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<mods:title id="A87389E25DFBCE19215C9CA033A422D3">A new species of Stygobromus Cope, 1872 (Amphipoda, Crangonyctidae) from a hypotelminorheic seepage spring in Washington, D. C., USA</mods:title>
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<mods:namePart id="61107861166268253BEDDEE1EEC4498E">Niemiller, Matthew L.</mods:namePart>
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<mods:affiliation id="297E9834CAF95F7FE50391C74C8A2B9A">Department of Biological Sciences, The University of Alabama in Huntsville, Huntsville, AL 35899, USA</mods:affiliation>
<mods:nameIdentifier id="831175E68F48031775AD4DA800108C9D" type="email">cavemander17@gmail.com</mods:nameIdentifier>
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<mods:namePart id="BE73E04C1CEB66936FBB29C63AF6B07A">Cannizzaro, Andrew G.</mods:namePart>
<mods:affiliation id="840F0290A917AA0CABE72E932E4998FD">Department of Biology, Miami University, Oxford, OH 45056, USA</mods:affiliation>
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<mods:namePart id="B9140BC6481D2576290F5DD132781332">Sawicki, Thomas R.</mods:namePart>
<mods:affiliation id="D83840278EFD2852BD84CDC2C84A7482">Department of Biological Sciences, Florida Agricultural and Mechanical University, Tallahassee, FL 32307, USA</mods:affiliation>
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<mods:namePart id="A36006F7E3A7DE758D4A98D7F330F07E">Culver, David C.</mods:namePart>
<mods:affiliation id="341F930481D3D7E7687A2471C27CB268">Department of Environmental Science, American University, Washington, DC 20016, USA</mods:affiliation>
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<mods:date id="A676BAEA68032AD904BD63166751E47B">2024</mods:date>
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<subSubSection id="593D46BE792AC805BFBDA7ED31900B59" pageId="0" pageNumber="117" type="nomenclature">
<paragraph id="2A4C06DA3813810B59C6C1D294224F66" pageId="0" pageNumber="117">
<taxonomicName id="FB61CA3470EF4DEA6A2412FC3FBFFD7E" LSID="https://zoobank.org/D66AA3F8-A53B-41A2-B16C-CA9486A39BC1" authority="Cannizzaro, Sawicki, &amp; Niemiller" authorityName="Cannizzaro, Sawicki, &amp; Niemiller" authorityYear="2024" class="Malacostraca" family="Crangonyctidae" genus="Stygobromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stygobromus anacostensis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="anacostensis" status="sp. nov.">Stygobromus anacostensis Cannizzaro, Sawicki, &amp; Niemiller</taxonomicName>
<taxonomicNameLabel id="8FE0F0F6908776AE1FB13875D8784B7A" pageId="0" pageNumber="117">sp. nov.</taxonomicNameLabel>
</paragraph>
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<paragraph id="828C13035CCA8D6E37A70BF45C471F80" pageId="0" pageNumber="117">
<figureCitation id="39617D3EBC30125CC5A20ABC7D0E2C15" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Stygobromus anacostensis sp. nov., habitus: A holotype male, 5.9 mm (USNM 1606902) B Allotype female, 5.3 mm (USNM 1606903). Scale bar: 1 mm." figureDoi="10.3897/subtbiol.48.112984.figure1" httpUri="https://binary.pensoft.net/fig/984740" pageId="0" pageNumber="117">Figs 1</figureCitation>
<figureCitation id="C54B59D2D2AC04702253DDCAB127B61E" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Stygobromus anacostensis sp. nov., Holotype male, 5.9 mm (USNM 1606902): A antenna 1 (single aesthetasc enlarged) C left mandible (palp omitted) D right mandible (lacinia mobilis enlarged). Paratype male, 5.7 mm (USNM 1606904): B antenna 2 (single calceolus enlarged). Scale bars: 0.5 mm (A, B); 0.25 mm (C, D)." figureDoi="10.3897/subtbiol.48.112984.figure2" httpUri="https://binary.pensoft.net/fig/984741" pageId="0" pageNumber="117">, 2</figureCitation>
<figureCitation id="C9DBDDFD6E77BB7CC908E0F9B01FA34B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Stygobromus anacostensis sp. nov., Paratype male, 5.7 mm (USNM 1606904): A upper lip D maxilla 2. Holotype male, 5.9 mm (USNM 1606902): B lower lip C maxilla 1 E maxilliped (distal margin of inner plate enlarged). Scale bars: 0.25 mm." figureDoi="10.3897/subtbiol.48.112984.figure3" httpUri="https://binary.pensoft.net/fig/984742" pageId="0" pageNumber="117">, 3</figureCitation>
<figureCitation id="74596D0A2D7E523CD7A0896AC5FF9372" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Stygobromus anacostensis sp. nov., Holotype male, 5.9 mm (USNM 1606902): A gnathopod 1 (palm and dactyl enlarged) B gnathopod 2 (rastellate seta, palm and dactyl enlarged). Scale bar: 0.5 mm." figureDoi="10.3897/subtbiol.48.112984.figure4" httpUri="https://binary.pensoft.net/fig/984743" pageId="0" pageNumber="117">, 4</figureCitation>
<figureCitation id="2F36341DEB194483DBE4FD2567E47DD4" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Stygobromus anacostensis sp. nov., Holotype male, 5.9 mm (USNM 1606902): A pereopod 3 B pereopod 4 C pereopod 5 D pereopod 6 E pereopod 7 F bifurcate sternal gill located on somites 6 and 7. Scale bar: 0.5 mm." figureDoi="10.3897/subtbiol.48.112984.figure5" httpUri="https://binary.pensoft.net/fig/984744" pageId="0" pageNumber="117">, 5</figureCitation>
<figureCitation id="E8DD1189F357E18FC01674954B84596B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Stygobromus anacostensis sp. nov., Holotype male, 5.9 mm (USNM 1606902): A epimera 1 - 3 B pleopod 1 (coupling spines and clothes pin seta enlarged) C uropod 1 (posteromedial protuberance enlarged) D uropod 2 E uropod 3 F telson. Scale bars: 0.25 mm." figureDoi="10.3897/subtbiol.48.112984.figure6" httpUri="https://binary.pensoft.net/fig/984745" pageId="0" pageNumber="117">, 6</figureCitation>
<figureCitation id="5BA8C5746F3B512C14F05FB1C4EF304B" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Stygobromus anacostensis sp. nov., Allotype female, 5.3 mm (USNM 1606903): A antenna 2 B gnathopod 1 (palm and dactyl enlarged) C gnathopod 2 (rastellate seta, palm and dactyl enlarged). Scale bars: 0.25 mm (A); 0.5 mm (B, C)." figureDoi="10.3897/subtbiol.48.112984.figure7" httpUri="https://binary.pensoft.net/fig/984746" pageId="0" pageNumber="117">, 7</figureCitation>
<figureCitation id="FA069FB5F9AAF3FA4A25A6374592431D" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Stygobromus anacostensis sp. nov., Allotype female, 5.3 mm (USNM 1606903): A coxa and basis of pereopod 7 showing placement of bifurcate sternal gill B uropod 1 C uropod 2 D uropod 3 E telson. Scale bars: 0.5 mm (A-C); 0.25 mm (D, E)." figureDoi="10.3897/subtbiol.48.112984.figure8" httpUri="https://binary.pensoft.net/fig/984747" pageId="0" pageNumber="117">, 8</figureCitation>
</paragraph>
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<subSubSection id="C019C001B52CCA89A8D30FBA83A5B8FE" pageId="0" pageNumber="117" type="type material">
<paragraph id="523CC6B76AC19A3B2D75FD7EC576D742" pageId="0" pageNumber="117">Type material.</paragraph>
<paragraph id="7E4085FA326EBA06B14E255A0843E3FE" pageId="0" pageNumber="117">
<emphasis id="6B68DBD9E647CC972D5CEA163E98A53E" bold="true" italics="true" pageId="0" pageNumber="117">Holotype</emphasis>
: male 5.9 mm, from USA, Washington, District of Columbia, Anacostia Park, (
<geoCoordinate id="92970CE31B6171E9954C6C7AB3B35D2E" degrees="38.83059" direction="north" orientation="latitude" precision="1" value="38.83059">38.83059°N</geoCoordinate>
, - 
<geoCoordinate id="59654AA4F6BF3CD961765E002004F2BC" degrees="76.9995" direction="west" orientation="longitude" precision="5" value="-76.9995">76.9995°W</geoCoordinate>
), deposited in the collection of the United States National Museum of Natural History, Smithsonian Institution, Washington, D.C (USNM 1606902); female allotype 5.3 mm (USNM 1606903). Holotype male and female allotype collected 18 October 2021 by Lizzy Sartain.
</paragraph>
<paragraph id="FC5AB063E5071DA654876B1844FF8597" pageId="0" pageNumber="117">
<emphasis id="406293BF48CEF0D46E1A21A0A9A45E9B" bold="true" italics="true" pageId="0" pageNumber="117">Paratypes</emphasis>
: 1 male (USNM 1606904) and 2 females (USNM 1606905-1606906) collected on 18 October 2021 by Lizzy Sartain from type locality; 1 female collected on 20 September 2021 by Lizzy Sartain from the type locality (USNM 1606907).
</paragraph>
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<subSubSection id="61E0C48881A6CCA53391845C7CD1FF12" pageId="0" pageNumber="117" type="etymology">
<paragraph id="3368C192BF27876D8C9AEFF368812D03" pageId="0" pageNumber="117">Etymology.</paragraph>
<paragraph id="513B197D7107C35B0083807716243E80" pageId="0" pageNumber="117">
The specific epithet 
<emphasis id="1CECD4B7F190A8F44C11FEBC93EC844B" italics="true" pageId="0" pageNumber="117">anacostensis</emphasis>
refers to its occurrence in Anacostia neighborhood in Washington, D.C., USA. It is part of the Anacostia River drainage.
</paragraph>
</subSubSection>
<subSubSection id="20C5D35E146F415EBD2E958454A41D74" pageId="0" pageNumber="117" type="type locality">
<paragraph id="BD995151D2F1A1E9C74AFB522C910AF3" pageId="0" pageNumber="117">Type locality.</paragraph>
<paragraph id="FEE00F11C011F3610A6DB903E0983E05" pageId="0" pageNumber="117">
USA. Washington, District of Columbia, hypotelminorheic seepage spring in a highly urbanized area that emerges from a small, 2-m high rockface ca. 5 m from Malcolm X Avenue SE in Shepherd Parkway (Figs 
<figureCitation id="FA70888F607774DE04452DCAD306D183" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Distribution of Stygobromus anacostensis sp. nov. and other S. tenuis group species in the Washington D. C. area, USA" figureDoi="10.3897/subtbiol.48.112984.figure9" httpUri="https://binary.pensoft.net/fig/984748" pageId="0" pageNumber="117">9</figureCitation>
, 
<figureCitation id="C738EC4632B824EB6CED8EA6CD469DC5" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. The type locality of S. anacostensis is a small hypotelminorheic seepage spring just off of Malcolm X Avenue, Shepherd Parkway, Washington, D. C., USA. Photograph by Jenna Keany." figureDoi="10.3897/subtbiol.48.112984.figure10" httpUri="https://binary.pensoft.net/fig/984749" pageId="0" pageNumber="117">10</figureCitation>
, 
<geoCoordinate id="8A6AC4408BC6A8FD2AF2025CDA69696D" degrees="38.83059" direction="north" orientation="latitude" precision="1" value="38.83059">38.83059°N</geoCoordinate>
, - 
<geoCoordinate id="B4A055DD1E87BCCA967ED9E9BC203BAB" degrees="76.9995" direction="west" orientation="longitude" precision="5" value="-76.9995">76.9995°W</geoCoordinate>
). Shepherd Parkway is part of National Capital Parks East. Most individuals were collected in the water flowing over moss-covered rocks. A few individuals were also present in decaying leaves at the base, a more usual hypotelminorheic habitat (
<bibRefCitation id="A429EB79B0791858E6C0792869BFEC5E" author="Culver, D" journalOrPublisher="Subterranean Biology" pageId="0" pageNumber="117" pagination="1 - 7" refId="B10" refString="Culver, D, Pipan, T, Gottstein, S, 2006. Hypotelminorheic-A unique freshwater habitat. Subterranean Biology 4: 1 - 7" title="Hypotelminorheic-A unique freshwater habitat." volume="4" year="2006">Culver et al. 2006</bibRefCitation>
, 
<bibRefCitation id="DE8EF5080D3BA3DBBFD1F9DE5F9A944B" DOI="https://doi.org/10.1656/045.019.m901" author="Culver, DC" journalOrPublisher="Northeastern Naturalist" pageId="0" pageNumber="117" pagination="1 - 42" refId="B11" refString="Culver, DC, Holsinger, JR, Feller, DJ, 2012. The fauna of seepage springs and other shallow subterranean habitats in the mid-Atlantic piedmont and coastal Plain. Northeastern Naturalist 19: 1 - 42, DOI: https://doi.org/10.1656/045.019.m901" title="The fauna of seepage springs and other shallow subterranean habitats in the mid-Atlantic piedmont and coastal Plain." url="https://doi.org/10.1656/045.019.m901" volume="19" year="2012">2012</bibRefCitation>
). The site is at the extreme tip of Shepherd Parkway, a unit of National Capital East (National Park Service). The width of park land is about 20 m and is bordered by Malcolm X Avenue. The site was discovered when a park ranger noticed extensive ice on the adjacent sidewalk resulting from flow from the seep.
</paragraph>
</subSubSection>
<subSubSection id="114B00385D1C9D496055D0BB099B13F1" pageId="0" pageNumber="117" type="diagnosis">
<paragraph id="FEE99E5841F17E8D1A175EE81E3E7434" pageId="0" pageNumber="117">Diagnosis.</paragraph>
<paragraph id="D01814CF4F4E6A9281D9CC05CECEC81F" pageId="0" pageNumber="117">
Small stygomorphic species distinguished from other members of the 
<taxonomicName id="9FBFFC35159E008DDDF3E8B5BD7330C9" baseAuthorityName="S. I. Smith" baseAuthorityYear="1874" class="Malacostraca" family="Crangonyctidae" genus="Stygobromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stygobromus tenuis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis id="8B2C70FC226195A6987F95EF8190FEBC" italics="true" pageId="0" pageNumber="117">Stygobromus tenuis</emphasis>
</taxonomicName>
group by size, largest male 5.9 mm, largest female 5.3 mm and as follows: 
<taxonomicName id="CFC3F2E1F23DE90DA73C78B0973EAED0" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. tenuis subsp. tenuis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="subSpecies" species="tenuis" subSpecies="tenuis">
<emphasis id="FCD5138ACE2EA97C1D2D7D5DF3EA8E31" italics="true" pageId="0" pageNumber="117">S. tenuis tenuis</emphasis>
</taxonomicName>
- by antenna 2 subequal to or shorter than antenna 1; 
<taxonomicName id="4F3FB041EA8F209806D1F4A1EBFC9F17" baseAuthorityName="Holsinger" baseAuthorityYear="1967" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. tenuis subsp. potomacus" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="subSpecies" species="tenuis" subSpecies="potomacus">
<emphasis id="9BB31B196085A68CE37046366ABE6C03" italics="true" pageId="0" pageNumber="117">S. tenuis potomacus</emphasis>
</taxonomicName>
- only 2 C-setae on mandibular palp and up to 8 E-setae; 
<taxonomicName id="C2BBFFD7374DC6E125C4F458A4FBE76C" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. allegheniensis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="allegheniensis">
<emphasis id="C4D9C9410C4078D760F3FB13895D6E6F" italics="true" pageId="0" pageNumber="117">S. allegheniensis</emphasis>
</taxonomicName>
- pereopods 5-7 basis posterior margin weakly convex, and telson tapering distally; 
<taxonomicName id="EEEACF24C2660BDB480AF916AB26DB97" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. hayi" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="hayi">
<emphasis id="D462D25A0D549D52A439D0218CEA4BDA" italics="true" pageId="0" pageNumber="117">S. hayi</emphasis>
</taxonomicName>
- by significantly less spinose uropods 1 and 2, and telson with significantly fewer apical robust setae. Further distinguished from all 
<taxonomicName id="EFBEF8E168A2ADE1A5548F8C7AE3C5E9" baseAuthorityName="S. I. Smith" baseAuthorityYear="1874" class="Malacostraca" family="Crangonyctidae" genus="Stygobromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stygobromus tenuis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis id="1E46A944DD6E24E61931AF4871CC678F" italics="true" pageId="0" pageNumber="117">Stygobromus tenuis</emphasis>
</taxonomicName>
group species by gnathopods 1 and 2 with a serrate blade-like edge running the length of both palms, and by the posterodistal margin of gnathopod 2 carpus possessing rastellate seta(e).
</paragraph>
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<subSubSection id="AEBF7D4848905552B65D13AF061B1BE5" pageId="0" pageNumber="117" type="description">
<paragraph id="149E2BBF8A83021372EB23BBD20938BF" pageId="0" pageNumber="117">Description.</paragraph>
<paragraph id="4638DBE9BDCAFDD6396A292C139B57A0" pageId="0" pageNumber="117">
<emphasis id="F1AE3E356F1054F51C4EEFEDAACD7AA2" bold="true" pageId="0" pageNumber="117">Male</emphasis>
: holotype, USNM 1606902 (Fig. 
<figureCitation id="C1DD101E6FF140AAE253EC9A5ECE375B" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Stygobromus anacostensis sp. nov., habitus: A holotype male, 5.9 mm (USNM 1606902) B Allotype female, 5.3 mm (USNM 1606903). Scale bar: 1 mm." figureDoi="10.3897/subtbiol.48.112984.figure1" httpUri="https://binary.pensoft.net/fig/984740" pageId="0" pageNumber="117">1A</figureCitation>
); Size 5.9 mm.
</paragraph>
<caption id="032E09F5973756720F40EA1446F654F4" doi="10.3897/subtbiol.48.112984.figure1" httpUri="https://binary.pensoft.net/fig/984740" pageId="0" pageNumber="117" start="Figure 1" startId="F1">
<paragraph id="F729CA56C79CD7C7DE9FC0AA1072D428" pageId="0" pageNumber="117">
<emphasis id="CCD4F8FBE84A89CBBDA18E76E097A930" bold="true" pageId="0" pageNumber="117">Figure 1.</emphasis>
<taxonomicName id="467FC247F09EB6300C25FF23F92F7432" authorityName="Cannizzaro, Sawicki, &amp; Niemiller" authorityYear="2024" class="Malacostraca" family="Crangonyctidae" genus="Stygobromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stygobromus anacostensis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="anacostensis">
<emphasis id="B0826BC413CBDFBAEEE096EDACAD13A2" italics="true" pageId="0" pageNumber="117">Stygobromus anacostensis</emphasis>
</taxonomicName>
sp. nov., habitus: 
<emphasis id="86668D582E31A975CD291BA07244F9A6" bold="true" pageId="0" pageNumber="117">A</emphasis>
holotype male, 5.9 mm (USNM 1606902) 
<emphasis id="5B7BF8506A7237066B7CD96FF4B74C2F" bold="true" pageId="0" pageNumber="117">B</emphasis>
Allotype female, 5.3 mm (USNM 1606903). Scale bar: 1 mm.
</paragraph>
</caption>
<paragraph id="E8FB4010DF3EC9414CE36BBB8554CE90" pageId="0" pageNumber="117">
<emphasis id="906B333302674560C9589EF54B1C714E" bold="true" pageId="0" pageNumber="117">
<emphasis id="C11B84F35EDE27597AD0333C9AD35299" italics="true" pageId="0" pageNumber="117">Antennae</emphasis>
.
</emphasis>
Antenna 1 (Fig. 
<figureCitation id="15FCC9698242FE34E84FE40A541E6F03" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Stygobromus anacostensis sp. nov., Holotype male, 5.9 mm (USNM 1606902): A antenna 1 (single aesthetasc enlarged) C left mandible (palp omitted) D right mandible (lacinia mobilis enlarged). Paratype male, 5.7 mm (USNM 1606904): B antenna 2 (single calceolus enlarged). Scale bars: 0.5 mm (A, B); 0.25 mm (C, D)." figureDoi="10.3897/subtbiol.48.112984.figure2" httpUri="https://binary.pensoft.net/fig/984741" pageId="0" pageNumber="117">2A</figureCitation>
): 45% body length, 60% length of antenna 2 (in paratype (USNM 1606904); primary flagellum with 18 segments, aesthetascs on most segments, aesthetascs shorter than respective segments; accessory flagellum 2-segmented, reaching beyond first segment in length.
</paragraph>
<caption id="8029B81A3CE84F1D6B2927568107E3F7" doi="10.3897/subtbiol.48.112984.figure2" httpUri="https://binary.pensoft.net/fig/984741" pageId="0" pageNumber="117" start="Figure 2" startId="F2">
<paragraph id="84F4F387CA8330D0D77815A9F2D592BF" pageId="0" pageNumber="117">
<emphasis id="B80D8A82A80706E2A80F8E53F24325FE" bold="true" pageId="0" pageNumber="117">Figure 2.</emphasis>
<taxonomicName id="CEC88EDA20F67B5B99C1416334ECCC71" authorityName="Cannizzaro, Sawicki, &amp; Niemiller" authorityYear="2024" class="Malacostraca" family="Crangonyctidae" genus="Stygobromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stygobromus anacostensis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="anacostensis">
<emphasis id="832220DC3292CE8072546B62153F446C" italics="true" pageId="0" pageNumber="117">Stygobromus anacostensis</emphasis>
</taxonomicName>
sp. nov., Holotype male, 5.9 mm (USNM 1606902): 
<emphasis id="CC93A361D9513B365EB2F68EA7F76B4C" bold="true" pageId="0" pageNumber="117">A</emphasis>
antenna 1 (single aesthetasc enlarged) 
<emphasis id="745D33BF6EBF45688F14615D308F2C61" bold="true" pageId="0" pageNumber="117">C</emphasis>
left mandible (palp omitted) 
<emphasis id="F2E0189FCA5ED4B74715681A799DC35B" bold="true" pageId="0" pageNumber="117">D</emphasis>
right mandible (lacinia mobilis enlarged). Paratype male, 5.7 mm (USNM 1606904): 
<emphasis id="88A396751CFC281A332A3F44375480F2" bold="true" pageId="0" pageNumber="117">B</emphasis>
antenna 2 (single calceolus enlarged). Scale bars: 0.5 mm (
<emphasis id="F0BA6F365EE676CE2260AF0B345188F5" bold="true" pageId="0" pageNumber="117">A, B</emphasis>
); 0.25 mm (
<emphasis id="0CDA77C0E4EF85DDE61B4B29879D9597" bold="true" pageId="0" pageNumber="117">C, D</emphasis>
).
</paragraph>
</caption>
<paragraph id="D102AC01C22A06748DF8CCA64C2CDFC9" pageId="0" pageNumber="117">
Antenna 2 (Fig. 
<figureCitation id="3874F5854AFCEB5163BF20BFA58BD370" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Stygobromus anacostensis sp. nov., Holotype male, 5.9 mm (USNM 1606902): A antenna 1 (single aesthetasc enlarged) C left mandible (palp omitted) D right mandible (lacinia mobilis enlarged). Paratype male, 5.7 mm (USNM 1606904): B antenna 2 (single calceolus enlarged). Scale bars: 0.5 mm (A, B); 0.25 mm (C, D)." figureDoi="10.3897/subtbiol.48.112984.figure2" httpUri="https://binary.pensoft.net/fig/984741" pageId="0" pageNumber="117">2B</figureCitation>
): damaged in holotype, description based on paratype (USNM 1606904); gland cone distinct; peduncle 80% length of flagellum, with weak plumose setules concentrated on postero and anterodistal margins of segments 4 and 5, peduncle segment 4 subequal in length to 5; flagellum 12-segmented, segment 5 with robust seta on anterodistal margin and segments 6 and 7 with robust seta placed along posterodistal margins, small calceoli-like structures apically on flagellar segments 5-12.
</paragraph>
<paragraph id="C40CF13C4E47BE9EE58A2D74E1BA856E" pageId="0" pageNumber="117">
<emphasis id="40270298A1CF04ECDE74216CD449D8EA" bold="true" italics="true" pageId="0" pageNumber="117">Mouthparts</emphasis>
(Figs 
<figureCitation id="BFE1F0954688A10E94A8D0E6DAB44D50" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Stygobromus anacostensis sp. nov., Holotype male, 5.9 mm (USNM 1606902): A antenna 1 (single aesthetasc enlarged) C left mandible (palp omitted) D right mandible (lacinia mobilis enlarged). Paratype male, 5.7 mm (USNM 1606904): B antenna 2 (single calceolus enlarged). Scale bars: 0.5 mm (A, B); 0.25 mm (C, D)." figureDoi="10.3897/subtbiol.48.112984.figure2" httpUri="https://binary.pensoft.net/fig/984741" pageId="0" pageNumber="117">2C, D</figureCitation>
, 
<figureCitation id="BB4B4F0E5E0BE37B6FEF2B3B9F32B4C5" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Stygobromus anacostensis sp. nov., Paratype male, 5.7 mm (USNM 1606904): A upper lip D maxilla 2. Holotype male, 5.9 mm (USNM 1606902): B lower lip C maxilla 1 E maxilliped (distal margin of inner plate enlarged). Scale bars: 0.25 mm." figureDoi="10.3897/subtbiol.48.112984.figure3" httpUri="https://binary.pensoft.net/fig/984742" pageId="0" pageNumber="117">3</figureCitation>
). Mandibles: left mandible (Fig. 
<figureCitation id="D76E618A8A368FFE1F4FC2D9858DFD92" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Stygobromus anacostensis sp. nov., Holotype male, 5.9 mm (USNM 1606902): A antenna 1 (single aesthetasc enlarged) C left mandible (palp omitted) D right mandible (lacinia mobilis enlarged). Paratype male, 5.7 mm (USNM 1606904): B antenna 2 (single calceolus enlarged). Scale bars: 0.5 mm (A, B); 0.25 mm (C, D)." figureDoi="10.3897/subtbiol.48.112984.figure2" httpUri="https://binary.pensoft.net/fig/984741" pageId="0" pageNumber="117">2C</figureCitation>
) incisor 5-dentate, lacinia mobilis 5-dentate, with 7 robust serrate and numerous plumose accessory setae; molar process reduced with simple seta; palp with 3 segments, second segment 85% length of third, with inner margin bearing 8 setae and sparse fine setae; segment 3 with 2 C-setae, 5 E-setae, 1 B-seta, and 8 plumose D-setae, lacking A-setae; face of article covered in numerous, fine pubescent setae.
</paragraph>
<paragraph id="F66165F06F5786E355B6A3C678259740" pageId="0" pageNumber="117">
Right Mandible (Fig. 
<figureCitation id="EAB63E7E218BC6DE7A2E86A45028F73C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Stygobromus anacostensis sp. nov., Holotype male, 5.9 mm (USNM 1606902): A antenna 1 (single aesthetasc enlarged) C left mandible (palp omitted) D right mandible (lacinia mobilis enlarged). Paratype male, 5.7 mm (USNM 1606904): B antenna 2 (single calceolus enlarged). Scale bars: 0.5 mm (A, B); 0.25 mm (C, D)." figureDoi="10.3897/subtbiol.48.112984.figure2" httpUri="https://binary.pensoft.net/fig/984741" pageId="0" pageNumber="117">2D</figureCitation>
): incisor 4-dentate, lacinia mobilis bifurcate, both lobes with numerous protuberances; accessory setae row with 4 robust, serrate setae and numerous plumose setae; molar process reduced with simple seta. Palp with 3 articles, relative articles length and setation patterns as in left mandible.
</paragraph>
<paragraph id="FEC3E61BEA786F68EE459965EC806D52" pageId="0" pageNumber="117">
Upper Lip (Fig. 
<figureCitation id="36A18FBE33D96A9907173E2DDCA096AA" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Stygobromus anacostensis sp. nov., Paratype male, 5.7 mm (USNM 1606904): A upper lip D maxilla 2. Holotype male, 5.9 mm (USNM 1606902): B lower lip C maxilla 1 E maxilliped (distal margin of inner plate enlarged). Scale bars: 0.25 mm." figureDoi="10.3897/subtbiol.48.112984.figure3" httpUri="https://binary.pensoft.net/fig/984742" pageId="0" pageNumber="117">3A</figureCitation>
): rounded, apical margin of labrum with numerous fine setae. Lower Lip (Fig. 
<figureCitation id="234A9263A60CF04935975D5411A895CB" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Stygobromus anacostensis sp. nov., Paratype male, 5.7 mm (USNM 1606904): A upper lip D maxilla 2. Holotype male, 5.9 mm (USNM 1606902): B lower lip C maxilla 1 E maxilliped (distal margin of inner plate enlarged). Scale bars: 0.25 mm." figureDoi="10.3897/subtbiol.48.112984.figure3" httpUri="https://binary.pensoft.net/fig/984742" pageId="0" pageNumber="117">3B</figureCitation>
): inner lobes distinct; outer margin of outer lobe sparsely covered in fine setae; inner margin of outer lobe heavily setose.
</paragraph>
<caption id="468F5FC2E09AE843109E9C4524628B02" doi="10.3897/subtbiol.48.112984.figure3" httpUri="https://binary.pensoft.net/fig/984742" pageId="0" pageNumber="117" start="Figure 3" startId="F3">
<paragraph id="ABA6540D6256A7BE0D03CCC72494BD67" pageId="0" pageNumber="117">
<emphasis id="BEF7B399609C7C507410CB298B64A5C1" bold="true" pageId="0" pageNumber="117">Figure 3.</emphasis>
<taxonomicName id="23502BD5F775D88DBF66AD7E3AED089A" authorityName="Cannizzaro, Sawicki, &amp; Niemiller" authorityYear="2024" class="Malacostraca" family="Crangonyctidae" genus="Stygobromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stygobromus anacostensis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="anacostensis">
<emphasis id="C1CD1D200F1F563C441809C4EF3214DF" italics="true" pageId="0" pageNumber="117">Stygobromus anacostensis</emphasis>
</taxonomicName>
sp. nov., Paratype male, 5.7 mm (USNM 1606904): 
<emphasis id="21688C0EC3B23BCA6CF8D648EC418C67" bold="true" pageId="0" pageNumber="117">A</emphasis>
upper lip 
<emphasis id="503803127C7E3172EB3F22D2BA5CA441" bold="true" pageId="0" pageNumber="117">D</emphasis>
maxilla 2. Holotype male, 5.9 mm (USNM 1606902): 
<emphasis id="3B2CA7A52B64FE58B7A01B4349DE9035" bold="true" pageId="0" pageNumber="117">B</emphasis>
lower lip 
<emphasis id="705D312602F05603D8B13EFE088E7E94" bold="true" pageId="0" pageNumber="117">C</emphasis>
maxilla 1 
<emphasis id="FF6FB723AFB9ED5EB8A1CA8EF52976FD" bold="true" pageId="0" pageNumber="117">E</emphasis>
maxilliped (distal margin of inner plate enlarged). Scale bars: 0.25 mm.
</paragraph>
</caption>
<paragraph id="E7D48B7C4FE5C446F9484C1F6F33716F" pageId="0" pageNumber="117">
Maxilla 1 (Fig. 
<figureCitation id="E3911F339F9480771C99400CDD8D5F20" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Stygobromus anacostensis sp. nov., Paratype male, 5.7 mm (USNM 1606904): A upper lip D maxilla 2. Holotype male, 5.9 mm (USNM 1606902): B lower lip C maxilla 1 E maxilliped (distal margin of inner plate enlarged). Scale bars: 0.25 mm." figureDoi="10.3897/subtbiol.48.112984.figure3" httpUri="https://binary.pensoft.net/fig/984742" pageId="0" pageNumber="117">3C</figureCitation>
): missing in holotype, description based on paratype (USNM 1606904); inner plate with 4 plumose marginal setae and numerous fine, pubescent setae covering entire plate; outer plate with 7 apical comb spines, pubescence covering inner margin, decreasing laterally and proximally; palp with 2 segments, distal segment covered in pubescence; subapical margin of distal article with 3 long setae, apical margin with 4 setae.
</paragraph>
<paragraph id="82D43DFF1D1D148CB7D2692FCE17207A" pageId="0" pageNumber="117">
Maxilla 2 (Fig. 
<figureCitation id="12C895FE7887D8F1920F95FA84C7C2B6" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Stygobromus anacostensis sp. nov., Paratype male, 5.7 mm (USNM 1606904): A upper lip D maxilla 2. Holotype male, 5.9 mm (USNM 1606902): B lower lip C maxilla 1 E maxilliped (distal margin of inner plate enlarged). Scale bars: 0.25 mm." figureDoi="10.3897/subtbiol.48.112984.figure3" httpUri="https://binary.pensoft.net/fig/984742" pageId="0" pageNumber="117">3D</figureCitation>
): missing in holotype, description based on paratype (USNM 1606904); both inner and outer plates covered in pubescent setae; outer plate not as wide as inner plate, not narrowing distally, with numerous distal setae; inner plate narrowing slightly distally, with numerous apical setae and 3 large plumose facial setae.
</paragraph>
<paragraph id="D8821224F9CC4FFFC12AE8908CE366EC" pageId="0" pageNumber="117">
Maxilliped (Fig. 
<figureCitation id="BEF70738A9B0099F91F88555C35FE392" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Stygobromus anacostensis sp. nov., Paratype male, 5.7 mm (USNM 1606904): A upper lip D maxilla 2. Holotype male, 5.9 mm (USNM 1606902): B lower lip C maxilla 1 E maxilliped (distal margin of inner plate enlarged). Scale bars: 0.25 mm." figureDoi="10.3897/subtbiol.48.112984.figure3" httpUri="https://binary.pensoft.net/fig/984742" pageId="0" pageNumber="117">3E</figureCitation>
): inner plate shorter than outer plate, with 4 naked cuspidate setae 3 setae along apical margin, surface of plate covered in fine pubescence; outer plate armed with numerous setae covering inner and apical margins; palp second segment with numerous marginal setae, third article with numerous marginal/submarginal setae; dactyl with 2 outer setae and 2 inner setae.
</paragraph>
<paragraph id="B53F9483F1437E1FCB5BC0AA7D7C88C7" pageId="0" pageNumber="117">
<emphasis id="A0B3E2871466432CA5B081E4E32D6EE9" bold="true" pageId="0" pageNumber="117">
<emphasis id="F4971B4E63D6B11BA2A0969750E75E3E" italics="true" pageId="0" pageNumber="117">Gnathopods</emphasis>
.
</emphasis>
Gnathopod 1 (Fig. 
<figureCitation id="B763114243A0E9D54B491ED098D7ADBF" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Stygobromus anacostensis sp. nov., Holotype male, 5.9 mm (USNM 1606902): A gnathopod 1 (palm and dactyl enlarged) B gnathopod 2 (rastellate seta, palm and dactyl enlarged). Scale bar: 0.5 mm." figureDoi="10.3897/subtbiol.48.112984.figure4" httpUri="https://binary.pensoft.net/fig/984743" pageId="0" pageNumber="117">4A</figureCitation>
): coxal plate with 3 apical setae; basis with long setae inserted along anterior, and posterior margins, small patch of pubescence on posterodistal corner; ischium with 4 setae and pubescence along posterior margin; merus weakly pubescent along posterior surface, numerous distal setae, and robust seta along anterior margin; carpus approximately 50% length of propodus with robust seta along anterior margin and a group of setae on anterodistal margin, one of which is approximately 50% length of propodus, posterior margin with single group of plumose setae and 6 submarginal setae directed distally; propodus 1.3 
<normalizedToken id="F11EE4D195843B2F79BF6BE5969B1BB0" originalValue="×">x</normalizedToken>
longer than broad, with 1 marginal anterior seta, 4 superior medial setae, with middle group of medial setae paired, 4 setae inserted at anterodistal corner, 6 inferior medial setae and numerous plumose posterior setae; palm oblique, concave, with serrate blade-like edge running the length, 7 outer and 7 inner bifid robust setae, 6 outer setae, and inner seta; inner margin of defining angle with 3 bifid robust setae, outer margin with 4 bifid robust setae; dactylus with outer seta and 7 short setae covering the entire inner margin and 3 setae placed along the inner margin at base of nail.
</paragraph>
<caption id="9A89107E9325D99BD94808CEA55BD6FA" doi="10.3897/subtbiol.48.112984.figure4" httpUri="https://binary.pensoft.net/fig/984743" pageId="0" pageNumber="117" start="Figure 4" startId="F4">
<paragraph id="FBDA8A902B063634BD4E0FE447AA6E54" pageId="0" pageNumber="117">
<emphasis id="4656F34A61F8CF05D79B586706BA2DB2" bold="true" pageId="0" pageNumber="117">Figure 4.</emphasis>
<taxonomicName id="B4336A5E6B2418C85852092DB9E5D6A7" authorityName="Cannizzaro, Sawicki, &amp; Niemiller" authorityYear="2024" class="Malacostraca" family="Crangonyctidae" genus="Stygobromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stygobromus anacostensis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="anacostensis">
<emphasis id="ED33EC6EBF7F5DF002BD22DD289B5830" italics="true" pageId="0" pageNumber="117">Stygobromus anacostensis</emphasis>
</taxonomicName>
sp. nov., Holotype male, 5.9 mm (USNM 1606902): 
<emphasis id="12636C7CC9A89D46868CC93C3BA88609" bold="true" pageId="0" pageNumber="117">A</emphasis>
gnathopod 1 (palm and dactyl enlarged) 
<emphasis id="66ADA2D9B9CA934A9F58E2B4C33ED37D" bold="true" pageId="0" pageNumber="117">B</emphasis>
gnathopod 2 (rastellate seta, palm and dactyl enlarged). Scale bar: 0.5 mm.
</paragraph>
</caption>
<paragraph id="656DA1B60293C588D44FD5DEEEC9904D" pageId="0" pageNumber="117">
Gnathopod 2 (Fig. 
<figureCitation id="62AB415569EE112FE2B610DB7A61555E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Stygobromus anacostensis sp. nov., Holotype male, 5.9 mm (USNM 1606902): A gnathopod 1 (palm and dactyl enlarged) B gnathopod 2 (rastellate seta, palm and dactyl enlarged). Scale bar: 0.5 mm." figureDoi="10.3897/subtbiol.48.112984.figure4" httpUri="https://binary.pensoft.net/fig/984743" pageId="0" pageNumber="117">4B</figureCitation>
): coxal plate with 4 apical setae and facial seta; basis with long setae inserted along anterior, and posterior margins, small patch of pubescence on posterodistal corner; ischium with 3 setae and pubescence along posterior margin; merus with pubescence covering posterior surface and 4 posterodistal setae and robust seta along anterior margin; carpus approximately 75% length of propodus, with robust seta along anterior margin and two setae on anterodistal margin, one of which is approximately 33% length of propodus, posterior margin with 4 groups of plumose setae, distal-most bearing 3 rastellate setae, and 3 submarginal setae directed distally; propodus 1.3 
<normalizedToken id="78B5EE39F5BB3DC76328F9548638DBC4" originalValue="×">x</normalizedToken>
longer than broad, with marginal anterior seta, 5 superior medial setae, distal-most paired, 5 setae inserted at anterodistal corner, 5 inferior medial setae, proximal-most paired, and 8 groups of plumose setae along posterior margin; palm oblique, straight, with serrate blade-like edge running the length, 5 outer and 5 inner bifid robust setae, 5 outer setae, and 2 inner setae; inner margin of defining angle with 6 bifid robust setae, outer margin with 5 bifid robust setae; dactylus with outer seta and seta placed along the inner margin at base of nail.
</paragraph>
<paragraph id="8ED045EDDBCB04559F568EC7EB83CE53" pageId="0" pageNumber="117">
<emphasis id="71042CFB89E5BA805E3777745B02CD52" bold="true" pageId="0" pageNumber="117">
<emphasis id="FF80E83D4DA4BBEB9ADB5DEC2BDF7BB4" italics="true" pageId="0" pageNumber="117">Pereopods</emphasis>
.
</emphasis>
Pereopod 3 (Fig. 
<figureCitation id="8B98B5EEECAF69B670F06CFA86AB6A73" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Stygobromus anacostensis sp. nov., Holotype male, 5.9 mm (USNM 1606902): A pereopod 3 B pereopod 4 C pereopod 5 D pereopod 6 E pereopod 7 F bifurcate sternal gill located on somites 6 and 7. Scale bar: 0.5 mm." figureDoi="10.3897/subtbiol.48.112984.figure5" httpUri="https://binary.pensoft.net/fig/984744" pageId="0" pageNumber="117">5A</figureCitation>
): coxal plate with 5 apical setae; merus 1.4 
<normalizedToken id="8EF2217D47FB07D2F2150F9EDE2410B5" originalValue="×">x</normalizedToken>
longer than carpus, carpus approximately 85% of propodus in length; dactylus approximately 50% length of propodus, with plumose seta on posterior margin, 2 setae along anterior margin followed by thin seta on medial margin.
</paragraph>
<caption id="19CBA624CD729D74B3556D3A4829DAEF" doi="10.3897/subtbiol.48.112984.figure5" httpUri="https://binary.pensoft.net/fig/984744" pageId="0" pageNumber="117" start="Figure 5" startId="F5">
<paragraph id="ED74DF18A0FBDC69CF89BE32F5691449" pageId="0" pageNumber="117">
<emphasis id="921018F3187E4F9C1E56B63BFECBA7DD" bold="true" pageId="0" pageNumber="117">Figure 5.</emphasis>
<taxonomicName id="12AEB64ACAC4404C6BBFDDC08E84CCA0" authorityName="Cannizzaro, Sawicki, &amp; Niemiller" authorityYear="2024" class="Malacostraca" family="Crangonyctidae" genus="Stygobromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stygobromus anacostensis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="anacostensis">
<emphasis id="EDE0CB203E082D72F62793CCD0912079" italics="true" pageId="0" pageNumber="117">Stygobromus anacostensis</emphasis>
</taxonomicName>
sp. nov., Holotype male, 5.9 mm (USNM 1606902): 
<emphasis id="B2A1B141DCDD7ABA8A51AC7667B94B3C" bold="true" pageId="0" pageNumber="117">A</emphasis>
pereopod 3 
<emphasis id="BB83DCDCD73C9DEC8C1E81FE391CAFD3" bold="true" pageId="0" pageNumber="117">B</emphasis>
pereopod 4 
<emphasis id="4149A236E32552933F620FE0E0A25C55" bold="true" pageId="0" pageNumber="117">C</emphasis>
pereopod 5 
<emphasis id="592E91E20B7A4B1E484F8C28D645D233" bold="true" pageId="0" pageNumber="117">D</emphasis>
pereopod 6 
<emphasis id="0FF8DECE0C514C51B96405B77674A88C" bold="true" pageId="0" pageNumber="117">E</emphasis>
pereopod 7 
<emphasis id="C5DC074D4814550756C2057D1AC95969" bold="true" pageId="0" pageNumber="117">F</emphasis>
bifurcate sternal gill located on somites 6 and 7. Scale bar: 0.5 mm.
</paragraph>
</caption>
<paragraph id="46F96910B985665FC4879A20F9512F00" pageId="0" pageNumber="117">
Pereopod 4 (Fig. 
<figureCitation id="CBE31261CF023753C1D8AC17E9CF2083" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Stygobromus anacostensis sp. nov., Holotype male, 5.9 mm (USNM 1606902): A pereopod 3 B pereopod 4 C pereopod 5 D pereopod 6 E pereopod 7 F bifurcate sternal gill located on somites 6 and 7. Scale bar: 0.5 mm." figureDoi="10.3897/subtbiol.48.112984.figure5" httpUri="https://binary.pensoft.net/fig/984744" pageId="0" pageNumber="117">5B</figureCitation>
): subequal to pereopod 3 in length; coxal plate armed with 4 anterior and 3 posterior apical setae; merus approximately 1.6 
<normalizedToken id="0732E78D27ABC5BC544777E4D383AE9D" originalValue="×">x</normalizedToken>
longer than carpus; carpus approximately 60% length of propodus; dactylus approximately 40% length of propodus, setation as in pereopod 3.
</paragraph>
<paragraph id="3C0354489396D1D2647A7D6AECB01546" pageId="0" pageNumber="117">
Pereopod 5 (Fig. 
<figureCitation id="C1CDB25FCAAAF2D42318A58D53A45442" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Stygobromus anacostensis sp. nov., Holotype male, 5.9 mm (USNM 1606902): A pereopod 3 B pereopod 4 C pereopod 5 D pereopod 6 E pereopod 7 F bifurcate sternal gill located on somites 6 and 7. Scale bar: 0.5 mm." figureDoi="10.3897/subtbiol.48.112984.figure5" httpUri="https://binary.pensoft.net/fig/984744" pageId="0" pageNumber="117">5C</figureCitation>
): coxal plate large, bilobate with distinct anterior and posterior lobes, posterior lobe with 4 robust setae on distal corner; basis posterior margin weakly convex with 9 shallow serrations, anterior margin with 6 split-tipped robust setae and 3 distal split-tipped setae; merus subequal in length to carpus; carpus subequal to propodus, dactylus approximately 50% length of propodus, setation as in pereopod 4.
</paragraph>
<paragraph id="FF12CE8AAB6E0A035E99AF1AB805DD4A" pageId="0" pageNumber="117">
Pereopod 6 (Fig. 
<figureCitation id="F5E3D7D1B945E0F6052DBA5F7F89086E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Stygobromus anacostensis sp. nov., Holotype male, 5.9 mm (USNM 1606902): A pereopod 3 B pereopod 4 C pereopod 5 D pereopod 6 E pereopod 7 F bifurcate sternal gill located on somites 6 and 7. Scale bar: 0.5 mm." figureDoi="10.3897/subtbiol.48.112984.figure5" httpUri="https://binary.pensoft.net/fig/984744" pageId="0" pageNumber="117">5D</figureCitation>
): coxal plate bilobate, with weakly produced anterior lobe, posterior lobe bearing 2 robust apical setae; basis posterior margin weakly convex with 8 serrations, anterior margin with 5 split-tipped robust setae, and 3 robust setae at anterodistal corner; merus approximately 1.2 
<normalizedToken id="0E7667885759FA8AF5BAF0085B5E2655" originalValue="×">x</normalizedToken>
length of carpus; carpus approximately 90% of propodus in length, dactylus approximately 50% length of propodus, setation as in pereopod 5.
</paragraph>
<paragraph id="08E0D50CCEC34519DBF02CF98BEA0F60" pageId="0" pageNumber="117">
Pereopod 7 (Fig. 
<figureCitation id="4BD5DD2ED11C8AABF64183F6D67913E2" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Stygobromus anacostensis sp. nov., Holotype male, 5.9 mm (USNM 1606902): A pereopod 3 B pereopod 4 C pereopod 5 D pereopod 6 E pereopod 7 F bifurcate sternal gill located on somites 6 and 7. Scale bar: 0.5 mm." figureDoi="10.3897/subtbiol.48.112984.figure5" httpUri="https://binary.pensoft.net/fig/984744" pageId="0" pageNumber="117">5E</figureCitation>
): coxal plate small, subtriangular, with 4 posterior setae; basis posterior margin weakly convex with 10 serrations and straight distal corner, anterior margin with 8 split-tipped robust setae, and 2 robust setae at anterodistal corner; merus subequal in length to carpus; carpus approximately 80% length of propodus, dactylus approximately 40% length of propodus, setation as in pereopods 5, 6.
</paragraph>
<paragraph id="BC04F051AA086290E433D34024CDCD89" pageId="0" pageNumber="117">
<emphasis id="DD8BADD3167747C22BBF5A39B7FD4852" bold="true" italics="true" pageId="0" pageNumber="117">Gills</emphasis>
(Fig. 
<figureCitation id="B01ED84B1C5AABB5D8770DB341FB5AB9" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Stygobromus anacostensis sp. nov., Holotype male, 5.9 mm (USNM 1606902): A pereopod 3 B pereopod 4 C pereopod 5 D pereopod 6 E pereopod 7 F bifurcate sternal gill located on somites 6 and 7. Scale bar: 0.5 mm." figureDoi="10.3897/subtbiol.48.112984.figure5" httpUri="https://binary.pensoft.net/fig/984744" pageId="0" pageNumber="117">5F</figureCitation>
). coxal gills on somites 2-6, somites 6 and 7 with bifurcate sternal gills.
</paragraph>
<paragraph id="CB270796047681DA254BA2AC79133298" pageId="0" pageNumber="117">
<emphasis id="CDE708EFA90604F39627BB50F4931182" bold="true" pageId="0" pageNumber="117">
<emphasis id="20A7DC50558B7B20C996B10851C5F03B" italics="true" pageId="0" pageNumber="117">Pleon</emphasis>
.
</emphasis>
Epimera (Fig. 
<figureCitation id="56C2A9D7B7377390836B46AB3AF68D6B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Stygobromus anacostensis sp. nov., Holotype male, 5.9 mm (USNM 1606902): A epimera 1 - 3 B pleopod 1 (coupling spines and clothes pin seta enlarged) C uropod 1 (posteromedial protuberance enlarged) D uropod 2 E uropod 3 F telson. Scale bars: 0.25 mm." figureDoi="10.3897/subtbiol.48.112984.figure6" httpUri="https://binary.pensoft.net/fig/984745" pageId="0" pageNumber="117">6A</figureCitation>
): first epimeron ventral margin with robust seta, distoposterior corner rounded, posterior margin with 2 setae. Second epimeron ventral margin with 3 robust setae, distoposterior corner rounded, posterior margin with 2 setae. Third epimeron ventral margin with 3 robust setae, distoposterior corner rounded, posterior margin with 2 setae.
</paragraph>
<caption id="576FA5258D2E5C74F70A47F52245C20B" doi="10.3897/subtbiol.48.112984.figure6" httpUri="https://binary.pensoft.net/fig/984745" pageId="0" pageNumber="117" start="Figure 6" startId="F6">
<paragraph id="67387667F32D645E43CF1A1BCE0DBE64" pageId="0" pageNumber="117">
<emphasis id="D5EE1D8247243B961609B881CCB0C0B9" bold="true" pageId="0" pageNumber="117">Figure 6.</emphasis>
<taxonomicName id="13E08A0934452C2B0548E4E6EC306176" authorityName="Cannizzaro, Sawicki, &amp; Niemiller" authorityYear="2024" class="Malacostraca" family="Crangonyctidae" genus="Stygobromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stygobromus anacostensis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="anacostensis">
<emphasis id="790E869BE27B3DFBE9AD7A6AC77C731F" italics="true" pageId="0" pageNumber="117">Stygobromus anacostensis</emphasis>
</taxonomicName>
sp. nov., Holotype male, 5.9 mm (USNM 1606902): 
<emphasis id="9FBEE02371F671801705D1DE2B1EE2C7" bold="true" pageId="0" pageNumber="117">A</emphasis>
epimera 1-3 
<emphasis id="E231725538C818D19D7D2FFFBBA0977B" bold="true" pageId="0" pageNumber="117">B</emphasis>
pleopod 1 (coupling spines and clothes pin seta enlarged) 
<emphasis id="16AA80EFA8A33165714CCAE0C5E0C6BF" bold="true" pageId="0" pageNumber="117">C</emphasis>
uropod 1 (posteromedial protuberance enlarged) 
<emphasis id="4375F58587C1BED7332B221F4359AA7F" bold="true" pageId="0" pageNumber="117">D</emphasis>
uropod 2 
<emphasis id="6A4587D91A92E921E9DF4E0FC419DD68" bold="true" pageId="0" pageNumber="117">E</emphasis>
uropod 3 
<emphasis id="1E2BEE0C6307BC45721581E6D5A5C0CE" bold="true" pageId="0" pageNumber="117">F</emphasis>
telson. Scale bars: 0.25 mm.
</paragraph>
</caption>
<paragraph id="4D54D75D87873C7E4A17E125FDD1B4C7" pageId="0" pageNumber="117">
Pleopods: pleopod 1 (Fig. 
<figureCitation id="575788301D3BD8095C4E0E84EC425422" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Stygobromus anacostensis sp. nov., Holotype male, 5.9 mm (USNM 1606902): A epimera 1 - 3 B pleopod 1 (coupling spines and clothes pin seta enlarged) C uropod 1 (posteromedial protuberance enlarged) D uropod 2 E uropod 3 F telson. Scale bars: 0.25 mm." figureDoi="10.3897/subtbiol.48.112984.figure6" httpUri="https://binary.pensoft.net/fig/984745" pageId="0" pageNumber="117">6B</figureCitation>
) peduncle lacking setae, with 2 coupling hooks; outer, inner rami with 8 and 11 segments respectively, basal segment of outer ramus with clothes-pin setae. Pleopod 2 peduncle lacking setae, with 2 coupling hooks; outer, inner rami with 7, 11 segments respectively, basal segment of outer ramus with clothes-pin setae. Pleopod 3 outer, inner rami with 7, 7 segments respectively, basal segment of outer ramus with clothes-pin setae.
</paragraph>
<paragraph id="1F0D671E31657AFB25A4A7D9698C9564" pageId="0" pageNumber="117">
<emphasis id="E53E2C38D5EAECCE573917A8B71A0905" bold="true" pageId="0" pageNumber="117">
<emphasis id="5F9BACCDA558AA91DCF9C26887516C76" italics="true" pageId="0" pageNumber="117">Urosome</emphasis>
.
</emphasis>
Mostly bare, with sparse setae covering dorsal surface. Uropod 1 (Fig. 
<figureCitation id="ADD26F592AF723DA050B9AAB71A970C0" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Stygobromus anacostensis sp. nov., Holotype male, 5.9 mm (USNM 1606902): A epimera 1 - 3 B pleopod 1 (coupling spines and clothes pin seta enlarged) C uropod 1 (posteromedial protuberance enlarged) D uropod 2 E uropod 3 F telson. Scale bars: 0.25 mm." figureDoi="10.3897/subtbiol.48.112984.figure6" httpUri="https://binary.pensoft.net/fig/984745" pageId="0" pageNumber="117">6C</figureCitation>
): peduncle 1.4 
<normalizedToken id="CCA4CFCB86018F24EC5A797D6D013587" originalValue="×">x</normalizedToken>
inner ramus in length, with 8 outer robust setae and inner robust seta(e), posteromedial margin with distinct protuberance approximately 20% of inner ramus in length, dorsal margin weakly serrate; outer ramus approximately 80% length of inner, with 2 inner and outer robust setae and 4 apical robust setae; inner ramus possessing 3 outer and two inner robust setae, and 5 apical robust setae.
</paragraph>
<paragraph id="84B94C635BC01BF75E2494A7CAA6E2C9" pageId="0" pageNumber="117">
Uropod 2 (Fig. 
<figureCitation id="25CC62805544F3EFC53446D613E05A77" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Stygobromus anacostensis sp. nov., Holotype male, 5.9 mm (USNM 1606902): A epimera 1 - 3 B pleopod 1 (coupling spines and clothes pin seta enlarged) C uropod 1 (posteromedial protuberance enlarged) D uropod 2 E uropod 3 F telson. Scale bars: 0.25 mm." figureDoi="10.3897/subtbiol.48.112984.figure6" httpUri="https://binary.pensoft.net/fig/984745" pageId="0" pageNumber="117">6D</figureCitation>
): peduncle subequal in length to inner ramus, with 2 outer robust setae and inner robust seta; outer ramus approximately 88% length of inner ramus without robust setae along the inner and outer margins, and 4 apical robust setae; inner ramus with 2 outer and 2 inner robust setae, with 5 apical robust setae.
</paragraph>
<paragraph id="7F868CB367DDCD58050542B2CC9C3557" pageId="0" pageNumber="117">
Uropod 3 (Fig. 
<figureCitation id="21615BE709AAC9135CAB55B4478FE878" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Stygobromus anacostensis sp. nov., Holotype male, 5.9 mm (USNM 1606902): A epimera 1 - 3 B pleopod 1 (coupling spines and clothes pin seta enlarged) C uropod 1 (posteromedial protuberance enlarged) D uropod 2 E uropod 3 F telson. Scale bars: 0.25 mm." figureDoi="10.3897/subtbiol.48.112984.figure6" httpUri="https://binary.pensoft.net/fig/984745" pageId="0" pageNumber="117">6E</figureCitation>
): small, shorter than telson, uniramous; peduncle 2 
<normalizedToken id="F2BF2FEB258E6E7999E79C4497F3ACDB" originalValue="×">x</normalizedToken>
length of ramus; ramus with 3 apical robust setae.
</paragraph>
<paragraph id="84664D46B04520713BF0F02A4ED9A494" pageId="0" pageNumber="117">
<emphasis id="87A89886B7257A5287196480CB52DEC9" bold="true" italics="true" pageId="0" pageNumber="117">Telson</emphasis>
(Fig. 
<figureCitation id="D993524DFB52F27771F1BB9DAC04DFD9" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Stygobromus anacostensis sp. nov., Holotype male, 5.9 mm (USNM 1606902): A epimera 1 - 3 B pleopod 1 (coupling spines and clothes pin seta enlarged) C uropod 1 (posteromedial protuberance enlarged) D uropod 2 E uropod 3 F telson. Scale bars: 0.25 mm." figureDoi="10.3897/subtbiol.48.112984.figure6" httpUri="https://binary.pensoft.net/fig/984745" pageId="0" pageNumber="117">6F</figureCitation>
). Telson entire, elongated, 1.5 
<normalizedToken id="83237977FC5FA8DA5F36778F226F234E" originalValue="×">x</normalizedToken>
longer than broad, weakly tapering distally; apex with 10 robust setae, and plumose seta, 2 plumose setae arise dorsolaterally from both outer margins.
</paragraph>
<paragraph id="14EF3DA8C8F919E299C44B19F70F8F02" pageId="0" pageNumber="117">
<emphasis id="BD9D283F6629B0AF6262B12EFDB38C76" bold="true" pageId="0" pageNumber="117">Female</emphasis>
: allotype USNM 1606903 (Fig. 
<figureCitation id="131863C231D2916A5C81DEC968DF2802" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Stygobromus anacostensis sp. nov., habitus: A holotype male, 5.9 mm (USNM 1606902) B Allotype female, 5.3 mm (USNM 1606903). Scale bar: 1 mm." figureDoi="10.3897/subtbiol.48.112984.figure1" httpUri="https://binary.pensoft.net/fig/984740" pageId="0" pageNumber="117">1B</figureCitation>
); Size 5.3 mm. Differing from male in several points, including, antennae; gnathopod shape and setation; uropods 1 and 2 shape and setation. Structures not described below are as in male.
</paragraph>
<paragraph id="010DF55AE5FF062E073613B8FD1BBBED" pageId="0" pageNumber="117">
<emphasis id="EF2CBBC94652D189FAE35EE5BFD8E6D3" bold="true" pageId="0" pageNumber="117">
<emphasis id="BFEB734F74CC35B83790C9DDC2A3415D" italics="true" pageId="0" pageNumber="117">Antennae</emphasis>
.
</emphasis>
Antenna 1 (not illustrated, but see Fig. 
<figureCitation id="15C265B3A5352120645049A83FC4FB61" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Stygobromus anacostensis sp. nov., habitus: A holotype male, 5.9 mm (USNM 1606902) B Allotype female, 5.3 mm (USNM 1606903). Scale bar: 1 mm." figureDoi="10.3897/subtbiol.48.112984.figure1" httpUri="https://binary.pensoft.net/fig/984740" pageId="0" pageNumber="117">1B</figureCitation>
): 50% body length, 1.5 
<normalizedToken id="FEFE5E8AC3ED3C40BE226EC8D1A8A4FA" originalValue="×">x</normalizedToken>
longer than antenna 2; peduncle, flagellum lacking robust setae; primary flagellum with 16 segments. Antenna 2 (Fig. 
<figureCitation id="A48D4619DFAFDA2F1F8F1428EFDEF345" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Stygobromus anacostensis sp. nov., Allotype female, 5.3 mm (USNM 1606903): A antenna 2 B gnathopod 1 (palm and dactyl enlarged) C gnathopod 2 (rastellate seta, palm and dactyl enlarged). Scale bars: 0.25 mm (A); 0.5 mm (B, C)." figureDoi="10.3897/subtbiol.48.112984.figure7" httpUri="https://binary.pensoft.net/fig/984746" pageId="0" pageNumber="117">7A</figureCitation>
): gland cone distinct; peduncle 1.5 
<normalizedToken id="8A8ECEA796DAEA59D1E79EF9B12D2ADE" originalValue="×">x</normalizedToken>
longer than flagellum, with robust setae anteriorly, laterally on segments 3, 4, peduncle segment 4 subequal in length to segment 5; flagellum 7-segmented, without small calceoli-like structures apically on distal flagellar segments.
</paragraph>
<paragraph id="8B385165434DE017DA7CE675049EA99A" pageId="0" pageNumber="117">
<emphasis id="6FCF4D5FC1BE39041BD41DF40092EA43" bold="true" pageId="0" pageNumber="117">
<emphasis id="5E9A69E09D5511B370CC4324B8B423B1" italics="true" pageId="0" pageNumber="117">Gnathopods</emphasis>
.
</emphasis>
Gnathopod 1 (Fig. 
<figureCitation id="3AEB58AB0B1665A9C4EECA5A283E2C02" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Stygobromus anacostensis sp. nov., Allotype female, 5.3 mm (USNM 1606903): A antenna 2 B gnathopod 1 (palm and dactyl enlarged) C gnathopod 2 (rastellate seta, palm and dactyl enlarged). Scale bars: 0.25 mm (A); 0.5 mm (B, C)." figureDoi="10.3897/subtbiol.48.112984.figure7" httpUri="https://binary.pensoft.net/fig/984746" pageId="0" pageNumber="117">7B</figureCitation>
): coxal plate with 3 apical and 2 facial setae; ischium with 2 setae and pubescence along posterior margin; carpus approximately 40% length of propodus with robust seta along anterior margin and a group of setae on anterodistal margin, one of which is approximately 50% length of propodus, posterior margin with single group of plumose setae and 4 submarginal setae directed distally; propodus 1.25 
<normalizedToken id="87E1C3552900F89997AD0DBEAB351067" originalValue="×">x</normalizedToken>
longer than broad, with 1 marginal anterior seta, 3 superior medial setae, 3 setae inserted at anterodistal corner, 3 inferior medial setae and numerous plumose posterior setae; palm oblique, straight, with serrate blade-like edge running the length, 5 outer and 5 inner bifid robust setae, 4 outer setae, and inner seta; inner margin of defining angle with 3 bifid robust setae, outer margin with 4 bifid robust setae; dactylus with outer seta and 4 short setae covering the inner margin and 2 setae placed along the inner margin at base of nail.
</paragraph>
<caption id="23FD9B7BAD315AF5EBA9EA43E52E3766" doi="10.3897/subtbiol.48.112984.figure7" httpUri="https://binary.pensoft.net/fig/984746" pageId="0" pageNumber="117" start="Figure 7" startId="F7">
<paragraph id="E6AF29C83A7F6259540A5BE263D7A303" pageId="0" pageNumber="117">
<emphasis id="9A28963738CB6DCEBB8D3AA39C8CD4B9" bold="true" pageId="0" pageNumber="117">Figure 7.</emphasis>
<taxonomicName id="93F5DCECDC244AE1A4B91B2DA438EF1F" authorityName="Cannizzaro, Sawicki, &amp; Niemiller" authorityYear="2024" class="Malacostraca" family="Crangonyctidae" genus="Stygobromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stygobromus anacostensis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="anacostensis">
<emphasis id="561C7E1E457A6F708741487EB20526E0" italics="true" pageId="0" pageNumber="117">Stygobromus anacostensis</emphasis>
</taxonomicName>
sp. nov., Allotype female, 5.3 mm (USNM 1606903): 
<emphasis id="FD326349257DB962349B9802DAC6F73C" bold="true" pageId="0" pageNumber="117">A</emphasis>
antenna 2 
<emphasis id="9270CF935537D0AE05C469FE1AE569AC" bold="true" pageId="0" pageNumber="117">B</emphasis>
gnathopod 1 (palm and dactyl enlarged) 
<emphasis id="E985AE0D9EE71AD8634866A3F71666D9" bold="true" pageId="0" pageNumber="117">C</emphasis>
gnathopod 2 (rastellate seta, palm and dactyl enlarged). Scale bars: 0.25 mm (
<emphasis id="2723A446D2FDE7CD53DD9140AF3541DD" bold="true" pageId="0" pageNumber="117">A</emphasis>
); 0.5 mm (
<emphasis id="2F61A4A6CAF128D54A4D710DECD60824" bold="true" pageId="0" pageNumber="117">B, C</emphasis>
).
</paragraph>
</caption>
<paragraph id="6FD4A85F877D7ADC5FDAE45A12A18A64" pageId="0" pageNumber="117">
Gnathopod 2 (Fig. 
<figureCitation id="A24713AC3B0662D06E41AC1FEE0389C6" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Stygobromus anacostensis sp. nov., Allotype female, 5.3 mm (USNM 1606903): A antenna 2 B gnathopod 1 (palm and dactyl enlarged) C gnathopod 2 (rastellate seta, palm and dactyl enlarged). Scale bars: 0.25 mm (A); 0.5 mm (B, C)." figureDoi="10.3897/subtbiol.48.112984.figure7" httpUri="https://binary.pensoft.net/fig/984746" pageId="0" pageNumber="117">7C</figureCitation>
): coxal plate with 6 apical setae and 2 facial setae; ischium with 2 setae and pubescence along posterior margin; merus with pubescence covering posterior surface and 4 posterodistal setae, without robust seta along anterior margin, and two robust setae along distal margin; carpus subequal in length to propodus, with robust seta along anterior margin and two setae on anterodistal margin, one of which is approximately 33% length of propodus, posterior margin with 3 groups of plumose setae, distal-most bearing rastellate seta, and 3 submarginal setae directed distally; propodus 1.1 
<normalizedToken id="9E1FE76EC45551A91B78B05687B9CAD3" originalValue="×">x</normalizedToken>
longer than broad, with marginal anterior seta, 3 superior medial setae, 4 setae inserted at anterodistal corner, 4 inferior medial setae, and 5 groups of plumose setae along posterior margin; palm oblique, straight, with serrate blade-like edge running the length, 3 outer and 3 inner bifid robust setae, 3 outer setae, and 1 inner seta; inner margin of defining angle with 4 bifid robust setae, outer margin with 4 bifid robust setae; dactylus with outer seta and 4 short setae covering the inner margin and seta placed along the inner margin at base of nail.
</paragraph>
<paragraph id="2C849BC7CD8C940F33E362D69B9D35BC" pageId="0" pageNumber="117">
<emphasis id="0FFE572CFFAF6BFF9F1036F64B346E5C" bold="true" pageId="0" pageNumber="117">
<emphasis id="325D59EEB29E49475781CE5FB4082E40" italics="true" pageId="0" pageNumber="117">Gills and brood plates</emphasis>
.
</emphasis>
Gills as in male with coxal gills on somites 2-6, somites 6 and 7 with bifurcate sternal gills (Fig. 
<figureCitation id="2626E15E0303084E9F6D2D9B76C4163B" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Stygobromus anacostensis sp. nov., Allotype female, 5.3 mm (USNM 1606903): A coxa and basis of pereopod 7 showing placement of bifurcate sternal gill B uropod 1 C uropod 2 D uropod 3 E telson. Scale bars: 0.5 mm (A-C); 0.25 mm (D, E)." figureDoi="10.3897/subtbiol.48.112984.figure8" httpUri="https://binary.pensoft.net/fig/984747" pageId="0" pageNumber="117">8A</figureCitation>
illustrates somite 7). Brood plates early in development in allotype, present on somites 2-5.
</paragraph>
<caption id="3B83062529619C61C623B9C06E84076B" doi="10.3897/subtbiol.48.112984.figure8" httpUri="https://binary.pensoft.net/fig/984747" pageId="0" pageNumber="117" start="Figure 8" startId="F8">
<paragraph id="81069B3FD538B9EAAFDF5AD55E80546A" pageId="0" pageNumber="117">
<emphasis id="8C1441F10E307A393272CAFAEEB7CDC7" bold="true" pageId="0" pageNumber="117">Figure 8.</emphasis>
<taxonomicName id="09A3AF221DA8B10DCAFE46AABDC439A0" authorityName="Cannizzaro, Sawicki, &amp; Niemiller" authorityYear="2024" class="Malacostraca" family="Crangonyctidae" genus="Stygobromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stygobromus anacostensis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="anacostensis">
<emphasis id="2DDEE85F5171D570DA4B0E9034F668D4" italics="true" pageId="0" pageNumber="117">Stygobromus anacostensis</emphasis>
</taxonomicName>
sp. nov., Allotype female, 5.3 mm (USNM 1606903): 
<emphasis id="4FB520652756C59E26730AB81B246DDE" bold="true" pageId="0" pageNumber="117">A</emphasis>
coxa and basis of pereopod 7 showing placement of bifurcate sternal gill 
<emphasis id="C4E23D05756FF676D095BA0E3DFD3272" bold="true" pageId="0" pageNumber="117">B</emphasis>
uropod 1 
<emphasis id="5FF1FC785F68FCC3E89D41416759892E" bold="true" pageId="0" pageNumber="117">C</emphasis>
uropod 2 
<emphasis id="5ABF9126402D21FB1E866D304FB32C6A" bold="true" pageId="0" pageNumber="117">D</emphasis>
uropod 3 
<emphasis id="EAF7B9FFD8791073913914094A3730DF" bold="true" pageId="0" pageNumber="117">E</emphasis>
telson. Scale bars: 0.5 mm (
<emphasis id="E49F47306AA0EA980807B23CBE23741A" bold="true" pageId="0" pageNumber="117">A-C</emphasis>
); 0.25 mm (
<emphasis id="2A96FCC4309E03F6535598A4CCCB6613" bold="true" pageId="0" pageNumber="117">D, E</emphasis>
).
</paragraph>
</caption>
<paragraph id="5A180910F621D67CCFA42550791EC4ED" pageId="0" pageNumber="117">
<emphasis id="FC7F65EE48B4F1A9884E75B291E7F925" bold="true" pageId="0" pageNumber="117">
<emphasis id="F0C8853B205EEC8601A35E2DD4437FC1" italics="true" pageId="0" pageNumber="117">Urosome</emphasis>
.
</emphasis>
Uropod 1 (Fig. 
<figureCitation id="ACA09A231B33D6984441383637EF3745" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Stygobromus anacostensis sp. nov., Allotype female, 5.3 mm (USNM 1606903): A coxa and basis of pereopod 7 showing placement of bifurcate sternal gill B uropod 1 C uropod 2 D uropod 3 E telson. Scale bars: 0.5 mm (A-C); 0.25 mm (D, E)." figureDoi="10.3897/subtbiol.48.112984.figure8" httpUri="https://binary.pensoft.net/fig/984747" pageId="0" pageNumber="117">8B</figureCitation>
): peduncle 1.5 
<normalizedToken id="21996539CDA842E13FDDFE73F495DDA5" originalValue="×">x</normalizedToken>
length of inner ramus, with 6 outer robust setae and inner robust seta(e), posteromedial margin lacking protuberance; outer ramus approximately 90% length of inner, with 1 inner and outer robust seta, and 4 apical robust setae; inner ramus possessing 2 outer and inner robust seta(e), and 5 apical robust setae.
</paragraph>
<paragraph id="6AEEAA1466BC3F395DCB45851B6E7379" pageId="0" pageNumber="117">
Uropod 2 (Fig. 
<figureCitation id="5C32517E3A4AAE012D63FC1012317349" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Stygobromus anacostensis sp. nov., Allotype female, 5.3 mm (USNM 1606903): A coxa and basis of pereopod 7 showing placement of bifurcate sternal gill B uropod 1 C uropod 2 D uropod 3 E telson. Scale bars: 0.5 mm (A-C); 0.25 mm (D, E)." figureDoi="10.3897/subtbiol.48.112984.figure8" httpUri="https://binary.pensoft.net/fig/984747" pageId="0" pageNumber="117">8C</figureCitation>
): peduncle subequal in length to inner ramus, with 2 outer robust setae and inner robust seta; outer ramus approximately 66% length of inner ramus with outer robust seta, and 4 apical robust setae, inner robust setae lacking.
</paragraph>
<paragraph id="7321D1AB556F521DFC8A8BD06B313349" pageId="0" pageNumber="117">
Uropod 3 (Fig. 
<figureCitation id="765C8E574E47C1D6173E547C22BC9EAB" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Stygobromus anacostensis sp. nov., Allotype female, 5.3 mm (USNM 1606903): A coxa and basis of pereopod 7 showing placement of bifurcate sternal gill B uropod 1 C uropod 2 D uropod 3 E telson. Scale bars: 0.5 mm (A-C); 0.25 mm (D, E)." figureDoi="10.3897/subtbiol.48.112984.figure8" httpUri="https://binary.pensoft.net/fig/984747" pageId="0" pageNumber="117">8D</figureCitation>
): small, shorter than telson, uniramous; peduncle 1.5 
<normalizedToken id="5724D09724A1FBC48115C1D48D1F66C5" originalValue="×">x</normalizedToken>
length of ramus; ramus with 4 apical robust setae.
</paragraph>
<paragraph id="962A9AD49B5DDE965087CEC1D64FCFAB" pageId="0" pageNumber="117">
<emphasis id="E38657E02D33552D22CCA1AE7DC367C1" bold="true" italics="true" pageId="0" pageNumber="117">Telson</emphasis>
(Fig. 
<figureCitation id="EC49959764C00DA4647D40827F779228" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Stygobromus anacostensis sp. nov., Allotype female, 5.3 mm (USNM 1606903): A coxa and basis of pereopod 7 showing placement of bifurcate sternal gill B uropod 1 C uropod 2 D uropod 3 E telson. Scale bars: 0.5 mm (A-C); 0.25 mm (D, E)." figureDoi="10.3897/subtbiol.48.112984.figure8" httpUri="https://binary.pensoft.net/fig/984747" pageId="0" pageNumber="117">8E</figureCitation>
). Telson entire, elongated, 1.5 
<normalizedToken id="1B82B3F20A7B0891C65738DF73E2A409" originalValue="×">x</normalizedToken>
longer than broad, weakly tapering distally; apex with 9 robust setae, 2 plumose setae arise dorsolaterally from both outer margins.
</paragraph>
</subSubSection>
<subSubSection id="C10BE0A2F6BEF85EFDEAB2259C9B4365" pageId="0" pageNumber="117" type="variation">
<paragraph id="FD04E8874F72C0BC511E413A7D737E62" pageId="0" pageNumber="117">Variation.</paragraph>
<paragraph id="D02D92F3C1334DAD6F18D11EE7D4F8E2" pageId="0" pageNumber="117">
The new species was shown to vary slightly in several morphological characteristics, particularly between males and females (Table 
<tableCitation id="BB189F650E138A2672A39257DCA315A9" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Variation in morphological characters among select specimens of Stygobromus anacostensis sp. nov. examined." httpUri="http://table.plazi.org/id/1CC5227CFE65500225D6CC81B4B08EF3" pageId="0" pageNumber="117" tableUuid="1CC5227CFE65500225D6CC81B4B08EF3">3</tableCitation>
).
</paragraph>
<caption id="1CC5227CFE65500225D6CC81B4B08EF3" ID-Table-UUID="1CC5227CFE65500225D6CC81B4B08EF3" httpUri="http://table.plazi.org/id/1CC5227CFE65500225D6CC81B4B08EF3" pageId="0" pageNumber="117" start="Table 3" startId="T3">
<paragraph id="DF6D119D660AA26F5EE94AB7A4913C1E" pageId="0" pageNumber="117">
<emphasis id="4F68EBCD68C130CFC2017C886454A371" bold="true" pageId="0" pageNumber="117">Table 3.</emphasis>
Variation in morphological characters among select specimens of 
<taxonomicName id="6F0A1BC782EC7266BCBF3C235908C928" authorityName="Cannizzaro, Sawicki, &amp; Niemiller" authorityYear="2024" class="Malacostraca" family="Crangonyctidae" genus="Stygobromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stygobromus anacostensis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="anacostensis">
<emphasis id="74AE746C881D1A1A61EB0C8669F5C22D" italics="true" pageId="0" pageNumber="117">Stygobromus anacostensis</emphasis>
</taxonomicName>
sp. nov. examined.
</paragraph>
</caption>
<paragraph id="D626515469F1BA44904ACC8E5E4699CE" pageId="0" pageNumber="117">
<table id="BBB8A12426A44845B7543670CB9B26FB" pageId="0" pageNumber="117">
<tr id="2D793F75E5D5CC400C05C57122CF693D" pageId="0" pageNumber="117">
<th id="36D5F6A43FA985D5B68F3E5F3E82A8CB" colspan="1" pageId="0" pageNumber="117" rowspan="1">Character</th>
<th id="6CB068CEEE16B62224971EB927E1B781" colspan="1" pageId="0" pageNumber="117" rowspan="1">Holotype Male USNM 1606902</th>
<th id="18F853043B547A30F9F7D00DB244E017" colspan="1" pageId="0" pageNumber="117" rowspan="1">Paratype Male USNM 1606904</th>
<th id="8E511000406B3FE43B1AD6982704179E" colspan="1" pageId="0" pageNumber="117" rowspan="1">Allotype Female USNM 1606903</th>
<th id="A049AD92773F37DA3E8554D4859FF395" colspan="1" pageId="0" pageNumber="117" rowspan="1">Paratype Female USNM 1606906</th>
<th id="DACD65142EE084CB48E81CEDA79EE75E" colspan="1" pageId="0" pageNumber="117" rowspan="1">Paratype Female USNM 1606905</th>
</tr>
<tr id="0C1D75EFDC38CE0C1935E6DA917BDF6A" pageId="0" pageNumber="117">
<td id="1AC36AA7B166262534489F176DF70FA3" colspan="1" pageId="0" pageNumber="117" rowspan="1">
<emphasis id="0F4CC1E91E54A99F5216872A2D8B31D6" bold="true" pageId="0" pageNumber="117">Body size</emphasis>
</td>
<td id="AFD0CE91DD67D576E8720E8287F08039" colspan="1" pageId="0" pageNumber="117" rowspan="1">5.9 mm</td>
<td id="097562AEF74757D771DFDF6129E66F41" colspan="1" pageId="0" pageNumber="117" rowspan="1">6.7 mm</td>
<td id="CA8B82E6B928091CB8F998C02DB88961" colspan="1" pageId="0" pageNumber="117" rowspan="1">5.3 mm</td>
<td id="5DE8F0D7F10CCE1F236D137C991ED67C" colspan="1" pageId="0" pageNumber="117" rowspan="1">4.8 mm</td>
<td id="4E9FD447E3EC6066B195DE367F87E80A" colspan="1" pageId="0" pageNumber="117" rowspan="1">4.8 mm</td>
</tr>
<tr id="99382C4559C25BBA2A1ED0A993E0FA3E" pageId="0" pageNumber="117">
<td id="360CFA8CCBD77C22B3583466696299C2" colspan="6" pageId="0" pageNumber="117" rowspan="1">
<emphasis id="56674CD039DBAA6AEE18063D5F11E4C4" bold="true" pageId="0" pageNumber="117">Antenna 1</emphasis>
</td>
</tr>
<tr id="848A60BB6A9E5099804140C235D7CD5E" pageId="0" pageNumber="117">
<td id="14C120CC65847308DF6DF5B09D78BAB8" colspan="1" pageId="0" pageNumber="117" rowspan="1">Flagellar segments</td>
<td id="1C132159C1F136031636391A98CA854E" colspan="1" pageId="0" pageNumber="117" rowspan="1">18</td>
<td id="6F5316630258CB13312886B475E71240" colspan="1" pageId="0" pageNumber="117" rowspan="1">21</td>
<td id="038C740B90AD3C6FFA5E569A3B42C0A9" colspan="1" pageId="0" pageNumber="117" rowspan="1">16</td>
<td id="670D8CC11A53302CAD3D0C61D6E67343" colspan="1" pageId="0" pageNumber="117" rowspan="1">16</td>
<td id="F0990991F9D8881F55E8485F11343FF1" colspan="1" pageId="0" pageNumber="117" rowspan="1">12</td>
</tr>
<tr id="B07F0A16150EB694A257E87040E8B5E7" pageId="0" pageNumber="117">
<td id="258E930A1658BB550529B2A102563A87" colspan="1" pageId="0" pageNumber="117" rowspan="1">Accessory flagellum</td>
<td id="364133693036FF8F2C96B990BFFE343B" colspan="1" pageId="0" pageNumber="117" rowspan="1">&gt; than 1st flagellar segment</td>
<td id="F486F19CAA6EB29AFA2C076AAEFE99CE" colspan="1" pageId="0" pageNumber="117" rowspan="1">&gt; than 2nd flagellar segment</td>
<td id="40FFA3415051DCC15A2601C9FE10041B" colspan="1" pageId="0" pageNumber="117" rowspan="1">&gt; than 1st flagellar segment</td>
<td id="78A5CB5D78021871939945EA59F68F0D" colspan="1" pageId="0" pageNumber="117" rowspan="1">&gt; than 2nd flagellar segment</td>
<td id="F626FC23C2C1C584ECBD71F6969DAE49" colspan="1" pageId="0" pageNumber="117" rowspan="1">sub equal to second flagellar segment</td>
</tr>
<tr id="5670829FE69E9585C90A6D58405C98FE" pageId="0" pageNumber="117">
<td id="1E33D3E3CA0D8328B901475514FC17B3" colspan="6" pageId="0" pageNumber="117" rowspan="1">
<emphasis id="9458CC8DFA44A2484DC0B06BE88C4E10" bold="true" pageId="0" pageNumber="117">Antenna 2</emphasis>
</td>
</tr>
<tr id="63B054C70D6CB931E4CE6826462191F5" pageId="0" pageNumber="117">
<td id="DC5F6414D1F33F060AB7B66831018063" colspan="1" pageId="0" pageNumber="117" rowspan="1">Peduncle Segments 4 to 5 length</td>
<td id="7BC6B18344982E8285602C64C6E54808" colspan="1" pageId="0" pageNumber="117" rowspan="1">
1.04 
<normalizedToken id="0DD9BFBB5B603C6C0E9E6FA8F57A8340" originalValue="×">x</normalizedToken>
</td>
<td id="B7E4962B2700176C912C76BE28439070" colspan="1" pageId="0" pageNumber="117" rowspan="1">97%</td>
<td id="E980C901B8E2FC7C5E65FF8CF7D77C2B" colspan="1" pageId="0" pageNumber="117" rowspan="1">
1.15 
<normalizedToken id="7A77FD9CCE6BE2B3882FE83ED7B4A4F2" originalValue="×">x</normalizedToken>
</td>
<td id="9691A4668628D68466767C3AF6D7036D" colspan="1" pageId="0" pageNumber="117" rowspan="1">
1.12 
<normalizedToken id="E96CB1A7A0A99A796C97480E6C72397A" originalValue="×">x</normalizedToken>
</td>
<td id="4CAC97057487091AC2EB487BBF8A4BFB" colspan="1" pageId="0" pageNumber="117" rowspan="1">
1.13 
<normalizedToken id="62BCC52E87ABF246CCEAC5C03BFEC730" originalValue="×">x</normalizedToken>
</td>
</tr>
<tr id="B1763AE23B6EB0234A4E9CD36431B25F" pageId="0" pageNumber="117">
<td id="6D614EA4395470F1D93E4463C46EEFC4" colspan="1" pageId="0" pageNumber="117" rowspan="1">Flagellar segments</td>
<td id="1B38768729BA64CFD8787302BFAD26E5" colspan="1" pageId="0" pageNumber="117" rowspan="1">unknown</td>
<td id="752A3B87977DC637C29BC1CD0727AA01" colspan="1" pageId="0" pageNumber="117" rowspan="1">12</td>
<td id="8E7438960736451326D225B2368BFAB1" colspan="1" pageId="0" pageNumber="117" rowspan="1">7</td>
<td id="06D1C70CBDCB49AE67D5E8B3066BDC14" colspan="1" pageId="0" pageNumber="117" rowspan="1">7</td>
<td id="CB3DC492B77BF94C40C34BC7DFEC4EBA" colspan="1" pageId="0" pageNumber="117" rowspan="1">7</td>
</tr>
<tr id="25D53E37DBFD828AEBD87CB2D564F52F" pageId="0" pageNumber="117">
<td id="94BFE57BBC7912EAA10EF83107D9AF20" colspan="6" pageId="0" pageNumber="117" rowspan="1">
<emphasis id="1E7C3574307ADA4C050C4646CD973072" bold="true" pageId="0" pageNumber="117">Left Mandible</emphasis>
</td>
</tr>
<tr id="A007175DF724AC1F0B0F3B9B8248F8C5" pageId="0" pageNumber="117">
<td id="DF93A7321A3F2190BEA39026C54A573F" colspan="1" pageId="0" pageNumber="117" rowspan="1">Palp segment 2 setae</td>
<td id="04F0723030EB9B0B6457A8AB0F281599" colspan="1" pageId="0" pageNumber="117" rowspan="1">8</td>
<td id="76C42A50C81222CB62C9E5A8F375FCC0" colspan="1" pageId="0" pageNumber="117" rowspan="1">7</td>
<td id="0233531526E7FD536FACA7D41871A499" colspan="1" pageId="0" pageNumber="117" rowspan="1">5</td>
<td id="A0A0DD5B5DFEA8038EB27F42EA31E7A0" colspan="1" pageId="0" pageNumber="117" rowspan="1">3</td>
<td id="259D00DA0FDF21348A6DA7C05337E285" colspan="1" pageId="0" pageNumber="117" rowspan="1">4</td>
</tr>
<tr id="9D3FE3A6AA0F9879B1990A1B50B09EFA" pageId="0" pageNumber="117">
<td id="2961C8DDD5FE1104CD60774A0CC56848" colspan="1" pageId="0" pageNumber="117" rowspan="1">E-setae</td>
<td id="BA8E9134C14F13437793B7B245B64F7B" colspan="1" pageId="0" pageNumber="117" rowspan="1">5</td>
<td id="179C513C7D5D462C31D9560D2CB19415" colspan="1" pageId="0" pageNumber="117" rowspan="1">5</td>
<td id="453BA26E1E0FABE0E154C29693937DB9" colspan="1" pageId="0" pageNumber="117" rowspan="1">5</td>
<td id="84EF1D607042488E9C4D888C3504B233" colspan="1" pageId="0" pageNumber="117" rowspan="1">4</td>
<td id="D1471CBB186C2DBD09CFD5351EAB3ABE" colspan="1" pageId="0" pageNumber="117" rowspan="1">3</td>
</tr>
<tr id="F34581D2AA323F6BED4C7CE4C9593A89" pageId="0" pageNumber="117">
<td id="0B3FB4E3D2208BADDABEFFBC3774CFC5" colspan="1" pageId="0" pageNumber="117" rowspan="1">D-setae</td>
<td id="ADBC8471FABAB8B179D1CD1D8602B7CE" colspan="1" pageId="0" pageNumber="117" rowspan="1">8</td>
<td id="8310D844EE8DE057310C2A10B908162B" colspan="1" pageId="0" pageNumber="117" rowspan="1">8</td>
<td id="AEB165EDD9042BC48C07AC07303B2D7C" colspan="1" pageId="0" pageNumber="117" rowspan="1">8</td>
<td id="218DB5E4B017A49832272ED51D2BF099" colspan="1" pageId="0" pageNumber="117" rowspan="1">8</td>
<td id="60EDD1A3812DFCBBCD6789AF8855116C" colspan="1" pageId="0" pageNumber="117" rowspan="1">5</td>
</tr>
<tr id="398AE44F53B780D9D32E0C8EEEE09A5A" pageId="0" pageNumber="117">
<td id="4CFDF0C85460050EBB3BFC6446E72E2E" colspan="6" pageId="0" pageNumber="117" rowspan="1">
<emphasis id="65A6814BC3DC5E55D30FD9CD4481C1ED" bold="true" pageId="0" pageNumber="117">Right Mantible</emphasis>
</td>
</tr>
<tr id="03A9DDE4B4B4E6B8868A3B580E0B030B" pageId="0" pageNumber="117">
<td id="A3B4E865FB8B9D5AA87C697A2A51A3C8" colspan="1" pageId="0" pageNumber="117" rowspan="1">Palp 2nd segment setae</td>
<td id="D69035A8D5CFAED10066B7CB579676CB" colspan="1" pageId="0" pageNumber="117" rowspan="1">8</td>
<td id="5EC95D48B2C2957724CACC2A26A39A47" colspan="1" pageId="0" pageNumber="117" rowspan="1">5</td>
<td id="2FE4C2143B4D8C977399A1D1A95BF21C" colspan="1" pageId="0" pageNumber="117" rowspan="1">5</td>
<td id="58EE46F22D1FC4A5CCBB1B924075D7E0" colspan="1" pageId="0" pageNumber="117" rowspan="1">4</td>
<td id="9A577CF2DDEAA1ED8CBE65AC986836BC" colspan="1" pageId="0" pageNumber="117" rowspan="1">4</td>
</tr>
<tr id="A576D8B943D3525D281D3D053198407B" pageId="0" pageNumber="117">
<td id="CA1966A3306E3253B1B66EB16456884D" colspan="1" pageId="0" pageNumber="117" rowspan="1">E-setae</td>
<td id="24F9462103CACBA78959880AAD579706" colspan="1" pageId="0" pageNumber="117" rowspan="1">5</td>
<td id="380CEE457D66B547698ABCCF338EFBE7" colspan="1" pageId="0" pageNumber="117" rowspan="1">5</td>
<td id="33F2DE8879273D1C0DE27E07E4A9743C" colspan="1" pageId="0" pageNumber="117" rowspan="1">4</td>
<td id="200EE82756949123B5A8199F399B0354" colspan="1" pageId="0" pageNumber="117" rowspan="1">4</td>
<td id="7CFEADAE80F70FD8D888FFFFB0E1BB9F" colspan="1" pageId="0" pageNumber="117" rowspan="1">3</td>
</tr>
<tr id="F3744F960F2D836E7D8BFA0E507F8903" pageId="0" pageNumber="117">
<td id="1FC67E01BA334E68C6AEAF620DE083A5" colspan="1" pageId="0" pageNumber="117" rowspan="1">D-setae</td>
<td id="909A50FA79D6FB9C6DC15995CE0831B7" colspan="1" pageId="0" pageNumber="117" rowspan="1">8</td>
<td id="9D8759CCEEFB36A4216566F6CCD2A61C" colspan="1" pageId="0" pageNumber="117" rowspan="1">8</td>
<td id="D0A5594A6369ED4AEB5A1C0F9E2B98DC" colspan="1" pageId="0" pageNumber="117" rowspan="1">8</td>
<td id="3C43E82E8F1D3D810B6DF8CBA86E6DC0" colspan="1" pageId="0" pageNumber="117" rowspan="1">7</td>
<td id="3C6027D28379DEE4FCB9C317EB699C9A" colspan="1" pageId="0" pageNumber="117" rowspan="1">5</td>
</tr>
<tr id="D18C42F314E0E9D7AD159E5223AF1604" pageId="0" pageNumber="117">
<td id="F9431F4EDB9C39C648CB8401DE2B0E9C" colspan="6" pageId="0" pageNumber="117" rowspan="1">
<emphasis id="260322306D0D2B195F07C774DC241212" bold="true" pageId="0" pageNumber="117">Maxilla 1</emphasis>
</td>
</tr>
<tr id="3F995359BE7F20D28D990B683C2F5E85" pageId="0" pageNumber="117">
<td id="626C7AC3BB3F110354EC35D138B24FFE" colspan="1" pageId="0" pageNumber="117" rowspan="1">Inner plate marginal setae</td>
<td id="EAC90F32F7B052752F4F7B4FF0F3131B" colspan="1" pageId="0" pageNumber="117" rowspan="1">unknown</td>
<td id="376DEC9B137D093E890291059DE4ED28" colspan="1" pageId="0" pageNumber="117" rowspan="1">4</td>
<td id="F4CED2CCCD3CDB22C09570CEC820272E" colspan="1" pageId="0" pageNumber="117" rowspan="1">4</td>
<td id="B0F29DDA020C07F0E98BE10590BF10DB" colspan="1" pageId="0" pageNumber="117" rowspan="1">3</td>
<td id="EDC30342D4059D42A5581E61E3059F25" colspan="1" pageId="0" pageNumber="117" rowspan="1">3</td>
</tr>
<tr id="E6325FCDFA8F404909249B967113E219" pageId="0" pageNumber="117">
<td id="C0A1CB90CFE5F39730F1C7C1B209EC48" colspan="1" pageId="0" pageNumber="117" rowspan="1">Palp subapical, apical setae</td>
<td id="F7593AAE07695200D30431CDB457CD9F" colspan="1" pageId="0" pageNumber="117" rowspan="1">unknown</td>
<td id="5616EAA5188BF314E0B62C547B26F2C1" colspan="1" pageId="0" pageNumber="117" rowspan="1">3, 4</td>
<td id="09D6BD4A525D10C404154AADCA07911A" colspan="1" pageId="0" pageNumber="117" rowspan="1">2, 4</td>
<td id="E68A6BCE3C25112993E5AD2FB66DC746" colspan="1" pageId="0" pageNumber="117" rowspan="1">2, 4</td>
<td id="344DAD685F7C4A9E69FC17D11286C08F" colspan="1" pageId="0" pageNumber="117" rowspan="1">4, 2</td>
</tr>
<tr id="0C3F866FD8CC337FE1FF3BB1C9877983" pageId="0" pageNumber="117">
<td id="8B2DF616134B325128FE71319180BAFF" colspan="6" pageId="0" pageNumber="117" rowspan="1">
<emphasis id="E69B220FE5AE6E2FF07C5DADEB3C802B" bold="true" pageId="0" pageNumber="117">Maxilla 2</emphasis>
</td>
</tr>
<tr id="BEEAF0BCDD3B52A389CCA4F844D84334" pageId="0" pageNumber="117">
<td id="DF96BBC0E503F56768ECD42B38019BCE" colspan="1" pageId="0" pageNumber="117" rowspan="1">Mx 2 inner plate facial setae</td>
<td id="FCDD660E9F572F9E1031286B491431CB" colspan="1" pageId="0" pageNumber="117" rowspan="1">unknown</td>
<td id="E444A9D2B7C707877077B1B940844550" colspan="1" pageId="0" pageNumber="117" rowspan="1">3</td>
<td id="03C2E3C72C42FAC6DBA696E9344BF56C" colspan="1" pageId="0" pageNumber="117" rowspan="1">2 or 3</td>
<td id="2190FADF0C2DD4AFC1BFED569B42FD23" colspan="1" pageId="0" pageNumber="117" rowspan="1">2 or 3</td>
<td id="9CE344447470FF4392C6ABFB50A3D638" colspan="1" pageId="0" pageNumber="117" rowspan="1">2</td>
</tr>
<tr id="96106D299FE5A8C24A2FBEEE421F102F" pageId="0" pageNumber="117">
<td id="EADE84E6BBA6AAFF5AB0257D9F40316E" colspan="6" pageId="0" pageNumber="117" rowspan="1">
<emphasis id="0A023B4FCF1ADCE0EBA25265C4FFA76B" bold="true" pageId="0" pageNumber="117">Ganthopod 1</emphasis>
</td>
</tr>
<tr id="3F1448566FC359B59B3FBAD4CCCB2240" pageId="0" pageNumber="117">
<td id="3E6CBB522939554E43789E419A69372F" colspan="1" pageId="0" pageNumber="117" rowspan="1">Ischium posterior setae</td>
<td id="66BDA18F79C346F89130963F806C4CDB" colspan="1" pageId="0" pageNumber="117" rowspan="1">4</td>
<td id="64600CF0047E90D6980A6ADD1727A33B" colspan="1" pageId="0" pageNumber="117" rowspan="1">4</td>
<td id="A09C04CA220F4C2CEF29FD37AE57C0A8" colspan="1" pageId="0" pageNumber="117" rowspan="1">2</td>
<td id="2D65545249418E9C4D82D73AD5F1AC6C" colspan="1" pageId="0" pageNumber="117" rowspan="1">3</td>
<td id="AC5B7B2FE10F4F720EA414E4D6CD26BF" colspan="1" pageId="0" pageNumber="117" rowspan="1">3</td>
</tr>
<tr id="06023E11EE070F66831AE85998157D28" pageId="0" pageNumber="117">
<td id="A4857668B4F9888C71BFEFAA5AB3B18B" colspan="1" pageId="0" pageNumber="117" rowspan="1">Carpus to propdus length</td>
<td id="4B6EC98CDD5CF752B9BBF030E82A6980" colspan="1" pageId="0" pageNumber="117" rowspan="1">40%</td>
<td id="942EE0FED5EF00B196EC50C34E9A82A5" colspan="1" pageId="0" pageNumber="117" rowspan="1">40%</td>
<td id="8D6596913E6BD50CA503D93807C23806" colspan="1" pageId="0" pageNumber="117" rowspan="1">43%</td>
<td id="F9354892E15AF2545F8CD2F5755C7BE2" colspan="1" pageId="0" pageNumber="117" rowspan="1">46%</td>
<td id="2D1F335E595ED36C87B0A380F1208851" colspan="1" pageId="0" pageNumber="117" rowspan="1">45%</td>
</tr>
<tr id="A42D3EE1D587EDE68A675F534B33B098" pageId="0" pageNumber="117">
<td id="EA6C27F4907D9A92BC68A06275F62C3C" colspan="1" pageId="0" pageNumber="117" rowspan="1">Carpus submarginal setae</td>
<td id="A6F04EA37693E882030D211B4A55EC10" colspan="1" pageId="0" pageNumber="117" rowspan="1">6</td>
<td id="2FF6BBBC271BE56C4BF139C7C42488D6" colspan="1" pageId="0" pageNumber="117" rowspan="1">5</td>
<td id="42CE213DF58B9997071763A1D2190233" colspan="1" pageId="0" pageNumber="117" rowspan="1">4</td>
<td id="2FA3B281CF3CD0BCB1C051FDD97A5406" colspan="1" pageId="0" pageNumber="117" rowspan="1">4</td>
<td id="0760B242F2E0CC80625107535997DF69" colspan="1" pageId="0" pageNumber="117" rowspan="1">5</td>
</tr>
<tr id="201F7DC5370228832FF48921AF8443E5" pageId="0" pageNumber="117">
<td id="9308ADD335128B0F42C37ACDD6D98CA9" colspan="1" pageId="0" pageNumber="117" rowspan="1">Propodus superior, inferior medial setae</td>
<td id="2C839F5A0EE638F053B4F523284E84C6" colspan="1" pageId="0" pageNumber="117" rowspan="1">4, 6</td>
<td id="A60C56C2ACA9460F659E650483819CC0" colspan="1" pageId="0" pageNumber="117" rowspan="1">2, 4</td>
<td id="E72D3A6141A7E7B88AF2195A053C0AFB" colspan="1" pageId="0" pageNumber="117" rowspan="1">3, 3</td>
<td id="7959919C9E6960BE9E176D836802B7CD" colspan="1" pageId="0" pageNumber="117" rowspan="1">3, 3</td>
<td id="50AF0F93570C1AE4EC892AA770D51547" colspan="1" pageId="0" pageNumber="117" rowspan="1">2, 4</td>
</tr>
<tr id="89E6F1770CD4C42FF712CFBD565039EE" pageId="0" pageNumber="117">
<td id="C3E661DC3A5BB21693CDED33E4A47506" colspan="1" pageId="0" pageNumber="117" rowspan="1">Palm inner, outer bifid setae</td>
<td id="49C4E3A1B9E45FADF0C9A6733D191A21" colspan="1" pageId="0" pageNumber="117" rowspan="1">7, 7</td>
<td id="583A8053DDA297423078B8C2740A710C" colspan="1" pageId="0" pageNumber="117" rowspan="1">7, 6</td>
<td id="67C557165A15BB0A9B431C2FFC898E5D" colspan="1" pageId="0" pageNumber="117" rowspan="1">5, 5</td>
<td id="8AFBF27C72BAABD4ABF47092C5265830" colspan="1" pageId="0" pageNumber="117" rowspan="1">4, 4</td>
<td id="9ED47CD6D1AEFFE38EBBEDCF258521DA" colspan="1" pageId="0" pageNumber="117" rowspan="1">5, 3</td>
</tr>
<tr id="40E883E83312975605187BD740B31898" pageId="0" pageNumber="117">
<td id="3DE97E0995461EDC02E5EC2A4C733291" colspan="6" pageId="0" pageNumber="117" rowspan="1">
<emphasis id="70D3AC1EEF3FBC7FC33D0BC441F070A9" bold="true" pageId="0" pageNumber="117">Ganthopod 2</emphasis>
</td>
</tr>
<tr id="D9A044F4869FF5CD861969C2D8850AEF" pageId="0" pageNumber="117">
<td id="08710FE3DF4DD9B9A1BD3C2A4F908EBA" colspan="1" pageId="0" pageNumber="117" rowspan="1">Coxal plate apical, facial setae</td>
<td id="D87DB5CBE6B29BE11566C19378840CB6" colspan="1" pageId="0" pageNumber="117" rowspan="1">4,1</td>
<td id="DC4718CE6CD61A2254D1DBAFA6A4346F" colspan="1" pageId="0" pageNumber="117" rowspan="1">4,1</td>
<td id="CACAF450360C38AE968063EAA622A220" colspan="1" pageId="0" pageNumber="117" rowspan="1">6,2</td>
<td id="B3BFE033EFDD9BF788DAE895A091B2D3" colspan="1" pageId="0" pageNumber="117" rowspan="1">4,1</td>
<td id="BF1C93461C6A0B3CF4981660E0D3EA24" colspan="1" pageId="0" pageNumber="117" rowspan="1">3,0</td>
</tr>
<tr id="C264EA63976DB1777D070F14BD08541B" pageId="0" pageNumber="117">
<td id="0482CABCB0988604611A53D589205501" colspan="1" pageId="0" pageNumber="117" rowspan="1">Ischium posterior setae</td>
<td id="1F376F3EF6F9613DDA63B64C59030825" colspan="1" pageId="0" pageNumber="117" rowspan="1">3</td>
<td id="0B4F1227B998A4E0794BF993858CA5ED" colspan="1" pageId="0" pageNumber="117" rowspan="1">4</td>
<td id="0B332D19F186EB08A84768C55BBF68A8" colspan="1" pageId="0" pageNumber="117" rowspan="1">2</td>
<td id="558BBB57C208E1F451C9AE7619F00F04" colspan="1" pageId="0" pageNumber="117" rowspan="1">2</td>
<td id="17F202FAFB9455FF7F2EA58C62C76A18" colspan="1" pageId="0" pageNumber="117" rowspan="1">3</td>
</tr>
<tr id="A80E6B3A16D06E7FA8F5B1319A635C65" pageId="0" pageNumber="117">
<td id="7F74CF28ECCB4D5F5AEF66DC35F54E3F" colspan="1" pageId="0" pageNumber="117" rowspan="1">Merus anterior margin robust seta</td>
<td id="ED368D720A1394B3C13FFAA3E2A26383" colspan="1" pageId="0" pageNumber="117" rowspan="1">1</td>
<td id="3E084F0ADA71D0A7AD58EB5E8C82AD89" colspan="1" pageId="0" pageNumber="117" rowspan="1">1</td>
<td id="7A471FC61362676FA451E141BB022BAE" colspan="1" pageId="0" pageNumber="117" rowspan="1">0</td>
<td id="BF63FD61B39D7C2E92338EFC6C12AC02" colspan="1" pageId="0" pageNumber="117" rowspan="1">0</td>
<td id="A86D7261A63624F4D027AA9093D984CB" colspan="1" pageId="0" pageNumber="117" rowspan="1">0</td>
</tr>
<tr id="2FCD2BD398346F3945A5C04CAACDAF0E" pageId="0" pageNumber="117">
<td id="F25FB9E6D7ABF1938F01B7BE407D0142" colspan="1" pageId="0" pageNumber="117" rowspan="1">Carpus to propdus length</td>
<td id="C27CD4C823A814793DFE5F875D6E28D1" colspan="1" pageId="0" pageNumber="117" rowspan="1">71%</td>
<td id="07AD905A2C84CF4A7B443A439AB98823" colspan="1" pageId="0" pageNumber="117" rowspan="1">64%</td>
<td id="7E11E29D2476520489E0141C3A694E47" colspan="1" pageId="0" pageNumber="117" rowspan="1">82%</td>
<td id="86AEC1E1286AF74DF0319D9B37C8F47A" colspan="1" pageId="0" pageNumber="117" rowspan="1">79%</td>
<td id="6038E92B7975E5C053D665D53B01CD1B" colspan="1" pageId="0" pageNumber="117" rowspan="1">84%</td>
</tr>
<tr id="4E0E024E777AFA996893131597DC76D9" pageId="0" pageNumber="117">
<td id="E9698EA50DD5F344AC42A68B0372D623" colspan="1" pageId="0" pageNumber="117" rowspan="1">Carpus rastellate setae</td>
<td id="97086887CB0E4E5F0C3AECB47920AE1A" colspan="1" pageId="0" pageNumber="117" rowspan="1">3</td>
<td id="61D420C12CFA0829FABDF975DB1611A2" colspan="1" pageId="0" pageNumber="117" rowspan="1">3</td>
<td id="4940CCC1C2D0297EED6EB4273744AC4C" colspan="1" pageId="0" pageNumber="117" rowspan="1">1</td>
<td id="686A133FA7061BD4B7CF05073D90522B" colspan="1" pageId="0" pageNumber="117" rowspan="1">3</td>
<td id="393D66887B0CD7DAB73F2ACE60875CDD" colspan="1" pageId="0" pageNumber="117" rowspan="1">0</td>
</tr>
<tr id="9D1064BE7F314FAC50131C634DFC1B49" pageId="0" pageNumber="117">
<td id="429147A6994AE8B9CDB09FDEE94B00F2" colspan="1" pageId="0" pageNumber="117" rowspan="1">Propodus superior, inferior medial setae</td>
<td id="76F026E91CE774D151BBED66B52650CD" colspan="1" pageId="0" pageNumber="117" rowspan="1">5, 5</td>
<td id="6E9990D2F02CDD0620D7498EA72DB6FA" colspan="1" pageId="0" pageNumber="117" rowspan="1">3, 4</td>
<td id="97F3A6F238A39C394579003FB3BBA3DC" colspan="1" pageId="0" pageNumber="117" rowspan="1">3, 4</td>
<td id="98EC7D3B138C3A9533D18E5EFC984268" colspan="1" pageId="0" pageNumber="117" rowspan="1">3, 3</td>
<td id="2E5CF36AFD93364AD3CC95A4718A92A6" colspan="1" pageId="0" pageNumber="117" rowspan="1">3, 3</td>
</tr>
<tr id="4268A1E675E2294D18A76B73DDEA88B3" pageId="0" pageNumber="117">
<td id="3589BC6118FE86E617B0B08FD2C10FDA" colspan="1" pageId="0" pageNumber="117" rowspan="1">Palm inner, outer bifid setae</td>
<td id="83DFFDDDE7EBFF55620D6502990F7878" colspan="1" pageId="0" pageNumber="117" rowspan="1">5, 5</td>
<td id="A6D8CAE9B07072E63FB8589C5C3226C9" colspan="1" pageId="0" pageNumber="117" rowspan="1">5, 6</td>
<td id="5176B836939861254C0BADD208AA4FDD" colspan="1" pageId="0" pageNumber="117" rowspan="1">3, 3</td>
<td id="EBAC8CDC77A8EC15DE8061F0EF993140" colspan="1" pageId="0" pageNumber="117" rowspan="1">3, 3</td>
<td id="7F98A8E8CB3DDA9C9C8672D61BC07723" colspan="1" pageId="0" pageNumber="117" rowspan="1">3, 4</td>
</tr>
<tr id="219B72085F48808F8B8E47BF68E9740F" pageId="0" pageNumber="117">
<td id="3A95097C460715FAFCB277CFDAE36469" colspan="1" pageId="0" pageNumber="117" rowspan="1">Dactylus inner setae</td>
<td id="01BF179D954FD2A2B594E65865993751" colspan="1" pageId="0" pageNumber="117" rowspan="1">1</td>
<td id="E1E9EA8DA2B115D9CC63721353E13DAC" colspan="1" pageId="0" pageNumber="117" rowspan="1">1</td>
<td id="7C277EFFD052FE1BC71321D3BEFDAF5B" colspan="1" pageId="0" pageNumber="117" rowspan="1">5</td>
<td id="781B4B1949841716B044CE5715022DF0" colspan="1" pageId="0" pageNumber="117" rowspan="1">4</td>
<td id="5873CCE01E7A300AFE62C64980974B65" colspan="1" pageId="0" pageNumber="117" rowspan="1">1</td>
</tr>
<tr id="184DEEAF0699FFAE5BF3CE9820B9A2A4" pageId="0" pageNumber="117">
<td id="6C8D4DB4B14AF5FA9A232EA863783EE7" colspan="6" pageId="0" pageNumber="117" rowspan="1">
<emphasis id="22654F69F5577F8F59012D1782E5E63A" bold="true" pageId="0" pageNumber="117">Pereopod 5</emphasis>
</td>
</tr>
<tr id="3A8ABC7C5F1F09993B097F11179A5AAB" pageId="0" pageNumber="117">
<td id="0EE448BC9642485DCBD3B163F093CCF0" colspan="1" pageId="0" pageNumber="117" rowspan="1">Coxal plate anterior apical setae</td>
<td id="06AF08D8D3829DDB9C177FD6E44A8A8E" colspan="1" pageId="0" pageNumber="117" rowspan="1">0</td>
<td id="1B59581F2806012D7768647D1B4468CD" colspan="1" pageId="0" pageNumber="117" rowspan="1">0</td>
<td id="B6981980A732728D02346600688A8CFB" colspan="1" pageId="0" pageNumber="117" rowspan="1">2</td>
<td id="407C90839CC4DB308108510DA0B4F9D6" colspan="1" pageId="0" pageNumber="117" rowspan="1">1 to 2</td>
<td id="0A8347FDD8F5335A042616758480B915" colspan="1" pageId="0" pageNumber="117" rowspan="1">2</td>
</tr>
<tr id="22EB9DA0C4ADEC57B4D2FCACC0298F71" pageId="0" pageNumber="117">
<td id="01C492C96034E7478BFBB0A4B4E5E89A" colspan="1" pageId="0" pageNumber="117" rowspan="1">Basis posterior serrations</td>
<td id="EF8D741686D4B4E44C2A07413E481004" colspan="1" pageId="0" pageNumber="117" rowspan="1">9</td>
<td id="7B53C2B3D25375A632C56DD66977DA76" colspan="1" pageId="0" pageNumber="117" rowspan="1">11</td>
<td id="54998B5F414F74FA77199D7620629BBF" colspan="1" pageId="0" pageNumber="117" rowspan="1">8</td>
<td id="638B00A053CE857257C822021F71A309" colspan="1" pageId="0" pageNumber="117" rowspan="1">8</td>
<td id="083EBE9DA9963713297FAB3928577F9F" colspan="1" pageId="0" pageNumber="117" rowspan="1">8</td>
</tr>
<tr id="FFFF98A04E1D3C2F9409EB404908FF27" pageId="0" pageNumber="117">
<td id="8DF33132C626B477ED39FB5A39EE2DA7" colspan="6" pageId="0" pageNumber="117" rowspan="1">
<emphasis id="2A82E751A35AA12D6B38DCCF4AE99EC9" bold="true" pageId="0" pageNumber="117">Pereopod 7</emphasis>
</td>
</tr>
<tr id="2B66D434DA6E789DBA20B13712B7EF47" pageId="0" pageNumber="117">
<td id="8C42AE9C58ADAA4D5B0CB2A818BFBC6B" colspan="1" pageId="0" pageNumber="117" rowspan="1">Coxal plate posterior apical setae</td>
<td id="3E1869AF5DF6E88814F2C97DFD7446FE" colspan="1" pageId="0" pageNumber="117" rowspan="1">4</td>
<td id="2A184EC6F0793733A9F9DAAE39B8962B" colspan="1" pageId="0" pageNumber="117" rowspan="1">3</td>
<td id="629022C257D41312F61C8EFF69FD3627" colspan="1" pageId="0" pageNumber="117" rowspan="1">4</td>
<td id="D2F0A0CCED4F7FEC964C606B3E0F9D93" colspan="1" pageId="0" pageNumber="117" rowspan="1">3</td>
<td id="1E3D3587B75E6F2F3969309F92B3DC04" colspan="1" pageId="0" pageNumber="117" rowspan="1">2</td>
</tr>
<tr id="41596F2136A5BC0BC6B08865EEA07318" pageId="0" pageNumber="117">
<td id="FCF807D68324ED209796E54F986E22F8" colspan="1" pageId="0" pageNumber="117" rowspan="1">Basis anterior setae</td>
<td id="013AC9B6585007351D0F2808BF7B024F" colspan="1" pageId="0" pageNumber="117" rowspan="1">8</td>
<td id="532BCBDBF72DFF9D6ACA4B1CBDF570F3" colspan="1" pageId="0" pageNumber="117" rowspan="1">7</td>
<td id="6AACB258D5C3C82B4CDD071C4E7753D8" colspan="1" pageId="0" pageNumber="117" rowspan="1">5</td>
<td id="1595AB7EEB7481F5BEEDEE5CABF49014" colspan="1" pageId="0" pageNumber="117" rowspan="1">5</td>
<td id="42F2DE620A101F39D5C5D7DAF6D3E9F5" colspan="1" pageId="0" pageNumber="117" rowspan="1">4</td>
</tr>
<tr id="77C1DF9F1BB1A01C6EEB25E2E67F4482" pageId="0" pageNumber="117">
<td id="D49C8AC94DB2212843B79290C2D1BCE0" colspan="6" pageId="0" pageNumber="117" rowspan="1">
<emphasis id="42135025295F25B24FD7B14DDD0D4476" bold="true" pageId="0" pageNumber="117">Epimera</emphasis>
</td>
</tr>
<tr id="8FC89F715B15E398D56541D4445E2BCB" pageId="0" pageNumber="117">
<td id="107B99E97109D8FBAAEFC66B3E058776" colspan="1" pageId="0" pageNumber="117" rowspan="1">Epimeron 2 ventral, posterior setae</td>
<td id="83919EDD2AE49FDAEA138C551DB5779B" colspan="1" pageId="0" pageNumber="117" rowspan="1">3, 2</td>
<td id="EAD897690BDDC6CE878B7AAF76C021CB" colspan="1" pageId="0" pageNumber="117" rowspan="1">unknown</td>
<td id="41911F35C11D7130D7D88938430F9C51" colspan="1" pageId="0" pageNumber="117" rowspan="1">3, 2</td>
<td id="E7EEC955967BA47585E6B5BA891B2127" colspan="1" pageId="0" pageNumber="117" rowspan="1">2, 2</td>
<td id="FC9BC99AA3E8738C0B42AC0C3E4F9E31" colspan="1" pageId="0" pageNumber="117" rowspan="1">2, 4</td>
</tr>
<tr id="CC0A3BF59614AC834BDA19F69F8A1F7C" pageId="0" pageNumber="117">
<td id="F3861CCFF52E47C19154D553FDB79F8E" colspan="6" pageId="0" pageNumber="117" rowspan="1">
<emphasis id="D44CC65894C09B9EC46E904F08A8810E" bold="true" pageId="0" pageNumber="117">Uropods</emphasis>
</td>
</tr>
<tr id="792E703F67FF42236CF7D49CF6A15457" pageId="0" pageNumber="117">
<td id="1F8AC0604E27B239B35963F1835692B3" colspan="1" pageId="0" pageNumber="117" rowspan="1">Uropod 1 peduncle outer, inner setae</td>
<td id="2A87F4478AACC89E58C1E936A538A854" colspan="1" pageId="0" pageNumber="117" rowspan="1">8, 1</td>
<td id="65E78C9E855BE04B5D591B280456FD4B" colspan="1" pageId="0" pageNumber="117" rowspan="1">10, 1</td>
<td id="23E6ACE31EB2A545D6270E6EB238AD12" colspan="1" pageId="0" pageNumber="117" rowspan="1">6, 1</td>
<td id="EFFB93025D0A7FEF467E7A4A5E6C4369" colspan="1" pageId="0" pageNumber="117" rowspan="1">9, 1</td>
<td id="465DA3A677655250AB17206F47B4248F" colspan="1" pageId="0" pageNumber="117" rowspan="1">8, 1</td>
</tr>
<tr id="FEB289614AE743FE474A5DB876063E31" pageId="0" pageNumber="117">
<td id="DE4ACE08185D848B2EEB26A65DD93D88" colspan="1" pageId="0" pageNumber="117" rowspan="1">Uropod 2 peduncle outer, inner setae</td>
<td id="14C126CFFF573FA179C457F9E963CE6E" colspan="1" pageId="0" pageNumber="117" rowspan="1">2, 2</td>
<td id="3BFC650E0CAA794407C850C708C1568E" colspan="1" pageId="0" pageNumber="117" rowspan="1">3, 1</td>
<td id="D2C59EC6DA6C2BCDBF5D40DF564236D3" colspan="1" pageId="0" pageNumber="117" rowspan="1">2, 1</td>
<td id="0F8BAB39AFC7CCA23961735E1039DCC2" colspan="1" pageId="0" pageNumber="117" rowspan="1">2, 1</td>
<td id="57068131C71A4FBBE0CD22C6A612D477" colspan="1" pageId="0" pageNumber="117" rowspan="1">2, 1</td>
</tr>
<tr id="98C55DD9E15BDD75A61A10F85A9B9B4C" pageId="0" pageNumber="117">
<td id="51DCDC3695B2B14F88C332E609AC2A86" colspan="1" pageId="0" pageNumber="117" rowspan="1">Uropod 2 outer ramus outer, inner setae</td>
<td id="FC494EA3DCB0037B63903CDEF3A7107F" colspan="1" pageId="0" pageNumber="117" rowspan="1">0, 0</td>
<td id="3FBABCE52EA672799A83C1BB8D867621" colspan="1" pageId="0" pageNumber="117" rowspan="1">1, 0</td>
<td id="4EC6E2AE2CFC1D15838CE468B8589DBA" colspan="1" pageId="0" pageNumber="117" rowspan="1">1, 0</td>
<td id="79DB6BB5368D8084A567507B6F753F74" colspan="1" pageId="0" pageNumber="117" rowspan="1">1, 0</td>
<td id="275573B077653974324B4BDAF0C8A6F6" colspan="1" pageId="0" pageNumber="117" rowspan="1">0, 0</td>
</tr>
<tr id="0E9401D1FC6AFA59763C38A59EEE9D01" pageId="0" pageNumber="117">
<td id="CA01D09F13C964384B3ACBE36113F70A" colspan="1" pageId="0" pageNumber="117" rowspan="1">Uropod 2 outer ramus apical setae</td>
<td id="924E3441CB37F2BF079192A510CA1E20" colspan="1" pageId="0" pageNumber="117" rowspan="1">4</td>
<td id="3B6F5A9DAD52D781C50D1CA59A40FDA5" colspan="1" pageId="0" pageNumber="117" rowspan="1">3</td>
<td id="5A1871DFF7BC738051DA4889CF9291AD" colspan="1" pageId="0" pageNumber="117" rowspan="1">4</td>
<td id="CEA97B3F63C3197A963CBB569428BBF9" colspan="1" pageId="0" pageNumber="117" rowspan="1">4</td>
<td id="D0EDB129439CE692BC8581C7F14A1DF2" colspan="1" pageId="0" pageNumber="117" rowspan="1">5</td>
</tr>
<tr id="FE0E711C2692CA54F2E33774D96A751A" pageId="0" pageNumber="117">
<td id="D8938ADA6EB24A2730EBD520DBC45EC2" colspan="1" pageId="0" pageNumber="117" rowspan="1">Uropod 2 inner ramus outer, inner setae</td>
<td id="A067AFBF42C427B00508748ACE5C7717" colspan="1" pageId="0" pageNumber="117" rowspan="1">2, 2</td>
<td id="67207E670FADD4B71027ED8F80767EC3" colspan="1" pageId="0" pageNumber="117" rowspan="1">2, 1</td>
<td id="CB1D4FE45C858A8B80472D90DC0691BC" colspan="1" pageId="0" pageNumber="117" rowspan="1">1, 1</td>
<td id="D2247D420CF0EE028CD28F5178D080F4" colspan="1" pageId="0" pageNumber="117" rowspan="1">1, 1</td>
<td id="F2994D094D93648CB1D946D988181013" colspan="1" pageId="0" pageNumber="117" rowspan="1">1, 1</td>
</tr>
<tr id="787B19FB4EB5A1F9299F29FB2C212F63" pageId="0" pageNumber="117">
<td id="19BF8D54A15D81D8EE09BB95C2EE4994" colspan="1" pageId="0" pageNumber="117" rowspan="1">Uropod 3 ramus setae</td>
<td id="BBB8F82D837AF2AC5DE264AC67F06414" colspan="1" pageId="0" pageNumber="117" rowspan="1">3</td>
<td id="2197F0D4A7F4944B7C0833EDF4942006" colspan="1" pageId="0" pageNumber="117" rowspan="1">4</td>
<td id="7A445A450FA9AE1EA0F4514B13E8A124" colspan="1" pageId="0" pageNumber="117" rowspan="1">4</td>
<td id="8A12E50758B44D2C612990AA91C7FF1A" colspan="1" pageId="0" pageNumber="117" rowspan="1">3</td>
<td id="7B0FEA1B818AAD135D8233E0EA3D2D19" colspan="1" pageId="0" pageNumber="117" rowspan="1">2</td>
</tr>
<tr id="34F539C57DBB8A5AAC78F978420C32B8" pageId="0" pageNumber="117">
<td id="7BF5195E22789FD28DB80AFD5D9D2867" colspan="1" pageId="0" pageNumber="117" rowspan="1">
<emphasis id="FBEE3471E11C5BDF23C03ABBDFB2A8FD" bold="true" pageId="0" pageNumber="117">Telson</emphasis>
apical robust setae
</td>
<td id="D3BDB6D47E5BCDDC0153463B15A0512E" colspan="1" pageId="0" pageNumber="117" rowspan="1">10</td>
<td id="2AC21E8674DDE70672EF7FF0DAF274F2" colspan="1" pageId="0" pageNumber="117" rowspan="1">10</td>
<td id="D8BF9572FA616D928EA057CBAABE6A24" colspan="1" pageId="0" pageNumber="117" rowspan="1">9</td>
<td id="2EABCC77AEB26BDB3EDEB7B3C8FFA6EF" colspan="1" pageId="0" pageNumber="117" rowspan="1">9</td>
<td id="7D2E6241D40278D4AF4C0E1F8AC779BA" colspan="1" pageId="0" pageNumber="117" rowspan="1">8</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection id="0BCED9DF9083F223BC7F2E4528224FBF" pageId="0" pageNumber="117" type="molecular diagnosis">
<paragraph id="D207061D214323934A2DE29F2EE55673" pageId="0" pageNumber="117">Molecular diagnosis.</paragraph>
<paragraph id="69E0980B16AF62C8AD6D2BB7A1DEC36A" pageId="0" pageNumber="117">
Average uncorrected pairwise genetic distance at the mitochondrial 
<emphasis id="61A43044AB3D5956AADEAA7E0C16356A" italics="true" pageId="0" pageNumber="117">co1</emphasis>
locus between 
<taxonomicName id="92EFE6BFBBEDE6B0E84B5692E73A3016" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. anacostensis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="anacostensis">
<emphasis id="48E7EBD1051A12B6E2E3A29F540F70BD" italics="true" pageId="0" pageNumber="117">S. anacostensis</emphasis>
</taxonomicName>
and the most closely related populations of 
<taxonomicName id="1CD075DC9F74B44E04F539E272E52106" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. t. subsp. potomacus" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="subSpecies" species="t." subSpecies="potomacus">
<emphasis id="E120AFB27D69639DB4E650148EF8BF65" italics="true" pageId="0" pageNumber="117">S. t. potomacus</emphasis>
</taxonomicName>
sampled at Caledon State Park is 6.5%, with 32 fixed mutations separating the two taxa. Between 
<taxonomicName id="E9D9DAE380EFFA2BCF3BF79D1E970C6C" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. anacostensis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="anacostensis">
<emphasis id="9CC5E97477835F4E9BD4CDEE0C81DBFA" italics="true" pageId="0" pageNumber="117">S. anacostensis</emphasis>
</taxonomicName>
and the closest 
<taxonomicName id="C2568A20C9F3A8CEAAA6AFAC038DFA6A" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. t. subsp. potomacus" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="subSpecies" species="t." subSpecies="potomacus">
<emphasis id="AA26D85B7DAB49EF6CE37432248ECAF2" italics="true" pageId="0" pageNumber="117">S. t. potomacus</emphasis>
</taxonomicName>
population (seepage spring near Malcolm X Ave in Anacostia Park; SP101), p-distance is 12.6%, with 67 fixed mutations. P-distance at the mitochondrial 
<emphasis id="8D818F67C31E2D68B6C558C9F78AC9E8" italics="true" pageId="0" pageNumber="117">16s</emphasis>
locus between 
<taxonomicName id="844F8B2A39274433E3120063DF3C2D0C" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. anacostensis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="anacostensis">
<emphasis id="E09DAAF50E6DF371619CE60EF115FE86" italics="true" pageId="0" pageNumber="117">S. anacostensis</emphasis>
</taxonomicName>
and the populations of 
<taxonomicName id="AD71801F74839D55468E7D7D27559122" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. t. subsp. potomacus" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="subSpecies" species="t." subSpecies="potomacus">
<emphasis id="DD09B23648F1E2F6FCB668B1D42AF767" italics="true" pageId="0" pageNumber="117">S. t. potomacus</emphasis>
</taxonomicName>
sampled at Caledon State Park is 2.3%, with eight fixed mutations. Nuclear loci exhibited low levels of variation among all 
<taxonomicName id="DFDFDA89F766675A85357F4F3A2D5822" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. tenuis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis id="240E747E235654376BE394A8828F9CCD" italics="true" pageId="0" pageNumber="117">S. tenuis</emphasis>
</taxonomicName>
species group taxa sampled; however, some diagnostic genetic variation was noted. Two fixed mutations in the 
<emphasis id="EE4611B9CEC3354E73F0EFE413356495" italics="true" pageId="0" pageNumber="117">h3</emphasis>
locus and one fixed mutation in the 
<emphasis id="DABE2A3E28C8C844A260ACD8393AAFE4" italics="true" pageId="0" pageNumber="117">18s</emphasis>
locus exist between 
<taxonomicName id="4D8C917795922510A8AE810CE7753B6F" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. anacostensis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="anacostensis">
<emphasis id="0DE242964C9DB3DCAA598638855780D2" italics="true" pageId="0" pageNumber="117">S. anacostensis</emphasis>
</taxonomicName>
and the closest 
<taxonomicName id="A3363A4A6BF17A080F23AB9D452C8695" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. t. subsp. potomacus" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="subSpecies" species="t." subSpecies="potomacus">
<emphasis id="4DEEF825C32D7F093A590BE4BEB67F59" italics="true" pageId="0" pageNumber="117">S. t. potomacus</emphasis>
</taxonomicName>
population (SP101).
</paragraph>
</subSubSection>
<subSubSection id="95CE0198F932E7A28FB32B73A560807C" pageId="0" pageNumber="117" type="distribution">
<paragraph id="D52B86B8A8F7EEF9FC0398BB1719E732" pageId="0" pageNumber="117">Geographical distribution.</paragraph>
<paragraph id="159F600F2DF183DA9C3D336BE51AA63B" pageId="0" pageNumber="117">
The species is known to date only from the type locality in Shepherd Parkway, which is a 1200-acre national park located on the southern bank of the Anacostia River just upstream from where the river flows into the Potomac River (Fig. 
<figureCitation id="92332602BF4C0F4E660A0B5CC6D4BCF3" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Distribution of Stygobromus anacostensis sp. nov. and other S. tenuis group species in the Washington D. C. area, USA" figureDoi="10.3897/subtbiol.48.112984.figure9" httpUri="https://binary.pensoft.net/fig/984748" pageId="0" pageNumber="117">9</figureCitation>
). Shepherd Parkway is part of National Capital Parks East (NACE).
</paragraph>
<caption id="7459451F2545ABDB0C954E3FC368B872" doi="10.3897/subtbiol.48.112984.figure9" httpUri="https://binary.pensoft.net/fig/984748" pageId="0" pageNumber="117" start="Figure 9" startId="F9">
<paragraph id="7059BB709BFAE8349BEEE169C8AF6424" pageId="0" pageNumber="117">
<emphasis id="ED8A39C956D680F324FDB3D17AF38532" bold="true" pageId="0" pageNumber="117">Figure 9.</emphasis>
Distribution of 
<taxonomicName id="F6CDB0EB4642F607D0948F1EB6B6AE55" authorityName="Cannizzaro, Sawicki, &amp; Niemiller" authorityYear="2024" class="Malacostraca" family="Crangonyctidae" genus="Stygobromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stygobromus anacostensis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="anacostensis">
<emphasis id="26E8EA0B1179E4CE2811B4569D6AA436" italics="true" pageId="0" pageNumber="117">Stygobromus anacostensis</emphasis>
</taxonomicName>
sp. nov. and other 
<taxonomicName id="6314260B5726F5BC80C0A06B230284A7" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. tenuis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis id="64EB96AFD73F246EE33296C21ACDD01D" italics="true" pageId="0" pageNumber="117">S. tenuis</emphasis>
</taxonomicName>
group species in the Washington D.C. area, USA
</paragraph>
</caption>
</subSubSection>
<subSubSection id="9280FDC42AF3DB8C0010C75344C9ABC0" pageId="0" pageNumber="117" type="habitat">
<paragraph id="0B10D491C5BC1DBA7A0339E133066992" pageId="0" pageNumber="117">Habitat and ecology.</paragraph>
<paragraph id="6FCC94514C6BAC644F3FFC04C03BC8EE" pageId="0" pageNumber="117">
Like other species of 
<taxonomicName id="D24C1CCEC92639E24F478C920C2C11A3" authorityName="Cope" authorityYear="1872" class="Malacostraca" family="Crangonyctidae" genus="Stygobromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stygobromus" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B9382DEA14CD052658B736CAAAEE6266" italics="true" pageId="0" pageNumber="117">Stygobromus</emphasis>
</taxonomicName>
, 
<taxonomicName id="085A30D757A62631854E3D5FA2F37F54" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. anacostensis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="anacostensis">
<emphasis id="EA2D3445A8C1657EC82D82D170A9E584" italics="true" pageId="0" pageNumber="117">S. anacostensis</emphasis>
</taxonomicName>
is a stygobiotic species occurring in groundwater habitats. All specimens have been collected from a seepage spring just off Malcolm X Avenue SE that marks the resurgence of hypotelminorheic groundwater at the surface (Fig. 
<figureCitation id="7ABF2674A1067B3E44A888C0CBAE9330" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. The type locality of S. anacostensis is a small hypotelminorheic seepage spring just off of Malcolm X Avenue, Shepherd Parkway, Washington, D. C., USA. Photograph by Jenna Keany." figureDoi="10.3897/subtbiol.48.112984.figure10" httpUri="https://binary.pensoft.net/fig/984749" pageId="0" pageNumber="117">10</figureCitation>
). Amphipods have been observed and collected from underneath moss-covered rocks, moss, and leaf litter on the small, 2-m high rockface as well as the small pool of the seepage spring. The seepage spring possesses water throughout most of the year Little is known regarding the ecology and life history currently. 
<taxonomicName id="F48DEDC4FEEBD8E98F6210DFA488390A" authorityName="Cannizzaro, Sawicki, &amp; Niemiller" authorityYear="2024" class="Malacostraca" family="Crangonyctidae" genus="Stygobromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stygobromus anacostensis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="anacostensis">
<emphasis id="A81CD72C32238159FFAD34BE00587313" italics="true" pageId="0" pageNumber="117">Stygobromus anacostensis</emphasis>
</taxonomicName>
co-occurs with the groundwater isopod 
<taxonomicName id="E6FD467DA0F02D54615494495461B2C7" class="Malacostraca" family="Asellidae" genus="Conasellus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Conasellus" order="Isopoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="9E7E8E7444DF0D489C88FD23696EBD98" italics="true" pageId="0" pageNumber="117">Conasellus</emphasis>
</taxonomicName>
(= 
<taxonomicName id="0BDEE2AA3B7E56AA87058E85E98666C2" authorityName="Packard" authorityYear="1871" class="Malacostraca" family="Asellidae" genus="Caecidotea" kingdom="Animalia" lsidName="Caecidotea" order="Isopoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="E0366B354F5CEF9FA82295E4F4731EA5" italics="true" pageId="0" pageNumber="117">Caecidotea</emphasis>
</taxonomicName>
) 
<taxonomicName id="8BFC701C21FFF4699FCC334EC9A6BB4B" authorityName="Holsinger" authorityYear="1978" class="Malacostraca" family="Crangonyctidae" genus="Stygobromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stygobromus kenki" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="kenki">
<emphasis id="8CB3A39446BD7E03834E6EEB5CAFBE38" italics="true" pageId="0" pageNumber="117">Stygobromus kenki</emphasis>
</taxonomicName>
(Bowman, 1967).
</paragraph>
<caption id="E73C0B1C8ADD367D5C1D9AB987BE6C54" doi="10.3897/subtbiol.48.112984.figure10" httpUri="https://binary.pensoft.net/fig/984749" pageId="0" pageNumber="117" start="Figure 10" startId="F10">
<paragraph id="D5434CCB398AE35E7F2FACEB282A6764" pageId="0" pageNumber="117">
<emphasis id="86FCF5435D02A8FAAC7C43A4BACB6E4F" bold="true" pageId="0" pageNumber="117">Figure 10.</emphasis>
The type locality of 
<taxonomicName id="DC4A6E4A6984581EE57E00EC97BC15B7" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. anacostensis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="anacostensis">
<emphasis id="F7F6070D461CDCF00FB93C9FEE4F289B" italics="true" pageId="0" pageNumber="117">S. anacostensis</emphasis>
</taxonomicName>
is a small hypotelminorheic seepage spring just off of Malcolm X Avenue, Shepherd Parkway, Washington, D.C., USA. Photograph by Jenna Keany.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="EA2C472D878A32F2E3C131F0781C39CC" pageId="0" pageNumber="117" type="conservation">
<paragraph id="67A58E818AFEC3B9A459B654ADF99BEE" pageId="0" pageNumber="117">Conservation.</paragraph>
<paragraph id="F58D4C62EAF41701F6594E0F9602BF6C" pageId="0" pageNumber="117">
<taxonomicName id="9DC1C52E7A83187A2E93396143656CA2" authorityName="Cannizzaro, Sawicki, &amp; Niemiller" authorityYear="2024" class="Malacostraca" family="Crangonyctidae" genus="Stygobromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stygobromus anacostensis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="anacostensis">
<emphasis id="2A2A5E73158E9185A12C868C9F04E262" italics="true" pageId="0" pageNumber="117">Stygobromus anacostensis</emphasis>
</taxonomicName>
is known only from the type locality. The NatureServe conservation rank calculated is Critically Imperiled (G1). Under IUCN Red List criteria, 
<taxonomicName id="2078B888540F042174AA4D409C1448AB" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. anacostensis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="anacostensis">
<emphasis id="90A3E5E4EEA40A6EDAD6BD4DE4C96F74" italics="true" pageId="0" pageNumber="117">S. anacostensis</emphasis>
</taxonomicName>
was assessed as Critically Endangered (CR B1) because of an extremely small EOO and AOO (known from a single small seep) in an urban area. Major threats to the species include increased risk of human intrusion and disturbance, habitat degradation, and pollution. The type locality population is offered some protection by occurring on National Park Service land, but the area controlled by the NPS is very narrow, and the site is highly vulnerable to road salt as well as any attempt to 
<normalizedToken id="301EB0160875B592547C7FE41EBE37B3" originalValue="“improve”">&quot;improve&quot;</normalizedToken>
the drainage in the vicinity of the sidewalk.
</paragraph>
</subSubSection>
<subSubSection id="3494738C91040C258DC48BDC55CEC4B3" pageId="0" pageNumber="117" type="genetic and phylogenetic analyses">
<paragraph id="522C8044BD40F52C372D1DBCD34D20C2" pageId="0" pageNumber="117">Genetic and phylogenetic analyses</paragraph>
<paragraph id="D0CC181600DE7A1A55323CD0851573B7" pageId="0" pageNumber="117">
We amplified in total 2,738 bp of five loci. Uncorrected mtDNA p-distance between 
<taxonomicName id="F9301279568256F9B7F0FFA638FBBC76" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. anacostensis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="anacostensis">
<emphasis id="E6AF217C7F050E625E5DC3E582B8D77E" italics="true" pageId="0" pageNumber="117">S. anacostensis</emphasis>
</taxonomicName>
and populations of 
<taxonomicName id="14192AFD4287248C56D499A031CE814B" baseAuthorityName="Holsinger" baseAuthorityYear="1967" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. tenuis subsp. potomacus" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="subSpecies" species="tenuis" subSpecies="potomacus">
<emphasis id="69DE53EB97E624165EB9048EBAEEA4D0" italics="true" pageId="0" pageNumber="117">S. tenuis potomacus</emphasis>
</taxonomicName>
at Caledon State Park (SP104 and SP105) was 6.5% and 12.6% between 
<taxonomicName id="6C24C9F08AC4879ACA41C5387FE214A9" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. anacostensis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="anacostensis">
<emphasis id="8CE39909626AD6BB1D0D304613CA0183" italics="true" pageId="0" pageNumber="117">S. anacostensis</emphasis>
</taxonomicName>
and the nearest 
<taxonomicName id="D617F33AC14A127962D7E1433F74DEE0" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. t. subsp. potomacus" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="subSpecies" species="t." subSpecies="potomacus">
<emphasis id="A592F78BBEB96389A100569256EAEFBF" italics="true" pageId="0" pageNumber="117">S. t. potomacus</emphasis>
</taxonomicName>
population sampled in Anacostia Park (SP101). Average uncorrected nucDNA p-distance was substantially lower, averaging 0.001 between 
<taxonomicName id="EFD5B65DA42E8E78F0B0E5F9E4867815" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. anacostensis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="anacostensis">
<emphasis id="FE3334DE4B022740A084E003FD9B5FF6" italics="true" pageId="0" pageNumber="117">S. anacostensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="92CF2013A218A2887FB6B8226BFD4443" baseAuthorityName="Holsinger" baseAuthorityYear="1967" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. tenuis subsp. potomacus" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="subSpecies" species="tenuis" subSpecies="potomacus">
<emphasis id="A42A38F2DDD669BA0B966467F9BF4C76" italics="true" pageId="0" pageNumber="117">S. tenuis potomacus</emphasis>
</taxonomicName>
at Caledon State Park (SP104 and SP105), and 0.004 between 
<taxonomicName id="2616F93A75117434A0CFB6636798637C" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. anacostensis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="anacostensis">
<emphasis id="72CD62353F49318A9B10A24B86A23BC5" italics="true" pageId="0" pageNumber="117">S. anacostensis</emphasis>
</taxonomicName>
and the nearest 
<taxonomicName id="5AA559C94A6AE16AFCC98B9BC8E8220F" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. t. subsp. potomacus" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="subSpecies" species="t." subSpecies="potomacus">
<emphasis id="B0777B9CD4062E2B5A80370C295FEECF" italics="true" pageId="0" pageNumber="117">S. t. potomacus</emphasis>
</taxonomicName>
population sampled in Anacostia Park (SP101).
</paragraph>
<paragraph id="1E09FDC2F62B15914AEF364D99C8BFDD" pageId="0" pageNumber="117">
The optimal substitution models for first, second, and third positions of 
<emphasis id="EBC89D1A9F82AAF08F81561A423239D7" italics="true" pageId="0" pageNumber="117">co1</emphasis>
were TrNef+I (
<bibRefCitation id="A08D6245EC120463D6FFD15D53E15121" author="Tamura, K" journalOrPublisher="Molecular Biology and Evolution" pageId="0" pageNumber="117" pagination="512 - 526" refId="B63" refString="Tamura, K, Nei, M, 1993. Estimation of the number of nucleotide substitutions in the control region of mitochondrial DNA in humans and chimpanzees. Molecular Biology and Evolution 10: 512 - 526" title="Estimation of the number of nucleotide substitutions in the control region of mitochondrial DNA in humans and chimpanzees." volume="10" year="1993">Tamura and Nei 1993</bibRefCitation>
), F81 (Felsenstein 1981), and K81+I (
<bibRefCitation id="59006F5E80E893A3E17027717BBE438C" DOI="https://doi.org/10.1073/pnas.78.1.454" author="Kimura, M" journalOrPublisher="Proceedings of the National Academy of Sciences" pageId="0" pageNumber="117" pagination="454 - 458" refId="B46" refString="Kimura, M, 1981. Estimation of evolutionary distances between homologous nucleotide sequences. Proceedings of the National Academy of Sciences 78: 454 - 458, DOI: https://doi.org/10.1073/pnas.78.1.454" title="Estimation of evolutionary distances between homologous nucleotide sequences." url="https://doi.org/10.1073/pnas.78.1.454" volume="78" year="1981">Kimura 1981</bibRefCitation>
), respectively. The optimal substitution model was HKY + I + G (
<bibRefCitation id="32E782D2855D4EDEC602702782608357" DOI="https://doi.org/10.1007/BF02101694" author="Hasegawa, M" journalOrPublisher="Journal of Molecular Evolution" pageId="0" pageNumber="117" pagination="160 - 174" refId="B26" refString="Hasegawa, M, Kishino, K, Yano, T, 1985. Dating the human-ape splitting by a molecular clock of mitochondrial DNA. Journal of Molecular Evolution 22: 160 - 174, DOI: https://doi.org/10.1007/BF02101694" title="Dating the human-ape splitting by a molecular clock of mitochondrial DNA." url="https://doi.org/10.1007/BF02101694" volume="22" year="1985">Hasegawa et al. 1985</bibRefCitation>
) for 
<emphasis id="02FCA049A59C52BA1230D5F457DA03F9" italics="true" pageId="0" pageNumber="117">16s</emphasis>
, K80 (
<bibRefCitation id="EBDB2F3872B3A2617709AF70764D2B99" DOI="https://doi.org/10.1007/BF01731581" author="Kimura, M" journalOrPublisher="Journal of Molecular Evolution" pageId="0" pageNumber="117" pagination="111 - 120" refId="B45" refString="Kimura, M, 1980. A simple method for estimating evolutionary rate of base substitutions through comparative studies of nucleotide sequences. Journal of Molecular Evolution 16: 111 - 120, DOI: https://doi.org/10.1007/BF01731581" title="A simple method for estimating evolutionary rate of base substitutions through comparative studies of nucleotide sequences." url="https://doi.org/10.1007/BF01731581" volume="16" year="1980">Kimura 1980</bibRefCitation>
) for 
<emphasis id="48CDB7BDC5BCB0BBA19F1067C7F75D04" italics="true" pageId="0" pageNumber="117">18s</emphasis>
, TIM2+I for 
<emphasis id="356C1FA02BE8D946D599B7F061ABFD31" italics="true" pageId="0" pageNumber="117">28s</emphasis>
, and JC (Jukes and Cantor 1980) for 
<emphasis id="5974EA93898A9238E372B648CEDE6E97" italics="true" pageId="0" pageNumber="117">h3</emphasis>
. Phylogenetic tree topologies obtained for ML and Bayesian inference were highly similar. Phylogenetic trees reconstructed using both ML and Bayesian inference for the mtDNA (
<emphasis id="86278A011271ECC31DE6026760CEBBD9" italics="true" pageId="0" pageNumber="117">co1</emphasis>
+ 
<emphasis id="21223A1C6E1DEBC504867C8DEE5E92A2" italics="true" pageId="0" pageNumber="117">16s</emphasis>
; Fig. 
<figureCitation id="4E9542EA5EA5543073808842B6AABEAD" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Maximum-likelihood phylogeny and species delimitations of Stygobromus anacostensis and other S. tenuis species group taxa for the mtDNA dataset (co 1 + 16 s loci). Asterisk represents bootstrap node support greater than 90. Colored bars represented hypothesized MOTU groupings (i. e., species) based on corresponding delimitation analyses." figureDoi="10.3897/subtbiol.48.112984.figure11" httpUri="https://binary.pensoft.net/fig/984750" pageId="0" pageNumber="117">11</figureCitation>
) and mtDNA+nucDNA datasets (Fig. 
<figureCitation id="4B20B57A8964DDE14B187D78A4808C46" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Maximum-likelihood phylogenies of of Stygobromus anacostensis and other S. tenuis species group taxa for the (A) mtDNA + nucDNA dataset (co 1 + 16 s + 18 s + 28 s + h 3 loci) and (B) nucDNA dataset (18 s + 28 s + h 3 loci). Asterisk represents bootstrap node support greater than 90." figureDoi="10.3897/subtbiol.48.112984.figure12" httpUri="https://binary.pensoft.net/fig/984751" pageId="0" pageNumber="117">12A</figureCitation>
) delimited individuals of 
<taxonomicName id="C954F9C38660DDD212DEA078E1481F34" authorityName="Cannizzaro, Sawicki, &amp; Niemiller" authorityYear="2024" class="Malacostraca" family="Crangonyctidae" genus="Stygobromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stygobromus anacostensis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="anacostensis">
<emphasis id="6BDBE6A110B516D3AB54ADDA052C58D5" italics="true" pageId="0" pageNumber="117">Stygobromus anacostensis</emphasis>
</taxonomicName>
from the type locality as distinct from other populations of the 
<taxonomicName id="87DBF80436523C10542CEC5AFE95705C" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. tenuis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis id="53F29F3556D54E2AF83BB8ADACDC2FDA" italics="true" pageId="0" pageNumber="117">S. tenuis</emphasis>
</taxonomicName>
species group sampled with high bootstrap support. Populations of 
<taxonomicName id="525F1115A60DF136DCE09C812F992C9C" baseAuthorityName="Holsinger" baseAuthorityYear="1967" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. tenuis subsp. potomacus" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="subSpecies" species="tenuis" subSpecies="potomacus">
<emphasis id="49BAB79116A4F8D30A8F704414BBFB5C" italics="true" pageId="0" pageNumber="117">S. tenuis potomacus</emphasis>
</taxonomicName>
did not form a monophyletic group for any dataset. Nuclear loci exhibited low levels of variation among all 
<taxonomicName id="33B614286D8F41CF5D4E3D59C777A15A" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. tenuis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis id="46B04DEAB3C533EF15780ECEEA110662" italics="true" pageId="0" pageNumber="117">S. tenuis</emphasis>
</taxonomicName>
species group taxa sampled (Figs 
<figureCitation id="CCCE2B13FAA01A41BAA55D2F9217DC0C" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Maximum-likelihood phylogenies of of Stygobromus anacostensis and other S. tenuis species group taxa for the (A) mtDNA + nucDNA dataset (co 1 + 16 s + 18 s + 28 s + h 3 loci) and (B) nucDNA dataset (18 s + 28 s + h 3 loci). Asterisk represents bootstrap node support greater than 90." figureDoi="10.3897/subtbiol.48.112984.figure12" httpUri="https://binary.pensoft.net/fig/984751" pageId="0" pageNumber="117">12B</figureCitation>
, 
<figureCitation id="56DBD6E1D5FD8086EA2A01F7FC7DDD36" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Median joining networks for nuclear loci (18 s, 28 s, and h 3) generated in PopART v 1.7." figureDoi="10.3897/subtbiol.48.112984.figure13" httpUri="https://binary.pensoft.net/fig/984752" pageId="0" pageNumber="117">13</figureCitation>
). Two fixed mutations in the 
<emphasis id="86A136536F2302820F24B2922331A3AB" italics="true" pageId="0" pageNumber="117">h3</emphasis>
locus and one fixed mutation in the 
<emphasis id="77E0AED73DE03DA2BEE1BB0A52509119" italics="true" pageId="0" pageNumber="117">18s</emphasis>
locus exist between 
<taxonomicName id="37C875EB2D79C73EDA1E85AA42A559ED" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. anacostensis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="anacostensis">
<emphasis id="E1EA548FAA8429F89669BD1BBD12A4FA" italics="true" pageId="0" pageNumber="117">S. anacostensis</emphasis>
</taxonomicName>
and the closest 
<taxonomicName id="684A336E9D6E1FED713046234E9260AF" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. t. subsp. potomacus" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="subSpecies" species="t." subSpecies="potomacus">
<emphasis id="0C6656ACB0E96614723B240BD5A6AB92" italics="true" pageId="0" pageNumber="117">S. t. potomacus</emphasis>
</taxonomicName>
population (SP101).
</paragraph>
<caption id="20E6FD12206E2CF05B7B32DEDA0D8B47" doi="10.3897/subtbiol.48.112984.figure11" httpUri="https://binary.pensoft.net/fig/984750" pageId="0" pageNumber="117" start="Figure 11" startId="F11">
<paragraph id="E50B767644CA294C34B1A6B7E2794188" pageId="0" pageNumber="117">
<emphasis id="BDCE7463935B92F9544BFFC8F338353C" bold="true" pageId="0" pageNumber="117">Figure 11.</emphasis>
Maximum-likelihood phylogeny and species delimitations of 
<taxonomicName id="D9D5A7104B0F617671973416C05A5EAB" authorityName="Cannizzaro, Sawicki, &amp; Niemiller" authorityYear="2024" class="Malacostraca" family="Crangonyctidae" genus="Stygobromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stygobromus anacostensis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="anacostensis">
<emphasis id="C0144F9256E59CE188F65ABD231BB1BD" italics="true" pageId="0" pageNumber="117">Stygobromus anacostensis</emphasis>
</taxonomicName>
and other 
<taxonomicName id="AED11F62E4640D9A9BE54D089D2EA4BB" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. tenuis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis id="D6195E1F62AE5E3BF54863C46905259E" italics="true" pageId="0" pageNumber="117">S. tenuis</emphasis>
</taxonomicName>
species group taxa for the mtDNA dataset (
<emphasis id="A229898626025F2037B9124AAD814EDA" italics="true" pageId="0" pageNumber="117">co1</emphasis>
+ 
<emphasis id="DECFB1C3C6A63EA3070CB0239518F016" italics="true" pageId="0" pageNumber="117">16s</emphasis>
loci). Asterisk represents bootstrap node support greater than 90. Colored bars represented hypothesized MOTU groupings (i.e., species) based on corresponding delimitation analyses.
</paragraph>
</caption>
<caption id="3988F900284CA57BEC9F9CDC8EF9055E" doi="10.3897/subtbiol.48.112984.figure12" httpUri="https://binary.pensoft.net/fig/984751" pageId="0" pageNumber="117" start="Figure 12" startId="F12">
<paragraph id="F6D448617953BECEC918A86BE40F2C22" pageId="0" pageNumber="117">
<emphasis id="BFC29B57B987F4B912072DF8639E0495" bold="true" pageId="0" pageNumber="117">Figure 12.</emphasis>
Maximum-likelihood phylogenies of of 
<taxonomicName id="9B909358651B40E723DF3F255D51B053" authorityName="Cannizzaro, Sawicki, &amp; Niemiller" authorityYear="2024" class="Malacostraca" family="Crangonyctidae" genus="Stygobromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stygobromus anacostensis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="anacostensis">
<emphasis id="167F43D98F7F72DD774A6E89A09BDF1E" italics="true" pageId="0" pageNumber="117">Stygobromus anacostensis</emphasis>
</taxonomicName>
and other 
<taxonomicName id="0E9B8D3A8E619F320E168AE76188A14A" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. tenuis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis id="1CBAFEB19B7154A90C10C3A18AC99183" italics="true" pageId="0" pageNumber="117">S. tenuis</emphasis>
</taxonomicName>
species group taxa for the (
<emphasis id="03C493D8CDED161C036151CEE876D48F" bold="true" pageId="0" pageNumber="117">A</emphasis>
) mtDNA+nucDNA dataset (
<emphasis id="DA8262687884835CF78BFF963665B558" italics="true" pageId="0" pageNumber="117">co1</emphasis>
+ 
<emphasis id="3A100DE75F81E9828E947B0029E8073F" italics="true" pageId="0" pageNumber="117">16s</emphasis>
+ 
<emphasis id="F5522B3BFC0FB0D93C0A48574DD2275F" italics="true" pageId="0" pageNumber="117">18s</emphasis>
+ 
<emphasis id="23871777CA64E9B1D637442D90693C6D" italics="true" pageId="0" pageNumber="117">28s</emphasis>
+ 
<emphasis id="9C752AAC2DCCC5295F0151095CC2B009" italics="true" pageId="0" pageNumber="117">h3</emphasis>
loci) and (
<emphasis id="6EB438D0468AFEB6FC7F9B519BA2A1DB" bold="true" pageId="0" pageNumber="117">B</emphasis>
) nucDNA dataset (
<emphasis id="DBE92E5BA00B8E7D672E40AB19A3C563" italics="true" pageId="0" pageNumber="117">18s</emphasis>
+ 
<emphasis id="5CDFEF6C1E5817F02DDFD54E148FAA76" italics="true" pageId="0" pageNumber="117">28s</emphasis>
+ 
<emphasis id="ADC8E191D9C1CB5C51D356FF3E13A7B3" italics="true" pageId="0" pageNumber="117">h3</emphasis>
loci). Asterisk represents bootstrap node support greater than 90.
</paragraph>
</caption>
<caption id="4649F7E7EB4D42009B4D26F39D174F5D" doi="10.3897/subtbiol.48.112984.figure13" httpUri="https://binary.pensoft.net/fig/984752" pageId="0" pageNumber="117" start="Figure 13" startId="F13">
<paragraph id="E92CB197A822DC50E78B9AAD56587D82" pageId="0" pageNumber="117">
<emphasis id="25D45763DC6E32173265BAEBBC2DC076" bold="true" pageId="0" pageNumber="117">Figure 13.</emphasis>
Median joining networks for nuclear loci (
<emphasis id="B67893C4F47AA9A3649576197ADAD7FB" italics="true" pageId="0" pageNumber="117">18s</emphasis>
, 
<emphasis id="A082E5EB41F962FAB351B35D7D21D08A" italics="true" pageId="0" pageNumber="117">28s</emphasis>
, and 
<emphasis id="D68285AB74C5CACB36A42E328EEF1DA0" italics="true" pageId="0" pageNumber="117">h3</emphasis>
) generated in PopART v1.7.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="DEDFF9F22882E1C7B1EAF6A77F12C0FE" pageId="0" pageNumber="117" type="species delimitation">
<paragraph id="A8AC41667EEF7B7EA4B80DD55A158161" pageId="0" pageNumber="117">Species delimitation</paragraph>
<paragraph id="44304ED06B5AF674769941CAE990B30F" pageId="0" pageNumber="117">
For the mtDNA dataset (Fig. 
<figureCitation id="A5EFA5F728CA5C724E7F0520B88409FE" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Maximum-likelihood phylogeny and species delimitations of Stygobromus anacostensis and other S. tenuis species group taxa for the mtDNA dataset (co 1 + 16 s loci). Asterisk represents bootstrap node support greater than 90. Colored bars represented hypothesized MOTU groupings (i. e., species) based on corresponding delimitation analyses." figureDoi="10.3897/subtbiol.48.112984.figure11" httpUri="https://binary.pensoft.net/fig/984750" pageId="0" pageNumber="117">11</figureCitation>
), the ABGD approach resulted in nine MOTUs, with convergence of initial and recursive partitions at prior intraspecific divergence (P) = 0.028, which remained stable until P = 0.0359. The PTP approach yielded the same MOTU delimitations. All 
<taxonomicName id="A534F56BD04F7C2D03296B88D9B455CC" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. anacostensis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="anacostensis">
<emphasis id="F12C1EB334B5D5AAA1118BC281C85CA7" italics="true" pageId="0" pageNumber="117">S. anacostensis</emphasis>
</taxonomicName>
samples formed a MOTU, while several 
<taxonomicName id="491191F2744C5B95B7F6C258E008F20F" baseAuthorityName="Holsinger" baseAuthorityYear="1967" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. tenuis subsp. potomacus" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="subSpecies" species="tenuis" subSpecies="potomacus">
<emphasis id="35DEED5C2F854B22745B99DA92C8EC5D" italics="true" pageId="0" pageNumber="117">S. tenuis potomacus</emphasis>
</taxonomicName>
populations were delimited as distinct MOTUs. The mPTP approach estimated seven MOTUs, with highly similar designations to the ABGD delimitations. 
<taxonomicName id="CBAB10BBD9FEE704BE57CB7EB3F0DDB5" authorityName="Cannizzaro, Sawicki, &amp; Niemiller" authorityYear="2024" class="Malacostraca" family="Crangonyctidae" genus="Stygobromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stygobromus anacostensis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="anacostensis">
<emphasis id="E3EFD1A19A72502E1EEEA8370E37E478" italics="true" pageId="0" pageNumber="117">Stygobromus anacostensis</emphasis>
</taxonomicName>
individuals were grouped as a single MOTU, as were several 
<taxonomicName id="11F02ED512DF18BC82BB50BB2A6AD840" baseAuthorityName="Holsinger" baseAuthorityYear="1967" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. tenuis subsp. potomacus" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="subSpecies" species="tenuis" subSpecies="potomacus">
<emphasis id="17AEFDFB967665EC8D4397D9B3ADC057" italics="true" pageId="0" pageNumber="117">S. tenuis potomacus</emphasis>
</taxonomicName>
populations. 
<taxonomicName id="5FFA23D2C80175E59EE43177941A1EED" baseAuthorityName="Holsinger" baseAuthorityYear="1967" class="Malacostraca" family="Crangonyctidae" genus="Stygobromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stygobromus allegheniensis" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="allegheniensis">
<emphasis id="AC2474AEA60A5611CDFDF8C9977CD0FE" italics="true" pageId="0" pageNumber="117">Stygobromus allegheniensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="74667C21C96CCEC85C75B485B6991705" class="Crustacea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="S. hayi" order="Amphipoda" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="hayi">
<emphasis id="0158D2F10C7B96BA737578AAE966113C" italics="true" pageId="0" pageNumber="117">S. hayi</emphasis>
</taxonomicName>
were grouped into a single MOTU.
</paragraph>
</subSubSection>
</treatment>
</document>