId="26" pageNumber="27">8G</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A31504CFF841230FD2AFB890F6A" box="[1072,1111,756,782]" captionStart="Fig" captionStartId="23.[189,232,1508,1534]" captionTargetId="figure@24.[189,1398,260,2026]" captionTargetPageId="24" captionText="Fig. 9 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov. A. Lateral overview. B. Ventral overview. C. Mouth cone. D. Introvert sector 8. E. Segments 1 to 3, dorsolateral view; note that laterodorsal sensory spots are missing in this particular specimen. F. Segments 1 to 3, ventral view. G. Detail showing lateral side of segment 8 without any indication of glandular cell outlets type 2. H. Segments 8 to 9, lateral view. I. Detail showing midlateral to lateroventral parts of segments 5 and 6. J. ♂, segments 10 to 11, dorsolateral view. K. ♂, segments 10 to 11, ventral view. L. ♀, segments 10 to 11, laterodorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdss = middorsal sensory spot; mlss = midlateral sensory spot; oos = outer oral style; pdss = paradorsal sensory spot; pe = penile spines; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlgco1 = ventrolateral glandular cell outlet type 1; vlss = ventrolateral sensory spot; vlt = ventrolateral tube; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817841" httpUri="https://zenodo.org/record/3817841/files/figure.png" pageId="26" pageNumber="27">9H</figureCitation>
|
||
). Sensory spots present in paradorsal, laterodorsal (
|
||
<figureCitation id="E6D02A31504CFF841407FCC6FDB60F56" box="[519,616,792,818]" captionStart="Fig" captionStartId="23.[189,232,1508,1534]" captionTargetId="figure@24.[189,1398,260,2026]" captionTargetPageId="24" captionText="Fig. 9 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov. A. Lateral overview. B. Ventral overview. C. Mouth cone. D. Introvert sector 8. E. Segments 1 to 3, dorsolateral view; note that laterodorsal sensory spots are missing in this particular specimen. F. Segments 1 to 3, ventral view. G. Detail showing lateral side of segment 8 without any indication of glandular cell outlets type 2. H. Segments 8 to 9, lateral view. I. Detail showing midlateral to lateroventral parts of segments 5 and 6. J. ♂, segments 10 to 11, dorsolateral view. K. ♂, segments 10 to 11, ventral view. L. ♀, segments 10 to 11, laterodorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdss = middorsal sensory spot; mlss = midlateral sensory spot; oos = outer oral style; pdss = paradorsal sensory spot; pe = penile spines; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlgco1 = ventrolateral glandular cell outlet type 1; vlss = ventrolateral sensory spot; vlt = ventrolateral tube; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817841" httpUri="https://zenodo.org/record/3817841/files/figure.png" pageId="26" pageNumber="27">Fig. 9H</figureCitation>
|
||
), and ventrolateral positions, and glandular cell outlets
|
||
<typeStatus id="A1508816504CFF84130AFCC6FAE30F56" box="[1290,1341,792,818]" pageId="26" pageNumber="27">type</typeStatus>
|
||
<quantity id="B9139B51504CFF841347FCC6FAA90F56" box="[1351,1399,792,818]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="26" pageNumber="27" unit="in" value="1.0">1 in</quantity>
|
||
paradorsal (
|
||
<figureCitation id="E6D02A31504CFF84174BFCE2FE750F32" box="[331,427,828,854]" captionStart="Fig" captionStartId="22.[189,232,1611,1637]" captionTargetBox="[256,1332,265,1599]" captionTargetId="figure@22.[256,1332,265,1599]" captionTargetPageId="22" captionText="Fig. 8. Light micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov., holotype, ♀ (A–D, H) (NHMD-223916), paratype, ♂ (E–F) (NHMD-223918) and paratype, ♀ (G) (NHMD-223920). A. Ventral overview. B. Segments 1 to 5, dorsal view. C. Segments 1 to 5, ventral view. D. Segments 6 to 8, dorsal view. E. Segments 6 to 9, ventral view. F. Detail showing sublateral glandular cell outlets type 2 on segments 1 and 2. G. Segments 8 to 9, lateral view. H. Segments 10 to 11 in female, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; pdgco1 = paradorsal glandular cell outlet type 1; pdss = paradorsal sensory spot; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817839" httpUri="https://zenodo.org/record/3817839/files/figure.png" pageId="26" pageNumber="27">Fig. 8D</figureCitation>
|
||
) and paraventral (
|
||
<figureCitation id="E6D02A31504CFF84148EFCE2FD350F32" box="[654,747,828,854]" captionStart="Fig" captionStartId="22.[189,232,1611,1637]" captionTargetBox="[256,1332,265,1599]" captionTargetId="figure@22.[256,1332,265,1599]" captionTargetPageId="22" captionText="Fig. 8. Light micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov., holotype, ♀ (A–D, H) (NHMD-223916), paratype, ♂ (E–F) (NHMD-223918) and paratype, ♀ (G) (NHMD-223920). A. Ventral overview. B. Segments 1 to 5, dorsal view. C. Segments 1 to 5, ventral view. D. Segments 6 to 8, dorsal view. E. Segments 6 to 9, ventral view. F. Detail showing sublateral glandular cell outlets type 2 on segments 1 and 2. G. Segments 8 to 9, lateral view. H. Segments 10 to 11 in female, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; pdgco1 = paradorsal glandular cell outlet type 1; pdss = paradorsal sensory spot; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817839" httpUri="https://zenodo.org/record/3817839/files/figure.png" pageId="26" pageNumber="27">Fig. 8E</figureCitation>
|
||
) positions. Small rounded sieve plates are present in lateral accessory positions (
|
||
<figureCitation id="E6D02A31504CFF841407FC81FDB40F1E" box="[519,618,863,890]" captionStart="Fig" captionStartId="21.[189,232,1753,1779]" captionTargetBox="[315,1235,269,1735]" captionTargetId="figure@21.[315,1273,265,1735]" captionTargetPageId="21" captionText="Fig. 7. Line art illustrations of Echinoderes hamiltonorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♂, segments 10 to 11, dorsal view. D. ♂, segments 10 to 11, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817837" httpUri="https://zenodo.org/record/3817837/files/figure.png" pageId="26" pageNumber="27">Figs 7B</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A31504CFF841478FC81FD1C0F1D" box="[632,706,863,889]" captionStart="Fig" captionStartId="22.[189,232,1611,1637]" captionTargetBox="[256,1332,265,1599]" captionTargetId="figure@22.[256,1332,265,1599]" captionTargetPageId="22" captionText="Fig. 8. Light micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov., holotype, ♀ (A–D, H) (NHMD-223916), paratype, ♂ (E–F) (NHMD-223918) and paratype, ♀ (G) (NHMD-223920). A. Ventral overview. B. Segments 1 to 5, dorsal view. C. Segments 1 to 5, ventral view. D. Segments 6 to 8, dorsal view. E. Segments 6 to 9, ventral view. F. Detail showing sublateral glandular cell outlets type 2 on segments 1 and 2. G. Segments 8 to 9, lateral view. H. Segments 10 to 11 in female, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; pdgco1 = paradorsal glandular cell outlet type 1; pdss = paradorsal sensory spot; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817839" httpUri="https://zenodo.org/record/3817839/files/figure.png" pageId="26" pageNumber="27">8E, G</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A31504CFF8414D0FC81FD290F1D" box="[720,759,863,889]" captionStart="Fig" captionStartId="23.[189,232,1508,1534]" captionTargetId="figure@24.[189,1398,260,2026]" captionTargetPageId="24" captionText="Fig. 9 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov. A. Lateral overview. B. Ventral overview. C. Mouth cone. D. Introvert sector 8. E. Segments 1 to 3, dorsolateral view; note that laterodorsal sensory spots are missing in this particular specimen. F. Segments 1 to 3, ventral view. G. Detail showing lateral side of segment 8 without any indication of glandular cell outlets type 2. H. Segments 8 to 9, lateral view. I. Detail showing midlateral to lateroventral parts of segments 5 and 6. J. ♂, segments 10 to 11, dorsolateral view. K. ♂, segments 10 to 11, ventral view. L. ♀, segments 10 to 11, laterodorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdss = middorsal sensory spot; mlss = midlateral sensory spot; oos = outer oral style; pdss = paradorsal sensory spot; pe = penile spines; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlgco1 = ventrolateral glandular cell outlet type 1; vlss = ventrolateral sensory spot; vlt = ventrolateral tube; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817841" httpUri="https://zenodo.org/record/3817841/files/figure.png" pageId="26" pageNumber="27">9H</figureCitation>
|
||
). Pachycycli and pectinate fringe of posterior margin as on preceding segment; cuticular hair covering also as on preceding segment, but with subdorsal to middorsal areas covered with short, filiform, hair-like extensions.
|
||
</paragraph>
|
||
<paragraph id="7E5436B4504CFF8416BDFC37FBAE0898" blockId="26.[189,1398,1001,1277]" pageId="26" pageNumber="27">
|
||
Segment 10 with laterodorsal tubes near posterior segment margin (
|
||
<figureCitation id="E6D02A31504CFF8415D3FC37FB880867" box="[979,1110,1001,1027]" captionStart="Fig" captionStartId="21.[189,232,1753,1779]" captionTargetBox="[315,1235,269,1735]" captionTargetId="figure@21.[315,1273,265,1735]" captionTargetPageId="21" captionText="Fig. 7. Line art illustrations of Echinoderes hamiltonorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♂, segments 10 to 11, dorsal view. D. ♂, segments 10 to 11, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817837" httpUri="https://zenodo.org/record/3817837/files/figure.png" pageId="26" pageNumber="27">Figs 7A, C</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A31504CFF841262FC37FB7E0867" box="[1122,1184,1001,1027]" captionStart="Fig" captionStartId="23.[189,232,1508,1534]" captionTargetId="figure@24.[189,1398,260,2026]" captionTargetPageId="24" captionText="Fig. 9 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov. A. Lateral overview. B. Ventral overview. C. Mouth cone. D. Introvert sector 8. E. Segments 1 to 3, dorsolateral view; note that laterodorsal sensory spots are missing in this particular specimen. F. Segments 1 to 3, ventral view. G. Detail showing lateral side of segment 8 without any indication of glandular cell outlets type 2. H. Segments 8 to 9, lateral view. I. Detail showing midlateral to lateroventral parts of segments 5 and 6. J. ♂, segments 10 to 11, dorsolateral view. K. ♂, segments 10 to 11, ventral view. L. ♀, segments 10 to 11, laterodorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdss = middorsal sensory spot; mlss = midlateral sensory spot; oos = outer oral style; pdss = paradorsal sensory spot; pe = penile spines; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlgco1 = ventrolateral glandular cell outlet type 1; vlss = ventrolateral sensory spot; vlt = ventrolateral tube; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817841" httpUri="https://zenodo.org/record/3817841/files/figure.png" pageId="26" pageNumber="27">9J, L</figureCitation>
|
||
); tubes are minute and do not show sexual dimorphism. Sensory spots present in paradorsal and ventrolateral positions (
|
||
<figureCitation id="E6D02A31504CFF8416C6FBEEFED3082E" box="[198,269,1072,1098]" captionStart="Fig" captionStartId="23.[189,232,1508,1534]" captionTargetId="figure@24.[189,1398,260,2026]" captionTargetPageId="24" captionText="Fig. 9 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov. A. Lateral overview. B. Ventral overview. C. Mouth cone. D. Introvert sector 8. E. Segments 1 to 3, dorsolateral view; note that laterodorsal sensory spots are missing in this particular specimen. F. Segments 1 to 3, ventral view. G. Detail showing lateral side of segment 8 without any indication of glandular cell outlets type 2. H. Segments 8 to 9, lateral view. I. Detail showing midlateral to lateroventral parts of segments 5 and 6. J. ♂, segments 10 to 11, dorsolateral view. K. ♂, segments 10 to 11, ventral view. L. ♀, segments 10 to 11, laterodorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdss = middorsal sensory spot; mlss = midlateral sensory spot; oos = outer oral style; pdss = paradorsal sensory spot; pe = penile spines; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlgco1 = ventrolateral glandular cell outlet type 1; vlss = ventrolateral sensory spot; vlt = ventrolateral tube; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817841" httpUri="https://zenodo.org/record/3817841/files/figure.png" pageId="26" pageNumber="27">Fig. 9</figureCitation>
|
||
J–L). Glandular cell outlets
|
||
<typeStatus id="A1508816504CFF841460FBEFFD4D082F" box="[608,659,1073,1099]" pageId="26" pageNumber="27">type</typeStatus>
|
||
1 present as two middorsal ones, and one pair in paraventral positions. The posterior segment margin of the tergal plate is straight, terminating in a pectinate fringe with rather short fringe tips. The margins of the sternal plates have slightly longer fringe tips, and extend midventrally into a broad comb that almost reaches the margin of the terminal segment (
|
||
<figureCitation id="E6D02A31504CFF84130CFB45FAB408D1" box="[1292,1386,1179,1205]" captionStart="Fig" captionStartId="23.[189,232,1508,1534]" captionTargetId="figure@24.[189,1398,260,2026]" captionTargetPageId="24" captionText="Fig. 9 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov. A. Lateral overview. B. Ventral overview. C. Mouth cone. D. Introvert sector 8. E. Segments 1 to 3, dorsolateral view; note that laterodorsal sensory spots are missing in this particular specimen. F. Segments 1 to 3, ventral view. G. Detail showing lateral side of segment 8 without any indication of glandular cell outlets type 2. H. Segments 8 to 9, lateral view. I. Detail showing midlateral to lateroventral parts of segments 5 and 6. J. ♂, segments 10 to 11, dorsolateral view. K. ♂, segments 10 to 11, ventral view. L. ♀, segments 10 to 11, laterodorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdss = middorsal sensory spot; mlss = midlateral sensory spot; oos = outer oral style; pdss = paradorsal sensory spot; pe = penile spines; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlgco1 = ventrolateral glandular cell outlet type 1; vlss = ventrolateral sensory spot; vlt = ventrolateral tube; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817841" httpUri="https://zenodo.org/record/3817841/files/figure.png" pageId="26" pageNumber="27">Fig. 9K</figureCitation>
|
||
). Pachycycli as on preceding segment; cuticular hair covering also as on preceding segment, but with short, filiform, hair-like extensions present in paradorsal to middorsal areas only.
|
||
</paragraph>
|
||
<paragraph id="7E5436B4504CFF8416BDFAFBFC4F0A38" blockId="26.[189,1399,1316,1629]" pageId="26" pageNumber="27">
|
||
Segment 11 with lateral terminal spines (
|
||
<figureCitation id="E6D02A31504CFF841496FAFAFD00095B" box="[662,734,1316,1343]" captionStart="Fig" captionStartId="21.[189,232,1753,1779]" captionTargetBox="[315,1235,269,1735]" captionTargetId="figure@21.[315,1273,265,1735]" captionTargetPageId="21" captionText="Fig. 7. Line art illustrations of Echinoderes hamiltonorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♂, segments 10 to 11, dorsal view. D. ♂, segments 10 to 11, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817837" httpUri="https://zenodo.org/record/3817837/files/figure.png" pageId="26" pageNumber="27">Figs 7</figureCitation>
|
||
A–B, 8A, H, 9L). Males with three pairs of penile spines; dorsal and ventral penile spines are thin, flexible tubes, whereas the median ones are thicker, conical, and stout (
|
||
<figureCitation id="E6D02A31504CFF84173CFAB2FE5609E2" box="[316,392,1388,1414]" captionStart="Fig" captionStartId="21.[189,232,1753,1779]" captionTargetBox="[315,1235,269,1735]" captionTargetId="figure@21.[315,1273,265,1735]" captionTargetPageId="21" captionText="Fig. 7. Line art illustrations of Echinoderes hamiltonorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♂, segments 10 to 11, dorsal view. D. ♂, segments 10 to 11, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817837" httpUri="https://zenodo.org/record/3817837/files/figure.png" pageId="26" pageNumber="27">Figs 7</figureCitation>
|
||
C–D, 9J–L); females with thin and relatively short lateral terminal accessory spines (
|
||
<figureCitation id="E6D02A31504CFF8416C5FA4EFECF09CE" box="[197,273,1424,1450]" captionStart="Fig" captionStartId="21.[189,232,1753,1779]" captionTargetBox="[315,1235,269,1735]" captionTargetId="figure@21.[315,1273,265,1735]" captionTargetPageId="21" captionText="Fig. 7. Line art illustrations of Echinoderes hamiltonorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♂, segments 10 to 11, dorsal view. D. ♂, segments 10 to 11, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817837" httpUri="https://zenodo.org/record/3817837/files/figure.png" pageId="26" pageNumber="27">Figs 7</figureCitation>
|
||
A–B, 8H, 9L). Sensory spots without marginal hairs present in paradorsal positions (
|
||
<figureCitation id="E6D02A31504CFF841312FA4EFAB609CE" box="[1298,1384,1424,1450]" captionStart="Fig" captionStartId="23.[189,232,1508,1534]" captionTargetId="figure@24.[189,1398,260,2026]" captionTargetPageId="24" captionText="Fig. 9 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov. A. Lateral overview. B. Ventral overview. C. Mouth cone. D. Introvert sector 8. E. Segments 1 to 3, dorsolateral view; note that laterodorsal sensory spots are missing in this particular specimen. F. Segments 1 to 3, ventral view. G. Detail showing lateral side of segment 8 without any indication of glandular cell outlets type 2. H. Segments 8 to 9, lateral view. I. Detail showing midlateral to lateroventral parts of segments 5 and 6. J. ♂, segments 10 to 11, dorsolateral view. K. ♂, segments 10 to 11, ventral view. L. ♀, segments 10 to 11, laterodorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdss = middorsal sensory spot; mlss = midlateral sensory spot; oos = outer oral style; pdss = paradorsal sensory spot; pe = penile spines; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlgco1 = ventrolateral glandular cell outlet type 1; vlss = ventrolateral sensory spot; vlt = ventrolateral tube; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817841" httpUri="https://zenodo.org/record/3817841/files/figure.png" pageId="26" pageNumber="27">Fig. 9J</figureCitation>
|
||
). Glandular cell outlets
|
||
<typeStatus id="A1508816504CFF8417CAFA6AFE2309AA" box="[458,509,1460,1486]" pageId="26" pageNumber="27">type</typeStatus>
|
||
1 not observed. The segment is completely devoid of cuticular hairs, but densely covered with short, filiform hair-like extensions, in middorsal to laterodorsal areas, and in ventromedial to midventral areas. Tergal extensions are short, with oblique margins that end in a short lateral point; lateral tips with long, extended fringe tips that looks like setae (
|
||
<figureCitation id="E6D02A31504CFF841242F9C0FB500A5D" box="[1090,1166,1566,1593]" captionStart="Fig" captionStartId="21.[189,232,1753,1779]" captionTargetBox="[315,1235,269,1735]" captionTargetId="figure@21.[315,1273,265,1735]" captionTargetPageId="21" captionText="Fig. 7. Line art illustrations of Echinoderes hamiltonorum sp. nov. A. ♀, dorsal view. B. ♀, ventral view. C. ♂, segments 10 to 11, dorsal view. D. ♂, segments 10 to 11, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mds = middorsal spine; mlss = midlateral sensory spot; pdgco1 = paradorsal glandular cell outlet type 1; pe = penile spines; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817837" httpUri="https://zenodo.org/record/3817837/files/figure.png" pageId="26" pageNumber="27">Figs 7</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A31504CFF84129AF9C1FB1E0A5D" box="[1178,1216,1567,1593]" captionStart="Fig" captionStartId="22.[189,232,1611,1637]" captionTargetBox="[256,1332,265,1599]" captionTargetId="figure@22.[256,1332,265,1599]" captionTargetPageId="22" captionText="Fig. 8. Light micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov., holotype, ♀ (A–D, H) (NHMD-223916), paratype, ♂ (E–F) (NHMD-223918) and paratype, ♀ (G) (NHMD-223920). A. Ventral overview. B. Segments 1 to 5, dorsal view. C. Segments 1 to 5, ventral view. D. Segments 6 to 8, dorsal view. E. Segments 6 to 9, ventral view. F. Detail showing sublateral glandular cell outlets type 2 on segments 1 and 2. G. Segments 8 to 9, lateral view. H. Segments 10 to 11 in female, ventral view. Abbreviations: eft = extended fringe tips; ldgco2 = laterodorsal glandular cell outlet type 2; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdgco1 = middorsal glandular cell outlet type 1; mds = middorsal spine; pdgco1 = paradorsal glandular cell outlet type 1; pdss = paradorsal sensory spot; pvgco1 = paraventral glandular cell outlet type 1; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; te = tergal extensions; vlt = ventrolateral tube." figureDoi="http://doi.org/10.5281/zenodo.3817839" httpUri="https://zenodo.org/record/3817839/files/figure.png" pageId="26" pageNumber="27">8H</figureCitation>
|
||
,
|
||
<figureCitation id="E6D02A31504CFF8412CDF9C1FB050A5D" box="[1229,1243,1567,1593]" captionStart="Fig" captionStartId="23.[189,232,1508,1534]" captionTargetId="figure@24.[189,1398,260,2026]" captionTargetPageId="24" captionText="Fig. 9 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes hamiltonorum sp. nov. A. Lateral overview. B. Ventral overview. C. Mouth cone. D. Introvert sector 8. E. Segments 1 to 3, dorsolateral view; note that laterodorsal sensory spots are missing in this particular specimen. F. Segments 1 to 3, ventral view. G. Detail showing lateral side of segment 8 without any indication of glandular cell outlets type 2. H. Segments 8 to 9, lateral view. I. Detail showing midlateral to lateroventral parts of segments 5 and 6. J. ♂, segments 10 to 11, dorsolateral view. K. ♂, segments 10 to 11, ventral view. L. ♀, segments 10 to 11, laterodorsal view. Abbreviations: ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; ldt = laterodorsal tube; ltas = lateral terminal accessory spine; lts = lateral terminal spine; lvs = lateroventral spine; lvt = lateroventral tube; mdss = middorsal sensory spot; mlss = midlateral sensory spot; oos = outer oral style; pdss = paradorsal sensory spot; pe = penile spines; psp = primary spinoscalid; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; sp = spinoscalid followed by introvert ring number; te = tergal extensions; tr = trichoscalid; vlgco1 = ventrolateral glandular cell outlet type 1; vlss = ventrolateral sensory spot; vlt = ventrolateral tube; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817841" httpUri="https://zenodo.org/record/3817841/files/figure.png" pageId="26" pageNumber="27">9</figureCitation>
|
||
J–K). Sternal plates shorter than tergal ones, and without tips or extensions.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="36F1653F504CFF8516BDF95AFE8309D0" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="discussion">
|
||
<paragraph id="7E5436B4504CFF8416BDF95AFEF10AFA" blockId="26.[189,303,1668,1694]" box="[189,303,1668,1694]" pageId="26" pageNumber="27">
|
||
<heading id="251C81D8504CFF8416BDF95AFEF10AFA" bold="true" box="[189,303,1668,1694]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
|
||
<emphasis id="4C9FEAA6504CFF8416BDF95AFEF10AFA" bold="true" box="[189,303,1668,1694]" pageId="26" pageNumber="27">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="7E5436B4504CFF8516BDF96DFDE80DF5" blockId="26.[189,1399,1715,2027]" lastBlockId="27.[189,1399,267,401]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
|
||
<taxonomicName id="B9EB4D37504CFF8416BDF96DFDDE0AA9" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[189,512,1715,1741]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="26" pageNumber="27" phylum="Cephalorhyncha" rank="species" species="hamiltonorum" status="sp. nov.">
|
||
<emphasis id="4C9FEAA6504CFF8416BDF96DFDDE0AA9" box="[189,512,1715,1741]" italics="true" pageId="26" pageNumber="27">Echinoderes hamiltonorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD504CFF84140BF96AFDB40AAA" box="[523,618,1716,1742]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is the first species known to show intraspecific variation in the presence or absence of glandular cell outlets
|
||
<typeStatus id="A1508816504CFF8414D7F909FCD40A95" box="[727,778,1751,1777]" pageId="26" pageNumber="27">type</typeStatus>
|
||
2 on segment 8. During the initial work, specimens with and without outlets on segment 8 were actually kept separate and considered as two potentially different species. However, since all meristic data overlapped between the two morphotypes, and since all other cuticular characters, including sensory spots, appeared to occur consistently in all specimens (but note that one specimen lacked laterodorsal sensory spots on segment 1), it was finally concluded that all specimens were conspecific. This was also done in the light of recent observations on other
|
||
<taxonomicName id="B9EB4D37504CFF8416BDF873FE910BA3" authorityName="Claparede" authorityYear="1863" box="[189,335,1965,1991]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="26" pageNumber="27" phylum="Cephalorhyncha" rank="genus">
|
||
<emphasis id="4C9FEAA6504CFF8416BDF873FE910BA3" box="[189,335,1965,1991]" italics="true" pageId="26" pageNumber="27">Echinoderes</emphasis>
|
||
</taxonomicName>
|
||
where the occurrence of tubes displayed intraspecific variation (see Grzelak & Sørensen in press; Sørensen 2018). However, until now this variation has seemed to be restricted to tubes only, and this is the first report of intraspecific variation in the occurrence of glandular cell outlets
|
||
<typeStatus id="A1508816504DFF8512DAFED2FAD30D42" box="[1242,1293,268,294]" pageId="27" pageNumber="28">type</typeStatus>
|
||
2. It can obviously not be ruled out that the dimorphism is reflected in the molecular sequences as well, and that the two morphotypes represent two semi-cryptic, sympatric species, but for now it seems most correct to consider them as one species.
|
||
</paragraph>
|
||
<paragraph id="7E5436B4504DFF8516BDFE67FD040ECE" blockId="27.[189,1399,441,682]" pageId="27" pageNumber="28">
|
||
The new species has the second most common spine pattern, and shares the presence of middorsal spines on segments 4, 6 and 8 combined with lateroventral spines on segments 6 to 9 with twenty additional species of
|
||
<taxonomicName id="B9EB4D37504DFF8517B2FDDFFD9A0E7F" authorityName="Claparede" authorityYear="1863" box="[434,580,513,539]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="27" pageNumber="28" phylum="Cephalorhyncha" rank="genus">
|
||
<emphasis id="4C9FEAA6504DFF8517B2FDDFFD9A0E7F" box="[434,580,513,539]" italics="true" pageId="27" pageNumber="28">Echinoderes</emphasis>
|
||
</taxonomicName>
|
||
. However,
|
||
<taxonomicName id="B9EB4D37504DFF8514C9FDDCFC510E7F" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[713,911,513,539]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="27" pageNumber="28" phylum="Cephalorhyncha" rank="species" species="hamiltonorum" status="sp. nov.">
|
||
<emphasis id="4C9FEAA6504DFF8514C9FDDCFC510E7F" box="[713,911,513,539]" italics="true" pageId="27" pageNumber="28">E. hamiltonorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD504DFF851595FDDFFC310E7F" box="[917,1007,513,539]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is the only species with this spine pattern that also shows glandular cell outlets
|
||
<typeStatus id="A1508816504DFF8514C9FDFBFD220E5B" box="[713,764,549,575]" pageId="27" pageNumber="28">type</typeStatus>
|
||
<quantity id="B9139B51504DFF851502FDFBFCF00E5B" box="[770,814,549,575]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="27" pageNumber="28" unit="in" value="2.0">2 in</quantity>
|
||
subdorsal, laterodorsal and sublateral positions of segment 2, and tubes in ventrolateral positions. The ventrolateral tubes are truly very hard to visualize in LM, but even if they are confused with glandular cell outlets
|
||
<typeStatus id="A1508816504DFF851581FDB3FC6A0EE3" box="[897,948,621,647]" pageId="27" pageNumber="28">type</typeStatus>
|
||
2, there would still be no other species with this particular combination of characters.
|
||
</paragraph>
|
||
<paragraph id="7E5436B4504DFF8516BDFD0CFD810813" blockId="27.[189,1399,722,1143]" pageId="27" pageNumber="28">
|
||
<taxonomicName id="B9EB4D37504DFF8516BDFD0CFDD60E88" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[189,520,722,748]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="27" pageNumber="28" phylum="Cephalorhyncha" rank="species" species="hamiltonorum" status="sp. nov.">
|
||
<emphasis id="4C9FEAA6504DFF8516BDFD0CFDD60E88" box="[189,520,722,748]" italics="true" pageId="27" pageNumber="28">Echinoderes hamiltonorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD504DFF85141CFD0DFD5D0E89" box="[540,643,723,749]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
|
||
probably shows most resemblance to
|
||
<taxonomicName id="B9EB4D37504DFF85128BFD0DFAEA0E88" box="[1163,1332,722,748]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="27" pageNumber="28" phylum="Cephalorhyncha" rank="species" species="hakaiensis">
|
||
<emphasis id="4C9FEAA6504DFF85128BFD0DFAEA0E88" box="[1163,1332,722,748]" italics="true" pageId="27" pageNumber="28">E. hakaiensis</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="1A7A4B45504DFF8516BDFD29FE750F74" author="Herranz M. & Yangel E. & Leander B." box="[189,427,758,785]" pageId="27" pageNumber="28" pagination="303 - 325" refId="ref36456" refString="Herranz M., Yangel E. & Leander B. 2018. Echinoderes hakaiensis sp. nov.: a new mud dragon (Kinorhyncha, Echinoderidae) from the northeastern Pacific Ocean with the redescription of Echinoderes pennaki Higgins, 1960. Marine Biodiversity 48: 303 - 325. https: // doi. org / 10.1007 / s 12526 - 017 - 0726 - z" type="journal article" year="2018">
|
||
Herranz
|
||
<emphasis id="4C9FEAA6504DFF851725FD29FEBC0F74" box="[293,354,758,784]" italics="true" pageId="27" pageNumber="28">et al.</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
).
|
||
<taxonomicName id="B9EB4D37504DFF8517C0FD28FD090F74" box="[448,727,758,784]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="27" pageNumber="28" phylum="Cephalorhyncha" rank="species" species="hakaiensis">
|
||
<emphasis id="4C9FEAA6504DFF8517C0FD28FD090F74" box="[448,727,758,784]" italics="true" pageId="27" pageNumber="28">Echinoderes hakaiensis</emphasis>
|
||
</taxonomicName>
|
||
is the only other species with the same spine pattern as
|
||
<taxonomicName id="B9EB4D37504DFF8516BDFCC5FE5B0F50" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[189,389,794,820]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="27" pageNumber="28" phylum="Cephalorhyncha" rank="species" species="hamiltonorum" status="sp. nov.">
|
||
<emphasis id="4C9FEAA6504DFF8516BDFCC5FE5B0F50" box="[189,389,794,820]" italics="true" pageId="27" pageNumber="28">E. hamiltonorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD504DFF85178DFCC5FE370F51" box="[397,489,795,821]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
|
||
that also shows cuticular structures in subdorsal, laterodorsal, sublateral and ventrolateral positions on segment 2.
|
||
<taxonomicName id="B9EB4D37504DFF851471FCE0FC580F3C" box="[625,902,830,856]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="27" pageNumber="28" phylum="Cephalorhyncha" rank="species" species="hakaiensis">
|
||
<emphasis id="4C9FEAA6504DFF851471FCE0FC580F3C" box="[625,902,830,856]" italics="true" pageId="27" pageNumber="28">Echinoderes hakaiensis</emphasis>
|
||
</taxonomicName>
|
||
has tubes though, instead of glandular cell outlets
|
||
<typeStatus id="A1508816504DFF851715FCBCFE960F18" box="[277,328,866,892]" pageId="27" pageNumber="28">type</typeStatus>
|
||
2, but these structures may sometimes be difficult to distinguish under LM.
|
||
<taxonomicName id="B9EB4D37504DFF8512E5FCBCFEE70FC4" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="27" pageNumber="28" phylum="Cephalorhyncha" rank="species" species="hakaiensis">
|
||
<emphasis id="4C9FEAA6504DFF8512E5FCBCFEE70FC4" italics="true" pageId="27" pageNumber="28">Echinoderes hakaiensis</emphasis>
|
||
</taxonomicName>
|
||
is also larger (the smallest
|
||
<taxonomicName id="B9EB4D37504DFF85147BFC59FCC60FC4" box="[635,792,902,928]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="27" pageNumber="28" phylum="Cephalorhyncha" rank="species" species="hakaiensis">
|
||
<emphasis id="4C9FEAA6504DFF85147BFC59FCC60FC4" box="[635,792,902,928]" italics="true" pageId="27" pageNumber="28">E. hakaiensis</emphasis>
|
||
</taxonomicName>
|
||
has the same length as the largest
|
||
<taxonomicName id="B9EB4D37504DFF8512AFFC59FAA80FC4" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[1199,1398,902,928]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="27" pageNumber="28" phylum="Cephalorhyncha" rank="species" species="hamiltonorum" status="sp. nov.">
|
||
<emphasis id="4C9FEAA6504DFF8512AFFC59FAA80FC4" box="[1199,1398,902,928]" italics="true" pageId="27" pageNumber="28">E. hamiltonorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD504DFF8516BDFC74FEC10FA0" box="[189,287,938,964]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
|
||
), and its tergal extensions appear conspicuously thicker, even though they are rather short.
|
||
<taxonomicName id="B9EB4D37504DFF8516BDFC13FE0B0F83" box="[189,469,973,999]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="27" pageNumber="28" phylum="Cephalorhyncha" rank="species" species="hakaiensis">
|
||
<emphasis id="4C9FEAA6504DFF8516BDFC13FE0B0F83" box="[189,469,973,999]" italics="true" pageId="27" pageNumber="28">Echinoderes hakaiensis</emphasis>
|
||
</taxonomicName>
|
||
furthermore has lateral accessory tubes on segment 8, and even though they could be confused with the
|
||
<typeStatus id="A1508816504DFF85140EFC2CFD9F0868" box="[526,577,1010,1036]" pageId="27" pageNumber="28">type</typeStatus>
|
||
2 sublateral glandular cell outlets of
|
||
<taxonomicName id="B9EB4D37504DFF851207FC2CFB0E086F" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[1031,1232,1009,1035]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="27" pageNumber="28" phylum="Cephalorhyncha" rank="species" species="hamiltonorum" status="sp. nov.">
|
||
<emphasis id="4C9FEAA6504DFF851207FC2CFB0E086F" box="[1031,1232,1009,1035]" italics="true" pageId="27" pageNumber="28">E. hamiltonorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD504DFF8512DAFC2CFAE40868" box="[1242,1338,1010,1036]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, it is usually easier to see the actual tubes on segment 8, compared to tubes on other segments. Finally,
|
||
<taxonomicName id="B9EB4D37504DFF8516BDFBE4FE5B0837" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[189,389,1081,1107]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="27" pageNumber="28" phylum="Cephalorhyncha" rank="species" species="hamiltonorum" status="sp. nov.">
|
||
<emphasis id="4C9FEAA6504DFF8516BDFBE4FE5B0837" box="[189,389,1081,1107]" italics="true" pageId="27" pageNumber="28">E. hamiltonorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD504DFF85178EFBE4FE350830" box="[398,491,1082,1108]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has its rather rare sublateral glandular cell outlets
|
||
<typeStatus id="A1508816504DFF851249FBE4FBA20830" box="[1097,1148,1082,1108]" pageId="27" pageNumber="28">type</typeStatus>
|
||
2 on segment 1, and
|
||
<taxonomicName id="B9EB4D37504DFF8516BDFB80FE840813" box="[189,346,1117,1143]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="27" pageNumber="28" phylum="Cephalorhyncha" rank="species" species="hakaiensis">
|
||
<emphasis id="4C9FEAA6504DFF8516BDFB80FE840813" box="[189,346,1117,1143]" italics="true" pageId="27" pageNumber="28">E. hakaiensis</emphasis>
|
||
</taxonomicName>
|
||
has no such structure.
|
||
</paragraph>
|
||
<paragraph id="7E5436B4504DFF8516BDFB7EFE8309D0" blockId="27.[189,1399,1183,1460]" pageId="27" pageNumber="28">
|
||
In fact, the presence of glandular cell outlets
|
||
<typeStatus id="A1508816504DFF8514CBFB7EFD2008DE" box="[715,766,1184,1210]" pageId="27" pageNumber="28">type</typeStatus>
|
||
2 on segment
|
||
<quantity id="B9139B51504DFF8515A7FB41FC0A08DD" box="[935,980,1183,1209]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="27" pageNumber="28" unit="in" value="1.0">1 in</quantity>
|
||
any
|
||
<taxonomicName id="B9EB4D37504DFF85120CFB41FB4008DD" authorityName="Claparede" authorityYear="1863" box="[1036,1182,1183,1209]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="27" pageNumber="28" phylum="Cephalorhyncha" rank="genus">
|
||
<emphasis id="4C9FEAA6504DFF85120CFB41FB4008DD" box="[1036,1182,1183,1209]" italics="true" pageId="27" pageNumber="28">Echinoderes</emphasis>
|
||
</taxonomicName>
|
||
was only reported very recently. The first report was on
|
||
<taxonomicName id="B9EB4D37504DFF851489FB1AFCE908B9" box="[649,823,1220,1245]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="27" pageNumber="28" phylum="Cephalorhyncha" rank="species" species="unispinosus">
|
||
<emphasis id="4C9FEAA6504DFF851489FB1AFCE908B9" box="[649,823,1220,1245]" italics="true" pageId="27" pageNumber="28">E. unispinosus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="1A7A4B45504DFF851549FB1DFB8508B9" author="Yamasaki H. & Neuhaus B. & George K. H." box="[841,1115,1219,1245]" pageId="27" pageNumber="28" pagination="541 - 566" refId="ref38773" refString="Yamasaki H., Neuhaus B. & George K. H. 2018 b. New species of Echinoderes (Kinorhyncha: Cyclorhagida) from Mediterranean Seamounts and from the deep-sea floor in the Northeast Atlantic Ocean, including notes on two undescribed species. Zootaxa 4378: 541 - 566. https: // doi. org / 10.11646 / zootaxa. 4387.3.8" type="journal article" year="2018">
|
||
Yamasaki
|
||
<emphasis id="4C9FEAA6504DFF8515C7FB1AFBDB08B9" box="[967,1029,1219,1245]" italics="true" pageId="27" pageNumber="28">et al.</emphasis>
|
||
2018b
|
||
</bibRefCitation>
|
||
, but see also
|
||
<figureCitation id="E6D02A31504DFF851303FB1DFAB008BA" box="[1283,1390,1219,1246]" captionStart="Fig" captionStartId="64.[189,232,1648,1674]" captionTargetId="figure@63.[189,1399,347,2018]" captionTargetPageId="63" captionText="Fig. 25 (opposite page). Scanning electron micrographs showing overviews and details of Echinoderes cf. unispinosus. A. Lateroventral overview. B. Dorsolateral overview. C. Segments 1 to 2, laterodorsal view. D. Segments 1 to 2, ventral view. E. Segments 1 to 3, laterodorsal view. F. Segments 2 to 3, subdorsal view G. Segments 1 to 4, dorsal view. H. Segments 3 to 6, ventral view. I. Segments 5 to 7, lateral view. J. Segments 8 to 9, lateral view. K. Segments 8 to 9, ventral view. L. ♂, segments 10 to 11, ventral view. Abbreviations: lagco2 = lateral accessory glandular cell outlet type 2; ldgco2 = laterodorsal glandular cell outlet type 2; ldss = laterodorsal sensory spot; lts = lateral terminal spine; lvs = lateroventral spine; mds = middorsal spine; mdss = middorsal sensory spot; mlgco2 = laterodorsal glandular cell outlet type 2; pe = penile spine; sdgco2 = subdorsal glandular cell outlet type 2; sdss = subdorsal sensory spot; si = sieve plate; slgco2 = sublateral glandular cell outlet type 2; slss = sublateral sensory spot; te = tergal extensions; vlss = ventrolateral sensory spot; vmss = ventromedial sensory spot." figureDoi="http://doi.org/10.5281/zenodo.3817867" httpUri="https://zenodo.org/record/3817867/files/figure.png" pageId="27" pageNumber="28">Fig. 25C</figureCitation>
|
||
) and an undescribed species that was addressed in the same contribution (
|
||
<bibRefCitation id="1A7A4B45504DFF85121AFB39FAF70966" author="Yamasaki H. & Neuhaus B. & George K. H." box="[1050,1321,1255,1282]" pageId="27" pageNumber="28" pagination="541 - 566" refId="ref38773" refString="Yamasaki H., Neuhaus B. & George K. H. 2018 b. New species of Echinoderes (Kinorhyncha: Cyclorhagida) from Mediterranean Seamounts and from the deep-sea floor in the Northeast Atlantic Ocean, including notes on two undescribed species. Zootaxa 4378: 541 - 566. https: // doi. org / 10.11646 / zootaxa. 4387.3.8" type="journal article" year="2018">
|
||
Yamasaki
|
||
<emphasis id="4C9FEAA6504DFF851297FB36FB0D0965" box="[1175,1235,1255,1281]" italics="true" pageId="27" pageNumber="28">et al.</emphasis>
|
||
2018b
|
||
</bibRefCitation>
|
||
). Both species have glandular cell outlets
|
||
<typeStatus id="A1508816504DFF851450FAD5FD5D0941" box="[592,643,1291,1317]" pageId="27" pageNumber="28">type</typeStatus>
|
||
<quantity id="B9139B51504DFF851489FAD5FD6B0941" box="[649,693,1291,1317]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="27" pageNumber="28" unit="in" value="2.0">2 in</quantity>
|
||
midlateral positions on segment 1. Outlets in these positions were also found during the present study in
|
||
<taxonomicName id="B9EB4D37504DFF8514CEFAF1FC9E092C" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[718,832,1327,1352]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="27" pageNumber="28" phylum="Cephalorhyncha" rank="species" species="anniae" status="sp. nov.">
|
||
<emphasis id="4C9FEAA6504DFF8514CEFAF1FC9E092C" box="[718,832,1327,1352]" italics="true" pageId="27" pageNumber="28">E. anniae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD504DFF851549FAF1FC76092D" box="[841,936,1327,1353]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, and the only other known example of glandular cell outlets
|
||
<typeStatus id="A1508816504DFF8517BFFA8DFE2C0909" box="[447,498,1363,1389]" pageId="27" pageNumber="28">type</typeStatus>
|
||
2 on segment 1 is from
|
||
<taxonomicName id="B9EB4D37504DFF851518FA8DFC3E0908" authority="Sørensen & Rohal & Thistle, 2018" authorityName="Sørensen & Rohal & Thistle" authorityYear="2018" box="[792,992,1362,1388]" class="Kinorhyncha" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Cyclorhagida" pageId="27" pageNumber="28" phylum="Cephalorhyncha" rank="species" species="hamiltonorum" status="sp. nov.">
|
||
<emphasis id="4C9FEAA6504DFF851518FA8DFC3E0908" box="[792,992,1362,1388]" italics="true" pageId="27" pageNumber="28">E. hamiltonorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57AC57DD504DFF8515E9FA8DFB9B0909" box="[1001,1093,1363,1389]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
|
||
However, unique for this species, the outlets are displaced to sublateral positions, rather than midlateral positions as found in the other species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |