treatments-xml/data/C9/7A/B1/C97AB10B4B33FFF662D27399FDC9683E.xml

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<document id="E22432B3E3EEEC8C6A43C0D2C263B72E" ID-DOI="10.37828/em.2020.35.10" ID-ISSN="2336-9744" ID-Zenodo-Dep="13232490" ID-ZooBank="urn:lsid:zoobank.org:pub:F63DAA57-8D7A-4EC7-9FA0-602943FED1FB" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722913961471" checkinUser="felipe" docAuthor="Georgiev, Dilian &amp; Glöer, Peter" docDate="2020" docId="C97AB10B4B33FFF662D27399FDC9683E" docLanguage="en" docName="EcolMontenegrina.35.129-137.pdf" docOrigin="Ecologica Montenegrina 35" docSource="http://dx.doi.org/10.37828/em.2020.35.10" docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.2:EcolMontenegrina.2017-.journal_article" docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE" docStyleName="EcolMontenegrina.2017-.journal_article" docStyleVersion="2" docTitle="Grossuana beroni Georgiev &amp; Glöer, 2020, n. sp." docType="treatment" docUuid="F8C2DAFC-0535-4E8E-A891-6F9ECFFA88AC" docUuidSource="ZooBank" docVersion="2" lastPageNumber="133" masterDocId="3543C9734B31FFF262457045FFEE693E" masterDocTitle="Three new hydrobiid species from Thassos Island (Greece) with a re-description of Amnicola charpentieri Roth, 1855" masterLastPageNumber="137" masterPageNumber="129" pageNumber="131" updateTime="1722916995419" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="244A8646E3DBCA79B14EF294AAE9B9CF">Three new hydrobiid species from Thassos Island (Greece) with a re-description of Amnicola charpentieri Roth, 1855</mods:title>
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<mods:name id="7CAE60F57D1A443FA18F9706F3572B17" type="personal">
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<mods:roleTerm id="EC6C2802E568C97D5FFFB464D7C84716">Author</mods:roleTerm>
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<mods:namePart id="44F9488400E0F3B2AFB5DB898A328BDD">Georgiev, Dilian</mods:namePart>
<mods:affiliation id="15388A250697CA8636710B2CFC93B514">Department of Ecology and Nature Conservation, Faculty of Biology, Paisii Hilendarski University of Plovdiv, 24 Tsar Assen Str., 4000 Plovdiv, Bulgaria</mods:affiliation>
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<mods:namePart id="804DBCEA468AD4C23C3C9C5A3587FE68">Glöer, Peter</mods:namePart>
<mods:affiliation id="8D22139D1E3ACF40B845FE0A67493175">Biodiversity Research Laboratory, Schulstrasse 3, D- 25491 Hetlingen, Germany Corresponding author. E-mail: diliangeorgiev @ abv. bg</mods:affiliation>
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<mods:title id="EA4224FD79580A15A3E8A254298CCFB7">Ecologica Montenegrina</mods:title>
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<mods:date id="389F910ED91F8C785C3667798178DC51">2020</mods:date>
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<mods:number id="D3284EEA00B4A6CCE0471B5D2C76A6B0">2020-10-22</mods:number>
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<mods:url id="2FA5EB4DC6755F139B4B6C3158DF52F0">http://dx.doi.org/10.37828/em.2020.35.10</mods:url>
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<mods:classification id="962FF77133E4EDDCB17AB592BEA9260B">journal article</mods:classification>
<mods:identifier id="666956C5860A5BD7D13D6C44AD3C06F7" type="DOI">10.37828/em.2020.35.10</mods:identifier>
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<treatment id="C97AB10B4B33FFF662D27399FDC9683E" ID-DOI="http://doi.org/10.5281/zenodo.13232604" ID-Zenodo-Dep="13232604" LSID="urn:lsid:zoobank.org:act:F8C2DAFC-0535-4E8E-A891-6F9ECFFA88AC" httpUri="http://treatment.plazi.org/id/C97AB10B4B33FFF662D27399FDC9683E" lastPageId="4" lastPageNumber="133" pageId="2" pageNumber="131">
<subSubSection id="09C953964B33FFF062D27399FB696D2A" pageId="2" pageNumber="131" type="nomenclature">
<paragraph id="416C001D4B33FFF062D27399FD976AC8" blockId="2.[151,1159,987,1044]" box="[151,633,987,1014]" pageId="2" pageNumber="131">
<heading id="1A24B7714B33FFF062D27399FD976AC8" box="[151,633,987,1014]" fontSize="11" level="2" pageId="2" pageNumber="131" reason="2">
<emphasis id="73A7DC0F4B33FFF062D27399FE556AC8" bold="true" box="[151,443,988,1014]" pageId="2" pageNumber="131">
<taxonomicName id="86D37B9E4B33FFF062D27399FE9F6AC8" ID-CoL="7RG9W" authorityName="Georgiev &amp; Glöer" authorityYear="2020" box="[151,369,988,1014]" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="2" pageNumber="131" phylum="Mollusca" rank="species" species="beroni" status="sp. nov.">
<emphasis id="73A7DC0F4B33FFF062D27399FE9F6AC8" bold="true" box="[151,369,988,1014]" italics="true" pageId="2" pageNumber="131">Grossuana beroni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="689461744B33FFF0633D7399FE556AC8" box="[376,443,988,1014]" pageId="2" pageNumber="131" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="D9E81C984B33FFF0638E739EFDF36AC8" box="[459,541,987,1014]" captionStart="Figure 2" captionStartId="2.[151,226,729,753]" captionTargetBox="[151,1437,223,711]" captionTargetId="figure-21@2.[151,1437,223,711]" captionTargetPageId="2" captionText="Figure 2. The sampling sites of the species under discussion on Thassos. 1: Rachoni village: type locality of Bythinella rachonica n. sp., localiy of Grossuana beroni n. sp.; 2: Drakotrype cave, Panagia village: localiy of Grossuana beroni n. sp.; 3: type localiy of Grossuana beroni; 4: type locality of Grossuana thasia n. sp., 5: Pernes Mountain, type locality of Bythinella reischuetzi n. sp., 6: Hymettos, type locality of Bythinella charpentieri (Roth, 1855)." figureDoi="http://doi.org/10.5281/zenodo.13232496" httpUri="https://zenodo.org/record/13232496/files/figure.png" pageId="2" pageNumber="131">Figs. 2</figureCitation>
,
<figureCitation id="D9E81C984B33FFF0606F739EFDD46AC8" box="[554,570,987,1014]" captionStart="Figures 3-5. 3" captionStartId="3.[151,236,1449,1473]" captionTargetBox="[151,1439,404,1431]" captionTargetId="figure-76@3.[151,1439,404,1431]" captionTargetPageId="3" captionText="Figures 3-5. 3: Type locality of Grossuana beroni n. sp., water source south of Skala Potamia near the round road of the island. 4: Type locality of Grossuana thasia n. sp., a spring south of Kinira near the round road of the island. 5: Type locality of Bythinella rachonica n. sp." figureDoi="http://doi.org/10.5281/zenodo.13232498" httpUri="https://zenodo.org/record/13232498/files/figure.png" pageId="2" pageNumber="131">3</figureCitation>
,
<figureCitation id="D9E81C984B33FFF06002739EFD816AC8" box="[583,623,987,1014]" captionStart="Figures 6-10" captionStartId="3.[151,236,1972,1996]" captionTargetBox="[151,1444,1565,1953]" captionTargetId="figure-136@3.[151,1444,1565,1953]" captionTargetPageId="3" captionText="Figures 6-10. The Grossuana spp. 6-7: G. beroni n. sp. (paratypes), 8: penis of G. beroni n. sp. 9: G. thasia n. sp (holotype), 10: penis of G. thasia n. sp." figureDoi="http://doi.org/10.5281/zenodo.13232500" httpUri="https://zenodo.org/record/13232500/files/figure.png" pageId="2" pageNumber="131">6-8</figureCitation>
)
</heading>
</paragraph>
<paragraph id="416C001D4B33FFF062D273B9FB696D2A" blockId="2.[151,1159,987,1044]" box="[151,1159,1020,1044]" pageId="2" pageNumber="131">
https://zoobank.org/
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urn:lsid:zoobank.org:act:
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</subSubSection>
<subSubSection id="09C953964B33FFF062D2747AFBE26C94" pageId="2" pageNumber="131" type="materials_examined">
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<emphasis id="73A7DC0F4B33FFF062D2747AFE9E6D67" bold="true" box="[151,368,1087,1113]" pageId="2" pageNumber="131">Material studied:</emphasis>
</paragraph>
<paragraph id="416C001D4B33FFF062B07425FB566D44" blockId="2.[151,1436,1087,1349]" box="[245,1208,1119,1146]" pageId="2" pageNumber="131">
<emphasis id="73A7DC0F4B33FFF062B07425FE9F6D44" bold="true" box="[245,369,1120,1146]" pageId="2" pageNumber="131">
<typeStatus id="9E68BEBF4B33FFF062B07425FE826D44" box="[245,364,1120,1146]" pageId="2" pageNumber="131" type="holotype">Holotype</typeStatus>
:
</emphasis>
shell height
<quantity id="862BADF84B33FFF0604D741AFD8A6D44" box="[520,612,1119,1146]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="131" unit="mm" value="1.7">1.7 mm</quantity>
, width
<quantity id="862BADF84B33FFF060FF741AFCF96D44" box="[698,791,1119,1146]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="131" unit="mm" value="1.2">1.2 mm</quantity>
, from type locality (ZMH 140789).
</paragraph>
<paragraph id="416C001D4B33FFF062B074C4FEA86D80" blockId="2.[151,1436,1087,1349]" pageId="2" pageNumber="131">
<materialsCitation id="F1BB0A404B33FFF062B074C4FC9D6DA2" box="[245,883,1153,1180]" collectionCode="ZMH" pageId="2" pageNumber="131" specimenCode="ZMH 140790" specimenCount="43" typeStatus="paratype">
<emphasis id="73A7DC0F4B33FFF062B074C4FE926DA5" bold="true" box="[245,380,1153,1179]" pageId="2" pageNumber="131">
<typeStatus id="9E68BEBF4B33FFF062B074C4FE996DA5" box="[245,375,1153,1179]" pageId="2" pageNumber="131" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="57D5CB944B33FFF063CD74C4FDC46DA2" box="[392,554,1153,1180]" count="43" pageId="2" pageNumber="131" type="generic">43 specimens</specimenCount>
in ethanol (
<specimenCode id="1175A8664B33FFF0608574C4FC826DA2" box="[704,876,1153,1180]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="2" pageNumber="131" type="Museum">ZMH 140790</specimenCode>
)
</materialsCitation>
,
<specimenCount id="57D5CB944B33FFF061C674C4FBF86DA2" box="[899,1046,1153,1180]" count="5" pageId="2" pageNumber="131" type="generic">5 specimens</specimenCount>
in coll. Glöer,
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in coll. Georgiev.
</paragraph>
<paragraph id="416C001D4B33FFF062B07481FBEA6C7B" blockId="2.[151,1436,1087,1349]" pageId="2" pageNumber="131">
<materialsCitation id="F1BB0A404B33FFF062B07481FBEE6C7B" collectionCode="ZMH" collectorName="Gloer" country="Greece" county="Thassos Island" elevation="124" latitude="40.75822" location="Panagia village" longLatPrecision="1" longitude="24.638138" municipality="Drakotrypa cave" pageId="2" pageNumber="131" specimenCode="ZMH 140792, ZMH 140791" specimenCount="50" stateProvince="Voreio Aigaio">
Additional localities: (1)
<collectingCountry id="39C4408D4B33FFF060587481FD9D6DE1" box="[541,627,1220,1247]" name="Greece" pageId="2" pageNumber="131">Greece</collectingCountry>
,
<collectingCounty id="A80D78914B33FFF0603A7481FCDE6DE1" box="[639,816,1220,1247]" pageId="2" pageNumber="131">Thassos Island</collectingCounty>
, spring below
<collectingMunicipality id="A1089A674B33FFF061997481FB4F6DE1" box="[988,1185,1220,1247]" pageId="2" pageNumber="131">Drakotrypa cave</collectingMunicipality>
,
<location id="440C56C64B33FFF066EB7481FA8C6DE1" LSID="urn:lsid:plazi:treatment:C97AB10B4B33FFF662D27399FDC9683E:440C56C64B33FFF066EB7481FA8C6DE1" box="[1198,1378,1220,1247]" country="Greece" county="Thassos Island" latitude="40.75822" longLatPrecision="1" longitude="24.638138" municipality="Drakotrypa cave" name="Panagia village" pageId="2" pageNumber="131" stateProvince="Voreio Aigaio">Panagia village</location>
<geoCoordinate id="24E766DA4B33FFF0672C7481FF1D6C3F" degrees="40" direction="north" minutes="44" orientation="latitude" pageId="2" pageNumber="131" precision="1" seconds="06.3" value="40.735085">N40 44 06.3</geoCoordinate>
<geoCoordinate id="24E766DA4B33FFF062B874A3FE7A6C3F" box="[253,404,1254,1281]" degrees="24" direction="east" minutes="44" orientation="longitude" pageId="2" pageNumber="131" precision="1" seconds="02.2" value="24.733944">E24 44 02.2</geoCoordinate>
,
<elevation id="CAFEE72E4B33FFF063E774A3FDDE6C3F" box="[418,560,1254,1281]" metricMagnitude="2" metricUnit="m" metricValue="1.24" pageId="2" pageNumber="131" unit="m" value="124.0">
<quantity id="862BADF84B33FFF063E774A3FE1E6C3F" box="[418,496,1254,1281]" metricMagnitude="2" metricUnit="m" metricValue="1.24" pageId="2" pageNumber="131" unit="m" value="124.0">124 m</quantity>
a.s.l.
</elevation>
(
<specimenCode id="1175A8664B33FFF0600774A3FD026C3F" box="[578,748,1254,1281]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="2" pageNumber="131" type="Museum">ZMH 140792</specimenCode>
:
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,
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coll.
<collectorName id="EC2665CB4B33FFF066CD74A3FB236C3F" box="[1160,1229,1254,1281]" pageId="2" pageNumber="131">Glöer</collectorName>
) and (2)
<collectingCountry id="39C4408D4B33FFF0670574A3FA786C3F" box="[1344,1430,1254,1281]" name="Greece" pageId="2" pageNumber="131">Greece</collectingCountry>
,
<location id="440C56C64B33FFF062D2754DFEA96C1D" LSID="urn:lsid:plazi:treatment:C97AB10B4B33FFF662D27399FDC9683E:440C56C64B33FFF062D2754DFEA96C1D" box="[151,327,1288,1315]" country="Greece" county="Thassos Island" latitude="40.75822" longLatPrecision="1" longitude="24.638138" municipality="Drakotrypa cave" name="Thassos Island" pageId="2" pageNumber="131" stateProvince="Voreio Aigaio">Thassos Island</location>
,
<collectingRegion id="8317CEFF4B33FFF06311754DFE126C1D" box="[340,508,1288,1315]" country="Greece" name="Voreio Aigaio" pageId="2" pageNumber="131">North Aegean</collectingRegion>
, a big spring in the yard of the church of
<location id="440C56C64B33FFF061A1754DFB4E6C1D" LSID="urn:lsid:plazi:treatment:C97AB10B4B33FFF662D27399FDC9683E:440C56C64B33FFF061A1754DFB4E6C1D" box="[996,1184,1288,1315]" country="Greece" county="Thassos Island" latitude="40.75822" longLatPrecision="1" longitude="24.638138" municipality="Drakotrypa cave" name="Rachoni village" pageId="2" pageNumber="131" stateProvince="Voreio Aigaio">Rachoni village</location>
,
<geoCoordinate id="24E766DA4B33FFF066E9754DFAAE6C1D" box="[1196,1344,1288,1315]" degrees="40" direction="north" minutes="45" orientation="latitude" pageId="2" pageNumber="131" precision="1" seconds="29.6" value="40.75822">N40 45 29.6</geoCoordinate>
<geoCoordinate id="24E766DA4B33FFF0670D754DFF236C7B" degrees="24" direction="east" minutes="38" orientation="longitude" pageId="2" pageNumber="131" precision="1" seconds="17.3" value="24.638138">E24 38 17.3</geoCoordinate>
,
<elevation id="CAFEE72E4B33FFF0629C756FFE8C6C7B" box="[217,354,1322,1349]" metricMagnitude="2" metricUnit="m" metricValue="1.0699999999999998" pageId="2" pageNumber="131" unit="m" value="107.0">
<quantity id="862BADF84B33FFF0629C756FFECA6C7B" box="[217,292,1322,1349]" metricMagnitude="2" metricUnit="m" metricValue="1.0699999999999998" pageId="2" pageNumber="131" unit="m" value="107.0">107 m</quantity>
a.s.l.
</elevation>
(
<specimenCode id="1175A8664B33FFF06334756FFDF96C7B" box="[369,535,1322,1349]" collectionCode="ZMH" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="2" pageNumber="131" type="Museum">ZMH 140791</specimenCode>
:
<specimenCount id="57D5CB944B33FFF06067756FFD5B6C7B" box="[546,693,1322,1349]" count="5" pageId="2" pageNumber="131" type="generic">5 specimens</specimenCount>
,
<specimenCount id="57D5CB944B33FFF06085756FFCBE6C7B" box="[704,848,1322,1349]" count="4" pageId="2" pageNumber="131" type="generic">4 specimens</specimenCount>
in coll.
<collectorName id="EC2665CB4B33FFF061F5756FFC166C7B" box="[944,1016,1322,1349]" pageId="2" pageNumber="131">Glöer</collectorName>
)
</materialsCitation>
.
</paragraph>
<paragraph id="416C001D4B33FFF062D2752BFBE26C94" blockId="2.[151,1436,1389,1450]" pageId="2" pageNumber="131">
<emphasis id="73A7DC0F4B33FFF062D2752BFEAB6CB6" bold="true" box="[151,325,1390,1416]" pageId="2" pageNumber="131">Type locality:</emphasis>
<materialsCitation id="F1BB0A404B33FFF0630A7528FBE66C94" country="Greece" elevation="115" latitude="40.68364" location="Skala Potamia" longLatPrecision="1" longitude="24.771305" municipality="Thassos Island" pageId="2" pageNumber="131" specimenCount="1" stateProvince="Voreio Aigaio">
<collectingCountry id="39C4408D4B33FFF0630A7528FE4B6CB6" box="[335,421,1389,1416]" name="Greece" pageId="2" pageNumber="131">Greece</collectingCountry>
,
<collectingMunicipality id="A1089A674B33FFF063F67528FD8B6CB6" box="[435,613,1389,1416]" pageId="2" pageNumber="131">Thassos Island</collectingMunicipality>
,
<collectingRegion id="8317CEFF4B33FFF060367528FCF36CB6" box="[627,797,1389,1416]" country="Greece" name="Voreio Aigaio" pageId="2" pageNumber="131">North Aegean</collectingRegion>
, a spring (water source) south of
<location id="440C56C64B33FFF066F17528FAB06CB6" LSID="urn:lsid:plazi:treatment:C97AB10B4B33FFF662D27399FDC9683E:440C56C64B33FFF066F17528FAB06CB6" box="[1204,1374,1389,1416]" country="Greece" latitude="40.68364" longLatPrecision="1" longitude="24.771305" municipality="Thassos Island" name="Skala Potamia" pageId="2" pageNumber="131" stateProvince="Voreio Aigaio">Skala Potamia</location>
near the round road of the island,
<geoCoordinate id="24E766DA4B33FFF063AC75CAFD956C94" box="[489,635,1423,1450]" degrees="40" direction="north" minutes="41" orientation="latitude" pageId="2" pageNumber="131" precision="1" seconds="01.1" value="40.68364">N40 41 01.1</geoCoordinate>
<geoCoordinate id="24E766DA4B33FFF060C775CAFCFD6C94" box="[642,787,1423,1450]" degrees="24" direction="east" minutes="46" orientation="longitude" pageId="2" pageNumber="131" precision="1" seconds="16.7" value="24.771305">E24 46 16.7</geoCoordinate>
,
<elevation id="CAFEE72E4B33FFF0615A75CAFC496C94" box="[799,935,1423,1450]" metricMagnitude="2" metricUnit="m" metricValue="1.15" pageId="2" pageNumber="131" unit="m" value="115.0">
<quantity id="862BADF84B33FFF0615A75CAFC866C94" box="[799,872,1423,1450]" metricMagnitude="2" metricUnit="m" metricValue="1.15" pageId="2" pageNumber="131" unit="m" value="115.0">115 m</quantity>
a.s.l.
</elevation>
(
<figureCitation id="D9E81C984B33FFF061F275CAFC156C94" box="[951,1019,1423,1450]" captionStart="Figure 1" captionStartId="1.[151,226,1019,1043]" captionTargetBox="[393,1196,606,1000]" captionTargetId="figure-224@1.[390,1196,606,1000]" captionTargetPageId="1" captionText="Figure 1. Faksimile of the original description of Amnicola charpentieri Roth, 1855" figureDoi="http://doi.org/10.5281/zenodo.13232494" httpUri="https://zenodo.org/record/13232494/files/figure.png" pageId="2" pageNumber="131">Fig. 1</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="09C953964B33FFF062D27596FBD86F31" pageId="2" pageNumber="131" type="etymology">
<paragraph id="416C001D4B33FFF062D27596FBD86F31" blockId="2.[151,1436,1490,1551]" pageId="2" pageNumber="131">
<emphasis id="73A7DC0F4B33FFF062D27596FEC66CD3" bold="true" box="[151,296,1491,1517]" pageId="2" pageNumber="131">Etymology:</emphasis>
Named in honor of Prof. Petar Beron (National Natural History Museum -
<collectingRegion id="8317CEFF4B33FFF0669D7597FAF66CD3" box="[1240,1304,1490,1517]" country="Bulgaria" name="Sofiya-Grad" pageId="2" pageNumber="131">Sofia</collectingRegion>
) who paid attention on this island and inspired D. Georgiev in its young age by his books.
</paragraph>
</subSubSection>
<subSubSection id="09C953964B33FFF162D2767DFECC683E" lastPageId="3" lastPageNumber="132" pageId="2" pageNumber="131" type="description">
<paragraph id="416C001D4B33FFF062D2767DFAD66E23" blockId="2.[151,1437,1591,1821]" pageId="2" pageNumber="131">
<emphasis id="73A7DC0F4B33FFF062D2767DFEDC6F6C" bold="true" box="[151,306,1592,1618]" pageId="2" pageNumber="131">Description:</emphasis>
Shell: The corneous, small shell is ovate with a prominent body whorl. The 4-4.5 whorls are convex with a weak suture, with smooth surface and fine growth lines. The aperture is rounded ovate with a sharp periostome thickened at the columella. The umbilicus is closed (
<figureCitation id="D9E81C984B33FFF061AB763EFBD46FA8" box="[1006,1082,1659,1686]" captionStart="Figure 2" captionStartId="2.[151,226,729,753]" captionTargetBox="[151,1437,223,711]" captionTargetId="figure-21@2.[151,1437,223,711]" captionTargetPageId="2" captionText="Figure 2. The sampling sites of the species under discussion on Thassos. 1: Rachoni village: type locality of Bythinella rachonica n. sp., localiy of Grossuana beroni n. sp.; 2: Drakotrype cave, Panagia village: localiy of Grossuana beroni n. sp.; 3: type localiy of Grossuana beroni; 4: type locality of Grossuana thasia n. sp., 5: Pernes Mountain, type locality of Bythinella reischuetzi n. sp., 6: Hymettos, type locality of Bythinella charpentieri (Roth, 1855)." figureDoi="http://doi.org/10.5281/zenodo.13232496" httpUri="https://zenodo.org/record/13232496/files/figure.png" pageId="2" pageNumber="131">Fig. 2</figureCitation>
). The operculum is dark red. Animal: The animal is mostly dirty whitish or gray colored. The mantle is white or gray with a black wide stripe in its middle. The snout and tentacles are whitish or gray. Small patches of black pigment are visible around eyes and at the base of the snout. Penis: The penis is nearly regularly broad with a long tapered penis-tip and a bilobed outgrowth in the middle part on the left side. There is a dark spot near the penis-tip.
</paragraph>
<paragraph id="416C001D4B33FFF162D27703FECC683E" blockId="2.[151,1436,1861,1990]" lastBlockId="3.[151,1436,195,256]" lastPageId="3" lastPageNumber="132" pageId="2" pageNumber="131">
<emphasis id="73A7DC0F4B33FFF062D27703FE256E5E" bold="true" box="[151,459,1862,1888]" pageId="2" pageNumber="131">Differentiating features:</emphasis>
<taxonomicName id="86D37B9E4B33FFF063927703FD5E6E5E" authorityName="Georgiev &amp; Glöer" authorityYear="2020" box="[471,688,1861,1888]" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="2" pageNumber="131" phylum="Mollusca" rank="species" species="beroni" status="sp. nov.">
<emphasis id="73A7DC0F4B33FFF063927703FD5E6E5E" box="[471,688,1861,1888]" italics="true" pageId="2" pageNumber="131">Grossuana beroni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="689461744B33FFF060F97703FCEE6E5E" box="[700,768,1862,1888]" pageId="2" pageNumber="131" rank="species">n. sp.</taxonomicNameLabel>
is most similar to
<taxonomicName id="86D37B9E4B33FFF061B57703FF326EBC" authority="(Westerlund, 1881)" baseAuthorityName="Westerlund" baseAuthorityYear="1881" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="2" pageNumber="131" phylum="Mollusca" rank="species" species="marginata">
<emphasis id="73A7DC0F4B33FFF061B57703FB166E61" box="[1008,1272,1862,1888]" italics="true" pageId="2" pageNumber="131">Grossuana marginata</emphasis>
(Westerlund, 1881)
</taxonomicName>
from which it differs by more regularly growing whorls, sharper and thinner periostome and lighter soft body pigmentation. The new species is also similar to
<taxonomicName id="86D37B9E4B33FFF0612F77CFFE6A6EF8" authority="Gloer, Reuselaars &amp; Papavasileiou, 2018" authorityName="Gloer, Reuselaars &amp; Papavasileiou" authorityYear="2018" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="2" pageNumber="131" phylum="Mollusca" rank="species" species="sidironerensis">
<emphasis id="73A7DC0F4B33FFF0612F77CFFB756E9A" box="[874,1179,1929,1956]" italics="true" pageId="2" pageNumber="131">Grossuana sidironerensis</emphasis>
<bibRefCitation id="25427DEC4B33FFF066E377CCFE6A6EF8" author="Gloer, P. &amp; Reuselaars, R. &amp; Papavasileiou, K." pageId="2" pageNumber="131" pagination="14 - 19" refId="ref4461" refString="Gloer, P., Reuselaars, R. &amp; Papavasileiou, K. (2018) Grossuana Radoman, 1973 from Macedonia (Greece) (Gastropoda: Truncatelloidea) with the description of three new species. Ecologica Montenegrina, 17, 14 - 19. https: // doi. org / 10.37828 / em. 2018.17.2" type="journal article" year="2018">Glöer, Reuselaars &amp; Papavasileiou, 2018</bibRefCitation>
</taxonomicName>
described from the area of Drama at the mainland of
<collectingCountry id="39C4408D4B33FFF0664177EEFBB86EF8" box="[1028,1110,1963,1990]" name="Greece" pageId="2" pageNumber="131">Greece</collectingCountry>
(
<bibRefCitation id="25427DEC4B33FFF0662377EEFAD76EF8" author="Gloer, P. &amp; Reuselaars, R. &amp; Papavasileiou, K." box="[1126,1337,1963,1990]" pageId="2" pageNumber="131" pagination="14 - 19" refId="ref4461" refString="Gloer, P., Reuselaars, R. &amp; Papavasileiou, K. (2018) Grossuana Radoman, 1973 from Macedonia (Greece) (Gastropoda: Truncatelloidea) with the description of three new species. Ecologica Montenegrina, 17, 14 - 19. https: // doi. org / 10.37828 / em. 2018.17.2" type="journal article" year="2018">Glöer et al., 2018</bibRefCitation>
) but the last species has a penis with shorter tip, black colored mantle, and its whorls are fast growing (as in
<taxonomicName id="86D37B9E4B32FFF1673A7081FEFF683E" baseAuthorityName="Westerlund" baseAuthorityYear="1881" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="132" phylum="Mollusca" rank="species" species="marginata">
<emphasis id="73A7DC0F4B32FFF1673A7081FEFF683E" italics="true" pageId="3" pageNumber="132">G. marginata</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="09C953964B32FFF662D2716CFDC9683E" lastPageId="4" lastPageNumber="133" pageId="3" pageNumber="132" type="distribution">
<paragraph id="416C001D4B32FFF162D2716CFCDE68B9" blockId="3.[151,1436,297,391]" pageId="3" pageNumber="132">
<emphasis id="73A7DC0F4B32FFF162D2716CFED4687D" bold="true" box="[151,314,297,323]" pageId="3" pageNumber="132">Distribution:</emphasis>
Known only from few springs on Thassos Island: a spring (water source) south of Skala Potamia near the round road of the island (
<typeStatus id="9E68BEBF4B32FFF160CB710FFD2C685B" box="[654,706,330,357]" pageId="3" pageNumber="132">type</typeStatus>
locality), spring below Drakotrypa cave, Panagia village, and a big spring in the yard of the church of Rachoni village.
</paragraph>
<caption id="15AC50954B32FFF162D275ECFD866CC0" ID-DOI="http://doi.org/10.5281/zenodo.13232498" ID-Zenodo-Dep="13232498" httpUri="https://zenodo.org/record/13232498/files/figure.png" pageId="3" pageNumber="132" startId="3.[151,236,1449,1473]" targetBox="[151,1439,404,1431]" targetPageId="3" targetType="figure">
<paragraph id="416C001D4B32FFF162D275ECFD866CC0" blockId="3.[151,1436,1449,1535]" pageId="3" pageNumber="132">
<emphasis id="73A7DC0F4B32FFF162D275ECFEF66CFF" bold="true" box="[151,280,1449,1473]" pageId="3" pageNumber="132">Figures 3-5</emphasis>
.
<emphasis id="73A7DC0F4B32FFF1636D75ECFEDB6CFF" bold="true" box="[296,309,1449,1473]" pageId="3" pageNumber="132">3</emphasis>
: Type locality of
<taxonomicName id="86D37B9E4B32FFF163BE75ECFD516CFF" authorityName="Georgiev &amp; Glöer" authorityYear="2020" box="[507,703,1449,1473]" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="132" phylum="Mollusca" rank="species" species="beroni" status="sp. nov.">
<emphasis id="73A7DC0F4B32FFF163BE75ECFD516CFF" box="[507,703,1449,1473]" italics="true" pageId="3" pageNumber="132">Grossuana beroni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="689461744B32FFF1608D75ECFCEA6CFE" box="[712,772,1449,1472]" pageId="3" pageNumber="132" rank="species">n. sp.</taxonomicNameLabel>
, water source south of Skala Potamia near the round road of the island.
<emphasis id="73A7DC0F4B32FFF163557582FEF36CE1" bold="true" box="[272,285,1479,1503]" pageId="3" pageNumber="132">4</emphasis>
: Type locality of
<taxonomicName id="86D37B9E4B32FFF163A2758DFD486CE1" authorityName="Georgiev &amp; Glöer" authorityYear="2020" box="[487,678,1479,1503]" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="132" phylum="Mollusca" rank="species" species="thasia" status="sp. nov.">
<emphasis id="73A7DC0F4B32FFF163A2758DFD486CE1" box="[487,678,1479,1503]" italics="true" pageId="3" pageNumber="132">Grossuana thasia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="689461744B32FFF160F5758DFD036CE1" box="[688,749,1480,1503]" pageId="3" pageNumber="132" rank="species">n. sp.</taxonomicNameLabel>
, a spring south of Kinira near the round road of the island.
<emphasis id="73A7DC0F4B32FFF167C27582FA7A6CE1" bold="true" box="[1415,1428,1479,1503]" pageId="3" pageNumber="132">5</emphasis>
: Type locality of
<taxonomicName id="86D37B9E4B32FFF1630D75A3FDC66CC0" authorityName="Georgiev &amp; Glöer" authorityYear="2020" box="[328,552,1510,1534]" class="Gastropoda" family="Bythinellidae" genus="Bythinella" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="132" phylum="Mollusca" rank="species" species="rachonica" status="sp. nov.">
<emphasis id="73A7DC0F4B32FFF1630D75A3FDC66CC0" box="[328,552,1510,1534]" italics="true" pageId="3" pageNumber="132">Bythinella rachonica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="689461744B32FFF1606A75A2FD866CC0" box="[559,616,1511,1534]" pageId="3" pageNumber="132" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</caption>
<caption id="15AC50954B32FFF162D277F1FDD46ED5" ID-DOI="http://doi.org/10.5281/zenodo.13232500" ID-Zenodo-Dep="13232500" httpUri="https://zenodo.org/record/13232500/files/figure.png" pageId="3" pageNumber="132" startId="3.[151,236,1972,1996]" targetBox="[151,1444,1565,1953]" targetPageId="3" targetType="figure">
<paragraph id="416C001D4B32FFF162D277F1FDD46ED5" blockId="3.[151,1436,1972,2027]" pageId="3" pageNumber="132">
<emphasis id="73A7DC0F4B32FFF162D277F1FEC06EF2" bold="true" box="[151,302,1972,1996]" pageId="3" pageNumber="132">Figures 6-10.</emphasis>
The
<taxonomicName id="86D37B9E4B32FFF1632977F0FDC16EF2" box="[364,559,1972,1996]" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="132" phylum="Mollusca" rank="species" species="undefined-6">
<emphasis id="73A7DC0F4B32FFF1632977F0FE0F6EF2" box="[364,481,1973,1996]" italics="true" pageId="3" pageNumber="132">Grossuana</emphasis>
spp.
<emphasis id="73A7DC0F4B32FFF1606777F1FDC16EF2" bold="true" box="[546,559,1972,1996]" pageId="3" pageNumber="132">6</emphasis>
</taxonomicName>
<emphasis id="73A7DC0F4B32FFF1606A77F1FDAB6EF2" bold="true" box="[559,581,1972,1996]" pageId="3" pageNumber="132">-7</emphasis>
:
<taxonomicName id="86D37B9E4B32FFF1601277F0FD2F6EF2" authorityName="Georgiev &amp; Glöer" authorityYear="2020" box="[599,705,1972,1996]" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="132" phylum="Mollusca" rank="species" species="beroni" status="n. sp.">
<emphasis id="73A7DC0F4B32FFF1601277F0FD2F6EF2" box="[599,705,1972,1996]" italics="true" pageId="3" pageNumber="132">G. beroni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="689461744B32FFF1608E77F0FCE66EF2" box="[715,776,1973,1996]" pageId="3" pageNumber="132" rank="species">n. sp.</taxonomicNameLabel>
(paratypes),
<emphasis id="73A7DC0F4B32FFF161DE77F1FC466EF2" bold="true" box="[923,936,1972,1996]" pageId="3" pageNumber="132">8</emphasis>
: penis of
<taxonomicName id="86D37B9E4B32FFF1665877F0FB696EF2" authorityName="Georgiev &amp; Glöer" authorityYear="2020" box="[1053,1159,1972,1996]" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="132" phylum="Mollusca" rank="species" species="beroni" status="n. sp.">
<emphasis id="73A7DC0F4B32FFF1665877F0FB696EF2" box="[1053,1159,1972,1996]" italics="true" pageId="3" pageNumber="132">G. beroni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="689461744B32FFF166D477F0FB206EF2" box="[1169,1230,1973,1996]" pageId="3" pageNumber="132" rank="species">n. sp.</taxonomicNameLabel>
<emphasis id="73A7DC0F4B32FFF1669D77F1FB0B6EF2" bold="true" box="[1240,1253,1972,1996]" pageId="3" pageNumber="132">9</emphasis>
:
<taxonomicName id="86D37B9E4B32FFF166B277F0FAB26EF2" authorityName="Georgiev &amp; Glöer" authorityYear="2020" box="[1271,1372,1972,1996]" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="132" phylum="Mollusca" rank="species" species="thasia">
<emphasis id="73A7DC0F4B32FFF166B277F0FAB26EF2" box="[1271,1372,1972,1996]" italics="true" pageId="3" pageNumber="132">G. thasia</emphasis>
</taxonomicName>
n. sp (holotype),
<emphasis id="73A7DC0F4B32FFF163517796FEC16ED5" bold="true" box="[276,303,2003,2027]" pageId="3" pageNumber="132">10</emphasis>
: penis of
<taxonomicName id="86D37B9E4B32FFF163DD7791FE146ED5" authorityName="Georgiev &amp; Glöer" authorityYear="2020" box="[408,506,2003,2027]" class="Gastropoda" family="Hydrobiidae" genus="Grossuana" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="132" phylum="Mollusca" rank="species" species="thasia" status="n. sp.">
<emphasis id="73A7DC0F4B32FFF163DD7791FE146ED5" box="[408,506,2003,2027]" italics="true" pageId="3" pageNumber="132">G. thasia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="689461744B32FFF160447791FDD46ED5" box="[513,570,2004,2027]" pageId="3" pageNumber="132" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="416C001D4B35FFF662D27081FDC9683E" blockId="4.[151,1436,195,256]" pageId="4" pageNumber="133">
<emphasis id="73A7DC0F4B35FFF662D27081FEEF69E0" bold="true" box="[151,257,196,222]" pageId="4" pageNumber="133">Habitat:</emphasis>
Karst spring in a mixed broad leaf forest (mainly
<taxonomicName id="86D37B9E4B35FFF6611C7086FBD469E0" box="[857,1082,195,222]" class="Magnoliopsida" family="Platanaceae" genus="Platanus" kingdom="Plantae" order="Proteales" pageId="4" pageNumber="133" phylum="Tracheophyta" rank="species" species="orientalis">
<emphasis id="73A7DC0F4B35FFF6611C7086FBD469E0" box="[857,1082,195,222]" italics="true" pageId="4" pageNumber="133">Platanus orientalis</emphasis>
</taxonomicName>
and
<taxonomicName id="86D37B9E4B35FFF666337081FAF869E0" box="[1142,1302,195,222]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="133" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="73A7DC0F4B35FFF666337081FB3469E0" box="[1142,1242,196,222]" italics="true" pageId="4" pageNumber="133">Quercus</emphasis>
spp.
</taxonomicName>
). Found on moss, dry leaves, sand and stones.
</paragraph>
</subSubSection>
</treatment>
</document>