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<document id="E4EBE959A672EF43C9A810CCF3F7DA10" ID-CLB-Dataset="26654" ID-DOI="10.11646/zootaxa.4612.2.1" ID-GBIF-Dataset="54df4dc5-0117-46a7-ae72-ccd9cfd5f707" ID-ISSN="1175-5326" ID-Zenodo-Dep="3234244" ID-ZooBank="221DBD46-29B1-48C4-BB4B-2C6DE2905EAE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1559107401948" checkinUser="plazi" docAuthor="Kottelat, Maurice" docDate="2019" docId="76100E3319278A00ACC7FEA2FE01CA56" docLanguage="en" docName="zootaxa.4612.2.1.pdf" docOrigin="Zootaxa 4612 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhyacoschistura larreci Kottelat 2019, new species" docType="treatment" docVersion="10" lastPageNumber="161" masterDocId="8A29764B192F8A0AAC50FFC4FF91CE61" masterDocTitle="Rhyacoschistura larreci, a new genus and species of loach from Laos and redescription of R. suber (Teleostei: Nemacheilidae)" masterLastPageNumber="170" masterPageNumber="151" pageNumber="159" updateTime="1698732185724" updateUser="ExternalLinkService">
<mods:mods id="001D68C0648D6A25F217B861C7FFE17A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="E73FA265F2CBADCEC14433807786876B">Rhyacoschistura larreci, a new genus and species of loach from Laos and redescription of R. suber (Teleostei: Nemacheilidae)</mods:title>
</mods:titleInfo>
<mods:name id="3BE317AAD61E310FFD00711940E9F858" type="personal">
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<mods:namePart id="9DE9F1CB29D263C7C62309471392AB2D">Kottelat, Maurice</mods:namePart>
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<mods:title id="46044E68E53078CE61070DBCA75A2015">Zootaxa</mods:title>
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<mods:part id="7C916724EF993AB2753421C4F00E2C1C">
<mods:date id="9F4AD6DA6129BDB5158F0DF663172E73">2019</mods:date>
<mods:detail id="5EE83DB51EB1C61C850425F9DA927CC5" type="pubDate">
<mods:number id="B0BB2C6194C0C52E0CBFF0D474C09865">2019-05-29</mods:number>
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<mods:number id="25862801A69631E64C9EAC325CD82B25">4612</mods:number>
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<mods:classification id="92708897150931220C4CAB39642C14FE">journal article</mods:classification>
<mods:identifier id="AEAE0866C5415A946C44FCEBABC3D746" type="CLB-Dataset">26654</mods:identifier>
<mods:identifier id="E47EE1B280482318F00EBD3D4EB88B05" type="DOI">10.11646/zootaxa.4612.2.1</mods:identifier>
<mods:identifier id="842A39B9518BF97BEABD9FEAFF640163" type="GBIF-Dataset">54df4dc5-0117-46a7-ae72-ccd9cfd5f707</mods:identifier>
<mods:identifier id="350679661B4BFB5643883DEF17CB6FFD" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
<treatment id="76100E3319278A00ACC7FEA2FE01CA56" ID-GBIF-Taxon="157293820" LSID="urn:lsid:plazi:treatment:76100E3319278A00ACC7FEA2FE01CA56" httpUri="http://treatment.plazi.org/id/76100E3319278A00ACC7FEA2FE01CA56" lastPageId="10" lastPageNumber="161" pageId="8" pageNumber="159">
<subSubSection id="B6A3ECAE19278A02ACC7FEA2FDC6CFE1" box="[151,599,358,385]" pageId="8" pageNumber="159" type="nomenclature">
<paragraph id="FE06BF2519278A02ACC7FEA2FDC6CFE1" blockId="8.[151,599,358,421]" box="[151,599,358,385]" pageId="8" pageNumber="159">
<heading id="A54E084919278A02ACC7FEA2FDC6CFE1" bold="true" box="[151,599,358,385]" fontSize="11" level="1" pageId="8" pageNumber="159" reason="1">
<emphasis id="CCCD633719278A02ACC7FEA2FDC6CFE1" bold="true" box="[151,599,358,385]" pageId="8" pageNumber="159">
<taxonomicName id="39B9C4A619278A02ACC7FEA2FE28CFE0" authority="Kottelat, 2019" authorityName="Kottelat" authorityYear="2019" box="[151,441,358,385]" class="Actinopterygii" family="Nemacheilidae" genus="Rhyacoschistura" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="159" phylum="Chordata" rank="species" species="larreci" status="sp. nov.">
<emphasis id="CCCD633719278A02ACC7FEA2FE28CFE0" bold="true" box="[151,441,358,385]" italics="true" pageId="8" pageNumber="159">Rhyacoschistura larreci</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="D7FEDE4C19278A02AD97FEA3FDC6CFE1" box="[455,599,358,385]" pageId="8" pageNumber="159" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B6A3ECAE19278A02ACC7FE4EFE83CFC4" box="[151,274,394,421]" pageId="8" pageNumber="159" type="description">
<paragraph id="FE06BF2519278A02ACC7FE4EFE83CFC4" blockId="8.[151,599,358,421]" box="[151,274,394,421]" pageId="8" pageNumber="159">
(
<figureCitation id="6682A3A019278A02ACCFFE4EFE9BCFC4" box="[159,266,394,421]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStartId-0="5.[151,250,998,1023]" captionStartId-1="5.[151,250,1802,1827]" captionStartId-2="6.[151,250,1986,2011]" captionTargetBox-0="[173,1412,217,909]" captionTargetBox-1="[153,1435,1056,1736]" captionTargetBox-2="[174,1421,177,1950]" captionTargetId-0="figure@5.[153,1438,177,940]" captionTargetId-1="figure@5.[151,1436,1055,1737]" captionTargetId-2="figure@6.[164,1423,177,1950]" captionTargetPageId-0="5" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURE 4. Rhyacoschistura larreci, MHNG 2727.011, holotype, 57.1 mm SL; Laos: Xayaburi: Nam Houng watershed." captionText-1="FIGURE 5. Rhyacoschistura larreci, CMK 28037, paratype, 57.8 mm SL; Laos: Xayaburi: Nam Houng watershed; immediately after fixation." captionText-2="FIGURE 6. Rhyacoschistura larreci, CMK 28037, paratypes; Laos: Xayaburi: Nam Houng watershed; a, 23.9 mm SL; b, 26.5 mm SL; c, 29.2 mm SL; d, 63.6 mm SL; e, 68.8 mm SL." httpUri-0="https://zenodo.org/record/3234250/files/figure.png" httpUri-1="https://zenodo.org/record/3234252/files/figure.png" httpUri-2="https://zenodo.org/record/3234254/files/figure.png" pageId="8" pageNumber="159">Figs. 4–6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B6A3ECAE19278A02ACC7FE16FB80CC39" pageId="8" pageNumber="159" type="materials_examined">
<paragraph id="FE06BF2519278A02ACC7FE16FD1FCC55" blockId="8.[151,1437,465,2041]" pageId="8" pageNumber="159">
<materialsCitation id="4ED1B57819278A02ACC7FE16FD1FCC55" ID-GBIF-Occurrence="2244538996" collectingDate="2018-12-12" collectionCode="MHNG" collectorName="M. Kottelat &amp; T. Phommavong" country="Laos" county="Xayaburi" elevation="528" latitude="19.538916" location="Xayaburi Province" longLatPrecision="1" longitude="101.49708" municipality="Hongsa" pageId="8" pageNumber="159" specimenCode="MHNG 2727.011" specimenCount="1" typeStatus="holotype">
<emphasis id="CCCD633719278A02ACC7FE16FE9ACF8D" bold="true" box="[151,267,466,492]" pageId="8" pageNumber="159">
<typeStatus id="2102018719278A02ACC7FE16FE97CF8D" box="[151,262,466,492]" pageId="8" pageNumber="159" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="AE1F175E19278A02AD44FE16FE4CCF8D" box="[276,477,466,492]" collectionCode="MHNG" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="8" pageNumber="159" type="Museum">MHNG 2727.011</specimenCode>
,
<quantity id="394112C019278A02ADBAFE15FDC1CF8C" box="[490,592,465,493]" metricMagnitude="-2" metricUnit="m" metricValue="5.71" pageId="8" pageNumber="159" unit="mm" value="57.1">57.1 mm</quantity>
SL;
<collectingCountry id="86AEFFB519278A02AEDAFE17FD54CF8C" box="[650,709,467,493]" name="Laos" pageId="8" pageNumber="159">Laos</collectingCountry>
:
<location id="FB66E9FE19278A02AE81FE16FC3BCF8D" LSID="urn:lsid:plazi:treatment:76100E3319278A00ACC7FEA2FE01CA56:FB66E9FE19278A02AE81FE16FC3BCF8D" box="[721,938,466,492]" country="Laos" county="Xayaburi" latitude="19.538916" longLatPrecision="1" longitude="101.49708" municipality="Hongsa" name="Xayaburi Province" pageId="8" pageNumber="159">Xayaburi Province</location>
:
<location id="FB66E9FE19278A02AFE6FE17FBDBCF8D" LSID="urn:lsid:plazi:treatment:76100E3319278A00ACC7FEA2FE01CA56:FB66E9FE19278A02AFE6FE17FBDBCF8D" box="[950,1098,466,493]" country="Laos" county="Xayaburi" latitude="19.538916" longLatPrecision="1" longitude="101.49708" municipality="Hongsa" name="Houay Plong" pageId="8" pageNumber="159">Houay Plong</location>
at bridge
<locationDeviation id="EFB4A60719278A02A891FE16FED4CC71" location="South of Ban Houy Chouang" pageId="8" pageNumber="159">
<quantity id="394112C019278A02A891FE16FB6DCF8D" box="[1217,1276,466,492]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="159" unit="km" value="4.0">4 km</quantity>
South of Ban Houy Chouang
</locationDeviation>
, itself
<quantity id="394112C019278A02ADC0FE32FE49CC71" box="[400,472,502,528]" metricMagnitude="4" metricUnit="m" metricValue="3.3" pageId="8" pageNumber="159" unit="km" value="33.0">33 km</quantity>
from
<collectingMunicipality id="1E62255F19278A02AE4EFE33FDE5CC70" box="[542,628,503,529]" pageId="8" pageNumber="159">Hongsa</collectingMunicipality>
on road to
<collectingCounty id="1767C7A919278A02AEA5FE32FCF0CC71" box="[757,865,502,528]" pageId="8" pageNumber="159">Xayaburi</collectingCounty>
;
<elevation id="7594581619278A02AF3DFE31FC46CC71" box="[877,983,501,528]" metricMagnitude="2" metricUnit="m" metricValue="5.279999999999999" pageId="8" pageNumber="159" unit="m" value="528.0">
<quantity id="394112C019278A02AF3DFE31FC46CC71" box="[877,983,501,528]" metricMagnitude="2" metricUnit="m" metricValue="5.279999999999999" pageId="8" pageNumber="159" unit="m" value="528.0">528 masl</quantity>
</elevation>
;
<geoCoordinate id="9B8DD9E219278A02AFB3FE32FB10CC71" box="[995,1153,502,528]" degrees="19" direction="north" minutes="32" orientation="latitude" pageId="8" pageNumber="159" precision="1" seconds="20.1" value="19.538916">19°32’20.1”N</geoCoordinate>
<geoCoordinate id="9B8DD9E219278A02A8D9FE31FAA4CC71" box="[1161,1333,501,528]" degrees="101" direction="east" minutes="29" orientation="longitude" pageId="8" pageNumber="159" precision="1" seconds="49.5" value="101.49708">101°29’49.5”E</geoCoordinate>
;
<collectorName id="534CDAF319278A02A911FE33FF56CC55" pageId="8" pageNumber="159">M. Kottelat</collectorName>
&amp;
<collectorName id="534CDAF319278A02ACBBFDDFFE36CC55" box="[235,423,538,565]" pageId="8" pageNumber="159">T. Phommavong</collectorName>
,
<collectingDate id="9A43600D19278A02ADE1FDDEFD1BCC55" box="[433,650,538,564]" pageId="8" pageNumber="159" value="2018-12-12">
<date id="8A0799E519278A02ADE1FDDEFD1BCC55" box="[433,650,538,564]" pageId="8" pageNumber="159" value="2018-12-12">12 December 2018</date>
</collectingDate>
.
</materialsCitation>
</paragraph>
<paragraph id="FE06BF2519278A02AC97FDFAFB80CC39" blockId="8.[151,1437,465,2041]" box="[199,1041,573,601]" pageId="8" pageNumber="159">
<materialsCitation id="4ED1B57819278A02AC97FDFAFB9DCC39" ID-GBIF-Occurrence="2244538995" box="[199,1036,573,601]" collectionCode="ZRC" pageId="8" pageNumber="159" specimenCode="8" specimenCount="1" typeStatus="paratype">
<emphasis id="CCCD633719278A02AC97FDFAFED7CC39" bold="true" box="[199,326,574,600]" pageId="8" pageNumber="159">
<typeStatus id="2102018719278A02AC97FDFAFED0CC39" box="[199,321,574,600]" pageId="8" pageNumber="159" type="paratype">Paratypes</typeStatus>
.
</emphasis>
CMK 28037, 48;
<collectionCode id="98A827E019278A02AE43FDFAFDD8CC39" box="[531,585,574,600]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="159">ZRC</collectionCode>
60828,
<specimenCode id="AE1F175E19278A02AEF4FDFAFD25CC39" box="[676,692,574,600]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="159">8</specimenCode>
;
<quantity id="394112C019278A02AE91FDF9FCF5CC38" box="[705,868,573,601]" metricMagnitude="-2" metricUnit="m" metricValue="4.765" metricValueMax="6.88" metricValueMin="2.65" pageId="8" pageNumber="159" unit="mm" value="47.65" valueMax="68.8" valueMin="26.5">26.5–68.8 mm</quantity>
SL; same data
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B6A3ECAE19278A02AC97FDA5FAC8CC89" pageId="8" pageNumber="159" type="diagnosis">
<paragraph id="FE06BF2519278A02AC97FDA5FAC8CC89" blockId="8.[151,1437,465,2041]" pageId="8" pageNumber="159">
<emphasis id="CCCD633719278A02AC97FDA5FED3CC1D" bold="true" box="[199,322,609,636]" pageId="8" pageNumber="159">Diagnosis.</emphasis>
<taxonomicName id="39B9C4A619278A02AD19FDA6FDC6CC1D" authorityName="Kottelat" authorityYear="2019" box="[329,599,610,636]" class="Actinopterygii" family="Nemacheilidae" genus="Rhyacoschistura" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="159" phylum="Chordata" rank="species" species="larreci">
<emphasis id="CCCD633719278A02AD19FDA6FDC6CC1D" box="[329,599,610,636]" italics="true" pageId="8" pageNumber="159">Rhyacoschistura larreci</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="39B9C4A619278A02AF06FDA7FC25CC1D" baseAuthorityName="Kottelat" baseAuthorityYear="2000" box="[854,948,610,636]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="159" phylum="Chordata" rank="species" species="suber">
<emphasis id="CCCD633719278A02AF06FDA7FC25CC1D" box="[854,948,610,636]" italics="true" pageId="8" pageNumber="159">R. suber</emphasis>
</taxonomicName>
by its colour pattern (flank with numerous narrow slanted bars, very irregularly organised and shaped, more or less connected, or sometimes covering whole flank); suborbital flap present in males (vs. absent); on dorsal surface of pectoral fin, small tubercles along branched rays 1–3 and on membrane behind branched ray 1 (vs. tubercles on unculiferous pad behind branched ray 1).
</paragraph>
</subSubSection>
<subSubSection id="B6A3ECAE19278A00AC97FD35FAA9CD76" lastPageId="10" lastPageNumber="161" pageId="8" pageNumber="159" type="description">
<paragraph id="FE06BF2519278A02AC97FD35FA12CD85" blockId="8.[151,1437,465,2041]" pageId="8" pageNumber="159">
<emphasis id="CCCD633719278A02AC97FD35FEC8CD6D" bold="true" box="[199,345,753,780]" pageId="8" pageNumber="159">Description.</emphasis>
See
<figureCitation id="6682A3A019278A02ADDDFD36FD80CD6D" box="[397,529,754,780]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStartId-0="5.[151,250,998,1023]" captionStartId-1="5.[151,250,1802,1827]" captionStartId-2="6.[151,250,1986,2011]" captionTargetBox-0="[173,1412,217,909]" captionTargetBox-1="[153,1435,1056,1736]" captionTargetBox-2="[174,1421,177,1950]" captionTargetId-0="figure@5.[153,1438,177,940]" captionTargetId-1="figure@5.[151,1436,1055,1737]" captionTargetId-2="figure@6.[164,1423,177,1950]" captionTargetPageId-0="5" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURE 4. Rhyacoschistura larreci, MHNG 2727.011, holotype, 57.1 mm SL; Laos: Xayaburi: Nam Houng watershed." captionText-1="FIGURE 5. Rhyacoschistura larreci, CMK 28037, paratype, 57.8 mm SL; Laos: Xayaburi: Nam Houng watershed; immediately after fixation." captionText-2="FIGURE 6. Rhyacoschistura larreci, CMK 28037, paratypes; Laos: Xayaburi: Nam Houng watershed; a, 23.9 mm SL; b, 26.5 mm SL; c, 29.2 mm SL; d, 63.6 mm SL; e, 68.8 mm SL." httpUri-0="https://zenodo.org/record/3234250/files/figure.png" httpUri-1="https://zenodo.org/record/3234252/files/figure.png" httpUri-2="https://zenodo.org/record/3234254/files/figure.png" pageId="8" pageNumber="159">Figures 4–6</figureCitation>
for general appearance and
<tableCitation id="B33B8A9E19278A02AF18FD36FC0BCD6D" box="[840,922,754,780]" captionStart="TABLE 1" captionStartId="9.[151,239,370,394]" captionTargetPageId="9" captionText="TABLE 1. Morphometric data of type material of Rhycoshistura larreci (n=7). Range and mean include holotype data." httpUri="http://table.plazi.org/id/AAC6EFAD19268A03ACC7FEB6FA1CCFEA" pageId="8" pageNumber="159" tableUuid="AAC6EFAD19268A03ACC7FEB6FA1CCFEA">Table 1</tableCitation>
for morphometric data of
<typeStatus id="2102018719278A02A8EDFD36FA8ECD6D" box="[1213,1311,754,780]" pageId="8" pageNumber="159" type="holotype">holotype</typeStatus>
and 6 largest
<typeStatus id="2102018719278A02ACEEFCD3FEBCCD50" box="[190,301,791,817]" pageId="8" pageNumber="159" type="paratype">paratypes</typeStatus>
. A moderately elongate nemacheilid with body depth slowly increasing up to slightly in front of dorsal-fin origin. Behind dorsal fin, body depth almost uniform until caudal-fin base. Dorsal profile with a very slight concavity between head and body. Head from about circular anteriorly to compressed posteriorly. Interorbital area slightly convex. In lateral view, eye flush with dorsal profile of head. Cheeks not swollen. Snout rounded in dorsal and lateral view. Depth of caudal peduncle 1.1–1.3 times in its length, depth uniform. Dorsal ridge on posterior fourth of post-dorsal area. Ventral ridge on posterior half of caudal peduncle. Largest recorded size
<quantity id="394112C019278A02A8A0FC0EFAC5CD84" box="[1264,1364,970,997]" metricMagnitude="-2" metricUnit="m" metricValue="6.88" pageId="8" pageNumber="159" unit="mm" value="68.8">68.8 mm</quantity>
SL.
</paragraph>
<paragraph id="FE06BF2519278A02AC97FC2AFDFCCB48" blockId="8.[151,1437,465,2041]" pageId="8" pageNumber="159">Dorsal fin with 4 unbranched and 7½ branched rays; distal margin convex. Second branched ray longest. Pectoral fin with 1 unbranched and 10* (6) or 11 (1) branched rays, rounded and reaching half of distance to pelvic-fin base in females, slightly falcate and reaching about ⅔ of distance to pelvic-fin base in males. No axillary pectoral lobe. Pelvic fin with 1 unbranched and 7 branched rays; reaching about halfway of distance to anal-fin origin; falcate, posterior margin straight; origin at vertical through dorsal-fin origin; axillary lobe present, entirely free. Anus situated about 1.5–2 eye diameters in front of anal fin, behind posterior extremity of pelvic fin. Anal fin with 3 unbranched and 5½ branched rays; distal margin straight. Caudal fin with 8+8* branched rays (one specimen with 7+8 rays is probably anomalous); dorsal and ventral procurrent rays cannot be counted; emarginate, lobes rounded, upper lobe slightly longer than lower one.</paragraph>
<paragraph id="FE06BF2519278A02AC97FAF6FD8CCBD8" blockId="8.[151,1437,465,2041]" pageId="8" pageNumber="159">Body entirely covered by scales, including predorsal area and belly between pectoral-fin bases. All scales very distinct. Lateral line complete in large specimens, interrupted in some smaller ones, with 103–122 pores (difficult to count with accuracy). Cephalic lateral line system with 6 supraorbital, 4 + 10–11 infraorbital, 9 preoperculo-mandibular and 3 supratemporal pores.</paragraph>
<paragraph id="FE06BF2519278A02AC97FA06FABCC8F0" blockId="8.[151,1437,465,2041]" pageId="8" pageNumber="159">
Anterior naris pierced on front side of a pointed flap-like tube. Posterior naris adjacent to anterior one. Mouth arched, gape about twice as wide as long (
<figureCitation id="6682A3A019278A02AE22FA22FD29C861" box="[626,696,1510,1536]" captionStart="FIGURE 2" captionStartId="3.[151,250,734,759]" captionTargetId="figure@3.[350,1237,192,684]" captionTargetPageId="3" captionText="FIGURE 2. Rhyacoschistura larreci, CMK 28034, 68.8 mm SL; mouth." httpUri="https://zenodo.org/record/3387006/files/figure.png" pageId="8" pageNumber="159">Fig. 2</figureCitation>
). Lips thick, fleshy. Upper lip with a median notch and short, fine wrinkles. Processus dentiformis present. Lower lip with median interruption; median part with 1–2 shallow sulci, lateral part with fine, shallow wrinkles. Frenum of lower lip wide, median part of lower lip not attached to jaw. Tip of lower jaw not exposed. No median notch on lower jaw. Inner rostral barbel reaching beyond corner of mouth; outer one reaching beyond posterior margin of eye. Maxillary barbel reaching middle of postorbital area.
</paragraph>
<paragraph id="FE06BF2519278A02AC97F95EFAB0C8B8" blockId="8.[151,1437,465,2041]" pageId="8" pageNumber="159">
Intestine with a loop behind stomach (
<figureCitation id="6682A3A019278A02AE3EF95EFD2EC8D4" box="[622,703,1690,1717]" captionStart="FIGURE 7" captionStartId="7.[151,250,1139,1164]" captionTargetBox="[238,1361,243,1060]" captionTargetId="figure@7.[153,1438,177,1083]" captionTargetPageId="7" captionText="FIGURE 7. Rhyacoschistura larreci, CMK 28034; a, 49.8 mm SL; digestive tract; b, 50.2 mm SL; air bladder capsule." httpUri="https://zenodo.org/record/3234256/files/figure.png" pageId="8" pageNumber="159">Fig. 7a</figureCitation>
). Air bladder without posterior chamber visible in abdominal cavity. Two halves of bony capsule of anterior chamber of air bladder connected by a manubrium (
<figureCitation id="6682A3A019278A02A891F97AFA85C8B8" box="[1217,1300,1726,1753]" captionStart="FIGURE 7" captionStartId="7.[151,250,1139,1164]" captionTargetBox="[238,1361,243,1060]" captionTargetId="figure@7.[153,1438,177,1083]" captionTargetPageId="7" captionText="FIGURE 7. Rhyacoschistura larreci, CMK 28034; a, 49.8 mm SL; digestive tract; b, 50.2 mm SL; air bladder capsule." httpUri="https://zenodo.org/record/3234256/files/figure.png" pageId="8" pageNumber="159">Fig. 7b</figureCitation>
).
</paragraph>
<paragraph id="FE06BF2519278A02AC97F925FF44C998" blockId="8.[151,1437,465,2041]" pageId="8" pageNumber="159">
<emphasis id="CCCD633719278A02AC97F925FE26C89D" bold="true" box="[199,439,1761,1788]" pageId="8" pageNumber="159">Sexual dimorphism.</emphasis>
Males with globulous suborbital flap, with small tubercles along posterior edge; females with neither suborbital flap nor suborbital groove. In males, pectoral fin rigid, slightly curled upwards (
<figureCitation id="6682A3A019278A02A96BF8C2FA1CC941" box="[1339,1421,1798,1824]" captionStart="FIGURE 1" captionStartId="2.[151,250,1783,1808]" captionTargetBox="[153,1425,179,1726]" captionTargetId="figure@2.[153,1425,179,1726]" captionTargetPageId="2" captionText="FIGURE 1. Rhyacoschistura larreci, left pectoral fin, dorsal view, of: a, male, MHNG 2727.011, holotype, 57.1 mm SL; and b, female, CMK 28034, 68.8 mm SL." httpUri="https://zenodo.org/record/3234246/files/figure.png" pageId="8" pageNumber="159">Fig. 1a</figureCitation>
). Membrane between unbranched and first branched rays narrower distally than at midlength. First branched ray about 4 times wider than adjacent rays. After branching point, anterior branch wider than posterior one, becoming narrower again at level of branching point of posterior branch. Anterior branch unbranched. Posterior branch branched again. No membrane between branches. On dorsal surface of first branched rays, on distal half of rays and adjacent part of membrane behind posterior branch, small tubercles organised in about 6–7 irregular longitudinal rows.
</paragraph>
<paragraph id="FE06BF2519268A03AC97FF5DFD17CE9A" blockId="9.[151,1437,153,324]" pageId="9" pageNumber="160">Branched ray 2: posterior branch branched again; no membrane between branches. Branched rays 3–8 branched twice; membrane narrow or absent between branching points 1 and 2. A few small tubercles along posterior edge of branched ray 2 and sometimes behind ray 3.</paragraph>
<paragraph id="FE06BF2519268A03AC97FEC1FBFACF22" blockId="9.[151,1437,153,324]" pageId="9" pageNumber="160">
In female, first branched ray only slightly wider than adjacent rays; branching of rays as in males, but all rays and branches connected by membranes (
<figureCitation id="6682A3A019268A03AE0DFEEDFD3ECF25" box="[605,687,297,324]" captionStart="FIGURE 1" captionStartId="2.[151,250,1783,1808]" captionTargetBox="[153,1425,179,1726]" captionTargetId="figure@2.[153,1425,179,1726]" captionTargetPageId="2" captionText="FIGURE 1. Rhyacoschistura larreci, left pectoral fin, dorsal view, of: a, male, MHNG 2727.011, holotype, 57.1 mm SL; and b, female, CMK 28034, 68.8 mm SL." httpUri="https://zenodo.org/record/3234246/files/figure.png" pageId="9" pageNumber="160">Fig. 1b</figureCitation>
). No tubercles on rays and membranes.
</paragraph>
<caption id="AAC6EFAD19268A03ACC7FEB6FA1CCFEA" ID-Table-UUID="AAC6EFAD19268A03ACC7FEB6FA1CCFEA" box="[151,1421,370,395]" httpUri="http://table.plazi.org/id/AAC6EFAD19268A03ACC7FEB6FA1CCFEA" pageId="9" pageNumber="160" targetBox="[153,1335,407,1717]" targetIsTable="true" targetPageId="9">
<paragraph id="FE06BF2519268A03ACC7FEB6FA1CCFEA" blockId="9.[151,1421,370,395]" box="[151,1421,370,395]" pageId="9" pageNumber="160">
<emphasis id="CCCD633719268A03ACC7FEB6FE9BCFEB" bold="true" box="[151,266,370,394]" pageId="9" pageNumber="160">TABLE 1.</emphasis>
Morphometric data of type material of
<emphasis id="CCCD633719268A03AEE1FEB7FC09CFEA" box="[689,920,371,395]" italics="true" pageId="9" pageNumber="160">
Rhycoshistura
<taxonomicName id="39B9C4A619268A03AF00FEB7FC09CFEA" authorityName="Kottelat" authorityYear="2019" box="[848,920,371,395]" class="Actinopterygii" family="Nemacheilidae" genus="Rhyacoschistura" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="160" phylum="Chordata" rank="species" species="larreci">larreci</taxonomicName>
</emphasis>
(n=7). Range and mean include holotype data.
</paragraph>
</caption>
<paragraph id="FE06BF2519268A03AF57FE53FA86C8D4" pageId="9" pageNumber="160">
<table id="8CB94D85192675F5ACC9FE53FAA6C8D4" box="[153,1335,407,1717]" gridcols="4" gridrows="35" pageId="9" pageNumber="160">
<tr id="4089BD67192675F5ACC9FE53FAA6CFD1" box="[153,1335,407,432]" gridrow="0" pageId="9" pageNumber="160" rowspan-0="1">
<th id="0358D41B192675F5AF57FE53FCF1CFD1" box="[775,864,407,432]" gridcol="1" gridrow="0" pageId="9" pageNumber="160">holotype</th>
<th id="0358D41B192675F5A857FE53FB17CFD1" box="[1031,1158,407,432]" gridcol="2" gridrow="0" pageId="9" pageNumber="160">range</th>
<th id="0358D41B192675F5A8AEFE53FAA6CFD1" box="[1278,1335,407,432]" gridcol="3" gridrow="0" pageId="9" pageNumber="160">mean</th>
</tr>
<tr id="4089BD67192675F5ACC9FE79FAA6CFB7" box="[153,1335,445,470]" gridrow="1" pageId="9" pageNumber="160" rowspan-3="1">
<th id="0358D41B192675F5ACC9FE79FE66CFB7" box="[153,503,445,470]" gridcol="0" gridrow="1" pageId="9" pageNumber="160">Standard lenth (mm)</th>
<td id="0358D41B192675F5AF57FE79FCF1CFB7" box="[775,864,445,470]" gridcol="1" gridrow="1" pageId="9" pageNumber="160">57.1</td>
<td id="0358D41B192675F5A857FE79FB17CFB7" box="[1031,1158,445,470]" gridcol="2" gridrow="1" pageId="9" pageNumber="160">52.3–68.8</td>
</tr>
<tr id="4089BD67192675F5ACC9FE27FAA6CF9D" box="[153,1335,483,508]" gridrow="2" pageId="9" pageNumber="160" rowspan-3="1">
<th id="0358D41B192675F5ACC9FE27FE66CF9D" box="[153,503,483,508]" gridcol="0" gridrow="2" pageId="9" pageNumber="160">Total lenth (mm)</th>
<td id="0358D41B192675F5AF57FE27FCF1CF9D" box="[775,864,483,508]" gridcol="1" gridrow="2" pageId="9" pageNumber="160">69.8</td>
<td id="0358D41B192675F5A857FE27FB17CF9D" box="[1031,1158,483,508]" gridcol="2" gridrow="2" pageId="9" pageNumber="160">64.1–82.7</td>
</tr>
<tr id="4089BD67192675F5ACC9FDCCFAA6CC40" box="[153,1335,520,545]" gridrow="3" pageId="9" pageNumber="160" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="0358D41B192675F5ACC9FDCCFE66CC40" box="[153,503,520,545]" gridcol="0" gridrow="3" pageId="9" pageNumber="160">In percent of standard length</th>
</tr>
<tr id="4089BD67192675F5ACC9FDEAFAA6CC26" box="[153,1335,558,583]" gridrow="4" pageId="9" pageNumber="160">
<th id="0358D41B192675F5ACC9FDEAFE66CC26" box="[153,503,558,583]" gridcol="0" gridrow="4" pageId="9" pageNumber="160">Total length</th>
<td id="0358D41B192675F5AF57FDEAFCF1CC26" box="[775,864,558,583]" gridcol="1" gridrow="4" pageId="9" pageNumber="160">122.3</td>
<td id="0358D41B192675F5A857FDEAFB17CC26" box="[1031,1158,558,583]" gridcol="2" gridrow="4" pageId="9" pageNumber="160">120.3–123.7</td>
<td id="0358D41B192675F5A8AEFDEAFAA6CC26" box="[1278,1335,558,583]" gridcol="3" gridrow="4" pageId="9" pageNumber="160">122.2</td>
</tr>
<tr id="4089BD67192675F5ACC9FD90FAA6CC0C" box="[153,1335,596,621]" gridrow="5" pageId="9" pageNumber="160">
<th id="0358D41B192675F5ACC9FD90FE66CC0C" box="[153,503,596,621]" gridcol="0" gridrow="5" pageId="9" pageNumber="160">Head length (dorsal)</th>
<td id="0358D41B192675F5AF57FD90FCF1CC0C" box="[775,864,596,621]" gridcol="1" gridrow="5" pageId="9" pageNumber="160">20.7</td>
<td id="0358D41B192675F5A857FD90FB17CC0C" box="[1031,1158,596,621]" gridcol="2" gridrow="5" pageId="9" pageNumber="160">19.3–21.5</td>
<td id="0358D41B192675F5A8AEFD90FAA6CC0C" box="[1278,1335,596,621]" gridcol="3" gridrow="5" pageId="9" pageNumber="160">20.3</td>
</tr>
<tr id="4089BD67192675F5ACC9FDBEFAA6CCF2" box="[153,1335,634,659]" gridrow="6" pageId="9" pageNumber="160">
<th id="0358D41B192675F5ACC9FDBEFE66CCF2" box="[153,503,634,659]" gridcol="0" gridrow="6" pageId="9" pageNumber="160">Head length (lateral)</th>
<td id="0358D41B192675F5AF57FDBEFCF1CCF2" box="[775,864,634,659]" gridcol="1" gridrow="6" pageId="9" pageNumber="160">23.9</td>
<td id="0358D41B192675F5A857FDBEFB17CCF2" box="[1031,1158,634,659]" gridcol="2" gridrow="6" pageId="9" pageNumber="160">22.6–24.1</td>
<td id="0358D41B192675F5A8AEFDBEFAA6CCF2" box="[1278,1335,634,659]" gridcol="3" gridrow="6" pageId="9" pageNumber="160">23.4</td>
</tr>
<tr id="4089BD67192675F5ACC9FD64FAA6CCD8" box="[153,1335,672,697]" gridrow="7" pageId="9" pageNumber="160">
<th id="0358D41B192675F5ACC9FD64FE66CCD8" box="[153,503,672,697]" gridcol="0" gridrow="7" pageId="9" pageNumber="160">Predorsal length</th>
<td id="0358D41B192675F5AF57FD64FCF1CCD8" box="[775,864,672,697]" gridcol="1" gridrow="7" pageId="9" pageNumber="160">56.2</td>
<td id="0358D41B192675F5A857FD64FB17CCD8" box="[1031,1158,672,697]" gridcol="2" gridrow="7" pageId="9" pageNumber="160">54.3–56.8</td>
<td id="0358D41B192675F5A8AEFD64FAA6CCD8" box="[1278,1335,672,697]" gridcol="3" gridrow="7" pageId="9" pageNumber="160">55.5</td>
</tr>
<tr id="4089BD67192675F5ACC9FD01FAA6CCBF" box="[153,1335,709,734]" gridrow="8" pageId="9" pageNumber="160">
<th id="0358D41B192675F5ACC9FD01FE66CCBF" box="[153,503,709,734]" gridcol="0" gridrow="8" pageId="9" pageNumber="160">Prepelvic length</th>
<td id="0358D41B192675F5AF57FD01FCF1CCBF" box="[775,864,709,734]" gridcol="1" gridrow="8" pageId="9" pageNumber="160">53.0</td>
<td id="0358D41B192675F5A857FD01FB17CCBF" box="[1031,1158,709,734]" gridcol="2" gridrow="8" pageId="9" pageNumber="160">50.6–53.0</td>
<td id="0358D41B192675F5A8AEFD01FAA6CCBF" box="[1278,1335,709,734]" gridcol="3" gridrow="8" pageId="9" pageNumber="160">51.6</td>
</tr>
<tr id="4089BD67192675F5ACC9FD2FFAA6CD65" box="[153,1335,747,772]" gridrow="9" pageId="9" pageNumber="160">
<th id="0358D41B192675F5ACC9FD2FFE66CD65" box="[153,503,747,772]" gridcol="0" gridrow="9" pageId="9" pageNumber="160">Pre-anus length</th>
<td id="0358D41B192675F5AF57FD2FFCF1CD65" box="[775,864,747,772]" gridcol="1" gridrow="9" pageId="9" pageNumber="160">73.3</td>
<td id="0358D41B192675F5A857FD2FFB17CD65" box="[1031,1158,747,772]" gridcol="2" gridrow="9" pageId="9" pageNumber="160">72.6–73.9</td>
<td id="0358D41B192675F5A8AEFD2FFAA6CD65" box="[1278,1335,747,772]" gridcol="3" gridrow="9" pageId="9" pageNumber="160">73.3</td>
</tr>
<tr id="4089BD67192675F5ACC9FCD5FAA6CD4B" box="[153,1335,785,810]" gridrow="10" pageId="9" pageNumber="160">
<th id="0358D41B192675F5ACC9FCD5FE66CD4B" box="[153,503,785,810]" gridcol="0" gridrow="10" pageId="9" pageNumber="160">Pre-anal length</th>
<td id="0358D41B192675F5AF57FCD5FCF1CD4B" box="[775,864,785,810]" gridcol="1" gridrow="10" pageId="9" pageNumber="160">78.0</td>
<td id="0358D41B192675F5A857FCD5FB17CD4B" box="[1031,1158,785,810]" gridcol="2" gridrow="10" pageId="9" pageNumber="160">77.8–80.1</td>
<td id="0358D41B192675F5A8AEFCD5FAA6CD4B" box="[1278,1335,785,810]" gridcol="3" gridrow="10" pageId="9" pageNumber="160">78.5</td>
</tr>
<tr id="4089BD67192675F5ACC9FCF3FAA6CD31" box="[153,1335,823,848]" gridrow="11" pageId="9" pageNumber="160">
<th id="0358D41B192675F5ACC9FCF3FE66CD31" box="[153,503,823,848]" gridcol="0" gridrow="11" pageId="9" pageNumber="160">Head depth</th>
<td id="0358D41B192675F5AF57FCF3FCF1CD31" box="[775,864,823,848]" gridcol="1" gridrow="11" pageId="9" pageNumber="160">13.4</td>
<td id="0358D41B192675F5A857FCF3FB17CD31" box="[1031,1158,823,848]" gridcol="2" gridrow="11" pageId="9" pageNumber="160">12.6–13.4</td>
<td id="0358D41B192675F5A8AEFCF3FAA6CD31" box="[1278,1335,823,848]" gridcol="3" gridrow="11" pageId="9" pageNumber="160">13.0</td>
</tr>
<tr id="4089BD67192675F5ACC9FC98FAA6CD14" box="[153,1335,860,885]" gridrow="12" pageId="9" pageNumber="160">
<th id="0358D41B192675F5ACC9FC98FE66CD14" box="[153,503,860,885]" gridcol="0" gridrow="12" pageId="9" pageNumber="160">Body depth at dorsal-fin origin</th>
<td id="0358D41B192675F5AF57FC98FCF1CD14" box="[775,864,860,885]" gridcol="1" gridrow="12" pageId="9" pageNumber="160">17.1</td>
<td id="0358D41B192675F5A857FC98FB17CD14" box="[1031,1158,860,885]" gridcol="2" gridrow="12" pageId="9" pageNumber="160">17.1–18.2</td>
<td id="0358D41B192675F5A8AEFC98FAA6CD14" box="[1278,1335,860,885]" gridcol="3" gridrow="12" pageId="9" pageNumber="160">17.8</td>
</tr>
<tr id="4089BD67192675F5ACC9FC46FAA6CDFA" box="[153,1335,898,923]" gridrow="13" pageId="9" pageNumber="160">
<th id="0358D41B192675F5ACC9FC46FE66CDFA" box="[153,503,898,923]" gridcol="0" gridrow="13" pageId="9" pageNumber="160">Depth of caudal peduncle</th>
<td id="0358D41B192675F5AF57FC46FCF1CDFA" box="[775,864,898,923]" gridcol="1" gridrow="13" pageId="9" pageNumber="160">13.3</td>
<td id="0358D41B192675F5A857FC46FB17CDFA" box="[1031,1158,898,923]" gridcol="2" gridrow="13" pageId="9" pageNumber="160">13.0–13.8</td>
<td id="0358D41B192675F5A8AEFC46FAA6CDFA" box="[1278,1335,898,923]" gridcol="3" gridrow="13" pageId="9" pageNumber="160">13.5</td>
</tr>
<tr id="4089BD67192675F5ACC9FC6CFAA6CDA0" box="[153,1335,936,961]" gridrow="14" pageId="9" pageNumber="160">
<th id="0358D41B192675F5ACC9FC6CFE66CDA0" box="[153,503,936,961]" gridcol="0" gridrow="14" pageId="9" pageNumber="160">Length of caudal peduncle</th>
<td id="0358D41B192675F5AF57FC6CFCF1CDA0" box="[775,864,936,961]" gridcol="1" gridrow="14" pageId="9" pageNumber="160">16.6</td>
<td id="0358D41B192675F5A857FC6CFB17CDA0" box="[1031,1158,936,961]" gridcol="2" gridrow="14" pageId="9" pageNumber="160">15.1–16.9</td>
<td id="0358D41B192675F5A8AEFC6CFAA6CDA0" box="[1278,1335,936,961]" gridcol="3" gridrow="14" pageId="9" pageNumber="160">15.8</td>
</tr>
<tr id="4089BD67192675F5ACC9FC0AFAA6CD86" box="[153,1335,974,999]" gridrow="15" pageId="9" pageNumber="160">
<th id="0358D41B192675F5ACC9FC0AFE66CD86" box="[153,503,974,999]" gridcol="0" gridrow="15" pageId="9" pageNumber="160">Head width</th>
<td id="0358D41B192675F5AF57FC0AFCF1CD86" box="[775,864,974,999]" gridcol="1" gridrow="15" pageId="9" pageNumber="160">17.5</td>
<td id="0358D41B192675F5A857FC0AFB17CD86" box="[1031,1158,974,999]" gridcol="2" gridrow="15" pageId="9" pageNumber="160">15.8–17.5</td>
<td id="0358D41B192675F5A8AEFC0AFAA6CD86" box="[1278,1335,974,999]" gridcol="3" gridrow="15" pageId="9" pageNumber="160">16.5</td>
</tr>
<tr id="4089BD67192675F5ACC9FC30FAA6CA6C" box="[153,1335,1012,1037]" gridrow="16" pageId="9" pageNumber="160">
<th id="0358D41B192675F5ACC9FC30FE66CA6C" box="[153,503,1012,1037]" gridcol="0" gridrow="16" pageId="9" pageNumber="160">Body width at dorsal-fin origin</th>
<td id="0358D41B192675F5AF57FC30FCF1CA6C" box="[775,864,1012,1037]" gridcol="1" gridrow="16" pageId="9" pageNumber="160">13.9</td>
<td id="0358D41B192675F5A857FC30FB17CA6C" box="[1031,1158,1012,1037]" gridcol="2" gridrow="16" pageId="9" pageNumber="160">13.8–14.3</td>
<td id="0358D41B192675F5A8AEFC30FAA6CA6C" box="[1278,1335,1012,1037]" gridcol="3" gridrow="16" pageId="9" pageNumber="160">14.0</td>
</tr>
<tr id="4089BD67192675F5ACC9FBDDFAA6CA53" box="[153,1335,1049,1074]" gridrow="17" pageId="9" pageNumber="160">
<th id="0358D41B192675F5ACC9FBDDFE66CA53" box="[153,503,1049,1074]" gridcol="0" gridrow="17" pageId="9" pageNumber="160">Snout length</th>
<td id="0358D41B192675F5AF57FBDDFCF1CA53" box="[775,864,1049,1074]" gridcol="1" gridrow="17" pageId="9" pageNumber="160">10.2</td>
<td id="0358D41B192675F5A857FBDDFB17CA53" box="[1031,1158,1049,1074]" gridcol="2" gridrow="17" pageId="9" pageNumber="160">9.6–10.3</td>
<td id="0358D41B192675F5A8AEFBDDFAA6CA53" box="[1278,1335,1049,1074]" gridcol="3" gridrow="17" pageId="9" pageNumber="160">10.1</td>
</tr>
<tr id="4089BD67192675F5ACC9FBFBFAA6CA39" box="[153,1335,1087,1112]" gridrow="18" pageId="9" pageNumber="160">
<th id="0358D41B192675F5ACC9FBFBFE66CA39" box="[153,503,1087,1112]" gridcol="0" gridrow="18" pageId="9" pageNumber="160">Eye diameter</th>
<td id="0358D41B192675F5AF57FBFBFCF1CA39" box="[775,864,1087,1112]" gridcol="1" gridrow="18" pageId="9" pageNumber="160">3.7</td>
<td id="0358D41B192675F5A857FBFBFB17CA39" box="[1031,1158,1087,1112]" gridcol="2" gridrow="18" pageId="9" pageNumber="160">3.1–3.7</td>
<td id="0358D41B192675F5A8AEFBFBFAA6CA39" box="[1278,1335,1087,1112]" gridcol="3" gridrow="18" pageId="9" pageNumber="160">3.4</td>
</tr>
<tr id="4089BD67192675F5ACC9FBA1FAA6CA1F" box="[153,1335,1125,1150]" gridrow="19" pageId="9" pageNumber="160">
<th id="0358D41B192675F5ACC9FBA1FE66CA1F" box="[153,503,1125,1150]" gridcol="0" gridrow="19" pageId="9" pageNumber="160">Interorbital width</th>
<td id="0358D41B192675F5AF57FBA1FCF1CA1F" box="[775,864,1125,1150]" gridcol="1" gridrow="19" pageId="9" pageNumber="160">8.4</td>
<td id="0358D41B192675F5A857FBA1FB17CA1F" box="[1031,1158,1125,1150]" gridcol="2" gridrow="19" pageId="9" pageNumber="160">7.5–8.6</td>
<td id="0358D41B192675F5A8AEFBA1FAA6CA1F" box="[1278,1335,1125,1150]" gridcol="3" gridrow="19" pageId="9" pageNumber="160">8.2</td>
</tr>
<tr id="4089BD67192675F5ACC9FB4FFAA6CAC5" box="[153,1335,1163,1188]" gridrow="20" pageId="9" pageNumber="160">
<th id="0358D41B192675F5ACC9FB4FFE66CAC5" box="[153,503,1163,1188]" gridcol="0" gridrow="20" pageId="9" pageNumber="160">Length of dorsal fin</th>
<td id="0358D41B192675F5AF57FB4FFCF1CAC5" box="[775,864,1163,1188]" gridcol="1" gridrow="20" pageId="9" pageNumber="160">19.1</td>
<td id="0358D41B192675F5A857FB4FFB17CAC5" box="[1031,1158,1163,1188]" gridcol="2" gridrow="20" pageId="9" pageNumber="160">17.0–20.6</td>
<td id="0358D41B192675F5A8AEFB4FFAA6CAC5" box="[1278,1335,1163,1188]" gridcol="3" gridrow="20" pageId="9" pageNumber="160">18.3</td>
</tr>
<tr id="4089BD67192675F5ACC9FB75FAA6CAAB" box="[153,1335,1201,1226]" gridrow="21" pageId="9" pageNumber="160">
<th id="0358D41B192675F5ACC9FB75FE66CAAB" box="[153,503,1201,1226]" gridcol="0" gridrow="21" pageId="9" pageNumber="160">Length of upper caudal-fin lobe</th>
<td id="0358D41B192675F5AF57FB75FCF1CAAB" box="[775,864,1201,1226]" gridcol="1" gridrow="21" pageId="9" pageNumber="160">22.0</td>
<td id="0358D41B192675F5A857FB75FB17CAAB" box="[1031,1158,1201,1226]" gridcol="2" gridrow="21" pageId="9" pageNumber="160">20.3–23.8</td>
<td id="0358D41B192675F5A8AEFB75FAA6CAAB" box="[1278,1335,1201,1226]" gridcol="3" gridrow="21" pageId="9" pageNumber="160">22.0</td>
</tr>
<tr id="4089BD67192675F5ACC9FB12FAA6CA91" box="[153,1335,1238,1264]" gridrow="22" pageId="9" pageNumber="160">
<th id="0358D41B192675F5ACC9FB12FE66CA91" box="[153,503,1238,1264]" gridcol="0" gridrow="22" pageId="9" pageNumber="160">Length of median caudal-fin rays</th>
<td id="0358D41B192675F5AF57FB12FCF1CA91" box="[775,864,1238,1264]" gridcol="1" gridrow="22" pageId="9" pageNumber="160">18.5</td>
<td id="0358D41B192675F5A857FB12FB17CA91" box="[1031,1158,1238,1264]" gridcol="2" gridrow="22" pageId="9" pageNumber="160">17.3–19.5</td>
<td id="0358D41B192675F5A8AEFB12FAA6CA91" box="[1278,1335,1238,1264]" gridcol="3" gridrow="22" pageId="9" pageNumber="160">18.1</td>
</tr>
<tr id="4089BD67192675F5ACC9FB38FAA6CB74" box="[153,1335,1276,1301]" gridrow="23" pageId="9" pageNumber="160">
<th id="0358D41B192675F5ACC9FB38FE66CB74" box="[153,503,1276,1301]" gridcol="0" gridrow="23" pageId="9" pageNumber="160">Length of lower caudal-fin lobe</th>
<td id="0358D41B192675F5AF57FB38FCF1CB74" box="[775,864,1276,1301]" gridcol="1" gridrow="23" pageId="9" pageNumber="160">23.0</td>
<td id="0358D41B192675F5A857FB38FB17CB74" box="[1031,1158,1276,1301]" gridcol="2" gridrow="23" pageId="9" pageNumber="160">21.3–24.5</td>
<td id="0358D41B192675F5A8AEFB38FAA6CB74" box="[1278,1335,1276,1301]" gridcol="3" gridrow="23" pageId="9" pageNumber="160">22.7</td>
</tr>
<tr id="4089BD67192675F5ACC9FAE6FAA6CB5A" box="[153,1335,1314,1339]" gridrow="24" pageId="9" pageNumber="160">
<th id="0358D41B192675F5ACC9FAE6FE66CB5A" box="[153,503,1314,1339]" gridcol="0" gridrow="24" pageId="9" pageNumber="160">Length of anal fin</th>
<td id="0358D41B192675F5AF57FAE6FCF1CB5A" box="[775,864,1314,1339]" gridcol="1" gridrow="24" pageId="9" pageNumber="160">18.8</td>
<td id="0358D41B192675F5A857FAE6FB17CB5A" box="[1031,1158,1314,1339]" gridcol="2" gridrow="24" pageId="9" pageNumber="160">16.3–19.3</td>
<td id="0358D41B192675F5A8AEFAE6FAA6CB5A" box="[1278,1335,1314,1339]" gridcol="3" gridrow="24" pageId="9" pageNumber="160">18.0</td>
</tr>
<tr id="4089BD67192675F5ACC9FA8CFAA6CB00" box="[153,1335,1352,1377]" gridrow="25" pageId="9" pageNumber="160">
<th id="0358D41B192675F5ACC9FA8CFE66CB00" box="[153,503,1352,1377]" gridcol="0" gridrow="25" pageId="9" pageNumber="160">Length of pelvic fin</th>
<td id="0358D41B192675F5AF57FA8CFCF1CB00" box="[775,864,1352,1377]" gridcol="1" gridrow="25" pageId="9" pageNumber="160">18.8</td>
<td id="0358D41B192675F5A857FA8CFB17CB00" box="[1031,1158,1352,1377]" gridcol="2" gridrow="25" pageId="9" pageNumber="160">15.7–19.4</td>
<td id="0358D41B192675F5A8AEFA8CFAA6CB00" box="[1278,1335,1352,1377]" gridcol="3" gridrow="25" pageId="9" pageNumber="160">18.3</td>
</tr>
<tr id="4089BD67192675F5ACC9FAAAFAA6CBE6" box="[153,1335,1390,1415]" gridrow="26" pageId="9" pageNumber="160">
<th id="0358D41B192675F5ACC9FAAAFE66CBE6" box="[153,503,1390,1415]" gridcol="0" gridrow="26" pageId="9" pageNumber="160">Length of pectoral fin</th>
<td id="0358D41B192675F5AF57FAAAFCF1CBE6" box="[775,864,1390,1415]" gridcol="1" gridrow="26" pageId="9" pageNumber="160">22.0</td>
<td id="0358D41B192675F5A857FAAAFB17CBE6" box="[1031,1158,1390,1415]" gridcol="2" gridrow="26" pageId="9" pageNumber="160">17.7–22.0</td>
<td id="0358D41B192675F5A8AEFAAAFAA6CBE6" box="[1278,1335,1390,1415]" gridcol="3" gridrow="26" pageId="9" pageNumber="160">20.7</td>
</tr>
<tr id="4089BD67192675F5ACC9FA57FAA6CBCD" box="[153,1335,1427,1452]" gridrow="27" pageId="9" pageNumber="160" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="0358D41B192675F5ACC9FA57FE66CBCD" box="[153,503,1427,1452]" gridcol="0" gridrow="27" pageId="9" pageNumber="160">In percent of dorsal head length</th>
</tr>
<tr id="4089BD67192675F5ACC9FA7DFAA6CBB3" box="[153,1335,1465,1490]" gridrow="28" pageId="9" pageNumber="160">
<th id="0358D41B192675F5ACC9FA7DFE66CBB3" box="[153,503,1465,1490]" gridcol="0" gridrow="28" pageId="9" pageNumber="160">Snout length</th>
<td id="0358D41B192675F5AF57FA7DFCF1CBB3" box="[775,864,1465,1490]" gridcol="1" gridrow="28" pageId="9" pageNumber="160">49</td>
<td id="0358D41B192675F5A857FA7DFB17CBB3" box="[1031,1158,1465,1490]" gridcol="2" gridrow="28" pageId="9" pageNumber="160">47–52</td>
<td id="0358D41B192675F5A8AEFA7DFAA6CBB3" box="[1278,1335,1465,1490]" gridcol="3" gridrow="28" pageId="9" pageNumber="160">50</td>
</tr>
<tr id="4089BD67192675F5ACC9FA1BFAA6CB99" box="[153,1335,1503,1528]" gridrow="29" pageId="9" pageNumber="160">
<th id="0358D41B192675F5ACC9FA1BFE66CB99" box="[153,503,1503,1528]" gridcol="0" gridrow="29" pageId="9" pageNumber="160">Eye diameter</th>
<td id="0358D41B192675F5AF57FA1BFCF1CB99" box="[775,864,1503,1528]" gridcol="1" gridrow="29" pageId="9" pageNumber="160">18</td>
<td id="0358D41B192675F5A857FA1BFB17CB99" box="[1031,1158,1503,1528]" gridcol="2" gridrow="29" pageId="9" pageNumber="160">16–18</td>
<td id="0358D41B192675F5A8AEFA1BFAA6CB99" box="[1278,1335,1503,1528]" gridcol="3" gridrow="29" pageId="9" pageNumber="160">17</td>
</tr>
<tr id="4089BD67192675F5ACC9F9C1FAA6C87F" box="[153,1335,1541,1566]" gridrow="30" pageId="9" pageNumber="160">
<th id="0358D41B192675F5ACC9F9C1FE66C87F" box="[153,503,1541,1566]" gridcol="0" gridrow="30" pageId="9" pageNumber="160">Interorbital width</th>
<td id="0358D41B192675F5AF57F9C1FCF1C87F" box="[775,864,1541,1566]" gridcol="1" gridrow="30" pageId="9" pageNumber="160">41</td>
<td id="0358D41B192675F5A857F9C1FB17C87F" box="[1031,1158,1541,1566]" gridcol="2" gridrow="30" pageId="9" pageNumber="160">38–43</td>
<td id="0358D41B192675F5A8AEF9C1FAA6C87F" box="[1278,1335,1541,1566]" gridcol="3" gridrow="30" pageId="9" pageNumber="160">41</td>
</tr>
<tr id="4089BD67192675F5ACC9F9EFFAA6C825" box="[153,1335,1579,1604]" gridrow="31" pageId="9" pageNumber="160" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="0358D41B192675F5ACC9F9EFFE66C825" box="[153,503,1579,1604]" gridcol="0" gridrow="31" pageId="9" pageNumber="160">In percent of lateral head length</th>
</tr>
<tr id="4089BD67192675F5ACC9F994FAA6C808" box="[153,1335,1616,1641]" gridrow="32" pageId="9" pageNumber="160">
<th id="0358D41B192675F5ACC9F994FE66C808" box="[153,503,1616,1641]" gridcol="0" gridrow="32" pageId="9" pageNumber="160">Snout length</th>
<td id="0358D41B192675F5AF57F994FCF1C808" box="[775,864,1616,1641]" gridcol="1" gridrow="32" pageId="9" pageNumber="160">43</td>
<td id="0358D41B192675F5A857F994FB17C808" box="[1031,1158,1616,1641]" gridcol="2" gridrow="32" pageId="9" pageNumber="160">42–44</td>
<td id="0358D41B192675F5A8AEF994FAA6C808" box="[1278,1335,1616,1641]" gridcol="3" gridrow="32" pageId="9" pageNumber="160">43</td>
</tr>
<tr id="4089BD67192675F5ACC9F9B2FAA6C8EE" box="[153,1335,1654,1679]" gridrow="33" pageId="9" pageNumber="160">
<th id="0358D41B192675F5ACC9F9B2FE66C8EE" box="[153,503,1654,1679]" gridcol="0" gridrow="33" pageId="9" pageNumber="160">Eye diameter</th>
<td id="0358D41B192675F5AF57F9B2FCF1C8EE" box="[775,864,1654,1679]" gridcol="1" gridrow="33" pageId="9" pageNumber="160">15</td>
<td id="0358D41B192675F5A857F9B2FB17C8EE" box="[1031,1158,1654,1679]" gridcol="2" gridrow="33" pageId="9" pageNumber="160">14–15</td>
<td id="0358D41B192675F5A8AEF9B2FAA6C8EE" box="[1278,1335,1654,1679]" gridcol="3" gridrow="33" pageId="9" pageNumber="160">15</td>
</tr>
<tr id="4089BD67192675F5ACC9F958FAA6C8D4" box="[153,1335,1692,1717]" gridrow="34" pageId="9" pageNumber="160">
<th id="0358D41B192675F5ACC9F958FE66C8D4" box="[153,503,1692,1717]" gridcol="0" gridrow="34" pageId="9" pageNumber="160">Interorbital width</th>
<td id="0358D41B192675F5AF57F958FCF1C8D4" box="[775,864,1692,1717]" gridcol="1" gridrow="34" pageId="9" pageNumber="160">35</td>
<td id="0358D41B192675F5A857F958FB17C8D4" box="[1031,1158,1692,1717]" gridcol="2" gridrow="34" pageId="9" pageNumber="160">33–37</td>
<td id="0358D41B192675F5A8AEF958FAA6C8D4" box="[1278,1335,1692,1717]" gridcol="3" gridrow="34" pageId="9" pageNumber="160">35</td>
</tr>
</table>
</paragraph>
<paragraph id="FE06BF2519268A03AC97F922FF7DC9B9" blockId="9.[151,1437,1766,2045]" pageId="9" pageNumber="160">
<emphasis id="CCCD633719268A03AC97F922FEDEC960" bold="true" box="[199,335,1766,1793]" pageId="9" pageNumber="160">Coloration.</emphasis>
After 2 weeks in formalin (
<figureCitation id="6682A3A019268A03AED3F922FD43C960" box="[643,722,1766,1793]" captionStart="FIGURE 4" captionStartId="5.[151,250,998,1023]" captionTargetBox="[173,1412,217,909]" captionTargetId="figure@5.[153,1438,177,940]" captionTargetPageId="5" captionText="FIGURE 4. Rhyacoschistura larreci, MHNG 2727.011, holotype, 57.1 mm SL; Laos: Xayaburi: Nam Houng watershed." httpUri="https://zenodo.org/record/3234250/files/figure.png" pageId="9" pageNumber="160">Figs. 4</figureCitation>
,
<figureCitation id="6682A3A019268A03AE8EF922FD7FC960" box="[734,750,1766,1793]" captionStart="FIGURE 6" captionStartId="6.[151,250,1986,2011]" captionTargetBox="[174,1421,177,1950]" captionTargetId="figure@6.[164,1423,177,1950]" captionTargetPageId="6" captionText="FIGURE 6. Rhyacoschistura larreci, CMK 28037, paratypes; Laos: Xayaburi: Nam Houng watershed; a, 23.9 mm SL; b, 26.5 mm SL; c, 29.2 mm SL; d, 63.6 mm SL; e, 68.8 mm SL." httpUri="https://zenodo.org/record/3234254/files/figure.png" pageId="9" pageNumber="160">6</figureCitation>
). Body background colour beige to pale brown, darker posteriorly, paler on belly; except otherwise stated, all markings darker grey to dark brown. Head dark brown to blackish on top and side, with a variable pattern of pale to whitish patches, typically organised in: a large blotch on nape, and one on each side of it anteriorly; a v-shaped mark between eyes and another one between nostrils; a median longitudinal line on snout and one between eye and base of outer rostral barbel; and a white patch behind and posterior to eye. Rostral barbels dark brown; inner ones whitish distally. Maxillary barbels whitish with a narrow black area at base.
</paragraph>
<paragraph id="FE06BF2519268A00AC97F827FE2DCF22" blockId="9.[151,1437,1766,2045]" lastBlockId="10.[151,1437,153,1079]" lastPageId="10" lastPageNumber="161" pageId="9" pageNumber="160">
In large specimens (
<figureCitation id="6682A3A019268A03ADF7F826FE67C99C" box="[423,502,2018,2045]" captionStart="FIGURE 4" captionStartId="5.[151,250,998,1023]" captionTargetBox="[173,1412,217,909]" captionTargetId="figure@5.[153,1438,177,940]" captionTargetPageId="5" captionText="FIGURE 4. Rhyacoschistura larreci, MHNG 2727.011, holotype, 57.1 mm SL; Laos: Xayaburi: Nam Houng watershed." httpUri="https://zenodo.org/record/3234250/files/figure.png" pageId="9" pageNumber="160">
Figs.
<geoCoordinate id="9B8DD9E219268A03ADB6F827FE67C99C" box="[486,502,2019,2045]" degrees="4, 6" direction="east" orientation="longitude" pageId="9" pageNumber="160" precision="5555" value="4.6">4</geoCoordinate>
</figureCitation>
,
<figureCitation id="6682A3A019268A03AE51F826FD8EC99C" box="[513,543,2018,2045]" captionStart="FIGURE 6" captionStartId="6.[151,250,1986,2011]" captionTargetBox="[174,1421,177,1950]" captionTargetId="figure@6.[164,1423,177,1950]" captionTargetPageId="6" captionText="FIGURE 6. Rhyacoschistura larreci, CMK 28037, paratypes; Laos: Xayaburi: Nam Houng watershed; a, 23.9 mm SL; b, 26.5 mm SL; c, 29.2 mm SL; d, 63.6 mm SL; e, 68.8 mm SL." httpUri="https://zenodo.org/record/3234254/files/figure.png" pageId="9" pageNumber="160">6e</figureCitation>
), flank with numerous narrow slanted bars, extending downwards to level of in- sertion of pectoral fin, very irregularly organised and shaped, more or less connected, or sometimes covering whole flank. In smallest specimens, flank covered by dark pigments, denser on dorsal half (
<figureCitation id="6682A3A019258A00A81FFF79FB30CEB9" box="[1103,1185,189,216]" captionStart="FIGURE 6" captionStartId="6.[151,250,1986,2011]" captionTargetBox="[174,1421,177,1950]" captionTargetId="figure@6.[164,1423,177,1950]" captionTargetPageId="6" captionText="FIGURE 6. Rhyacoschistura larreci, CMK 28037, paratypes; Laos: Xayaburi: Nam Houng watershed; a, 23.9 mm SL; b, 26.5 mm SL; c, 29.2 mm SL; d, 63.6 mm SL; e, 68.8 mm SL." httpUri="https://zenodo.org/record/3234254/files/figure.png" pageId="10" pageNumber="161">Fig. 6a</figureCitation>
); with increasing size, pigments become denser at midheight of flank, forming a midlateral band from head to caudal-fin base (
<figureCitation id="6682A3A019258A00A971FF25FAF2CE9D" box="[1313,1379,225,252]" captionStart="FIGURE 6" captionStartId="6.[151,250,1986,2011]" captionTargetBox="[174,1421,177,1950]" captionTargetId="figure@6.[164,1423,177,1950]" captionTargetPageId="6" captionText="FIGURE 6. Rhyacoschistura larreci, CMK 28037, paratypes; Laos: Xayaburi: Nam Houng watershed; a, 23.9 mm SL; b, 26.5 mm SL; c, 29.2 mm SL; d, 63.6 mm SL; e, 68.8 mm SL." httpUri="https://zenodo.org/record/3234254/files/figure.png" pageId="10" pageNumber="161">Fig. 6</figureCitation>
b–c), later dissociated into irregular bars (
<figureCitation id="6682A3A019258A00AE75FEC1FDE9CF41" box="[549,632,261,288]" captionStart="FIGURE 6" captionStartId="6.[151,250,1986,2011]" captionTargetBox="[174,1421,177,1950]" captionTargetId="figure@6.[164,1423,177,1950]" captionTargetPageId="6" captionText="FIGURE 6. Rhyacoschistura larreci, CMK 28037, paratypes; Laos: Xayaburi: Nam Houng watershed; a, 23.9 mm SL; b, 26.5 mm SL; c, 29.2 mm SL; d, 63.6 mm SL; e, 68.8 mm SL." httpUri="https://zenodo.org/record/3234254/files/figure.png" pageId="10" pageNumber="161">Fig. 6d</figureCitation>
). Back crossed by narrow bars, which are often distinct and merging into a dark background colour.
</paragraph>
<paragraph id="FE06BF2519258A00AC97FE89FD78CFEA" blockId="10.[151,1437,153,1079]" pageId="10" pageNumber="161">Black bar at caudal base arched, extending on base of rays, almost complete, except along upper and lower extremities. A narrow pale band in front of black bar.</paragraph>
<paragraph id="FE06BF2519258A00AC97FE51FEF2CC7A" blockId="10.[151,1437,153,1079]" pageId="10" pageNumber="161">Dorsal fin hyaline; in smaller specimens with a black spot at base of unbranched ray and first branched ray; in larger specimens black band extending along entire base. A black mark at midlength of membrane covering unbranched rays. A few black pigments along rays and between segments at level of first branching point, forming a faint row of spots.</paragraph>
<paragraph id="FE06BF2519258A00AC97FDE1FDF1CC5E" blockId="10.[151,1437,153,1079]" box="[199,608,549,575]" pageId="10" pageNumber="161">Caudal, anal and pelvic fins hyaline.</paragraph>
<paragraph id="FE06BF2519258A00AC97FD8DFBFFCCE6" blockId="10.[151,1437,153,1079]" pageId="10" pageNumber="161">Pectoral fin hyaline; black pigments along middle posterior edge of unbranched and branched rays 1–2, and their branches, forming faint longitudinal marks, more intense in males than in females.</paragraph>
<paragraph id="FE06BF2519258A00AC97FD55FEE7CC95" blockId="10.[151,1437,153,1079]" pageId="10" pageNumber="161">
In life (
<figureCitation id="6682A3A019258A00AD4BFD55FECECCCA" box="[283,351,656,683]" captionStart="FIGURE 5" captionStartId="5.[151,250,1802,1827]" captionTargetBox="[153,1435,1056,1736]" captionTargetId="figure@5.[151,1436,1055,1737]" captionTargetPageId="5" captionText="FIGURE 5. Rhyacoschistura larreci, CMK 28037, paratype, 57.8 mm SL; Laos: Xayaburi: Nam Houng watershed; immediately after fixation." httpUri="https://zenodo.org/record/3234252/files/figure.png" pageId="10" pageNumber="161">Fig. 5</figureCitation>
): background colour pale grey, marks dark brown, bar at caudal-fin base and markings on dorsal fin black. Fins yellowish, pectoral fin more intense. (Sampling happened in late afternoon and in rain, therefore vague information).
</paragraph>
<paragraph id="FE06BF2519258A00AC97FD39FAA9CD76" blockId="10.[151,1437,153,1079]" box="[199,1336,764,791]" pageId="10" pageNumber="161">
<emphasis id="CCCD633719258A00AC97FD39FE03CD76" bold="true" box="[199,402,764,791]" pageId="10" pageNumber="161">Notes on biology.</emphasis>
Two dissected females (CMK 28037, 50.2,
<quantity id="394112C019258A00AFD0FD39FC75CD76" box="[896,996,765,791]" metricMagnitude="-2" metricUnit="m" metricValue="6.29" pageId="10" pageNumber="161" unit="mm" value="62.9">62.9 mm</quantity>
SL) did not have ripe ovaries.
</paragraph>
</subSubSection>
<subSubSection id="B6A3ECAE19258A00AC97FCE4FC82CDAA" pageId="10" pageNumber="161" type="distribution">
<paragraph id="FE06BF2519258A00AC97FCE4FC82CDAA" blockId="10.[151,1437,153,1079]" pageId="10" pageNumber="161">
<emphasis id="CCCD633719258A00AC97FCE4FEF0CD5A" bold="true" box="[199,353,800,827]" pageId="10" pageNumber="161">Distribution.</emphasis>
<taxonomicName id="39B9C4A619258A00AD3AFCE5FDE9CD5A" authorityName="Kottelat" authorityYear="2019" box="[362,632,801,827]" class="Actinopterygii" family="Nemacheilidae" genus="Rhyacoschistura" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="161" phylum="Chordata" rank="species" species="larreci">
<emphasis id="CCCD633719258A00AD3AFCE5FDE9CD5A" box="[362,632,801,827]" italics="true" pageId="10" pageNumber="161">Rhyacoschistura larreci</emphasis>
</taxonomicName>
is presently known only from the
<typeStatus id="2102018719258A00A859FCE5FBA8CD5A" box="[1033,1081,801,827]" pageId="10" pageNumber="161">type</typeStatus>
locality, Houay Plong, a small mountain creek tributary of Nam Met, itself a tributary of Nam Houng, itself a tributary of Mekong. It was obtained in a very small stream, about
<quantity id="394112C019258A00ADB9FCADFD84CDE2" box="[489,533,873,899]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="161" unit="m" value="1.0">1 m</quantity>
wide and
<quantity id="394112C019258A00AEDDFCADFD42CDE2" box="[653,723,873,899]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="161" unit="cm" value="20.0">20 cm</quantity>
deep, among stones and boulders. It is probably present in the uppermost parts of many other tributaries of the Nam Met, and possibly adjacent streams, but time was not available to reach and sample more sites with similar morphology.
</paragraph>
</subSubSection>
<subSubSection id="B6A3ECAE19258A00AC97FC11FE01CA56" pageId="10" pageNumber="161" type="etymology">
<paragraph id="FE06BF2519258A00AC97FC11FE01CA56" blockId="10.[151,1437,153,1079]" pageId="10" pageNumber="161">
<emphasis id="CCCD633719258A00AC97FC11FEDFCD8E" bold="true" box="[199,334,981,1007]" pageId="10" pageNumber="161">Etymology.</emphasis>
Named for LARReC, Living Aquatic Resources Research Center,
<collectingRegion id="3C7D71C719258A00A811FC11FB3ECD8E" box="[1089,1199,981,1007]" country="Laos" name="Viangchan" pageId="10" pageNumber="161">Vientiane</collectingRegion>
, for its 20th anniversary, and appreciation to several of its staff for 20 years of collaboration in the field. Treated as a masculine noun in genitive, indeclinable.
</paragraph>
</subSubSection>
</treatment>
</document>
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