386 lines
52 KiB
XML
386 lines
52 KiB
XML
<document id="3D723BA011C925FFE8CF1DE6126B5937" ID-DOI="10.26107/RBZ-2019-0038" ID-ISSN="2345-7600" ID-Zenodo-Dep="4577132" ID-ZooBank="02C676DE-6038-4277-A68A-39214E7B7E51" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614784115311" checkinUser="carolina" docAuthor="Yan, Bee, Lee & Ng, Peter K. L." docDate="2019" docId="792987CCFFCBFFD3FEFC04BFFD3CF9B2" docLanguage="en" docName="RafflesBZool.67.490-497.pdf" docOrigin="Raffles Bulletin of Zoology 67" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Hyastenus tabolongi Yan & Lee & Ng 2019, new species" docType="treatment" docVersion="3" lastPageNumber="496" masterDocId="8510FFB4FFCEFFD5FFF30063FFF4FFA4" masterDocTitle="On the identity of Hyastenus inermis (Rathbun, 1911), and description of a new species from Sulawesi, Indonesia (Crustacea: Decapoda: Majoidea: Epialtidae)" masterLastPageNumber="497" masterPageNumber="490" pageNumber="495" updateTime="1698878593994" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="624560133304B667A9E4CADC383A6F0C">On the identity of Hyastenus inermis (Rathbun, 1911), and description of a new species from Sulawesi, Indonesia (Crustacea: Decapoda: Majoidea: Epialtidae)</mods:title>
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<mods:name id="07AC8009A5127E3152FD3DC614AB600B" type="personal">
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<mods:namePart id="AAE3EDD5B5769B2230791208A4659036">Yan, Bee</mods:namePart>
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<mods:namePart id="510984CD4958607E9A91B570964868A3">Lee</mods:namePart>
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<mods:name id="E3753ED9D89BE1E3588C435B5F0EA9DF" type="personal">
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<mods:roleTerm id="B6E191580676AD961120356E76F77DBB">Author</mods:roleTerm>
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<mods:namePart id="06199C470547032A7CB30CA3C642B2CE">Ng, Peter K. L.</mods:namePart>
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<mods:title id="A3C5B4651DCEFA7A553494FF36A84CF9">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="7581E381329E49F72388480F55D117B3">
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<mods:date id="7359EBA46799E2AD99BCCA27C4226893">2019</mods:date>
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<mods:number id="62968AE2A7F96FF5615B7095460117DD">2019-10-29</mods:number>
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<mods:identifier id="4B980DFF5779F6E8BF4FBDA17058B852" type="DOI">10.26107/RBZ-2019-0038</mods:identifier>
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<mods:identifier id="224A19632AA31B014E7C0976F9004E44" type="ISSN">2345-7600</mods:identifier>
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<subSubSection id="B99A6551FFCBFFD0FEFC04BFFD77FB57" box="[271,643,1243,1268]" pageId="5" pageNumber="495" type="nomenclature">
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<paragraph id="F13F36DAFFCBFFD0FEFC04BFFD77FB57" blockId="5.[271,643,1243,1300]" box="[271,643,1243,1268]" pageId="5" pageNumber="495">
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<emphasis id="C3F4EAC8FFCBFFD0FEFC04BFFD77FB57" bold="true" box="[271,643,1243,1268]" pageId="5" pageNumber="495">
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<taxonomicName id="36804D59FFCBFFD0FEFC04BFFE07FB57" ID-CoL="84GWZ" authority="Yan & Lee & Ng, 2019" authorityName="Yan & Lee & Ng" authorityYear="2019" box="[271,499,1243,1267]" class="Malacostraca" family="Epialtidae" genus="Hyastenus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="495" phylum="Arthropoda" rank="species" species="tabolongi" status="sp. nov.">
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<emphasis id="C3F4EAC8FFCBFFD0FEFC04BFFE07FB57" bold="true" box="[271,499,1243,1267]" italics="true" pageId="5" pageNumber="495">Hyastenus tabolongi</emphasis>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="B99A6551FFCBFFD0FE91049FFDD9FAB0" box="[354,557,1276,1300]" pageId="5" pageNumber="495" type="description">
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<paragraph id="F13F36DAFFCBFFD0FE91049FFDD9FAB0" blockId="5.[271,643,1243,1300]" box="[354,557,1276,1300]" pageId="5" pageNumber="495">
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(
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<figureCitation id="69BB2A5FFFCBFFD0FE99049FFE1AFAB7" box="[362,494,1276,1300]" captionStart="Fig" captionStartId="2.[113,148,2035,2057]" captionTargetBox="[137,1428,190,2020]" captionTargetId="figure-16@2.[119,1430,176,2023]" captionTargetPageId="2" captionText="Fig. 2. Drawing of left G1 and G2. A–F, Hyastenus inermis (Rathbun, 1911), holotype male (15.5 × 10.6 mm) (USNM 41402), Indian Ocean; G, H, H. tabolongi, new species, holotype male (10.7 × 6.6 mm) (MZB Cru 5010), north Sulawesi. A, G, ventral view; B, ventral view of distal portion; C, H, dorsal view; D, dorsal view of distal portion; E, ventral view of G2; F, dorsal view of G2. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4577138" httpUri="https://zenodo.org/record/4577138/files/figure.png" pageId="5" pageNumber="495">Figs. 2G, H</figureCitation>
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,
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<figureCitation id="69BB2A5FFFCBFFD0FE09049FFDFDFAB0" box="[506,521,1276,1300]" captionStart="Fig" captionStartId="3.[136,171,1936,1958]" captionTargetBox="[136,1474,286,1920]" captionTargetId="figure-7@3.[136,1474,286,1920]" captionTargetPageId="3" captionText="Fig. 3. Hyastenus tabolongi, new species, A–C, holotype male (10.7 × 6.6 mm) (MZB Cru 5010), north Sulawesi; D–F, paratype female (12.3 × 7.2 mm) (ZRC 2019.1324), north Sulawesi. A, D, overall dorsal view of carapace; B, E, overall ventral view; C, F, lateral view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4577142" httpUri="https://zenodo.org/record/4577142/files/figure.png" pageId="5" pageNumber="495">3</figureCitation>
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,
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<figureCitation id="69BB2A5FFFCBFFD0FDE4049FFDD1FAB0" box="[535,549,1276,1300]" captionStart="Fig" captionStartId="5.[136,171,1138,1160]" captionTargetBox="[136,1474,190,1123]" captionTargetId="figure-41@5.[136,1474,190,1123]" captionTargetPageId="5" captionText="Fig. 4. Hyastenus tabolongi, new species, observed in situ from Bunaken, Lekuan on different dates. A, B, photographed in November 2011; C, D, photographed in July 2018. Photographs by: AC Podzorski" figureDoi="http://doi.org/10.5281/zenodo.4577144" httpUri="https://zenodo.org/record/4577144/files/figure.png" pageId="5" pageNumber="495">4</figureCitation>
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)
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<subSubSection id="B99A6551FFCBFFD0FF7B0558FE67F970" pageId="5" pageNumber="495" type="materials_examined">
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<paragraph id="F13F36DAFFCBFFD0FF7B0558FE67F970" blockId="5.[136,775,1339,1748]" pageId="5" pageNumber="495">
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<emphasis id="C3F4EAC8FFCBFFD0FF7B0558FE91FAF7" bold="true" box="[136,357,1339,1363]" pageId="5" pageNumber="495">Material examined.</emphasis>
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<materialsCitation id="41E83C87FFCBFFD0FE98055FFE63FA30" collectionCode="MZB" country="Indonesia" location="Cru" pageId="5" pageNumber="495" specimenCount="1" stateProvince="Sulawesi Utara" typeStatus="holotype">
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<typeStatus id="2E3B8878FFCBFFD0FE98055FFE26FAF0" box="[363,466,1340,1364]" pageId="5" pageNumber="495" type="holotype">Holotype</typeStatus>
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: male (10.7 ×
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) (
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<collectionCode id="9791AE1FFFCBFFD0FD38055FFCF2FAF0" box="[715,774,1340,1364]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="5" pageNumber="495" type="Museum">MZB</collectionCode>
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<location id="F45F6001FFCBFFD0FF7B053FFF44FAD0" LSID="urn:lsid:plazi:treatment:792987CCFFCBFFD3FEFC04BFFD3CF9B2:F45F6001FFCBFFD0FF7B053FFF44FAD0" box="[136,176,1372,1396]" country="Indonesia" name="Cru" pageId="5" pageNumber="495" stateProvince="Sulawesi Utara">Cru</location>
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5010),
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<collectingRegion id="3344F838FFCBFFD0FEFB053FFE5BFAD0" box="[264,431,1372,1396]" country="Indonesia" name="Sulawesi Utara" pageId="5" pageNumber="495">North Sulawesi</collectingRegion>
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[=
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<collectingRegion id="3344F838FFCBFFD0FE28053FFD70FAD7" box="[475,644,1372,1396]" country="Indonesia" name="Sulawesi Utara" pageId="5" pageNumber="495">Sulawesi Utara</collectingRegion>
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, coll. local divers, 2000s
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</materialsCitation>
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.
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<materialsCitation id="41E83C87FFCBFFD0FE57051FFDFCF990" collectingDate="2014-12-05" collectionCode="ZRC" country="Indonesia" latitude="1.4186667" location="Poh Poh" longLatPrecision="1" longitude="124.6298" pageId="5" pageNumber="495" specimenCode="ZRC 2019.1324, ZRC 2019.1325" specimenCount="1" specimenCount-female="1" stateProvince="Sulawesi Utara" typeStatus="paratype">
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<typeStatus id="2E3B8878FFCBFFD0FE57051FFDE5FA30" box="[420,529,1404,1428]" pageId="5" pageNumber="495" type="paratype">Paratypes</typeStatus>
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:
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(12.3 ×
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) (
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<specimenCode id="A1269EA1FFCBFFD0FF3505FFFE82FA10" box="[198,374,1436,1460]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="5" pageNumber="495">ZRC 2019.1324</specimenCode>
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),
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<location id="F45F6001FFCBFFD0FE7605FFFE14FA10" LSID="urn:lsid:plazi:treatment:792987CCFFCBFFD3FEFC04BFFD3CF9B2:F45F6001FFCBFFD0FE7605FFFE14FA10" box="[389,480,1436,1460]" country="Indonesia" latitude="1.4186667" longLatPrecision="1" longitude="124.6298" name="Poh Poh" pageId="5" pageNumber="495" stateProvince="Sulawesi Utara">Poh Poh</location>
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, Tanahwanko village,
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<collectingRegion id="3344F838FFCBFFD0FD3A05FFFF1FFA70" country="Indonesia" name="Sulawesi Utara" pageId="5" pageNumber="495">North Sulawesi</collectingRegion>
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,
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<collectingCountry id="8997764AFFCBFFD0FF0B05DFFE96FA70" box="[248,354,1468,1492]" name="Indonesia" pageId="5" pageNumber="495">Indonesia</collectingCountry>
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,
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<geoCoordinate id="94B4501DFFCBFFD0FE0505DFFD7AFA70" box="[502,654,1468,1492]" degrees="124" direction="east" minutes="37.788" orientation="longitude" pageId="5" pageNumber="495" precision="1" value="124.6298">124°37.788’E</geoCoordinate>
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, coll. local divers,
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<collectingDate id="957AE9F2FFCBFFD0FF2905BFFE55FA50" box="[218,417,1500,1524]" pageId="5" pageNumber="495" value="2014-12-05">5 December 2014</collectingDate>
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</date>
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; 1 ovigerous female (10.6 ×
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) (
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),
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[=
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],
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<collectingCountry id="8997764AFFCBFFD0FF7B067FFF06F990" box="[136,242,1564,1588]" name="Indonesia" pageId="5" pageNumber="495">Indonesia</collectingCountry>
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, coll. local divers, 2000s
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</materialsCitation>
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;
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<specimenCount id="E786FD53FFCBFFD0FDE6067FFD76F990" box="[533,642,1564,1588]" pageId="5" pageNumber="495" type="juvenile">1 juvenile</specimenCount>
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female (5.3 ×
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) (
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<specimenCode id="A1269EA1FFCBFFD0FEFE065FFE35F9F0" box="[269,449,1596,1620]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="5" pageNumber="495">ZRC 2019.1326</specimenCode>
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),
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<collectingRegion id="3344F838FFCBFFD0FE26065FFD88F9F0" box="[469,636,1596,1620]" country="Indonesia" name="Sulawesi Utara" pageId="5" pageNumber="495">North Sulawesi</collectingRegion>
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[=
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<collectingRegion id="3344F838FFCBFFD0FD54065FFF3CF9D7" country="Indonesia" name="Sulawesi Utara" pageId="5" pageNumber="495">Sulawesi Utara</collectingRegion>
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],
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, coll. local divers, 2000s
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</materialsCitation>
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;
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(9.8 ×
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),
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<specimenCount id="E786FD53FFCBFFD0FF05061FFEA2F930" box="[246,342,1660,1684]" pageId="5" pageNumber="495" type="female">2 female</specimenCount>
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(10.0 ×
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<quantity id="36789B3FFFCBFFD0FE49061FFDE5F937" box="[442,529,1660,1684]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="5" pageNumber="495" unit="mm" value="5.8">5.8 mm</quantity>
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, 5.0 ×
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) (
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<specimenCode id="A1269EA1FFCBFFD0FD22061FFF0AF910" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="5" pageNumber="495">ZRC 2019.1327</specimenCode>
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),
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[=
|
||
<collectingRegion id="3344F838FFCBFFD0FE2D06FFFD72F917" box="[478,646,1692,1716]" country="Indonesia" name="Sulawesi Utara" pageId="5" pageNumber="495">Sulawesi Utara</collectingRegion>
|
||
],
|
||
<collectingCountry id="8997764AFFCBFFD0FD6B06FFFCF6F910" box="[664,770,1692,1716]" name="Indonesia" pageId="5" pageNumber="495">Indonesia</collectingCountry>
|
||
, coll. local divers, 2000s
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B99A6551FFCBFFD0FF7B0698FA36F9B0" pageId="5" pageNumber="495" type="diagnosis">
|
||
<paragraph id="F13F36DAFFCBFFD0FF7B0698FA36F9B0" blockId="5.[136,775,1787,2100]" lastBlockId="5.[835,1474,1244,1556]" pageId="5" pageNumber="495">
|
||
<emphasis id="C3F4EAC8FFCBFFD0FF7B0698FF0EF8B7" bold="true" box="[136,250,1787,1811]" pageId="5" pageNumber="495">Diagnosis.</emphasis>
|
||
Carapace pyriform, regions defined. Pseudorostral spines straight, divergent. Supraorbital eave forming rectangular lobe; preorbital angle rounded, antorbital angle rounded, outer margin slightly constricted; orbital hiatus between supraorbital and postorbital lobes form narrow U-shape; postorbital lobe cup-like. Carapace lacking granules; gastric region prominently swollen; small tubercle on mesogastric region; epibranchial region swollen, slightly arched; cardiac region swollen; intestinal region swollen with 3 slightly swollen areas; 2 on metabranchial region and 1 medially on intestinal region (
|
||
<figureCitation id="69BB2A5FFFCBFFD0FB2204BFFABFFB50" box="[1233,1355,1244,1268]" captionStart="Fig" captionStartId="3.[136,171,1936,1958]" captionTargetBox="[136,1474,286,1920]" captionTargetId="figure-7@3.[136,1474,286,1920]" captionTargetPageId="3" captionText="Fig. 3. Hyastenus tabolongi, new species, A–C, holotype male (10.7 × 6.6 mm) (MZB Cru 5010), north Sulawesi; D–F, paratype female (12.3 × 7.2 mm) (ZRC 2019.1324), north Sulawesi. A, D, overall dorsal view of carapace; B, E, overall ventral view; C, F, lateral view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4577142" httpUri="https://zenodo.org/record/4577142/files/figure.png" pageId="5" pageNumber="495">Fig. 3A, D</figureCitation>
|
||
). Antennal flagellum shorter than pseudorostral spine. Basal antennal article longer than broad, distolateral angle of article sharp, outer margin distinctly constricted medially. Pterygostomial region with single granule on outer margin (
|
||
<figureCitation id="69BB2A5FFFCBFFD0FAC8053FFA42FAD0" box="[1339,1462,1372,1396]" captionStart="Fig" captionStartId="3.[136,171,1936,1958]" captionTargetBox="[136,1474,286,1920]" captionTargetId="figure-7@3.[136,1474,286,1920]" captionTargetPageId="3" captionText="Fig. 3. Hyastenus tabolongi, new species, A–C, holotype male (10.7 × 6.6 mm) (MZB Cru 5010), north Sulawesi; D–F, paratype female (12.3 × 7.2 mm) (ZRC 2019.1324), north Sulawesi. A, D, overall dorsal view of carapace; B, E, overall ventral view; C, F, lateral view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4577142" httpUri="https://zenodo.org/record/4577142/files/figure.png" pageId="5" pageNumber="495">Fig. 3B, E</figureCitation>
|
||
). Cheliped slender; surface smooth (
|
||
<figureCitation id="69BB2A5FFFCBFFD0FB44051FFAD9FA30" box="[1207,1325,1404,1428]" captionStart="Fig" captionStartId="3.[136,171,1936,1958]" captionTargetBox="[136,1474,286,1920]" captionTargetId="figure-7@3.[136,1474,286,1920]" captionTargetPageId="3" captionText="Fig. 3. Hyastenus tabolongi, new species, A–C, holotype male (10.7 × 6.6 mm) (MZB Cru 5010), north Sulawesi; D–F, paratype female (12.3 × 7.2 mm) (ZRC 2019.1324), north Sulawesi. A, D, overall dorsal view of carapace; B, E, overall ventral view; C, F, lateral view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4577142" httpUri="https://zenodo.org/record/4577142/files/figure.png" pageId="5" pageNumber="495">Fig. 3A, D</figureCitation>
|
||
). Ambulatory legs slender; P2 longest (
|
||
<figureCitation id="69BB2A5FFFCBFFD0FBBF05FFFB35FA10" box="[1100,1217,1436,1460]" captionStart="Fig" captionStartId="3.[136,171,1936,1958]" captionTargetBox="[136,1474,286,1920]" captionTargetId="figure-7@3.[136,1474,286,1920]" captionTargetPageId="3" captionText="Fig. 3. Hyastenus tabolongi, new species, A–C, holotype male (10.7 × 6.6 mm) (MZB Cru 5010), north Sulawesi; D–F, paratype female (12.3 × 7.2 mm) (ZRC 2019.1324), north Sulawesi. A, D, overall dorsal view of carapace; B, E, overall ventral view; C, F, lateral view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4577142" httpUri="https://zenodo.org/record/4577142/files/figure.png" pageId="5" pageNumber="495">Fig. 3A, D</figureCitation>
|
||
). Male thoracic sternum slightly concave anteriorly; sternites 3, 4 with lateral margin constricted. Male pleon slender, triangular, telson triangular (
|
||
<figureCitation id="69BB2A5FFFCBFFD0FCB8059FFC54F9B0" box="[843,928,1532,1556]" captionStart="Fig" captionStartId="3.[136,171,1936,1958]" captionTargetBox="[136,1474,286,1920]" captionTargetId="figure-7@3.[136,1474,286,1920]" captionTargetPageId="3" captionText="Fig. 3. Hyastenus tabolongi, new species, A–C, holotype male (10.7 × 6.6 mm) (MZB Cru 5010), north Sulawesi; D–F, paratype female (12.3 × 7.2 mm) (ZRC 2019.1324), north Sulawesi. A, D, overall dorsal view of carapace; B, E, overall ventral view; C, F, lateral view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4577142" httpUri="https://zenodo.org/record/4577142/files/figure.png" pageId="5" pageNumber="495">Fig. 3B</figureCitation>
|
||
). G1 slightly curved, distal tip sharp (
|
||
<figureCitation id="69BB2A5FFFCBFFD0FACE059FFA42F9B0" box="[1341,1462,1532,1556]" captionStart="Fig" captionStartId="2.[113,148,2035,2057]" captionTargetBox="[137,1428,190,2020]" captionTargetId="figure-16@2.[119,1430,176,2023]" captionTargetPageId="2" captionText="Fig. 2. Drawing of left G1 and G2. A–F, Hyastenus inermis (Rathbun, 1911), holotype male (15.5 × 10.6 mm) (USNM 41402), Indian Ocean; G, H, H. tabolongi, new species, holotype male (10.7 × 6.6 mm) (MZB Cru 5010), north Sulawesi. A, G, ventral view; B, ventral view of distal portion; C, H, dorsal view; D, dorsal view of distal portion; E, ventral view of G2; F, dorsal view of G2. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4577138" httpUri="https://zenodo.org/record/4577138/files/figure.png" pageId="5" pageNumber="495">Fig. 2G, H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B99A6551FFCBFFD0FCB0065FFA37F931" pageId="5" pageNumber="495" type="etymology">
|
||
<paragraph id="F13F36DAFFCBFFD0FCB0065FFA37F931" blockId="5.[835,1475,1596,1685]" pageId="5" pageNumber="495">
|
||
<emphasis id="C3F4EAC8FFCBFFD0FCB0065FFC33F9F0" bold="true" box="[835,967,1596,1620]" pageId="5" pageNumber="495">Etymology.</emphasis>
|
||
This species is named after Maxi Tabolong, a senior dive guide who first showed the crab to Andrew Podzorski, who was then able to photograph it over the years.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B99A6551FFCBFFD0FCB006DFFAFAF970" box="[835,1294,1724,1748]" pageId="5" pageNumber="495" type="materials_examined">
|
||
<paragraph id="F13F36DAFFCBFFD0FCB006DFFAFAF970" blockId="5.[835,1294,1724,1748]" box="[835,1294,1724,1748]" pageId="5" pageNumber="495">
|
||
<materialsCitation id="41E83C87FFCBFFD0FCB006DFFAFDF970" box="[835,1289,1724,1748]" country="Indonesia" location="North Sulawesi" pageId="5" pageNumber="495" specimenCount="1" stateProvince="Sulawesi Utara" typeStatus="holotype">
|
||
<emphasis id="C3F4EAC8FFCBFFD0FCB006DFFC14F970" bold="true" box="[835,992,1724,1748]" pageId="5" pageNumber="495">
|
||
<typeStatus id="2E3B8878FFCBFFD0FCB006DFFC89F970" box="[835,893,1724,1748]" pageId="5" pageNumber="495">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<collectingRegion id="3344F838FFCBFFD0FC1B06DFFB66F970" box="[1000,1170,1724,1748]" country="Indonesia" name="Sulawesi Utara" pageId="5" pageNumber="495">North Sulawesi</collectingRegion>
|
||
,
|
||
<collectingCountry id="8997764AFFCBFFD0FB6D06DFFAFDF970" box="[1182,1289,1724,1748]" name="Indonesia" pageId="5" pageNumber="495">Indonesia</collectingCountry>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B99A6551FFCBFFD3FCB0069FFD51FE32" lastPageId="6" lastPageNumber="496" pageId="5" pageNumber="495" type="discussion">
|
||
<paragraph id="F13F36DAFFCBFFD3FCB0069FFD51FE32" blockId="5.[835,1475,1788,2100]" lastBlockId="6.[112,753,190,406]" lastPageId="6" lastPageNumber="496" pageId="5" pageNumber="495">
|
||
<emphasis id="C3F4EAC8FFCBFFD0FCB0069FFC46F8B0" bold="true" box="[835,946,1788,1812]" pageId="5" pageNumber="495">Remarks.</emphasis>
|
||
<taxonomicName id="36804D59FFCBFFD0FC44069EFB65F8B0" authority="Yan & Lee & Ng, 2019" authorityName="Yan & Lee & Ng" authorityYear="2019" box="[951,1169,1788,1812]" class="Malacostraca" family="Epialtidae" genus="Hyastenus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="495" phylum="Arthropoda" rank="species" species="tabolongi" status="sp. nov.">
|
||
<emphasis id="C3F4EAC8FFCBFFD0FC44069EFB65F8B0" box="[951,1169,1788,1812]" italics="true" pageId="5" pageNumber="495">Hyastenus tabolongi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="D8C757B3FFCBFFD0FB6E069EFAEBF8B0" box="[1181,1311,1788,1812]" pageId="5" pageNumber="495" rank="species">new species</taxonomicNameLabel>
|
||
, is most similar to
|
||
<taxonomicName id="36804D59FFCBFFD0FCAE077EFC3FF890" baseAuthorityName="Rathbun" baseAuthorityYear="1911" box="[861,971,1821,1844]" class="Malacostraca" family="Epialtidae" genus="Hyastenus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="495" phylum="Arthropoda" rank="species" species="inermis">
|
||
<emphasis id="C3F4EAC8FFCBFFD0FCAE077EFC84F890" box="[861,880,1821,1844]" italics="true" pageId="5" pageNumber="495">H</emphasis>
|
||
.
|
||
<emphasis id="C3F4EAC8FFCBFFD0FC8F077EFC3FF890" box="[892,971,1821,1844]" italics="true" pageId="5" pageNumber="495">inermis</emphasis>
|
||
</taxonomicName>
|
||
, with both species completely lacking the lateral branchial spines and possessing a swollen gastric region (
|
||
<figureCitation id="69BB2A5FFFCBFFD0FCB8073FFC60F8D0" box="[843,916,1884,1908]" captionStart="Fig" captionStartId="1.[136,171,1463,1485]" captionTargetBox="[136,1474,189,1448]" captionTargetId="figure-250@1.[136,1474,189,1448]" captionTargetPageId="1" captionText="Fig. 1. Hyastenus inermis (Rathbun, 1911), holotype male (15.5 × 10.6 mm) (USNM 41402), Indian Ocean. A, overall dorsal view of carapace; B, overall ventral view; C, lateral view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4577134" httpUri="https://zenodo.org/record/4577134/files/figure.png" pageId="5" pageNumber="495">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="69BB2A5FFFCBFFD0FC51073FFC45F8D0" box="[930,945,1884,1908]" captionStart="Fig" captionStartId="3.[136,171,1936,1958]" captionTargetBox="[136,1474,286,1920]" captionTargetId="figure-7@3.[136,1474,286,1920]" captionTargetPageId="3" captionText="Fig. 3. Hyastenus tabolongi, new species, A–C, holotype male (10.7 × 6.6 mm) (MZB Cru 5010), north Sulawesi; D–F, paratype female (12.3 × 7.2 mm) (ZRC 2019.1324), north Sulawesi. A, D, overall dorsal view of carapace; B, E, overall ventral view; C, F, lateral view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4577142" httpUri="https://zenodo.org/record/4577142/files/figure.png" pageId="5" pageNumber="495">3</figureCitation>
|
||
). The gastric region in
|
||
<taxonomicName id="36804D59FFCBFFD0FB5B073EFADAF8D0" authority="Yan & Lee & Ng, 2019" authorityName="Yan & Lee & Ng" authorityYear="2019" box="[1192,1326,1884,1908]" class="Malacostraca" family="Epialtidae" genus="Hyastenus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="495" phylum="Arthropoda" rank="species" species="tabolongi" status="sp. nov.">
|
||
<emphasis id="C3F4EAC8FFCBFFD0FB5B073EFB4FF8D0" box="[1192,1211,1885,1908]" italics="true" pageId="5" pageNumber="495">H</emphasis>
|
||
.
|
||
<emphasis id="C3F4EAC8FFCBFFD0FB3B073FFADAF8D0" box="[1224,1326,1884,1908]" italics="true" pageId="5" pageNumber="495">tabolongi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="D8C757B3FFCBFFD0FAC8073EFA4AF8D0" box="[1339,1470,1884,1908]" pageId="5" pageNumber="495" rank="species">new species</taxonomicNameLabel>
|
||
, however, is prominently more swollen (
|
||
<figureCitation id="69BB2A5FFFCBFFD0FB0D071FFA4EF830" box="[1278,1466,1916,1940]" captionStart="Fig" captionStartId="3.[136,171,1936,1958]" captionTargetBox="[136,1474,286,1920]" captionTargetId="figure-7@3.[136,1474,286,1920]" captionTargetPageId="3" captionText="Fig. 3. Hyastenus tabolongi, new species, A–C, holotype male (10.7 × 6.6 mm) (MZB Cru 5010), north Sulawesi; D–F, paratype female (12.3 × 7.2 mm) (ZRC 2019.1324), north Sulawesi. A, D, overall dorsal view of carapace; B, E, overall ventral view; C, F, lateral view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4577142" httpUri="https://zenodo.org/record/4577142/files/figure.png" pageId="5" pageNumber="495">Fig. 3A, C, D, F</figureCitation>
|
||
) (versus distinctly less swollen in
|
||
<taxonomicName id="36804D59FFCBFFD0FB4607FEFADDF810" baseAuthorityName="Rathbun" baseAuthorityYear="1911" box="[1205,1321,1949,1972]" class="Malacostraca" family="Epialtidae" genus="Hyastenus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="495" phylum="Arthropoda" rank="species" species="inermis">
|
||
<emphasis id="C3F4EAC8FFCBFFD0FB4607FEFB3CF810" box="[1205,1224,1949,1972]" italics="true" pageId="5" pageNumber="495">H</emphasis>
|
||
.
|
||
<emphasis id="C3F4EAC8FFCBFFD0FB2A07FEFADDF810" box="[1241,1321,1949,1972]" italics="true" pageId="5" pageNumber="495">inermis</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="69BB2A5FFFCBFFD0FAC907FFFA41F810" box="[1338,1461,1948,1972]" captionStart="Fig" captionStartId="1.[136,171,1463,1485]" captionTargetBox="[136,1474,189,1448]" captionTargetId="figure-250@1.[136,1474,189,1448]" captionTargetPageId="1" captionText="Fig. 1. Hyastenus inermis (Rathbun, 1911), holotype male (15.5 × 10.6 mm) (USNM 41402), Indian Ocean. A, overall dorsal view of carapace; B, overall ventral view; C, lateral view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4577134" httpUri="https://zenodo.org/record/4577134/files/figure.png" pageId="5" pageNumber="495">Fig. 1A, C</figureCitation>
|
||
); the posterolateral region of the carapace is proportionately narrower (
|
||
<figureCitation id="69BB2A5FFFCBFFD0FC4607BFFBDBF850" box="[949,1071,2012,2036]" captionStart="Fig" captionStartId="3.[136,171,1936,1958]" captionTargetBox="[136,1474,286,1920]" captionTargetId="figure-7@3.[136,1474,286,1920]" captionTargetPageId="3" captionText="Fig. 3. Hyastenus tabolongi, new species, A–C, holotype male (10.7 × 6.6 mm) (MZB Cru 5010), north Sulawesi; D–F, paratype female (12.3 × 7.2 mm) (ZRC 2019.1324), north Sulawesi. A, D, overall dorsal view of carapace; B, E, overall ventral view; C, F, lateral view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4577142" httpUri="https://zenodo.org/record/4577142/files/figure.png" pageId="5" pageNumber="495">Fig. 3A, D</figureCitation>
|
||
) (versus the posterolateral region of carapace wider in
|
||
<taxonomicName id="36804D59FFCBFFD0FC0E079EFB9EF7B0" baseAuthorityName="Rathbun" baseAuthorityYear="1911" box="[1021,1130,2045,2068]" class="Malacostraca" family="Epialtidae" genus="Hyastenus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="495" phylum="Arthropoda" rank="species" species="inermis">
|
||
<emphasis id="C3F4EAC8FFCBFFD0FC0E079EFBE4F7B0" box="[1021,1040,2045,2068]" italics="true" pageId="5" pageNumber="495">H</emphasis>
|
||
.
|
||
<emphasis id="C3F4EAC8FFCBFFD0FBEF079EFB9EF7B0" box="[1052,1130,2045,2068]" italics="true" pageId="5" pageNumber="495">inermis</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="69BB2A5FFFCBFFD0FB85079FFB32F7B0" box="[1142,1222,2044,2068]" captionStart="Fig" captionStartId="1.[136,171,1463,1485]" captionTargetBox="[136,1474,189,1448]" captionTargetId="figure-250@1.[136,1474,189,1448]" captionTargetPageId="1" captionText="Fig. 1. Hyastenus inermis (Rathbun, 1911), holotype male (15.5 × 10.6 mm) (USNM 41402), Indian Ocean. A, overall dorsal view of carapace; B, overall ventral view; C, lateral view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4577134" httpUri="https://zenodo.org/record/4577134/files/figure.png" pageId="5" pageNumber="495">Fig. 1A</figureCitation>
|
||
); the outer margin of the basal antennal article is more constricted (
|
||
<figureCitation id="69BB2A5FFFCBFFD0FB08087FFA9CF790" box="[1275,1384,2076,2100]" captionStart="Fig" captionStartId="3.[136,171,1936,1958]" captionTargetBox="[136,1474,286,1920]" captionTargetId="figure-7@3.[136,1474,286,1920]" captionTargetPageId="3" captionText="Fig. 3. Hyastenus tabolongi, new species, A–C, holotype male (10.7 × 6.6 mm) (MZB Cru 5010), north Sulawesi; D–F, paratype female (12.3 × 7.2 mm) (ZRC 2019.1324), north Sulawesi. A, D, overall dorsal view of carapace; B, E, overall ventral view; C, F, lateral view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4577142" httpUri="https://zenodo.org/record/4577142/files/figure.png" pageId="5" pageNumber="495">Fig. 3B, E</figureCitation>
|
||
) (versus the outer margin of the basal antennal article is relatively less constricted in
|
||
<taxonomicName id="36804D59FFC8FFD3FEB100BCFE42FF52" baseAuthorityName="Rathbun" baseAuthorityYear="1911" box="[322,438,223,246]" class="Malacostraca" family="Epialtidae" genus="Hyastenus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="496" phylum="Arthropoda" rank="species" species="inermis">
|
||
<emphasis id="C3F4EAC8FFC8FFD3FEB100BCFEA1FF52" box="[322,341,223,246]" italics="true" pageId="6" pageNumber="496">H</emphasis>
|
||
.
|
||
<emphasis id="C3F4EAC8FFC8FFD3FE9500BCFE42FF52" box="[358,438,223,246]" italics="true" pageId="6" pageNumber="496">inermis</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="69BB2A5FFFC8FFD3FE3B00BDFDE9FF52" box="[456,541,222,246]" captionStart="Fig" captionStartId="1.[136,171,1463,1485]" captionTargetBox="[136,1474,189,1448]" captionTargetId="figure-250@1.[136,1474,189,1448]" captionTargetPageId="1" captionText="Fig. 1. Hyastenus inermis (Rathbun, 1911), holotype male (15.5 × 10.6 mm) (USNM 41402), Indian Ocean. A, overall dorsal view of carapace; B, overall ventral view; C, lateral view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4577134" httpUri="https://zenodo.org/record/4577134/files/figure.png" pageId="6" pageNumber="496">Fig. 1B</figureCitation>
|
||
); the chelipeds are proportionately more slender and smooth (
|
||
<figureCitation id="69BB2A5FFFC8FFD3FDDA009DFD6CFEB2" box="[553,664,254,278]" captionStart="Fig" captionStartId="3.[136,171,1936,1958]" captionTargetBox="[136,1474,286,1920]" captionTargetId="figure-7@3.[136,1474,286,1920]" captionTargetPageId="3" captionText="Fig. 3. Hyastenus tabolongi, new species, A–C, holotype male (10.7 × 6.6 mm) (MZB Cru 5010), north Sulawesi; D–F, paratype female (12.3 × 7.2 mm) (ZRC 2019.1324), north Sulawesi. A, D, overall dorsal view of carapace; B, E, overall ventral view; C, F, lateral view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4577142" httpUri="https://zenodo.org/record/4577142/files/figure.png" pageId="6" pageNumber="496">Fig. 3A, D</figureCitation>
|
||
) (versus the chelipeds propodus slightly swollen and granulate in
|
||
<taxonomicName id="36804D59FFC8FFD3FD24017CFF35FEF2" baseAuthorityName="Rathbun" baseAuthorityYear="1911" class="Malacostraca" family="Epialtidae" genus="Hyastenus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="496" phylum="Arthropoda" rank="species" species="inermis">
|
||
<emphasis id="C3F4EAC8FFC8FFD3FD24017CFD1EFE92" box="[727,746,287,310]" italics="true" pageId="6" pageNumber="496">H</emphasis>
|
||
.
|
||
<emphasis id="C3F4EAC8FFC8FFD3FF82015CFF35FEF2" box="[113,193,319,342]" italics="true" pageId="6" pageNumber="496">inermis</emphasis>
|
||
</taxonomicName>
|
||
with;
|
||
<figureCitation id="69BB2A5FFFC8FFD3FEF9015DFE95FEF2" box="[266,353,318,342]" captionStart="Fig" captionStartId="1.[136,171,1463,1485]" captionTargetBox="[136,1474,189,1448]" captionTargetId="figure-250@1.[136,1474,189,1448]" captionTargetPageId="1" captionText="Fig. 1. Hyastenus inermis (Rathbun, 1911), holotype male (15.5 × 10.6 mm) (USNM 41402), Indian Ocean. A, overall dorsal view of carapace; B, overall ventral view; C, lateral view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4577134" httpUri="https://zenodo.org/record/4577134/files/figure.png" pageId="6" pageNumber="496">Fig. 1A</figureCitation>
|
||
); and the G1 is slightly curved with the distal tip sharp (
|
||
<figureCitation id="69BB2A5FFFC8FFD3FEA7013DFE25FED2" box="[340,465,350,374]" captionStart="Fig" captionStartId="2.[113,148,2035,2057]" captionTargetBox="[137,1428,190,2020]" captionTargetId="figure-16@2.[119,1430,176,2023]" captionTargetPageId="2" captionText="Fig. 2. Drawing of left G1 and G2. A–F, Hyastenus inermis (Rathbun, 1911), holotype male (15.5 × 10.6 mm) (USNM 41402), Indian Ocean; G, H, H. tabolongi, new species, holotype male (10.7 × 6.6 mm) (MZB Cru 5010), north Sulawesi. A, G, ventral view; B, ventral view of distal portion; C, H, dorsal view; D, dorsal view of distal portion; E, ventral view of G2; F, dorsal view of G2. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4577138" httpUri="https://zenodo.org/record/4577138/files/figure.png" pageId="6" pageNumber="496">Fig. 2G, H</figureCitation>
|
||
) (versus G1 straight with ear-like lobe at distal tip in
|
||
<taxonomicName id="36804D59FFC8FFD3FE56011CFDE1FE32" baseAuthorityName="Rathbun" baseAuthorityYear="1911" box="[421,533,383,406]" class="Malacostraca" family="Epialtidae" genus="Hyastenus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="496" phylum="Arthropoda" rank="species" species="inermis">
|
||
<emphasis id="C3F4EAC8FFC8FFD3FE56011CFE42FE32" box="[421,438,383,406]" italics="true" pageId="6" pageNumber="496">H</emphasis>
|
||
.
|
||
<emphasis id="C3F4EAC8FFC8FFD3FE36011CFDE1FE32" box="[453,533,383,406]" italics="true" pageId="6" pageNumber="496">inermis</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="69BB2A5FFFC8FFD3FDD6011DFD91FE32" box="[549,613,382,406]" captionStart="Fig" captionStartId="2.[113,148,2035,2057]" captionTargetBox="[137,1428,190,2020]" captionTargetId="figure-16@2.[119,1430,176,2023]" captionTargetPageId="2" captionText="Fig. 2. Drawing of left G1 and G2. A–F, Hyastenus inermis (Rathbun, 1911), holotype male (15.5 × 10.6 mm) (USNM 41402), Indian Ocean; G, H, H. tabolongi, new species, holotype male (10.7 × 6.6 mm) (MZB Cru 5010), north Sulawesi. A, G, ventral view; B, ventral view of distal portion; C, H, dorsal view; D, dorsal view of distal portion; E, ventral view of G2; F, dorsal view of G2. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4577138" httpUri="https://zenodo.org/record/4577138/files/figure.png" pageId="6" pageNumber="496">Fig. 2</figureCitation>
|
||
A–F).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B99A6551FFC8FFD3FF8001DDFD1CFA12" pageId="6" pageNumber="496" type="biology_ecology">
|
||
<paragraph id="F13F36DAFFC8FFD3FF8001DDFD1CFA12" blockId="6.[112,754,446,1462]" pageId="6" pageNumber="496">
|
||
<emphasis id="C3F4EAC8FFC8FFD3FF8001DDFF22FE72" bold="true" box="[115,214,446,470]" pageId="6" pageNumber="496">Ecology.</emphasis>
|
||
Andrew Podzorski (pers. comm.) observed and photographed
|
||
<taxonomicName id="36804D59FFC8FFD3FEE301BCFE6FFE52" authority="Yan & Lee & Ng, 2019" authorityName="Yan & Lee & Ng" authorityYear="2019" box="[272,411,478,502]" class="Malacostraca" family="Epialtidae" genus="Hyastenus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="496" phylum="Arthropoda" rank="species" species="tabolongi" status="sp. nov.">
|
||
<emphasis id="C3F4EAC8FFC8FFD3FEE301BCFED7FE52" box="[272,291,479,502]" italics="true" pageId="6" pageNumber="496">H</emphasis>
|
||
.
|
||
<emphasis id="C3F4EAC8FFC8FFD3FEC701BDFE6FFE52" box="[308,411,478,502]" italics="true" pageId="6" pageNumber="496">tabolongi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="D8C757B3FFC8FFD3FE5F01BCFDC7FE52" box="[428,563,478,502]" pageId="6" pageNumber="496" rank="species">new species</taxonomicNameLabel>
|
||
, from three sites in
|
||
<collectingCountry id="8997764AFFC8FFD3FF7D019DFF0CFDB2" box="[142,248,510,534]" name="Indonesia" pageId="6" pageNumber="496">Indonesia</collectingCountry>
|
||
, namely, the islands of Bunaken and Poh Poh, Tombariri in
|
||
<collectingRegion id="3344F838FFC8FFD3FEF1027DFE5EFD92" box="[258,426,542,566]" country="Indonesia" name="Sulawesi Utara" pageId="6" pageNumber="496">North Sulawesi</collectingRegion>
|
||
, and Pulau Molana in
|
||
<collectingCountry id="8997764AFFC8FFD3FD68027DFD1BFD92" box="[667,751,542,566]" name="Indonesia" pageId="6" pageNumber="496">Ambon</collectingCountry>
|
||
. His many excellent photographs leave no doubt that his specimens are conspecific with the
|
||
<typeStatus id="2E3B8878FFC8FFD3FDE1023CFDB6FDD3" box="[530,578,607,631]" pageId="6" pageNumber="496">type</typeStatus>
|
||
material of
|
||
<taxonomicName id="36804D59FFC8FFD3FD25023CFF23FD32" authority="Yan & Lee & Ng, 2019" authorityName="Yan & Lee & Ng" authorityYear="2019" class="Malacostraca" family="Epialtidae" genus="Hyastenus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="496" phylum="Arthropoda" rank="species" species="tabolongi" status="sp. nov.">
|
||
<emphasis id="C3F4EAC8FFC8FFD3FD25023CFD1DFDD2" box="[726,745,607,630]" italics="true" pageId="6" pageNumber="496">H</emphasis>
|
||
.
|
||
<emphasis id="C3F4EAC8FFC8FFD3FF82021DFF23FD32" box="[113,215,638,662]" italics="true" pageId="6" pageNumber="496">tabolongi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="D8C757B3FFC8FFD3FF16021CFE9CFD32" box="[229,360,638,662]" pageId="6" pageNumber="496" rank="species">new species</taxonomicNameLabel>
|
||
. Amongst the three sites, the species is frequently observed on the island of Bunaken in
|
||
<collectingRegion id="3344F838FFC8FFD3FD4102FDFF25FD72" country="Indonesia" name="Sulawesi Utara" pageId="6" pageNumber="496">North Sulawesi</collectingRegion>
|
||
(
|
||
<figureCitation id="69BB2A5FFFC8FFD3FF1302DDFED5FD72" box="[224,289,702,726]" captionStart="Fig" captionStartId="5.[136,171,1138,1160]" captionTargetBox="[136,1474,190,1123]" captionTargetId="figure-41@5.[136,1474,190,1123]" captionTargetPageId="5" captionText="Fig. 4. Hyastenus tabolongi, new species, observed in situ from Bunaken, Lekuan on different dates. A, B, photographed in November 2011; C, D, photographed in July 2018. Photographs by: AC Podzorski" figureDoi="http://doi.org/10.5281/zenodo.4577144" httpUri="https://zenodo.org/record/4577144/files/figure.png" pageId="6" pageNumber="496">Fig. 4</figureCitation>
|
||
). Despite this,
|
||
<taxonomicName id="36804D59FFC8FFD3FE4D02DCFDB1FD72" authority="Yan & Lee & Ng, 2019" authorityName="Yan & Lee & Ng" authorityYear="2019" box="[446,581,702,726]" class="Malacostraca" family="Epialtidae" genus="Hyastenus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="496" phylum="Arthropoda" rank="species" species="tabolongi" status="sp. nov.">
|
||
<emphasis id="C3F4EAC8FFC8FFD3FE4D02DCFE25FD72" box="[446,465,703,726]" italics="true" pageId="6" pageNumber="496">H</emphasis>
|
||
.
|
||
<emphasis id="C3F4EAC8FFC8FFD3FE2C02DDFDB1FD72" box="[479,581,702,726]" italics="true" pageId="6" pageNumber="496">tabolongi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="D8C757B3FFC8FFD3FDA102DCFD21FD72" box="[594,725,702,726]" pageId="6" pageNumber="496" rank="species">new species</taxonomicNameLabel>
|
||
, is not commonly observed on the south side and only once on the north side of the Bunaken island.
|
||
<taxonomicName id="36804D59FFC8FFD3FDFE029CFD1EFCB2" authority="Yan & Lee & Ng, 2019" authorityName="Yan & Lee & Ng" authorityYear="2019" box="[525,746,766,790]" class="Malacostraca" family="Epialtidae" genus="Hyastenus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="496" phylum="Arthropoda" rank="species" species="tabolongi" status="sp. nov.">
|
||
<emphasis id="C3F4EAC8FFC8FFD3FDFE029CFD1EFCB2" box="[525,746,766,790]" italics="true" pageId="6" pageNumber="496">Hyastenus tabolongi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="D8C757B3FFC8FFD3FF82037CFF0EFC92" box="[113,250,798,822]" pageId="6" pageNumber="496" rank="species">new species</taxonomicNameLabel>
|
||
, has also been photographed from Poh Poh, Tombariri,
|
||
<collectingRegion id="3344F838FFC8FFD3FF1E035DFE61FCF2" box="[237,405,830,854]" country="Indonesia" name="Sulawesi Utara" pageId="6" pageNumber="496">North Sulawesi</collectingRegion>
|
||
(
|
||
<geoCoordinate id="94B4501DFFC8FFD3FE55035EFDD9FCF2" box="[422,557,829,854]" degrees="1" direction="north" minutes="25" orientation="latitude" pageId="6" pageNumber="496" precision="1" seconds="6.00" value="1.4183333">1°25′6.00″N</geoCoordinate>
|
||
<geoCoordinate id="94B4501DFFC8FFD3FDC5035EFD11FCF2" box="[566,741,829,854]" degrees="124" direction="east" minutes="37" orientation="longitude" pageId="6" pageNumber="496" precision="1" seconds="48.95" value="124.630264">124°37′48.95″E</geoCoordinate>
|
||
). In
|
||
<collectingCountry id="8997764AFFC8FFD3FF63033DFF17FCD2" box="[144,227,862,886]" name="Indonesia" pageId="6" pageNumber="496">Ambon</collectingCountry>
|
||
, it has also been photographed at Pulau Molana (
|
||
<geoCoordinate id="94B4501DFFC8FFD3FF88031EFEF6FC32" box="[123,258,893,918]" degrees="3" direction="south" minutes="39" orientation="latitude" pageId="6" pageNumber="496" precision="1" seconds="5.43" value="-3.6515083">3°39′5.43″S</geoCoordinate>
|
||
<geoCoordinate id="94B4501DFFC8FFD3FEFE031EFE36FC32" box="[269,450,893,918]" degrees="128" direction="east" minutes="36" orientation="longitude" pageId="6" pageNumber="496" precision="1" seconds="53.88" value="128.61497">128°36′53.88″E</geoCoordinate>
|
||
). Andrew Podzorski (pers. comm.) notes: “The habitat of this crab is very specialised and to date, absolutely consistent. It is found exclusively on a sparsely branched species of black
|
||
<taxonomicName id="36804D59FFC8FFD3FE0B03BDFEC7FBB2" authority="Tischbierek, 1930" authorityName="Tischbierek" authorityYear="1930" class="Anthozoa" family="Epizoanthidae" genus="Epizoanthus" kingdom="Animalia" order="Zoantharia" pageId="6" pageNumber="496" phylum="Cnidaria" rank="species" species="illoricatus">
|
||
<emphasis id="C3F4EAC8FFC8FFD3FE0B03BDFD04FC52" box="[504,752,990,1014]" italics="true" pageId="6" pageNumber="496">Epizoanthus illoricatus</emphasis>
|
||
<bibRefCitation id="95114B2BFFC8FFD3FF82039DFEC7FBB2" author="Tischbierek H" box="[113,307,1022,1046]" pageId="6" pageNumber="496" pagination="91 - 95" refId="ref5348" refString="Tischbierek H (1930) Zoanthiden auf Wurmrohren. Zoologischer Anzeiger, 91: 91 - 95." type="journal article" year="1930">Tischbierek, 1930</bibRefCitation>
|
||
</taxonomicName>
|
||
[Hexacorallia:
|
||
<taxonomicName id="36804D59FFC8FFD3FE2D039DFD89FBB2" box="[478,637,1022,1046]" class="Anthozoa" family="Epizoanthidae" kingdom="Animalia" order="Zoantharia" pageId="6" pageNumber="496" phylum="Cnidaria" rank="family">Epizoanthidae</taxonomicName>
|
||
]. The crab places many polyps of
|
||
<taxonomicName id="36804D59FFC8FFD3FE87047DFD84FB92" authorityName="Tischbierek" authorityYear="1930" box="[372,624,1054,1078]" class="Anthozoa" family="Epizoanthidae" genus="Epizoanthus" kingdom="Animalia" order="Zoantharia" pageId="6" pageNumber="496" phylum="Cnidaria" rank="species" species="illoricatus">
|
||
<emphasis id="C3F4EAC8FFC8FFD3FE87047DFD84FB92" box="[372,624,1054,1078]" italics="true" pageId="6" pageNumber="496">Epizoanthus illoricatus</emphasis>
|
||
</taxonomicName>
|
||
all over its body, so much so it is almost impossible to distinguish the crab from the colony (
|
||
<figureCitation id="69BB2A5FFFC8FFD3FE94043DFE49FBD2" box="[359,445,1118,1142]" captionStart="Fig" captionStartId="5.[136,171,1138,1160]" captionTargetBox="[136,1474,190,1123]" captionTargetId="figure-41@5.[136,1474,190,1123]" captionTargetPageId="5" captionText="Fig. 4. Hyastenus tabolongi, new species, observed in situ from Bunaken, Lekuan on different dates. A, B, photographed in November 2011; C, D, photographed in July 2018. Photographs by: AC Podzorski" figureDoi="http://doi.org/10.5281/zenodo.4577144" httpUri="https://zenodo.org/record/4577144/files/figure.png" pageId="6" pageNumber="496">Fig. 4B</figureCitation>
|
||
). The only suggestion that a crab is there, is a thickening of the branch. This species of
|
||
<taxonomicName id="36804D59FFC8FFD3FF8004FDFE9CFB12" authorityName="Tischbierek" authorityYear="1930" box="[115,360,1182,1206]" class="Anthozoa" family="Epizoanthidae" genus="Epizoanthus" kingdom="Animalia" order="Zoantharia" pageId="6" pageNumber="496" phylum="Cnidaria" rank="species" species="illoricatus">
|
||
<emphasis id="C3F4EAC8FFC8FFD3FF8004FDFE9CFB12" box="[115,360,1182,1206]" italics="true" pageId="6" pageNumber="496">Epizoanthus illoricatus</emphasis>
|
||
</taxonomicName>
|
||
is common across western
|
||
<collectingCountry id="8997764AFFC8FFD3FD7004FDFD19FB12" box="[643,749,1182,1206]" name="Indonesia" pageId="6" pageNumber="496">Indonesia</collectingCountry>
|
||
, growing up to
|
||
<quantity id="36789B3FFFC8FFD3FEEB04DDFEAAFB72" box="[280,350,1214,1238]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="496" unit="cm" value="15.0">15 cm</quantity>
|
||
high, and generally occurs in areas that are influenced by strong currents. The zoanthid can also be found on flat rocky bottoms and on drop-offs. The only locations where the crab lives on the colonies, is on dropoffs, where the zoanthid grows in small hollows or caves on the face of the drop-offs. Here, the colonies are protected from direct currents.
|
||
<taxonomicName id="36804D59FFC8FFD3FEA7051CFE37FA32" authorityName="White" authorityYear="1847" box="[340,451,1407,1430]" class="Malacostraca" family="Epialtidae" genus="Hyastenus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="496" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C3F4EAC8FFC8FFD3FEA7051CFE37FA32" box="[340,451,1407,1430]" italics="true" pageId="6" pageNumber="496">Hyastenus</emphasis>
|
||
</taxonomicName>
|
||
specimens have been found from depths of
|
||
<quantity id="36789B3FFFC8FFD3FEE805FDFE62FA12" box="[283,406,1438,1462]" metricMagnitude="1" metricUnit="m" metricValue="2.05" metricValueMax="3.0" metricValueMin="1.1" pageId="6" pageNumber="496" unit="m" value="20.5" valueMax="30.0" valueMin="11.0">11 to 30 m</quantity>
|
||
, but most often around
|
||
<specimenCount id="E786FD53FFC8FFD3FD6E05FDFD29FA12" box="[669,733,1438,1462]" pageId="6" pageNumber="496" type="generic">
|
||
<quantity id="36789B3FFFC8FFD3FD6E05FDFD2DFA12" box="[669,729,1438,1462]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="496" unit="m" value="20.0">20 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
”
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B99A6551FFC8FFD3FF8205BDFD3CF9B2" pageId="6" pageNumber="496" type="distribution">
|
||
<paragraph id="F13F36DAFFC8FFD3FF8205BDFD3CF9B2" blockId="6.[113,752,1502,1558]" pageId="6" pageNumber="496">
|
||
<emphasis id="C3F4EAC8FFC8FFD3FF8205BDFEF0FA52" bold="true" box="[113,260,1502,1526]" pageId="6" pageNumber="496">Distribution.</emphasis>
|
||
Currently only known from its
|
||
<typeStatus id="2E3B8878FFC8FFD3FD9005BCFD65FA53" box="[611,657,1503,1527]" pageId="6" pageNumber="496">type</typeStatus>
|
||
locality,
|
||
<collectingRegion id="3344F838FFC8FFD3FF80059DFEE3F9B2" box="[115,279,1534,1558]" country="Indonesia" name="Sulawesi Utara" pageId="6" pageNumber="496">north Sulawesi</collectingRegion>
|
||
, and Pulau Molana,
|
||
<collectingCountry id="8997764AFFC8FFD3FE0A059DFDB8F9B2" box="[505,588,1534,1558]" name="Indonesia" pageId="6" pageNumber="496">Ambon</collectingCountry>
|
||
,
|
||
<collectingCountry id="8997764AFFC8FFD3FDAA059DFD30F9B2" box="[601,708,1534,1558]" name="Indonesia" pageId="6" pageNumber="496">Indonesia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |