plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/B3/6D/06/B36D0606FFD7AC1FFF28FC99FC78FF2D.xml

388 lines
55 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="FFC0439B1431CA23065EBBE89165029F" ID-DOI="10.11646/zootaxa.5492.3.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="13234851" ID-ZooBank="8F81F053-DE78-4684-BCC9-875AE0C41615" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722932859467" checkinUser="plazi" docAuthor="Schmidt, Joachim &amp; Merene, Yeshitla" docDate="2024" docId="B36D0606FFD7AC1FFF28FC99FC78FF2D" docLanguage="en" docName="zootaxa.5492.3.3.pdf" docOrigin="Zootaxa 5492 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5492.3.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trechus (Abyssinotus) apertus Schmidt &amp; Merene 2024, sp. n." docType="treatment" docVersion="2" lastPageNumber="346" masterDocId="4F547E7EFFD6AC1BFFBFFFDAFFC3FFF8" masterDocTitle="Trechus species from Mt. Choke of northern Ethiopia related to T. niloticus (Quéinnec &amp; Ollivier) with notable male genital morphology (Carabidae: Trechini)" masterLastPageNumber="355" masterPageNumber="343" pageNumber="344" updateTime="1722933277969" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="5B1AE5D3D3AD6A61FD8EC69421729762" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0393E411868C0AD0E8BC5D02B6314055">
<mods:title id="2D404D811B0F3D44057A747CD5224DA7">Trechus species from Mt. Choke of northern Ethiopia related to T. niloticus (Quéinnec &amp; Ollivier) with notable male genital morphology (Carabidae: Trechini)</mods:title>
</mods:titleInfo>
<mods:name id="8B6FD463D238A7D74A9A82600526A826" type="personal">
<mods:role id="BB5EBE3D3B038A29BB5EF1D4CB73D5F1">
<mods:roleTerm id="AB203BF7A351D9FE437287E31648485D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9EE0F2C7C63A8338510AE39B752804B9">Schmidt, Joachim</mods:namePart>
<mods:affiliation id="AF28F213118E1C6CDEE6AEF6F49692AC">General and Systematic Zoology, University of Rostock, Germany</mods:affiliation>
</mods:name>
<mods:name id="9AFEEDBC76C18E7E6425248C71A2DC67" type="personal">
<mods:role id="90DA576E5D93C3172C1C4563BFBB55EB">
<mods:roleTerm id="21DADCF6F9216ACAECA57968098C5213">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EABAE4A7F233514C0D935C506BEFE52A">Merene, Yeshitla</mods:namePart>
<mods:affiliation id="9E62DC6213F78319F84727F5E828162D">Department of Zoological Sciences, Addis Ababa University, Ethiopia &amp; Amhara Agricultural Research Institute, Bahir Dar, Ethiopia</mods:affiliation>
</mods:name>
<mods:typeOfResource id="062592BA6A88BD4AF285847770F8691B">text</mods:typeOfResource>
<mods:relatedItem id="3BF65D4E0CDA3DA07C1385A80441FDE3" type="host">
<mods:titleInfo id="DB3482619DFB0DECE2C24491794798C9">
<mods:title id="24D137286BB9535FC4DBD311E9C1EA6C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="1BEE8776F328FF6E861E3EF4624956AA">
<mods:date id="703DA6B07D8E7D4F4FC7F8EDD4EA4B20">2024</mods:date>
<mods:detail id="A3B77D357AAD9C4110513B641ECCEEFF" type="pubDate">
<mods:number id="3980C09E377F042013DCFBF650399765">2024-08-06</mods:number>
</mods:detail>
<mods:detail id="E401888692D3F6F73A6F8A1C4C93F48D" type="volume">
<mods:number id="98941CC15B5DFF90858F3C6245E05D6E">5492</mods:number>
</mods:detail>
<mods:detail id="274AAD2F1528723DB2D075078E08F8A6" type="issue">
<mods:number id="75A5BE8DFE89524EDD59D615B5F4C2CF">3</mods:number>
</mods:detail>
<mods:extent id="C3555AFD4EF143F1AC8CB42CF9189B8A" unit="page">
<mods:start id="7049801CD43FAF7B6A4AB690823193EB">343</mods:start>
<mods:end id="93ABD7E24584204677721A532C28B217">355</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="538B84A8DC4036B6F9AA3D86006C3F2C">
<mods:url id="5561AC07079039FD4986EEDCDA3C7DD3">http://dx.doi.org/10.11646/zootaxa.5492.3.3</mods:url>
</mods:location>
<mods:classification id="CFA8777556E673C19C957D3044A45915">journal article</mods:classification>
<mods:identifier id="BD2074CF0C3304B4A300A9C7DF363444" type="DOI">10.11646/zootaxa.5492.3.3</mods:identifier>
<mods:identifier id="72A19C321F97CEBB54A16F5053B68626" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="6071C3D5E10D6647F5D806F32D4252A1" type="Zenodo-Dep">13234851</mods:identifier>
<mods:identifier id="1889F5C9CEA0B9E87082B3EDAFBAE79A" type="ZooBank">8F81F053-DE78-4684-BCC9-875AE0C41615</mods:identifier>
</mods:mods>
<treatment id="B36D0606FFD7AC1FFF28FC99FC78FF2D" ID-DOI="http://doi.org/10.5281/zenodo.13234859" ID-Zenodo-Dep="13234859" LSID="urn:lsid:plazi:treatment:B36D0606FFD7AC1FFF28FC99FC78FF2D" httpUri="http://treatment.plazi.org/id/B36D0606FFD7AC1FFF28FC99FC78FF2D" lastPageId="4" lastPageNumber="346" pageId="1" pageNumber="344">
<subSubSection id="73DEE49BFFD7AC1AFF28FC99FD04FCA5" box="[151,711,835,862]" pageId="1" pageNumber="344" type="nomenclature">
<paragraph id="3B7BB710FFD7AC1AFF28FC99FD04FCA5" blockId="1.[151,711,835,897]" box="[151,711,835,862]" pageId="1" pageNumber="344">
<heading id="6033007CFFD7AC1AFF28FC99FD04FCA5" bold="true" box="[151,711,835,862]" fontSize="11" level="1" pageId="1" pageNumber="344" reason="1">
<emphasis id="09B06B02FFD7AC1AFF28FC99FD04FCA5" bold="true" box="[151,711,835,862]" pageId="1" pageNumber="344">
<taxonomicName id="FCC4CC93FFD7AC1AFF28FC99FDBBFCA5" authority="Schmidt" authorityName="Schmidt &amp; Merene" authorityYear="2024" box="[151,632,835,862]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="344" phylum="Arthropoda" rank="species" species="apertus" status="sp. nov." subGenus="Abyssinotus">
<emphasis id="09B06B02FFD7AC1AFF28FC99FF3AFCA6" bold="true" box="[151,249,835,862]" italics="true" pageId="1" pageNumber="344">Trechus</emphasis>
(
<emphasis id="09B06B02FFD7AC1AFEB4FC99FE5FFCA6" bold="true" box="[267,412,835,862]" italics="true" pageId="1" pageNumber="344">Abyssinotus</emphasis>
)
<emphasis id="09B06B02FFD7AC1AFE12FC99FDC4FCA5" bold="true" box="[429,519,835,861]" italics="true" pageId="1" pageNumber="344">apertus</emphasis>
Schmidt
</taxonomicName>
,
<taxonomicNameLabel id="1283D679FFD7AC1AFD3AFC99FD04FCA5" box="[645,711,835,861]" pageId="1" pageNumber="344" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="73DEE49BFFD7AC1AFF28FCBDFF1AFC79" box="[151,217,871,897]" pageId="1" pageNumber="344" type="description">
<paragraph id="3B7BB710FFD7AC1AFF28FCBDFF1AFC79" blockId="1.[151,711,835,897]" box="[151,217,871,897]" pageId="1" pageNumber="344">
<figureCitation id="A3FFAB95FFD7AC1AFF28FCBDFF1AFC79" box="[151,217,871,897]" captionStart="FIGURE 1" captionStartId="2.[152,255,1856,1880]" captionTargetBox="[183,1401,183,1825]" captionTargetId="figure-23@2.[183,1404,179,1830]" captionTargetPageId="2" captionText="FIGURE 1. Trechus (Abyssinotus) apertus Schmidt, sp. n., holotype (Fig. 1A) and details of paratypes (Fig. 1B–H). A, habitus; the white circles point to the insertions of the supraorbital setae, pronotal lateral setae, parascutellar seta, elytral discal setae, and setae of the umbilicate series. B, right mandible, ventral aspect. C, left mandible, ventral aspect. D, labrum, dorsal aspect. E, left maxilla. F, left labial palpus and ligula, dorsal aspect; the arrows point to the three setae inserting on the penultimate palpomere. G, aedeagus, left lateral view. H, aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13234853" httpUri="https://zenodo.org/record/13234853/files/figure.png" pageId="1" pageNumber="344">Fig. 1</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="73DEE49BFFD7AC1AFF28FC75FC24FBCD" pageId="1" pageNumber="344" type="materials_examined">
<paragraph id="3B7BB710FFD7AC1AFF28FC75FC24FBCD" blockId="1.[151,1437,943,1834]" pageId="1" pageNumber="344">
<emphasis id="09B06B02FFD7AC1AFF28FC75FE81FC31" bold="true" box="[151,322,943,969]" pageId="1" pageNumber="344">Type material.</emphasis>
<materialsCitation id="8BACBD4DFFD7AC1AFEF9FC75FC3DFC15" collectingDate="2019-02-23" collectionCode="CSCHM" collectorName="D. Hauth &amp; Hauth, J &amp; Yeshitla M. &amp; Yitbarek, W." country="Ethiopia" elevation="3370" latitude="10.635278" location="Mt. Choke" longLatPrecision="21" longitude="37.764168" pageId="1" pageNumber="344" specimenCount="1" specimenCount-male="1" stateProvince="Amara" typeStatus="holotype">
<typeStatus id="E47F09B2FFD7AC1AFEF9FC75FE70FC31" box="[326,435,943,969]" pageId="1" pageNumber="344" type="holotype">Holotype</typeStatus>
:
<specimenCount id="2DC27C99FFD7AC1AFE04FC6AFE11FC32" box="[443,466,944,970]" count="1" pageId="1" pageNumber="344" type="male">♂</specimenCount>
, with label data:
<collectingCountry id="43D3F780FFD7AC1AFD39FC75FD2BFC31" box="[646,744,943,969]" name="Ethiopia" pageId="1" pageNumber="344">Ethiopia</collectingCountry>
,
<collectingRegion id="F90079F2FFD7AC1AFD50FC75FC8FFC31" box="[751,844,943,969]" country="Ethiopia" name="Amara" pageId="1" pageNumber="344">Amhara</collectingRegion>
, W-slope
<location id="3E1BE1CBFFD7AC1AFC0AFC6AFBEEFC31" LSID="urn:lsid:plazi:treatment:B36D0606FFD7AC1FFF28FC99FC78FF2D:3E1BE1CBFFD7AC1AFC0AFC6AFBEEFC31" box="[949,1069,943,970]" country="Ethiopia" latitude="10.635278" longLatPrecision="21" longitude="37.764168" name="Mt. Choke" pageId="1" pageNumber="344" stateProvince="Amara">Mt.Choke</location>
,
<quantity id="FC3C1AF5FFD7AC1AFB89FC75FB49FC32" box="[1078,1162,943,970]" metricMagnitude="3" metricUnit="m" metricValue="3.37" pageId="1" pageNumber="344" unit="m" value="3370.0">
<elevation id="B0E95023FFD7AC1AFB89FC75FB49FC32" box="[1078,1162,943,970]" metricMagnitude="3" metricUnit="m" metricValue="3.37" pageId="1" pageNumber="344" unit="m" value="3370.0">3370 m</elevation>
</quantity>
,
<date id="4F7A91D0FFD7AC1AFB2CFC75FAC8FC31" box="[1171,1291,943,969]" pageId="1" pageNumber="344" value="2019-02-23">
<collectingDate id="5F3E6838FFD7AC1AFB2CFC75FAC8FC31" box="[1171,1291,943,969]" pageId="1" pageNumber="344" value="2019-02-23">23.II.2019</collectingDate>
</date>
,
<geoCoordinate id="5EF0D1D7FFD7AC1AFAACFC75FA5FFC31" box="[1299,1436,943,969]" degrees="10" direction="north" minutes="38" orientation="latitude" pageId="1" pageNumber="344" precision="15" seconds="07" value="10.635278">10°38’07”N</geoCoordinate>
<geoCoordinate id="5EF0D1D7FFD7AC1AFF28FC09FEE3FC16" box="[151,288,979,1006]" degrees="37" direction="east" minutes="45" orientation="longitude" pageId="1" pageNumber="344" precision="15" seconds="51" value="37.764168">37°45’51”E</geoCoordinate>
, leg.
<collectorName id="9631D2C6FFD7AC1AFEE0FC0EFE09FC15" box="[351,458,979,1006]" pageId="1" pageNumber="344">D. Hauth</collectorName>
, J. Schmidt,
<collectorName id="9631D2C6FFD7AC1AFDE2FC09FD21FC16" box="[605,738,979,1006]" pageId="1" pageNumber="344">Yeshitla M.</collectorName>
,
<collectorName id="9631D2C6FFD7AC1AFD50FC09FCBDFC16" box="[751,894,979,1006]" pageId="1" pageNumber="344">Yitbarek, W.</collectorName>
(
<collectionCode id="5DD52FD5FFD7AC1AFC31FC09FC34FC15" box="[910,1015,979,1005]" pageId="1" pageNumber="344">CSCHM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="8BACBD4DFFD7AC1AFBB5FC0EFE0CFBE9" collectingDate="2019-02-23" collectionCode="CSCHM" collectorName="D. Hauth &amp; Hauth, J &amp; Yeshitla M. &amp; Yitbarek, W." country="Ethiopia" elevation="3370" latitude="10.635278" location="Mt. Choke" longLatPrecision="21" longitude="37.764168" pageId="1" pageNumber="344" specimenCount="15" specimenCount-female="7" specimenCount-male="8" stateProvince="Amara" typeStatus="paratype">
<typeStatus id="E47F09B2FFD7AC1AFBB5FC0EFBB8FC16" box="[1034,1147,980,1006]" pageId="1" pageNumber="344" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="2DC27C99FFD7AC1AFB37FC09FB09FC16" box="[1160,1226,979,1006]" count="8" pageId="1" pageNumber="344" type="male">8 ♂♂</specimenCount>
,
<specimenCount id="2DC27C99FFD7AC1AFB67FC0EFAD9FC16" box="[1240,1306,980,1006]" count="7" pageId="1" pageNumber="344" type="female">7 ♀♀</specimenCount>
, with same data as holotype (
<collectionCode id="5DD52FD5FFD7AC1AFEE0FC2DFE0BFBE9" box="[351,456,1015,1041]" pageId="1" pageNumber="344">CSCHM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="8BACBD4DFFD7AC1AFE64FC2DFC27FBCD" collectingDate="2019-02-22" collectionCode="NHMAA, CAF, CSCHM" collectorName="D. Hauth &amp; Hauth, J &amp; Yeshitla M. &amp; Yitbarek, W." country="Ethiopia" elevation="3450" latitude="10.635833" location="Mt. Choke" longLatPrecision="21" longitude="37.768333" pageId="1" pageNumber="344" specimenCount="9" specimenCount-female="5" specimenCount-male="4" stateProvince="Amara" typeStatus="paratype">
<specimenCount id="2DC27C99FFD7AC1AFE64FC2DFDDFFBEA" box="[475,540,1015,1042]" count="4" pageId="1" pageNumber="344" type="male">4 ♂♂</specimenCount>
,
<specimenCount id="2DC27C99FFD7AC1AFD96FC2DFDA9FBEA" box="[553,618,1015,1042]" count="5" pageId="1" pageNumber="344" type="female">5 ♀♀</specimenCount>
, W-slope
<location id="3E1BE1CBFFD7AC1AFD62FC22FC9AFBE9" LSID="urn:lsid:plazi:treatment:B36D0606FFD7AC1FFF28FC99FC78FF2D:3E1BE1CBFFD7AC1AFD62FC22FC9AFBE9" box="[733,857,1015,1042]" country="Ethiopia" latitude="10.635833" longLatPrecision="21" longitude="37.768333" name="Mt. Choke" pageId="1" pageNumber="344" stateProvince="Amara">Mt. Choke</location>
,
<quantity id="FC3C1AF5FFD7AC1AFCDAFC2DFC7EFBEA" box="[869,957,1015,1042]" metricMagnitude="3" metricUnit="m" metricValue="3.45" pageId="1" pageNumber="344" unit="m" value="3450.0">
<elevation id="B0E95023FFD7AC1AFCDAFC2DFC7EFBEA" box="[869,957,1015,1042]" metricMagnitude="3" metricUnit="m" metricValue="3.45" pageId="1" pageNumber="344" unit="m" value="3450.0">3450 m</elevation>
</quantity>
,
<date id="4F7A91D0FFD7AC1AFC76FC2DFB82FBE9" box="[969,1089,1015,1041]" pageId="1" pageNumber="344" value="2019-02-22">
<collectingDate id="5F3E6838FFD7AC1AFC76FC2DFB82FBE9" box="[969,1089,1015,1041]" pageId="1" pageNumber="344" value="2019-02-22">22.II.2019</collectingDate>
</date>
,
<geoCoordinate id="5EF0D1D7FFD7AC1AFBF2FC2DFB15FBE9" box="[1101,1238,1015,1041]" degrees="10" direction="north" minutes="38" orientation="latitude" pageId="1" pageNumber="344" precision="15" seconds="09" value="10.635833">10°38’09”N</geoCoordinate>
<geoCoordinate id="5EF0D1D7FFD7AC1AFB61FC2DFAA5FBE9" box="[1246,1382,1015,1041]" degrees="37" direction="east" minutes="46" orientation="longitude" pageId="1" pageNumber="344" precision="15" seconds="06" value="37.768333">37°46’06”E</geoCoordinate>
, leg.
<collectorName id="9631D2C6FFD7AC1AFF28FBC6FEC3FBCD" box="[151,256,1051,1078]" pageId="1" pageNumber="344">D. Hauth</collectorName>
, J. Schmidt,
<collectorName id="9631D2C6FFD7AC1AFE30FBC1FDD0FBCE" box="[399,531,1051,1078]" pageId="1" pageNumber="344">Yeshitla M.</collectorName>
,
<collectorName id="9631D2C6FFD7AC1AFDA2FBC1FD68FBCE" box="[541,683,1051,1078]" pageId="1" pageNumber="344">Yitbarek, W.</collectorName>
(
<collectionCode id="5DD52FD5FFD7AC1AFD06FBC1FCE4FBCD" box="[697,807,1051,1077]" pageId="1" pageNumber="344">NHMAA</collectionCode>
,
<collectionCode id="5DD52FD5FFD7AC1AFC8DFBC1FCAAFBCD" box="[818,873,1051,1077]" country="China" lsid="urn:lsid:biocol.org:col:12802" name="Chinese Academy of Forestry" pageId="1" pageNumber="344" type="Herbarium">CAF</collectionCode>
,
<collectionCode id="5DD52FD5FFD7AC1AFCCBFBC1FC18FBCD" box="[884,987,1051,1077]" pageId="1" pageNumber="344">CSCHM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="73DEE49BFFD7AC1AFF78FBE5FCCDFB85" pageId="1" pageNumber="344" type="etymology">
<paragraph id="3B7BB710FFD7AC1AFF78FBE5FCCDFB85" blockId="1.[151,1437,943,1834]" pageId="1" pageNumber="344">
<emphasis id="09B06B02FFD7AC1AFF78FBE5FE8BFBA1" bold="true" box="[199,328,1087,1113]" pageId="1" pageNumber="344">Etymology</emphasis>
. The species’ epithet refers to the extraordinary wide dorsal opening of the aedeagal median lobe which is a morphological peculiarity of the new species.
</paragraph>
</subSubSection>
<subSubSection id="73DEE49BFFD7AC18FF78FB5DFAB7FB3D" lastPageId="3" lastPageNumber="346" pageId="1" pageNumber="344" type="description">
<paragraph id="3B7BB710FFD7AC1AFF78FB5DFD45FB5A" blockId="1.[151,1437,943,1834]" box="[199,646,1159,1186]" pageId="1" pageNumber="344">
<emphasis id="09B06B02FFD7AC1AFF78FB5DFE92FB59" bold="true" box="[199,337,1159,1185]" pageId="1" pageNumber="344">Description</emphasis>
. Body length:
<quantity id="FC3C1AF5FFD7AC1AFE47FB5DFD41FB5A" box="[504,642,1159,1186]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="1" pageNumber="344" unit="mm" value="2.65" valueMax="2.8" valueMin="2.5">2.5–2.8 mm</quantity>
.
</paragraph>
<paragraph id="3B7BB710FFD7AC1AFF78FB71FA4CFB11" blockId="1.[151,1437,943,1834]" pageId="1" pageNumber="344">Proportions (n = 12): PW/HW = 1.35–1.44 (Ø = 1.41); PW/PL = 1.37–1.45 (Ø = 1.40); PW/PBW = 1.18–1.22 (Ø = 1.20); PBW/PAW = 1.18–1.24 (Ø = 1.21); EW/PW = 1.38–1.46 (Ø = 1.43); EL/EW = 1.24–1.30 (Ø = 1.27).</paragraph>
<paragraph id="3B7BB710FFD7AC1AFF78FB29FC4EFAAD" blockId="1.[151,1437,943,1834]" pageId="1" pageNumber="344">
Colour: Head, pronotum and elytra blackish brown; mandibles, palps, antennal base and legs yellowish brown; antennae slightly darkened beginning from second antennomere; in most specimens 2
<superScript id="CCB11A58FFD7AC1AFB2EFACDFB62FADD" attach="left" box="[1169,1185,1303,1317]" fontSize="6" pageId="1" pageNumber="344">nd</superScript>
maxillary palpomere somewhat darkened; elytral apical margin very narrowly yellowish.
</paragraph>
<paragraph id="3B7BB710FFD7AC1AFF78FA85FE5CFA1D" blockId="1.[151,1437,943,1834]" pageId="1" pageNumber="344">Microsculpture: Sculpticells on frons and supraorbital area large, moderately impressed, almost isodiametric; sculpticells on clypeus very finely impressed, slightly transverse. Sculpticells on pronotal disc smaller and finer impressed than on head disc, slightly transverse. Elytra with large and deeply impressed, almost isodiametric sculpticells throughout.</paragraph>
<paragraph id="3B7BB710FFD7AC1AFF78FA35FDD3F8D1" blockId="1.[151,1437,943,1834]" pageId="1" pageNumber="344">
Head: Moderately robust. Mandibles short, with right mandible tridentate where all teeth are almost the same length, without diastema, without recognizable separation of premolar and retinaculum (
<figureCitation id="A3FFAB95FFD7AC1AFB6FF9C9FAEAF9D5" box="[1232,1321,1555,1581]" captionStart="FIGURE 1" captionStartId="2.[152,255,1856,1880]" captionTargetBox="[183,1401,183,1825]" captionTargetId="figure-23@2.[183,1404,179,1830]" captionTargetPageId="2" captionText="FIGURE 1. Trechus (Abyssinotus) apertus Schmidt, sp. n., holotype (Fig. 1A) and details of paratypes (Fig. 1B–H). A, habitus; the white circles point to the insertions of the supraorbital setae, pronotal lateral setae, parascutellar seta, elytral discal setae, and setae of the umbilicate series. B, right mandible, ventral aspect. C, left mandible, ventral aspect. D, labrum, dorsal aspect. E, left maxilla. F, left labial palpus and ligula, dorsal aspect; the arrows point to the three setae inserting on the penultimate palpomere. G, aedeagus, left lateral view. H, aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13234853" httpUri="https://zenodo.org/record/13234853/files/figure.png" pageId="1" pageNumber="344">Fig. 1B</figureCitation>
); tooth of left mandible with two blunt cusps (
<figureCitation id="A3FFAB95FFD7AC1AFD80F9EDFD5AF9A9" box="[575,665,1591,1617]" captionStart="FIGURE 1" captionStartId="2.[152,255,1856,1880]" captionTargetBox="[183,1401,183,1825]" captionTargetId="figure-23@2.[183,1404,179,1830]" captionTargetPageId="2" captionText="FIGURE 1. Trechus (Abyssinotus) apertus Schmidt, sp. n., holotype (Fig. 1A) and details of paratypes (Fig. 1B–H). A, habitus; the white circles point to the insertions of the supraorbital setae, pronotal lateral setae, parascutellar seta, elytral discal setae, and setae of the umbilicate series. B, right mandible, ventral aspect. C, left mandible, ventral aspect. D, labrum, dorsal aspect. E, left maxilla. F, left labial palpus and ligula, dorsal aspect; the arrows point to the three setae inserting on the penultimate palpomere. G, aedeagus, left lateral view. H, aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13234853" httpUri="https://zenodo.org/record/13234853/files/figure.png" pageId="1" pageNumber="344">Fig. 1C</figureCitation>
). Penultimate labial palpomere with three setae (
<figureCitation id="A3FFAB95FFD7AC1AFB6BF9EDFAE8F9A9" box="[1236,1323,1591,1617]" captionStart="FIGURE 1" captionStartId="2.[152,255,1856,1880]" captionTargetBox="[183,1401,183,1825]" captionTargetId="figure-23@2.[183,1404,179,1830]" captionTargetPageId="2" captionText="FIGURE 1. Trechus (Abyssinotus) apertus Schmidt, sp. n., holotype (Fig. 1A) and details of paratypes (Fig. 1B–H). A, habitus; the white circles point to the insertions of the supraorbital setae, pronotal lateral setae, parascutellar seta, elytral discal setae, and setae of the umbilicate series. B, right mandible, ventral aspect. C, left mandible, ventral aspect. D, labrum, dorsal aspect. E, left maxilla. F, left labial palpus and ligula, dorsal aspect; the arrows point to the three setae inserting on the penultimate palpomere. G, aedeagus, left lateral view. H, aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13234853" httpUri="https://zenodo.org/record/13234853/files/figure.png" pageId="1" pageNumber="344">Fig. 1F</figureCitation>
). Labrum with apical margin moderately emarginate, with six setae near apical margin (
<figureCitation id="A3FFAB95FFD7AC1AFBA2F981FBBAF98D" box="[1053,1145,1627,1653]" captionStart="FIGURE 1" captionStartId="2.[152,255,1856,1880]" captionTargetBox="[183,1401,183,1825]" captionTargetId="figure-23@2.[183,1404,179,1830]" captionTargetPageId="2" captionText="FIGURE 1. Trechus (Abyssinotus) apertus Schmidt, sp. n., holotype (Fig. 1A) and details of paratypes (Fig. 1B–H). A, habitus; the white circles point to the insertions of the supraorbital setae, pronotal lateral setae, parascutellar seta, elytral discal setae, and setae of the umbilicate series. B, right mandible, ventral aspect. C, left mandible, ventral aspect. D, labrum, dorsal aspect. E, left maxilla. F, left labial palpus and ligula, dorsal aspect; the arrows point to the three setae inserting on the penultimate palpomere. G, aedeagus, left lateral view. H, aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13234853" httpUri="https://zenodo.org/record/13234853/files/figure.png" pageId="1" pageNumber="344">Fig. 1D</figureCitation>
). Clypeus each side with two setae. Compound eyes rather small, slightly convex; tempora about ¾ of length of eyes, markedly convex, distinctly wrinkled to the neck. Dorsal surface of head with supraorbital area and frons markedly convex, and with supraorbital furrows deeply impressed, evenly curved. Two supraorbital setae and one suborbital seta each side. Tempora distinctly pubescent. Antennae short, with second antennomere about as long as scape, and with third antennomere shorter than second.
</paragraph>
<caption id="6FBBE798FFD4AC19FF27F89AFD13F810" ID-DOI="http://doi.org/10.5281/zenodo.13234853" ID-Zenodo-Dep="13234853" httpUri="https://zenodo.org/record/13234853/files/figure.png" pageId="2" pageNumber="345" startId="2.[152,255,1856,1880]" targetBox="[183,1401,183,1825]" targetPageId="2" targetType="figure">
<paragraph id="3B7BB710FFD4AC19FF27F89AFD13F810" blockId="2.[151,1437,1856,2024]" pageId="2" pageNumber="345">
<emphasis id="09B06B02FFD4AC19FF27F89AFEDBF8A0" bold="true" box="[152,280,1856,1880]" pageId="2" pageNumber="345">FIGURE 1.</emphasis>
<taxonomicName id="FCC4CC93FFD4AC19FEA2F89AFD6AF8A0" authority="Schmidt" authorityName="Schmidt &amp; Merene" authorityYear="2024" box="[285,681,1856,1880]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="345" phylum="Arthropoda" rank="species" species="apertus" status="sp. nov." subGenus="Abyssinotus">
<emphasis id="09B06B02FFD4AC19FEA2F89AFEAFF8A0" box="[285,364,1856,1880]" italics="true" pageId="2" pageNumber="345">Trechus</emphasis>
(
<emphasis id="09B06B02FFD4AC19FEC5F89AFE30F8A0" box="[378,499,1856,1880]" italics="true" pageId="2" pageNumber="345">Abyssinotus</emphasis>
)
<emphasis id="09B06B02FFD4AC19FDBEF89BFD8EF8A0" box="[513,589,1857,1880]" italics="true" pageId="2" pageNumber="345">apertus</emphasis>
Schmidt
</taxonomicName>
,
<taxonomicNameLabel id="1283D679FFD4AC19FD0CF89BFD2AF8A0" box="[691,745,1857,1880]" pageId="2" pageNumber="345" rank="species">sp. n.</taxonomicNameLabel>
, holotype (Fig. 1A) and details of paratypes (Fig. 1B–H).
<emphasis id="09B06B02FFD4AC19FA91F89AFA83F8A0" bold="true" box="[1326,1344,1856,1880]" pageId="2" pageNumber="345">A</emphasis>
, habitus; the white circles point to the insertions of the supraorbital setae, pronotal lateral setae, parascutellar seta, elytral discal setae, and setae of the umbilicate series.
<emphasis id="09B06B02FFD4AC19FE7BF852FE16F858" bold="true" box="[452,469,1928,1952]" pageId="2" pageNumber="345">B</emphasis>
, right mandible, ventral aspect.
<emphasis id="09B06B02FFD4AC19FCA8F852FCEAF858" bold="true" box="[791,809,1928,1952]" pageId="2" pageNumber="345">C</emphasis>
, left mandible, ventral aspect.
<emphasis id="09B06B02FFD4AC19FBE1F852FBB3F858" bold="true" box="[1118,1136,1928,1952]" pageId="2" pageNumber="345">D</emphasis>
, labrum, dorsal aspect.
<emphasis id="09B06B02FFD4AC19FAE1F852FAACF858" bold="true" box="[1374,1391,1928,1952]" pageId="2" pageNumber="345">E</emphasis>
, left maxilla.
<emphasis id="09B06B02FFD4AC19FF50F876FF3DF83C" bold="true" box="[239,254,1964,1988]" pageId="2" pageNumber="345">F</emphasis>
, left labial palpus and ligula, dorsal aspect; the arrows point to the three setae inserting on the penultimate palpomere.
<emphasis id="09B06B02FFD4AC19FF28F80AFF68F810" bold="true" box="[151,171,2000,2024]" pageId="2" pageNumber="345">G</emphasis>
, aedeagus, left lateral view.
<emphasis id="09B06B02FFD4AC19FE74F80AFE1CF810" bold="true" box="[459,479,2000,2024]" pageId="2" pageNumber="345">H</emphasis>
, aedeagus, dorsal view.
</paragraph>
</caption>
<paragraph id="3B7BB710FFD5AC18FF78FF4DFD45FE55" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="346">
Pronotum: Large and transverse, sub-discoidal, broadest slightly before middle, with base distinctly broader than apex; disc moderately convex. Anterior margin slightly concave (almost straight in middle), with anterior angles slightly protruded anteriorly, rounded; basal margin convex, with laterobasal angles markedly shifted anteriorly; lateral margin almost evenly rounded throughout; laterobasal angles very obtuse, indistinct (
<figureCitation id="A3FFAB95FFD5AC18FB7DFED9FADDFEE5" box="[1218,1310,259,285]" captionStart="FIGURE 1" captionStartId="2.[152,255,1856,1880]" captionTargetBox="[183,1401,183,1825]" captionTargetId="figure-23@2.[183,1404,179,1830]" captionTargetPageId="2" captionText="FIGURE 1. Trechus (Abyssinotus) apertus Schmidt, sp. n., holotype (Fig. 1A) and details of paratypes (Fig. 1B–H). A, habitus; the white circles point to the insertions of the supraorbital setae, pronotal lateral setae, parascutellar seta, elytral discal setae, and setae of the umbilicate series. B, right mandible, ventral aspect. C, left mandible, ventral aspect. D, labrum, dorsal aspect. E, left maxilla. F, left labial palpus and ligula, dorsal aspect; the arrows point to the three setae inserting on the penultimate palpomere. G, aedeagus, left lateral view. H, aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13234853" httpUri="https://zenodo.org/record/13234853/files/figure.png" pageId="3" pageNumber="346">Fig. 1A</figureCitation>
). Marginal gutter narrow, slightly widened near laterobasal angle, extending towards laterobasal foveae. Median longitudinal impression very finely incised, disappearing near apex; anterior and posterior transverse impressions indistinct, smooth; laterobasal foveae small, moderately deep, smooth. Lateral and laterobasal setae present, with the former situated at the end of anterior pronotal third.
</paragraph>
<paragraph id="3B7BB710FFD5AC18FF78FE6DFD1EFE29" blockId="3.[151,1437,151,2013]" box="[199,733,439,465]" pageId="3" pageNumber="346">Pro- and mesepisternum: Glabrous and smooth.</paragraph>
<paragraph id="3B7BB710FFD5AC18FF78FE01FCF5FCC1" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="346">
Pterothorax: Elytra short, ovate, with disc convex, in dorsal view broadest in middle, humerus rounded but distinct, subapical sinuation indistinct, apex rounded. Striae 1–3 and anterior portion of stria 8 slightly impressed, punctate, external striae finer, 6–7 sometimes indistinct, stria 8 distinctly deepened before apex; parascutellar stria very short, deep, isolated. Recurrent preapical stria short, deep, with outer curvature directed towards the fifth stria. Parascutellar seta present; three discal setae located in the 3rd interval, adjoined to the 3rd stria (
<figureCitation id="A3FFAB95FFD5AC18FB6AFDB1FAECFD7D" box="[1237,1327,619,645]" captionStart="FIGURE 1" captionStartId="2.[152,255,1856,1880]" captionTargetBox="[183,1401,183,1825]" captionTargetId="figure-23@2.[183,1404,179,1830]" captionTargetPageId="2" captionText="FIGURE 1. Trechus (Abyssinotus) apertus Schmidt, sp. n., holotype (Fig. 1A) and details of paratypes (Fig. 1B–H). A, habitus; the white circles point to the insertions of the supraorbital setae, pronotal lateral setae, parascutellar seta, elytral discal setae, and setae of the umbilicate series. B, right mandible, ventral aspect. C, left mandible, ventral aspect. D, labrum, dorsal aspect. E, left maxilla. F, left labial palpus and ligula, dorsal aspect; the arrows point to the three setae inserting on the penultimate palpomere. G, aedeagus, left lateral view. H, aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13234853" httpUri="https://zenodo.org/record/13234853/files/figure.png" pageId="3" pageNumber="346">Fig. 1A</figureCitation>
): anterior seta located near the end of the anterior elytral 7th; middle seta located anterior of elytral middle, near the end of elytral 3/7; posterior seta (= subapical seta) located about 1/8 of elytral length from elytral apex; seta of the recurrent stria isolated, removed from this stria by distance of the diameter of the pore (
<figureCitation id="A3FFAB95FFD5AC18FBADFD0DFBAFFD09" box="[1042,1132,727,753]" captionStart="FIGURE 1" captionStartId="2.[152,255,1856,1880]" captionTargetBox="[183,1401,183,1825]" captionTargetId="figure-23@2.[183,1404,179,1830]" captionTargetPageId="2" captionText="FIGURE 1. Trechus (Abyssinotus) apertus Schmidt, sp. n., holotype (Fig. 1A) and details of paratypes (Fig. 1B–H). A, habitus; the white circles point to the insertions of the supraorbital setae, pronotal lateral setae, parascutellar seta, elytral discal setae, and setae of the umbilicate series. B, right mandible, ventral aspect. C, left mandible, ventral aspect. D, labrum, dorsal aspect. E, left maxilla. F, left labial palpus and ligula, dorsal aspect; the arrows point to the three setae inserting on the penultimate palpomere. G, aedeagus, left lateral view. H, aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13234853" httpUri="https://zenodo.org/record/13234853/files/figure.png" pageId="3" pageNumber="346">Fig. 1A</figureCitation>
). Number and positions of the setae of the marginal umbilicate series typical of
<taxonomicName id="FCC4CC93FFD5AC18FD5FFD21FCFBFCED" authorityName="Clairville" authorityYear="1806" box="[736,824,763,789]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="346" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="09B06B02FFD5AC18FD5FFD21FCFBFCED" box="[736,824,763,789]" italics="true" pageId="3" pageNumber="346">Trechus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1283D679FFD5AC18FC81FD21FCA6FCED" box="[830,869,763,789]" pageId="3" pageNumber="346" sensu="lato">s. l.</taxonomicNameLabel>
(
<figureCitation id="A3FFAB95FFD5AC18FCCCFD21FC0FFCED" box="[883,972,763,789]" captionStart="FIGURE 1" captionStartId="2.[152,255,1856,1880]" captionTargetBox="[183,1401,183,1825]" captionTargetId="figure-23@2.[183,1404,179,1830]" captionTargetPageId="2" captionText="FIGURE 1. Trechus (Abyssinotus) apertus Schmidt, sp. n., holotype (Fig. 1A) and details of paratypes (Fig. 1B–H). A, habitus; the white circles point to the insertions of the supraorbital setae, pronotal lateral setae, parascutellar seta, elytral discal setae, and setae of the umbilicate series. B, right mandible, ventral aspect. C, left mandible, ventral aspect. D, labrum, dorsal aspect. E, left maxilla. F, left labial palpus and ligula, dorsal aspect; the arrows point to the three setae inserting on the penultimate palpomere. G, aedeagus, left lateral view. H, aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13234853" httpUri="https://zenodo.org/record/13234853/files/figure.png" pageId="3" pageNumber="346">Fig. 1A</figureCitation>
). Metepisternum very short, glabrous and smooth, with outer margin about as long as anterior margin.
</paragraph>
<paragraph id="3B7BB710FFD5AC18FF78FC99FD6DFC5D" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="346">
Legs: Moderately short and robust. Protibia dilated towards apex, straight on external margin, its dorsal surface with very flat longitudinal groove without micro-setae on anterior surface. Two basal protarsomeres of males dilated and dentoid at the inner apical border (
<figureCitation id="A3FFAB95FFD5AC18FDF6FC51FD63FC5D" box="[585,672,907,933]" captionStart="FIGURE 1" captionStartId="2.[152,255,1856,1880]" captionTargetBox="[183,1401,183,1825]" captionTargetId="figure-23@2.[183,1404,179,1830]" captionTargetPageId="2" captionText="FIGURE 1. Trechus (Abyssinotus) apertus Schmidt, sp. n., holotype (Fig. 1A) and details of paratypes (Fig. 1B–H). A, habitus; the white circles point to the insertions of the supraorbital setae, pronotal lateral setae, parascutellar seta, elytral discal setae, and setae of the umbilicate series. B, right mandible, ventral aspect. C, left mandible, ventral aspect. D, labrum, dorsal aspect. E, left maxilla. F, left labial palpus and ligula, dorsal aspect; the arrows point to the three setae inserting on the penultimate palpomere. G, aedeagus, left lateral view. H, aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13234853" httpUri="https://zenodo.org/record/13234853/files/figure.png" pageId="3" pageNumber="346">Fig. 1C</figureCitation>
).
</paragraph>
<paragraph id="3B7BB710FFD5AC18FF78FC75FAB7FB3D" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="346">
Male genitalia: EL/AL= 2.00–2.23 (Ø = 2.11; n = 8). Aedeagal median lobe robust, in lateral view markedly bent in basal third, almost straight towards apex, with ventral margin slightly convex in middle, ventrally bent towards apex, the latter with a small upwardly bent hook (
<figureCitation id="A3FFAB95FFD5AC18FD7DFC2DFCD8FBE9" box="[706,795,1015,1041]" captionStart="FIGURE 1" captionStartId="2.[152,255,1856,1880]" captionTargetBox="[183,1401,183,1825]" captionTargetId="figure-23@2.[183,1404,179,1830]" captionTargetPageId="2" captionText="FIGURE 1. Trechus (Abyssinotus) apertus Schmidt, sp. n., holotype (Fig. 1A) and details of paratypes (Fig. 1B–H). A, habitus; the white circles point to the insertions of the supraorbital setae, pronotal lateral setae, parascutellar seta, elytral discal setae, and setae of the umbilicate series. B, right mandible, ventral aspect. C, left mandible, ventral aspect. D, labrum, dorsal aspect. E, left maxilla. F, left labial palpus and ligula, dorsal aspect; the arrows point to the three setae inserting on the penultimate palpomere. G, aedeagus, left lateral view. H, aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13234853" httpUri="https://zenodo.org/record/13234853/files/figure.png" pageId="3" pageNumber="346">Fig. 1G</figureCitation>
); in dorsal view broad in basal half, sinusoidal, narrowed from middle to apex (
<figureCitation id="A3FFAB95FFD5AC18FE2DFBC1FE29FBCD" box="[402,490,1051,1077]" captionStart="FIGURE 1" captionStartId="2.[152,255,1856,1880]" captionTargetBox="[183,1401,183,1825]" captionTargetId="figure-23@2.[183,1404,179,1830]" captionTargetPageId="2" captionText="FIGURE 1. Trechus (Abyssinotus) apertus Schmidt, sp. n., holotype (Fig. 1A) and details of paratypes (Fig. 1B–H). A, habitus; the white circles point to the insertions of the supraorbital setae, pronotal lateral setae, parascutellar seta, elytral discal setae, and setae of the umbilicate series. B, right mandible, ventral aspect. C, left mandible, ventral aspect. D, labrum, dorsal aspect. E, left maxilla. F, left labial palpus and ligula, dorsal aspect; the arrows point to the three setae inserting on the penultimate palpomere. G, aedeagus, left lateral view. H, aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13234853" httpUri="https://zenodo.org/record/13234853/files/figure.png" pageId="3" pageNumber="346">Fig. 1H</figureCitation>
). Dorsal surface of median lobe not sclerotized apart from the basal quarter of the basal bulb near the basal opening which is slightly sclerotized; sagittal aileron absent. Endophallus symmetrical in dorsal view, without distinct copulatory pieces, densely covered by rows of tiny scales along dorsal side of the inactive internal sac from its base to apex, and with a complex folding structure and large sclerotized scales near base of internal sac. Parameres similar in size and shape, moderately short, each with three or four apical setae.
</paragraph>
</subSubSection>
<subSubSection id="73DEE49BFFD5AC18FF78FB14FB55F8FD" pageId="3" pageNumber="346" type="diagnosis">
<paragraph id="3B7BB710FFD5AC18FF78FB14FB55F8FD" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="346">
<emphasis id="09B06B02FFD5AC18FF78FB14FE0AFB10" bold="true" box="[199,457,1230,1256]" pageId="3" pageNumber="346">Differential diagnosis</emphasis>
.
<taxonomicName id="FCC4CC93FFD5AC18FE64FB15FD52FB10" authorityName="Schmidt &amp; Merene" authorityYear="2024" box="[475,657,1231,1257]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="346" phylum="Arthropoda" rank="species" species="apertus" status="sp. nov.">
<emphasis id="09B06B02FFD5AC18FE64FB15FD52FB10" box="[475,657,1231,1257]" italics="true" pageId="3" pageNumber="346">Trechus apertus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1283D679FFD5AC18FD24FB15FD19FB11" box="[667,730,1231,1257]" pageId="3" pageNumber="346" rank="species">sp. n.</taxonomicNameLabel>
is distinguished from all previously described
<taxonomicName id="FCC4CC93FFD5AC18FB42FB15FA96FB11" authorityName="Clairville" authorityYear="1806" box="[1277,1365,1231,1257]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="346" phylum="Arthropoda" rank="genus">
<emphasis id="09B06B02FFD5AC18FB42FB15FA96FB11" box="[1277,1365,1231,1257]" italics="true" pageId="3" pageNumber="346">Trechus</emphasis>
</taxonomicName>
(s. l.) species of
<collectingCountry id="43D3F780FFD5AC18FEAEFB29FEB0FAF5" box="[273,371,1267,1293]" name="Ethiopia" pageId="3" pageNumber="346">Ethiopia</collectingCountry>
, by the aedeagal median lobe ostium which extends basally across three quarters of the basal bulb. Not any Ethiopian Trechina species was known so far that has a dorsal opening that includes parts of the basal bulb. In external characters, the new species differs additionally from all the small to medium-sized trechine species from Mt. Choke as follows: from
<taxonomicName id="FCC4CC93FFD5AC18FDAFFA85FC32FA81" authority="(Queinnec &amp; Ollivier)" baseAuthorityName="Queinnec &amp; Ollivier" box="[528,1009,1375,1401]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="346" phylum="Arthropoda" rank="species" species="abyssinicus">
<emphasis id="09B06B02FFD5AC18FDAFFA85FD2CFA81" box="[528,751,1375,1401]" italics="true" pageId="3" pageNumber="346">Trechus abyssinicus</emphasis>
(Quéinnec &amp; Ollivier)
</taxonomicName>
,
<taxonomicName id="FCC4CC93FFD5AC18FC44FA85FA5BFA81" authority="(Queinnec &amp; Ollivier)" baseAuthorityName="Queinnec &amp; Ollivier" box="[1019,1432,1375,1401]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="346" phylum="Arthropoda" rank="species" species="afroalpinus">
<emphasis id="09B06B02FFD5AC18FC44FA85FB55FA81" box="[1019,1174,1375,1401]" italics="true" pageId="3" pageNumber="346">T. afroalpinus</emphasis>
(Quéinnec &amp; Ollivier)
</taxonomicName>
,
<taxonomicName id="FCC4CC93FFD5AC18FF28FA59FE30FA65" authority="Ortuno &amp; Novoa" authorityName="Ortuno &amp; Novoa" authorityYear="2011" box="[151,499,1411,1437]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="346" phylum="Arthropoda" rank="species" species="amharicus">
<emphasis id="09B06B02FFD5AC18FF28FA59FEE9FA65" box="[151,298,1411,1437]" italics="true" pageId="3" pageNumber="346">T. amharicus</emphasis>
Ortuño &amp; Novoa
</taxonomicName>
and
<taxonomicName id="FCC4CC93FFD5AC18FD93FA59FC64FA65" authority="(Queinnec &amp; Ollivier)" baseAuthorityName="Queinnec &amp; Ollivier" box="[556,935,1411,1437]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="346" phylum="Arthropoda" rank="species" species="lobeliae">
<emphasis id="09B06B02FFD5AC18FD93FA59FD60FA65" box="[556,675,1411,1437]" italics="true" pageId="3" pageNumber="346">T. lobeliae</emphasis>
(Quéinnec &amp; Ollivier)
</taxonomicName>
in two basal protarsomeres dilated in males (one in all the latter species), from
<taxonomicName id="FCC4CC93FFD5AC18FD9FFA7DFD71FA39" authorityName="Ortuno &amp; Novoa" authorityYear="2011" box="[544,690,1447,1473]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="346" phylum="Arthropoda" rank="species" species="amharicus">
<emphasis id="09B06B02FFD5AC18FD9FFA7DFD71FA39" box="[544,690,1447,1473]" italics="true" pageId="3" pageNumber="346">T. amharicus</emphasis>
</taxonomicName>
additionally in the presence of the middle seta of the elytral discal series (absent in
<taxonomicName id="FCC4CC93FFD5AC18FEE6FA11FE2FFA1D" authorityName="Ortuno &amp; Novoa" authorityYear="2011" box="[345,492,1483,1509]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="346" phylum="Arthropoda" rank="species" species="amharicus">
<emphasis id="09B06B02FFD5AC18FEE6FA11FE2FFA1D" box="[345,492,1483,1509]" italics="true" pageId="3" pageNumber="346">T. amharicus</emphasis>
</taxonomicName>
), from
<taxonomicName id="FCC4CC93FFD5AC18FDFAFA11FD7EFA1D" box="[581,701,1483,1509]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="346" phylum="Arthropoda" rank="species" species="lobeliae">
<emphasis id="09B06B02FFD5AC18FDFAFA11FD7EFA1D" box="[581,701,1483,1509]" italics="true" pageId="3" pageNumber="346">T. lobeliae</emphasis>
</taxonomicName>
additionally in the smaller body size and the almost completely rounded pronotal laterobasal angles (rectangular in
<taxonomicName id="FCC4CC93FFD5AC18FD63FA35FC85F9F1" box="[732,838,1519,1545]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="346" phylum="Arthropoda" rank="species" species="lobelia">
<emphasis id="09B06B02FFD5AC18FD63FA35FC85F9F1" box="[732,838,1519,1545]" italics="true" pageId="3" pageNumber="346">T. lobelia</emphasis>
</taxonomicName>
), and from
<taxonomicName id="FCC4CC93FFD5AC18FC73FA35FBAAF9F1" box="[972,1129,1519,1545]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="346" phylum="Arthropoda" rank="species" species="abyssinicus">
<emphasis id="09B06B02FFD5AC18FC73FA35FBAAF9F1" box="[972,1129,1519,1545]" italics="true" pageId="3" pageNumber="346">T. abyssinicus</emphasis>
</taxonomicName>
,
<taxonomicName id="FCC4CC93FFD5AC18FBC8FA35FAD0F9F1" box="[1143,1299,1519,1545]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="346" phylum="Arthropoda" rank="species" species="afroalpinus">
<emphasis id="09B06B02FFD5AC18FBC8FA35FAD0F9F1" box="[1143,1299,1519,1545]" italics="true" pageId="3" pageNumber="346">T. afroalpinus</emphasis>
</taxonomicName>
,
<taxonomicName id="FCC4CC93FFD5AC18FA9DFA35FAB7F9F0" authorityName="Schmidt &amp; Merene" authorityYear="2024" box="[1314,1396,1519,1544]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="346" phylum="Arthropoda" rank="species" species="igori" status="sp. n.">
<emphasis id="09B06B02FFD5AC18FA9DFA35FAB7F9F0" box="[1314,1396,1519,1544]" italics="true" pageId="3" pageNumber="346">T. igori</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1283D679FFD5AC18FAC3FA35FF6EF9D5" pageId="3" pageNumber="346" rank="species">sp. n.</taxonomicNameLabel>
,
<taxonomicName id="FCC4CC93FFD5AC18FF03F9C9FE87F9D5" authorityName="Schmidt &amp; Merene" authorityYear="2024" box="[188,324,1555,1581]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="346" phylum="Arthropoda" rank="species" species="kniphofia" status="sp. n.">
<emphasis id="09B06B02FFD5AC18FF03F9C9FE87F9D5" box="[188,324,1555,1581]" italics="true" pageId="3" pageNumber="346">T. kniphofia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1283D679FFD5AC18FEF2F9C9FE4EF9D5" box="[333,397,1555,1581]" pageId="3" pageNumber="346" rank="species">sp. n.</taxonomicNameLabel>
,
<taxonomicName id="FCC4CC93FFD5AC18FE23F9C9FCCFF9D5" authority="(Queinnec &amp; Ollivier)" baseAuthorityName="Queinnec &amp; Ollivier" baseAuthorityYear="2021" box="[412,780,1555,1581]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="346" phylum="Arthropoda" rank="species" species="reebae">
<emphasis id="09B06B02FFD5AC18FE23F9C9FDC6F9D5" box="[412,517,1555,1581]" italics="true" pageId="3" pageNumber="346">T. reebae</emphasis>
(Quéinnec &amp; Ollivier)
</taxonomicName>
and
<taxonomicName id="FCC4CC93FFD5AC18FCF8F9C9FC04F9D5" authorityName="Schmidt &amp; Merene" authorityYear="2024" box="[839,967,1555,1581]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="346" phylum="Arthropoda" rank="species" species="yitbareki" status="sp. n.">
<emphasis id="09B06B02FFD5AC18FCF8F9C9FC04F9D5" box="[839,967,1555,1581]" italics="true" pageId="3" pageNumber="346">T. yitbareki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1283D679FFD5AC18FC6FF9C9FBCDF9D5" box="[976,1038,1555,1581]" pageId="3" pageNumber="346" rank="species">sp. n.</taxonomicNameLabel>
in the more deeply impressed and distinctly punctate elytral striae (finely impressed and smooth or indistinctly punctate in all the latter species). In habitus,
<taxonomicName id="FCC4CC93FFD5AC18FF48F981FEA4F98C" authorityName="Schmidt &amp; Merene" authorityYear="2024" box="[247,359,1627,1652]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="346" phylum="Arthropoda" rank="species" species="apertus" status="sp. n.">
<emphasis id="09B06B02FFD5AC18FF48F981FEA4F98C" box="[247,359,1627,1652]" italics="true" pageId="3" pageNumber="346">T. apertus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1283D679FFD5AC18FED1F981FE68F98D" box="[366,427,1627,1653]" pageId="3" pageNumber="346" rank="species">sp. n.</taxonomicNameLabel>
is very similar to
<taxonomicName id="FCC4CC93FFD5AC18FDC6F981FC27F98D" authority="Queinnec &amp; Ollivier" authorityName="Queinnec &amp; Ollivier" box="[633,996,1627,1653]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="346" phylum="Arthropoda" rank="species" species="niloticus">
<emphasis id="09B06B02FFD5AC18FDC6F981FD36F98D" box="[633,757,1627,1653]" italics="true" pageId="3" pageNumber="346">T. niloticus</emphasis>
Quéinnec &amp; Ollivier
</taxonomicName>
which was described from the western crater rim of Mt. Choke. However, in
<taxonomicName id="FCC4CC93FFD5AC18FDFAF9A5FD02F961" box="[581,705,1663,1689]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="346" phylum="Arthropoda" rank="species" species="niloticus">
<emphasis id="09B06B02FFD5AC18FDFAF9A5FD02F961" box="[581,705,1663,1689]" italics="true" pageId="3" pageNumber="346">T. niloticus</emphasis>
</taxonomicName>
the dorsal opening of the aedeagal median lobe do not reach the basal bulb (see
<bibRefCitation id="5F55CAE1FFD5AC18FEF8F979FDF5F945" author="Queinnec, E. &amp; Ollivier, E. &amp; Reeb, C." box="[327,566,1699,1725]" pageId="3" pageNumber="346" pagination="1 - 70" refId="ref9659" refString="Queinnec, E., Ollivier, E. &amp; Reeb, C. (2021) Insights on Ethiopian montane ground beetle biodiversity: Taxonomic study of afro-alpine and sub-alpine Trechini (Coleoptera: Carabidae: Trechinae). Journal of Insect Biodiversity, 27 (1), 1 - 70. https: // doi. org / 10.12976 / jib / 2021.27.1.1" type="journal article" year="2021">
Quéinnec
<emphasis id="09B06B02FFD5AC18FE05F979FE30F945" box="[442,499,1699,1725]" italics="true" pageId="3" pageNumber="346">et al.</emphasis>
2021
</bibRefCitation>
: Fig. 26B, C). In addition, in
<taxonomicName id="FCC4CC93FFD5AC18FC3AF979FBC2F945" box="[901,1025,1699,1725]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="346" phylum="Arthropoda" rank="species" species="niloticus">
<emphasis id="09B06B02FFD5AC18FC3AF979FBC2F945" box="[901,1025,1699,1725]" italics="true" pageId="3" pageNumber="346">T. niloticus</emphasis>
</taxonomicName>
, the median lobe is more elongated, with ventral margin straight in middle (convex in
<taxonomicName id="FCC4CC93FFD5AC18FD7FF91DFCECF918" authorityName="Schmidt &amp; Merene" authorityYear="2024" box="[704,815,1735,1760]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="346" phylum="Arthropoda" rank="species" species="apertus" status="sp. n.">
<emphasis id="09B06B02FFD5AC18FD7FF91DFCECF918" box="[704,815,1735,1760]" italics="true" pageId="3" pageNumber="346">T. apertus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1283D679FFD5AC18FC8AF91DFCB1F919" box="[821,882,1735,1761]" pageId="3" pageNumber="346" rank="species">sp. n.</taxonomicNameLabel>
), its apical lamella is much longer, markedly bent ventrally (short and slightly bent ventrally in
<taxonomicName id="FCC4CC93FFD5AC18FD2CF931FCC1F8FC" authorityName="Schmidt &amp; Merene" authorityYear="2024" box="[659,770,1771,1796]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="346" phylum="Arthropoda" rank="species" species="apertus" status="sp. n.">
<emphasis id="09B06B02FFD5AC18FD2CF931FCC1F8FC" box="[659,770,1771,1796]" italics="true" pageId="3" pageNumber="346">T. apertus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1283D679FFD5AC18FCB6F931FC85F8FD" box="[777,838,1771,1797]" pageId="3" pageNumber="346" rank="species">sp. n.</taxonomicNameLabel>
), the parameres are slenderer.
</paragraph>
</subSubSection>
<subSubSection id="73DEE49BFFD5AC1FFF78F8D4FC78FF2D" lastPageId="4" lastPageNumber="347" pageId="3" pageNumber="346" type="distribution">
<paragraph id="3B7BB710FFD5AC18FF78F8D4FC4EF8B5" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="346">
<emphasis id="09B06B02FFD5AC18FF78F8D4FE9AF8D0" bold="true" box="[199,345,1806,1832]" pageId="3" pageNumber="346">Distribution</emphasis>
. Endemic to Mt. Choke in northern
<collectingCountry id="43D3F780FFD5AC18FD46F8D5FC9FF8D1" box="[761,860,1807,1833]" name="Ethiopia" pageId="3" pageNumber="346">Ethiopia</collectingCountry>
. The new species was only found near the top of a mountain ridge that extends from the crater rim to the west (
<figureCitation id="A3FFAB95FFD5AC18FC85F8E9FCBCF8B5" box="[826,895,1843,1869]" captionStart="FIGURE 4" captionStartId="9.[152,255,1559,1583]" captionTargetBox="[151,1436,561,1535]" captionTargetId="figure-219@9.[151,1436,561,1535]" captionTargetPageId="9" captionText="FIGURE 4. Topographic map of Mt. Choke (highest point marked by a white triangle), showing sampling localities of six Trechus (Abyssinotus) species of the niloticus group (white circles; the numbers refer to the different species). 1, T. apertus Schmidt, sp. n. 2, T. reebae (Quéinnec &amp; Ollivier). 3, T. yitbareki Schmidt, sp. n. 4, T. kniphofia Schmidt, sp. n. 5, T. igori Schmidt, sp. n. 6, T. niloticus (Quéinnec &amp; Ollivier). Sampling data of T. reebae and T. niloticus published by Quéinnec et al. (2021) are considered. Base map was downloaded from Topographic-map.com." figureDoi="http://doi.org/10.5281/zenodo.13234861" httpUri="https://zenodo.org/record/13234861/files/figure.png" pageId="3" pageNumber="346">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="3B7BB710FFD5AC1FFF78F88CFC78FF2D" blockId="3.[151,1437,151,2013]" lastBlockId="4.[151,1436,151,213]" lastPageId="4" lastPageNumber="347" pageId="3" pageNumber="346">
<emphasis id="09B06B02FFD5AC18FF78F88CFE75F888" bold="true" box="[199,438,1878,1904]" pageId="3" pageNumber="346">Habitat preferences</emphasis>
. All specimens of
<taxonomicName id="FCC4CC93FFD5AC18FD29F88DFCCAF888" authorityName="Schmidt &amp; Merene" authorityYear="2024" box="[662,777,1879,1904]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="346" phylum="Arthropoda" rank="species" species="apertus" status="sp. n.">
<emphasis id="09B06B02FFD5AC18FD29F88DFCCAF888" box="[662,777,1879,1904]" italics="true" pageId="3" pageNumber="346">T. apertus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1283D679FFD5AC18FCABF88DFC90F889" box="[788,851,1879,1905]" pageId="3" pageNumber="346" rank="species">sp. n.</taxonomicNameLabel>
were sifted from the humus-rich soil under dense vegetation alongside a small brook. The brook has its source near the top of the mountain ridge and flows to the north, at an elevation of
<quantity id="FC3C1AF5FFD5AC18FE62F844FDBEF841" box="[477,637,1950,1977]" metricMagnitude="3" metricUnit="m" metricValue="3.41" metricValueMax="3.45" metricValueMin="3.37" pageId="3" pageNumber="346" unit="m" value="3410.0" valueMax="3450.0" valueMin="3370.0">3370–3450 m</quantity>
. No specimens were found on the pastures and in the bushland which bordered the brook. Based on these collecting circumstances, the new species can be considered hygrophilic and cold-adapted. As potentially natural vegetation, a moist afromontane forest would have formed at the site where the new species was found, but here it has given way to a cultivated steppe.
</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue View git blame Copy permalink