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<document id="244DEF9AC179BAE1ACE339DB24B8ED86" ID-DOI="10.5252/adansonia2024v46a7" ID-ISSN="1639-4798" ID-Zenodo-Dep="11058077" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="guilherme" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1713947152188" checkinUser="plazi" docAuthor="Bordbar, Firouzeh &amp; Mirtadzadini, Mansour" docDate="2024" docId="B4697C4FFF87FFDEFC6725E0FBB47795" docLanguage="en" docName="adansonia.2024.46.7.pdf" docOrigin="Adansonia (3) 46 (7)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/adansonia2024v46a7.pdf" docStyle="DocumentStyle:2FE19AC740987067EC2A6C89A85F9648.8:Adansonia.2018-.journal_article" docStyleId="2FE19AC740987067EC2A6C89A85F9648" docStyleName="Adansonia.2018-.journal_article" docStyleVersion="8" docTitle="Satureja sahendica Bornm." docType="treatment" docVersion="4" lastPageNumber="60" masterDocId="48500437FF96FFCFFFBC2417FF81713B" masterDocTitle="A morphological re-evaluation of the taxonomic status of Satureja L. (Lamiaceae, Nepetoideae, Mentheae) from flora of Iran" masterLastPageNumber="63" masterPageNumber="45" pageNumber="60" updateTime="1714137651825" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="1991582B0A2E984398A2A22BAEA96361">A morphological re-evaluation of the taxonomic status of Satureja L. (Lamiaceae, Nepetoideae, Mentheae) from flora of Iran</mods:title>
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<mods:namePart id="F503064EB396587E673793B96E708592">Bordbar, Firouzeh</mods:namePart>
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<mods:namePart id="B621B05F3FCA2FACE1DD8E50DFC6A064">Mirtadzadini, Mansour</mods:namePart>
<mods:affiliation id="F9F7E3567963088739222F9F1322B76E">Department of Biology, Faculty of Science, Shahid Bahonar University of Kerman, Kerman (Iran) bordbar @ uk. ac. ir, fbordbar 2000 @ yahoo. com (corresponding author) mirtadz @ uk. ac. ir</mods:affiliation>
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<mods:title id="DA77EF21A0F2F43C717D66BC0F134C2B">Adansonia</mods:title>
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<taxonomicName id="FBC0B6DAFF87FFDEFC6725E0FA83732A" ID-CoL="79QGJ" authority="Bornm." authorityName="Bornm." box="[987,1282,503,530]" class="Magnoliopsida" family="Lamiaceae" genus="Satureja" kingdom="Plantae" order="Lamiales" pageId="17" pageNumber="60" phylum="Tracheophyta" rank="species" species="sahendica">
<emphasis id="0EB4114BFF87FFDEFC6725E0FB277329" bold="true" box="[987,1190,503,530]" italics="true" pageId="17" pageNumber="60">Satureja sahendica</emphasis>
Bornm.
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</heading>
</paragraph>
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<subSubSection id="74DA9ED2FF87FFDEFB812600FC18723A" pageId="17" pageNumber="60" type="description">
<paragraph id="3C7FCD59FF87FFDEFB812600FB21730A" blockId="17.[987,1282,503,561]" box="[1085,1184,535,561]" pageId="17" pageNumber="60">
(
<figureCitation id="A4FBD1DCFF87FFDEFBFA2600FB16730A" box="[1094,1175,535,561]" captionStart="FIG" captionStartId="12.[132,143,1473,1490]" captionTargetBox="[132,1455,215,1431]" captionTargetId="figure-209@12.[132,1455,215,1431]" captionTargetPageId="12" captionText="FIG. 2. — Habit and close-up of inflorescence of Satureja L.species from flora of Iran:A, S.kermanica Payandeh,Bordbar &amp; Mirtadz.(Bordbar 1956); B, C, S. khuzistanica Jamzad (Mirtadzadini 3805); D, S. khuzistanica (Mirtadzadini 3804); E, S. macrantha C.A.Mey. (Bordbar 1970); F, S. macrantha (Mirtadzadini 2036); G, S. macrosiphoniaBornm.(Mirtadzadini &amp; Bordbar 5156); H, S. macrosiphoniaBornm.(Mirtadzadini &amp; Bordbar 4157); I, S.mutica Fisch. &amp; C.A.Mey. (Mirtadzadini 3822); J, S. mutica (Mirtadzadini 3832); K, S. sahendica Bornm. (Mirtadzadini 1984)." figureDoi="http://doi.org/10.5281/zenodo.11058081" httpUri="https://zenodo.org/record/11058081/files/figure.png" pageId="17" pageNumber="60">Fig. 2K</figureCitation>
)
</paragraph>
<paragraph id="3C7FCD59FF87FFDEFCFF2647FB03738C" blockId="17.[811,1456,592,1076]" pageId="17" pageNumber="60">
<bibCitation id="E9EAC535FF87FFDEFCFF2647FBC473B9" author="Verhandlungen der Zoologisch-Botanischen Gesellschaft" journal="Wien" page="160" pageId="17" pageNumber="60" score="0.0340967026758743" volume="60" year="1910">
<emphasis id="0EB4114BFF87FFDEFCFF2647FAD47353" box="[835,1365,592,616]" italics="true" pageId="17" pageNumber="60">
Verhandlungen der Zoologisch-Botanischen
<bibCitation id="E9EAC535FF87FFDEFB542647FAD47353" box="[1256,1365,592,616]" journal="Gesellschaft in Wien" page="160" pageId="17" pageNumber="60" score="30.208167052571333" volume="60" year="1910">Gesellschaft</bibCitation>
</emphasis>
in Wien 60: 160 (
<bibRefCitation id="5851B0A8FF87FFDEFC30267DFBBA73B9" author="BORNMULLER J." box="[908,1083,618,642]" pageId="17" pageNumber="60" pagination="61 - 194" refId="ref17789" refString="BORNMULLER J. 1910. - Bearbeitung der von J. A. Knapp im nordwestlichen Persien gesammelten pflanzen. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien 60: 61 - 194." type="journal article" year="1910">Bornmüller 1910</bibRefCitation>
)
</bibCitation>
. —
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Type:
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<collectingCountry id="44D78DC9FF87FFDEFB0F267DFB5C73B9" box="[1203,1245,618,642]" name="Iran" pageId="17" pageNumber="60">Iran</collectingCountry>
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• Persia,
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,
<location id="391F9B82FF87FFDEFC922692FB3373A6" LSID="urn:lsid:plazi:treatment:B4697C4FFF87FFDEFC6725E0FBB47795:391F9B82FF87FFDEFC922692FB3373A6" box="[814,1202,645,669]" country="Iran" name="in declivibus borealibus montis Sahend" pageId="17" pageNumber="60">in declivibus borealibus montis Sahend</location>
;
<date id="487EEB99FF87FFDEFB012692FABB73A6" box="[1213,1338,645,669]" pageId="17" pageNumber="60" value="1884-07-30">
<collectingDate id="583A1271FF87FFDEFB012692FABB73A6" box="[1213,1338,645,669]" pageId="17" pageNumber="60" value="1884-07-30">30.VII.1884</collectingDate>
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;
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s.n.
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;
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,
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[
<specimenCode id="6C666522FF87FFDEFC222688FBA5738C" box="[926,1060,671,695]" collectionCode="WU" country="USA" name="Wayland University" pageId="17" pageNumber="60">WU0043454</specimenCode>
image!])
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.
</paragraph>
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. — Flowering July to November, fruiting October to November.
</paragraph>
</subSubSection>
<subSubSection id="74DA9ED2FF87FFDEFC91270EFBB5750F" pageId="17" pageNumber="60" type="distribution">
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. — This is an endemic species distributed on the rocky slopes of the Irano-Turanian region of NW of
<collectingCountry id="44D78DC9FF87FFDEFCFB2759FCEE725E" box="[839,879,846,869]" name="Iran" pageId="17" pageNumber="60">Iran</collectingCountry>
(
<figureCitation id="A4FBD1DCFF87FFDEFCC12759FC48725D" box="[893,969,846,870]" captionStart="FIG" captionStartId="18.[132,143,1161,1178]" captionTargetBox="[221,1379,204,1132]" captionTargetId="graphics-517@18.[18,979,636,1118]" captionText="FIG. 5. — Distribution map of Satureja L. species in Iran: A, S. macrantha C.A.Mey.; B, S. macrosiphonoa (Coss.) Maire; C, S. sahendica Bornm.; D, S. mutica Fisch. &amp; C.A.Mey. (circle), S. longiflora Boiss. &amp; Hausskn. (square)." figureDoi="http://doi.org/10.5281/zenodo.11072765" httpUri="https://zenodo.org/record/11072765/files/figure.png" pageId="17" pageNumber="60">Fig. 5C</figureCitation>
).
</paragraph>
<paragraph id="3C7FCD59FF87FFDEFC912769FBB5750F" blockId="17.[811,1456,592,1076]" pageId="17" pageNumber="60">
<materialsCitation id="8CA8C704FF87FFDEFC912769FAAB72F0" collectingDate="2016-07-08" collectingDateMax="2016-07-08" collectingDateMin="1984-08-08" collectionCode="MIR" collectorName="Mirtadzadini" country="Iran" county="Mt. Nour" elevation="2459" latitude="35.810833" location="Qaraghaj" longLatPrecision="19" longitude="49.14639" municipality="Mirtadzadini" pageId="17" pageNumber="60" specimenCount="1" stateProvince="East Azerbaijan">
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. —
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<collectingCountry id="44D78DC9FF87FFDEFB9F2769FBCD72AD" box="[1059,1100,894,918]" name="Iran" pageId="17" pageNumber="60">Iran</collectingCountry>
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• NW Iran,
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, south of
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, south west of
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;
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,
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; alt.
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;
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</date>
;
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<collectorName id="9135A88FFF87FFDEFB9027A5FB2D72F1" box="[1068,1196,946,970]" pageId="17" pageNumber="60">Mirtadzadini</collectorName>
1984
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(
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)
</materialsCitation>
•
<materialsCitation id="8CA8C704FF87FFDEFA8227A5FB9E72C4" collectingDate="1984-08-08" collectingDateMax="2016-07-08" collectingDateMin="1984-08-08" collectionCode="IRAN" collectorName="Terme &amp; Moussavi" country="Iran" county="Mt. Nour" elevation="2275" latitude="35.810833" location="Kandovan" longLatPrecision="19" longitude="49.14639" municipality="Osku" pageId="17" pageNumber="60" specimenCount="1" stateProvince="Azarbayjan-e Sharqi">
<collectingMunicipality id="DC1B5723FF87FFDEFA8227A5FAF772F1" box="[1342,1398,946,970]" pageId="17" pageNumber="60">Osku</collectingMunicipality>
,
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,
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,
<collectingCounty id="D51EB5D5FF87FFDEFC0927DAFB9672DF" box="[949,1047,973,996]" pageId="17" pageNumber="60">Mt. Nour</collectingCounty>
: alt.
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</quantity>
;
<date id="487EEB99FF87FFDEFB6D27DAFAD472DE" box="[1233,1365,973,997]" pageId="17" pageNumber="60" value="1984-08-08">
<collectingDate id="583A1271FF87FFDEFB6D27DAFAD472DE" box="[1233,1365,973,997]" pageId="17" pageNumber="60" value="1984-08-08">08.VIII.1984</collectingDate>
</date>
;
<emphasis id="0EB4114BFF87FFDEFAE027D9FC4B72C4" italics="true" pageId="17" pageNumber="60">
<collectorName id="9135A88FFF87FFDEFAE027D9FA1472DE" box="[1372,1429,974,997]" pageId="17" pageNumber="60">Terme</collectorName>
&amp;
<collectorName id="9135A88FFF87FFDEFC9127FFFC0572C4" box="[813,900,1000,1023]" pageId="17" pageNumber="60">Moussavi</collectorName>
25038
</emphasis>
(
<collectionCode id="5AD1559CFF87FFDEFC6527F0FB9972C4" box="[985,1048,999,1023]" pageId="17" pageNumber="60">IRAN</collectionCode>
)
</materialsCitation>
•
<materialsCitation id="8CA8C704FF87FFDEFB8427F0FBAE750F" collectingDate="2012-08-10" collectingDateMax="2016-07-08" collectingDateMin="1984-08-08" collectionCode="MIR" collectorName="Mirtadzadini" country="Iran" county="Mt. Nour" elevation="1716" latitude="35.810833" location="Kiseh-Jin to Dashtak" longLatPrecision="19" longitude="49.14639" municipality="Mirtadzadini" pageId="17" pageNumber="60" specimenCount="1" stateProvince="Qazvin Prov.">
<collectingRegion id="FE0403BBFF87FFDEFB8427F0FB3A72C4" box="[1080,1211,999,1023]" country="Iran" name="Qazvin" pageId="17" pageNumber="60">Qazvin Prov.</collectingRegion>
,
<quantity id="FB3860BCFF87FFDEFB7927F0FB7872C4" box="[1221,1273,999,1023]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="60" unit="km" value="5.0">
<locationDeviation id="2DCDD47BFF87FFDEFB7927F0FB7872C4" box="[1221,1273,999,1023]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="60" unit="km" value="5.0">5 km</locationDeviation>
</quantity>
from
<location id="391F9B82FF87FFDEFA8427F0FC047521" LSID="urn:lsid:plazi:treatment:B4697C4FFF87FFDEFC6725E0FBB47795:391F9B82FF87FFDEFA8427F0FC047521" country="Iran" county="Mt. Nour" latitude="35.810833" longLatPrecision="19" longitude="49.14639" municipality="Mirtadzadini" name="Kiseh-Jin to Dashtak" pageId="17" pageNumber="60" stateProvince="Qazvin Prov.">Kiseh-Jin to Dashtak</location>
;
<geoCoordinate id="59F4AB9EFF87FFDEFC2C2015FB8E7521" box="[912,1039,1026,1050]" degrees="35" direction="north" minutes="48" orientation="latitude" pageId="17" pageNumber="60" precision="15" seconds="39" value="35.810833">35°48’39”N</geoCoordinate>
,
<geoCoordinate id="59F4AB9EFF87FFDEFBA02015FB077521" box="[1052,1158,1026,1050]" degrees="49" direction="east" minutes="8" orientation="longitude" pageId="17" pageNumber="60" precision="15" seconds="47" value="49.14639">49°8’47”E</geoCoordinate>
; alt.
<quantity id="FB3860BCFF87FFDEFB032015FA927522" box="[1215,1299,1026,1050]" metricMagnitude="3" metricUnit="m" metricValue="1.716" pageId="17" pageNumber="60" unit="m" value="1716.0">
<elevation id="B7ED2A6AFF87FFDEFB032015FA927522" box="[1215,1299,1026,1050]" metricMagnitude="3" metricUnit="m" metricValue="1.716" pageId="17" pageNumber="60" unit="m" value="1716.0">1716 m</elevation>
</quantity>
;
<date id="487EEB99FF87FFDEFAA32015FA2A7521" box="[1311,1451,1026,1050]" pageId="17" pageNumber="60" value="2012-08-10">
<collectingDate id="583A1271FF87FFDEFAA32015FA2A7521" box="[1311,1451,1026,1050]" pageId="17" pageNumber="60" value="2012-08-10">10.VIII.2012</collectingDate>
</date>
;
<emphasis id="0EB4114BFF87FFDEFC92200BFC69750F" box="[814,1000,1052,1076]" italics="true" pageId="17" pageNumber="60">
<collectorName id="9135A88FFF87FFDEFC92200BFC2F750F" box="[814,942,1052,1076]" pageId="17" pageNumber="60">Mirtadzadini</collectorName>
1985
</emphasis>
(
<collectionCode id="5AD1559CFF87FFDEFC48200BFBA8750F" box="[1012,1065,1052,1076]" collectionName="MIR" pageId="17" pageNumber="60">MIR</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="74DA9ED2FF87FFDEFC912041FA307735" pageId="17" pageNumber="60" type="description">
<paragraph id="3C7FCD59FF87FFDEFC912041FC3C7555" blockId="17.[810,1457,1110,1551]" box="[813,957,1110,1136]" pageId="17" pageNumber="60">
<smallCapsWord id="3A995B85FF87FFDEFC912041FC3C7555" baselines="1129,1129" box="[813,957,1110,1136]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="17" pageNumber="60">DESCRIPTION</smallCapsWord>
</paragraph>
<paragraph id="3C7FCD59FF87FFDEFC912062FA307735" blockId="17.[810,1457,1110,1551]" pageId="17" pageNumber="60">
Aromatic perennial,
<quantity id="FB3860BCFF87FFDEFBBF2062FBEB75B4" box="[1027,1130,1141,1168]" metricMagnitude="-1" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="17" pageNumber="60" unit="cm" value="22.5" valueMax="25.0" valueMin="20.0">20-25 cm</quantity>
high, stems erect-procumbent, covered with white short hairs and sessile glands. Leaves sessile, flattened or conduplicate, linear-oblong to lanceolate,
<quantity id="FB3860BCFF87FFDEFAF120A2FA2F75F4" box="[1357,1454,1205,1231]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="17" pageNumber="60" unit="mm" value="10.0" valueMax="12.0" valueMin="8.0">8-12 mm</quantity>
long,
<quantity id="FB3860BCFF87FFDEFCDB20C2FC3E75D4" box="[871,959,1237,1263]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="17" pageNumber="60" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1-2 mm</quantity>
wide, attenuate at base, apex obtusiusculate or acute, with short simple hairs over sessile glands, floral leaves similar to the cauline leaves in shape. Verticillasters distant or approximate, 1-6 flowers in verticillaster, flowers often sessile or with a short pedicel,
<emphasis id="0EB4114BFF87FFDEFB922142FBBF7455" box="[1070,1086,1365,1390]" italics="true" pageId="17" pageNumber="60">c.</emphasis>
<quantity id="FB3860BCFF87FFDEFBF82143FB087454" box="[1092,1161,1364,1391]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="60" unit="mm" value="1.0">1 mm</quantity>
. Calyx green, campanulate-tubular, covered with short hairs and sessile glands on the outside,
<quantity id="FB3860BCFF87FFDEFCDD2183FC387494" box="[865,953,1428,1455]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="17" pageNumber="60" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4-6 mm</quantity>
, superior teeth triangular-subulate, inferior teeth lanceolate-subulate. Corolla white,
<quantity id="FB3860BCFF87FFDEFB1B21A3FA8F74F4" box="[1191,1294,1460,1487]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="17" pageNumber="60" unit="mm" value="10.0" valueMax="12.0" valueMin="8.0">8-12 mm</quantity>
long. Stamens exserted from the tube. Style exserted from the tube. Nutlets elliptic,brown to dark brown,
<quantity id="FB3860BCFF87FFDEFBFB21E3FB407735" box="[1095,1217,1524,1551]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="17" pageNumber="60" unit="mm" value="1.35" valueMax="1.5" valueMin="1.2">1.2-1.5 mm</quantity>
long,
<quantity id="FB3860BCFF87FFDEFB4021E3FAF47735" box="[1276,1397,1524,1550]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="17" pageNumber="60" unit="mm" value="0.75" valueMax="0.8" valueMin="0.7">0.7-0.8mm</quantity>
wide.
</paragraph>
</subSubSection>
<subSubSection id="74DA9ED2FF87FFDEFC912223FBB47795" pageId="17" pageNumber="60" type="discussion">
<paragraph id="3C7FCD59FF87FFDEFC912223FC107777" blockId="17.[813,1457,1588,1710]" box="[813,913,1588,1614]" pageId="17" pageNumber="60">
<smallCapsWord id="3A995B85FF87FFDEFC912223FC107777" baselines="1607,1607" box="[813,913,1588,1614]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Remarks" pageId="17" pageNumber="60">REMARKS</smallCapsWord>
</paragraph>
<paragraph id="3C7FCD59FF87FFDEFC912243FBB47795" blockId="17.[813,1457,1588,1710]" pageId="17" pageNumber="60">
<taxonomicName id="FBC0B6DAFF87FFDEFC912243FC2F7756" authorityName="Bornm. (Mirtadzadini" authorityYear="1984" box="[813,942,1619,1646]" class="Magnoliopsida" family="Lamiaceae" genus="Satureja" kingdom="Plantae" order="Lamiales" pageId="17" pageNumber="60" phylum="Tracheophyta" rank="species" species="sahendica">
<emphasis id="0EB4114BFF87FFDEFC912243FC2F7756" box="[813,942,1619,1646]" italics="true" pageId="17" pageNumber="60">S. sahendica</emphasis>
</taxonomicName>
is morphologically close to
<taxonomicName id="FBC0B6DAFF87FFDEFB5B2243FACD7756" box="[1255,1356,1620,1646]" class="Magnoliopsida" family="Lamiaceae" genus="Satureja" kingdom="Plantae" order="Lamiales" pageId="17" pageNumber="60" phylum="Tracheophyta" rank="species" species="mutica">
<emphasis id="0EB4114BFF87FFDEFB5B2243FACD7756" box="[1255,1356,1620,1646]" italics="true" pageId="17" pageNumber="60">S. mutica</emphasis>
</taxonomicName>
but they are recognized by the corolla tube obviously exserted from the calyx in
<taxonomicName id="FBC0B6DAFF87FFDEFC132283FBAE7796" authorityName="Bornm. (Mirtadzadini" authorityYear="1984" box="[943,1071,1683,1710]" class="Magnoliopsida" family="Lamiaceae" genus="Satureja" kingdom="Plantae" order="Lamiales" pageId="17" pageNumber="60" phylum="Tracheophyta" rank="species" species="sahendica">
<emphasis id="0EB4114BFF87FFDEFC132283FBAE7796" box="[943,1071,1683,1710]" italics="true" pageId="17" pageNumber="60">S. sahendica</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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