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<document ID-CLB-Dataset="22826" ID-DOI="http://dx.doi.org/10.3897/alpento.4.50703" ID-GBIF-Dataset="a9e0336c-85fe-4d7c-8998-fadb8b1e4a29" ID-Pensoft-Pub="2535-0889-4-39" ID-Pensoft-UUID="73E3CBB8841D53E98BB78BA05DDC69C9" ID-ZooBank="D9502F1AAEC04B0F845C87D86632AF17" ModsDocID="2535-0889-4-39" checkinTime="1587713590471" checkinUser="pensoft" docAuthor="Huemer, Peter" docDate="2020" docId="767C6E4F29F85CE0A3C15512EFC736B1" docLanguage="en" docName="AlpEntomol 4: 39-63" docOrigin="Alpine Entomology 4" docSource="http://dx.doi.org/10.3897/alpento.4.50703" docTitle="Caryocolum messneri Huemer 2020, sp. nov." docType="treatment" docUuid="BEDFE053-00A6-4904-8F94-E0EF8AF4E716" docUuidSource="ZooBank" docVersion="6" id="73E3CBB8841D53E98BB78BA05DDC69C9" lastPageNumber="39" masterDocId="73E3CBB8841D53E98BB78BA05DDC69C9" masterDocTitle="Integrative revision of the Caryocolum schleichi species group - a striking example of a temporally changing species concept (Lepidoptera, Gelechiidae)" masterLastPageNumber="63" masterPageNumber="39" pageNumber="39" updateTime="1732751505677" updateUser="ExternalLinkService">
<mods:mods id="354A4B4839D51281933F2A230806369A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A3953A066026305AAC6A7ACA709E0940">
<mods:title id="5E443D3FEAD1FF9982BD7FD9AB1D5873">Integrative revision of the Caryocolum schleichi species group - a striking example of a temporally changing species concept (Lepidoptera, Gelechiidae)</mods:title>
</mods:titleInfo>
<mods:name id="BF2D3FA618D5E62D2FADCE553DD79E2D" type="personal">
<mods:role id="11C0A036A596A3972103D01439DEFDB2">
<mods:roleTerm id="308E98AC4E6A47E778B0C07F3DD35C74">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CF0DE36BA8B1B327DF0D518AFD52A800">Huemer, Peter</mods:namePart>
<mods:affiliation id="8E20033039306BB81B24BA0B47E62727">Tiroler Landesmuseen Betriebgsges. m. b. H., Sammlungs- und Forschungszentrum, Naturwissenschaftliche Sammlungen, Krajnc-Str. 1, A- 6060 Hall in Tirol, Austria</mods:affiliation>
<mods:nameIdentifier id="95F6968A53C61384E4D529E344AA81DD" type="orcid">https://orcid.org/0000-0002-0630-545X</mods:nameIdentifier>
<mods:nameIdentifier id="5453CFF968AE251A4CE1039659E8608F" type="email">p.huemer@tiroler-landesmuseen.at</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="2A5D831E0ACDEB1BC738C645027020E1">text</mods:typeOfResource>
<mods:relatedItem id="4964366E28E0D57C5C114637A4752316" type="host">
<mods:titleInfo id="BA615040A976698BC4A3AD4F581BA096">
<mods:title id="9018C53BD80A0ACA4099C0216E19D14E">Alpine Entomology</mods:title>
</mods:titleInfo>
<mods:part id="9002A571D7B3C3F2C17D0F93710962E4">
<mods:date id="328D84A89E3208C7A1CB122A4E55EA00">2020</mods:date>
<mods:detail id="B74901ADD061228C7D5D9B784DD90310" type="volume">
<mods:number id="2E019FA5373E9A36E95ECB4C81964D5C">4</mods:number>
</mods:detail>
<mods:extent id="C1F6BBB66B1C7CB1BA0059966131484F" unit="page">
<mods:start id="489FA88CC0B6C6DF3C9F171F56FF6035">39</mods:start>
<mods:end id="11B3E0877A2FA01A4D03968A1EDF24EF">63</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="3A59CE94BC2C4B6274F5343FF928BA4F">
<mods:url id="51CAE508CDAEE08255611F32DE660F14">http://dx.doi.org/10.3897/alpento.4.50703</mods:url>
</mods:location>
<mods:classification id="F1BA98FAB743EFDC709F97A912B54CCC">journal article</mods:classification>
<mods:identifier id="095B5FEE6CB52C990DB8546BB1E64005" type="DOI">http://dx.doi.org/10.3897/alpento.4.50703</mods:identifier>
<mods:identifier id="FDC4FCDC4A601A6D54B5E20E0FD32D9E" type="Pensoft-Pub">2535-0889-4-39</mods:identifier>
<mods:identifier id="6F1D40D3D5C18549F44BDB381272A712" type="ZooBank">D9502F1AAEC04B0F845C87D86632AF17</mods:identifier>
<mods:identifier id="65D0E2CD5EF83A7F99770C17FDA2751B" type="Pensoft-UUID">73E3CBB8841D53E98BB78BA05DDC69C9</mods:identifier>
</mods:mods>
<treatment id="767C6E4F29F85CE0A3C15512EFC736B1" ID-GBIF-Taxon="163737596" LSID="urn:lsid:zoobank.org:act:BEDFE053-00A6-4904-8F94-E0EF8AF4E716" httpUri="http://treatment.plazi.org/id/767C6E4F29F85CE0A3C15512EFC736B1" lastPageNumber="39" pageId="0" pageNumber="39" scope_family="Gelechiidae" scope_order="Lepidoptera">
<subSubSection id="3DEA368274CD204D0A011347E069DD7E" pageId="0" pageNumber="39" type="nomenclature">
<paragraph id="B19E896217820FC7D133E736A14B84AF" pageId="0" pageNumber="39">
<taxonomicName id="63A0C3BA25594813CE3A41C7010B9C80" ID-CoL="98ZS4" LSID="767C6E4F-29F8-5CE0-A3C1-5512EFC736B1" authority="Huemer, 2020" authorityName="Huemer" authorityYear="2020" class="Insecta" family="Gelechiidae" genus="Caryocolum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caryocolum messneri" order="Lepidoptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="messneri" status="sp. nov.">Caryocolum messneri</taxonomicName>
<taxonomicNameLabel id="D29D28B76CF3D7B0D8650E6D5FC925E5" pageId="0" pageNumber="39">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="539A5A05C7309909774668BF5A0CDEB1" pageId="0" pageNumber="39" type="materials_examined">
<paragraph id="CE9484C15C65C60CC2AE5D4FA51C86A4" pageId="0" pageNumber="39">Material examined.</paragraph>
<paragraph id="8655CF16D06C093C2AD6563B4963B48A" pageId="0" pageNumber="39">
<materialsCitation id="5F137AF2238741D0CBFA557FA2EDFC8F" ID-GBIF-Occurrence="2609799210" collectingDate="2020-01-01" collectingDateMax="2020-12-31" collectingDateMin="2020-01-01" collectionCode="TLMF" collectorName="Huemer &amp; P. Huemer" country="Italy" elevation="0" latitude="46.67417" location="Suedtirol / Schnals" longitude="10.945" municipality="Suedtirol / Schnals" specimenCode="TLMF Lep 16694, GEL 1255" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="BAD94870524A0464EA1315D07770DC62" bold="true" pageId="0" pageNumber="39">
<typeStatus id="6D3BCFB1CB4CB6DA13D5AFAD003D3BA3">Holotype</typeStatus>
</emphasis>
<specimenCount id="4C0EE1DF2842AEEC7EA93B03E854C51C" type="male">♂</specimenCount>
, &quot;
<collectingCountry id="A873E085A05D38F897F7DF95989219C1">ITALIA</collectingCountry>
sept.
<collectingMunicipality id="7C275B611BE4BCC99021625A47F54019">
<normalizedToken id="16E1768E3A3EC121B854EBDFDD7E5C3D" originalValue="Südtirol">Suedtirol</normalizedToken>
/ Schnals
</collectingMunicipality>
, Neu-Ratteis, / Fuchsberg,
<elevation id="B37D3127FCB158672491EA9C87E3A871">980 m</elevation>
/
<geoCoordinate id="7ED3865FC95ABC41BB268EBA9D776D28">10°56'42&quot;E</geoCoordinate>
,
<geoCoordinate id="A6F0D12943D0748709452F2740C22950">46°40'27&quot;N</geoCoordinate>
/
<elevation id="2FC7856751343F3A971619D8820F24D7">650 m,</elevation>
<collectingDate id="D14314214EDDA7E72FC04B85EF2782A7">28.8.2014</collectingDate>
/ leg.
<collectorName id="521BC0302EDDA307D7BBA761750D7288">Huemer</collectorName>
&quot; &quot;DNA Barcode /
<specimenCode id="9CD44166FC7C4C7B32872944E4CB25B7">TLMF Lep 16694</specimenCode>
&quot; &quot;
<collectorName id="7B704ED937733DA85959E962743D6A62">P. Huemer</collectorName>
/
<specimenCode id="4270F002CAE2291B048FC3FA0DE45F9A">GEL 1255</specimenCode>
♂&quot; (
<collectionCode id="581FE3CE43488ACE92049AB3C540A85A">TLMF</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="BA38881A085B4D25FEF20922ABE3BFF0" pageId="0" pageNumber="39">
<emphasis id="D95DE775B40EED0D98DCE344012808D7" bold="true" pageId="0" pageNumber="39">Paratypes</emphasis>
: Italy: 1 ♂, same data as holotype, but DNA Barcode TLMF Lep 16694; 1 ♂, 3 ♀♀,
<normalizedToken id="68DFDFDB1BF46CA59CAF4B049E030578" originalValue="Südtirol">Suedtirol</normalizedToken>
province, Schnalstal, unt. Ladurner, 650 m, 31 Aug. 2019, leg. Huemer, DNA Barcodes TLMF Lep 23981, 23982, 4 gen. slides in glycerin; 1 ♂, Trento province, Villamontagna, 600 m, 17 Aug. 1982, leg. Burmann, gen. slide GU 86/035 ♂ P. Huemer (all TLMF). Bulgaria: 1 ♀, Trigrad env., 1240 m, 4 Aug. 2013, leg. Karsholt &amp; Zlatkov, DNA Barcode TLMF Lep 21297 (ZMUC). Greece: 1 ♀, Epirus, Mitsikeli Mt. near Joannina, 1400 m,
<geoCoordinate id="61B54843F5B761CF5E718E1B4E773FD2" degrees="39" direction="north" minutes="46" orientation="latitude" precision="15" seconds="15" value="39.770832">39°46'15&quot;N</geoCoordinate>
,
<geoCoordinate id="DC22A0FB03DAC91A9943387C404B9CF4" degrees="20" direction="east" minutes="48" orientation="longitude" precision="15" seconds="20" value="20.805555">20°48'20&quot;E</geoCoordinate>
, 8 Jul. 2005, leg. Mayr (RCTM).
</paragraph>
<caption id="E43CD7E7D743897E0F849AAFCE4BEB17" doi="10.3897/alpento.4.50703.figures36-39" httpUri="https://binary.pensoft.net/fig/403197" pageId="0" pageNumber="39" start="Figures 36–39" startId="F12">
<paragraph id="72D2F9194E5ED7C4B737C8EABA7B2EF4" pageId="0" pageNumber="39">
<emphasis id="B0E1D372B0E618BEC69A4B4F22A99AC5" bold="true" pageId="0" pageNumber="39">Figures 36-39.</emphasis>
Female genitalia (segment VIII).
<emphasis id="CEE9750B911B5D3E927BF8F5B826FD5A" bold="true" pageId="0" pageNumber="39">36</emphasis>
<taxonomicName id="F29EC794DB8284F692A03975E29AF5F5" baseAuthorityName="Christoph" baseAuthorityYear="1872" class="Insecta" family="Gelechiidae" genus="Caryocolum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caryocolum schleichi" order="Lepidoptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="schleichi">
<emphasis id="A3E9ACA831BF1F54ADEEFBF6681B9F2F" italics="true" pageId="0" pageNumber="39">Caryocolum schleichi</emphasis>
</taxonomicName>
, Syria, slide GU 86/277 P. Huemer;
<emphasis id="679A8E2D5976203842506CB191E11389" bold="true" pageId="0" pageNumber="39">37</emphasis>
<taxonomicName id="3494568C3B76E43F55CF8D3EDD8E5DE8" lsidName="C. dianthella" pageId="0" pageNumber="39" rank="species" species="dianthella">
<emphasis id="970AA57B87E632787B670EE23C672DAC" italics="true" pageId="0" pageNumber="39">C. dianthella</emphasis>
</taxonomicName>
, Spain, slide GEL 1285 P. Huemer;
<emphasis id="95564D14F981343CA58FABD2E7393C18" bold="true" pageId="0" pageNumber="39">38</emphasis>
<taxonomicName id="1B79DF437CD9A2FDA2DDFB4465076277" lsidName="C. improvisella" pageId="0" pageNumber="39" rank="species" species="improvisella">
<emphasis id="E8B7ABCF9904F52219DD67FCB184C120" italics="true" pageId="0" pageNumber="39">C. improvisella</emphasis>
</taxonomicName>
, Italy, slide GEL 1291 P. Huemer;
<emphasis id="4BEFDA0B150388DE005B4C281EE9705E" bold="true" pageId="0" pageNumber="39">39</emphasis>
<taxonomicName id="BE921B49AEFA08F55A2A964BF9206426" lsidName="C. arenariella" pageId="0" pageNumber="39" rank="species" species="arenariella">
<emphasis id="6A530E17E6EB0E685293F8D31921C909" italics="true" pageId="0" pageNumber="39">C. arenariella</emphasis>
</taxonomicName>
, Sweden, slide GEL 1292 P. Huemer.
</paragraph>
</caption>
<caption id="FAA30EB37DE2378CF3C11A1D6838C624" doi="10.3897/alpento.4.50703.figures40-42" httpUri="https://binary.pensoft.net/fig/403198" pageId="0" pageNumber="39" start="Figures 40–42" startId="F13">
<paragraph id="44F3E87A66C13F2BB5940DE06B147F7B" pageId="0" pageNumber="39">
<emphasis id="8D71F1A2C1FCBF377B3FA7962AD3CDE5" bold="true" pageId="0" pageNumber="39">Figures 40-42.</emphasis>
Female genitalia (segment VIII).
<emphasis id="11F8BD69C3E5BF603E229817C4753415" bold="true" pageId="0" pageNumber="39">40</emphasis>
<taxonomicName id="4D48A34A0D4EF5F9BD1D21B60A75AEA0" authorityName="Huemer" authorityYear="2020" class="Insecta" family="Gelechiidae" genus="Caryocolum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caryocolum messneri" order="Lepidoptera" pageId="0" pageNumber="39" phylum="Arthropoda" rank="species" species="messneri">
<emphasis id="159D8B3CC07241A2A0BD0516586E127D" italics="true" pageId="0" pageNumber="39">Caryocolum messneri</emphasis>
</taxonomicName>
, paratype, Italy, slide GEL 1295 P. Huemer;
<emphasis id="CD3FBAF05EE30780E9529440AF07590F" bold="true" pageId="0" pageNumber="39">41</emphasis>
<taxonomicName id="E3568E6E3820E630F85F8BA86E5013F6" lsidName="C. lamai" pageId="0" pageNumber="39" rank="species" species="lamai">
<emphasis id="8864C45957CCAB57E0250D1F895F8E35" italics="true" pageId="0" pageNumber="39">C. lamai</emphasis>
</taxonomicName>
, paratype, Italy, slide GEL 1286 P. Huemer;
<emphasis id="A48C9824A3ED267456B127A4D2C45D53" bold="true" pageId="0" pageNumber="39">42</emphasis>
<taxonomicName id="B1AC5F8D0F1C092B7E287AFE418E2DD6" lsidName="C. habeleri" pageId="0" pageNumber="39" rank="species" species="habeleri">
<emphasis id="AD214B9B7CD11F730C16E9AC82BA63D4" italics="true" pageId="0" pageNumber="39">C. habeleri</emphasis>
</taxonomicName>
, paratype, Germany, slide M 1703.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="3E7FD3DB4AB4FC9441FA959EFC869990" pageId="0" pageNumber="39" type="diagnosis">
<paragraph id="8D82088E3588C0EEBDB977B9E990F876" pageId="0" pageNumber="39">Diagnosis.</paragraph>
<paragraph id="3F52B4275E2BA17D583A5875B090FA57" pageId="0" pageNumber="39">
<taxonomicName id="79CD75BA8651077BF212FE720A8C3E86" lsidName="C. messneri" pageId="0" pageNumber="39" rank="species" species="messneri">
<emphasis id="14201702052710A14F95F55A9F1838D1" italics="true" pageId="0" pageNumber="39">C. messneri</emphasis>
</taxonomicName>
differs from
<taxonomicName id="01DDBE681ECD5BAF65E3022D5F8BAE39" lsidName="C. schleichi" pageId="0" pageNumber="39" rank="species" species="schleichi">
<emphasis id="0437D7BA858E44CF16061BC97CBFBD5E" italics="true" pageId="0" pageNumber="39">C. schleichi</emphasis>
</taxonomicName>
by the brown rather than cream-white head, thorax and tegulae. It is usually larger than
<taxonomicName id="ED7280A86EF5F397098A31FB59D1B5BA" lsidName="C. arenariella" pageId="0" pageNumber="39" rank="species" species="arenariella">
<emphasis id="B61FFADA14C5FDFEE2E33E7066252315" italics="true" pageId="0" pageNumber="39">C. arenariella</emphasis>
</taxonomicName>
and
<taxonomicName id="BFBB3CC7EB43BEBE31DA874147EA8C07" lsidName="C. lamai" pageId="0" pageNumber="39" rank="species" species="lamai">
<emphasis id="C449FCF17B46D18C8087392E6BB1ED7F" italics="true" pageId="0" pageNumber="39">C. lamai</emphasis>
</taxonomicName>
and the medial spot is less sharply delimited than in
<taxonomicName id="25C6698A9CEC0112F77009AB4A44A36A" lsidName="C. dianthella" pageId="0" pageNumber="39" rank="species" species="dianthella">
<emphasis id="E63422B071475B91095E870D858B2BC6" italics="true" pageId="0" pageNumber="39">C. dianthella</emphasis>
</taxonomicName>
. Otherwise,
<taxonomicName id="F9031E21E9E97B2AA6D3E64151E0D3E2" lsidName="C. messneri" pageId="0" pageNumber="39" rank="species" species="messneri">
<emphasis id="45A6DEA0D78BE8E716BA11164F0980D3" italics="true" pageId="0" pageNumber="39">C. messneri</emphasis>
</taxonomicName>
cannot be reliably separated from the other species of the complex by external characters, though the usually well developed rusty-brown mottling seems unique. The male genitalia are characterized in particular by the moderately long, pointed dorsal process of the valva, a character which is only shared with
<taxonomicName id="06B89D192A02DB8445887779BBF45E7E" lsidName="C. improvisella" pageId="0" pageNumber="39" rank="species" species="improvisella">
<emphasis id="F8E5F7D21D244A2564B2CE26BF15E473" italics="true" pageId="0" pageNumber="39">C. improvisella</emphasis>
</taxonomicName>
,
<taxonomicName id="1C7D404F88D4F8CAAC32BE490A9E49C5" lsidName="C. arenariella" pageId="0" pageNumber="39" rank="species" species="arenariella">
<emphasis id="21F3F19352474FA76AF8B3BA5D9E9D78" italics="true" pageId="0" pageNumber="39">C. arenariella</emphasis>
</taxonomicName>
, and
<taxonomicName id="D9B645E2B4E25514B5A7CC383A849713" lsidName="C. lamai" pageId="0" pageNumber="39" rank="species" species="lamai">
<emphasis id="32C9341C680B6E8C6AB577C475FAF4D8" italics="true" pageId="0" pageNumber="39">C. lamai</emphasis>
</taxonomicName>
. However, this process is distinctly shorter than in
<taxonomicName id="C60CFE22DB976A26030E9E703587D0FE" lsidName="C. arenariella" pageId="0" pageNumber="39" rank="species" species="arenariella">
<emphasis id="D68C76451D3C0EB97163A380C65621D4" italics="true" pageId="0" pageNumber="39">C. arenariella</emphasis>
</taxonomicName>
and
<taxonomicName id="2CEADA8772573EB8D4F4E9D2E26872B7" lsidName="C. lamai" pageId="0" pageNumber="39" rank="species" species="lamai">
<emphasis id="FFCEE836182001DED0A6773F2C1BEF17" italics="true" pageId="0" pageNumber="39">C. lamai</emphasis>
</taxonomicName>
. The species further differs from
<taxonomicName id="3DF3DD9E4C85C4B2796D58055174A86C" lsidName="C. improvisella" pageId="0" pageNumber="39" rank="species" species="improvisella">
<emphasis id="9BC19F1AAD44A3718B261F3FCD5F98DA" italics="true" pageId="0" pageNumber="39">C. improvisella</emphasis>
</taxonomicName>
in several characters such as the more slender valva with parallel outer edges and a distodorsal bulge, a shorter dorsal process, and the distinctly longer latero-medial processes of the vinculum. The female genitalia are hardly discernible from several other species of the
<taxonomicName id="F27DEBFE7E8C8072CF26B589862AE8AB" lsidName="C. schleichi" pageId="0" pageNumber="39" rank="species" species="schleichi">
<emphasis id="6A77C1D3D34221B9366BB97D3F466401" italics="true" pageId="0" pageNumber="39">C. schleichi</emphasis>
</taxonomicName>
species group though the shortly crescent-shaped base of the signum in combination with the moderately slender signum hook seem characteristic and differ from all other species except for
<taxonomicName id="380F7E3A8DEFF90F5D8790CC5DEEE6C1" lsidName="C. improvisella" pageId="0" pageNumber="39" rank="species" species="improvisella">
<emphasis id="7C18F6A5C38E6FC8D1B6AC8129746E45" italics="true" pageId="0" pageNumber="39">C. improvisella</emphasis>
</taxonomicName>
. However, the individual variation of these characters is insufficiently documented due to lack of material.
</paragraph>
</subSubSection>
<subSubSection id="D31D91F3C843608211D7CF37E672374E" pageId="0" pageNumber="39" type="description">
<paragraph id="DFD533D16E4B719681C7CA3813A75E21" pageId="0" pageNumber="39">Description.</paragraph>
<paragraph id="51C290581AD3526ED28087DB5A9250D5" pageId="0" pageNumber="39">
Adult (Fig.
<figureCitation id="45993AD7FDA88FCB4E982AEC699D05EB" captionStart="Figures 9–12" captionStartId="F4" captionText="Figures 9 - 12. Adults. 9 Caryocolum lamai, paratype, male, Italy; 10 C. lamai, paratype, male, Italy; 11 C. messneri, paratype, male, Italy; 12 C. habeleri, paratype, male, France." figureDoi="10.3897/alpento.4.50703.figures9-12" httpUri="https://binary.pensoft.net/fig/403189" pageId="0" pageNumber="39">11</figureCitation>
). Forewing length ♂ 5.5-6.0 mm, ♀ 5.0 mm. Head with dark grey-brown vertex, mottled with white, frons cream-white; first segment of labial palpus dark brown, second segment cream-white on inner and upper surface, ventral and outer surface predominantly dark brown, dark brown mixed with few whitish scales on segment three; antenna black, weakly ringed paler. Thorax and tegula dark brown anteriorly, posterior half cream with rusty brown mottling. Abdomen cream-white with some grey on ventral surface. Forewing dark brown, mottled with white on dorsal margin; basal two thirds with rusty-brown mottling, particularly white markings; white transverse fascia from fold to costa at one-fifth; white medial spot occasionally extended towards costa and dorsal margin; white costal and tornal spots separate; fringes basally dark brown, distal part paler. Hindwing light grey.
</paragraph>
</subSubSection>
<subSubSection id="AB8B05E5B6E46A9DA210E1BCA11A62DB" pageId="0" pageNumber="39" type="variation">
<paragraph id="D31EC722CB666B5031079546E07336C1" pageId="0" pageNumber="39">Variation.</paragraph>
<paragraph id="9FD7B0252695C7FFB964DC0CFA684B4F" pageId="0" pageNumber="39">In the limited available material, the extent of rusty-brown mottling shows some variation.</paragraph>
</subSubSection>
<subSubSection id="796051C29BAD430183DD27075D3FC7CA" pageId="0" pageNumber="39" type="male genitalia">
<paragraph id="E72C1323982F95A3D9E5D7356F0233E2" pageId="0" pageNumber="39">Male genitalia</paragraph>
<paragraph id="0CD4E25E203B7103CF6F6A92394C827F" pageId="0" pageNumber="39">
(Figs
<figureCitation id="E99E287EBCD29D57B094379B433EAF01" captionStart="Figures 17–20" captionStartId="F6" captionText="Figures 17 - 20. Male genitalia. 17 Caryocolum arenariella, Russia, slide GEL 1260 P. Huemer; 18 C. messneri, holotype, Italy, slide GEL 1255; 19 C. lamai, paratype, Italy, slide GEL 1258 P. Huemer; 21 C. habeleri, paratype, France, slide GEL 1289 P. Huemer." figureDoi="10.3897/alpento.4.50703.figures17-20" httpUri="https://binary.pensoft.net/fig/403191" pageId="0" pageNumber="39">18</figureCitation>
,
<figureCitation id="52F897303B5AC61D8C763EA6E392A173" captionStart="Figures 25–28" captionStartId="F8" captionText="Figures 25 - 28. Male genitalia (details vinculum-valva-complex). 25 Caryocolum arenariella, Russia, slide GEL 1260 P. Huemer; 26 C. messneri, holotype, Italy, slide GEL 1255; 27 C. lamai, paratype, Italy, slide GEL 1258 P. Huemer; 28 C. habeleri, paratype, France, slide GEL 1289 P. Huemer." figureDoi="10.3897/alpento.4.50703.figures25-28" httpUri="https://binary.pensoft.net/fig/403193" pageId="0" pageNumber="39">26</figureCitation>
). Uncus broadly sub-quadrangular, posterior corners rounded; lateral sclerites of gnathos distinct, medial part with large minutely spined culcitula; tegumen weakly widened anteriorly, with slightly emarginated anterior margin; transtilla sclerotized, longitudinally folded; pedunculus large, sub-triangular, with sclerotized inner edge; valva nearly straight, long, moderately slender distal part with parallel edges, except for bulged disto-dorsal edge, apex with two processes, stout and moderately long and pointed dorsal process extended to about middle of uncus, broadly digitate ventral process at about right-angles to and slightly longer than dorsal process; sacculus moderately long, slender, with parallel margins, apically pointed; posterior margin of vinculum deeply incised medially, with pair of moderately long digitate processes, pair of latero-medial processes moderately broad, digitate, distinctly projected; saccus slightly longer than valva, slender, gradually tapered to apex; phallus long, slender, nearly straight, apically with area of small cornuti.
</paragraph>
</subSubSection>
<subSubSection id="1ECA3A83FCE30FF667FC709AE7E340AE" pageId="0" pageNumber="39" type="female genitalia">
<paragraph id="D7F855781AFCAB349C466AE929B4D382" pageId="0" pageNumber="39">Female genitalia</paragraph>
<paragraph id="A467F444D7243453C5E810CC8CB71441" pageId="0" pageNumber="39">
(Figs
<figureCitation id="B96ACB80A116D9EB73529A27ED69A463" captionStart="Figures 33–35" captionStartId="F11" captionText="Figures 33 - 35. Female genitalia. 33 Caryocolum messneri, paratype, Italy, slide GEL 1295 P. Huemer; 34 C. lamai, paratype, Italy, slide GEL 1286 P. Huemer; 35 C. habeleri, paratype, Germany, slide M 1703." figureDoi="10.3897/alpento.4.50703.figures33-35" httpUri="https://binary.pensoft.net/fig/403196" pageId="0" pageNumber="39">33</figureCitation>
,
<figureCitation id="D868EFBC7F89EDCE71F7A890BA06E009" captionStart="Figures 40–42" captionStartId="F13" captionText="Figures 40 - 42. Female genitalia (segment VIII). 40 Caryocolum messneri, paratype, Italy, slide GEL 1295 P. Huemer; 41 C. lamai, paratype, Italy, slide GEL 1286 P. Huemer; 42 C. habeleri, paratype, Germany, slide M 1703." figureDoi="10.3897/alpento.4.50703.figures40-42" httpUri="https://binary.pensoft.net/fig/403198" pageId="0" pageNumber="39">40</figureCitation>
,
<figureCitation id="D31A2D19CE25BD67330BB3614B5D6B9C" captionStart="Figures 47–49" captionStartId="F15" captionText="Figures 47 - 49. Female genitalia (signum). 47 Caryocolum messneri, paratype, Italy, slide GEL 1295 P. Huemer; 48 C. lamai, paratype, Italy, slide GEL 1286 P. Huemer; 49 C. habeleri, paratype, Germany, slide M 1703." figureDoi="10.3897/alpento.4.50703.figures47-49" httpUri="https://binary.pensoft.net/fig/403200" pageId="0" pageNumber="39">47</figureCitation>
). Apophysis posterior almost five times length of apophysis anterior; segment VIII without processes, smooth; ostium bursae with short lateral folds; apophysis anterior about length of segment VIII; antrum short, about one-fifth length of apophysis anterior, funnel-shaped; posterior part of ductus bursae with pair of long lateral sclerites extending to about apex of apophysis anterior, membranous part of ductus bursae about length of segment VIII including apophysis anterior; signum on right side of entrance to sub-oval corpus bursae, with broad and shortly crescent-shaped base and moderately slender hook.
</paragraph>
</subSubSection>
<subSubSection id="03B74AEF12B74B8941FFA3D6961A37D3" pageId="0" pageNumber="39" type="molecular data">
<paragraph id="FAEC53A122F2CC3A3008CA98788122D9" pageId="0" pageNumber="39">Molecular data.</paragraph>
<paragraph id="627C0D16F3419140A94A7033BBABCF58" pageId="0" pageNumber="39">
BIN: BOLD:ACT3307. The intraspecific average distance of the barcode region is 1.18%, the maximum distance 1.77% (p-dist) (n = 4). The minimum distance to the nearest neighbor,
<taxonomicName id="170364C625916ACAA79C8833C63D6D87" lsidName="C. arenariella" pageId="0" pageNumber="39" rank="species" species="arenariella">
<emphasis id="38D42A902E806C67A1902264DAD5C4AC" italics="true" pageId="0" pageNumber="39">C. arenariella</emphasis>
</taxonomicName>
, is 2.25%.
</paragraph>
</subSubSection>
<subSubSection id="ED431CC75E318D8093170DC8692FF201" pageId="0" pageNumber="39" type="etymology">
<paragraph id="4C6B400CC0DDB8A567FC56F3A86D0BDA" pageId="0" pageNumber="39">Etymology.</paragraph>
<paragraph id="72B55D8133E12E2BE1323C5599D7953D" pageId="0" pageNumber="39">The species is named in honour of Reinhold Messner, the first climber to ascend all fourteen peaks above 8000 metres sea level and living in the neighborhood of the type locality of the new species.</paragraph>
</subSubSection>
<subSubSection id="B27A6762E90D458C5114E75041DA083D" pageId="0" pageNumber="39" type="distribution">
<paragraph id="D0859A3344C0BEC6106469DFAAAE5ABB" pageId="0" pageNumber="39">Distribution</paragraph>
<paragraph id="D873E7346032B8EACC30E20F265750A0" pageId="0" pageNumber="39">
(Fig.
<figureCitation id="151120E6C148B847E27ABA476985FA44" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution map of successfully sequenced material of the C. schleichi species group. Map created with SimpleMappr (http: // www. simplemappr. net)." figureDoi="10.3897/alpento.4.50703.figure1" httpUri="https://binary.pensoft.net/fig/403186" pageId="0" pageNumber="39">1</figureCitation>
). The species is currently only known from a restricted area in northern Italy (region of Trentino-Alto Adige) and from specimens collected in Slovenia, Bulgaria and Greece.
</paragraph>
</subSubSection>
<subSubSection id="BA9CF906476585E501465ACFAD7BC2DE" pageId="0" pageNumber="39" type="bionomics">
<paragraph id="D62C9BC5F424685754E1E185E26C4D2B" pageId="0" pageNumber="39">Bionomics.</paragraph>
<paragraph id="82BFB81385E47427937B7CB9FEF73E9C" pageId="0" pageNumber="39">
Host-plant and early stages are undescribed but it seems most likely that the species shows a similar behaviour as related taxa with a hostplant restriction to
<taxonomicName id="51154A9667EB59D53AFC96ABA4A28ACD" class="Magnoliopsida" family="Caryophyllaceae" genus="Dianthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dianthus" order="Caryophyllales" pageId="0" pageNumber="39" phylum="Tracheophyta" rank="genus">
<emphasis id="399E114AD188785A1D77D698A4A503C2" italics="true" pageId="0" pageNumber="39">Dianthus</emphasis>
</taxonomicName>
spp. At the type-locality
<taxonomicName id="6B160D114B21A29A86052D4EECE7A4E0" class="Magnoliopsida" family="Caryophyllaceae" genus="Dianthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dianthus sylvestris" order="Caryophyllales" pageId="0" pageNumber="39" phylum="Tracheophyta" rank="species" species="sylvestris">
<emphasis id="57C82C12E444882FC6AAD4AFFA7CF429" italics="true" pageId="0" pageNumber="39">Dianthus sylvestris</emphasis>
</taxonomicName>
Wulfen has been recorded. The adults have been found from mid to late August nearby rocks on siliceous soil where they were attracted to artificial light sources.
</paragraph>
</subSubSection>
</treatment>
</document>
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