treatments-xml/data/3D/7B/FE/3D7BFE0AFFA3FFCBFE6EFE87FD50FD8E.xml

<document id="E5DD8CC94C1701382AC05D8F47ABA3E2" ID-CLB-Dataset="7699" ID-DOI="10.5852/ejt.2021.739.1271" ID-GBIF-Dataset="99f6713d-6ef6-4bee-905b-b987df8832d4" ID-ISSN="2118-9773" ID-Zenodo-Dep="4610565" ID-ZooBank="81383BD5-6040-4156-BD09-A2704E5D9DA7" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615988829382" checkinUser="plazi" docAuthor="Wang, Wendy Y., Yong, Gordon W. J. &amp; Jaitrong, Weeyawat" docDate="2021" docId="3D7BFE0AFFA3FFCBFE6EFE87FD50FD8E" docLanguage="en" docName="ejt-739_117-157.pdf" docOrigin="European Journal of Taxonomy 739 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.739.1271" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Rhopalomastix robusta Wang &amp; Yong &amp; Jaitrong 2021, sp. nov." docType="treatment" docUuid="7FE9D47B-988F-40BB-9AEE-C208F52D6159" docUuidSource="ZooBank" docVersion="9" lastPageNumber="154" masterDocId="C1428672FF82FFEDFF9BFF8FFFEBFFB3" masterDocTitle="Revision of the elusive ant genus Rhopalomastix (Hymenoptera, Formicidae, Myrmicinae) in Thailand based on morphology and DNA barcodes, with descriptions of three new species" masterLastPageNumber="157" masterPageNumber="117" pageNumber="150" updateTime="1698933474431" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="6F3936E0CBCFE4310BC941CD891DEC29">Revision of the elusive ant genus Rhopalomastix (Hymenoptera, Formicidae, Myrmicinae) in Thailand based on morphology and DNA barcodes, with descriptions of three new species</mods:title>
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<mods:namePart id="77ACD676096AC7C11BD145281B9A8686">Wang, Wendy Y.</mods:namePart>
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<mods:affiliation id="E02FFE72B5F613B69926DEEBB1696B98">Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, Singapore 117377. Department of Biological Sciences, Faculty of Science, National University of Singapore, 16 Science Drive 4, Singapore 117558. Natural History Museum, National Science Museum, Technopolis, Khlong 5, Khlong Luang, Pathum Thani, 12120 Thailand.</mods:affiliation>
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<taxonomicName id="72D2349FFFA3FFCCFE6EFE87FC35FE90" authority="Wang &amp; Jaitrong" authorityName="Wang &amp; Yong &amp; Jaitrong" authorityYear="2021" box="[501,990,264,291]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="150" phylum="Arthropoda" rank="species" species="robusta" status="sp. nov.">
<emphasis id="87A6930EFFA3FFCCFE6EFE87FCE5FE90" bold="true" box="[501,782,264,291]" italics="true" pageId="33" pageNumber="150">Rhopalomastix robusta</emphasis>
Wang &amp; Jaitrong
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<paragraph id="B56D4F1CFFA3FFCCFEEEFEC0FB54FEDA" blockId="33.[366,1222,264,362]" box="[373,1215,335,362]" pageId="33" pageNumber="150">
<figureCitation id="2DE95399FFA3FFCCFEEEFEC0FE24FEDA" box="[373,463,335,361]" captionStart="Fig" captionStartId="5.[189,232,1816,1842]" captionTargetBox="[227,1361,973,1778]" captionTargetId="figure-170@5.[227,1361,973,1778]" captionTargetPageId="5" captionText="Fig. 2. Closeups of external mandibular surface for workers of 5 Thai species of Rhopalomastix Forel, 1900 (ZRC); white arrows indicate distinct striations on the external mandibular face. a. R. impithuksai Wang &amp; Jaitrong sp. nov., paratype (ZRC_ENT00007608). b. R. robusta Wang &amp; Jaitrong sp. nov., paratype (ZRC_ENT00013949). c. R. javana Wheeler, 1929, non-type (ZRC_ENT00000932). d. R. johorensis Wheeler, 1929, non-type (ZRC_ENT00007874). e. R. parva Wang &amp; Jaitrong sp. nov., paratype (ZRC_ENT00013948)." figureDoi="http://doi.org/10.5281/zenodo.4610567" httpUri="https://zenodo.org/record/4610567/files/figure.png" pageId="33" pageNumber="150">Figs 2b</figureCitation>
, 
<figureCitation id="2DE95399FFA3FFCCFE40FEC0FE13FEDA" box="[475,504,335,361]" captionStart="Fig" captionStartId="34.[189,232,1917,1943]" captionTargetBox="[227,1361,977,1884]" captionTargetId="figure-275@34.[227,1361,977,1884]" captionTargetPageId="34" captionText="Fig. 13. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, average-sized worker (ZRC_ ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Close-up of clypeus in profile. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610589" httpUri="https://zenodo.org/record/4610589/files/figure.png" pageId="33" pageNumber="150">13</figureCitation>
(average worker), 14 (large worker), 15 (small worker), 16a
</paragraph>
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<emphasis id="87A6930EFFA3FFCCFF26FE1FFEDDFE19" bold="true" box="[189,310,400,426]" pageId="33" pageNumber="150">Diagnosis</emphasis>
</heading>
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<paragraph id="B56D4F1CFFA3FFCCFF71FE30FEA7FE6A" blockId="33.[189,1400,447,1037]" box="[234,332,447,473]" pageId="33" pageNumber="150">
<emphasis id="87A6930EFFA3FFCCFF71FE30FEA7FE6A" bold="true" box="[234,332,447,473]" pageId="33" pageNumber="150">Worker</emphasis>
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<paragraph id="B56D4F1CFFA3FFCCFF26FE6DFBFCFBBF" blockId="33.[189,1400,447,1037]" pageId="33" pageNumber="150">
Relatively large size (small worker TL&gt; 
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; large worker TL&gt; 
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) with robust habitus; workers monomorphic with broad intranidal size variation. Head subsquarish, as long as wide or slightly wider than long; in small workers slightly longer than wide; distinctly narrowed anteriorly; narrow median furrow between frontal lobes deep and long, extending till or past transverse midline of head, narrower or constricted shortly before midlength; posterolateral corners roundly convex and laterally expanded forming somewhat rounded vertexal lobes (
<figureCitation id="2DE95399FFA3FFCCFD5BFD1DFCC2FD1E" box="[704,809,658,685]" captionStart="Fig" captionStartId="34.[189,232,1917,1943]" captionTargetBox="[227,1361,977,1884]" captionTargetId="figure-275@34.[227,1361,977,1884]" captionTargetPageId="34" captionText="Fig. 13. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, average-sized worker (ZRC_ ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Close-up of clypeus in profile. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610589" httpUri="https://zenodo.org/record/4610589/files/figure.png" pageId="33" pageNumber="150">Figs 13a</figureCitation>
, 
<figureCitation id="2DE95399FFA3FFCCFCAEFD1CFC89FD1E" box="[821,866,659,685]" captionStart="Fig" captionStartId="35.[189,232,1945,1971]" captionTargetBox="[227,1361,995,1902]" captionTargetId="figure-258@35.[227,1361,995,1902]" captionTargetPageId="35" captionText="Fig. 14. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, large worker (ZRC_ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610591" httpUri="https://zenodo.org/record/4610591/files/figure.png" pageId="33" pageNumber="150">14a</figureCitation>
, 
<figureCitation id="2DE95399FFA3FFCCFCF4FD1DFC77FD1E" box="[879,924,658,685]" captionStart="Fig" captionStartId="36.[189,232,1945,1971]" captionTargetBox="[227,1361,701,1911]" captionTargetId="figure-150@36.[227,1361,701,1911]" captionTargetPageId="36" captionText="Fig. 15. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, small worker (ZRC_ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610593" httpUri="https://zenodo.org/record/4610593/files/figure.png" pageId="33" pageNumber="150">15a</figureCitation>
). Anterior clypeal face in profile weakly and evenly convex, sometimes almost flat (
<figureCitation id="2DE95399FFA3FFCCFD58FD39FCC2FD63" box="[707,809,694,720]" captionStart="Fig" captionStartId="34.[189,232,1917,1943]" captionTargetBox="[227,1361,977,1884]" captionTargetId="figure-275@34.[227,1361,977,1884]" captionTargetPageId="34" captionText="Fig. 13. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, average-sized worker (ZRC_ ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Close-up of clypeus in profile. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610589" httpUri="https://zenodo.org/record/4610589/files/figure.png" pageId="33" pageNumber="150">Fig. 13c</figureCitation>
). In dorsal view, anterior margin of pronotal disc broadly convex with bluntly angulate median point, humeral corners obtusely angulate, lateral margins with distinct inflexions differentiating pronotum, mesonotum and propodeum; posterolateral corners of propodeum indistinct and rounded; posterior propodeal margin indistinctly concave (
<figureCitation id="2DE95399FFA3FFCCFB50FC90FADDFC8A" box="[1227,1334,799,825]" captionStart="Fig" captionStartId="34.[189,232,1917,1943]" captionTargetBox="[227,1361,977,1884]" captionTargetId="figure-275@34.[227,1361,977,1884]" captionTargetPageId="34" captionText="Fig. 13. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, average-sized worker (ZRC_ ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Close-up of clypeus in profile. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610589" httpUri="https://zenodo.org/record/4610589/files/figure.png" pageId="33" pageNumber="150">Figs 13b</figureCitation>
, 
<figureCitation id="2DE95399FFA3FFCCFAD8FC90FA9AFC8A" box="[1347,1393,799,825]" captionStart="Fig" captionStartId="35.[189,232,1945,1971]" captionTargetBox="[227,1361,995,1902]" captionTargetId="figure-258@35.[227,1361,995,1902]" captionTargetPageId="35" captionText="Fig. 14. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, large worker (ZRC_ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610591" httpUri="https://zenodo.org/record/4610591/files/figure.png" pageId="33" pageNumber="150">14b</figureCitation>
, 
<figureCitation id="2DE95399FFA3FFCCFF25FCCDFF06FCEE" box="[190,237,834,861]" captionStart="Fig" captionStartId="36.[189,232,1945,1971]" captionTargetBox="[227,1361,701,1911]" captionTargetId="figure-150@36.[227,1361,701,1911]" captionTargetPageId="36" captionText="Fig. 15. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, small worker (ZRC_ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610593" httpUri="https://zenodo.org/record/4610593/files/figure.png" pageId="33" pageNumber="150">15b</figureCitation>
). Posterior face of propodeum very weakly marginate, rounding smoothly into lateral propodeal face without distinct marginate edge. Median clypeal face coarsely carinulate, interspaces punctate and weakly shiny. Mandible with upper half of outer face finely striate, lower half superficially substriate (
<figureCitation id="2DE95399FFA3FFCCFF5DFC23FEF5FC74" box="[198,286,940,967]" captionStart="Fig" captionStartId="5.[189,232,1816,1842]" captionTargetBox="[227,1361,973,1778]" captionTargetId="figure-170@5.[227,1361,973,1778]" captionTargetPageId="5" captionText="Fig. 2. Closeups of external mandibular surface for workers of 5 Thai species of Rhopalomastix Forel, 1900 (ZRC); white arrows indicate distinct striations on the external mandibular face. a. R. impithuksai Wang &amp; Jaitrong sp. nov., paratype (ZRC_ENT00007608). b. R. robusta Wang &amp; Jaitrong sp. nov., paratype (ZRC_ENT00013949). c. R. javana Wheeler, 1929, non-type (ZRC_ENT00000932). d. R. johorensis Wheeler, 1929, non-type (ZRC_ENT00007874). e. R. parva Wang &amp; Jaitrong sp. nov., paratype (ZRC_ENT00013948)." figureDoi="http://doi.org/10.5281/zenodo.4610567" httpUri="https://zenodo.org/record/4610567/files/figure.png" pageId="33" pageNumber="150">Fig. 2b</figureCitation>
). Petiolar node and postpetiole each with two pairs of very long erect hairs. In larger workers, head, dorsum of mesosoma, and procoxa blackish or dark reddish brown, with rest of body generally uniformly lighter brown, tips of antennae and legs pale yellowish brown.
</paragraph>
</subSubSection>
<subSubSection id="FDC81C97FFA3FFCCFF26FBBBFB75FB13" pageId="33" pageNumber="150" type="etymology">
<paragraph id="B56D4F1CFFA3FFCCFF26FBBBFEAFFBFD" blockId="33.[189,324,1076,1102]" box="[189,324,1076,1102]" pageId="33" pageNumber="150">
<heading id="EE25F870FFA3FFCCFF26FBBBFEAFFBFD" bold="true" box="[189,324,1076,1102]" fontSize="11" level="3" pageId="33" pageNumber="150" reason="3">
<emphasis id="87A6930EFFA3FFCCFF26FBBBFEAFFBFD" bold="true" box="[189,324,1076,1102]" pageId="33" pageNumber="150">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="B56D4F1CFFA3FFCCFF26FBEDFB75FB13" blockId="33.[189,1398,1122,1184]" pageId="33" pageNumber="150">The species name is a both a direct reference to the robust habitus of the workers, and also a subtle allusion to the species’ intense dark brown colour resembling brewed robusta coffee.</paragraph>
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<subSubSection id="FDC81C97FFA3FFCCFF26FB48FD21F9D8" pageId="33" pageNumber="150" type="materials_examined">
<paragraph id="B56D4F1CFFA3FFCCFF26FB48FE41FB52" blockId="33.[189,426,1223,1249]" box="[189,426,1223,1249]" pageId="33" pageNumber="150">
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<emphasis id="87A6930EFFA3FFCCFF26FB48FE41FB52" bold="true" box="[189,426,1223,1249]" pageId="33" pageNumber="150">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="B56D4F1CFFA3FFCCFF71FB79FBD8FACA" blockId="33.[189,1398,1270,1402]" pageId="33" pageNumber="150">
<materialsCitation id="05BA4541FFA3FFCCFF71FB79FBD8FACA" ID-GBIF-Occurrence="3053841337" collectedFrom="hill evergreen forest, nest in bark of Toona ciliata M. Roem." collectingDate="2018-01-08" collectorName="W. Jaitrong" country="Thailand" county="Central Thailand" elevation="927" latitude="14.4372225" location="Ban Hin Subdistrict" longLatPrecision="21" longitude="101.37583" municipality="Muaeng District" pageId="33" pageNumber="150" specimenCode="THNHM-I-21782" specimenCount="1" stateProvince="Nakhon Nayok Province" typeStatus="holotype">
<emphasis id="87A6930EFFA3FFCCFF71FB79FEB7FAA3" bold="true" box="[234,348,1270,1296]" pageId="33" pageNumber="150">
<typeStatus id="6A69F1BEFFA3FFCCFF71FB79FEB7FAA3" box="[234,348,1270,1296]" pageId="33" pageNumber="150" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="CDC50F8CFFA3FFCCFF26FA96FEBEFA80" box="[189,341,1305,1331]" name="Thailand" pageId="33" pageNumber="150">THAILAND</collectingCountry>
• 
<specimenCount id="A3D48495FFA3FFCCFEF2FA96FE2BFA80" box="[361,448,1305,1331]" pageId="33" pageNumber="150" type="worker">worker</specimenCount>
; 
<collectingCounty id="5C0C3790FFA3FFCCFE52FA96FD65FA80" box="[457,654,1305,1331]" pageId="33" pageNumber="150">Central Thailand</collectingCounty>
, 
<collectingRegion id="771681FEFFA3FFCCFD03FA96FC53FA80" box="[664,952,1305,1331]" country="Thailand" name="Nakhon Nayok" pageId="33" pageNumber="150">Nakhon Nayok Province</collectingRegion>
, 
<collectingMunicipality id="5509D566FFA3FFCCFC5AFA96FB6AFA80" box="[961,1153,1305,1331]" pageId="33" pageNumber="150">Muaeng District</collectingMunicipality>
, 
<location id="B00D19C7FFA3FFCCFB11FA96FA99FA80" LSID="urn:lsid:plazi:treatment:3D7BFE0AFFA3FFCBFE6EFE87FD50FD8E:B00D19C7FFA3FFCCFB11FA96FA99FA80" box="[1162,1394,1305,1331]" country="Thailand" county="Central Thailand" latitude="14.4372225" longLatPrecision="21" longitude="101.37583" municipality="Muaeng District" name="Ban Hin Subdistrict" pageId="33" pageNumber="150" stateProvince="Nakhon Nayok Province">Ban Hin Subdistrict</location>
; 
<geoCoordinate id="D0E629DBFFA3FFCCFF26FAB5FEB9FAE4" box="[189,338,1338,1367]" degrees="14" direction="north" minutes="26" orientation="latitude" pageId="33" pageNumber="150" precision="15" seconds="14" value="14.4372225">14°26′14″ N</geoCoordinate>
, 
<geoCoordinate id="D0E629DBFFA3FFCCFEC6FAB5FE15FAE4" box="[349,510,1338,1367]" degrees="101" direction="east" minutes="22" orientation="longitude" pageId="33" pageNumber="150" precision="15" seconds="33" value="101.37583">101°22′33″ E</geoCoordinate>
; 
<elevation id="3EFFA82FFFA3FFCCFD92FAB3FD79FAE5" box="[521,658,1340,1367]" metricMagnitude="2" metricUnit="m" metricValue="9.27" pageId="33" pageNumber="150" unit="m" value="927.0">
<quantity id="722AE2F9FFA3FFCCFD92FAB3FDB8FAE4" box="[521,595,1340,1367]" metricMagnitude="2" metricUnit="m" metricValue="9.27" pageId="33" pageNumber="150" unit="m" value="927.0">927 m</quantity>
a.s.l.
</elevation>
; 
<date id="C16C69DCFFA3FFCCFD05FAB3FCCCFAE5" box="[670,807,1340,1367]" pageId="33" pageNumber="150" value="2018-01-08">
<collectingDate id="D1289034FFA3FFCCFD05FAB3FCCCFAE5" box="[670,807,1340,1367]" pageId="33" pageNumber="150" value="2018-01-08">8 Jan. 2018</collectingDate>
</date>
; 
<collectorName id="18272ACAFFA3FFCCFCAAFAB2FC5CFAE5" box="[817,951,1340,1367]" pageId="33" pageNumber="150">W. Jaitrong</collectorName>
leg.; 
<collectedFrom id="06CE0CE1FFA3FFCCFC6CFAB3FE12FAC9" pageId="33" pageNumber="150">
hill evergreen forest, nest in bark of 
<taxonomicName id="72D2349FFFA3FFCCFF46FAEFFE12FAC9" authority="M. Roem." authorityName="M. Roem." box="[221,505,1375,1402]" class="Magnoliopsida" family="Meliaceae" genus="Toona" kingdom="Plantae" order="Sapindales" pageId="33" pageNumber="150" phylum="Tracheophyta" rank="species" species="ciliata">
<emphasis id="87A6930EFFA3FFCCFF46FAEFFE9CFACA" box="[221,375,1375,1401]" italics="true" pageId="33" pageNumber="150">Toona ciliata</emphasis>
M. Roem.
</taxonomicName>
</collectedFrom>
; colony code: WJT080118-1; 
<specimenCode id="E574E767FFA3FFCCFCC2FAD0FBDBFACA" box="[857,1072,1375,1401]" pageId="33" pageNumber="150">THNHM-I-21782</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="B56D4F1CFFA3FFCCFF71FA2EFD21F9D8" blockId="33.[189,1399,1441,1643]" pageId="33" pageNumber="150">
<materialsCitation id="05BA4541FFA3FFCCFF71FA2EFA9CFA6D" ID-GBIF-Occurrence="3053841315" collectedFrom="hill evergreen forest, nest in bark of Toona ciliata M. Roem." collectingDate="2018-01-08" collectorName="W. Jaitrong" country="Thailand" county="Central Thailand" elevation="927" latitude="14.4372225" location="THAILAND" longLatPrecision="21" longitude="101.37583" municipality="Muaeng District" pageId="33" pageNumber="150" specimenCode="THNHM-I-21783 to THNHM-I-21801" specimenCount="19" specimenCount-worker="19" stateProvince="Nakhon Nayok" typeStatus="paratype">
<emphasis id="87A6930EFFA3FFCCFF71FA2EFE8CFA08" bold="true" box="[234,359,1441,1467]" pageId="33" pageNumber="150">
<typeStatus id="6A69F1BEFFA3FFCCFF71FA2EFE8CFA08" box="[234,359,1441,1467]" pageId="33" pageNumber="150" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="CDC50F8CFFA3FFCCFF26FA4BFEBEFA6D" box="[189,341,1476,1502]" name="Thailand" pageId="33" pageNumber="150">THAILAND</collectingCountry>
• 
<specimenCount id="A3D48495FFA3FFCCFEF7FA4BFE19FA6D" box="[364,498,1476,1502]" pageId="33" pageNumber="150" type="worker">19 workers</specimenCount>
; same collection data as for holotype; 
<specimenCode id="E574E767FFA3FFCCFC34FA4BFA9CFA6D" box="[943,1399,1476,1502]" pageId="33" pageNumber="150">THNHM-I-21783 to THNHM-I-21801</specimenCode>
</materialsCitation>
• 
<materialsCitation id="05BA4541FFA3FFCCFF4AFA68FB38F996" ID-GBIF-Occurrence="3053841307" collectedFrom="hill evergreen forest, nest in bark of Toona ciliata" collectingDate="2019-11-18" collectionCode="THNHM" collectorName="W. Wang &amp; W. Jaitrong" country="Thailand" county="Central Thailand" elevation="927" latitude="14.4372225" location="Ban Hin Subdistrict" longLatPrecision="21" longitude="101.37583" municipality="Muaeng District" pageId="33" pageNumber="150" specimenCount="10" specimenCount-worker="10" stateProvince="Nakhon Nayok" typeStatus="paratype">
<specimenCount id="A3D48495FFA3FFCCFF4AFA68FEB0F9B2" box="[209,347,1511,1537]" pageId="33" pageNumber="150" type="worker">10 workers</specimenCount>
; same collection data as for holotype; 
<date id="C16C69DCFFA3FFCCFCAAFA68FC33F9B2" box="[817,984,1511,1538]" pageId="33" pageNumber="150" value="2019-11-18">
<collectingDate id="D1289034FFA3FFCCFCAAFA68FC33F9B2" box="[817,984,1511,1538]" pageId="33" pageNumber="150" value="2019-11-18">18 Nov. 2019</collectingDate>
</date>
; 
<collectorName id="18272ACAFFA3FFCCFC7CFA67FBBEF9B1" box="[999,1109,1512,1538]" pageId="33" pageNumber="150">W. Wang</collectorName>
&amp; 
<collectorName id="18272ACAFFA3FFCCFBE4FA67FAE3F9B2" box="[1151,1288,1511,1538]" pageId="33" pageNumber="150">W. Jaitrong</collectorName>
leg.; 
<collectedFrom id="06CE0CE1FFA3FFCCFAD4FA68FD3FF997" pageId="33" pageNumber="150">
hill evergreen forest, nest in bark of 
<taxonomicName id="72D2349FFFA3FFCCFDA2F984FD3FF997" box="[569,724,1546,1572]" class="Magnoliopsida" family="Meliaceae" genus="Toona" kingdom="Plantae" order="Sapindales" pageId="33" pageNumber="150" phylum="Tracheophyta" rank="species" species="ciliata">
<emphasis id="87A6930EFFA3FFCCFDA2F984FD3FF997" box="[569,724,1546,1572]" italics="true" pageId="33" pageNumber="150">Toona ciliata</emphasis>
</taxonomicName>
</collectedFrom>
; colony code: WJW-THL19-007; 
<collectionCode id="D3C3D7D9FFA3FFCCFBFCF984FB38F996" box="[1127,1235,1547,1573]" pageId="33" pageNumber="150">THNHM</collectionCode>
</materialsCitation>
• 
<materialsCitation id="05BA4541FFA3FFCCFB70F984FD2EF9D8" ID-GBIF-Occurrence="3053841327" accessionNumber="MW267049, MW267051" collectedFrom="hill evergreen forest, nest in bark of Toona ciliata" collectingDate="2019-11-18" collectionCode="MW, ZRC" collectorName="GenBank" country="Thailand" county="Central Thailand" elevation="927" latitude="14.4372225" location="Ban Hin Subdistrict" longLatPrecision="21" longitude="101.37583" municipality="Muaeng District" pageId="33" pageNumber="150" specimenCode="ZRC_ ENT00013949.1 to ZRC_ENT00013949.77" specimenCount="77" specimenCount-worker="77" stateProvince="Nakhon Nayok" typeStatus="paratype">
<specimenCount id="A3D48495FFA3FFCCFB70F984FA99F997" box="[1259,1394,1546,1573]" pageId="33" pageNumber="150" type="worker">77 workers</specimenCount>
; same collection data as for preceding; GenBank: 
<accessionNumber id="AA81D2FFFFA3FFCCFCBDF9A2FC5FF9F4" box="[806,948,1581,1607]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MW267049" pageId="33" pageNumber="150">MW267049</accessionNumber>
to 
<accessionNumber id="AA81D2FFFFA3FFCCFC7FF9A2FB99F9FB" box="[996,1138,1581,1608]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MW267051" pageId="33" pageNumber="150">MW267051</accessionNumber>
(3 barcoded); 
<specimenCode id="E574E767FFA3FFCCFAB4F9A1FD2EF9D8" pageId="33" pageNumber="150">ZRC_ ENT00013949.1 to ZRC_ENT00013949.77</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="FDC81C97FFA3FFC8FF26F91DFDC5FD12" lastPageId="37" lastPageNumber="154" pageId="33" pageNumber="150" type="description">
<paragraph id="B56D4F1CFFA3FFCCFF26F91DFEA5F91F" blockId="33.[189,334,1682,1708]" box="[189,334,1682,1708]" pageId="33" pageNumber="150">
<heading id="EE25F870FFA3FFCCFF26F91DFEA5F91F" bold="true" box="[189,334,1682,1708]" fontSize="11" level="3" pageId="33" pageNumber="150" reason="3">
<emphasis id="87A6930EFFA3FFCCFF26F91DFEA5F91F" bold="true" box="[189,334,1682,1708]" pageId="33" pageNumber="150">Description</emphasis>
</heading>
</paragraph>
<paragraph id="B56D4F1CFFA3FFCCFF71F94EFEA7F968" blockId="33.[189,1399,1729,1826]" box="[234,332,1729,1755]" pageId="33" pageNumber="150">
<emphasis id="87A6930EFFA3FFCCFF71F94EFEA7F968" bold="true" box="[234,332,1729,1755]" pageId="33" pageNumber="150">Worker</emphasis>
</paragraph>
<paragraph id="B56D4F1CFFA3FFCCFF26F96BFC17F892" blockId="33.[189,1399,1729,1826]" pageId="33" pageNumber="150">
Measurements. 
<typeStatus id="6A69F1BEFFA3FFCCFEE7F96BFE02F94D" box="[380,489,1764,1790]" pageId="33" pageNumber="150" type="holotype">Holotype</typeStatus>
worker: EL 0.12; EW 0.08; HL 0.64; HW 0.66; ML 0.86; PronW 0.46; PtH 0.25; PtL 0.27; SL 0.28; TL 2.91; CI 103; PtHI 93; REL 18; SI 42.
</paragraph>
<paragraph id="B56D4F1CFFA3FFCCFF26F8C6FE2EF81A" blockId="33.[188,1398,1864,1961]" pageId="33" pageNumber="150">
<typeStatus id="6A69F1BEFFA3FFCCFF26F8C6FECFF8D0" box="[189,292,1865,1891]" pageId="33" pageNumber="150" type="paratype">Paratype</typeStatus>
workers (n = 12): EL 0.09–0.14; EW 0.06–0.08; HL 0.55–0.68; HW 0.52–0.70; ML 0.68–0.96; PronW 0.36–0.48; PtH 0.19–0.28; PtL 0.20–0.31; SL 0.24–0.31; TL 2.24–3.16; CI 95–103; PtHI 91–96; REL 17–20; SI 42–46.
</paragraph>
<paragraph id="B56D4F1CFFA0FFCEFF26FE86FD3BFE44" blockId="34.[187,1401,265,925]" lastBlockId="35.[188,1400,265,503]" lastPageId="35" lastPageNumber="152" pageId="34" pageNumber="151">
Relatively large sized for 
<taxonomicName id="72D2349FFFA0FFCFFD90FE86FD50FE90" authorityName="Forel" authorityYear="1900" box="[523,699,265,291]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="87A6930EFFA0FFCFFD90FE86FD50FE90" box="[523,699,265,291]" italics="true" pageId="34" pageNumber="151">Rhopalomastix</emphasis>
</taxonomicName>
(TL 2.24–3.16) with habitus generally robust; workers monomorphic with broad size variation. Head in full-face view subsquarish, as long as wide or slightly wider than long (CI 100–103), in small workers sometimes slightly longer than wide (
<figureCitation id="2DE95399FFA0FFCFFB0FFEC0FB1DFED9" box="[1172,1270,335,362]" captionStart="Fig" captionStartId="36.[189,232,1945,1971]" captionTargetBox="[227,1361,701,1911]" captionTargetId="figure-150@36.[227,1361,701,1911]" captionTargetPageId="36" captionText="Fig. 15. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, small worker (ZRC_ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610593" httpUri="https://zenodo.org/record/4610593/files/figure.png" pageId="34" pageNumber="151">Fig. 15a</figureCitation>
), distinctly narrowed anteriorly; narrow median furrow between frontal lobes deep and long, extending till or past transverse midline of head, narrower or constricted shortly before midlength; posterior margin with a broad but shallow median concavity, posterolateral corners roundly convex and laterally expanded to form somewhat rounded vertexal lobes, lateral margins of head broadly convex and weakly converging anteriorly, anterior clypeal margin with broadly and weakly convex median section (
<figureCitation id="2DE95399FFA0FFCFFB53FE70FADEFDA9" box="[1224,1333,511,538]" captionStart="Fig" captionStartId="34.[189,232,1917,1943]" captionTargetBox="[227,1361,977,1884]" captionTargetId="figure-275@34.[227,1361,977,1884]" captionTargetPageId="34" captionText="Fig. 13. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, average-sized worker (ZRC_ ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Close-up of clypeus in profile. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610589" httpUri="https://zenodo.org/record/4610589/files/figure.png" pageId="34" pageNumber="151">Figs 13a</figureCitation>
, 
<figureCitation id="2DE95399FFA0FFCFFADEFD8FFA9AFDA9" box="[1349,1393,512,538]" captionStart="Fig" captionStartId="35.[189,232,1945,1971]" captionTargetBox="[227,1361,995,1902]" captionTargetId="figure-258@35.[227,1361,995,1902]" captionTargetPageId="35" captionText="Fig. 14. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, large worker (ZRC_ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610591" httpUri="https://zenodo.org/record/4610591/files/figure.png" pageId="34" pageNumber="151">14a</figureCitation>
, 
<figureCitation id="2DE95399FFA0FFCFFF26FDADFF00FD8E" box="[189,235,546,573]" captionStart="Fig" captionStartId="36.[189,232,1945,1971]" captionTargetBox="[227,1361,701,1911]" captionTargetId="figure-150@36.[227,1361,701,1911]" captionTargetPageId="36" captionText="Fig. 15. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, small worker (ZRC_ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610593" httpUri="https://zenodo.org/record/4610593/files/figure.png" pageId="34" pageNumber="151">15a</figureCitation>
). Eye with 16–20 ommatidia in small workers, 22–32 ommatidia in larger workers. In lateral view, anterior face of clypeus weakly and evenly convex, sometimes almost flat (
<figureCitation id="2DE95399FFA0FFCFFBCDFDC9FB55FDD3" box="[1110,1214,582,608]" captionStart="Fig" captionStartId="34.[189,232,1917,1943]" captionTargetBox="[227,1361,977,1884]" captionTargetId="figure-275@34.[227,1361,977,1884]" captionTargetPageId="34" captionText="Fig. 13. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, average-sized worker (ZRC_ ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Close-up of clypeus in profile. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610589" httpUri="https://zenodo.org/record/4610589/files/figure.png" pageId="34" pageNumber="151">Fig. 13c</figureCitation>
). Mesosoma in lateral view box-shaped and subcylindrical, dorsal outline weakly and broadly convex, almost flat, propodeal junction roundly obtuse, propodeal declivity steep and shallowly concave (
<figureCitation id="2DE95399FFA0FFCFFB53FD03FADDFD15" box="[1224,1334,652,678]" captionStart="Fig" captionStartId="34.[189,232,1917,1943]" captionTargetBox="[227,1361,977,1884]" captionTargetId="figure-275@34.[227,1361,977,1884]" captionTargetPageId="34" captionText="Fig. 13. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, average-sized worker (ZRC_ ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Close-up of clypeus in profile. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610589" httpUri="https://zenodo.org/record/4610589/files/figure.png" pageId="34" pageNumber="151">Figs 13d</figureCitation>
, 
<figureCitation id="2DE95399FFA0FFCFFADEFD03FA99FD15" box="[1349,1394,652,678]" captionStart="Fig" captionStartId="35.[189,232,1945,1971]" captionTargetBox="[227,1361,995,1902]" captionTargetId="figure-258@35.[227,1361,995,1902]" captionTargetPageId="35" captionText="Fig. 14. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, large worker (ZRC_ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610591" httpUri="https://zenodo.org/record/4610591/files/figure.png" pageId="34" pageNumber="151">14c</figureCitation>
, 
<figureCitation id="2DE95399FFA0FFCFFF26FD20FF01FD79" box="[189,234,687,714]" captionStart="Fig" captionStartId="36.[189,232,1945,1971]" captionTargetBox="[227,1361,701,1911]" captionTargetId="figure-150@36.[227,1361,701,1911]" captionTargetPageId="36" captionText="Fig. 15. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, small worker (ZRC_ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610593" httpUri="https://zenodo.org/record/4610593/files/figure.png" pageId="34" pageNumber="151">15c</figureCitation>
); propodeal dorsum rounding smoothly into posterior declivitous face at indistinct edge. In dorsal view, mesosoma subcylindrical, anterior margin of pronotal disc broadly convex with bluntly angulate median point, humeral corners obtusely angulate, lateral margins with distinct inflexions differentiating pronotum, mesonotum and propodeum (
<figureCitation id="2DE95399FFA0FFCFFD36FC96FCF6FC80" box="[685,797,793,819]" captionStart="Fig" captionStartId="34.[189,232,1917,1943]" captionTargetBox="[227,1361,977,1884]" captionTargetId="figure-275@34.[227,1361,977,1884]" captionTargetPageId="34" captionText="Fig. 13. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, average-sized worker (ZRC_ ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Close-up of clypeus in profile. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610589" httpUri="https://zenodo.org/record/4610589/files/figure.png" pageId="34" pageNumber="151">Figs 13b</figureCitation>
, 
<figureCitation id="2DE95399FFA0FFCFFCB4FC96FCB6FC80" box="[815,861,793,819]" captionStart="Fig" captionStartId="35.[189,232,1945,1971]" captionTargetBox="[227,1361,995,1902]" captionTargetId="figure-258@35.[227,1361,995,1902]" captionTargetPageId="35" captionText="Fig. 14. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, large worker (ZRC_ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610591" httpUri="https://zenodo.org/record/4610591/files/figure.png" pageId="34" pageNumber="151">14b</figureCitation>
, 
<figureCitation id="2DE95399FFA0FFCFFCF4FC96FC74FC87" box="[879,927,793,820]" captionStart="Fig" captionStartId="36.[189,232,1945,1971]" captionTargetBox="[227,1361,701,1911]" captionTargetId="figure-150@36.[227,1361,701,1911]" captionTargetPageId="36" captionText="Fig. 15. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, small worker (ZRC_ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610593" httpUri="https://zenodo.org/record/4610593/files/figure.png" pageId="34" pageNumber="151">15b</figureCitation>
); in large workers, mesosoma weakly narrowed posteriorly (
<figureCitation id="2DE95399FFA0FFCFFE5FFCB3FDC0FCE4" box="[452,555,828,855]" captionStart="Fig" captionStartId="35.[189,232,1945,1971]" captionTargetBox="[227,1361,995,1902]" captionTargetId="figure-258@35.[227,1361,995,1902]" captionTargetPageId="35" captionText="Fig. 14. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, large worker (ZRC_ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610591" httpUri="https://zenodo.org/record/4610591/files/figure.png" pageId="34" pageNumber="151">Fig. 14b</figureCitation>
); lateral margins of propodeum broadly convex with indistinct rounded posterolateral corners, posterior propodeal margin weakly defined (
<figureCitation id="2DE95399FFA0FFCFFC41FCD0FBAEFCCA" box="[986,1093,863,889]" captionStart="Fig" captionStartId="34.[189,232,1917,1943]" captionTargetBox="[227,1361,977,1884]" captionTargetId="figure-275@34.[227,1361,977,1884]" captionTargetPageId="34" captionText="Fig. 13. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, average-sized worker (ZRC_ ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Close-up of clypeus in profile. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610589" httpUri="https://zenodo.org/record/4610589/files/figure.png" pageId="34" pageNumber="151">Figs 13b</figureCitation>
, 
<figureCitation id="2DE95399FFA0FFCFFBC8FCD0FB6AFCCA" box="[1107,1153,863,889]" captionStart="Fig" captionStartId="35.[189,232,1945,1971]" captionTargetBox="[227,1361,995,1902]" captionTargetId="figure-258@35.[227,1361,995,1902]" captionTargetPageId="35" captionText="Fig. 14. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, large worker (ZRC_ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610591" httpUri="https://zenodo.org/record/4610591/files/figure.png" pageId="34" pageNumber="151">14b</figureCitation>
, 
<figureCitation id="2DE95399FFA0FFCFFB15FCD0FB55FCC9" box="[1166,1214,863,890]" captionStart="Fig" captionStartId="36.[189,232,1945,1971]" captionTargetBox="[227,1361,701,1911]" captionTargetId="figure-150@36.[227,1361,701,1911]" captionTargetPageId="36" captionText="Fig. 15. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, small worker (ZRC_ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610593" httpUri="https://zenodo.org/record/4610593/files/figure.png" pageId="34" pageNumber="151">15b</figureCitation>
). Posterior face of propodeum very weakly marginate and almost flat, sometimes with upper half shallowly depressed, rounding smoothly into lateral propodeal face without distinct marginate edge. In lateral view, petiole inclined posteriorly and bell-shaped, slightly longer than high, anterior face steep and weakly concave, apex rounded and slightly higher than postpetiole, apical dorsum rounding into weakly convex posterior face at indistinct rounded edge, posterior face steeper and shorter than anterior face (
<figureCitation id="2DE95399FFA1FFCEFB15FEFDFB1CFE3F" box="[1166,1271,370,396]" captionStart="Fig" captionStartId="34.[189,232,1917,1943]" captionTargetBox="[227,1361,977,1884]" captionTargetId="figure-275@34.[227,1361,977,1884]" captionTargetPageId="34" captionText="Fig. 13. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, average-sized worker (ZRC_ ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Close-up of clypeus in profile. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610589" httpUri="https://zenodo.org/record/4610589/files/figure.png" pageId="35" pageNumber="152">Figs 13d</figureCitation>
, 
<figureCitation id="2DE95399FFA1FFCEFA99FEFCFAC4FE3E" box="[1282,1327,371,397]" captionStart="Fig" captionStartId="35.[189,232,1945,1971]" captionTargetBox="[227,1361,995,1902]" captionTargetId="figure-258@35.[227,1361,995,1902]" captionTargetPageId="35" captionText="Fig. 14. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, large worker (ZRC_ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610591" httpUri="https://zenodo.org/record/4610591/files/figure.png" pageId="35" pageNumber="152">14c</figureCitation>
, 
<figureCitation id="2DE95399FFA1FFCEFAA1FEFDFA83FE3E" box="[1338,1384,370,397]" captionStart="Fig" captionStartId="36.[189,232,1945,1971]" captionTargetBox="[227,1361,701,1911]" captionTargetId="figure-150@36.[227,1361,701,1911]" captionTargetPageId="36" captionText="Fig. 15. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, small worker (ZRC_ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610593" httpUri="https://zenodo.org/record/4610593/files/figure.png" pageId="35" pageNumber="152">15c</figureCitation>
). Petiole higher and longer than postpetiole, dorsal margin of postpetiole weakly convex, almost flat in large workers (
<figureCitation id="2DE95399FFA1FFCEFEF7FE36FE24FE60" box="[364,463,441,467]" captionStart="Fig" captionStartId="35.[189,232,1945,1971]" captionTargetBox="[227,1361,995,1902]" captionTargetId="figure-258@35.[227,1361,995,1902]" captionTargetPageId="35" captionText="Fig. 14. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, large worker (ZRC_ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610591" httpUri="https://zenodo.org/record/4610591/files/figure.png" pageId="35" pageNumber="152">Fig. 14c</figureCitation>
). In dorsal view, petiole suboval, wider than long; postpetiole globular, less than twice as wide as petiole (
<figureCitation id="2DE95399FFA1FFCEFE79FE53FDA7FE45" box="[482,588,476,502]" captionStart="Fig" captionStartId="34.[189,232,1917,1943]" captionTargetBox="[227,1361,977,1884]" captionTargetId="figure-275@34.[227,1361,977,1884]" captionTargetPageId="34" captionText="Fig. 13. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, average-sized worker (ZRC_ ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Close-up of clypeus in profile. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610589" httpUri="https://zenodo.org/record/4610589/files/figure.png" pageId="35" pageNumber="152">Figs 13b</figureCitation>
, 
<figureCitation id="2DE95399FFA1FFCEFDC2FE53FD6CFE45" box="[601,647,476,502]" captionStart="Fig" captionStartId="35.[189,232,1945,1971]" captionTargetBox="[227,1361,995,1902]" captionTargetId="figure-258@35.[227,1361,995,1902]" captionTargetPageId="35" captionText="Fig. 14. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, large worker (ZRC_ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610591" httpUri="https://zenodo.org/record/4610591/files/figure.png" pageId="35" pageNumber="152">14b</figureCitation>
, 
<figureCitation id="2DE95399FFA1FFCEFD08FE53FD28FE44" box="[659,707,476,503]" captionStart="Fig" captionStartId="36.[189,232,1945,1971]" captionTargetBox="[227,1361,701,1911]" captionTargetId="figure-150@36.[227,1361,701,1911]" captionTargetPageId="36" captionText="Fig. 15. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, small worker (ZRC_ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610593" httpUri="https://zenodo.org/record/4610593/files/figure.png" pageId="35" pageNumber="152">15b</figureCitation>
).
</paragraph>
<caption id="E1AD1F94FFA0FFCFFF26F8F2FE3AF86D" ID-DOI="http://doi.org/10.5281/zenodo.4610589" ID-Zenodo-Dep="4610589" httpUri="https://zenodo.org/record/4610589/files/figure.png" pageId="34" pageNumber="151" startId="34.[189,232,1917,1943]" targetBox="[227,1361,977,1884]" targetPageId="34">
<paragraph id="B56D4F1CFFA0FFCFFF26F8F2FE3AF86D" blockId="34.[189,1399,1917,2014]" pageId="34" pageNumber="151">
<emphasis id="87A6930EFFA0FFCFFF26F8F2FEF5F824" bold="true" box="[189,286,1917,1943]" pageId="34" pageNumber="151">Fig. 13.</emphasis>
<taxonomicName id="72D2349FFFA0FFCFFEB0F8F2FCF4F824" authority="Wang &amp; Jaitrong" authorityName="Wang &amp; Yong &amp; Jaitrong" authorityYear="2021" box="[299,799,1917,1944]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="151" phylum="Arthropoda" rank="species" species="robusta" status="sp. nov.">
<emphasis id="87A6930EFFA0FFCFFEB0F8F2FDABF824" box="[299,576,1917,1943]" italics="true" pageId="34" pageNumber="151">Rhopalomastix robusta</emphasis>
Wang &amp; Jaitrong
</taxonomicName>
<taxonomicNameLabel id="9C952E75FFA0FFCFFCB0F8F1FC65F82B" box="[811,910,1918,1944]" pageId="34" pageNumber="151" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, average-sized worker (ZRC_ ENT00013949). 
<emphasis id="87A6930EFFA0FFCFFE1DF82EFE7EF808" bold="true" box="[390,405,1953,1979]" pageId="34" pageNumber="151">a</emphasis>
. Head in full-face view. 
<emphasis id="87A6930EFFA0FFCFFD5FF82EFD3FF808" bold="true" box="[708,724,1953,1979]" pageId="34" pageNumber="151">b</emphasis>
. Body in dorsal view. 
<emphasis id="87A6930EFFA0FFCFFC71F82EFC1CF808" bold="true" box="[1002,1015,1953,1979]" pageId="34" pageNumber="151">c</emphasis>
. Close-up of clypeus in profile. 
<emphasis id="87A6930EFFA0FFCFFF26F84BFF26F86D" bold="true" box="[189,205,1988,2014]" pageId="34" pageNumber="151">d</emphasis>
. Body in lateral view.
</paragraph>
</caption>
<paragraph id="B56D4F1CFFA1FFC9FF26FD91FDF2FEDA" blockId="35.[187,1399,542,920]" lastBlockId="36.[189,1399,265,361]" lastPageId="36" lastPageNumber="153" pageId="35" pageNumber="152">
Dorsum of head finely striate and dully shining with scattered punctures, lateral face substriate-reticulate with scattered punctures and interspaces smooth and shining; ventral face largely finely reticulate with interspaces smooth and shining; median clypeal face coarsely carinulate, interspaces punctate and weakly shiny. Mandible in full-face view with upper half of outer face finely striate, lower half superficially substriate (
<figureCitation id="2DE95399FFA1FFCEFEDAFD25FE77FD76" box="[321,412,682,709]" captionStart="Fig" captionStartId="5.[189,232,1816,1842]" captionTargetBox="[227,1361,973,1778]" captionTargetId="figure-170@5.[227,1361,973,1778]" captionTargetPageId="5" captionText="Fig. 2. Closeups of external mandibular surface for workers of 5 Thai species of Rhopalomastix Forel, 1900 (ZRC); white arrows indicate distinct striations on the external mandibular face. a. R. impithuksai Wang &amp; Jaitrong sp. nov., paratype (ZRC_ENT00007608). b. R. robusta Wang &amp; Jaitrong sp. nov., paratype (ZRC_ENT00013949). c. R. javana Wheeler, 1929, non-type (ZRC_ENT00000932). d. R. johorensis Wheeler, 1929, non-type (ZRC_ENT00007874). e. R. parva Wang &amp; Jaitrong sp. nov., paratype (ZRC_ENT00013948)." figureDoi="http://doi.org/10.5281/zenodo.4610567" httpUri="https://zenodo.org/record/4610567/files/figure.png" pageId="35" pageNumber="152">Fig. 2b</figureCitation>
), in larger workers striae on mandible more coarse, area immediately adjacent to masticatory margin punctate. Lateral face of mesosoma largely substriate-reticulate with interspaces smooth and shining (
<figureCitation id="2DE95399FFA1FFCEFE2FFD7EFDF4FCB8" box="[436,543,753,779]" captionStart="Fig" captionStartId="34.[189,232,1917,1943]" captionTargetBox="[227,1361,977,1884]" captionTargetId="figure-275@34.[227,1361,977,1884]" captionTargetPageId="34" captionText="Fig. 13. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, average-sized worker (ZRC_ ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Close-up of clypeus in profile. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610589" httpUri="https://zenodo.org/record/4610589/files/figure.png" pageId="35" pageNumber="152">Figs 13d</figureCitation>
, 
<figureCitation id="2DE95399FFA1FFCEFDB7FD7EFDB2FCB8" box="[556,601,753,779]" captionStart="Fig" captionStartId="35.[189,232,1945,1971]" captionTargetBox="[227,1361,995,1902]" captionTargetId="figure-258@35.[227,1361,995,1902]" captionTargetPageId="35" captionText="Fig. 14. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, large worker (ZRC_ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610591" httpUri="https://zenodo.org/record/4610591/files/figure.png" pageId="35" pageNumber="152">14c</figureCitation>
, 
<figureCitation id="2DE95399FFA1FFCEFDFDFD7EFD78FCBF" box="[614,659,753,780]" captionStart="Fig" captionStartId="36.[189,232,1945,1971]" captionTargetBox="[227,1361,701,1911]" captionTargetId="figure-150@36.[227,1361,701,1911]" captionTargetPageId="36" captionText="Fig. 15. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, small worker (ZRC_ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610593" httpUri="https://zenodo.org/record/4610593/files/figure.png" pageId="35" pageNumber="152">15c</figureCitation>
); dorsum of mesosoma finely striate and shiny (
<figureCitation id="2DE95399FFA1FFCEFB57FD7EFADDFCB8" box="[1228,1334,753,779]" captionStart="Fig" captionStartId="34.[189,232,1917,1943]" captionTargetBox="[227,1361,977,1884]" captionTargetId="figure-275@34.[227,1361,977,1884]" captionTargetPageId="34" captionText="Fig. 13. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, average-sized worker (ZRC_ ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Close-up of clypeus in profile. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610589" httpUri="https://zenodo.org/record/4610589/files/figure.png" pageId="35" pageNumber="152">Figs 13b</figureCitation>
, 
<figureCitation id="2DE95399FFA1FFCEFAD8FD7EFA9AFCB8" box="[1347,1393,753,779]" captionStart="Fig" captionStartId="35.[189,232,1945,1971]" captionTargetBox="[227,1361,995,1902]" captionTargetId="figure-258@35.[227,1361,995,1902]" captionTargetPageId="35" captionText="Fig. 14. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, large worker (ZRC_ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610591" httpUri="https://zenodo.org/record/4610591/files/figure.png" pageId="35" pageNumber="152">14b</figureCitation>
, 
<figureCitation id="2DE95399FFA1FFCEFF26FC9BFF07FC9C" box="[189,236,788,815]" captionStart="Fig" captionStartId="36.[189,232,1945,1971]" captionTargetBox="[227,1361,701,1911]" captionTargetId="figure-150@36.[227,1361,701,1911]" captionTargetPageId="36" captionText="Fig. 15. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, small worker (ZRC_ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610593" httpUri="https://zenodo.org/record/4610593/files/figure.png" pageId="35" pageNumber="152">15b</figureCitation>
); posterior face of propodeum superficially and transversely substriate-reticulate with interspaces smooth and shining. Coxae and femora distinctly but superficially reticulate with interspaces smooth and shining. Lateral faces of petiole and postpetiole superficially substriate-reticulate with interspaces smooth and shining, apicodorsal faces largely superficially reticulate with interspaces smooth and shining, apicalmost point of petiole entirely smooth and shining. Gaster largely superficially reticulate with interspaces smooth and shining; sculpture slightly stronger around base of first gastral tergite where it articulates with postpetiole.
</paragraph>
<caption id="E1AD1F94FFA1FFCEFF26F816FBFCF865" ID-DOI="http://doi.org/10.5281/zenodo.4610591" ID-Zenodo-Dep="4610591" httpUri="https://zenodo.org/record/4610591/files/figure.png" pageId="35" pageNumber="152" startId="35.[189,232,1945,1971]" targetBox="[227,1361,995,1902]" targetPageId="35">
<paragraph id="B56D4F1CFFA1FFCEFF26F816FBFCF865" blockId="35.[187,1399,1945,2007]" pageId="35" pageNumber="152">
<emphasis id="87A6930EFFA1FFCEFF26F816FEFCF800" bold="true" box="[189,279,1945,1971]" pageId="35" pageNumber="152">Fig. 14.</emphasis>
<taxonomicName id="72D2349FFFA1FFCEFE87F816FD18F800" authority="Wang &amp; Jaitrong" authorityName="Wang &amp; Yong &amp; Jaitrong" authorityYear="2021" box="[284,755,1945,1972]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="152" phylum="Arthropoda" rank="species" species="robusta" status="sp. nov.">
<emphasis id="87A6930EFFA1FFCEFE87F816FDC1F800" box="[284,554,1945,1971]" italics="true" pageId="35" pageNumber="152">Rhopalomastix robusta</emphasis>
Wang &amp; Jaitrong
</taxonomicName>
<taxonomicNameLabel id="9C952E75FFA1FFCEFD63F815FCB8F807" box="[760,851,1946,1972]" pageId="35" pageNumber="152" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, large worker (ZRC_ENT00013949). 
<emphasis id="87A6930EFFA1FFCEFF20F833FF21F865" bold="true" box="[187,202,1980,2006]" pageId="35" pageNumber="152">a</emphasis>
. Head in full-face view. 
<emphasis id="87A6930EFFA1FFCEFE75F833FE15F865" bold="true" box="[494,510,1980,2006]" pageId="35" pageNumber="152">b</emphasis>
. Body in dorsal view. 
<emphasis id="87A6930EFFA1FFCEFC9CF833FCFFF865" bold="true" box="[775,788,1980,2006]" pageId="35" pageNumber="152">c</emphasis>
. Body in lateral view.
</paragraph>
</caption>
<paragraph id="B56D4F1CFFA6FFC8FF26FE1EFAC6FE45" blockId="36.[189,1399,401,639]" lastBlockId="37.[188,1399,265,503]" lastPageId="37" lastPageNumber="154" pageId="36" pageNumber="153">
Dorsum and ventral face of head with fairly dense though well-spaced short standing and suberect hairs interspersed with sparse long erect hairs (
<figureCitation id="2DE95399FFA6FFC9FD65FE3BFC81FE7D" box="[766,874,436,462]" captionStart="Fig" captionStartId="34.[189,232,1917,1943]" captionTargetBox="[227,1361,977,1884]" captionTargetId="figure-275@34.[227,1361,977,1884]" captionTargetPageId="34" captionText="Fig. 13. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, average-sized worker (ZRC_ ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Close-up of clypeus in profile. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610589" httpUri="https://zenodo.org/record/4610589/files/figure.png" pageId="36" pageNumber="153">Figs 13a</figureCitation>
, 
<figureCitation id="2DE95399FFA6FFC9FCE1FE3BFC4CFE7D" box="[890,935,436,462]" captionStart="Fig" captionStartId="35.[189,232,1945,1971]" captionTargetBox="[227,1361,995,1902]" captionTargetId="figure-258@35.[227,1361,995,1902]" captionTargetPageId="35" captionText="Fig. 14. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, large worker (ZRC_ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610591" httpUri="https://zenodo.org/record/4610591/files/figure.png" pageId="36" pageNumber="153">14a</figureCitation>
, 
<figureCitation id="2DE95399FFA6FFC9FC2CFE3BFC0EFE7C" box="[951,997,436,463]" captionStart="Fig" captionStartId="36.[189,232,1945,1971]" captionTargetBox="[227,1361,701,1911]" captionTargetId="figure-150@36.[227,1361,701,1911]" captionTargetPageId="36" captionText="Fig. 15. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, small worker (ZRC_ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610593" httpUri="https://zenodo.org/record/4610593/files/figure.png" pageId="36" pageNumber="153">15a</figureCitation>
); lateral face with short suberect and subdecumbent hairs. Mesosoma dorsum with scattered short standing hairs, more dense close to dorsolateral margin (
<figureCitation id="2DE95399FFA6FFC9FE23FE75FDCEFDA7" box="[440,549,506,532]" captionStart="Fig" captionStartId="34.[189,232,1917,1943]" captionTargetBox="[227,1361,977,1884]" captionTargetId="figure-275@34.[227,1361,977,1884]" captionTargetPageId="34" captionText="Fig. 13. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, average-sized worker (ZRC_ ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Close-up of clypeus in profile. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610589" httpUri="https://zenodo.org/record/4610589/files/figure.png" pageId="36" pageNumber="153">Figs 13b</figureCitation>
, 
<figureCitation id="2DE95399FFA6FFC9FDAEFE75FD88FDA7" box="[565,611,506,532]" captionStart="Fig" captionStartId="35.[189,232,1945,1971]" captionTargetBox="[227,1361,995,1902]" captionTargetId="figure-258@35.[227,1361,995,1902]" captionTargetPageId="35" captionText="Fig. 14. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, large worker (ZRC_ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610591" httpUri="https://zenodo.org/record/4610591/files/figure.png" pageId="36" pageNumber="153">14b</figureCitation>
, 
<figureCitation id="2DE95399FFA6FFC9FDE8FE75FD49FDA6" box="[627,674,506,533]" captionStart="Fig" captionStartId="36.[189,232,1945,1971]" captionTargetBox="[227,1361,701,1911]" captionTargetId="figure-150@36.[227,1361,701,1911]" captionTargetPageId="36" captionText="Fig. 15. Rhopalomastix robusta Wang &amp; Jaitrong sp. nov., paratype, small worker (ZRC_ENT00013949). a. Head in full-face view. b. Body in dorsal view. c. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610593" httpUri="https://zenodo.org/record/4610593/files/figure.png" pageId="36" pageNumber="153">15b</figureCitation>
); paired long erect hairs sparsely present along dorsolateral margin, with at least one pair each flanking the anterior margin of pronotal disc, pronotum, mesonotum and propodeum; large workers sometimes with 2–3 pairs of long erect hairs on anterior margin of pronotal disc, and/or 2–3 pairs of long erect hairs flanking pronotum and propodeum. Anterior rising face of petiole with relatively sparse (compared to head) short decumbent and subdecumbent hairs, slightly more dense closer to apex; lateral petiolar face with 1–2 very long erect hairs and standing hairs slightly longer than those on anterior face, mostly densely along posterolateral margin; apex of petiolar node with sparse standing hairs longer than those on anterior face, and two pairs of very long erect hairs; posterior petiolar face almost hairless or with sparse short standing hairs. Postpetiole with fairly dense erect and suberect hairs longer than those on anterior petiolar face, and two pairs of much longer erect hairs. Gaster pilose with dense erect and suberect hairs, interspersed with sparse long erect hairs.
</paragraph>
<caption id="E1AD1F94FFA6FFC9FF26F816FBFCF865" ID-DOI="http://doi.org/10.5281/zenodo.4610593" ID-Zenodo-Dep="4610593" httpUri="https://zenodo.org/record/4610593/files/figure.png" pageId="36" pageNumber="153" startId="36.[189,232,1945,1971]" targetBox="[227,1361,701,1911]" targetPageId="36">
<paragraph id="B56D4F1CFFA6FFC9FF26F816FBFCF865" blockId="36.[186,1399,1945,2007]" pageId="36" pageNumber="153">
<emphasis id="87A6930EFFA6FFC9FF26F816FEFCF800" bold="true" box="[189,279,1945,1971]" pageId="36" pageNumber="153">Fig. 15.</emphasis>
<taxonomicName id="72D2349FFFA6FFC9FE80F816FD1BF800" authority="Wang &amp; Jaitrong" authorityName="Wang &amp; Yong &amp; Jaitrong" authorityYear="2021" box="[283,752,1945,1972]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="153" phylum="Arthropoda" rank="species" species="robusta" status="sp. nov.">
<emphasis id="87A6930EFFA6FFC9FE80F816FDC2F800" box="[283,553,1945,1971]" italics="true" pageId="36" pageNumber="153">Rhopalomastix robusta</emphasis>
Wang &amp; Jaitrong
</taxonomicName>
<taxonomicNameLabel id="9C952E75FFA6FFC9FD6FF815FCA4F807" box="[756,847,1946,1972]" pageId="36" pageNumber="153" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, small worker (ZRC_ENT00013949). 
<emphasis id="87A6930EFFA6FFC9FF21F833FF22F865" bold="true" box="[186,201,1980,2006]" pageId="36" pageNumber="153">a</emphasis>
. Head in full-face view. 
<emphasis id="87A6930EFFA6FFC9FE75F833FE15F865" bold="true" box="[494,510,1980,2006]" pageId="36" pageNumber="153">b</emphasis>
. Body in dorsal view. 
<emphasis id="87A6930EFFA6FFC9FC9CF833FCFFF865" bold="true" box="[775,788,1980,2006]" pageId="36" pageNumber="153">c</emphasis>
. Body in lateral view.
</paragraph>
</caption>
<paragraph id="B56D4F1CFFA7FFC8FF26FD91FDC5FD12" blockId="37.[187,1399,542,673]" pageId="37" pageNumber="154">Head and dorsum of mesosoma generally dark brown with rest of body mostly uniformly lighter brown. In larger workers, head, dorsum of mesosoma, and procoxa blackish or dark reddish brown; petiolar apex and dorsum of postpetiole generally darker brown than their respective lateral faces. Tips of antennae and legs pale yellowish-brown.</paragraph>
</subSubSection>
<subSubSection id="FDC81C97FFA7FFC8FF26FD47FCA0FC2D" pageId="37" pageNumber="154" type="distribution">
<paragraph id="B56D4F1CFFA7FFC8FF26FD47FDEFFD51" blockId="37.[189,516,712,739]" box="[189,516,712,739]" pageId="37" pageNumber="154">
<heading id="EE25F870FFA7FFC8FF26FD47FDEFFD51" bold="true" box="[189,516,712,739]" fontSize="11" level="3" pageId="37" pageNumber="154" reason="3">
<emphasis id="87A6930EFFA7FFC8FF26FD47FDEFFD51" bold="true" box="[189,516,712,739]" pageId="37" pageNumber="154">Distribution and binomics</emphasis>
</heading>
</paragraph>
<paragraph id="B56D4F1CFFA7FFC8FF26FD78FCA0FC2D" blockId="37.[188,1400,759,927]" pageId="37" pageNumber="154">
<collectingCountry id="CDC50F8CFFA7FFC8FF26FD78FECEFCA2" box="[189,293,759,785]" name="Thailand" pageId="37" pageNumber="154">Thailand</collectingCountry>
(
<figureCitation id="2DE95399FFA7FFC8FEACFD78FE76FCA1" box="[311,413,759,786]" captionStart="Fig" captionStartId="37.[191,234,1923,1949]" captionTargetBox="[224,1364,1287,1873]" captionTargetId="figure-368@37.[224,1364,1287,1873]" captionTargetPageId="37" captionText="Fig. 16. Distribution maps of species of Rhopalomastix Forel, 1900 in Thailand. a. New species: R. impithuksai Wang &amp; Jaitrong sp. nov., R. parva Wang &amp; Jaitrong sp. nov., R. robusta Wang &amp; Jaitrong sp. nov. b. R. javana Wheeler, 1929 and R. johorensis Wheeler, 1929." figureDoi="http://doi.org/10.5281/zenodo.4610595" httpUri="https://zenodo.org/record/4610595/files/figure.png" pageId="37" pageNumber="154">Fig. 16a</figureCitation>
). This species is thus far only known from its 
<typeStatus id="6A69F1BEFFA7FFC8FC5DFD77FC12FCA1" box="[966,1017,760,786]" pageId="37" pageNumber="154">type</typeStatus>
series, collected from relatively good secondary forest at moderately high elevation (
<quantity id="722AE2F9FFA7FFC8FCAAFC95FC69FC86" box="[817,898,794,821]" metricMagnitude="2" metricUnit="m" metricValue="9.27" pageId="37" pageNumber="154" unit="m" value="927.0">927 m</quantity>
). The colony was found in bark of a large dipterocarp tree, 
<taxonomicName id="72D2349FFFA7FFC8FE19FCB1FDF0FCE4" box="[386,539,829,855]" class="Magnoliopsida" family="Meliaceae" genus="Toona" kingdom="Plantae" order="Sapindales" pageId="37" pageNumber="154" phylum="Tracheophyta" rank="species" species="ciliata">
<emphasis id="87A6930EFFA7FFC8FE19FCB1FDF0FCE4" box="[386,539,829,855]" italics="true" pageId="37" pageNumber="154">Toona ciliata</emphasis>
</taxonomicName>
, located near the edge of the forest. Members of the colony seemed rather gregarious when found, with numerous individuals moving quickly en masse as a collective group to seek cover when exposed upon removal of a bark layer.
</paragraph>
</subSubSection>
<subSubSection id="FDC81C97FFA7FFCBFF26FC49FD50FD8E" lastPageId="38" lastPageNumber="155" pageId="37" pageNumber="154" type="discussion">
<paragraph id="B56D4F1CFFA7FFC8FF26FC49FEC4FC53" blockId="37.[189,303,966,992]" box="[189,303,966,992]" pageId="37" pageNumber="154">
<heading id="EE25F870FFA7FFC8FF26FC49FEC4FC53" bold="true" box="[189,303,966,992]" fontSize="11" level="3" pageId="37" pageNumber="154" reason="3">
<emphasis id="87A6930EFFA7FFC8FF26FC49FEC4FC53" bold="true" box="[189,303,966,992]" pageId="37" pageNumber="154">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="B56D4F1CFFA7FFCBFF26FC7BFD50FD8E" blockId="37.[186,1401,1012,1214]" lastBlockId="38.[189,1401,265,573]" lastPageId="38" lastPageNumber="155" pageId="37" pageNumber="154">
Smaller workers of 
<taxonomicName id="72D2349FFFA7FFC8FE00FC7AFD37FBBD" authority="Wang &amp; Jaitrong" authorityName="Wang &amp; Yong &amp; Jaitrong" authorityYear="2021" box="[411,732,1012,1038]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="154" phylum="Arthropoda" rank="species" species="robusta" status="sp. nov.">
<emphasis id="87A6930EFFA7FFC8FE00FC7AFDF9FBBD" box="[411,530,1012,1038]" italics="true" pageId="37" pageNumber="154">R. robusta</emphasis>
Wang &amp; Jaitrong
</taxonomicName>
<taxonomicNameLabel id="9C952E75FFA7FFC8FD7BFC7BFCD3FBBD" box="[736,824,1012,1038]" pageId="37" pageNumber="154" rank="species">sp. nov.</taxonomicNameLabel>
are similar to large workers of both 
<taxonomicName id="72D2349FFFA7FFC8FB56FC7AFE6AFB82" authority="Wang &amp; Jaitrong" authorityName="Wang &amp; Yong &amp; Jaitrong" authorityYear="2021" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="154" phylum="Arthropoda" rank="species" species="impithuksai" status="sp. nov.">
<emphasis id="87A6930EFFA7FFC8FB56FC7AFA9FFBBD" box="[1229,1396,1012,1038]" italics="true" pageId="37" pageNumber="154">R. impithuksai</emphasis>
Wang &amp; Jaitrong
</taxonomicName>
<taxonomicNameLabel id="9C952E75FFA7FFC8FE1CFB97FE0AFB81" box="[391,481,1048,1074]" pageId="37" pageNumber="154" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="72D2349FFFA7FFC8FD81FB97FD58FB82" authorityName="Wheeler" authorityYear="1929" box="[538,691,1047,1073]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="154" phylum="Arthropoda" rank="species" species="johorensis">
<emphasis id="87A6930EFFA7FFC8FD81FB97FD58FB82" box="[538,691,1047,1073]" italics="true" pageId="37" pageNumber="154">R. johorensis</emphasis>
</taxonomicName>
. The first two species can be differentiated by the following: 1) striations on dorsum of head uniformly dense from frontal lobes to posterior margin in 
<taxonomicName id="72D2349FFFA7FFC8FB66FBB4FE6DFBC4" authority="Wang &amp; Jaitrong" authorityName="Wang &amp; Yong &amp; Jaitrong" authorityYear="2021" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="154" phylum="Arthropoda" rank="species" species="robusta" status="sp. nov.">
<emphasis id="87A6930EFFA7FFC8FB66FBB4FA9CFBE7" box="[1277,1399,1082,1108]" italics="true" pageId="37" pageNumber="154">R. robusta</emphasis>
Wang &amp; Jaitrong
</taxonomicName>
<taxonomicNameLabel id="9C952E75FFA7FFC8FE17FBD1FE02FBCB" box="[396,489,1118,1144]" pageId="37" pageNumber="154" rank="species">sp. nov.</taxonomicNameLabel>
, but in 
<taxonomicName id="72D2349FFFA7FFC8FDA0FBD1FC5AFBC4" authority="Wang &amp; Jaitrong" authorityName="Wang &amp; Yong &amp; Jaitrong" authorityYear="2021" box="[571,945,1117,1144]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="154" phylum="Arthropoda" rank="species" species="impithuksai" status="sp. nov.">
<emphasis id="87A6930EFFA7FFC8FDA0FBD1FD0FFBC4" box="[571,740,1117,1143]" italics="true" pageId="37" pageNumber="154">R. impithuksai</emphasis>
Wang &amp; Jaitrong
</taxonomicName>
<taxonomicNameLabel id="9C952E75FFA7FFC8FC2CFBD1FBFFFBCB" box="[951,1044,1118,1144]" pageId="37" pageNumber="154" rank="species">sp. nov.</taxonomicNameLabel>
, striations on head dorsum are weaker and more fragmented in posterior half; 2) outline of clypeus in lateral view–entirely flat or weakly convex with no angulate edge in 
<taxonomicName id="72D2349FFFA7FFC8FDD4FB2AFC4BFB0D" authority="Wang &amp; Jaitrong" authorityName="Wang &amp; Yong &amp; Jaitrong" authorityYear="2021" box="[591,928,1188,1214]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="154" phylum="Arthropoda" rank="species" species="robusta" status="sp. nov.">
<emphasis id="87A6930EFFA7FFC8FDD4FB2AFD21FB0D" box="[591,714,1188,1214]" italics="true" pageId="37" pageNumber="154">R. robusta</emphasis>
Wang &amp; Jaitrong
</taxonomicName>
<taxonomicNameLabel id="9C952E75FFA7FFC8FC33FB2BFBE3FB0D" box="[936,1032,1188,1214]" pageId="37" pageNumber="154" rank="species">sp. nov.</taxonomicNameLabel>
, but with short weakly convex dorsal face separated from steep and almost flat anterior face by a weak angulate edge in 
<taxonomicName id="72D2349FFFA4FFCBFB56FE85FE6CFEF5" authority="Wang &amp; Jaitrong" authorityName="Wang &amp; Yong &amp; Jaitrong" authorityYear="2021" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="155" phylum="Arthropoda" rank="species" species="impithuksai" status="sp. nov.">
<emphasis id="87A6930EFFA4FFCBFB56FE85FA9CFE90" box="[1229,1399,265,291]" italics="true" pageId="38" pageNumber="155">R. impithuksai</emphasis>
Wang &amp; Jaitrong
</taxonomicName>
<taxonomicNameLabel id="9C952E75FFA4FFCBFE14FEA2FE05FEF4" box="[399,494,301,327]" pageId="38" pageNumber="155" rank="species">sp. nov.</taxonomicNameLabel>
; 3) head in full-face view as long as wide or sometimes wider than long for 
<taxonomicName id="72D2349FFFA4FFCBFF25FEDFFDE8FEDA" authority="Wang &amp; Jaitrong" authorityName="Wang &amp; Yong &amp; Jaitrong" authorityYear="2021" box="[190,515,335,362]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="155" phylum="Arthropoda" rank="species" species="robusta" status="sp. nov.">
<emphasis id="87A6930EFFA4FFCBFF25FEDFFEDCFEDA" box="[190,311,335,361]" italics="true" pageId="38" pageNumber="155">R. robusta</emphasis>
Wang &amp; Jaitrong
</taxonomicName>
<taxonomicNameLabel id="9C952E75FFA4FFCBFD91FEDFFD8CFED9" box="[522,615,336,362]" pageId="38" pageNumber="155" rank="species">sp. nov.</taxonomicNameLabel>
, whereas for 
<taxonomicName id="72D2349FFFA4FFCBFC99FEDFFB92FEDA" authority="Wang &amp; Jaitrong" authorityName="Wang &amp; Yong &amp; Jaitrong" authorityYear="2021" box="[770,1145,335,362]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="155" phylum="Arthropoda" rank="species" species="impithuksai" status="sp. nov.">
<emphasis id="87A6930EFFA4FFCBFC99FEDFFC40FEDA" box="[770,939,335,361]" italics="true" pageId="38" pageNumber="155">R. impithuksai</emphasis>
Wang &amp; Jaitrong
</taxonomicName>
<taxonomicNameLabel id="9C952E75FFA4FFCBFBE4FEDFFB37FED9" box="[1151,1244,336,362]" pageId="38" pageNumber="155" rank="species">sp. nov.</taxonomicNameLabel>
, head in fullface view always longer than wide; 4) median furrow on head dorsum starting from between frontal lobes quite long and relatively deep in 
<taxonomicName id="72D2349FFFA4FFCBFD1FFE18FC20FE03" authority="Wang &amp; Jaitrong" authorityName="Wang &amp; Yong &amp; Jaitrong" authorityYear="2021" box="[644,971,406,432]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="155" phylum="Arthropoda" rank="species" species="robusta" status="sp. nov.">
<emphasis id="87A6930EFFA4FFCBFD1FFE18FD16FE03" box="[644,765,406,432]" italics="true" pageId="38" pageNumber="155">R. robusta</emphasis>
Wang &amp; Jaitrong
</taxonomicName>
<taxonomicNameLabel id="9C952E75FFA4FFCBFC49FE19FBC4FE03" box="[978,1071,406,432]" pageId="38" pageNumber="155" rank="species">sp. nov.</taxonomicNameLabel>
, either extending till or past transverse midline of head, in 
<taxonomicName id="72D2349FFFA4FFCBFDB6FE35FC47FE60" authority="Wang &amp; Jaitrong" authorityName="Wang &amp; Yong &amp; Jaitrong" authorityYear="2021" box="[557,940,441,468]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="155" phylum="Arthropoda" rank="species" species="impithuksai" status="sp. nov.">
<emphasis id="87A6930EFFA4FFCBFDB6FE35FD3CFE60" box="[557,727,441,467]" italics="true" pageId="38" pageNumber="155">R. impithuksai</emphasis>
Wang &amp; Jaitrong
</taxonomicName>
<taxonomicNameLabel id="9C952E75FFA4FFCBFC2EFE35FBF9FE67" box="[949,1042,442,468]" pageId="38" pageNumber="155" rank="species">sp. nov.</taxonomicNameLabel>
the median furrow is shorter, usually narrower and shallower in its posterior half, often fading out before transverse midline of head. Points (3) and (4) can also be used to distinguish between 
<taxonomicName id="72D2349FFFA4FFCBFCEDFD8FFB2AFDAA" authority="Wang &amp; Jaitrong" authorityName="Wang &amp; Yong &amp; Jaitrong" authorityYear="2021" box="[886,1217,511,538]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="155" phylum="Arthropoda" rank="species" species="robusta" status="sp. nov.">
<emphasis id="87A6930EFFA4FFCBFCEDFD8FFC1AFDAA" box="[886,1009,511,537]" italics="true" pageId="38" pageNumber="155">R. robusta</emphasis>
Wang &amp; Jaitrong
</taxonomicName>
<taxonomicNameLabel id="9C952E75FFA4FFCBFB52FD8FFACEFDA9" box="[1225,1317,512,538]" pageId="38" pageNumber="155" rank="species">sp. nov.</taxonomicNameLabel>
(small workers) and 
<taxonomicName id="72D2349FFFA4FFCBFEC5FDACFE13FD8F" authorityName="Wheeler" authorityYear="1929" box="[350,504,546,572]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="155" phylum="Arthropoda" rank="species" species="johorensis">
<emphasis id="87A6930EFFA4FFCBFEC5FDACFE13FD8F" box="[350,504,546,572]" italics="true" pageId="38" pageNumber="155">R. johorensis</emphasis>
</taxonomicName>
(large workers).
</paragraph>
</subSubSection>
</treatment>
</document>