270 lines
28 KiB
XML
270 lines
28 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.974.54794" ID-GBIF-Dataset="5f0b0760-7a15-49c0-9e80-fc6747f8046e" ID-PMC="PMC7557337" ID-Pensoft-Pub="1313-2970-974-31" ID-Pensoft-UUID="20CAFA6BA01359369DA3F27F90E22660" ID-PubMed="33110378" ID-ZooBank="9B8888660F074DECAE7B88DFB0A4621C" ModsDocID="1313-2970-974-31" checkinTime="1602130124706" checkinUser="pensoft" docAuthor="Wood, Thomas James, Michez, Denis, Cejas, Diego, Lhomme, Patrick & Rasmont, Pierre" docDate="2020" docId="E9BC8D33FE6E54EF8B350FE4F44EBDB0" docLanguage="en" docName="ZooKeys 974: 31-92" docOrigin="ZooKeys 974" docSource="http://dx.doi.org/10.3897/zookeys.974.54794" docTitle="Andrena (Aciandrena) semiadesus Wood 2020, sp. nov." docType="treatment" docUuid="235512E8-91EA-4A84-91B8-41F7727AE452" docUuidSource="ZooBank" docVersion="4" id="20CAFA6BA01359369DA3F27F90E22660" lastPageNumber="31" masterDocId="20CAFA6BA01359369DA3F27F90E22660" masterDocTitle="An update and revision of the Andrena fauna of Morocco (Hymenoptera, Apoidea, Andrenidae) with the description of eleven new North African species" masterLastPageNumber="92" masterPageNumber="31" pageNumber="31" updateTime="1668169591898" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>An update and revision of the Andrena fauna of Morocco (Hymenoptera, Apoidea, Andrenidae) with the description of eleven new North African species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wood, Thomas James</mods:namePart>
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<mods:affiliation>Laboratoire de Zoologie, Universite de Mons, 7000, Mons, Belgium</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-5653-224X</mods:nameIdentifier>
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<mods:nameIdentifier type="email">thomasjames.wood@umons.ac.be</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Michez, Denis</mods:namePart>
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<mods:affiliation>Laboratoire de Zoologie, Universite de Mons, 7000, Mons, Belgium</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-8880-1838</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cejas, Diego</mods:namePart>
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<mods:affiliation>Laboratoire de Zoologie, Universite de Mons, 7000, Mons, Belgium</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lhomme, Patrick</mods:namePart>
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<mods:affiliation>Laboratoire de Zoologie, Universite de Mons, 7000, Mons, Belgium & International Center of Agricultural Research in the Dry Areas, Rabat, Morocco</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-6735-9104</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rasmont, Pierre</mods:namePart>
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<mods:affiliation>Laboratoire de Zoologie, Universite de Mons, 7000, Mons, Belgium</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>974</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>31</mods:start>
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<mods:end>92</mods:end>
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</mods:extent>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.974.54794</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.974.54794</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-974-31</mods:identifier>
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<mods:identifier type="Pensoft-UUID">20CAFA6BA01359369DA3F27F90E22660</mods:identifier>
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<treatment ID-GBIF-Taxon="168243705" LSID="urn:lsid:zoobank.org:act:235512E8-91EA-4A84-91B8-41F7727AE452" httpUri="http://treatment.plazi.org/id/E9BC8D33FE6E54EF8B350FE4F44EBDB0" lastPageNumber="31" pageId="0" pageNumber="31">
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<subSubSection pageId="0" pageNumber="31" type="nomenclature">
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<paragraph pageId="0" pageNumber="31">
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<taxonomicName LSID="E9BC8D33-FE6E-54EF-8B35-0FE4F44EBDB0" authority="Wood" authorityName="Wood" authorityYear="2020" class="Insecta" family="Andrenidae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena (Aciandrena) semiadesus" order="Hymenoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="semiadesus" status="sp. nov." subGenus="Aciandrena">Andrena (Aciandrena) semiadesus Wood</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="31">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figures 1–8" captionStartId="F1" captionText="Figures 1 - 8. Andrena (Aciandrena) semiadesus sp. nov. 1 female profile 2 female face 3 female dorsum 4 female tergites 5 male profile 6 male face 7 male tergites 8 male genitalia." figureDoi="10.3897/zookeys.974.54794.figures1-8" httpUri="https://binary.pensoft.net/fig/460642" pageId="0" pageNumber="31">Figures 1-8</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="31" type="materials_examined">
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<paragraph pageId="0" pageNumber="31">Material.</paragraph>
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<paragraph pageId="0" pageNumber="31">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="31">Holotype</emphasis>
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: Morocco:
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<normalizedToken originalValue="Fès-Meknès">Fes-Meknes</normalizedToken>
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, Laanoucer, 1456 m,
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<geoCoordinate degrees="33.6166" direction="north" orientation="latitude" precision="5" value="33.6166">33.6166N</geoCoordinate>
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, -
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<geoCoordinate degrees="4.7484" direction="west" orientation="longitude" precision="5" value="-4.7484">4.7484W</geoCoordinate>
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, 11-12.iv.2019, 1♂, white pan trap, leg. L. Hamroud & A. Sentil. Deposited in the
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<normalizedToken originalValue="OÖLM">OOELM</normalizedToken>
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.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="31">Paratypes</emphasis>
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: Morocco:
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<normalizedToken originalValue="Fès-Meknès">Fes-Meknes</normalizedToken>
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, Laanoucer, 1456 m, 11-12.iv.2019, 4♂, white and yellow pan traps, leg. L. Hamroud & A. Sentil;
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<normalizedToken originalValue="Drâa-Tafilalet">Draa-Tafilalet</normalizedToken>
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, 20 km W Boudnib, 9.iv.1995, 2♂, 2♀, leg. Ma. Halada. Paratypes are deposited at the
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<normalizedToken originalValue="OÖLM">OOELM</normalizedToken>
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, with a male and female retained in the personal collection of TJW.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="31" type="diagnosis">
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<paragraph pageId="0" pageNumber="31">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="31">
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The finely shagreened propodeal triangle, the narrow facial foveae, the absence of longitudinal striations on the clypeus, and the yellow-marked clypeus in the male place this bee in the
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<taxonomicName authorityName="Warncke" authorityYear="1968" class="Insecta" family="Andrenidae" genus="Aciandrena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Aciandrena" order="Hymenoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="31">Aciandrena</emphasis>
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</taxonomicName>
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. The classification of
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<taxonomicName authorityName="Warncke" authorityYear="1968" class="Insecta" family="Andrenidae" genus="Aciandrena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Aciandrena" order="Hymenoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="31">Aciandrena</emphasis>
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</taxonomicName>
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and its relationship to
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<taxonomicName authorityName="Ashmead" authorityYear="1899" class="Insecta" family="Andrenidae" genus="Micrandrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Micrandrena" order="Hymenoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="31">Micrandrena</emphasis>
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</taxonomicName>
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,
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<taxonomicName authorityName="Warncke" authorityYear="1968" class="Insecta" family="Andrenidae" genus="Graecandrena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Graecandrena" order="Hymenoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="31">Graecandrena</emphasis>
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</taxonomicName>
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, and
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<taxonomicName authorityName="Warncke" authorityYear="1968" class="Insecta" family="Andrenidae" genus="Distandrena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Distandrena" order="Hymenoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="genus">Distandrena</taxonomicName>
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is somewhat in flux, and the subgenus is currently polyphyletic (
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<bibRefCitation DOI="https://doi.org/10.1101/2020.06.09.103556" author="Pisanty, G" journalOrPublisher="Gounouilhou, Bordeaux" pageId="0" pageNumber="31" refId="B28" refString="Pisanty, G, Richter, R, Martin, T, Dettman, J, Cardinal, S, 2020. Molecular phylogeny and historical biogeography of andrenine bees (Hymenoptera: Andrenidae). bioRxiv. https://doi.org/10.1101/2020.06.09.103556" title="Molecular phylogeny and historical biogeography of andrenine bees (Hymenoptera: Andrenidae). bioRxiv." url="https://doi.org/10.1101/2020.06.09.103556" year="2020">Pisanty et al. 2020</bibRefCitation>
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). However, when excluding uncharacteristic taxa that are
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<taxonomicName class="Insecta" family="Andrenidae" genus="Distandrena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="currently" order="Hymenoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="currently">currently</taxonomicName>
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placed in the
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<taxonomicName authorityName="Warncke" authorityYear="1968" class="Insecta" family="Andrenidae" genus="Aciandrena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Aciandrena" order="Hymenoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="31">Aciandrena</emphasis>
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</taxonomicName>
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such as
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<taxonomicName lsidName="A. janthina" pageId="0" pageNumber="31" rank="species" species="janthina">
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<emphasis italics="true" pageId="0" pageNumber="31">A. janthina</emphasis>
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</taxonomicName>
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Warncke, 1975 that clearly belong elsewhere (
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<bibRefCitation DOI="https://doi.org/10.1101/2020.06.09.103556" author="Pisanty, G" journalOrPublisher="Gounouilhou, Bordeaux" pageId="0" pageNumber="31" refId="B28" refString="Pisanty, G, Richter, R, Martin, T, Dettman, J, Cardinal, S, 2020. Molecular phylogeny and historical biogeography of andrenine bees (Hymenoptera: Andrenidae). bioRxiv. https://doi.org/10.1101/2020.06.09.103556" title="Molecular phylogeny and historical biogeography of andrenine bees (Hymenoptera: Andrenidae). bioRxiv." url="https://doi.org/10.1101/2020.06.09.103556" year="2020">Pisanty et al. 2020</bibRefCitation>
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),
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<taxonomicName lsidName="A. semiadesus" pageId="0" pageNumber="31" rank="species" species="semiadesus">
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<emphasis italics="true" pageId="0" pageNumber="31">A. semiadesus</emphasis>
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</taxonomicName>
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meets the classical definition of
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<taxonomicName authorityName="Warncke" authorityYear="1968" class="Insecta" family="Andrenidae" genus="Aciandrena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Aciandrena" order="Hymenoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="31">Aciandrena</emphasis>
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</taxonomicName>
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. The male can be instantly separated from the other
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<taxonomicName authorityName="Warncke" authorityYear="1968" class="Insecta" family="Andrenidae" genus="Aciandrena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Aciandrena" order="Hymenoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="31">Aciandrena</emphasis>
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</taxonomicName>
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with yellow clypei because the yellow marking is diminished, it does not extend all the way to the clypeal margins, and the two dark spots that are usually found towards the centre of the clypeus in yellow-faced
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<taxonomicName authorityName="Warncke" authorityYear="1968" class="Insecta" family="Andrenidae" genus="Aciandrena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Aciandrena" order="Hymenoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="31">Aciandrena</emphasis>
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</taxonomicName>
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are absent as they instead are contiguous with black markings that extend in from the clypeal margins (Fig.
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<figureCitation captionStart="Figures 1–8" captionStartId="F1" captionText="Figures 1 - 8. Andrena (Aciandrena) semiadesus sp. nov. 1 female profile 2 female face 3 female dorsum 4 female tergites 5 male profile 6 male face 7 male tergites 8 male genitalia." figureDoi="10.3897/zookeys.974.54794.figures1-8" httpUri="https://binary.pensoft.net/fig/460642" pageId="0" pageNumber="31">6</figureCitation>
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). The genitalia are also noticeably more elongate, the capsule almost twice as long as wide, and with a broad penis valve, most other
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<taxonomicName authorityName="Warncke" authorityYear="1968" class="Insecta" family="Andrenidae" genus="Aciandrena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Aciandrena" order="Hymenoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="31">Aciandrena</emphasis>
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</taxonomicName>
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species with capsules only a little longer than wide and with a comparatively narrow penis valve (Figs
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<figureCitation captionStart="Figures 99–104" captionStartId="F17" captionText="Figures 99 - 104. Andrena (Aciandrena) fulica Warncke, 1974 99 male face 100 male genitalia. Andrena (Aciandrena) fulica astrella Warncke, 1975 (Morocco) 101 male face 102 male genitalia (Spain) 103 male face 104 male genitalia." figureDoi="10.3897/zookeys.974.54794.figures99-104" httpUri="https://binary.pensoft.net/fig/460658" pageId="0" pageNumber="31">100</figureCitation>
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,
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<figureCitation captionStart="Figures 99–104" captionStartId="F17" captionText="Figures 99 - 104. Andrena (Aciandrena) fulica Warncke, 1974 99 male face 100 male genitalia. Andrena (Aciandrena) fulica astrella Warncke, 1975 (Morocco) 101 male face 102 male genitalia (Spain) 103 male face 104 male genitalia." figureDoi="10.3897/zookeys.974.54794.figures99-104" httpUri="https://binary.pensoft.net/fig/460658" pageId="0" pageNumber="31">102</figureCitation>
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,
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<figureCitation captionStart="Figures 99–104" captionStartId="F17" captionText="Figures 99 - 104. Andrena (Aciandrena) fulica Warncke, 1974 99 male face 100 male genitalia. Andrena (Aciandrena) fulica astrella Warncke, 1975 (Morocco) 101 male face 102 male genitalia (Spain) 103 male face 104 male genitalia." figureDoi="10.3897/zookeys.974.54794.figures99-104" httpUri="https://binary.pensoft.net/fig/460658" pageId="0" pageNumber="31">104</figureCitation>
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,
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<figureCitation captionStart="Figures 125–130" captionStartId="F21" captionText="Figures 125 - 130. Andrena (Aciandrena) pratincola Warncke, 1974 (Morocco) 125 male face 126 male genitalia (Egypt, Paratype) 127 male face 128 male genitalia Andrena (Aciandrena) varicornis Perez, 1895 129 male face 130 male genitalia." figureDoi="10.3897/zookeys.974.54794.figures125-130" httpUri="https://binary.pensoft.net/fig/460662" pageId="0" pageNumber="31">126</figureCitation>
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,
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<figureCitation captionStart="Figures 125–130" captionStartId="F21" captionText="Figures 125 - 130. Andrena (Aciandrena) pratincola Warncke, 1974 (Morocco) 125 male face 126 male genitalia (Egypt, Paratype) 127 male face 128 male genitalia Andrena (Aciandrena) varicornis Perez, 1895 129 male face 130 male genitalia." figureDoi="10.3897/zookeys.974.54794.figures125-130" httpUri="https://binary.pensoft.net/fig/460662" pageId="0" pageNumber="31">128</figureCitation>
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,
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<figureCitation captionStart="Figures 125–130" captionStartId="F21" captionText="Figures 125 - 130. Andrena (Aciandrena) pratincola Warncke, 1974 (Morocco) 125 male face 126 male genitalia (Egypt, Paratype) 127 male face 128 male genitalia Andrena (Aciandrena) varicornis Perez, 1895 129 male face 130 male genitalia." figureDoi="10.3897/zookeys.974.54794.figures125-130" httpUri="https://binary.pensoft.net/fig/460662" pageId="0" pageNumber="31">130</figureCitation>
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;
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<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4189.3.3" author="Pisanty, G" journalOrPublisher="Zootaxa" pageId="0" pageNumber="31" pagination="485 - 515" refId="B29" refString="Pisanty, G, Scheuchl, E, Dorchin, N, 2016. Eight new species of Andrena Fabricius (Hymenoptera: Apoidea: Andrenidae) from Israel - a Mediterranean hotspot for wild bees. Zootaxa 4189: 485 - 515, DOI: https://doi.org/10.11646/zootaxa.4189.3.3" title="Eight new species of Andrena Fabricius (Hymenoptera: Apoidea: Andrenidae) from Israel - a Mediterranean hotspot for wild bees." url="https://doi.org/10.11646/zootaxa.4189.3.3" volume="4189" year="2016">Pisanty et al. 2016</bibRefCitation>
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). The female is most similar to
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<taxonomicName lsidName="A. (Graecandrena) totana" pageId="0" pageNumber="31" rank="species" species="totana" subGenus="Graecandrena">A. (Graecandrena) totana</taxonomicName>
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Warncke, 1974 as they both have a broad, shagreened, dull, moderately raised, and centrally slightly flattened sparsely punctate clypeus, but the tergites of
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<taxonomicName lsidName="A. semiadesus" pageId="0" pageNumber="31" rank="species" species="semiadesus">
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<emphasis italics="true" pageId="0" pageNumber="31">A. semiadesus</emphasis>
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</taxonomicName>
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are completely impunctate and the foveae are narrower.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="31" type="description">
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<paragraph pageId="0" pageNumber="31">Description.</paragraph>
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<paragraph pageId="0" pageNumber="31">
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<emphasis bold="true" pageId="0" pageNumber="31">Female</emphasis>
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: Body length 7 mm (Fig.
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<figureCitation captionStart="Figures 1–8" captionStartId="F1" captionText="Figures 1 - 8. Andrena (Aciandrena) semiadesus sp. nov. 1 female profile 2 female face 3 female dorsum 4 female tergites 5 male profile 6 male face 7 male tergites 8 male genitalia." figureDoi="10.3897/zookeys.974.54794.figures1-8" httpUri="https://binary.pensoft.net/fig/460642" pageId="0" pageNumber="31">1</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="31">Head</emphasis>
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: Black, as wide as long (Fig.
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<figureCitation captionStart="Figures 1–8" captionStartId="F1" captionText="Figures 1 - 8. Andrena (Aciandrena) semiadesus sp. nov. 1 female profile 2 female face 3 female dorsum 4 female tergites 5 male profile 6 male face 7 male tergites 8 male genitalia." figureDoi="10.3897/zookeys.974.54794.figures1-8" httpUri="https://binary.pensoft.net/fig/460642" pageId="0" pageNumber="31">2</figureCitation>
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). Clypeus broad, slightly arched, dull, strongly shagreened with exception of apical margin where shagreenation is weaker, therefore weakly shining. Sparsely and shallowly punctured, punctures separated by 2-3 puncture diameters. Process of labrum semi-circular, weakly shining. Foveae in lower half deep, narrow, less than half the width of an antenna, very close to inner margin of compound eye. In upper half foveae widen to approximately the width of an antenna, at their widest still occupying less than half the distance between top of compound eye and lateral ocellus. Face, gena, and vertex with moderate brownish to whitish hairs, the longest of these roughly equal to
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<normalizedToken originalValue="½">1/2</normalizedToken>
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the length of the scape. Antennae dark, A5-12 lightened orange below, A3 equalling A4+5 combined. Gena slightly wider than width of compound eye, weakly longitudinally striate. Ocelloccipital distance short, less than 1/3 width of lateral ocellus.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="31">Mesosoma</emphasis>
|
||
: Scutum and scutellum completely shagreened, weakly shining, punctures shallow and inconspicuous, punctures separated by 2-3 puncture diameters (Fig.
|
||
<figureCitation captionStart="Figures 1–8" captionStartId="F1" captionText="Figures 1 - 8. Andrena (Aciandrena) semiadesus sp. nov. 1 female profile 2 female face 3 female dorsum 4 female tergites 5 male profile 6 male face 7 male tergites 8 male genitalia." figureDoi="10.3897/zookeys.974.54794.figures1-8" httpUri="https://binary.pensoft.net/fig/460642" pageId="0" pageNumber="31">3</figureCitation>
|
||
). Propodeal triangle differentiated from rest of propodeum by larger and coarser shagreenation, weakly shining. Episternum and propodeum finely shagreened, dull. Episternum with sparse, long white hairs, these approaching
|
||
<normalizedToken originalValue="¾">3/4</normalizedToken>
|
||
of the length of the scape. Legs dark, tarsal segments becoming dark brown. Femoral and tibial scopa simple, white. Wings hyaline, venation light brown, stigma pale yellow, nervulus interstitial to slightly antefurcal.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="31">Metasoma</emphasis>
|
||
: Tergites dark, margins lightened yellow (Fig.
|
||
<figureCitation captionStart="Figures 1–8" captionStartId="F1" captionText="Figures 1 - 8. Andrena (Aciandrena) semiadesus sp. nov. 1 female profile 2 female face 3 female dorsum 4 female tergites 5 male profile 6 male face 7 male tergites 8 male genitalia." figureDoi="10.3897/zookeys.974.54794.figures1-8" httpUri="https://binary.pensoft.net/fig/460642" pageId="0" pageNumber="31">4</figureCitation>
|
||
). Tergal discs strongly microreticulate, punctation extremely sparse and obscured. T2-4 laterally with very weak fringes of white hairs. T5-6 with golden hairs flanking pygidial plate, this rounded with raised margin, shagreened, weakly shining.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="31">
|
||
<emphasis bold="true" pageId="0" pageNumber="31">Male.</emphasis>
|
||
Body length 6.5-7 mm (Fig.
|
||
<figureCitation captionStart="Figures 1–8" captionStartId="F1" captionText="Figures 1 - 8. Andrena (Aciandrena) semiadesus sp. nov. 1 female profile 2 female face 3 female dorsum 4 female tergites 5 male profile 6 male face 7 male tergites 8 male genitalia." figureDoi="10.3897/zookeys.974.54794.figures1-8" httpUri="https://binary.pensoft.net/fig/460642" pageId="0" pageNumber="31">5</figureCitation>
|
||
).
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="31">Head</emphasis>
|
||
: Black, wider than long. Clypeus slightly arched, ground colour black but with large yellow marking covering approximately 90% of surface (Fig.
|
||
<figureCitation captionStart="Figures 1–8" captionStartId="F1" captionText="Figures 1 - 8. Andrena (Aciandrena) semiadesus sp. nov. 1 female profile 2 female face 3 female dorsum 4 female tergites 5 male profile 6 male face 7 male tergites 8 male genitalia." figureDoi="10.3897/zookeys.974.54794.figures1-8" httpUri="https://binary.pensoft.net/fig/460642" pageId="0" pageNumber="31">6</figureCitation>
|
||
). Yellow marking does not extend to base of clypeus. Laterad of clypeal centre, yellow marking reduced, two black triangular markings extending in towards the centre, giving marking a broad, inverted T-shape. In basal half, clypeus dull and shagreened, becoming shiny in apical half. Clypeal punctures shallow and irregular, separated by 2-5 puncture diameters. Process of labrum rectangular, twice as broad and long, shiny. Face, gena, and vertex with long white hairs, longest on underside of head, equalling length of the scape. Scape and pedicel black, following antennal segments dark brown to black, A3 longer than A4, shorter than A4+5 combined. Gena and ocelloccipital distance as in female.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="31">Mesosoma</emphasis>
|
||
: Scutum and scutellum dull, strongly shagreened, sparsely and shallowly punctured, punctures separated by 1-4 puncture diameters. Propodeal triangle weakly marked, scarcely differentiated from propodeum, shagreened, slightly shining laterally. Mesosoma with white hairs, sparse on scutum, denser and longer on episternum, exceeding length of scape. Legs dark, only final tarsal segment slightly lightened to dark brown, pubescence white. Wings hyaline, venation brown, stigma brown, nervulus interstitial.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="31">Metasoma</emphasis>
|
||
: Tergites as in female, but margins more strongly lightened, transparent whitish apically (Fig.
|
||
<figureCitation captionStart="Figures 1–8" captionStartId="F1" captionText="Figures 1 - 8. Andrena (Aciandrena) semiadesus sp. nov. 1 female profile 2 female face 3 female dorsum 4 female tergites 5 male profile 6 male face 7 male tergites 8 male genitalia." figureDoi="10.3897/zookeys.974.54794.figures1-8" httpUri="https://binary.pensoft.net/fig/460642" pageId="0" pageNumber="31">7</figureCitation>
|
||
). Tergal margins laterally with loose white hair bands of irregular length, very widely interrupted. Genitalia simple (Fig.
|
||
<figureCitation captionStart="Figures 1–8" captionStartId="F1" captionText="Figures 1 - 8. Andrena (Aciandrena) semiadesus sp. nov. 1 female profile 2 female face 3 female dorsum 4 female tergites 5 male profile 6 male face 7 male tergites 8 male genitalia." figureDoi="10.3897/zookeys.974.54794.figures1-8" httpUri="https://binary.pensoft.net/fig/460642" pageId="0" pageNumber="31">8</figureCitation>
|
||
), gonocoxites forming rounded dorsal lobes, gonostyli long, tapering to rounded end.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.974.54794.figures1-8" httpUri="https://binary.pensoft.net/fig/460642" pageId="0" pageNumber="31" start="Figures 1–8" startId="F1">
|
||
<paragraph pageId="0" pageNumber="31">
|
||
<emphasis bold="true" pageId="0" pageNumber="31">Figures 1-8.</emphasis>
|
||
<taxonomicName authorityName="Wood" authorityYear="2020" class="Insecta" family="Andrenidae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena (Aciandrena) semiadesus" order="Hymenoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="semiadesus" subGenus="Aciandrena">Andrena (Aciandrena) semiadesus</taxonomicName>
|
||
sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="31">1</emphasis>
|
||
female profile
|
||
<emphasis bold="true" pageId="0" pageNumber="31">2</emphasis>
|
||
female face
|
||
<emphasis bold="true" pageId="0" pageNumber="31">3</emphasis>
|
||
female dorsum
|
||
<emphasis bold="true" pageId="0" pageNumber="31">4</emphasis>
|
||
female tergites
|
||
<emphasis bold="true" pageId="0" pageNumber="31">5</emphasis>
|
||
male profile
|
||
<emphasis bold="true" pageId="0" pageNumber="31">6</emphasis>
|
||
male face
|
||
<emphasis bold="true" pageId="0" pageNumber="31">7</emphasis>
|
||
male tergites
|
||
<emphasis bold="true" pageId="0" pageNumber="31">8</emphasis>
|
||
male genitalia.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="31" type="distribution">
|
||
<paragraph pageId="0" pageNumber="31">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="31">
|
||
Central and eastern Morocco (Fig.
|
||
<figureCitation captionStart="Figure 145" captionStartId="F24" captionText="Figure 145. Distribution of sampling locations of newly described species A Andrena hebescens (circles), Andrena semiadesus (squares) B Andrena tenebricorpus (diamond), Andrena niveofacies (circle), Andrena triangulivalvis (triangle) C Andrena sparsipunctata (circles), Andrena breviceps (diamonds) D Andrena acutidentis (squares), Andrena farinosoides (triangles), Andrena nigriclypeus (circle). Relief is indicated by shading, measured in meters above sea level." figureDoi="10.3897/zookeys.974.54794.figure145" httpUri="https://binary.pensoft.net/fig/460665" pageId="0" pageNumber="31">145a</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="31" type="floral preferences">
|
||
<paragraph pageId="0" pageNumber="31">Floral preferences.</paragraph>
|
||
<paragraph pageId="0" pageNumber="31">None recorded.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="31" type="etymology">
|
||
<paragraph pageId="0" pageNumber="31">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="31">
|
||
The name
|
||
<emphasis italics="true" pageId="0" pageNumber="31">semi</emphasis>
|
||
(partly or partially) +
|
||
<emphasis italics="true" pageId="0" pageNumber="31">adesus</emphasis>
|
||
(eaten, worn, eroded) was chosen to illustrate the clypeus of the male, where the central yellow marking is laterally diminished by intruding black marks.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |