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<document id="14CF97F5A3A8D8D00B438DA97AA6D15A" ID-DOI="10.11646/zootaxa.5492.3.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="13234851" ID-ZooBank="8F81F053-DE78-4684-BCC9-875AE0C41615" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722932859467" checkinUser="plazi" docAuthor="Schmidt, Joachim &amp; Merene, Yeshitla" docDate="2024" docId="B36D0606FFD1AC12FF28FAE0FE37FDE1" docLanguage="en" docName="zootaxa.5492.3.3.pdf" docOrigin="Zootaxa 5492 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5492.3.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trechus (Abyssinotus) igori Schmidt &amp; Merene 2024, sp. n." docType="treatment" docVersion="2" lastPageNumber="352" masterDocId="4F547E7EFFD6AC1BFFBFFFDAFFC3FFF8" masterDocTitle="Trechus species from Mt. Choke of northern Ethiopia related to T. niloticus (Quéinnec &amp; Ollivier) with notable male genital morphology (Carabidae: Trechini)" masterLastPageNumber="355" masterPageNumber="343" pageNumber="350" updateTime="1722933277969" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="52E07D6CAF6E619986C601CAF893DB60" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6B878D691938C1EC1F613E572CD369B9">
<mods:title id="F01736CA992C647404F2419A02AE3BCD">Trechus species from Mt. Choke of northern Ethiopia related to T. niloticus (Quéinnec &amp; Ollivier) with notable male genital morphology (Carabidae: Trechini)</mods:title>
</mods:titleInfo>
<mods:name id="31C68CB8723F76D18493DACB791B2C19" type="personal">
<mods:role id="127571F9C053C54C96A1709BB564F71E">
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</mods:role>
<mods:namePart id="1E110C2386B1EFA080E92891168AF5AD">Schmidt, Joachim</mods:namePart>
<mods:affiliation id="13F87AC637CDBC63B479A40A478D8F99">General and Systematic Zoology, University of Rostock, Germany</mods:affiliation>
</mods:name>
<mods:name id="81C6F70E5655E8C4AF0D5399A3DA648D" type="personal">
<mods:role id="F760CAAF7E15D09AA58B97AE8872BA7E">
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</mods:role>
<mods:namePart id="94BAA5D224EA53A3969BA9A0FD7BE2BF">Merene, Yeshitla</mods:namePart>
<mods:affiliation id="7F1E05ED833356A10F9A8E8D40F7FCB9">Department of Zoological Sciences, Addis Ababa University, Ethiopia &amp; Amhara Agricultural Research Institute, Bahir Dar, Ethiopia</mods:affiliation>
</mods:name>
<mods:typeOfResource id="5AF9CC7B23B732A506E3C9D7BF02AA8D">text</mods:typeOfResource>
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<mods:title id="E30218A4E310C81219ED2CE734B2F14E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="DB6AC82ED59BB56DE7D8B261BACF2B2F">
<mods:date id="FC82120D1871C07E0ECCAFCA8C6629A6">2024</mods:date>
<mods:detail id="0344E5E7674B28986C03A7A0231B2901" type="pubDate">
<mods:number id="A14A6AB2E0D34E8E31ECC30A3A7397F2">2024-08-06</mods:number>
</mods:detail>
<mods:detail id="1E778E1597A55846BEB8F8D526F0CA75" type="volume">
<mods:number id="45AE04BB6E5484C607B9728F424F46B3">5492</mods:number>
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<mods:detail id="6B26DBC5AC4405F57F42E87E96EE5AA4" type="issue">
<mods:number id="DB2BD6B3D389E28D9EF53C947297EA9E">3</mods:number>
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<mods:location id="6253D10C74CBD745EE0B97AB9CF43CF8">
<mods:url id="0F8571E7426E3FD2D476225EA9064E41">http://dx.doi.org/10.11646/zootaxa.5492.3.3</mods:url>
</mods:location>
<mods:classification id="9835614FBD57F02A3E0C4FE5D1CB9617">journal article</mods:classification>
<mods:identifier id="6FD40B8DA5CF38DD61EEBCABF77E6EDE" type="DOI">10.11646/zootaxa.5492.3.3</mods:identifier>
<mods:identifier id="2CF0FF91301AB362550D0D45BC58A6B0" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E01AD76474871F71F14B95DF24E728C9" type="Zenodo-Dep">13234851</mods:identifier>
<mods:identifier id="55357A09F594FA0D8E3A37B6B21BA5A1" type="ZooBank">8F81F053-DE78-4684-BCC9-875AE0C41615</mods:identifier>
</mods:mods>
<treatment id="B36D0606FFD1AC12FF28FAE0FE37FDE1" ID-DOI="http://doi.org/10.5281/zenodo.13234871" ID-Zenodo-Dep="13234871" LSID="urn:lsid:plazi:treatment:B36D0606FFD1AC12FF28FAE0FE37FDE1" httpUri="http://treatment.plazi.org/id/B36D0606FFD1AC12FF28FAE0FE37FDE1" lastPageId="9" lastPageNumber="352" pageId="7" pageNumber="350">
<subSubSection id="73DEE49BFFD1AC1CFF28FAE0FD65FAAD" box="[151,678,1338,1365]" pageId="7" pageNumber="350" type="nomenclature">
<paragraph id="3B7BB710FFD1AC1CFF28FAE0FD65FAAD" blockId="7.[151,678,1338,1401]" box="[151,678,1338,1365]" pageId="7" pageNumber="350">
<heading id="6033007CFFD1AC1CFF28FAE0FD65FAAD" bold="true" box="[151,678,1338,1365]" fontSize="11" level="1" pageId="7" pageNumber="350" reason="1">
<emphasis id="09B06B02FFD1AC1CFF28FAE0FD65FAAD" bold="true" box="[151,678,1338,1365]" pageId="7" pageNumber="350">
<taxonomicName id="FCC4CC93FFD1AC1CFF28FAE0FD94FAAC" authority="Schmidt" authorityName="Schmidt &amp; Merene" authorityYear="2024" box="[151,599,1338,1365]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="350" phylum="Arthropoda" rank="species" species="igori" status="sp. nov." subGenus="Abyssinotus">
<emphasis id="09B06B02FFD1AC1CFF28FAE0FF3AFAAD" bold="true" box="[151,249,1338,1365]" italics="true" pageId="7" pageNumber="350">Trechus</emphasis>
(
<emphasis id="09B06B02FFD1AC1CFEB4FAE0FE5FFAAD" bold="true" box="[267,412,1338,1365]" italics="true" pageId="7" pageNumber="350">Abyssinotus</emphasis>
)
<emphasis id="09B06B02FFD1AC1CFE12FAE1FE25FAAD" bold="true" box="[429,486,1339,1365]" italics="true" pageId="7" pageNumber="350">igori</emphasis>
Schmidt
</taxonomicName>
,
<taxonomicNameLabel id="1283D679FFD1AC1CFDDBFAE1FD65FAAD" box="[612,678,1339,1365]" pageId="7" pageNumber="350" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="73DEE49BFFD1AC1CFF28FA85FEFCFA81" box="[151,319,1375,1401]" pageId="7" pageNumber="350" type="description">
<paragraph id="3B7BB710FFD1AC1CFF28FA85FEFCFA81" blockId="7.[151,678,1338,1401]" box="[151,319,1375,1401]" pageId="7" pageNumber="350">
<figureCitation id="A3FFAB95FFD1AC1CFF28FA85FF2CFA81" box="[151,239,1375,1401]" captionStart="FIGURE 2" captionStartId="5.[152,255,1889,1913]" captionTargetBox="[280,1308,182,1866]" captionTargetId="figure-24@5.[280,1308,181,1866]" captionTargetPageId="5" captionText="FIGURE 2. Habitus photographs of Trechus (Abyssinotus) species; the white circles point to the insertions of the supraorbital setae, pronotal lateral setae, parascutellar seta, elytral discal setae, and setae of the umbilicate series. A, T. reebae (Quéinnec &amp; Ollivier, 2021), specimen from western crater valley. B, T. yitbareki Schmidt, sp. n., paratype. C, T. kniphofia Schmidt, sp. n., paratype. D, T. igori Schmidt, sp. n., paratype." figureDoi="http://doi.org/10.5281/zenodo.13234855" httpUri="https://zenodo.org/record/13234855/files/figure.png" pageId="7" pageNumber="350">Fig. 2D</figureCitation>
,
<figureCitation id="A3FFAB95FFD1AC1CFF44FA85FEFCFA81" box="[251,319,1375,1401]" captionStart="FIGURE 3" captionStartId="8.[152,255,1470,1494]" captionTargetBox="[152,1436,181,1445]" captionTargetId="figure-23@8.[152,1436,181,1446]" captionTargetPageId="8" captionText="FIGURE 3. Left protarsomeres (A–C), aedeagus in left lateral view (D–G) and dorsal view (H–K) of Trechus (Abyssinotus) species. A, D, I, T. reebae (Quéinnec &amp; Ollivier, 2021), specimen from western crater valley. B, E, J, T. kniphofia Schmidt, sp. n., paratype. C, F, H, T. yitbareki Schmidt, sp. n., paratype. G, K, T. igori Schmidt, sp. n., paratype." figureDoi="http://doi.org/10.5281/zenodo.13234857" httpUri="https://zenodo.org/record/13234857/files/figure.png" pageId="7" pageNumber="350">3G, K</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="73DEE49BFFD1AC1CFF28FA7DFE13F9F1" pageId="7" pageNumber="350" type="materials_examined">
<paragraph id="3B7BB710FFD1AC1CFF28FA7DFE13F9F1" blockId="7.[151,1437,1446,1977]" pageId="7" pageNumber="350">
<emphasis id="09B06B02FFD1AC1CFF28FA7DFE81FA38" bold="true" box="[151,322,1446,1473]" pageId="7" pageNumber="350">Type material.</emphasis>
<materialsCitation id="8BACBD4DFFD1AC1CFEF9FA7DFC3DFA1D" collectingDate="2019-02-23" collectionCode="CSCHM" collectorName="D. Hauth &amp; Hauth, J &amp; Yeshitla M. &amp; Yitbarek, W." country="Ethiopia" elevation="3370" latitude="10.635278" location="Mt. Choke" longLatPrecision="21" longitude="37.764168" pageId="7" pageNumber="350" specimenCount="1" specimenCount-male="1" stateProvince="Amara" typeStatus="holotype">
<typeStatus id="E47F09B2FFD1AC1CFEF9FA7DFE70FA39" box="[326,435,1447,1473]" pageId="7" pageNumber="350" type="holotype">Holotype</typeStatus>
:
<specimenCount id="2DC27C99FFD1AC1CFE04FA7DFE11FA39" box="[443,466,1447,1473]" count="1" pageId="7" pageNumber="350" type="male">♂</specimenCount>
, with label data:
<collectingCountry id="43D3F780FFD1AC1CFD39FA7DFD2BFA39" box="[646,744,1447,1473]" name="Ethiopia" pageId="7" pageNumber="350">Ethiopia</collectingCountry>
,
<collectingRegion id="F90079F2FFD1AC1CFD50FA7DFC8FFA39" box="[751,844,1447,1473]" country="Ethiopia" name="Amara" pageId="7" pageNumber="350">Amhara</collectingRegion>
, W-slope
<location id="3E1BE1CBFFD1AC1CFC0AFA7DFBEEFA39" LSID="urn:lsid:plazi:treatment:B36D0606FFD1AC12FF28FAE0FE37FDE1:3E1BE1CBFFD1AC1CFC0AFA7DFBEEFA39" box="[949,1069,1447,1473]" country="Ethiopia" latitude="10.635278" longLatPrecision="21" longitude="37.764168" name="Mt. Choke" pageId="7" pageNumber="350" stateProvince="Amara">Mt.Choke</location>
,
<quantity id="FC3C1AF5FFD1AC1CFB89FA7DFB49FA39" box="[1078,1162,1447,1473]" metricMagnitude="3" metricUnit="m" metricValue="3.37" pageId="7" pageNumber="350" unit="m" value="3370.0">
<elevation id="B0E95023FFD1AC1CFB89FA7DFB49FA39" box="[1078,1162,1447,1473]" metricMagnitude="3" metricUnit="m" metricValue="3.37" pageId="7" pageNumber="350" unit="m" value="3370.0">3370 m</elevation>
</quantity>
,
<date id="4F7A91D0FFD1AC1CFB2CFA7DFAC8FA39" box="[1171,1291,1447,1473]" pageId="7" pageNumber="350" value="2019-02-23">
<collectingDate id="5F3E6838FFD1AC1CFB2CFA7DFAC8FA39" box="[1171,1291,1447,1473]" pageId="7" pageNumber="350" value="2019-02-23">23.II.2019</collectingDate>
</date>
,
<geoCoordinate id="5EF0D1D7FFD1AC1CFAACFA7DFA5FFA39" box="[1299,1436,1447,1473]" degrees="10" direction="north" minutes="38" orientation="latitude" pageId="7" pageNumber="350" precision="15" seconds="07" value="10.635278">10°38’07”N</geoCoordinate>
<geoCoordinate id="5EF0D1D7FFD1AC1CFF28FA10FEE3FA1D" box="[151,288,1482,1509]" degrees="37" direction="east" minutes="45" orientation="longitude" pageId="7" pageNumber="350" precision="15" seconds="51" value="37.764168">37°45’51”E</geoCoordinate>
, leg.
<collectorName id="9631D2C6FFD1AC1CFEE0FA11FE09FA1D" box="[351,458,1483,1509]" pageId="7" pageNumber="350">D. Hauth</collectorName>
, J. Schmidt,
<collectorName id="9631D2C6FFD1AC1CFDE2FA11FD21FA1D" box="[605,738,1483,1509]" pageId="7" pageNumber="350">Yeshitla M.</collectorName>
,
<collectorName id="9631D2C6FFD1AC1CFD50FA11FCBDFA1D" box="[751,894,1483,1509]" pageId="7" pageNumber="350">Yitbarek, W.</collectorName>
(
<collectionCode id="5DD52FD5FFD1AC1CFC31FA11FC34FA1D" box="[910,1015,1483,1509]" pageId="7" pageNumber="350">CSCHM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="8BACBD4DFFD1AC1CFBB5FA11FE0EF9F1" collectingDate="2019-02-23" collectionCode="CSCHM" collectorName="D. Hauth &amp; Hauth, J &amp; Yeshitla M. &amp; Yitbarek, W." country="Ethiopia" elevation="3370" latitude="10.635278" location="Mt. Choke" longLatPrecision="21" longitude="37.764168" pageId="7" pageNumber="350" specimenCount="15" specimenCount-female="7" specimenCount-male="8" stateProvince="Amara" typeStatus="paratype">
<typeStatus id="E47F09B2FFD1AC1CFBB5FA11FBB8FA1D" box="[1034,1147,1483,1509]" pageId="7" pageNumber="350" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="2DC27C99FFD1AC1CFB37FA11FB09FA1D" box="[1160,1226,1483,1509]" count="8" pageId="7" pageNumber="350" type="male">8 ♂♂</specimenCount>
,
<specimenCount id="2DC27C99FFD1AC1CFB67FA11FAD9FA1D" box="[1240,1306,1483,1509]" count="7" pageId="7" pageNumber="350" type="female">7 ♀♀</specimenCount>
, with same data as holotype (
<collectionCode id="5DD52FD5FFD1AC1CFEE3FA35FE06F9F1" box="[348,453,1519,1545]" pageId="7" pageNumber="350">CSCHM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="73DEE49BFFD1AC1CFF78F9C9FC2DF9A9" pageId="7" pageNumber="350" type="etymology">
<paragraph id="3B7BB710FFD1AC1CFF78F9C9FC2DF9A9" blockId="7.[151,1437,1446,1977]" pageId="7" pageNumber="350">
<emphasis id="09B06B02FFD1AC1CFF78F9C9FE8BF9D5" bold="true" box="[199,328,1555,1581]" pageId="7" pageNumber="350">Etymology</emphasis>
. The new species is dedicated to my dear friend and colleague, the distinguished carabidologist and explorer of the high altitude environments, Igor A. Belousov, St. Petersburg.
</paragraph>
</subSubSection>
<subSubSection id="73DEE49BFFD1AC13FF78F980FF14F837" lastPageId="8" lastPageNumber="351" pageId="7" pageNumber="350" type="description">
<paragraph id="3B7BB710FFD1AC1CFF78F980FD45F98D" blockId="7.[151,1437,1446,1977]" box="[199,646,1626,1653]" pageId="7" pageNumber="350">
<emphasis id="09B06B02FFD1AC1CFF78F980FE92F98C" bold="true" box="[199,337,1626,1652]" pageId="7" pageNumber="350">Description</emphasis>
. Body length:
<quantity id="FC3C1AF5FFD1AC1CFE47F981FD41F98D" box="[504,642,1627,1653]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.3" metricValueMin="2.2" pageId="7" pageNumber="350" unit="mm" value="2.25" valueMax="2.3" valueMin="2.2">2.2–2.3 mm</quantity>
.
</paragraph>
<paragraph id="3B7BB710FFD1AC1CFF78F9A5FA4CF945" blockId="7.[151,1437,1446,1977]" pageId="7" pageNumber="350">Proportions (n = 10): PW/HW = 1.31–1.41 (Ø = 1.36); PW/PL = 1.41–1.50 (Ø = 1.44); PW/PBW = 1.23–1.30 (Ø = 1.27); PBW/PAW = 1.07–1.12 (Ø = 1.10); EW/PW = 1.44–1.53 (Ø = 1.50); EL/EW = 1.30–1.35 (Ø = 1.32).</paragraph>
<paragraph id="3B7BB710FFD1AC1CFF78F91DFD30F8FD" blockId="7.[151,1437,1446,1977]" pageId="7" pageNumber="350">Colour: Head capsule, pronotum and elytra dark brown, pronotum paler in some specimens; clypeus, labrum, mandibles, palps, antennae and legs yellowish brown.</paragraph>
<paragraph id="3B7BB710FFD1AC1CFF78F8D5FAE9F8B5" blockId="7.[151,1437,1446,1977]" pageId="7" pageNumber="350">Microsculpture: Sculpticells on frons and supraorbital area large, moderately impressed, almost isodiametric, slightly transverse on clypeus. Sculpticells on pronotal disc and elytra moderately impressed, transverse.</paragraph>
<paragraph id="3B7BB710FFD1AC1CFF78F88DFD45F889" blockId="7.[151,1437,1446,1977]" box="[199,646,1879,1905]" pageId="7" pageNumber="350">
Head: As described in
<taxonomicName id="FCC4CC93FFD1AC1CFE7AF88DFD80F889" authorityName="Schmidt &amp; Merene" authorityYear="2024" box="[453,579,1879,1905]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="350" phylum="Arthropoda" rank="species" species="yitbareki" status="sp. n.">
<emphasis id="09B06B02FFD1AC1CFE7AF88DFD80F889" box="[453,579,1879,1905]" italics="true" pageId="7" pageNumber="350">T. yitbareki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1283D679FFD1AC1CFDF5F88DFD45F889" box="[586,646,1879,1905]" pageId="7" pageNumber="350" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="3B7BB710FFD1AC1CFF78F8A1FB3BF841" blockId="7.[151,1437,1446,1977]" pageId="7" pageNumber="350">
Prothorax: Pronotum moderately large, transverse, sub-discoidal, with base slightly more markedly narrowed towards base than in the proceeding species. In all other characters as described in
<taxonomicName id="FCC4CC93FFD1AC1CFB88F845FB76F841" authorityName="Schmidt &amp; Merene" authorityYear="2024" box="[1079,1205,1951,1977]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="350" phylum="Arthropoda" rank="species" species="yitbareki" status="sp. n.">
<emphasis id="09B06B02FFD1AC1CFB88F845FB76F841" box="[1079,1205,1951,1977]" italics="true" pageId="7" pageNumber="350">T. yitbareki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1283D679FFD1AC1CFB03F845FB3BF841" box="[1212,1272,1951,1977]" pageId="7" pageNumber="350" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<caption id="6FBBE798FFDEAC13FF27FA64FB49F9E7" ID-DOI="http://doi.org/10.5281/zenodo.13234857" ID-Zenodo-Dep="13234857" httpUri="https://zenodo.org/record/13234857/files/figure.png" pageId="8" pageNumber="351" startId="8.[152,255,1470,1494]" targetBox="[152,1436,181,1445]" targetPageId="8" targetType="figure">
<paragraph id="3B7BB710FFDEAC13FF27FA64FB49F9E7" blockId="8.[151,1437,1470,1567]" pageId="8" pageNumber="351">
<emphasis id="09B06B02FFDEAC13FF27FA64FEDAFA2E" bold="true" box="[152,281,1470,1494]" pageId="8" pageNumber="351">FIGURE 3.</emphasis>
Left protarsomeres (A–C), aedeagus in left lateral view (D–G) and dorsal view (H–K) of
<taxonomicName id="FCC4CC93FFDEAC13FB04FA64FA5FFA2E" box="[1211,1436,1470,1494]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="351" phylum="Arthropoda" rank="subGenus" subGenus="Abyssinotus">
<emphasis id="09B06B02FFDEAC13FB04FA64FAC9FA2E" box="[1211,1290,1470,1494]" italics="true" pageId="8" pageNumber="351">Trechus</emphasis>
(
<emphasis id="09B06B02FFDEAC13FAA4FA64FA57FA2E" box="[1307,1428,1470,1494]" italics="true" pageId="8" pageNumber="351">Abyssinotus</emphasis>
)
</taxonomicName>
species.
<emphasis id="09B06B02FFDEAC13FF52FA38FF3CFA02" bold="true" box="[237,255,1506,1530]" pageId="8" pageNumber="351">A</emphasis>
,
<emphasis id="09B06B02FFDEAC13FEB3FA38FEDDFA02" bold="true" box="[268,286,1506,1530]" pageId="8" pageNumber="351">D</emphasis>
,
<emphasis id="09B06B02FFDEAC13FE94FA38FEF6FA02" bold="true" box="[299,309,1506,1530]" pageId="8" pageNumber="351">I</emphasis>
,
<taxonomicName id="FCC4CC93FFDEAC13FEFEFA39FD06FA02" authority="(Queinnec &amp; Ollivier, 2021)" baseAuthorityName="Queinnec &amp; Ollivier" baseAuthorityYear="2021" box="[321,709,1506,1530]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="351" phylum="Arthropoda" rank="species" species="reebae">
<emphasis id="09B06B02FFDEAC13FEFEFA39FE5DFA02" box="[321,414,1506,1530]" italics="true" pageId="8" pageNumber="351">T. reebae</emphasis>
(
<bibRefCitation id="5F55CAE1FFDEAC13FE14FA38FD7DFA02" author="Queinnec, E. &amp; Ollivier, E. &amp; Reeb, C." box="[427,702,1506,1530]" pageId="8" pageNumber="351" pagination="1 - 70" refId="ref9659" refString="Queinnec, E., Ollivier, E. &amp; Reeb, C. (2021) Insights on Ethiopian montane ground beetle biodiversity: Taxonomic study of afro-alpine and sub-alpine Trechini (Coleoptera: Carabidae: Trechinae). Journal of Insect Biodiversity, 27 (1), 1 - 70. https: // doi. org / 10.12976 / jib / 2021.27.1.1" type="journal article" year="2021">Quéinnec &amp; Ollivier, 2021</bibRefCitation>
)
</taxonomicName>
, specimen from western crater valley.
<emphasis id="09B06B02FFDEAC13FBF4FA38FB9FFA02" bold="true" box="[1099,1116,1506,1530]" pageId="8" pageNumber="351">B</emphasis>
,
<emphasis id="09B06B02FFDEAC13FBD6FA38FBB9FA02" bold="true" box="[1129,1146,1506,1530]" pageId="8" pageNumber="351">E</emphasis>
,
<emphasis id="09B06B02FFDEAC13FB39FA38FB50FA02" bold="true" box="[1158,1171,1506,1530]" pageId="8" pageNumber="351">J</emphasis>
,
<taxonomicName id="FCC4CC93FFDEAC13FB1FFA39FAB6FA02" authority="Schmidt" authorityName="Schmidt &amp; Merene" authorityYear="2024" box="[1184,1397,1506,1530]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="351" phylum="Arthropoda" rank="species" species="kniphofia" status="sp. n.">
<emphasis id="09B06B02FFDEAC13FB1FFA39FADBFA02" box="[1184,1304,1506,1530]" italics="true" pageId="8" pageNumber="351">T. kniphofia</emphasis>
Schmidt
</taxonomicName>
,
<taxonomicNameLabel id="1283D679FFDEAC13FAC0FA39FF6FF9E6" pageId="8" pageNumber="351" rank="species">sp. n.</taxonomicNameLabel>
, paratype.
<emphasis id="09B06B02FFDEAC13FEA4F9DCFEEEF9E6" bold="true" box="[283,301,1542,1566]" pageId="8" pageNumber="351">C</emphasis>
,
<emphasis id="09B06B02FFDEAC13FE85F9DCFE8AF9E6" bold="true" box="[314,329,1542,1566]" pageId="8" pageNumber="351">F</emphasis>
,
<emphasis id="09B06B02FFDEAC13FEE9F9DCFEA9F9E6" bold="true" box="[342,362,1542,1566]" pageId="8" pageNumber="351">H</emphasis>
,
<taxonomicName id="FCC4CC93FFDEAC13FEC9F9DDFD85F9E6" authority="Schmidt" authorityName="Schmidt &amp; Merene" authorityYear="2024" box="[374,582,1542,1566]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="351" phylum="Arthropoda" rank="species" species="yitbareki" status="sp. n.">
<emphasis id="09B06B02FFDEAC13FEC9F9DDFE2BF9E6" box="[374,488,1542,1566]" italics="true" pageId="8" pageNumber="351">T. yitbareki</emphasis>
Schmidt
</taxonomicName>
,
<taxonomicNameLabel id="1283D679FFDEAC13FDEFF9DDFD44F9E6" box="[592,647,1543,1566]" pageId="8" pageNumber="351" rank="species">sp. n.</taxonomicNameLabel>
, paratype.
<emphasis id="09B06B02FFDEAC13FD49F9DCFCC9F9E6" bold="true" box="[758,778,1542,1566]" pageId="8" pageNumber="351">G</emphasis>
,
<emphasis id="09B06B02FFDEAC13FCA8F9DCFCE8F9E6" bold="true" box="[791,811,1542,1566]" pageId="8" pageNumber="351">K</emphasis>
,
<taxonomicName id="FCC4CC93FFDEAC13FC88F9DDFC1CF9E6" authority="Schmidt" authorityName="Schmidt &amp; Merene" authorityYear="2024" box="[823,991,1542,1566]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="351" phylum="Arthropoda" rank="species" species="igori" status="sp. n.">
<emphasis id="09B06B02FFDEAC13FC88F9DDFC42F9E6" box="[823,897,1543,1566]" italics="true" pageId="8" pageNumber="351">T. igori</emphasis>
Schmidt
</taxonomicName>
,
<taxonomicNameLabel id="1283D679FFDEAC13FC56F9DDFBE3F9E6" box="[1001,1056,1543,1566]" pageId="8" pageNumber="351" rank="species">sp. n.</taxonomicNameLabel>
, paratype.
</paragraph>
</caption>
<paragraph id="3B7BB710FFDEAC13FF78F9AAFEB6F92B" blockId="8.[151,1437,1648,1999]" pageId="8" pageNumber="351">
Pterothorax: First stria absent in anterior 2/3, suggestively impressed before its apex, striae 2 and 3 very finely impressed, indistinct, external striae absent apart from the apical portion of stria 8. In all other characters as described in
<taxonomicName id="FCC4CC93FFDEAC13FF0BF963FEF1F92A" authorityName="Schmidt &amp; Merene" authorityYear="2024" box="[180,306,1720,1746]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="351" phylum="Arthropoda" rank="species" species="yitbareki" status="sp. n.">
<emphasis id="09B06B02FFDEAC13FF0BF963FEF1F92A" box="[180,306,1720,1746]" italics="true" pageId="8" pageNumber="351">T. yitbareki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1283D679FFDEAC13FE86F963FEB6F92B" box="[313,373,1721,1747]" pageId="8" pageNumber="351" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="3B7BB710FFDEAC13FF78F907FDD5F90F" blockId="8.[151,1437,1648,1999]" box="[199,534,1756,1783]" pageId="8" pageNumber="351">
Legs: As in
<taxonomicName id="FCC4CC93FFDEAC13FEF1F907FE10F90E" authorityName="Schmidt &amp; Merene" authorityYear="2024" box="[334,467,1756,1782]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="351" phylum="Arthropoda" rank="species" species="kniphofia" status="sp. n.">
<emphasis id="09B06B02FFDEAC13FEF1F907FE10F90E" box="[334,467,1756,1782]" italics="true" pageId="8" pageNumber="351">T. kniphofia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1283D679FFDEAC13FE65F907FDD5F90F" box="[474,534,1757,1783]" pageId="8" pageNumber="351" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph id="3B7BB710FFDEAC13FF78F8DAFF14F837" blockId="8.[151,1437,1648,1999]" pageId="8" pageNumber="351">
Male genitalia: EL/AL = 2.20–2.30 (Ø = 2.24; n = 8). Aedeagal median lobe moderately robust, rather stout, in lateral view evenly curved throughout, with apical lamella very short, very slightly sclerotized, very shortly bent upwardly (
<figureCitation id="A3FFAB95FFDEAC13FEAFF892FEA9F89A" box="[272,362,1864,1890]" captionStart="FIGURE 3" captionStartId="8.[152,255,1470,1494]" captionTargetBox="[152,1436,181,1445]" captionTargetId="figure-23@8.[152,1436,181,1446]" captionTargetPageId="8" captionText="FIGURE 3. Left protarsomeres (A–C), aedeagus in left lateral view (D–G) and dorsal view (H–K) of Trechus (Abyssinotus) species. A, D, I, T. reebae (Quéinnec &amp; Ollivier, 2021), specimen from western crater valley. B, E, J, T. kniphofia Schmidt, sp. n., paratype. C, F, H, T. yitbareki Schmidt, sp. n., paratype. G, K, T. igori Schmidt, sp. n., paratype." figureDoi="http://doi.org/10.5281/zenodo.13234857" httpUri="https://zenodo.org/record/13234857/files/figure.png" pageId="8" pageNumber="351">Fig. 3G</figureCitation>
). Median lobe in dorsal view with side margins almost parallel, shortly narrowed towards apical lamella (
<figureCitation id="A3FFAB95FFDEAC13FF45F8B6FE91F87F" box="[250,338,1900,1927]" captionStart="FIGURE 3" captionStartId="8.[152,255,1470,1494]" captionTargetBox="[152,1436,181,1445]" captionTargetId="figure-23@8.[152,1436,181,1446]" captionTargetPageId="8" captionText="FIGURE 3. Left protarsomeres (A–C), aedeagus in left lateral view (D–G) and dorsal view (H–K) of Trechus (Abyssinotus) species. A, D, I, T. reebae (Quéinnec &amp; Ollivier, 2021), specimen from western crater valley. B, E, J, T. kniphofia Schmidt, sp. n., paratype. C, F, H, T. yitbareki Schmidt, sp. n., paratype. G, K, T. igori Schmidt, sp. n., paratype." figureDoi="http://doi.org/10.5281/zenodo.13234857" httpUri="https://zenodo.org/record/13234857/files/figure.png" pageId="8" pageNumber="351">Fig. 3K</figureCitation>
). Dorsal opening extends basally near to the apical margin of basal bulb. Sagittal aileron absent. Endophallus as described in
<taxonomicName id="FCC4CC93FFDEAC13FE63F84BFDA0F852" authorityName="Schmidt &amp; Merene" authorityYear="2024" box="[476,611,1936,1962]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="351" phylum="Arthropoda" rank="species" species="kniphofia" status="sp. n.">
<emphasis id="09B06B02FFDEAC13FE63F84BFDA0F852" box="[476,611,1936,1962]" italics="true" pageId="8" pageNumber="351">T. kniphofia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1283D679FFDEAC13FDD5F84BFD64F853" box="[618,679,1937,1963]" pageId="8" pageNumber="351" rank="species">sp. n.</taxonomicNameLabel>
Parameres similar in size and shape, slender, each with 3–4 apical setae.
</paragraph>
</subSubSection>
<subSubSection id="73DEE49BFFDFAC12FF78FF4CFAEEFE9D" pageId="9" pageNumber="352" type="diagnosis">
<paragraph id="3B7BB710FFDFAC12FF78FF4CFAEEFE9D" blockId="9.[151,1437,150,537]" pageId="9" pageNumber="352">
<emphasis id="09B06B02FFDFAC12FF78FF4CFE07FF48" bold="true" box="[199,452,150,176]" pageId="9" pageNumber="352">Differential diagnosis</emphasis>
. On average,
<taxonomicName id="FCC4CC93FFDFAC12FDE4FF4DFD68FF48" authorityName="Schmidt &amp; Merene" authorityYear="2024" box="[603,683,151,176]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="352" phylum="Arthropoda" rank="species" species="igori" status="sp. n.">
<emphasis id="09B06B02FFDFAC12FDE4FF4DFD68FF48" box="[603,683,151,176]" italics="true" pageId="9" pageNumber="352">T. igori</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1283D679FFDFAC12FD0EFF4DFD28FF49" box="[689,747,151,177]" pageId="9" pageNumber="352" rank="species">sp. n.</taxonomicNameLabel>
is the smallest species of the
<taxonomicName id="FCC4CC93FFDFAC12FB96FF4DFB4DFF49" baseAuthorityName="Queinnec &amp; Ollivier" baseAuthorityYear="2021" box="[1065,1166,151,177]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="352" phylum="Arthropoda" rank="species" species="reebae">
<emphasis id="09B06B02FFDFAC12FB96FF4DFB4DFF49" box="[1065,1166,151,177]" italics="true" pageId="9" pageNumber="352">T. reebae</emphasis>
</taxonomicName>
group. It differs from all other species of the group by the transverse pattern of elytral microsculpture (isodiametric or somewhat irregularly shaped in the other species) and the very slightly sclerotized apex of the aedeagal median lobe. It additionally differs from
<taxonomicName id="FCC4CC93FFDFAC12FF6AFED9FE92FEE5" box="[213,337,259,285]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="352" phylum="Arthropoda" rank="species" species="niloticus">
<emphasis id="09B06B02FFDFAC12FF6AFED9FE92FEE5" box="[213,337,259,285]" italics="true" pageId="9" pageNumber="352">T. niloticus</emphasis>
</taxonomicName>
,
<taxonomicName id="FCC4CC93FFDFAC12FEDFFED9FE04FEE5" baseAuthorityName="Queinnec &amp; Ollivier" baseAuthorityYear="2021" box="[352,455,259,285]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="352" phylum="Arthropoda" rank="species" species="reebae">
<emphasis id="09B06B02FFDFAC12FEDFFED9FE04FEE5" box="[352,455,259,285]" italics="true" pageId="9" pageNumber="352">T. reebae</emphasis>
</taxonomicName>
and
<taxonomicName id="FCC4CC93FFDFAC12FE41FED9FDBEFEE5" authorityName="Schmidt &amp; Merene" authorityYear="2024" box="[510,637,259,285]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="352" phylum="Arthropoda" rank="species" species="yitbareki" status="sp. n.">
<emphasis id="09B06B02FFDFAC12FE41FED9FDBEFEE5" box="[510,637,259,285]" italics="true" pageId="9" pageNumber="352">T. yitbareki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1283D679FFDFAC12FD3BFED9FD03FEE5" box="[644,704,259,285]" pageId="9" pageNumber="352" rank="species">sp. n.</taxonomicNameLabel>
in the less markedly dilated basal protarsomeres in the male sex. Due to its small body length and shallower elytral striae,
<taxonomicName id="FCC4CC93FFDFAC12FCB3FEFDFC9FFEB8" authorityName="Schmidt &amp; Merene" authorityYear="2024" box="[780,860,295,320]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="352" phylum="Arthropoda" rank="species" species="igori" status="sp. n.">
<emphasis id="09B06B02FFDFAC12FCB3FEFDFC9FFEB8" box="[780,860,295,320]" italics="true" pageId="9" pageNumber="352">T. igori</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1283D679FFDFAC12FCDCFEFDFC5EFEB9" box="[867,925,295,321]" pageId="9" pageNumber="352" rank="species">sp. n.</taxonomicNameLabel>
is similar to
<taxonomicName id="FCC4CC93FFDFAC12FB94FEFDFB05FEB9" box="[1067,1222,295,321]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="352" phylum="Arthropoda" rank="species" species="abyssinicus">
<emphasis id="09B06B02FFDFAC12FB94FEFDFB05FEB9" box="[1067,1222,295,321]" italics="true" pageId="9" pageNumber="352">T. abyssinicus</emphasis>
</taxonomicName>
and
<taxonomicName id="FCC4CC93FFDFAC12FB44FEFDFA56FEB9" box="[1275,1429,295,321]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="352" phylum="Arthropoda" rank="species" species="afroalpinus">
<emphasis id="09B06B02FFDFAC12FB44FEFDFA56FEB9" box="[1275,1429,295,321]" italics="true" pageId="9" pageNumber="352">T. afroalpinus</emphasis>
</taxonomicName>
, however, it is easily distinguished by two basal protarsomeres dilated in males (one in the latter species).
</paragraph>
</subSubSection>
<subSubSection id="73DEE49BFFDFAC12FF78FEB4FE37FDE1" pageId="9" pageNumber="352" type="distribution">
<paragraph id="3B7BB710FFDFAC12FF78FEB4FE5DFE29" blockId="9.[151,1437,150,537]" pageId="9" pageNumber="352">
<emphasis id="09B06B02FFDFAC12FF78FEB4FE9AFE70" bold="true" box="[199,345,366,392]" pageId="9" pageNumber="352">Distribution</emphasis>
. Endemic to Mt. Choke in northern
<collectingCountry id="43D3F780FFDFAC12FD46FEB5FC9FFE71" box="[761,860,367,393]" name="Ethiopia" pageId="9" pageNumber="352">Ethiopia</collectingCountry>
. The new species was only found near the top of a mountain ridge that extends from the crater rim to the west were it occurs sympatric with
<taxonomicName id="FCC4CC93FFDFAC12FB27FE49FACAFE54" authorityName="Schmidt &amp; Merene" authorityYear="2024" box="[1176,1289,403,428]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="352" phylum="Arthropoda" rank="species" species="apertus" status="sp. n.">
<emphasis id="09B06B02FFDFAC12FB27FE49FACAFE54" box="[1176,1289,403,428]" italics="true" pageId="9" pageNumber="352">T. apertus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1283D679FFDFAC12FAADFE49FA8CFE55" box="[1298,1359,403,429]" pageId="9" pageNumber="352" rank="species">sp. n.</taxonomicNameLabel>
and
<taxonomicName id="FCC4CC93FFDFAC12FA37FE49FF39FE29" authorityName="Schmidt &amp; Merene" authorityYear="2024" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="352" phylum="Arthropoda" rank="species" species="yitbareki" status="sp. n.">
<emphasis id="09B06B02FFDFAC12FA37FE49FF39FE29" italics="true" pageId="9" pageNumber="352">T. yitbareki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1283D679FFDFAC12FEBEFE6DFEFEFE29" box="[257,317,439,465]" pageId="9" pageNumber="352" rank="species">sp. n.</taxonomicNameLabel>
(
<figureCitation id="A3FFAB95FFDFAC12FEF3FE6DFE52FE29" box="[332,401,439,465]" captionStart="FIGURE 4" captionStartId="9.[152,255,1559,1583]" captionTargetBox="[151,1436,561,1535]" captionTargetId="figure-219@9.[151,1436,561,1535]" captionTargetPageId="9" captionText="FIGURE 4. Topographic map of Mt. Choke (highest point marked by a white triangle), showing sampling localities of six Trechus (Abyssinotus) species of the niloticus group (white circles; the numbers refer to the different species). 1, T. apertus Schmidt, sp. n. 2, T. reebae (Quéinnec &amp; Ollivier). 3, T. yitbareki Schmidt, sp. n. 4, T. kniphofia Schmidt, sp. n. 5, T. igori Schmidt, sp. n. 6, T. niloticus (Quéinnec &amp; Ollivier). Sampling data of T. reebae and T. niloticus published by Quéinnec et al. (2021) are considered. Base map was downloaded from Topographic-map.com." figureDoi="http://doi.org/10.5281/zenodo.13234861" httpUri="https://zenodo.org/record/13234861/files/figure.png" pageId="9" pageNumber="352">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="3B7BB710FFDFAC12FF78FE00FE37FDE1" blockId="9.[151,1437,150,537]" pageId="9" pageNumber="352">
<emphasis id="09B06B02FFDFAC12FF78FE00FE76FE0C" bold="true" box="[199,437,474,500]" pageId="9" pageNumber="352">Habitat preferences</emphasis>
. The new species was found syntopic with
<taxonomicName id="FCC4CC93FFDFAC12FC12FE01FBDCFE0C" authorityName="Schmidt &amp; Merene" authorityYear="2024" box="[941,1055,475,500]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="352" phylum="Arthropoda" rank="species" species="apertus" status="sp. n.">
<emphasis id="09B06B02FFDFAC12FC12FE01FBDCFE0C" box="[941,1055,475,500]" italics="true" pageId="9" pageNumber="352">T. apertus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1283D679FFDFAC12FB97FE01FBA5FE0D" box="[1064,1126,475,501]" pageId="9" pageNumber="352" rank="species">sp. n.</taxonomicNameLabel>
and
<taxonomicName id="FCC4CC93FFDFAC12FB1FFE01FAE2FE0D" authorityName="Schmidt &amp; Merene" authorityYear="2024" box="[1184,1313,475,501]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="352" phylum="Arthropoda" rank="species" species="yitbareki" status="sp. n.">
<emphasis id="09B06B02FFDFAC12FB1FFE01FAE2FE0D" box="[1184,1313,475,501]" italics="true" pageId="9" pageNumber="352">T. yitbareki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1283D679FFDFAC12FA95FE01FAAAFE0D" box="[1322,1385,475,501]" pageId="9" pageNumber="352" rank="species">sp. n.</taxonomicNameLabel>
; see above for details of the habitat.
</paragraph>
</subSubSection>
</treatment>
</document>
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