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<document id="F11CBE96D82E3373FC68402D629BEB8E" ID-DOI="10.11646/phytotaxa.640.1.7" ID-ISSN="1179-3163" ID-Zenodo-Dep="13213500" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722601880092" checkinUser="felipe" docAuthor="Blasco, Freddie A., Tandang, Danilo N. &amp; Alejandro, Grecebio Jonathan D." docDate="2024" docId="9C6D9E66FFDCFFDBB3B57A48627FFEC5" docLanguage="en" docName="phytotaxa.640.1.7.pdf" docOrigin="Phytotaxa 640 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.640.1.7" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Argostemma mirae Blasco, Tandang &amp; Alejandro, sp. nov." docType="treatment" docVersion="3" lastPageNumber="69" masterDocId="6054E61EFFDDFFDFB33D7E7C6722FFE1" masterDocTitle="Argostemma mirae (Rubiaceae) a new species from Tago, Surigao del Sur, Mindanao Island, Philippines" masterLastPageNumber="70" masterPageNumber="65" pageNumber="66" updateTime="1722803603210" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="8216D5C40F7076144CBABB1FC84CBB11" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A257677BF95447317BCBBEE731A87DDB">
<mods:title id="48BF901FD03D60BBCF8E19620AD5EAA0">Argostemma mirae (Rubiaceae) a new species from Tago, Surigao del Sur, Mindanao Island, Philippines</mods:title>
</mods:titleInfo>
<mods:name id="848762560A93706BA229320031D393CF" type="personal">
<mods:role id="45B1A991A3CB46BA03C56E677E1C7466">
<mods:roleTerm id="81885A8CA971BB56123CE234E66B6BB7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A1B871DF9A10BC490C39A29B8EACF6F4">Blasco, Freddie A.</mods:namePart>
<mods:affiliation id="ACE01C1A3E10AA5B69D7A603864AB8A7">College of Arts &amp; Sciences Department, Saint Theresa College of Tandag, Tandag City 8300, Surigao del Sur, Philippines</mods:affiliation>
</mods:name>
<mods:name id="DF9E460AC5053A4FD0DB4ED705798766" type="personal">
<mods:role id="7D7A1F171BF290148F05A018861CCE60">
<mods:roleTerm id="792FAAE7A3802E09612EA1892B61039C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="342F3AAAA72E9FDEB21CA866894ED90C">Tandang, Danilo N.</mods:namePart>
<mods:affiliation id="FD7A89B738F67DC72843110803C396BD">Philippine National Herbarium, Botany and National Herbarium Division, National Museum of Natural History, National Museum of the Philippines, T. M. Kalaw St., Manila 1000, Philippines &amp; Biodiversity Program, Taiwan International Graduate Program, Academia Sinica and National Taiwan Normal University, Taipei, 11529, Taiwan &amp; Department of Life Science, National Taiwan Normal University, Taipei 11677, Taiwan &amp; Biodiversity Research Center, Academia Sinica, Taipei 11529, Taiwan</mods:affiliation>
</mods:name>
<mods:name id="280644765EF981FA4DE574F4AAA4BDD3" type="personal">
<mods:role id="895EF3A6D318EFB65B3E901DAF4BAD15">
<mods:roleTerm id="F142A17CEEA07847AEBAD7684D67813B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B3541146BE611FC54B5639C247D7D43F">Alejandro, Grecebio Jonathan D.</mods:namePart>
<mods:affiliation id="AC310D6AA5E1608BC9AFA43BE8475063">College of Science and Research Center for the Natural and Applied Science, University of Santo Tomas, España, Manila 1015, Philippines</mods:affiliation>
</mods:name>
<mods:typeOfResource id="437D22FF6A7DF1988463A915C68344AE">text</mods:typeOfResource>
<mods:relatedItem id="CCBC1D64F9A621586CBF18A67AD58A11" type="host">
<mods:titleInfo id="50FEB770BF98AB93E0EA727FC4DB117E">
<mods:title id="3BC24341EE284E7810FAD4429EA76491">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="9D917CBEA59327FAFCAF730B2A293BA4">
<mods:date id="B72D56398B7011FED7B9BB5E651A0D6A">2024</mods:date>
<mods:detail id="486451E7FE1BC91DAEC9EAF915F72567" type="pubDate">
<mods:number id="ADD4E608BD48A6D97BD5570672C35FE6">2024-03-07</mods:number>
</mods:detail>
<mods:detail id="313C7CF792A01DA4743F987D98A1233A" type="volume">
<mods:number id="101EDD843F7FF20B3FE387BF09758F74">640</mods:number>
</mods:detail>
<mods:detail id="A4D62B3FB73B61E6F42247B50B1D26DC" type="issue">
<mods:number id="CE6539AB77A0076722759FF8F1FB62D0">1</mods:number>
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<mods:start id="FE25DA235D05A6EEE15C3B2E7667DD0F">65</mods:start>
<mods:end id="935FB9D7C1C0CB0AE4F89D86BEC9FC4B">70</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="DEE9B029D051FFB694ED66C5DE117175">
<mods:url id="8DEAAA297761B44674887E77FA8CBD18">http://dx.doi.org/10.11646/phytotaxa.640.1.7</mods:url>
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<mods:classification id="37FBD0FB034C456FEC54F1941C331F23">journal article</mods:classification>
<mods:identifier id="7C1713C424837431C697FA21B4C2C04C" type="DOI">10.11646/phytotaxa.640.1.7</mods:identifier>
<mods:identifier id="C7DE30B6D224E57153DBF725A22C1196" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="3FB2E5865EB4478E453615C73D7D58F4" type="Zenodo-Dep">13213500</mods:identifier>
</mods:mods>
<treatment id="9C6D9E66FFDCFFDBB3B57A48627FFEC5" ID-DOI="http://doi.org/10.5281/zenodo.13213078" ID-Zenodo-Dep="13213078" LSID="urn:lsid:plazi:treatment:9C6D9E66FFDCFFDBB3B57A48627FFEC5" httpUri="http://treatment.plazi.org/id/9C6D9E66FFDCFFDBB3B57A48627FFEC5" lastPageId="4" lastPageNumber="69" pageId="1" pageNumber="66">
<subSubSection id="5CDE7CFBFFDCFFDEB3B57A4864B2FBAF" box="[136,912,1076,1103]" pageId="1" pageNumber="66" type="nomenclature">
<paragraph id="147B2F70FFDCFFDEB3B57A4864B2FBAF" blockId="1.[136,912,1076,1103]" box="[136,912,1076,1103]" pageId="1" pageNumber="66">
<heading id="4F33981CFFDCFFDEB3B57A4864B2FBAF" box="[136,912,1076,1103]" fontSize="11" level="2" pageId="1" pageNumber="66" reason="3">
<taxonomicName id="D3C454F3FFDCFFDEB3B57A4865EEFBAF" authority="Blasco, Tandang &amp; Alejandro" authorityName="Blasco, Tandang &amp; Alejandro" box="[136,716,1076,1102]" class="Magnoliopsida" family="Rubiaceae" genus="Argostemma" kingdom="Plantae" order="Gentianales" pageId="1" pageNumber="66" phylum="Tracheophyta" rank="species" species="mirae" status="sp. nov.">
<emphasis id="26B0F362FFDCFFDEB3B57A4865EEFBAF" bold="true" box="[136,716,1076,1102]" pageId="1" pageNumber="66">
<emphasis id="26B0F362FFDCFFDEB3B57A486646FBAF" bold="true" box="[136,356,1076,1102]" italics="true" pageId="1" pageNumber="66">Argostemma mirae</emphasis>
Blasco, Tandang &amp; Alejandro
</emphasis>
</taxonomicName>
,
<emphasis id="26B0F362FFDCFFDEB1E77A49640DFBAF" box="[730,815,1077,1102]" italics="true" pageId="1" pageNumber="66">
<taxonomicNameLabel id="3D834E19FFDCFFDEB1E77A49640DFBAF" box="[730,815,1077,1102]" pageId="1" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="8CFF33F5FFDCFFDEB0007A48645DFBAF" box="[829,895,1076,1102]" captionStart="FIGURE 1" captionStartId="2.[136,229,1659,1681]" captionTargetBox="[156,1431,192,1626]" captionTargetId="figure-25@2.[151,1436,187,1632]" captionTargetPageId="2" captionText="FIGURE 1. Argostemma mirae Blasco, Tandang &amp; Alejandro. A. Habit &amp; Habitat; B. Young flower; C. Matured flower; D. Fruit." figureDoi="http://doi.org/10.5281/zenodo.13213502" httpUri="https://zenodo.org/record/13213502/files/figure.png" pageId="1" pageNumber="66">Fig. 1</figureCitation>
).
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5CDE7CFBFFDCFFDEB3B57AFC63FDFB58" pageId="1" pageNumber="66" type="materials_examined">
<paragraph id="147B2F70FFDCFFDEB3B57AFC63FDFB58" blockId="1.[136,1451,1152,1210]" pageId="1" pageNumber="66">
<emphasis id="26B0F362FFDCFFDEB3B57AFC67F0FB74" bold="true" box="[136,210,1152,1173]" pageId="1" pageNumber="66">Type—</emphasis>
<collectingCountry id="6CD36FE0FFDCFFDEB3EF7AFC6641FB74" box="[210,355,1152,1173]" name="Philippines" pageId="1" pageNumber="66">PHILIPPINES</collectingCountry>
. Mindanao Island,
<collectingRegion id="D600E192FFDCFFDEB11E7AFC65E2FB74" box="[547,704,1152,1174]" country="Philippines" name="Surigao del Sur" pageId="1" pageNumber="66">Surigao del Sur</collectingRegion>
, Tago, Badong, Mt. Tingtingan, elev.
<emphasis id="26B0F362FFDCFFDEB7007AFC6378FB74" box="[1085,1114,1152,1173]" italics="true" pageId="1" pageNumber="66">ca.</emphasis>
<specimenCount id="02C2E4F9FFDCFFDEB75E7AFC63B6FB74" box="[1123,1172,1152,1173]" count="10" pageId="1" pageNumber="66" type="generic">
<quantity id="D33C8295FFDCFFDEB75E7AFC63B2FB74" box="[1123,1168,1152,1173]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="66" unit="m" value="10.0">10m</quantity>
.
</specimenCount>
It grows on a shady mossy limestone forest on a lower elevation.
<date id="607A09B0FFDCFFDEB1107AD8658FFB58" box="[557,685,1188,1209]" pageId="1" pageNumber="66" value="2022-05-22">22 May 2022</date>
<emphasis id="26B0F362FFDCFFDEB18E7AD8648BFB58" box="[691,937,1188,1209]" italics="true" pageId="1" pageNumber="66">
Freddie
<collectionCode id="72D5B7B5FFDCFFDEB03E7AD86437FB58" box="[771,789,1188,1209]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="1" pageNumber="66" type="Herbarium">A</collectionCode>
. Blasco 22-002
</emphasis>
(
<typeStatus id="CB7F91D2FFDCFFDEB08B7AD8632EFB5B" box="[950,1036,1188,1210]" pageId="1" pageNumber="66" type="holotype">holotype</typeStatus>
<collectionCode id="72D5B7B5FFDCFFDEB72C7AD86361FB58" box="[1041,1091,1188,1209]" country="Philippines" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15314" lsid="urn:lsid:biocol.org:col:15314" name="National Museum" pageId="1" pageNumber="66" type="Herbarium">PNH</collectionCode>
,
<typeStatus id="CB7F91D2FFDCFFDEB7707AD863B0FB5B" box="[1101,1170,1188,1210]" pageId="1" pageNumber="66" type="isotype">isotype</typeStatus>
<collectionCode id="72D5B7B5FFDCFFDEB7A47AD863FBFB58" box="[1177,1241,1188,1209]" pageId="1" pageNumber="66">USTH</collectionCode>
)
</paragraph>
</subSubSection>
<subSubSection id="5CDE7CFBFFDCFFDEB3B57A9463DBF84F" pageId="1" pageNumber="66" type="diagnosis">
<paragraph id="147B2F70FFDCFFDEB3B57A946204FA1F" blockId="1.[136,1452,1256,2075]" pageId="1" pageNumber="66">
<emphasis id="26B0F362FFDCFFDEB3B57A9467D9FAE3" bold="true" box="[136,251,1256,1282]" pageId="1" pageNumber="66">Diagnosis</emphasis>
—
<taxonomicName id="D3C454F3FFDCFFDEB22A7A9466C8FAE3" box="[279,490,1256,1282]" class="Magnoliopsida" family="Rubiaceae" genus="Argostemma" kingdom="Plantae" order="Gentianales" pageId="1" pageNumber="66" phylum="Tracheophyta" rank="species" species="mirae">
<emphasis id="26B0F362FFDCFFDEB22A7A9466C8FAE3" box="[279,490,1256,1282]" italics="true" pageId="1" pageNumber="66">Argostemma mirae</emphasis>
</taxonomicName>
closely resembles
<taxonomicName id="D3C454F3FFDCFFDEB1867A946490FAE3" authority="Elmer" authorityName="Elmer" authorityYear="1906" box="[699,946,1256,1282]" class="Magnoliopsida" family="Rubiaceae" genus="Argostemma" kingdom="Plantae" order="Gentianales" pageId="1" pageNumber="66" phylum="Tracheophyta" rank="species" species="solaniflorum">
<emphasis id="26B0F362FFDCFFDEB1867A94644AFAE3" box="[699,872,1256,1282]" italics="true" pageId="1" pageNumber="66">A. solaniflorum</emphasis>
Elmer
</taxonomicName>
over
<taxonomicName id="D3C454F3FFDCFFDEB0D27A9463CEFAE3" authority="Merrill" authorityName="Merrill" authorityYear="1922" box="[1007,1260,1256,1282]" class="Magnoliopsida" family="Rubiaceae" genus="Argostemma" kingdom="Plantae" order="Gentianales" pageId="1" pageNumber="66" phylum="Tracheophyta" rank="species" species="arachnosum">
<emphasis id="26B0F362FFDCFFDEB0D27A9463B5FAE3" box="[1007,1175,1256,1282]" italics="true" pageId="1" pageNumber="66">A. arachnosum</emphasis>
Merrill
</taxonomicName>
and
<taxonomicName id="D3C454F3FFDCFFDEB61C7A9467FCFAC7" authority="Valeton" authorityName="Valeton" authorityYear="1911" class="Magnoliopsida" family="Rubiaceae" genus="Argostemma" kingdom="Plantae" order="Gentianales" pageId="1" pageNumber="66" phylum="Tracheophyta" rank="species" species="distichum">
<emphasis id="26B0F362FFDCFFDEB61C7A946289FAE3" box="[1313,1451,1256,1282]" italics="true" pageId="1" pageNumber="66">A. distichum</emphasis>
Valeton
</taxonomicName>
in having erect, branching and pubescent stems, glabrous stipules, pubescent petioles, oblanceolate leaves, dark green lamina, abaxially pubescent leaf surface, acute base, margin entire, pubescent calyx lobes, corolla lobes surface abaxially pubescent, white corollas and pubescent pedicel but differs in having shorter stems
<quantity id="D33C8295FFDCFFDEB63E7B286255FA8E" box="[1283,1399,1364,1391]" metricMagnitude="-1" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="1" pageNumber="66" unit="cm" value="11.5" valueMax="13.0" valueMin="10.0">10–13 cm</quantity>
. (vs. 16), stipules broadly ovate (vs. oblong), smaller leaf blades 2.5–3. ×
<quantity id="D33C8295FFDCFFDEB0427B0464CEFA72" box="[895,1004,1400,1427]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="1" pageNumber="66" unit="cm" value="1.15" valueMax="1.3" valueMin="1.0">1– 1.3 cm</quantity>
(vs. 7 × 2), acuminate apex (vs. obtuse), adaxially glabrous leaf surface (vs. pubescent), lateral leaf veins 4–5 pairs (vs. 7–8), inflorescences 2 (vs. 5), 4 calyx lobes (vs. 5), 4 corolla lobes (vs. 5 star-shaped), corolla lobes surface adaxially glabrous (vs. pubescent), longer pedicel
<quantity id="D33C8295FFDCFFDEB3B57B986623FA1E" box="[136,257,1508,1535]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="1" pageNumber="66" unit="mm" value="16.0" valueMax="17.0" valueMin="15.0">15–17 mm</quantity>
(vs. short 8), anthers length
<quantity id="D33C8295FFDCFFDEB17C7B9865BCFA1E" box="[577,670,1508,1535]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="1" pageNumber="66" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
(vs. 7) and opening by longitudinal slits (vs. apical pore).
</paragraph>
<paragraph id="147B2F70FFDCFFDEB380787463E6F86B" blockId="1.[136,1452,1256,2075]" pageId="1" pageNumber="66">
Lithophytic herb,
<emphasis id="26B0F362FFDCFFDEB2B7787466E5F9C3" bold="true" box="[394,455,1544,1570]" pageId="1" pageNumber="66">Stem</emphasis>
,
<quantity id="D33C8295FFDCFFDEB2EB78746564F9C2" box="[470,582,1544,1571]" metricMagnitude="-1" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="1" pageNumber="66" unit="cm" value="11.5" valueMax="13.0" valueMin="10.0">10–13 cm</quantity>
tall,
<quantity id="D33C8295FFDCFFDEB1BF787465D4F9C2" box="[642,758,1544,1571]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="1" pageNumber="66" unit="mm" value="2.75" valueMax="3.0" valueMin="2.5">2.5–3 mm</quantity>
in diameter, pubescent, erect and branching,
<emphasis id="26B0F362FFDCFFDEB7C67874625BF9C3" bold="true" box="[1275,1401,1544,1570]" pageId="1" pageNumber="66">internodes</emphasis>
<quantity id="D33C8295FFDCFFDEB6BC78746796F9A6" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="1" pageNumber="66" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5–7 mm</quantity>
long, terete.
<emphasis id="26B0F362FFDCFFDEB27B78506687F9A7" bold="true" box="[326,421,1580,1606]" pageId="1" pageNumber="66">Stipules</emphasis>
persistent, broadly ovate, 5–6 ×
<quantity id="D33C8295FFDCFFDEB02978506451F9A6" box="[788,883,1580,1607]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="1" pageNumber="66" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
, glabrous,
<emphasis id="26B0F362FFDCFFDEB0D078506307F9A7" bold="true" box="[1005,1061,1580,1606]" pageId="1" pageNumber="66">apex</emphasis>
acuminate,
<emphasis id="26B0F362FFDCFFDEB79078506226F9A7" bold="true" box="[1197,1284,1580,1606]" pageId="1" pageNumber="66">margin</emphasis>
entire.
<emphasis id="26B0F362FFDCFFDEB66478506289F9A7" bold="true" box="[1369,1451,1580,1606]" pageId="1" pageNumber="66">Leaves</emphasis>
glabrous, glossy, oblanceolate,
<emphasis id="26B0F362FFDCFFDEB2D0782C651EF98B" bold="true" box="[493,572,1616,1642]" pageId="1" pageNumber="66">petiole</emphasis>
<quantity id="D33C8295FFDCFFDEB17B782C6584F98A" box="[582,678,1616,1643]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="1" pageNumber="66" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
long, terete, pubescent,
<emphasis id="26B0F362FFDCFFDEB0FC782C6320F98B" bold="true" box="[961,1026,1616,1642]" pageId="1" pageNumber="66">blade</emphasis>
2.5–3. ×
<quantity id="D33C8295FFDCFFDEB749782C63C0F98A" box="[1140,1250,1616,1643]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="1" pageNumber="66" unit="cm" value="1.15" valueMax="1.3" valueMin="1.0">1–1.3 cm</quantity>
,
<emphasis id="26B0F362FFDCFFDEB7CD782C620AF98B" bold="true" box="[1264,1320,1616,1642]" pageId="1" pageNumber="66">apex</emphasis>
acuminate,
<emphasis id="26B0F362FFDCFFDEB3B5780867FDF96F" bold="true" box="[136,223,1652,1678]" pageId="1" pageNumber="66">margin</emphasis>
entire, obliquely
<emphasis id="26B0F362FFDCFFDEB29478086505F96F" bold="true" box="[425,551,1652,1678]" pageId="1" pageNumber="66">acute base</emphasis>
, adaxial surface glossy, glabrous, dark green with prominent midrib and veins, abaxial surface light green and hairy with distinct pubescent midrib, lateral veins 4–5 pairs.
<emphasis id="26B0F362FFDCFFDEB79A78E46267F953" bold="true" box="[1191,1349,1688,1714]" pageId="1" pageNumber="66">Inflorescence</emphasis>
terminal to axillary, cymose, 2–flowered, peduncle pilose,
<quantity id="D33C8295FFDCFFDEB1FD78C0640DF936" box="[704,815,1724,1751]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="1" pageNumber="66" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">8–10 mm</quantity>
,
<emphasis id="26B0F362FFDCFFDEB00178C064A5F937" bold="true" box="[828,903,1724,1750]" pageId="1" pageNumber="66">bracts</emphasis>
3–4 ×
<quantity id="D33C8295FFDCFFDEB0E678C06318F936" box="[987,1082,1724,1751]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="1" pageNumber="66" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
persistent and pubescent, ovate, pedicel
<quantity id="D33C8295FFDCFFDEB3E2789C6674F91A" box="[223,342,1760,1787]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="1" pageNumber="66" unit="mm" value="16.0" valueMax="17.0" valueMin="15.0">15–17 mm</quantity>
long.
<emphasis id="26B0F362FFDCFFDEB2A7789C66C2F91B" bold="true" box="[410,480,1760,1786]" pageId="1" pageNumber="66">Calyx</emphasis>
lobes 4, triangular to narrowly triangular, gamosepalous, hypanthium, infundibuliform, pubescent.
<emphasis id="26B0F362FFDCFFDEB2387978667DF8FF" bold="true" box="[261,351,1796,1822]" pageId="1" pageNumber="66">Corolla</emphasis>
lobes 4, white, adaxially glabrous, abaxially pubescent 6–7 ×
<quantity id="D33C8295FFDCFFDEB72E79786353F8FE" box="[1043,1137,1796,1823]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="1" pageNumber="66" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3–5 mm</quantity>
, broadly ovate when young, narrowly triangular when mature, apex acuminate.
<emphasis id="26B0F362FFDCFFDEB1FF79546405F8A3" bold="true" box="[706,807,1832,1858]" pageId="1" pageNumber="66">Stamens</emphasis>
4,
<quantity id="D33C8295FFDCFFDEB07479546484F8A2" box="[841,934,1832,1859]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="1" pageNumber="66" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5–6 mm</quantity>
long, filaments short, anthers yellow,
<quantity id="D33C8295FFDCFFDEB6727954628EF8A2" box="[1359,1452,1832,1859]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="1" pageNumber="66" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
long.
<emphasis id="26B0F362FFDCFFDEB3F779306634F887" bold="true" box="[202,278,1868,1894]" pageId="1" pageNumber="66">Ovary</emphasis>
inferior, bilocular, ovules numerous, style
<quantity id="D33C8295FFDCFFDEB03C79306442F886" box="[769,864,1868,1895]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="1" pageNumber="66" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
long, stigma
<quantity id="D33C8295FFDCFFDEB0C07930637EF886" box="[1021,1116,1868,1895]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="1" pageNumber="66" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
long, simple, exserted.
<emphasis id="26B0F362FFDCFFDEB65079306289F887" bold="true" box="[1389,1451,1868,1894]" pageId="1" pageNumber="66">Fruit</emphasis>
obovoid, 5–6 ×
<quantity id="D33C8295FFDCFFDEB207790C66BBF86A" box="[314,409,1904,1931]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="1" pageNumber="66" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
, pubescent, crowned by a persistent calyx, opening by longitudinal slits.
</paragraph>
<paragraph id="147B2F70FFDCFFDEB38079E863DBF84F" blockId="1.[136,1452,1256,2075]" box="[189,1273,1940,1967]" pageId="1" pageNumber="66">
<emphasis id="26B0F362FFDCFFDEB38079E8661BF84F" bold="true" box="[189,313,1940,1966]" pageId="1" pageNumber="66">Phenology</emphasis>
—Observed flowering and fruiting in April to May when the dry season is at its peak.
</paragraph>
</subSubSection>
<subSubSection id="5CDE7CFBFFDCFFDEB38079C46562F7FB" pageId="1" pageNumber="66" type="etymology">
<paragraph id="147B2F70FFDCFFDEB38079C46562F7FB" blockId="1.[136,1452,1256,2075]" pageId="1" pageNumber="66">
<emphasis id="26B0F362FFDCFFDEB38079C4661CF833" bold="true" box="[189,318,1976,2002]" pageId="1" pageNumber="66">Etymology</emphasis>
—The species is named after Mrs. Evangeline A. Mira to honor her 45 years of service to Saint Theresa College of Tandag. Ma’am Mira, as she is fondly called by her students, colleagues and friends is a biology and language teacher and a dear friend.
</paragraph>
</subSubSection>
<caption id="40BB7FF8FFDFFFDDB3B5780762A0F970" ID-DOI="http://doi.org/10.5281/zenodo.13213502" ID-Zenodo-Dep="13213502" box="[136,1410,1659,1681]" httpUri="https://zenodo.org/record/13213502/files/figure.png" pageId="2" pageNumber="67" startId="2.[136,229,1659,1681]" targetBox="[156,1431,192,1626]" targetPageId="2" targetType="figure">
<paragraph id="147B2F70FFDFFFDDB3B5780762A0F970" blockId="2.[136,1410,1659,1681]" box="[136,1410,1659,1681]" pageId="2" pageNumber="67">
<emphasis id="26B0F362FFDFFFDDB3B5780767DCF971" bold="true" box="[136,254,1659,1681]" pageId="2" pageNumber="67">FIGURE 1.</emphasis>
<taxonomicName id="D3C454F3FFDFFFDDB23E7807649EF970" authority="Blasco, Tandang &amp; Alejandro. A. Habit &amp; Habitat" box="[259,956,1659,1681]" class="Magnoliopsida" family="Rubiaceae" genus="Argostemma" kingdom="Plantae" order="Gentianales" pageId="2" pageNumber="67" phylum="Tracheophyta" rank="species" species="mirae">
<emphasis id="26B0F362FFDFFFDDB23E780765D8F970" bold="true" box="[259,762,1659,1681]" pageId="2" pageNumber="67">
<emphasis id="26B0F362FFDFFFDDB23E780766E2F971" bold="true" box="[259,448,1659,1680]" italics="true" pageId="2" pageNumber="67">Argostemma mirae</emphasis>
Blasco, Tandang &amp; Alejandro.
</emphasis>
A. Habit &amp; Habitat
</taxonomicName>
; B. Young flower; C. Matured flower; D. Fruit.
</paragraph>
</caption>
<subSubSection id="5CDE7CFBFFDFFFDBB38078C3627FFEC5" lastPageId="4" lastPageNumber="69" pageId="2" pageNumber="67" type="distribution">
<paragraph id="147B2F70FFDFFFDDB38078C366A4F8C3" blockId="2.[136,1452,1727,2078]" pageId="2" pageNumber="67">
<emphasis id="26B0F362FFDFFFDDB38078C366CEF93B" bold="true" box="[189,492,1727,1754]" pageId="2" pageNumber="67">Distribution and Ecology</emphasis>
—Endemic to the province of
<collectingRegion id="D600E192FFDFFFDDB07978BC64D4F93B" box="[836,1014,1728,1754]" country="Philippines" name="Surigao del Sur" pageId="2" pageNumber="67">Surigao del Sur</collectingRegion>
, CARAGA Region Eastern Mindanao,
<collectingCountry id="6CD36FE0FFDFFFDDB3B578986628F91F" box="[136,266,1764,1790]" name="Philippines" pageId="2" pageNumber="67">Philippines</collectingCountry>
. It grows on a shady, mossy limestone forest on a lower elevation in Mt. Tingtingan, Badong, Tago,
<collectingRegion id="D600E192FFDFFFDDB669789867F8F8C3" country="Philippines" name="Surigao del Sur" pageId="2" pageNumber="67">Surigao del Sur</collectingRegion>
, elev.
<quantity id="D33C8295FFDFFFDDB21D7974667BF8C3" box="[288,345,1800,1826]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="67" unit="m" value="10.0">10 m</quantity>
asl.
</paragraph>
<paragraph id="147B2F70FFDFFFDCB380795063A0FF3D" blockId="2.[136,1452,1727,2078]" lastBlockId="3.[136,1451,158,221]" lastPageId="3" lastPageNumber="68" pageId="2" pageNumber="67">
<emphasis id="26B0F362FFDFFFDDB3807950663EF8A7" bold="true" box="[189,284,1836,1862]" pageId="2" pageNumber="67">Notes—</emphasis>
<taxonomicName id="D3C454F3FFDFFFDDB221795066D7F8A4" box="[284,501,1836,1862]" class="Magnoliopsida" family="Rubiaceae" genus="Argostemma" kingdom="Plantae" order="Gentianales" pageId="2" pageNumber="67" phylum="Tracheophyta" rank="species" species="mirae">
<emphasis id="26B0F362FFDFFFDDB221795066D7F8A4" box="[284,501,1836,1862]" italics="true" pageId="2" pageNumber="67">Argostemma mirae</emphasis>
</taxonomicName>
superficially resembles
<taxonomicName id="D3C454F3FFDFFFDDB02E79506327F8A7" authority="Valeton" authorityName="Valeton" authorityYear="1911" box="[787,1029,1836,1862]" class="Magnoliopsida" family="Rubiaceae" genus="Argostemma" kingdom="Plantae" order="Gentianales" pageId="2" pageNumber="67" phylum="Tracheophyta" rank="species" species="distichum">
<emphasis id="26B0F362FFDFFFDDB02E79506486F8A7" box="[787,932,1836,1862]" italics="true" pageId="2" pageNumber="67">A. distichum</emphasis>
Valeton
</taxonomicName>
in having branching stems, broadly ovate stipules, dark green lamina, entire margin, 4 glabrous white corollas, and longitudinal slits dehiscence. However, the new species is distinct in having erect and longer stems (vs. creeping), larger leaves, acuminate apex (vs. sharplyacute), numerous major leaf veins 4–5 (vs. 1–2), longer pedicel and anthers, and 2-flowered inflorescence (vs. multi-flowered 5–6). It is also comparable to
<taxonomicName id="D3C454F3FFDFFFDDB10779C0641AF837" authority="Merrill" authorityName="Merrill" authorityYear="1922" box="[570,824,1980,2006]" class="Magnoliopsida" family="Rubiaceae" genus="Argostemma" kingdom="Plantae" order="Gentianales" pageId="2" pageNumber="67" phylum="Tracheophyta" rank="species" species="arachnosum">
<emphasis id="26B0F362FFDFFFDDB10779C065C0F837" box="[570,738,1980,2006]" italics="true" pageId="2" pageNumber="67">A. arachnosum</emphasis>
Merrill
</taxonomicName>
in having pubescent stems, glabrous stipules, acuminate apex, pubescent calyx lobes, white corollas, and longitudinal slits dehiscence, but differs in having shorter erect stems (vs. creeping, solitary), dark green lamina (vs. light green with green sparse), acute leaf base (vs. cordate-rounded) longer pedicel
<quantity id="D33C8295FFDEFFDCB2127EE2668BFF58" box="[303,425,158,185]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="3" pageNumber="68" unit="mm" value="16.0" valueMax="17.0" valueMin="15.0">15–17 mm</quantity>
(vs.
<quantity id="D33C8295FFDEFFDCB2DC7EE26542FF59" box="[481,608,158,184]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="3" pageNumber="68" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">10–12 mm</quantity>
.), 2-flowered inflorescence (vs. multi-flowered 10–25), and 4 corolla lobes (vs. 5 star-shaped).
<tableCitation id="59461ACBFFDEFFDCB25E7EBE6694FF3D" box="[355,438,194,220]" captionStart="TABLE 1" captionStartId="3.[136,224,1133,1157]" captionText="TABLE 1. Comparative morphology among Argostemma mirae, A. arachnosum, A. distichum and A. solaniflorum." pageId="3" pageNumber="68">Table 1</tableCitation>
offers a thorough comparison of
<taxonomicName id="D3C454F3FFDEFFDCB0117EBF64AEFF3D" box="[812,908,195,221]" class="Magnoliopsida" family="Rubiaceae" genus="Argostemma" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="68" phylum="Tracheophyta" rank="species" species="mirae">
<emphasis id="26B0F362FFDEFFDCB0117EBF64AEFF3D" box="[812,908,195,221]" italics="true" pageId="3" pageNumber="68">A. mirae</emphasis>
</taxonomicName>
and its allied species.
</paragraph>
<caption id="40BB7FF8FFDEFFDCB3B57A5B64EBFBDC" ID-DOI="http://doi.org/10.5281/zenodo.13213504" ID-Zenodo-Dep="13213504" box="[136,969,1063,1085]" httpUri="https://zenodo.org/record/13213504/files/figure.png" pageId="3" pageNumber="68" startId="3.[136,229,1063,1085]" targetBox="[260,1326,260,1014]" targetPageId="3" targetType="figure">
<paragraph id="147B2F70FFDEFFDCB3B57A5B64EBFBDC" blockId="3.[136,969,1063,1085]" box="[136,969,1063,1085]" pageId="3" pageNumber="68">
<emphasis id="26B0F362FFDEFFDCB3B57A5B67DCFBDD" bold="true" box="[136,254,1063,1085]" pageId="3" pageNumber="68">FIGURE 2.</emphasis>
Map of Mindanao showing the distribution of
<taxonomicName id="D3C454F3FFDEFFDCB1827A546455FBDC" box="[703,887,1063,1085]" class="Magnoliopsida" family="Rubiaceae" genus="Argostemma" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="68" phylum="Tracheophyta" rank="species" species="mirae">Argostemma mirae</taxonomicName>
(green).
</paragraph>
</caption>
<caption id="40BB7FF8FFDEFFDCB3B57A11627AFB67" box="[136,1368,1133,1158]" pageId="3" pageNumber="68" startId="3.[136,224,1133,1157]" targetType="table">
<paragraph id="147B2F70FFDEFFDCB3B57A11627AFB67" blockId="3.[136,1368,1133,1158]" box="[136,1368,1133,1158]" pageId="3" pageNumber="68">
<emphasis id="26B0F362FFDEFFDCB3B57A1167D9FB64" bold="true" box="[136,251,1133,1157]" pageId="3" pageNumber="68">TABLE 1.</emphasis>
Comparative morphology among
<taxonomicName id="D3C454F3FFDEFFDCB1577A126417FB64" box="[618,821,1134,1157]" class="Magnoliopsida" family="Rubiaceae" genus="Argostemma" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="68" phylum="Tracheophyta" rank="species" species="mirae">
<emphasis id="26B0F362FFDEFFDCB1577A126417FB64" box="[618,821,1134,1157]" italics="true" pageId="3" pageNumber="68">Argostemma mirae</emphasis>
</taxonomicName>
,
<taxonomicName id="D3C454F3FFDEFFDCB07E7A1264C6FB67" authorityName="Merrill" authorityYear="1922" box="[835,996,1134,1158]" class="Magnoliopsida" family="Rubiaceae" genus="Argostemma" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="68" phylum="Tracheophyta" rank="species" species="arachnosum">
<emphasis id="26B0F362FFDEFFDCB07E7A1264C6FB67" box="[835,996,1134,1158]" italics="true" pageId="3" pageNumber="68">A. arachnosum</emphasis>
</taxonomicName>
,
<taxonomicName id="D3C454F3FFDEFFDCB0CC7A126355FB67" authorityName="Valeton" authorityYear="1911" box="[1009,1143,1134,1158]" class="Magnoliopsida" family="Rubiaceae" genus="Argostemma" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="68" phylum="Tracheophyta" rank="species" species="distichum">
<emphasis id="26B0F362FFDEFFDCB0CC7A126355FB67" box="[1009,1143,1134,1158]" italics="true" pageId="3" pageNumber="68">A. distichum</emphasis>
</taxonomicName>
and
<taxonomicName id="D3C454F3FFDEFFDCB7977A126276FB67" authorityName="Elmer" authorityYear="1906" box="[1194,1364,1134,1158]" class="Magnoliopsida" family="Rubiaceae" genus="Argostemma" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="68" phylum="Tracheophyta" rank="species" species="solaniflorum">A. solaniflorum</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="147B2F70FFDEFFDCB3AA7ADE6212F7F0" pageId="3" pageNumber="68">
<table id="66C4DDD0FFDE0020B3AA7ADD62BAF7F3" box="[151,1432,1185,2066]" gridcols="5" gridrows="16" pageId="3" pageNumber="68">
<tr id="AAF42D32FFDE0020B3AA7ADD62BAFB59" box="[151,1432,1185,1208]" gridrow="0" pageId="3" pageNumber="68">
<th id="E925444EFFDE0020B3AA7ADD665CFB59" box="[151,382,1185,1208]" gridcol="0" gridrow="0" pageId="3" pageNumber="68">
<emphasis id="26B0F362FFDEFFDCB3AA7ADE6621FB59" bold="true" box="[151,259,1186,1208]" pageId="3" pageNumber="68">Characters</emphasis>
</th>
<th id="E925444EFFDE0020B2A27ADD65A5FB59" box="[415,647,1185,1208]" gridcol="1" gridrow="0" pageId="3" pageNumber="68">
<taxonomicName id="D3C454F3FFDEFFDCB2997ADE66D7FB56" box="[420,501,1186,1207]" class="Magnoliopsida" family="Rubiaceae" genus="Argostemma" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="68" phylum="Tracheophyta" rank="species" species="mirae">
<emphasis id="26B0F362FFDEFFDCB2997ADE66D7FB56" bold="true" box="[420,501,1186,1207]" italics="true" pageId="3" pageNumber="68">A. mirae</emphasis>
</taxonomicName>
</th>
<th id="E925444EFFDE0020B1987ADD64B5FB59" box="[677,919,1185,1208]" gridcol="2" gridrow="0" pageId="3" pageNumber="68">
<taxonomicName id="D3C454F3FFDEFFDCB1987ADE6416FB56" authorityName="Merrill" authorityYear="1922" box="[677,820,1185,1207]" class="Magnoliopsida" family="Rubiaceae" genus="Argostemma" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="68" phylum="Tracheophyta" rank="species" species="arachnosum">
<emphasis id="26B0F362FFDEFFDCB1987ADE6416FB56" bold="true" box="[677,820,1185,1207]" italics="true" pageId="3" pageNumber="68">A. arachnosum</emphasis>
</taxonomicName>
</th>
<th id="E925444EFFDE0020B0867ADD6387FB59" box="[955,1189,1185,1208]" gridcol="3" gridrow="0" pageId="3" pageNumber="68">
<taxonomicName id="D3C454F3FFDEFFDCB0827ADE6314FB56" authorityName="Valeton" authorityYear="1911" box="[959,1078,1185,1207]" class="Magnoliopsida" family="Rubiaceae" genus="Argostemma" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="68" phylum="Tracheophyta" rank="species" species="distichum">
<emphasis id="26B0F362FFDEFFDCB0827ADE6314FB56" bold="true" box="[959,1078,1185,1207]" italics="true" pageId="3" pageNumber="68">A. distichum</emphasis>
</taxonomicName>
</th>
<th id="E925444EFFDE0020B7FB7ADD62BAFB59" box="[1222,1432,1185,1208]" gridcol="4" gridrow="0" pageId="3" pageNumber="68">
<taxonomicName id="D3C454F3FFDEFFDCB7F67ADE627DFB56" authority="Stems" authorityName="Stems" box="[1227,1375,1185,1207]" class="Magnoliopsida" family="Rubiaceae" genus="Argostemma" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="68" phylum="Tracheophyta" rank="species" species="solaniflorum">
<emphasis id="26B0F362FFDEFFDCB7F67ADE627DFB56" bold="true" box="[1227,1375,1185,1207]" italics="true" pageId="3" pageNumber="68">A. solaniflorum</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="AAF42D32FFDE0020B3AA7AA462BAFAEF" box="[151,1432,1240,1294]" gridrow="1" pageId="3" pageNumber="68">
<th id="E925444EFFDE0020B3AA7AA4665CFAEF" box="[151,382,1240,1294]" gridcol="0" gridrow="1" pageId="3" pageNumber="68">Stems and height in cm.</th>
<td id="E925444EFFDE0020B2A27AA465A5FAEF" box="[415,647,1240,1294]" gridcol="1" gridrow="1" pageId="3" pageNumber="68">
Erect, branching, pubescent.
<emphasis id="26B0F362FFDEFFDCB1387A84651CFAEC" bold="true" box="[517,574,1272,1293]" pageId="3" pageNumber="68">10–13</emphasis>
</td>
<td id="E925444EFFDE0020B1987AA464B5FAEF" box="[677,919,1240,1294]" gridcol="2" gridrow="1" pageId="3" pageNumber="68">Creeping, solitary, pubescent. 20</td>
<td id="E925444EFFDE0020B0867AA46387FAEF" box="[955,1189,1240,1294]" gridcol="3" gridrow="1" pageId="3" pageNumber="68">Creeping, branching, glabrous. 5–10</td>
<td id="E925444EFFDE0020B7FB7AA462BAFAEF" box="[1222,1432,1240,1294]" gridcol="4" gridrow="1" pageId="3" pageNumber="68">Erect, branching, pubescent. 16</td>
</tr>
<tr id="AAF42D32FFDE0020B3AA7B5A62BAFADD" box="[151,1432,1318,1340]" gridrow="2" pageId="3" pageNumber="68">
<th id="E925444EFFDE0020B3AA7B5A665CFADD" box="[151,382,1318,1340]" gridcol="0" gridrow="2" pageId="3" pageNumber="68">Stipules</th>
<td id="E925444EFFDE0020B2A27B5A65A5FADD" box="[415,647,1318,1340]" gridcol="1" gridrow="2" pageId="3" pageNumber="68">
<emphasis id="26B0F362FFDEFFDCB2A27B5A65A4FADA" bold="true" box="[415,646,1318,1339]" pageId="3" pageNumber="68">Broadly ovate, glabrous</emphasis>
</td>
<td id="E925444EFFDE0020B1987B5A64B5FADD" box="[677,919,1318,1340]" gridcol="2" gridrow="2" pageId="3" pageNumber="68">Oblong, glabrous</td>
<td id="E925444EFFDE0020B0867B5A6387FADD" box="[955,1189,1318,1340]" gridcol="3" gridrow="2" pageId="3" pageNumber="68">Broadly ovate, hairy</td>
<td id="E925444EFFDE0020B7FB7B5A62BAFADD" box="[1222,1432,1318,1340]" gridcol="4" gridrow="2" pageId="3" pageNumber="68">Oblong, glabrous</td>
</tr>
<tr id="AAF42D32FFDE0020B3AA7B2862BAFA4B" box="[151,1432,1364,1450]" gridrow="3" pageId="3" pageNumber="68">
<th id="E925444EFFDE0020B3AA7B28665CFA4B" box="[151,382,1364,1450]" gridcol="0" gridrow="3" pageId="3" pageNumber="68">Leaf blade lamina and shape</th>
<td id="E925444EFFDE0020B2A27B2865A5FA4B" box="[415,647,1364,1450]" gridcol="1" gridrow="3" pageId="3" pageNumber="68">Dark green. Oblanceolate</td>
<td id="E925444EFFDE0020B1987B2864B5FA4B" box="[677,919,1364,1450]" gridcol="2" gridrow="3" pageId="3" pageNumber="68">Light green with light green sparse. Oblong- elliptic</td>
<td id="E925444EFFDE0020B0867B286387FA4B" box="[955,1189,1364,1450]" gridcol="3" gridrow="3" pageId="3" pageNumber="68">Dark green. Ovate- lanceolate</td>
<td id="E925444EFFDE0020B7FB7B2862BAFA4B" box="[1222,1432,1364,1450]" gridcol="4" gridrow="3" pageId="3" pageNumber="68">Dark green. Oblanceolate</td>
</tr>
<tr id="AAF42D32FFDE0020B3AA7BBE62BAFA39" box="[151,1432,1474,1496]" gridrow="4" pageId="3" pageNumber="68">
<th id="E925444EFFDE0020B3AA7BBE665CFA39" box="[151,382,1474,1496]" gridcol="0" gridrow="4" pageId="3" pageNumber="68">Leaf blade size cm.</th>
<td id="E925444EFFDE0020B2A27BBE65A5FA39" box="[415,647,1474,1496]" gridcol="1" gridrow="4" pageId="3" pageNumber="68">
<emphasis id="26B0F362FFDEFFDCB2A27BBE653CFA36" bold="true" box="[415,542,1474,1495]" pageId="3" pageNumber="68">2.5–3 × 1–1.3</emphasis>
</td>
<td id="E925444EFFDE0020B1987BBE64B5FA39" box="[677,919,1474,1496]" gridcol="2" gridrow="4" pageId="3" pageNumber="68">4.5–8 × 1.8–2.5</td>
<td id="E925444EFFDE0020B0867BBE6387FA39" box="[955,1189,1474,1496]" gridcol="3" gridrow="4" pageId="3" pageNumber="68">1.8–2.5 × 0.5</td>
<td id="E925444EFFDE0020B7FB7BBE62BAFA39" box="[1222,1432,1474,1496]" gridcol="4" gridrow="4" pageId="3" pageNumber="68">7 × 2</td>
</tr>
<tr id="AAF42D32FFDE0020B3AA7B8C62BAF9E7" box="[151,1432,1520,1542]" gridrow="5" pageId="3" pageNumber="68">
<th id="E925444EFFDE0020B3AA7B8C665CF9E7" box="[151,382,1520,1542]" gridcol="0" gridrow="5" pageId="3" pageNumber="68">Leaf Apex</th>
<td id="E925444EFFDE0020B2A27B8C65A5F9E7" box="[415,647,1520,1542]" gridcol="1" gridrow="5" pageId="3" pageNumber="68">Acuminate</td>
<td id="E925444EFFDE0020B1987B8C64B5F9E7" box="[677,919,1520,1542]" gridcol="2" gridrow="5" pageId="3" pageNumber="68">Acuminate</td>
<td id="E925444EFFDE0020B0867B8C6387F9E7" box="[955,1189,1520,1542]" gridcol="3" gridrow="5" pageId="3" pageNumber="68">Sharply acute</td>
<td id="E925444EFFDE0020B7FB7B8C62BAF9E7" box="[1222,1432,1520,1542]" gridcol="4" gridrow="5" pageId="3" pageNumber="68">Obtuse</td>
</tr>
<tr id="AAF42D32FFDE0020B3AA786262BAF9B5" box="[151,1432,1566,1620]" gridrow="6" pageId="3" pageNumber="68">
<th id="E925444EFFDE0020B3AA7862665CF9B5" box="[151,382,1566,1620]" gridcol="0" gridrow="6" pageId="3" pageNumber="68">Leaf surfaces</th>
<td id="E925444EFFDE0020B2A2786265A5F9B5" box="[415,647,1566,1620]" gridcol="1" gridrow="6" pageId="3" pageNumber="68">Adaxially glabrous, abaxially pubescent</td>
<td id="E925444EFFDE0020B198786264B5F9B5" box="[677,919,1566,1620]" gridcol="2" gridrow="6" pageId="3" pageNumber="68">Adaxially glabrous, abaxially pubescent</td>
<td id="E925444EFFDE0020B08678626387F9B5" box="[955,1189,1566,1620]" gridcol="3" gridrow="6" pageId="3" pageNumber="68">Adaxially and abaxially pubescent</td>
<td id="E925444EFFDE0020B7FB786262BAF9B5" box="[1222,1432,1566,1620]" gridcol="4" gridrow="6" pageId="3" pageNumber="68">Adaxially and abaxially pubescent</td>
</tr>
<tr id="AAF42D32FFDE0020B3AA781062BAF963" box="[151,1432,1644,1666]" gridrow="7" pageId="3" pageNumber="68">
<th id="E925444EFFDE0020B3AA7810665CF963" box="[151,382,1644,1666]" gridcol="0" gridrow="7" pageId="3" pageNumber="68">Leaf Base</th>
<td id="E925444EFFDE0020B2A2781065A5F963" box="[415,647,1644,1666]" gridcol="1" gridrow="7" pageId="3" pageNumber="68">Acute</td>
<td id="E925444EFFDE0020B198781064B5F963" box="[677,919,1644,1666]" gridcol="2" gridrow="7" pageId="3" pageNumber="68">Cordate-rounded</td>
<td id="E925444EFFDE0020B08678106387F963" box="[955,1189,1644,1666]" gridcol="3" gridrow="7" pageId="3" pageNumber="68">Acute</td>
<td id="E925444EFFDE0020B7FB781062BAF963" box="[1222,1432,1644,1666]" gridcol="4" gridrow="7" pageId="3" pageNumber="68">Acute</td>
</tr>
<tr id="AAF42D32FFDE0020B3AA78E662BAF951" box="[151,1432,1690,1712]" gridrow="8" pageId="3" pageNumber="68">
<th id="E925444EFFDE0020B3AA78E6665CF951" box="[151,382,1690,1712]" gridcol="0" gridrow="8" pageId="3" pageNumber="68">Leaf Margin</th>
<td id="E925444EFFDE0020B2A278E665A5F951" box="[415,647,1690,1712]" gridcol="1" gridrow="8" pageId="3" pageNumber="68">Entire</td>
<td id="E925444EFFDE0020B19878E664B5F951" box="[677,919,1690,1712]" gridcol="2" gridrow="8" pageId="3" pageNumber="68">Sinuate</td>
<td id="E925444EFFDE0020B08678E66387F951" box="[955,1189,1690,1712]" gridcol="3" gridrow="8" pageId="3" pageNumber="68">Entire</td>
<td id="E925444EFFDE0020B7FB78E662BAF951" box="[1222,1432,1690,1712]" gridcol="4" gridrow="8" pageId="3" pageNumber="68">Entire</td>
</tr>
<tr id="AAF42D32FFDE0020B3AA78B462BAF93F" box="[151,1432,1736,1758]" gridrow="9" pageId="3" pageNumber="68">
<th id="E925444EFFDE0020B3AA78B4665CF93F" box="[151,382,1736,1758]" gridcol="0" gridrow="9" pageId="3" pageNumber="68">Lateral leaf veins in pairs</th>
<td id="E925444EFFDE0020B2A278B465A5F93F" box="[415,647,1736,1758]" gridcol="1" gridrow="9" pageId="3" pageNumber="68">
<emphasis id="26B0F362FFDEFFDCB2A278B466E0F93C" bold="true" box="[415,450,1736,1757]" pageId="3" pageNumber="68">4–5</emphasis>
</td>
<td id="E925444EFFDE0020B19878B464B5F93F" box="[677,919,1736,1758]" gridcol="2" gridrow="9" pageId="3" pageNumber="68">8</td>
<td id="E925444EFFDE0020B08678B46387F93F" box="[955,1189,1736,1758]" gridcol="3" gridrow="9" pageId="3" pageNumber="68">1–2</td>
<td id="E925444EFFDE0020B7FB78B462BAF93F" box="[1222,1432,1736,1758]" gridcol="4" gridrow="9" pageId="3" pageNumber="68">7–8</td>
</tr>
<tr id="AAF42D32FFDE0020B3AA788A62BAF8ED" box="[151,1432,1782,1804]" gridrow="10" pageId="3" pageNumber="68">
<th id="E925444EFFDE0020B3AA788A665CF8ED" box="[151,382,1782,1804]" gridcol="0" gridrow="10" pageId="3" pageNumber="68">Inflorescence</th>
<td id="E925444EFFDE0020B2A2788A65A5F8ED" box="[415,647,1782,1804]" gridcol="1" gridrow="10" pageId="3" pageNumber="68">
<emphasis id="26B0F362FFDEFFDCB2A2788A6688F8EA" bold="true" box="[415,426,1782,1803]" pageId="3" pageNumber="68">2</emphasis>
</td>
<td id="E925444EFFDE0020B198788A64B5F8ED" box="[677,919,1782,1804]" gridcol="2" gridrow="10" pageId="3" pageNumber="68">10–25</td>
<td id="E925444EFFDE0020B086788A6387F8ED" box="[955,1189,1782,1804]" gridcol="3" gridrow="10" pageId="3" pageNumber="68">5–6</td>
<td id="E925444EFFDE0020B7FB788A62BAF8ED" box="[1222,1432,1782,1804]" gridcol="4" gridrow="10" pageId="3" pageNumber="68">5</td>
</tr>
<tr id="AAF42D32FFDE0020B3AA795F62BAF8DB" box="[151,1432,1827,1850]" gridrow="11" pageId="3" pageNumber="68">
<th id="E925444EFFDE0020B3AA795F665CF8DB" box="[151,382,1827,1850]" gridcol="0" gridrow="11" pageId="3" pageNumber="68">Calyx lobes</th>
<td id="E925444EFFDE0020B2A2795F65A5F8DB" box="[415,647,1827,1850]" gridcol="1" gridrow="11" pageId="3" pageNumber="68">
<emphasis id="26B0F362FFDEFFDCB2A2795F6533F8D8" bold="true" box="[415,529,1827,1849]" pageId="3" pageNumber="68">4 pubescent</emphasis>
</td>
<td id="E925444EFFDE0020B198795F64B5F8DB" box="[677,919,1827,1850]" gridcol="2" gridrow="11" pageId="3" pageNumber="68">5 pubescent</td>
<td id="E925444EFFDE0020B086795F6387F8DB" box="[955,1189,1827,1850]" gridcol="3" gridrow="11" pageId="3" pageNumber="68">4 glabrous</td>
<td id="E925444EFFDE0020B7FB795F62BAF8DB" box="[1222,1432,1827,1850]" gridcol="4" gridrow="11" pageId="3" pageNumber="68">5 pubescent</td>
</tr>
<tr id="AAF42D32FFDE0020B3AA792E62BAF869" box="[151,1432,1874,1928]" gridrow="12" pageId="3" pageNumber="68">
<th id="E925444EFFDE0020B3AA792E665CF869" box="[151,382,1874,1928]" gridcol="0" gridrow="12" pageId="3" pageNumber="68">Corolla lobes</th>
<td id="E925444EFFDE0020B2A2792E65A5F869" box="[415,647,1874,1928]" gridcol="1" gridrow="12" pageId="3" pageNumber="68">4 glabrous both adaxially, and abaxially</td>
<td id="E925444EFFDE0020B198792E64B5F869" box="[677,919,1874,1928]" gridcol="2" gridrow="12" pageId="3" pageNumber="68">5 pubescent both adaxially and abaxially</td>
<td id="E925444EFFDE0020B086792E6387F869" box="[955,1189,1874,1928]" gridcol="3" gridrow="12" pageId="3" pageNumber="68">4 glabrous both adaxially, abaxially</td>
<td id="E925444EFFDE0020B7FB792E62BAF869" box="[1222,1432,1874,1928]" gridcol="4" gridrow="12" pageId="3" pageNumber="68">5 pubescent both adaxially and abaxially</td>
</tr>
<tr id="AAF42D32FFDE0020B3AA79DC62BAF857" box="[151,1432,1952,1974]" gridrow="13" pageId="3" pageNumber="68">
<th id="E925444EFFDE0020B3AA79DC665CF857" box="[151,382,1952,1974]" gridcol="0" gridrow="13" pageId="3" pageNumber="68">Pedicel mm.</th>
<td id="E925444EFFDE0020B2A279DC65A5F857" box="[415,647,1952,1974]" gridcol="1" gridrow="13" pageId="3" pageNumber="68">
<emphasis id="26B0F362FFDEFFDCB2A279DC66FAF854" bold="true" box="[415,472,1952,1973]" pageId="3" pageNumber="68">15–17</emphasis>
pubescent
</td>
<td id="E925444EFFDE0020B19879DC64B5F857" box="[677,919,1952,1974]" gridcol="2" gridrow="13" pageId="3" pageNumber="68">10–12 pubescent</td>
<td id="E925444EFFDE0020B08679DC6387F857" box="[955,1189,1952,1974]" gridcol="3" gridrow="13" pageId="3" pageNumber="68">3–5 pubescent</td>
<td id="E925444EFFDE0020B7FB79DC62BAF857" box="[1222,1432,1952,1974]" gridcol="4" gridrow="13" pageId="3" pageNumber="68">8 pubescent</td>
</tr>
<tr id="AAF42D32FFDE0020B3AA79B162BAF805" box="[151,1432,1997,2020]" gridrow="14" pageId="3" pageNumber="68">
<th id="E925444EFFDE0020B3AA79B1665CF805" box="[151,382,1997,2020]" gridcol="0" gridrow="14" pageId="3" pageNumber="68">Anthers length mm.</th>
<td id="E925444EFFDE0020B2A279B165A5F805" box="[415,647,1997,2020]" gridcol="1" gridrow="14" pageId="3" pageNumber="68">
<emphasis id="26B0F362FFDEFFDCB2A279B166E3F803" bold="true" box="[415,449,1997,2018]" pageId="3" pageNumber="68">3–4</emphasis>
</td>
<td id="E925444EFFDE0020B19879B164B5F805" box="[677,919,1997,2020]" gridcol="2" gridrow="14" pageId="3" pageNumber="68">10–12</td>
<td id="E925444EFFDE0020B08679B16387F805" box="[955,1189,1997,2020]" gridcol="3" gridrow="14" pageId="3" pageNumber="68">1–5</td>
<td id="E925444EFFDE0020B7FB79B162BAF805" box="[1222,1432,1997,2020]" gridcol="4" gridrow="14" pageId="3" pageNumber="68">7</td>
</tr>
<tr id="AAF42D32FFDE0020B3AA798062BAF7F3" box="[151,1432,2044,2066]" gridrow="15" pageId="3" pageNumber="68">
<th id="E925444EFFDE0020B3AA7980665CF7F3" box="[151,382,2044,2066]" gridcol="0" gridrow="15" pageId="3" pageNumber="68">Anthers dehiscence</th>
<td id="E925444EFFDE0020B2A2798065A5F7F3" box="[415,647,2044,2066]" gridcol="1" gridrow="15" pageId="3" pageNumber="68">Longitudinal slits</td>
<td id="E925444EFFDE0020B198798064B5F7F3" box="[677,919,2044,2066]" gridcol="2" gridrow="15" pageId="3" pageNumber="68">Longitudinal slits</td>
<td id="E925444EFFDE0020B08679806387F7F3" box="[955,1189,2044,2066]" gridcol="3" gridrow="15" pageId="3" pageNumber="68">Longitudinal slits</td>
<td id="E925444EFFDE0020B7FB798062BAF7F3" box="[1222,1432,2044,2066]" gridcol="4" gridrow="15" pageId="3" pageNumber="68">Apical pore</td>
</tr>
</table>
</paragraph>
<paragraph id="147B2F70FFD9FFDBB3807EE2627FFEC5" blockId="4.[136,1452,158,293]" pageId="4" pageNumber="69">
<emphasis id="26B0F362FFD9FFDBB3807EE2652EFF59" bold="true" box="[189,524,158,184]" pageId="4" pageNumber="69">Proposed IUCN category:—</emphasis>
Mt. Tingtingan has not been explored by any researchers due to the peace and order situation, which can make it hard to assess the conservation status of the species. The location is not a protected area. Although there is only one population currently known, there are several individuals found in a moist shady, mossy limestone in an undisturbed forest. We recommend a conservation status of data deficient [DD] (
<bibRefCitation id="70555281FFD9FFDBB7FD7F766273FEC5" author="IUCN Standards &amp; Petitions Committee" box="[1216,1361,266,292]" pageId="4" pageNumber="69" refId="ref3164" refString="IUCN Standards &amp; Petitions Committee. (2022) Guidelines for Using the IUCN Red List Categories and Criteria. Version 15.1. Available from: https: // www. iucnredlist. org / documents / RedListGuidelines. pdf (accessed: 6 March 2024)." type="url" year="2022">IUCN, 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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