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<mods:title id="84BED4F8310A38A00043148035B2ED18">Brachyuran crabs of the family Macrophthalmidae Dana, 1851 (Decapoda: Brachyura: Macrophthalmidae) of the Persian Gulf</mods:title>
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<mods:namePart id="62757A7BE7467DB85CAB984BE849D50C">Naderloo, Reza</mods:namePart>
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<mods:namePart id="E8CE65B3B5691A3A99D2E6585B194B7D">Türkay, Michael</mods:namePart>
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<mods:namePart id="8F0A981864134FADEB4DAA2F61A7AC73">Apel, Michael</mods:namePart>
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<treatment id="587287CE552BFFE6FF774B62FBE07ADC" ID-DOI="http://doi.org/10.5281/zenodo.5417620" ID-GBIF-Taxon="119410913" ID-Zenodo-Dep="5417620" LSID="urn:lsid:plazi:treatment:587287CE552BFFE6FF774B62FBE07ADC" httpUri="http://treatment.plazi.org/id/587287CE552BFFE6FF774B62FBE07ADC" lastPageId="22" lastPageNumber="23" pageId="18" pageNumber="19">
<subSubSection id="98C16553552BFFE2FF774B62FE357DA4" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="D06436D8552BFFE2FF774B62FCE37D83" blockId="18.[151,771,1905,1964]" box="[151,771,1905,1932]" pageId="18" pageNumber="19">
<heading id="8B2C81B4552BFFE2FF774B62FCE37D83" bold="true" box="[151,771,1905,1932]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
<taxonomicName id="17DB4D5B552BFFE2FF774B62FCE37D83" ID-CoL="3X864" authority="A. Milne-Edwards, 1873" authorityName="A. Milne-Edwards" authorityYear="1873" box="[151,771,1905,1932]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="graeffei">
<emphasis id="E2AFEACA552BFFE2FF774B62FCE37D83" bold="true" box="[151,771,1905,1932]" pageId="18" pageNumber="19">
<emphasis id="E2AFEACA552BFFE2FF774B62FE2C7D84" bold="true" box="[151,460,1906,1932]" italics="true" pageId="18" pageNumber="19">Macrophthalmus graeffei</emphasis>
<bibRefCitation id="B44A4B29552BFFE2FE334B61FCE37D83" author="Milne-Edwards" box="[467,771,1905,1931]" pageId="18" pageNumber="19" refString="Milne-Edwards, A. (1873) Description de quelques Crustaces nouveaux ou peu connus provenant du Musee de M. C. Godeffroy. Journal des Museum Godeffroy, 1 (4), 77 - 88 (253 - 264), pls. 12 - 13." type="journal article" year="1873">A. Milne-Edwards, 1873</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="D06436D8552BFFE2FF774B84FE357DA4" blockId="18.[151,771,1905,1964]" box="[151,469,1940,1964]" pageId="18" pageNumber="19">
(
<figureCitation id="48E02A5D552BFFE2FF7F4B84FF197DA4" box="[159,249,1940,1964]" captionStart="FIGURE 11" captionStartId="20.[151,250,1633,1656]" captionTargetBox="[193,1397,215,1595]" captionTargetId="figure@20.[171,1416,193,1612]" captionTargetPageId="20" captionText="FIGURE 11. Macrophthalmus graeffei A. Milne-Edwards 1873, male (ZMUC CRU- 1782): a, Posterior surface of carapace, left side; b, merus of male cheliped, inner surface; c, cheliped of male, outer surface; d, cheliped of male, inner surface; e, male abdomen." httpUri="https://zenodo.org/record/203109/files/figure.png" pageId="18" pageNumber="19">Figs. 11</figureCitation>
ae, 12ad, 17a, b)
</paragraph>
</subSubSection>
<subSubSection id="98C16553552BFFE3FF774BCDFAF67B01" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="reference_group">
<paragraph id="D06436D8552BFFE2FF774BCDFB947DFA" blockId="18.[151,1140,2012,2035]" box="[151,1140,2012,2035]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="F0CB11F6552BFFE2FF774BCDFB947DFA" box="[151,1140,2012,2035]" pageId="18" pageNumber="19">
<taxonomicName id="17DB4D5B552BFFE2FF774BCDFD297DFA" ID-CoL="3X864" authority="A. Milne-Edwards 1873: 257" authorityName="A. Milne-Edwards" authorityPageNumber="257" authorityYear="1873" box="[151,713,2012,2035]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="graeffei">
<emphasis id="E2AFEACA552BFFE2FF774BCDFE787DFB" box="[151,408,2013,2035]" italics="true" pageId="18" pageNumber="19">Macrophthalmus graeffei</emphasis>
<treatmentCitation id="517A10C9552BFFE2FE7E4BCDFD297DFA" author="Milne-Edwards" box="[414,713,2012,2034]" page="257" pageId="18" pageNumber="19" year="1873">
<bibRefCitation id="B44A4B29552BFFE2FE7E4BCDFD777DFA" author="Milne-Edwards" box="[414,663,2012,2034]" pageId="18" pageNumber="19" refString="Milne-Edwards, A. (1873) Description de quelques Crustaces nouveaux ou peu connus provenant du Musee de M. C. Godeffroy. Journal des Museum Godeffroy, 1 (4), 77 - 88 (253 - 264), pls. 12 - 13." type="journal article" year="1873">A. Milne-Edwards 1873</bibRefCitation>
: 257
</treatmentCitation>
</taxonomicName>
, pl. 2, fig. 5. —
<treatmentCitation id="517A10C9552BFFE2FC8F4BCCFBFD7DFA" author="Titgen" box="[879,1053,2012,2034]" page="253" pageId="18" pageNumber="19" year="1982">
<bibRefCitation id="B44A4B29552BFFE2FC8F4BCCFC0D7DFA" author="Titgen" box="[879,1005,2012,2034]" pageId="18" pageNumber="19" refString="Titgen, R. H. (1982) The Systematics and Ecology of the Decapods of Dubai, and their Zoogeographic Relationships to the Arabian Gulf and the Western Indian Ocean. Unpublished D. Phil. Thesis, Texas A &amp; M University." type="book" year="1982">Titgen 1982</bibRefCitation>
: 253
</treatmentCitation>
(in list).
</treatmentCitationGroup>
</paragraph>
<paragraph id="D06436D8552AFFE3FF774C88FD417AC4" blockId="19.[151,1436,152,266]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="F0CB11F6552AFFE3FF774C88FD417AC4" bridged="true" pageId="19" pageNumber="20">
<taxonomicName id="17DB4D5B552AFFE3FF774C88FDBF7AA6" ID-CoL="3X864" box="[151,607,152,174]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="graeffei" subGenus="Macrophthalmus">
<emphasis id="E2AFEACA552AFFE3FF774C88FDBF7AA6" box="[151,607,152,174]" italics="true" pageId="19" pageNumber="20">Macrophthalmus (Macrophthalmus) graeffei</emphasis>
</taxonomicName>
<treatmentCitation id="517A10C9552AFFE3FD694C89FCA37AA6" author="Barnes" box="[649,835,152,174]" page="225" pageId="19" pageNumber="20" year="1970">
<bibRefCitation id="B44A4B29552AFFE3FD694C89FCEE7AA5" author="Barnes" box="[649,782,152,174]" pageId="19" pageNumber="20" refString="Barnes, R. S. K. (1970) The species of Macrophthalmus in the collections of the British Museum (Natural History). Bulletin of the British Museum of Natural History (Zoology), 20, 203 - 251." type="journal article" year="1970">Barnes 1970</bibRefCitation>
: 225
</treatmentCitation>
; 1971: 13, 36 (in key), fig. 3; 1977: 276 (in key), 279 (in list); 2010: 34 (in key), 39. —
<treatmentCitation id="517A10C9552AFFE3FE1D4CA6FD7D7AC4" author="Apel" box="[509,669,182,204]" page="108" pageId="19" pageNumber="20" year="2001">
<bibRefCitation id="B44A4B29552AFFE3FE1D4CA6FD8A7AC4" author="Apel" box="[509,618,182,204]" pageId="19" pageNumber="20" refString="Apel, M. (2001) Taxonomie und Zoogeographie der Brachyura, Paguridea und Porcellanidae (Crustacea: Decapoda) des Persisch-Arabischen Golfes unpublished Ph. D. Thesis, Johann Wolfgang Goethe-Universitat, Frankfurt am Main 268 pp." type="book" year="2001">Apel 2001</bibRefCitation>
: 108
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="D06436D8552AFFE3FF774CC5FAF67B01" blockId="19.[151,1436,152,266]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="F0CB11F6552AFFE3FF774CC5FAF67B01" pageId="19" pageNumber="20">
<taxonomicName id="17DB4D5B552AFFE3FF774CC5FEA37AE3" ID-CoL="7PDBN" box="[151,323,213,235]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="E2AFEACA552AFFE3FF774CC5FEA37AE3" box="[151,323,213,235]" italics="true" pageId="19" pageNumber="20">Macrophthalmus</emphasis>
</taxonomicName>
(
<taxonomicName id="17DB4D5B552AFFE3FEB24CC6FDF67AE3" ID-CoL="3X85Y" authority="Stimpson" authorityName="Stimpson" box="[338,534,213,235]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="convexus">
<emphasis id="E2AFEACA552AFFE3FEB24CC6FE4F7AE3" box="[338,431,214,235]" italics="true" pageId="19" pageNumber="20">convexus</emphasis>
Stimpson
</taxonomicName>
?) —
<treatmentCitation id="517A10C9552AFFE3FDAD4CC5FCD07AE2" author="Stephensen" box="[589,816,213,235]" page="191" pageId="19" pageNumber="20" year="1945">
<bibRefCitation id="B44A4B29552AFFE3FDAD4CC5FD1D7AE3" author="Stephensen" box="[589,765,213,235]" pageId="19" pageNumber="20" refString="Stephensen, K. (1945) The Brachyura of the Iranian Gulf. Danish Scientific Investigations in Iran, Part IV. E. Munksgaard, Copenhagen, 57 - 237." type="book chapter" year="1945">Stephensen 1945</bibRefCitation>
: 191
</treatmentCitation>
. [not
<taxonomicName id="17DB4D5B552AFFE3FC884CC5FAC27AE3" ID-CoL="3X85Y" authority="Stimpson, 1858" authorityName="Stimpson" authorityYear="1858" box="[872,1314,213,235]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="convexus">
<emphasis id="E2AFEACA552AFFE3FC884CC5FB987AE3" box="[872,1144,213,235]" italics="true" pageId="19" pageNumber="20">Macrophthalmus convexus</emphasis>
Stimpson, 1858
</taxonomicName>
].
<taxonomicName id="17DB4D5B552AFFE3FF774CE4FE467B02" ID-CoL="3X85Y" box="[151,422,244,266]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="convexus">
<emphasis id="E2AFEACA552AFFE3FF774CE4FE467B02" box="[151,422,244,266]" italics="true" pageId="19" pageNumber="20">Macrophthalmus convexus</emphasis>
</taxonomicName>
<treatmentCitation id="517A10C9552AFFE3FE2C4CE3FD9E7B01" author="Guinot" box="[460,638,243,265]" page="282" pageId="19" pageNumber="20" year="1967">
<bibRefCitation id="B44A4B29552AFFE3FE2C4CE3FDAF7B01" author="Guinot" box="[460,591,243,265]" pageId="19" pageNumber="20" refString="Guinot, D. (1967) La faune carcinologique (Crustacea, Brachyura) de l'Ocean Indien occidental et de la Mer Rouge. Catalogue remarques biogeographiques et bibliographie. Memoires de l'Institut fondamental d'Afrique noire, 77, 235 - 352." type="journal article" year="1967">Guinot 1967</bibRefCitation>
: 282
</treatmentCitation>
(in list; partly). [not
<taxonomicName id="17DB4D5B552AFFE3FCB34CE4FAEC7B01" ID-CoL="3X85Y" authority="Stimpson, 1858" authorityName="Stimpson" authorityYear="1858" box="[851,1292,243,266]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="convexus">
<emphasis id="E2AFEACA552AFFE3FCB34CE4FB827B02" box="[851,1122,244,266]" italics="true" pageId="19" pageNumber="20">Macrophthalmus convexus</emphasis>
Stimpson, 1858
</taxonomicName>
].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="98C16553552AFFE3FF774D27FC4B780A" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="D06436D8552AFFE3FF774D27FE0F7B46" blockId="19.[151,1437,310,1882]" box="[151,495,310,335]" pageId="19" pageNumber="20">
<emphasis id="E2AFEACA552AFFE3FF774D27FED57B47" bold="true" box="[151,309,310,335]" pageId="19" pageNumber="20">
<typeStatus id="0F60887A552AFFE3FF774D27FF327B47" box="[151,210,311,335]" pageId="19" pageNumber="20">Type</typeStatus>
locality.
</emphasis>
Upolu (
<collectingCountry id="A8CC7648552AFFE3FE714D26FE037B46" box="[401,483,310,334]" name="Samoa" pageId="19" pageNumber="20">Samoa</collectingCountry>
).
</paragraph>
<paragraph id="D06436D8552AFFE3FF254D4AFDBD7BB2" blockId="19.[151,1437,310,1882]" pageId="19" pageNumber="20">
<emphasis id="E2AFEACA552AFFE3FF254D4AFE517B7B" bold="true" box="[197,433,346,371]" pageId="19" pageNumber="20">Examined material.</emphasis>
<emphasis id="E2AFEACA552AFFE3FE574D4CFDA77B7B" box="[439,583,347,371]" italics="true" pageId="19" pageNumber="20">Persian Gulf</emphasis>
:
<emphasis id="E2AFEACA552AFFE3FDB54D4CFD657B7B" box="[597,645,348,371]" italics="true" pageId="19" pageNumber="20">
<collectingCountry id="A8CC7648552AFFE3FDB54D4CFD657B7B" box="[597,645,348,371]" name="Iran" pageId="19" pageNumber="20">Iran</collectingCountry>
</emphasis>
:
<specimenCount id="C6DDFD51552AFFE3FD744D4AFD3E7B7A" box="[660,734,346,370]" pageId="19" pageNumber="20" type="male">1 male</specimenCount>
(CL = 14.8, CB =
<quantity id="17239B3D552AFFE3FC504D4AFBEA7B7A" box="[944,1034,346,370]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="19" pageNumber="20" unit="mm" value="8.4">8.4 mm</quantity>
),
<specimenCount id="C6DDFD51552AFFE3FBFC4D4AFB877B7A" box="[1052,1127,346,370]" pageId="19" pageNumber="20" type="male">1 male</specimenCount>
(juv.) (
<collectionCode id="B6CAAE1D552AFFE3FB584D4AFAEB7B7A" box="[1208,1291,346,370]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="19" pageNumber="20">ZMUC</collectionCode>
<collectionCode id="B6CAAE1D552AFFE3FAF14D4AFAB77B7A" box="[1297,1367,346,370]" pageId="19" pageNumber="20">CRU-</collectionCode>
1782),
<collectionCode id="B6CAAE1D552AFFE3FF774D6EFF2D7B9E" box="[151,205,382,406]" pageId="19" pageNumber="20">DSII</collectionCode>
St. 26, O, Kharg I., sand and shells,
<quantity id="17239B3D552AFFE3FD8D4D6EFD477B9E" box="[621,679,382,406]" metricMagnitude="1" metricUnit="m" metricValue="1.8" pageId="19" pageNumber="20" unit="m" value="18.0">18 m</quantity>
deep,
<date id="A4651018552AFFE3FD124D6EFC927B9E" box="[754,882,382,406]" pageId="19" pageNumber="20" value="1937-03-15">15.03.1937</date>
, G. Thorson, identified as
<taxonomicName id="17DB4D5B552AFFE3FB7B4D6EFAB97B9E" box="[1179,1369,382,406]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="E2AFEACA552AFFE3FB7B4D6EFAB97B9E" box="[1179,1369,382,406]" italics="true" pageId="19" pageNumber="20">Macrophthalmus</emphasis>
</taxonomicName>
(
<taxonomicName id="17DB4D5B552AFFE3FA8A4D90FEA97BB2" authority="Stimpson" authorityName="Stimpson" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="convexus">
<emphasis id="E2AFEACA552AFFE3FA8A4D90FF357BB3" italics="true" pageId="19" pageNumber="20">convexus</emphasis>
Stimpson
</taxonomicName>
?) by
<bibRefCitation id="B44A4B29552AFFE3FE654DB2FDB97BB2" author="Stephensen" box="[389,601,418,442]" pageId="19" pageNumber="20" refString="Stephensen, K. (1945) The Brachyura of the Iranian Gulf. Danish Scientific Investigations in Iran, Part IV. E. Munksgaard, Copenhagen, 57 - 237." type="book chapter" year="1945">Stephensen (1945)</bibRefCitation>
.
</paragraph>
<paragraph id="D06436D8552AFFE3FF254DD6FC4B780A" blockId="19.[151,1437,310,1882]" pageId="19" pageNumber="20">
<emphasis id="E2AFEACA552AFFE3FF254DD6FE397BD7" bold="true" box="[197,473,454,479]" pageId="19" pageNumber="20">Comparative material.</emphasis>
<emphasis id="E2AFEACA552AFFE3FE044DD6FDA37BD6" box="[484,579,454,478]" italics="true" pageId="19" pageNumber="20">Red Sea</emphasis>
:
<specimenCount id="C6DDFD51552AFFE3FDB64DD6FD267BD6" box="[598,710,454,478]" pageId="19" pageNumber="20" type="female">3 females</specimenCount>
(NHM 1934: 1:17: 133135, appendages are broken),
<collectingCountry id="A8CC7648552AFFE3FAAF4DD6FA787BD6" box="[1359,1432,454,478]" name="Sudan" pageId="19" pageNumber="20">Sudan</collectingCountry>
, Mersa Marja in Shab-ul-Shumur, coll. C. Crossland, det. R. D. Laurie.
</paragraph>
</subSubSection>
<subSubSection id="98C16553552AFFE4FF254E1EFC1E7D8C" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="description">
<paragraph id="D06436D8552AFFE3FF254E1EFB7B78BE" blockId="19.[151,1437,310,1882]" pageId="19" pageNumber="20">
<emphasis id="E2AFEACA552AFFE3FF254E1EFE93782F" bold="true" box="[197,371,526,551]" pageId="19" pageNumber="20">Redescription.</emphasis>
Carapace (
<figureCitation id="48E02A5D552AFFE3FE134E1EFDB2782E" box="[499,594,526,550]" captionStart="FIGURE 17. a, b" captionStartId="29.[151,250,1350,1373]" captionTargetBox="[211,1376,203,1316]" captionTargetId="figure@29.[199,1387,193,1329]" captionTargetPageId="29" captionText="FIGURE 17. a, b: Macrophthalmus graeffei A. Milne-Edwards 1873, male (UZM 1782), a, posterior surface; b, ventral surface; c, d: Macrophthalmus serenei Takeda &amp; Komai, 1991, male (SMF 36097), a, posterior surface; b, anterior surface; e, f: Macrophthalmus sinuspersici Naderloo &amp; Türkay, 2010, holotype, male (SMF 3690), e, posterior surface; f, ventral surface." httpUri="https://zenodo.org/record/203115/files/figure.png" pageId="19" pageNumber="20">Fig. 17a</figureCitation>
) distinctly wider than long (CB/CL = 1.84), moderately convex, posterior surface smooth; lump of small granules on epibranchial region, granules round; scarce granules on lateral regions. Regions relatively well defined; furrows delimiting gastric region remarkably deep, transverse furrow of epibranchial region nearly shallow depression. Front deflexed, narrow, about 0.12 times as wide as carapace, constricted medially; anterior edge smooth, very slightly concave, faintly bi-lobed, frontal furrow faint.
</paragraph>
<paragraph id="D06436D8552AFFE3FF254ED2FB577962" blockId="19.[151,1437,310,1882]" pageId="19" pageNumber="20">Lateral margin with three distinct teeth including exorbital angle; first tooth remarkably long, longer than second, elongatedly triangular, directed outwards, slightly directed forwards, its posterior margin smooth; second tooth about half as long as first one, triangular, directed outwards, separated from former by long V-shaped notch; third smallest, broadly triangular, directed outwards; posterolateral margin nearly straight, very slightly converging posteriorly, serrated with pointed granules, granules larger posteriorly; posterior ridge smooth.</paragraph>
<paragraph id="D06436D8552AFFE3FF254F66FD1179F2" blockId="19.[151,1437,310,1882]" pageId="19" pageNumber="20">
Eyestalk narrow (
<figureCitation id="48E02A5D552AFFE3FE6F4F66FE0B7986" box="[399,491,886,910]" captionStart="FIGURE 11" captionStartId="20.[151,250,1633,1656]" captionTargetBox="[193,1397,215,1595]" captionTargetId="figure@20.[171,1416,193,1612]" captionTargetPageId="20" captionText="FIGURE 11. Macrophthalmus graeffei A. Milne-Edwards 1873, male (ZMUC CRU- 1782): a, Posterior surface of carapace, left side; b, merus of male cheliped, inner surface; c, cheliped of male, outer surface; d, cheliped of male, inner surface; e, male abdomen." httpUri="https://zenodo.org/record/203109/files/figure.png" pageId="19" pageNumber="20">Figs. 11</figureCitation>
a, 17a, b), long, extending slightly less than half of cornea beyond exorbital angle, distal part of cornea with remarkably narrow bulge (
<figureCitation id="48E02A5D552AFFE3FD014F8AFCD179BA" box="[737,817,922,946]" captionStart="FIGURE 11" captionStartId="20.[151,250,1633,1656]" captionTargetBox="[193,1397,215,1595]" captionTargetId="figure@20.[171,1416,193,1612]" captionTargetPageId="20" captionText="FIGURE 11. Macrophthalmus graeffei A. Milne-Edwards 1873, male (ZMUC CRU- 1782): a, Posterior surface of carapace, left side; b, merus of male cheliped, inner surface; c, cheliped of male, outer surface; d, cheliped of male, inner surface; e, male abdomen." httpUri="https://zenodo.org/record/203109/files/figure.png" pageId="19" pageNumber="20">Fig. 11</figureCitation>
a); upper orbital margin slightly convex, gently sloping outward, regularly granular, granules very small, low blunt, smaller laterally; lower margin with granules, relatively large, pointed, of various size, smaller laterally.
</paragraph>
<paragraph id="D06436D8552AFFE3FF254816FDCC7E82" blockId="19.[151,1437,310,1882]" pageId="19" pageNumber="20">Third maxilliped large, ischium about twice as long as merus, inner margin of ischium with long setae, outer margin nearly straight, without setae, outer surface smooth; merus subquadrate, wide, about 1.7 times as wide as long, inner margin with long setae, outer margin proximally swollen, distally strongly converging, finely dentate, without setae, outer surface smooth.</paragraph>
<paragraph id="D06436D8552AFFE3FF254886FBD27C8A" blockId="19.[151,1437,310,1882]" pageId="19" pageNumber="20">
Chelipeds (
<figureCitation id="48E02A5D552AFFE3FEA54886FE417EA6" box="[325,417,1174,1198]" captionStart="FIGURE 11" captionStartId="20.[151,250,1633,1656]" captionTargetBox="[193,1397,215,1595]" captionTargetId="figure@20.[171,1416,193,1612]" captionTargetPageId="20" captionText="FIGURE 11. Macrophthalmus graeffei A. Milne-Edwards 1873, male (ZMUC CRU- 1782): a, Posterior surface of carapace, left side; b, merus of male cheliped, inner surface; c, cheliped of male, outer surface; d, cheliped of male, inner surface; e, male abdomen." httpUri="https://zenodo.org/record/203109/files/figure.png" pageId="19" pageNumber="20">Figs. 11</figureCitation>
c, d) nearly equal. Merus (
<figureCitation id="48E02A5D552AFFE3FD2D4886FCFC7EA6" box="[717,796,1174,1198]" captionStart="FIGURE 11" captionStartId="20.[151,250,1633,1656]" captionTargetBox="[193,1397,215,1595]" captionTargetId="figure@20.[171,1416,193,1612]" captionTargetPageId="20" captionText="FIGURE 11. Macrophthalmus graeffei A. Milne-Edwards 1873, male (ZMUC CRU- 1782): a, Posterior surface of carapace, left side; b, merus of male cheliped, inner surface; c, cheliped of male, outer surface; d, cheliped of male, inner surface; e, male abdomen." httpUri="https://zenodo.org/record/203109/files/figure.png" pageId="19" pageNumber="20">Fig. 11</figureCitation>
b) with outer margin bearing large denticles along entire length, denticles slightly larger medially; inner upper margin with few small granules, with long setae, continuing to inner surface; inner lower margin with small granules; distal margin with very small granules. Carpus outer surface with small granules, getting dense towards upper, lower surfaces, medially smooth; inner distal margin serrated with small granules. Palm relatively long, about 1.6 times as long as high in distal portion; outer surface (
<figureCitation id="48E02A5D552AFFE3FA8F4936FF527F6A" captionStart="FIGURE 11" captionStartId="20.[151,250,1633,1656]" captionTargetBox="[193,1397,215,1595]" captionTargetId="figure@20.[171,1416,193,1612]" captionTargetPageId="20" captionText="FIGURE 11. Macrophthalmus graeffei A. Milne-Edwards 1873, male (ZMUC CRU- 1782): a, Posterior surface of carapace, left side; b, merus of male cheliped, inner surface; c, cheliped of male, outer surface; d, cheliped of male, inner surface; e, male abdomen." httpUri="https://zenodo.org/record/203109/files/figure.png" pageId="19" pageNumber="20">Fig. 11</figureCitation>
c) with small granules except in lower distal portion at base of immovable finger, granules continuous to upper, lower margins, granules on upper margin slightly larger than those of lower margin; longitudinal ridge on lower portion, nearly straight, parallel to lower margin, running from proximal part to near end of immovable finger, finely granular, granules small; inner surface (
<figureCitation id="48E02A5D552AFFE3FD7A49A6FD147FC6" box="[666,756,1462,1486]" captionStart="FIGURE 11" captionStartId="20.[151,250,1633,1656]" captionTargetBox="[193,1397,215,1595]" captionTargetId="figure@20.[171,1416,193,1612]" captionTargetPageId="20" captionText="FIGURE 11. Macrophthalmus graeffei A. Milne-Edwards 1873, male (ZMUC CRU- 1782): a, Posterior surface of carapace, left side; b, merus of male cheliped, inner surface; c, cheliped of male, outer surface; d, cheliped of male, inner surface; e, male abdomen." httpUri="https://zenodo.org/record/203109/files/figure.png" pageId="19" pageNumber="20">Figs. 11</figureCitation>
d, 17b) without patch of setae, small granules on lower proximal portion, continuous medially to near base of fingers. Movable finger relatively long, curved strongly inward, upper margin with large denticles along proximal two thirds, larger distally, small granules proximally; cutting edge with small differentiated tooth subproximally, teeth along cutting edge small, round, long setae along inner lower surface in movable finger. Immovable finger short, cutting edge elevated medially, large teeth along it, long setae along inner surface; tip of cutting edge of both fingers remarkably narrowed.
</paragraph>
<paragraph id="D06436D8552AFFE3FF254A9EFF367D1A" blockId="19.[151,1437,310,1882]" pageId="19" pageNumber="20">Walking legs medium-size, relatively narrow; merus of third walking leg about 3.4 times as long as wide; merus of second, third with large subdistal tooth; anterior, posterior margins serrated; margins of carpus, dactylus smooth; dactylus of third walking leg about as long as propodus; margins of segments scarcely beset with long setae.</paragraph>
<paragraph id="D06436D8552AFFE3FF254B0EFEE37D52" blockId="19.[151,1437,310,1882]" pageId="19" pageNumber="20">Posteromedian margin of epistome moderately convex, anterior part of buccal cavity distinctly swollen, without ridge.</paragraph>
<caption id="84A46650552DFFE4FF774A71FF1B7CBC" httpUri="https://zenodo.org/record/203109/files/figure.png" pageId="20" pageNumber="21" targetBox="[193,1397,215,1595]" targetPageId="20">
<paragraph id="D06436D8552DFFE4FF774A71FF1B7CBC" blockId="20.[151,1436,1633,1716]" pageId="20" pageNumber="21">
<emphasis id="E2AFEACA552DFFE4FF774A71FEFF7C7F" bold="true" box="[151,287,1633,1656]" pageId="20" pageNumber="21">FIGURE 11.</emphasis>
<taxonomicName id="17DB4D5B552DFFE4FEC74A71FCCD7C7F" authority="A. Milne-Edwards 1873" authorityName="A. Milne-Edwards" authorityYear="1873" box="[295,813,1633,1655]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="graeffei">
<emphasis id="E2AFEACA552DFFE4FEC74A71FDCA7C7F" box="[295,554,1633,1655]" italics="true" pageId="20" pageNumber="21">Macrophthalmus graeffei</emphasis>
A. Milne-Edwards 1873
</taxonomicName>
, male (ZMUC CRU-1782): a, Posterior surface of carapace, left side; b, merus of male cheliped, inner surface; c, cheliped of male, outer surface; d, cheliped of male, inner surface; e, male abdomen.
</paragraph>
</caption>
<paragraph id="D06436D8552DFFE4FF254ACCFAE27D10" blockId="20.[151,1436,1756,2033]" pageId="20" pageNumber="21">
Male abdomen (
<figureCitation id="48E02A5D552DFFE4FE614ACCFE3F7CFC" box="[385,479,1756,1780]" captionStart="FIGURE 11" captionStartId="20.[151,250,1633,1656]" captionTargetBox="[193,1397,215,1595]" captionTargetId="figure@20.[171,1416,193,1612]" captionTargetPageId="20" captionText="FIGURE 11. Macrophthalmus graeffei A. Milne-Edwards 1873, male (ZMUC CRU- 1782): a, Posterior surface of carapace, left side; b, merus of male cheliped, inner surface; c, cheliped of male, outer surface; d, cheliped of male, inner surface; e, male abdomen." httpUri="https://zenodo.org/record/203109/files/figure.png" pageId="20" pageNumber="21">Figs. 11</figureCitation>
e, 17b) with segments 5, 6 of same length; lateral margin of segment 6 proximally remarkably swollen, distally very gently converging; telson semicircular, about as long as segment 6.
</paragraph>
<paragraph id="D06436D8552DFFE4FF254B34FC1E7D8C" blockId="20.[151,1436,1756,2033]" pageId="20" pageNumber="21">
Male G1 (
<figureCitation id="48E02A5D552DFFE4FEDE4B34FE7C7D34" box="[318,412,1828,1852]" captionStart="FIGURE 12" captionStartId="21.[151,250,1324,1347]" captionTargetBox="[182,1413,401,1283]" captionTargetId="figure@21.[151,1436,375,1303]" captionTargetPageId="21" captionText="FIGURE 12. Macrophthalmus graeffei A. Milne-Edwards 1873, male (ZMUC CRU- 1782): a, G 1 of male, ventral surface (right); b, apical part of G 1, ventral surface; c, apical part of G 1, dorsal surface; d, apical part of G 1, lateral surface." httpUri="https://zenodo.org/record/203110/files/figure.png" pageId="20" pageNumber="21">Figs. 12</figureCitation>
ad) relatively long, narrow, slightly curving outward medially; apical chitinous process short, narrow, directed outwards at 90°; distal opening (
<figureCitation id="48E02A5D552DFFE4FCEA4B58FCBB7D68" box="[778,859,1864,1888]" captionStart="FIGURE 12" captionStartId="21.[151,250,1324,1347]" captionTargetBox="[182,1413,401,1283]" captionTargetId="figure@21.[151,1436,375,1303]" captionTargetPageId="21" captionText="FIGURE 12. Macrophthalmus graeffei A. Milne-Edwards 1873, male (ZMUC CRU- 1782): a, G 1 of male, ventral surface (right); b, apical part of G 1, ventral surface; c, apical part of G 1, dorsal surface; d, apical part of G 1, lateral surface." httpUri="https://zenodo.org/record/203110/files/figure.png" pageId="20" pageNumber="21">Fig. 12</figureCitation>
d) on tip of chitinous process; dorsal palp remarkably large; long setae scarcely on apical portion, not concealing apical process.
</paragraph>
</subSubSection>
<subSubSection id="98C16553552DFFE5FF254B80FE1E7B6C" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="discussion">
<paragraph id="D06436D8552DFFE5FF254B80FE1E7B6C" blockId="20.[151,1436,1756,2033]" lastBlockId="21.[151,1436,151,356]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
<emphasis id="E2AFEACA552DFFE4FF254B80FED97DA0" bold="true" box="[197,313,1936,1960]" pageId="20" pageNumber="21">Remarks.</emphasis>
<bibRefCitation id="B44A4B29552DFFE4FEA04B80FDF07DA0" author="Stephensen" box="[320,528,1936,1960]" pageId="20" pageNumber="21" refString="Stephensen, K. (1945) The Brachyura of the Iranian Gulf. Danish Scientific Investigations in Iran, Part IV. E. Munksgaard, Copenhagen, 57 - 237." type="book chapter" year="1945">Stephensen (1945)</bibRefCitation>
recorded one male and one juvenile from the east coast of Kharg I. of
<collectingCountry id="A8CC7648552DFFE4FAC94B81FAB67DA0" box="[1321,1366,1937,1960]" name="Iran" pageId="20" pageNumber="21">Iran</collectingCountry>
in the Persian Gulf which he tentatively identified as
<emphasis id="E2AFEACA552DFFE4FD414BA5FD5F7DC4" box="[673,703,1973,1996]" italics="true" pageId="20" pageNumber="21">M.</emphasis>
(
<taxonomicName id="17DB4D5B552DFFE4FD2F4BA5FC4C7DC4" authority="Stimpson" authorityName="Stimpson" box="[719,940,1972,1996]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="convexus">
<emphasis id="E2AFEACA552DFFE4FD2F4BA5FCD67DC4" box="[719,822,1973,1996]" italics="true" pageId="20" pageNumber="21">convexus</emphasis>
Stimpson
</taxonomicName>
?).
<bibRefCitation id="B44A4B29552DFFE4FC284BA5FB877DC4" author="Barnes" box="[968,1127,1972,1996]" pageId="20" pageNumber="21" refString="Barnes, R. S. K. (1971) Biological results of the Snellius Expedition. XXIII. The Genus Macrophthalmus (Decapoda, Brachyura). Zoologische Verhandelingen, 115, 1 - 40." type="journal article" year="1971">Barnes (1971)</bibRefCitation>
cited this record in the synonymy of
<taxonomicName id="17DB4D5B552DFFE4FEEA4BC9FE667DF8" box="[266,390,2008,2032]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="graeffei">
<emphasis id="E2AFEACA552DFFE4FEEA4BC9FE667DF8" box="[266,390,2008,2032]" italics="true" pageId="20" pageNumber="21">M. graeffei</emphasis>
</taxonomicName>
.
<bibRefCitation id="B44A4B29552DFFE4FE744BC8FDFC7DF8" author="Apel" box="[404,540,2008,2032]" pageId="20" pageNumber="21" refString="Apel, M. (2001) Taxonomie und Zoogeographie der Brachyura, Paguridea und Porcellanidae (Crustacea: Decapoda) des Persisch-Arabischen Golfes unpublished Ph. D. Thesis, Johann Wolfgang Goethe-Universitat, Frankfurt am Main 268 pp." type="book" year="2001">Apel (2001)</bibRefCitation>
examined this specimen and found that it is identical with the
<taxonomicName id="17DB4D5B552DFFE5FB3A4BC9FE767AA7" authority="Barnes. Barnes (1971)" authorityName="Barnes. Barnes" authorityYear="1971" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" lastPageId="21" lastPageNumber="22" order="Decapoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="graeffei">
<emphasis id="E2AFEACA552DFFE4FB3A4BC9FB117DF8" box="[1242,1265,2009,2032]" italics="true" pageId="20" pageNumber="21">M</emphasis>
.
<emphasis id="E2AFEACA552DFFE4FB1F4BC8FAB67DF8" box="[1279,1366,2008,2032]" italics="true" pageId="20" pageNumber="21">graeffei</emphasis>
sensu
<bibRefCitation id="B44A4B29552CFFE5FF774C88FE767AA7" box="[151,406,151,175]" pageId="21" pageNumber="22" refString="Barnes, R. S. K. (1971) Biological results of the Snellius Expedition. XXIII. The Genus Macrophthalmus (Decapoda, Brachyura). Zoologische Verhandelingen, 115, 1 - 40." type="journal article">Barnes. Barnes (1971)</bibRefCitation>
</taxonomicName>
, however, did not examine the
<typeStatus id="0F60887A552CFFE5FD164C88FCC67AB8" box="[758,806,152,176]" pageId="21" pageNumber="22">type</typeStatus>
specimens of
<taxonomicName id="17DB4D5B552CFFE5FC294C89FBA67AB8" box="[969,1094,152,176]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="graeffei">
<emphasis id="E2AFEACA552CFFE5FC294C89FC007AB8" box="[969,992,153,176]" italics="true" pageId="21" pageNumber="22">M</emphasis>
.
<emphasis id="E2AFEACA552CFFE5FC0F4C88FBA67AB8" box="[1007,1094,152,176]" italics="true" pageId="21" pageNumber="22">graeffei</emphasis>
</taxonomicName>
, which had been described by
<bibRefCitation id="B44A4B29552CFFE5FF774CACFE597ADC" author="Milne-Edwards" box="[151,441,188,212]" pageId="21" pageNumber="22" refString="Milne-Edwards, A. (1873) Description de quelques Crustaces nouveaux ou peu connus provenant du Musee de M. C. Godeffroy. Journal des Museum Godeffroy, 1 (4), 77 - 88 (253 - 264), pls. 12 - 13." type="journal article" year="1873">A. Milne-Edwards (1873)</bibRefCitation>
from
<collectingCountry id="A8CC7648552CFFE5FE1E4CACFDAE7ADC" box="[510,590,188,212]" name="Samoa" pageId="21" pageNumber="22">Samoa</collectingCountry>
.
<bibRefCitation id="B44A4B29552CFFE5FDB84CADFD187ADC" author="Barnes" box="[600,760,188,212]" pageId="21" pageNumber="22" refString="Barnes, R. S. K. (1971) Biological results of the Snellius Expedition. XXIII. The Genus Macrophthalmus (Decapoda, Brachyura). Zoologische Verhandelingen, 115, 1 - 40." type="journal article" year="1971">Barnes (1971)</bibRefCitation>
record of this species is based on two males collected at the eastern Indian Ocean in South
<collectingCountry id="A8CC7648552CFFE5FE194CCFFD867AFF" box="[505,614,223,247]" name="Indonesia" pageId="21" pageNumber="22">Indonesia</collectingCountry>
(Timor Archipelago), and all following authors used his description for defining
<taxonomicName id="17DB4D5B552CFFE5FF1B4D15FE967B14" box="[251,374,260,284]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="graeffei">
<emphasis id="E2AFEACA552CFFE5FF1B4D15FEF27B14" box="[251,274,261,284]" italics="true" pageId="21" pageNumber="22">M</emphasis>
.
<emphasis id="E2AFEACA552CFFE5FEFF4D14FE967B14" box="[287,374,260,284]" italics="true" pageId="21" pageNumber="22">graeffei</emphasis>
</taxonomicName>
. Therefore, the taxonomic position of
<taxonomicName id="17DB4D5B552CFFE5FCC44D15FC7F7B14" box="[804,927,260,284]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="graeffei">
<emphasis id="E2AFEACA552CFFE5FCC44D15FC7F7B14" box="[804,927,260,284]" italics="true" pageId="21" pageNumber="22">M. graeffei</emphasis>
</taxonomicName>
is still not at all certain, and its identity needs to be established. The present material agrees fully with the description given by
<bibRefCitation id="B44A4B29552CFFE5FBCB4D38FB2E7B37" author="Barnes" box="[1067,1230,295,319]" pageId="21" pageNumber="22" refString="Barnes, R. S. K. (1971) Biological results of the Snellius Expedition. XXIII. The Genus Macrophthalmus (Decapoda, Brachyura). Zoologische Verhandelingen, 115, 1 - 40." type="journal article" year="1971">Barnes (1971)</bibRefCitation>
, and thus it is tentatively assigned to this species.
</paragraph>
</subSubSection>
<caption id="84A46650552CFFE5FF77493CFACA7F69" httpUri="https://zenodo.org/record/203110/files/figure.png" pageId="21" pageNumber="22" targetBox="[182,1413,401,1283]" targetPageId="21">
<paragraph id="D06436D8552CFFE5FF77493CFACA7F69" blockId="21.[151,1436,1324,1377]" pageId="21" pageNumber="22">
<emphasis id="E2AFEACA552CFFE5FF77493CFEC27F49" bold="true" box="[151,290,1324,1347]" pageId="21" pageNumber="22">FIGURE 12.</emphasis>
<taxonomicName id="17DB4D5B552CFFE5FECC493CFCD77F4A" authority="A. Milne-Edwards 1873" authorityName="A. Milne-Edwards" authorityYear="1873" box="[300,823,1324,1346]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="graeffei">
<emphasis id="E2AFEACA552CFFE5FECC493CFDD07F4A" box="[300,560,1324,1346]" italics="true" pageId="21" pageNumber="22">Macrophthalmus graeffei</emphasis>
A. Milne-Edwards 1873
</taxonomicName>
, male (ZMUC CRU-1782): a, G1 of male, ventral surface (right); b, apical part of G1, ventral surface; c, apical part of G1, dorsal surface; d, apical part of G1, lateral surface.
</paragraph>
</caption>
<subSubSection id="98C16553552CFFE5FF25499EFEDF7D3A" pageId="21" pageNumber="22" type="reference_group">
<paragraph id="D06436D8552CFFE5FF25499EFC2A7C52" blockId="21.[151,1436,1422,2022]" pageId="21" pageNumber="22">
<taxonomicName id="17DB4D5B552CFFE5FF25499EFE017FAE" ID-CoL="3X864" box="[197,481,1422,1446]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="graeffei">
<emphasis id="E2AFEACA552CFFE5FF25499EFE017FAE" box="[197,481,1422,1446]" italics="true" pageId="21" pageNumber="22">Macrophthalmus graeffei</emphasis>
</taxonomicName>
belongs to the subgenus
<taxonomicName id="17DB4D5B552CFFE5FD1D499EFB617FAE" ID-CoL="7PDBN" authority="Desmarest, 1823" authorityName="Desmarest" authorityYear="1823" box="[765,1153,1422,1446]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="E2AFEACA552CFFE5FD1D499EFC5B7FAE" box="[765,955,1422,1446]" italics="true" pageId="21" pageNumber="22">Macrophthalmus</emphasis>
Desmarest, 1823
</taxonomicName>
, which recently has been divided into four different subgroups (
<bibRefCitation id="B44A4B29552CFFE5FDB749A2FD0E7FC2" author="Barnes" box="[599,750,1458,1482]" pageId="21" pageNumber="22" refString="Barnes, R. S. K. (2010) A review of the sentinel and allied crabs (Crustacea: Brachyura: Macrophthalmidae), with particular references to the genus Macrophthalmus. Raffles Bulletin of Zoology, 58 (1), 31 - 49." type="journal article" year="2010">Barnes 2010</bibRefCitation>
).
<bibRefCitation id="B44A4B29552CFFE5FCE449A3FC487FC2" author="Barnes" box="[772,936,1458,1482]" pageId="21" pageNumber="22" refString="Barnes, R. S. K. (2010) A review of the sentinel and allied crabs (Crustacea: Brachyura: Macrophthalmidae), with particular references to the genus Macrophthalmus. Raffles Bulletin of Zoology, 58 (1), 31 - 49." type="journal article" year="2010">Barnes (2010)</bibRefCitation>
placed
<taxonomicName id="17DB4D5B552CFFE5FBE749A3FB677FC2" ID-CoL="3X864" box="[1031,1159,1458,1482]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="graeffei">
<emphasis id="E2AFEACA552CFFE5FBE749A3FBFE7FC2" box="[1031,1054,1459,1482]" italics="true" pageId="21" pageNumber="22">M</emphasis>
.
<emphasis id="E2AFEACA552CFFE5FBD049A2FB677FC2" box="[1072,1159,1458,1482]" italics="true" pageId="21" pageNumber="22">graeffei</emphasis>
</taxonomicName>
in the
<taxonomicName id="17DB4D5B552CFFE5FB0049A3FA737FC2" ID-CoL="3X86M" box="[1248,1427,1458,1482]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="telescopicus">
<emphasis id="E2AFEACA552CFFE5FB0049A3FA737FC2" box="[1248,1427,1458,1482]" italics="true" pageId="21" pageNumber="22">M. telescopicus</emphasis>
</taxonomicName>
- group, which is mainly characterized by long eyestalks with the cornea extended beyond the lateral carapace margin, the carapace being moderately broad with three lateral teeth, the front being relatively broad, the carpus of the male cheliped lacking spines on the distal margin, the fingers being short and the immovable finger not being deflexed. Most species of this group including
<taxonomicName id="17DB4D5B552CFFE5FD444A53FCC17C52" ID-CoL="3X864" box="[676,801,1602,1626]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="graeffei">
<emphasis id="E2AFEACA552CFFE5FD444A53FD5B7C52" box="[676,699,1603,1626]" italics="true" pageId="21" pageNumber="22">M</emphasis>
.
<emphasis id="E2AFEACA552CFFE5FD2A4A52FCC17C52" box="[714,801,1602,1626]" italics="true" pageId="21" pageNumber="22">graeffei</emphasis>
</taxonomicName>
are sublittoral.
</paragraph>
<paragraph id="D06436D8552CFFE5FF254A76FEDF7D3A" blockId="21.[151,1436,1422,2022]" pageId="21" pageNumber="22">
The only record of this species from the Persian Gulf is the two males (one juvenile) from Kharg I. collected by the Danish Scientific expedition (
<bibRefCitation id="B44A4B29552CFFE5FDF24A9AFD377CAA" author="Stephensen" box="[530,727,1674,1698]" pageId="21" pageNumber="22" refString="Stephensen, K. (1945) The Brachyura of the Iranian Gulf. Danish Scientific Investigations in Iran, Part IV. E. Munksgaard, Copenhagen, 57 - 237." type="book chapter" year="1945">Stephensen 1945</bibRefCitation>
).
<bibRefCitation id="B44A4B29552CFFE5FD0C4A9AFC957CAA" author="Apel" box="[748,885,1674,1698]" pageId="21" pageNumber="22" refString="Apel, M. (2001) Taxonomie und Zoogeographie der Brachyura, Paguridea und Porcellanidae (Crustacea: Decapoda) des Persisch-Arabischen Golfes unpublished Ph. D. Thesis, Johann Wolfgang Goethe-Universitat, Frankfurt am Main 268 pp." type="book" year="2001">Apel (2001)</bibRefCitation>
furthermore recorded one specimen from
<collectingCountry id="A8CC7648552CFFE5FAB74A9AFA7C7CAA" box="[1367,1436,1674,1698]" name="Oman" pageId="21" pageNumber="22">Oman</collectingCountry>
(“Djaraman, Mer dOman, Côte dArabie”) deposited in MNHN Paris. The
<typeStatus id="0F60887A552CFFE5FC154ABFFBC57CCF" box="[1013,1061,1711,1735]" pageId="21" pageNumber="22">type</typeStatus>
material of
<taxonomicName id="17DB4D5B552CFFE5FB514ABFFACE7CCE" ID-CoL="3X864" box="[1201,1326,1710,1734]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="graeffei">
<emphasis id="E2AFEACA552CFFE5FB514ABFFB287CCE" box="[1201,1224,1711,1734]" italics="true" pageId="21" pageNumber="22">M</emphasis>
.
<emphasis id="E2AFEACA552CFFE5FB374ABEFACE7CCE" box="[1239,1326,1710,1734]" italics="true" pageId="21" pageNumber="22">graeffei</emphasis>
</taxonomicName>
had been deposited in the Godeffroy Museum of Hamburg (
<date id="A4651018552CFFE5FD314AC2FCB47CE2" box="[721,852,1746,1770]" pageId="21" pageNumber="22" value="1861" valueMax="1885">18611885</date>
),
<collectingCountry id="A8CC7648552CFFE5FC884AC2FC327CE2" box="[872,978,1746,1770]" name="Germany" pageId="21" pageNumber="22">Germany</collectingCountry>
. In 1885, however, the zoological specimens of this museum were sold to different museums, and no information is available regarding the
<typeStatus id="0F60887A552CFFE5FAE84AE7FAD87D07" box="[1288,1336,1783,1807]" pageId="21" pageNumber="22">type</typeStatus>
material of that species.
</paragraph>
</subSubSection>
<subSubSection id="98C16553552CFFE6FF254B2DFBE07ADC" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="biology_ecology">
<paragraph id="D06436D8552CFFE5FF254B2DFE317DEE" blockId="21.[151,1436,1422,2022]" pageId="21" pageNumber="22">
<emphasis id="E2AFEACA552CFFE5FF254B2DFEC57D5E" bold="true" box="[197,293,1853,1878]" pageId="21" pageNumber="22">Biology.</emphasis>
<taxonomicName id="17DB4D5B552CFFE5FECC4B2EFDA87D5E" box="[300,584,1854,1878]" class="Malacostraca" family="Macrophthalmidae" genus="Macrophthalmus" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="graeffei">
<emphasis id="E2AFEACA552CFFE5FECC4B2EFDA87D5E" box="[300,584,1854,1878]" italics="true" pageId="21" pageNumber="22">Macrophthalmus graeffei</emphasis>
</taxonomicName>
is a relatively small-size species. The male specimen from the Persian Gulf is the largest specimen (CL = 14.8, CB =
<quantity id="17239B3D552CFFE5FD8A4B72FD217D72" box="[618,705,1890,1914]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="21" pageNumber="22" unit="mm" value="8.4">8.4 mm</quantity>
) that has so far been collected. The species is one of the rarest of the genus, even though the lack of material could be particularly due to its subtidal habitat (
<bibRefCitation id="B44A4B29552CFFE5FB444B96FA837D96" author="Lewinsohn" box="[1188,1379,1926,1950]" pageId="21" pageNumber="22" refString="Lewinsohn, C. (1977) Die Ocypodidae des Roten Meeres (Crustacea, Decapoda, Brachyura). Zoologische Verhandelingen, 152, 45 - 84." type="journal article" year="1977">Lewinsohn 1977</bibRefCitation>
) and the lack of adequate collections carried out. The specimen from Kharg I. was collected from a sandy bottom in
<quantity id="17239B3D552CFFE5FA604BBAFF4D7DEE" metricMagnitude="1" metricUnit="m" metricValue="1.8" pageId="21" pageNumber="22" unit="m" value="18.0">18 m</quantity>
depth (
<bibRefCitation id="B44A4B29552CFFE5FEE04BDEFE247DEE" author="Stephensen" box="[256,452,1998,2022]" pageId="21" pageNumber="22" refString="Stephensen, K. (1945) The Brachyura of the Iranian Gulf. Danish Scientific Investigations in Iran, Part IV. E. Munksgaard, Copenhagen, 57 - 237." type="book chapter" year="1945">Stephensen 1945</bibRefCitation>
).
</paragraph>
<paragraph id="D06436D8552FFFE6FF254C87FBE07ADC" blockId="22.[151,1437,151,212]" pageId="22" pageNumber="23">
<emphasis id="E2AFEACA552FFFE6FF254C87FDE37AB8" bold="true" box="[197,515,151,176]" pageId="22" pageNumber="23">Geographical distribution.</emphasis>
Wide Indo-West Pacific distribution: Red Sea, Gulf of
<collectingCountry id="A8CC7648552FFFE6FB914C87FB597AA7" box="[1137,1209,151,175]" name="Oman" pageId="22" pageNumber="23">Oman</collectingCountry>
, Persian Gulf,
<collectingCountry id="A8CC7648552FFFE6FA804C87FF317ADC" name="Indonesia" pageId="22" pageNumber="23">Indonesia</collectingCountry>
(Timor),
<collectingCountry id="A8CC7648552FFFE6FEDD4CACFE6D7ADC" box="[317,397,188,212]" name="Samoa" pageId="22" pageNumber="23">Samoa</collectingCountry>
; migrated to the Mediterranean through the Suez Canal.
</paragraph>
</subSubSection>
</treatment>
</document>