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<document id="901043A6772A9456058D81EDC111DC0A" ID-CLB-Dataset="7699" ID-DOI="10.5852/ejt.2021.739.1271" ID-GBIF-Dataset="99f6713d-6ef6-4bee-905b-b987df8832d4" ID-ISSN="2118-9773" ID-Zenodo-Dep="4610565" ID-ZooBank="81383BD5-6040-4156-BD09-A2704E5D9DA7" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615988829382" checkinUser="plazi" docAuthor="Wang, Wendy Y., Yong, Gordon W. J. &amp; Jaitrong, Weeyawat" docDate="2021" docId="3D7BFE0AFF89FFFEFDA2F95CFAA4FA39" docLanguage="en" docName="ejt-739_117-157.pdf" docOrigin="European Journal of Taxonomy 739 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.739.1271" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Rhopalomastix javana Wheeler 1929" docType="treatment" docVersion="10" lastPageNumber="136" masterDocId="C1428672FF82FFEDFF9BFF8FFFEBFFB3" masterDocTitle="Revision of the elusive ant genus Rhopalomastix (Hymenoptera, Formicidae, Myrmicinae) in Thailand based on morphology and DNA barcodes, with descriptions of three new species" masterLastPageNumber="157" masterPageNumber="117" pageNumber="128" updateTime="1698933474431" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="DBFCAB9C12D72BA112DF96916E103CD8">Revision of the elusive ant genus Rhopalomastix (Hymenoptera, Formicidae, Myrmicinae) in Thailand based on morphology and DNA barcodes, with descriptions of three new species</mods:title>
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<mods:namePart id="840213827A0DDCAD49C53B77011505AD">Wang, Wendy Y.</mods:namePart>
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<mods:affiliation id="A7E1F6C6A90C72A6DB0247F2E6868100">Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, Singapore 117377. Department of Biological Sciences, Faculty of Science, National University of Singapore, 16 Science Drive 4, Singapore 117558. Natural History Museum, National Science Museum, Technopolis, Khlong 5, Khlong Luang, Pathum Thani, 12120 Thailand.</mods:affiliation>
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<mods:namePart id="C813A1E800BE88FBAEC9044478EA686C">Yong, Gordon W. J.</mods:namePart>
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<mods:namePart id="785808C6B9ED1838290CF5CD01576C6A">Jaitrong, Weeyawat</mods:namePart>
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<taxonomicName id="72D2349FFF89FFE6FDA2F95CFC10F95E" authority="Wheeler, 1929" authorityName="Wheeler" authorityYear="1929" box="[569,1019,1747,1774]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="javana">
<emphasis id="87A6930EFF89FFE6FDA2F95CFCA3F95E" bold="true" box="[569,840,1747,1774]" italics="true" pageId="11" pageNumber="128">Rhopalomastix javana</emphasis>
<bibRefCitation id="D14332EDFF89FFE6FCD4F95CFC10F95E" author="Wheeler W. M." box="[847,1019,1747,1773]" pageId="11" pageNumber="128" pagination="95 - 101" refId="ref24237" refString="Wheeler W. M. 1929. The ant genus Rhopalomastix. Psyche 36 (2): 95 - 101." type="journal article" year="1929">Wheeler, 1929</bibRefCitation>
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<paragraph id="B56D4F1CFF89FFE6FF7EF979FAA5F8A3" blockId="11.[229,1358,1747,1809]" box="[229,1358,1782,1809]" pageId="11" pageNumber="128">
<figureCitation id="2DE95399FF89FFE6FF7EF979FED6F8A3" box="[229,317,1782,1808]" captionStart="Fig" captionStartId="5.[189,232,1816,1842]" captionTargetBox="[227,1361,973,1778]" captionTargetId="figure-170@5.[227,1361,973,1778]" captionTargetPageId="5" captionText="Fig. 2. Closeups of external mandibular surface for workers of 5 Thai species of Rhopalomastix Forel, 1900 (ZRC); white arrows indicate distinct striations on the external mandibular face. a. R. impithuksai Wang &amp; Jaitrong sp. nov., paratype (ZRC_ENT00007608). b. R. robusta Wang &amp; Jaitrong sp. nov., paratype (ZRC_ENT00013949). c. R. javana Wheeler, 1929, non-type (ZRC_ENT00000932). d. R. johorensis Wheeler, 1929, non-type (ZRC_ENT00007874). e. R. parva Wang &amp; Jaitrong sp. nov., paratype (ZRC_ENT00013948)." figureDoi="http://doi.org/10.5281/zenodo.4610567" httpUri="https://zenodo.org/record/4610567/files/figure.png" pageId="11" pageNumber="128">Figs 2c</figureCitation>
, 
<figureCitation id="2DE95399FF89FFE6FED1F979FEB2F8A2" box="[330,345,1782,1809]" captionStart="Fig" captionStartId="14.[189,232,1913,1939]" captionTargetBox="[227,1361,933,1860]" captionTargetId="figure-234@14.[227,1361,933,1860]" captionTargetPageId="14" captionText="Fig. 5. Rhopalomastix javana Wheeler, 1929, non-type, worker, West Thailand (ZRC_ENT00000932). a. Head in full-face view. b. Body in dorsal view. c. Close-up of clypeus and head in profile. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610573" httpUri="https://zenodo.org/record/4610573/files/figure.png" pageId="11" pageNumber="128">5</figureCitation>
(worker), 6 (queen), 7 (male), 16b. Also see 
<bibRefCitation id="D14332EDFF89FFE6FCF3F979FBA6F8A3" author="Wang W. Y. &amp; Yong G. W. &amp; Jaitrong W." box="[872,1101,1782,1808]" pageId="11" pageNumber="128" pagination="301 - 340" refId="ref24028" refString="Wang W. Y., Yong G. W. &amp; Jaitrong W. 2018 b. The ant genus Rhopalomastix (Hymenoptera: Formicidae: Myrmicinae) in Southeast Asia, with descriptions of four new species from Singapore based on morphology and DNA barcoding. Zootaxa 4532 (3): 301 - 340. https: // doi. org / 10.11646 / zootaxa. 4532.3.1" type="journal article" year="2018">
Wang 
<emphasis id="87A6930EFF89FFE6FC28F978FC05F8A3" box="[947,1006,1782,1808]" italics="true" pageId="11" pageNumber="128">et al.</emphasis>
(2018b
</bibRefCitation>
: figs 20–23) for male.
</paragraph>
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<subSubSection id="FDC81C97FF89FFE6FF26F8B8FB13F86B" pageId="11" pageNumber="128" type="reference_group">
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<treatmentCitation id="3473690DFF89FFE6FF26F8B8FCB4F8E2" author="Wheeler W. M." box="[189,863,1847,1873]" page="96" pageId="11" pageNumber="128" year="1929">
<taxonomicName id="72D2349FFF89FFE6FF26F8B8FCB4F8E2" ID-CoL="5KX5Y" authority="Wheeler, 1929: 96" authorityName="Wheeler" authorityPageNumber="96" authorityYear="1929" box="[189,863,1847,1873]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="128" phylum="Arthropoda" rank="subSpecies" species="rothneyi" subSpecies="javana">
<emphasis id="87A6930EFF89FFE6FF26F8B8FE3EF8E2" box="[189,469,1847,1873]" italics="true" pageId="11" pageNumber="128">Rhopalomastix rothneyi</emphasis>
subsp. 
<emphasis id="87A6930EFF89FFE6FDB5F8B7FD95F8E2" box="[558,638,1848,1873]" italics="true" pageId="11" pageNumber="128">javana</emphasis>
<bibRefCitation id="D14332EDFF89FFE6FD1FF8B8FCB4F8E2" author="Wheeler W. M." box="[644,863,1847,1873]" pageId="11" pageNumber="128" pagination="95 - 101" refId="ref24237" refString="Wheeler W. M. 1929. The ant genus Rhopalomastix. Psyche 36 (2): 95 - 101." type="journal article" year="1929">Wheeler, 1929: 96</bibRefCitation>
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, fig. 1 (w.q.m).
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<taxonomicName id="72D2349FFF89FFE6FF26F8F7FD6DF821" ID-CoL="5KX5Y" authorityName="Wheeler" authorityYear="1929" box="[189,646,1912,1938]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="128" phylum="Arthropoda" rank="subSpecies" species="rothneyi" subSpecies="javana">
<emphasis id="87A6930EFF89FFE6FF26F8F7FE33F821" box="[189,472,1912,1938]" italics="true" pageId="11" pageNumber="128">Rhopalomastix rothneyi</emphasis>
subsp. 
<emphasis id="87A6930EFF89FFE6FDADF8F6FD6DF821" box="[566,646,1913,1938]" italics="true" pageId="11" pageNumber="128">javana</emphasis>
</taxonomicName>
<emphasis id="87A6930EFF89FFE6FD14F8F6FD75F821" box="[655,670,1913,1938]" italics="true" pageId="11" pageNumber="128">–</emphasis>
<treatmentCitation id="3473690DFF89FFE6FD33F8F7FBF9F821" author="Chapman J. W. &amp; Capco S. R." box="[680,1042,1912,1939]" page="111" pageId="11" pageNumber="128" year="1951">
<bibRefCitation id="D14332EDFF89FFE6FD33F8F7FBF9F821" author="Chapman J. W. &amp; Capco S. R." box="[680,1042,1912,1939]" pageId="11" pageNumber="128" pagination="1 - 327" refId="ref23389" refString="Chapman J. W. &amp; Capco S. R. 1951. Check list of the ants (Hymenoptera: Formicidae) of Asia. Monographs of the Institute of Science and Technology, Manila 1: 1 - 327" type="journal article" year="1951">Chapman &amp; Capco 1951: 111</bibRefCitation>
</treatmentCitation>
. — 
<treatmentCitation id="3473690DFF89FFE6FBDCF8F7FAF5F821" author="Bolton B." box="[1095,1310,1912,1939]" page="377" pageId="11" pageNumber="128" year="1995">
<bibRefCitation id="D14332EDFF89FFE6FBDCF8F7FAF5F821" author="Bolton B." box="[1095,1310,1912,1939]" pageId="11" pageNumber="128" refId="ref23342" refString="Bolton B. 1995. A New General Catalogue of the Ants of the World. Harvard University Press, Cambridge, MA." type="book" year="1995">Bolton 1995: 377</bibRefCitation>
</treatmentCitation>
. — 
<treatmentCitation id="3473690DFF89FFE6FAC8F8F7FE8BF806" author="Xu Z." page="131" pageId="11" pageNumber="128" year="1999">
<bibRefCitation id="D14332EDFF89FFE6FAC8F8F7FE8BF806" author="Xu Z." pageId="11" pageNumber="128" pagination="129 - 136" refId="ref24259" refString="Xu Z. 1999. Systematic studies on the ant genera of Carebara, Rhopalomastix and Kartidris in China (Hymenoptera: Formicidae: Myrmicinae). Acta Biologica Plateau Sinica 14: 129 - 136." type="journal article" year="1999">Xu 1999: 131</bibRefCitation>
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(in key).
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</paragraph>
<paragraph id="B56D4F1CFF89FFE6FF26F832FB13F86B" blockId="11.[189,1399,1912,2008]" box="[189,1272,1981,2008]" pageId="11" pageNumber="128">
<treatmentCitationGroup id="95C26832FF89FFE6FF26F832FB13F86B" box="[189,1272,1981,2008]" pageId="11" pageNumber="128">
<emphasis id="87A6930EFF89FFE6FF26F832FE31F864" box="[189,474,1981,2007]" italics="true" pageId="11" pageNumber="128">
<taxonomicName id="72D2349FFF89FFE6FF26F832FE2FF864" authorityName="Wheeler" authorityYear="1929" box="[189,452,1981,2007]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="javana">Rhopalomastix javana</taxonomicName>
–
</emphasis>
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<bibRefCitation id="D14332EDFF89FFE6FE7AF831FD1DF864" author="Wang W. Y. &amp; Yong G. W. &amp; Jaitrong W." box="[481,758,1981,2008]" pageId="11" pageNumber="128" pagination="301 - 340" refId="ref24028" refString="Wang W. Y., Yong G. W. &amp; Jaitrong W. 2018 b. The ant genus Rhopalomastix (Hymenoptera: Formicidae: Myrmicinae) in Southeast Asia, with descriptions of four new species from Singapore based on morphology and DNA barcoding. Zootaxa 4532 (3): 301 - 340. https: // doi. org / 10.11646 / zootaxa. 4532.3.1" type="journal article" year="2018">
Wang 
<emphasis id="87A6930EFF89FFE6FDB7F831FD8CF864" box="[556,615,1981,2007]" italics="true" pageId="11" pageNumber="128">et al.</emphasis>
2018b: 310
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, figs 8–11 (raised to species, redescription).
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</paragraph>
</subSubSection>
<subSubSection id="FDC81C97FF8EFFE1FF26FE87FC7DF959" pageId="12" pageNumber="129" type="diagnosis">
<paragraph id="B56D4F1CFF8EFFE1FF26FE87FEDDFE91" blockId="12.[189,310,264,290]" box="[189,310,264,290]" pageId="12" pageNumber="129">
<heading id="EE25F870FF8EFFE1FF26FE87FEDDFE91" bold="true" box="[189,310,264,290]" fontSize="11" level="3" pageId="12" pageNumber="129" reason="3">
<emphasis id="87A6930EFF8EFFE1FF26FE87FEDDFE91" bold="true" box="[189,310,264,290]" pageId="12" pageNumber="129">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="B56D4F1CFF8EFFE1FF71FEB8FEA7FEE2" blockId="12.[187,1399,311,760]" box="[234,332,311,337]" pageId="12" pageNumber="129">
<emphasis id="87A6930EFF8EFFE1FF71FEB8FEA7FEE2" bold="true" box="[234,332,311,337]" pageId="12" pageNumber="129">Worker</emphasis>
</paragraph>
<paragraph id="B56D4F1CFF8EFFE1FF26FED5FA9CFD44" blockId="12.[187,1399,311,760]" pageId="12" pageNumber="129">
Workers monomorphic with little size variation (
<typeStatus id="6A69F1BEFF8EFFE1FCB8FED5FC8CFEC7" box="[803,871,346,372]" pageId="12" pageNumber="129">types</typeStatus>
: HL 0.48–0.52, HW 0.45–0.48; non-types HL 0.46–0.48, HW 0.41–0.44). Head in full-face view subrectangular, longer than wide, posterior margin with a shallow median concavity (
<figureCitation id="2DE95399FF8EFFE1FD5DFE2EFCCBFE08" box="[710,800,416,443]" captionStart="Fig" captionStartId="14.[189,232,1913,1939]" captionTargetBox="[227,1361,933,1860]" captionTargetId="figure-234@14.[227,1361,933,1860]" captionTargetPageId="14" captionText="Fig. 5. Rhopalomastix javana Wheeler, 1929, non-type, worker, West Thailand (ZRC_ENT00000932). a. Head in full-face view. b. Body in dorsal view. c. Close-up of clypeus and head in profile. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610573" httpUri="https://zenodo.org/record/4610573/files/figure.png" pageId="12" pageNumber="129">Fig. 5a</figureCitation>
). In profile, clypeus broadly and weakly convex (
<figureCitation id="2DE95399FF8EFFE1FF5DFE4BFEF7FE6C" box="[198,284,452,479]" captionStart="Fig" captionStartId="14.[189,232,1913,1939]" captionTargetBox="[227,1361,933,1860]" captionTargetId="figure-234@14.[227,1361,933,1860]" captionTargetPageId="14" captionText="Fig. 5. Rhopalomastix javana Wheeler, 1929, non-type, worker, West Thailand (ZRC_ENT00000932). a. Head in full-face view. b. Body in dorsal view. c. Close-up of clypeus and head in profile. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610573" httpUri="https://zenodo.org/record/4610573/files/figure.png" pageId="12" pageNumber="129">Fig. 5c</figureCitation>
); dorsum of head finely striate (
<figureCitation id="2DE95399FF8EFFE1FD00FE4BFD1AFE6C" box="[667,753,452,479]" captionStart="Fig" captionStartId="14.[189,232,1913,1939]" captionTargetBox="[227,1361,933,1860]" captionTargetId="figure-234@14.[227,1361,933,1860]" captionTargetPageId="14" captionText="Fig. 5. Rhopalomastix javana Wheeler, 1929, non-type, worker, West Thailand (ZRC_ENT00000932). a. Head in full-face view. b. Body in dorsal view. c. Close-up of clypeus and head in profile. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610573" httpUri="https://zenodo.org/record/4610573/files/figure.png" pageId="12" pageNumber="129">Fig. 5a</figureCitation>
); striations on posterior half of head generally weaker and more fragmented; ventral face of head largely superficially reticulate. In dorsal view, mesosoma subrectangular with almost parallel sides and weakly concave posterior propodeal margin (
<figureCitation id="2DE95399FF8EFFE1FA96FD85FA8CFD96" box="[1293,1383,522,549]" captionStart="Fig" captionStartId="14.[189,232,1913,1939]" captionTargetBox="[227,1361,933,1860]" captionTargetId="figure-234@14.[227,1361,933,1860]" captionTargetPageId="14" captionText="Fig. 5. Rhopalomastix javana Wheeler, 1929, non-type, worker, West Thailand (ZRC_ENT00000932). a. Head in full-face view. b. Body in dorsal view. c. Close-up of clypeus and head in profile. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610573" httpUri="https://zenodo.org/record/4610573/files/figure.png" pageId="12" pageNumber="129">Fig. 5b</figureCitation>
); posterolateral corners of propodeum bluntly angulate, slightly projecting posteriorly and distinctly, though shallowly, differentiated from inner posterior propodeal face. Propodeal junction in lateral view roundly obtuse (
<figureCitation id="2DE95399FF8EFFE1FEE7FDFBFE39FD3C" box="[380,466,628,655]" captionStart="Fig" captionStartId="14.[189,232,1913,1939]" captionTargetBox="[227,1361,933,1860]" captionTargetId="figure-234@14.[227,1361,933,1860]" captionTargetPageId="14" captionText="Fig. 5. Rhopalomastix javana Wheeler, 1929, non-type, worker, West Thailand (ZRC_ENT00000932). a. Head in full-face view. b. Body in dorsal view. c. Close-up of clypeus and head in profile. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610573" httpUri="https://zenodo.org/record/4610573/files/figure.png" pageId="12" pageNumber="129">Fig. 5d</figureCitation>
); posterior propodeal declivitous face weakly marginate and shallowly concave (
<figureCitation id="2DE95399FF8EFFE1FF5EFD18FEDDFD01" box="[197,310,663,690]" captionStart="Fig" captionStartId="14.[189,232,1913,1939]" captionTargetBox="[227,1361,933,1860]" captionTargetId="figure-234@14.[227,1361,933,1860]" captionTargetPageId="14" captionText="Fig. 5. Rhopalomastix javana Wheeler, 1929, non-type, worker, West Thailand (ZRC_ENT00000932). a. Head in full-face view. b. Body in dorsal view. c. Close-up of clypeus and head in profile. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610573" httpUri="https://zenodo.org/record/4610573/files/figure.png" pageId="12" pageNumber="129">Fig. 5b, d</figureCitation>
); propodeal dorsum differentiated from posterior face by weakly marginate edge.Anterior face of petiole weakly concave (
<figureCitation id="2DE95399FF8EFFE1FE67FD35FDB9FD66" box="[508,594,698,725]" captionStart="Fig" captionStartId="14.[189,232,1913,1939]" captionTargetBox="[227,1361,933,1860]" captionTargetId="figure-234@14.[227,1361,933,1860]" captionTargetPageId="14" captionText="Fig. 5. Rhopalomastix javana Wheeler, 1929, non-type, worker, West Thailand (ZRC_ENT00000932). a. Head in full-face view. b. Body in dorsal view. c. Close-up of clypeus and head in profile. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610573" httpUri="https://zenodo.org/record/4610573/files/figure.png" pageId="12" pageNumber="129">Fig. 5d</figureCitation>
), with fairly dense appressed and decumbent hairs. Head, mesosoma dorsum and gaster darker brown than rest of body; dorsum of head sometimes with blackish undertones.
</paragraph>
<paragraph id="B56D4F1CFF8EFFE1FF71FC90FED0FC8A" blockId="12.[187,1400,799,1283]" box="[234,315,799,825]" pageId="12" pageNumber="129">
<emphasis id="87A6930EFF8EFFE1FF71FC90FED0FC8A" bold="true" box="[234,315,799,825]" pageId="12" pageNumber="129">Queen</emphasis>
</paragraph>
<paragraph id="B56D4F1CFF8EFFE1FF26FCCDFC25FAB1" blockId="12.[187,1400,799,1283]" pageId="12" pageNumber="129">
Small-medium size relative to 
<taxonomicName id="72D2349FFF8EFFE1FDB2FCCCFD29FCEF" authorityName="Wheeler" authorityYear="1929" box="[553,706,834,860]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="129" phylum="Arthropoda" rank="species" species="johorensis">
<emphasis id="87A6930EFF8EFFE1FDB2FCCCFD29FCEF" box="[553,706,834,860]" italics="true" pageId="12" pageNumber="129">R. johorensis</emphasis>
</taxonomicName>
queen (non-types: HL 0.48–0.49, HW 0.44–0.45), head in full-face view subrectangular and slightly narrower anteriorly (
<figureCitation id="2DE95399FF8EFFE1FC38FCEAFC13FC33" box="[931,1016,869,896]" captionStart="Fig" captionStartId="16.[190,233,1926,1952]" captionTargetBox="[227,1361,1098,1871]" captionTargetId="figure-312@16.[227,1361,1097,1871]" captionTargetPageId="16" captionText="Fig. 6. Rhopalomastix javana Wheeler, 1929, non-type, queen, Northeast Thailand (THNHM-I-21751). a. Head in full-face view. b. Body in dorsal view. c. Close-up of mesosoma in dorsal view. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610575" httpUri="https://zenodo.org/record/4610575/files/figure.png" pageId="12" pageNumber="129">Fig. 6a</figureCitation>
). Anterior clypeal face in profile weakly and broadly convex, sometimes almost flat (
<figureCitation id="2DE95399FF8EFFE1FCA8FC07FC65FC10" box="[819,910,904,931]" captionStart="Fig" captionStartId="16.[190,233,1926,1952]" captionTargetBox="[227,1361,1098,1871]" captionTargetId="figure-312@16.[227,1361,1097,1871]" captionTargetPageId="16" captionText="Fig. 6. Rhopalomastix javana Wheeler, 1929, non-type, queen, Northeast Thailand (THNHM-I-21751). a. Head in full-face view. b. Body in dorsal view. c. Close-up of mesosoma in dorsal view. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610575" httpUri="https://zenodo.org/record/4610575/files/figure.png" pageId="12" pageNumber="129">Fig. 6d</figureCitation>
). Posterior half and ventral face of head largely smooth and shiny, with feeble striations on frons and vertex. Dorsal outline of mesosoma weakly and broadly convex; propodeal junction roundly obtuse and indistinct (
<figureCitation id="2DE95399FF8EFFE1FC61FC40FBB9FC59" box="[1018,1106,975,1002]" captionStart="Fig" captionStartId="16.[190,233,1926,1952]" captionTargetBox="[227,1361,1098,1871]" captionTargetId="figure-312@16.[227,1361,1097,1871]" captionTargetPageId="16" captionText="Fig. 6. Rhopalomastix javana Wheeler, 1929, non-type, queen, Northeast Thailand (THNHM-I-21751). a. Head in full-face view. b. Body in dorsal view. c. Close-up of mesosoma in dorsal view. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610575" httpUri="https://zenodo.org/record/4610575/files/figure.png" pageId="12" pageNumber="129">Fig. 6d</figureCitation>
). In dorsal view, anterior margin of pronotal disc broadly convex with roundly obtuse humeral corners; mesoscutum slightly wider than long; propodeum much wider than long; propodeal dorsum weakly convex and rounding into steep posterior declivity at indistinct rounded posterior edge; posterior propodeal face with shallow median concavity (
<figureCitation id="2DE95399FF8EFFE1FEDBFBD3FE7FFBC4" box="[320,404,1116,1143]" captionStart="Fig" captionStartId="16.[190,233,1926,1952]" captionTargetBox="[227,1361,1098,1871]" captionTargetId="figure-312@16.[227,1361,1097,1871]" captionTargetPageId="16" captionText="Fig. 6. Rhopalomastix javana Wheeler, 1929, non-type, queen, Northeast Thailand (THNHM-I-21751). a. Head in full-face view. b. Body in dorsal view. c. Close-up of mesosoma in dorsal view. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610575" httpUri="https://zenodo.org/record/4610575/files/figure.png" pageId="12" pageNumber="129">Fig. 6c</figureCitation>
). Dorsum of mesosoma largely finely striate and shiny; lateral-most triangular area of mesoscutum immediately above parascutellar carina mostly smooth and shining (
<figureCitation id="2DE95399FF8EFFE1FB1AFBF0FB33FB29" box="[1153,1240,1151,1178]" captionStart="Fig" captionStartId="16.[190,233,1926,1952]" captionTargetBox="[227,1361,1098,1871]" captionTargetId="figure-312@16.[227,1361,1097,1871]" captionTargetPageId="16" captionText="Fig. 6. Rhopalomastix javana Wheeler, 1929, non-type, queen, Northeast Thailand (THNHM-I-21751). a. Head in full-face view. b. Body in dorsal view. c. Close-up of mesosoma in dorsal view. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610575" httpUri="https://zenodo.org/record/4610575/files/figure.png" pageId="12" pageNumber="129">Fig. 6c</figureCitation>
); apicodorsal faces of petiole and postpetiole feebly and superficially reticulate(
<figureCitation id="2DE95399FF8EFFE1FC48FB2DFBAEFB0E" box="[979,1093,1186,1213]" captionStart="Fig" captionStartId="16.[190,233,1926,1952]" captionTargetBox="[227,1361,1098,1871]" captionTargetId="figure-312@16.[227,1361,1097,1871]" captionTargetPageId="16" captionText="Fig. 6. Rhopalomastix javana Wheeler, 1929, non-type, queen, Northeast Thailand (THNHM-I-21751). a. Head in full-face view. b. Body in dorsal view. c. Close-up of mesosoma in dorsal view. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610575" httpUri="https://zenodo.org/record/4610575/files/figure.png" pageId="12" pageNumber="129">Fig. 6b, d</figureCitation>
), each with a central area that is smooth and shining (
<figureCitation id="2DE95399FF8EFFE1FD93FB4AFD89FB53" box="[520,610,1221,1248]" captionStart="Fig" captionStartId="16.[190,233,1926,1952]" captionTargetBox="[227,1361,1098,1871]" captionTargetId="figure-312@16.[227,1361,1097,1871]" captionTargetPageId="16" captionText="Fig. 6. Rhopalomastix javana Wheeler, 1929, non-type, queen, Northeast Thailand (THNHM-I-21751). a. Head in full-face view. b. Body in dorsal view. c. Close-up of mesosoma in dorsal view. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610575" httpUri="https://zenodo.org/record/4610575/files/figure.png" pageId="12" pageNumber="129">Fig. 6b</figureCitation>
). Body largely uniformly dark brown; head, dorsum of mesosoma and gaster slightly darker brown, tibia and tarsus yellowish brown.
</paragraph>
<paragraph id="B56D4F1CFF8EFFE1FF71FAA5FEC1FAF7" blockId="12.[189,1399,1322,1770]" box="[234,298,1322,1348]" pageId="12" pageNumber="129">
<emphasis id="87A6930EFF8EFFE1FF71FAA5FEC1FAF7" bold="true" box="[234,298,1322,1348]" pageId="12" pageNumber="129">Male</emphasis>
</paragraph>
<paragraph id="B56D4F1CFF8EFFE1FF26FAC2FC7DF959" blockId="12.[189,1399,1322,1770]" pageId="12" pageNumber="129">
Small size (non-types: HL 0.39–0.41, HW 0.38–0.4). Head in full-face view sub-spherical, posterior margin broadly and strongly rounded (
<bibRefCitation id="D14332EDFF8EFFE1FD10FAFFFC84FA39" author="Wang W. Y. &amp; Yong G. W. &amp; Jaitrong W." box="[651,879,1392,1419]" pageId="12" pageNumber="129" pagination="301 - 340" refId="ref24028" refString="Wang W. Y., Yong G. W. &amp; Jaitrong W. 2018 b. The ant genus Rhopalomastix (Hymenoptera: Formicidae: Myrmicinae) in Southeast Asia, with descriptions of four new species from Singapore based on morphology and DNA barcoding. Zootaxa 4532 (3): 301 - 340. https: // doi. org / 10.11646 / zootaxa. 4532.3.1" type="journal article" year="2018">
Wang 
<emphasis id="87A6930EFF8EFFE1FD46FAFEFCFBFA39" box="[733,784,1392,1418]" italics="true" pageId="12" pageNumber="129">et al</emphasis>
. 2018b
</bibRefCitation>
: fig. 21); in lateral view, head with dorsum gently inclined posteriorly; clypeus in profile with distinct dorsal and anterior faces separated by a roundly angulate edge, dorsal face gently sloped downwards and nearly flat, anterior face steep and very weakly convex or flat (
<figureCitation id="2DE95399FF8EFFE1FD9FFA55FDB1FA47" box="[516,602,1498,1524]" captionStart="Fig" captionStartId="18.[189,232,1956,1982]" captionTargetBox="[258,1330,1395,1916]" captionTargetId="figure-340@18.[258,1330,1395,1916]" captionTargetPageId="18" captionText="Fig. 7. Rhopalomastix javana Wheeler, 1929, non-type, ♂, Singapore (ZRC_HYM_0000576). a. Head and clypeus in profile. b. Close-up of genitalia in profile." figureDoi="http://doi.org/10.5281/zenodo.4610577" httpUri="https://zenodo.org/record/4610577/files/figure.png" pageId="12" pageNumber="129">Fig. 7a</figureCitation>
). Median ocellus subcircular, less elongate and slightly smaller than lateral ocelli; frontal lobes distinctly raised and differentiated from frontal supraclypeal area; posterior apex of supraclypeal area bluntly rounded (
<bibRefCitation id="D14332EDFF8EFFE1FD54F9AFFC5EF989" author="Wang W. Y. &amp; Yong G. W. &amp; Jaitrong W." box="[719,949,1568,1594]" pageId="12" pageNumber="129" pagination="301 - 340" refId="ref24028" refString="Wang W. Y., Yong G. W. &amp; Jaitrong W. 2018 b. The ant genus Rhopalomastix (Hymenoptera: Formicidae: Myrmicinae) in Southeast Asia, with descriptions of four new species from Singapore based on morphology and DNA barcoding. Zootaxa 4532 (3): 301 - 340. https: // doi. org / 10.11646 / zootaxa. 4532.3.1" type="journal article" year="2018">
Wang 
<emphasis id="87A6930EFF8EFFE1FC84F9AEFCB6F989" box="[799,861,1568,1594]" italics="true" pageId="12" pageNumber="129">et al.</emphasis>
2018b
</bibRefCitation>
: fig. 21). In posterior view, posterior propodeal face indistinctly marginate, weakly differentiated from lateral propodeal face by indistinct rounded lateral margin. In lateral view, petiole short and subtrapezoidal, longer than high, anterior margin long, steep and weakly concave, apical dorsum weakly convex, rounding into short posterior face at indistinct rounded edge (
<bibRefCitation id="D14332EDFF8EFFE1FDB5F922FCECF974" author="Wang W. Y. &amp; Yong G. W. &amp; Jaitrong W." box="[558,775,1709,1735]" pageId="12" pageNumber="129" pagination="301 - 340" refId="ref24028" refString="Wang W. Y., Yong G. W. &amp; Jaitrong W. 2018 b. The ant genus Rhopalomastix (Hymenoptera: Formicidae: Myrmicinae) in Southeast Asia, with descriptions of four new species from Singapore based on morphology and DNA barcoding. Zootaxa 4532 (3): 301 - 340. https: // doi. org / 10.11646 / zootaxa. 4532.3.1" type="journal article" year="2018">
Wang 
<emphasis id="87A6930EFF8EFFE1FDE2F921FD59F974" box="[633,690,1709,1735]" italics="true" pageId="12" pageNumber="129">et al.</emphasis>
2018b
</bibRefCitation>
: fig. 20). Funiculus of antenna uniformly light brown; legs greyish brown with joints and tarsi pale brownish yellow.
</paragraph>
</subSubSection>
<subSubSection id="FDC81C97FF8EFFE0FF26F89EFB5FF9E4" lastPageId="13" lastPageNumber="130" pageId="12" pageNumber="129" type="materials_examined">
<paragraph id="B56D4F1CFF8EFFE1FF26F89EFE41F898" blockId="12.[189,426,1809,1835]" box="[189,426,1809,1835]" pageId="12" pageNumber="129">
<heading id="EE25F870FF8EFFE1FF26F89EFE41F898" bold="true" box="[189,426,1809,1835]" fontSize="11" level="3" pageId="12" pageNumber="129" reason="3">
<emphasis id="87A6930EFF8EFFE1FF26F89EFE41F898" bold="true" box="[189,426,1809,1835]" pageId="12" pageNumber="129">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="B56D4F1CFF8EFFE1FF71F8CFFED8F8E9" blockId="12.[189,1398,1856,1989]" box="[234,307,1856,1882]" pageId="12" pageNumber="129">
<emphasis id="87A6930EFF8EFFE1FF71F8CFFED8F8E9" bold="true" box="[234,307,1856,1882]" pageId="12" pageNumber="129">
<typeStatus id="6A69F1BEFF8EFFE1FF71F8CFFED8F8E9" box="[234,307,1856,1882]" pageId="12" pageNumber="129">Types</typeStatus>
</emphasis>
</paragraph>
<paragraph id="B56D4F1CFF8EFFE1FF26F8EBFC1EF877" blockId="12.[189,1398,1856,1989]" pageId="12" pageNumber="129">
<materialsCitation id="05BA4541FF8EFFE1FF26F8EBFC1AF877" ID-GBIF-Occurrence="3053841383" collectionCode="USNM" country="Indonesia" location="Bondowoso" municipality="Besoeki" pageId="12" pageNumber="129" specimenCount="2" specimenCount-worker="2" stateProvince="East Java" typeStatus="lectotype">
<collectingCountry id="CDC50F8CFF8EFFE1FF26F8EBFEB0F8CD" box="[189,347,1892,1918]" name="Indonesia" pageId="12" pageNumber="129">INDONESIA</collectingCountry>
• 
<typeStatus id="6A69F1BEFF8EFFE1FEE9F8ECFE35F8CE" box="[370,478,1891,1917]" pageId="12" pageNumber="129" type="lectotype">lectotype</typeStatus>
(top specimen on same pin) and 
<specimenCount id="A3D48495FF8EFFE1FCC0F8EBFB99F8CE" box="[859,1138,1891,1918]" pageId="12" pageNumber="129" type="worker" typeStatus="paralectotype">2 paralectotype workers</specimenCount>
of 
<taxonomicName id="72D2349FFF8EFFE1FB03F8EBFA85F8CE" authorityName="Wheeler" authorityYear="1929" box="[1176,1390,1891,1917]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="129" phylum="Arthropoda" rank="subSpecies" species="rothneyi" subSpecies="javana">
<emphasis id="87A6930EFF8EFFE1FB03F8EBFA85F8CE" box="[1176,1390,1891,1917]" italics="true" pageId="12" pageNumber="129">R. rothneyi javana</emphasis>
</taxonomicName>
; 
<collectingRegion id="771681FEFF8EFFE1FF26F808FEC4F812" box="[189,303,1927,1953]" country="Indonesia" name="Jawa Timur" pageId="12" pageNumber="129">East Java</collectingRegion>
, 
<collectingMunicipality id="5509D566FF8EFFE1FEA7F808FE75F812" box="[316,414,1927,1953]" pageId="12" pageNumber="129">Besoeki</collectingMunicipality>
, 
<location id="B00D19C7FF8EFFE1FE31F808FDD1F812" LSID="urn:lsid:plazi:treatment:3D7BFE0AFF89FFFEFDA2F95CFAA4FA39:B00D19C7FF8EFFE1FE31F808FDD1F812" box="[426,570,1927,1953]" country="Indonesia" municipality="Besoeki" name="Bondowoso" pageId="12" pageNumber="129" stateProvince="East Java">Bondowoso</location>
; 
<collectionCode id="D3C3D7D9FF8EFFE1FDDCF808FD77F812" box="[583,668,1927,1953]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="129" type="Museum">USNM</collectionCode>
MCZ.6.9.20783/SNM.595.31. Images generated by the same authors for a previous publication (i.e., 
<bibRefCitation id="D14332EDFF8EFFE1FD16F825FC8DF876" author="Wang W. Y. &amp; Yong G. W. &amp; Jaitrong W." box="[653,870,1962,1989]" pageId="12" pageNumber="129" pagination="301 - 340" refId="ref24028" refString="Wang W. Y., Yong G. W. &amp; Jaitrong W. 2018 b. The ant genus Rhopalomastix (Hymenoptera: Formicidae: Myrmicinae) in Southeast Asia, with descriptions of four new species from Singapore based on morphology and DNA barcoding. Zootaxa 4532 (3): 301 - 340. https: // doi. org / 10.11646 / zootaxa. 4532.3.1" type="journal article" year="2018">
Wang 
<emphasis id="87A6930EFF8EFFE1FD4CF824FCF9F877" box="[727,786,1962,1988]" italics="true" pageId="12" pageNumber="129">et al.</emphasis>
2018b
</bibRefCitation>
), examined
</materialsCitation>
.
</paragraph>
<paragraph id="B56D4F1CFF8FFFE0FF71FE87FE4DFE91" blockId="13.[189,1400,264,467]" box="[234,422,264,290]" pageId="13" pageNumber="130">
<emphasis id="87A6930EFF8FFFE0FF71FE87FE4DFE91" bold="true" box="[234,422,264,290]" pageId="13" pageNumber="130">Other material</emphasis>
</paragraph>
<paragraph id="B56D4F1CFF8FFFE0FF26FEA3FD3EFE60" blockId="13.[189,1400,264,467]" pageId="13" pageNumber="130">
<materialsCitation id="05BA4541FF8FFFE0FF26FEA3FC3BFED9" ID-GBIF-Occurrence="3053841329" collectingDate="2017-01-04" collectionCode="ZRC" collectorName="G. W. Yong" country="Singapore" latitude="1.40337" location="Mandai Track" longLatPrecision="1" longitude="103.77824" pageId="13" pageNumber="130" specimenCode="ZRC_HYM000576" specimenCount="21" specimenCount-queen="1" specimenCount-worker="18">
<collectingCountry id="CDC50F8CFF8FFFE0FF26FEA3FE89FEF5" box="[189,354,300,326]" name="Singapore" pageId="13" pageNumber="130">SINGAPORE</collectingCountry>
• 
<specimenCount id="A3D48495FF8FFFE0FEE4FEA3FDECFEF5" box="[383,519,300,326]" pageId="13" pageNumber="130" type="worker">18 workers</specimenCount>
, 
<specimenCount id="A3D48495FF8FFFE0FD8DFEA3FD9CFEF5" box="[534,631,300,326]" pageId="13" pageNumber="130" type="queen">1 queen</specimenCount>
, 
<specimenCount id="A3D48495FF8FFFE0FD1EFEA3FD2CFEFB" box="[645,711,300,328]" pageId="13" pageNumber="130">2 ♂♂</specimenCount>
; 
<location id="B00D19C7FF8FFFE0FD40FEA3FC6EFEF5" LSID="urn:lsid:plazi:treatment:3D7BFE0AFF89FFFEFDA2F95CFAA4FA39:B00D19C7FF8FFFE0FD40FEA3FC6EFEF5" box="[731,901,300,326]" country="Singapore" latitude="1.40337" longLatPrecision="1" longitude="103.77824" name="Mandai Track" pageId="13" pageNumber="130">Mandai Track</location>
; 
<geoCoordinate id="D0E629DBFF8FFFE0FC08FEA3FBF4FEF5" box="[915,1055,300,326]" degrees="1.40337" direction="north" orientation="latitude" pageId="13" pageNumber="130" precision="1" value="1.40337">1.40337° N</geoCoordinate>
, 
<geoCoordinate id="D0E629DBFF8FFFE0FBB5FEA3FB3EFEF5" box="[1070,1237,300,326]" degrees="103.77824" direction="east" orientation="longitude" pageId="13" pageNumber="130" precision="1" value="103.77824">103.77824° E</geoCoordinate>
; 
<date id="C16C69DCFF8FFFE0FB78FEA3FA99FEF5" box="[1251,1394,300,326]" pageId="13" pageNumber="130" value="2017-01-04">
<collectingDate id="D1289034FF8FFFE0FB78FEA3FA99FEF5" box="[1251,1394,300,326]" pageId="13" pageNumber="130" value="2017-01-04">4 Jan. 2017</collectingDate>
</date>
; 
<collectorName id="18272ACAFF8FFFE0FF26FEC0FED6FEDA" box="[189,317,335,361]" pageId="13" pageNumber="130">G.W. Yong</collectorName>
leg.; colony code: GY-SG17-RhoN; 
<specimenCode id="E574E767FF8FFFE0FD77FEC0FC3BFED9" box="[748,976,335,362]" pageId="13" pageNumber="130">ZRC_HYM000576</specimenCode>
</materialsCitation>
• 
<materialsCitation id="05BA4541FF8FFFE0FC73FEC0FE3EFE3E" ID-GBIF-Occurrence="3053841325" collectingDate="2017-01-04" collectionCode="THNHM" collectorName="G. W. Yong" country="Singapore" latitude="1.40337" location="Mandai Track" longLatPrecision="1" longitude="103.77824" pageId="13" pageNumber="130" specimenCount="8" specimenCount-worker="8">
<specimenCount id="A3D48495FF8FFFE0FC73FEC0FBB4FEDA" box="[1000,1119,335,361]" pageId="13" pageNumber="130" type="worker">8 workers</specimenCount>
; same collection data as for preceding; 
<collectionCode id="D3C3D7D9FF8FFFE0FEF2FEFCFE3EFE3E" box="[361,469,371,397]" pageId="13" pageNumber="130">THNHM</collectionCode>
</materialsCitation>
• 
<materialsCitation id="05BA4541FF8FFFE0FE75FEFDFD39FE60" ID-GBIF-Occurrence="3053841305" collectedFrom="nest in bark of Campnosperma auriculatum (Blume) Hook. f." collectingDate="2017-06-21" collectionCode="ZRC" collectorName="C. Peeters" country="Singapore" latitude="1.35449" location="Bukit Timah Nature Reserve" longLatPrecision="1" longitude="103.78211" pageId="13" pageNumber="130" specimenCode="ZRC_HYM_0001732" specimenCount="14" specimenCount-worker="12">
<specimenCount id="A3D48495FF8FFFE0FE75FEFDFD9FFE3F" box="[494,628,370,396]" pageId="13" pageNumber="130" type="worker">12 workers</specimenCount>
, 
<specimenCount id="A3D48495FF8FFFE0FD1AFEFDFD07FE3E" box="[641,748,370,397]" pageId="13" pageNumber="130">2 queens</specimenCount>
; 
<location id="B00D19C7FF8FFFE0FD63FEFDFBA7FE3E" LSID="urn:lsid:plazi:treatment:3D7BFE0AFF89FFFEFDA2F95CFAA4FA39:B00D19C7FF8FFFE0FD63FEFDFBA7FE3E" box="[760,1100,370,397]" country="Singapore" latitude="1.35449" longLatPrecision="1" longitude="103.78211" name="Bukit Timah Nature Reserve" pageId="13" pageNumber="130">Bukit Timah Nature Reserve</location>
; 
<geoCoordinate id="D0E629DBFF8FFFE0FBC2FEFDFB08FE3E" box="[1113,1251,370,397]" degrees="1.35449" direction="north" orientation="latitude" pageId="13" pageNumber="130" precision="1" value="1.35449">1.35449° N</geoCoordinate>
, 
<geoCoordinate id="D0E629DBFF8FFFE0FB74FEFDFF39FE03" degrees="103.78211" direction="east" orientation="longitude" pageId="13" pageNumber="130" precision="1" value="103.78211">103.78211° E</geoCoordinate>
; 
<date id="C16C69DCFF8FFFE0FF44FE1AFE90FE1C" box="[223,379,405,432]" pageId="13" pageNumber="130" value="2017-06-21">
<collectingDate id="D1289034FF8FFFE0FF44FE1AFE90FE1C" box="[223,379,405,432]" pageId="13" pageNumber="130" value="2017-06-21">21 Jun. 2017</collectingDate>
</date>
; 
<collectorName id="18272ACAFF8FFFE0FE12FE1AFDEAFE03" box="[393,513,405,432]" pageId="13" pageNumber="130">C. Peeters</collectorName>
leg.; 
<collectedFrom id="06CE0CE1FF8FFFE0FDDEFE19FAF0FE1C" box="[581,1307,405,432]" pageId="13" pageNumber="130">
nest in bark of 
<taxonomicName id="72D2349FFF8FFFE0FD61FE19FAE0FE1C" authority="(Blume) Hook." authorityName="Hook." baseAuthorityName="Blume" box="[762,1291,405,432]" class="Magnoliopsida" family="Anacardiaceae" genus="Campnosperma" kingdom="Plantae" order="Sapindales" pageId="13" pageNumber="130" phylum="Tracheophyta" rank="species" species="auriculatum">
<emphasis id="87A6930EFF8FFFE0FD61FE19FBA0FE1C" box="[762,1099,405,432]" italics="true" pageId="13" pageNumber="130">Campnosperma auriculatum</emphasis>
(Blume) Hook.
</taxonomicName>
f.
</collectedFrom>
; colony code: WW-SG17-015; 
<specimenCode id="E574E767FF8FFFE0FE57FE36FD39FE60" box="[460,722,441,467]" pageId="13" pageNumber="130">ZRC_HYM_0001732</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="B56D4F1CFF8FFFE0FF26FE75FB5FF9E4" blockId="13.[189,1399,506,1624]" pageId="13" pageNumber="130">
<materialsCitation id="05BA4541FF8FFFE0FF26FE75FEACFDE8" ID-GBIF-Occurrence="3053841384" collectedFrom="mango tree" collectingDate="2018-03-11" collectionCode="THNHM" collectorName="W. Jaitrong" country="Thailand" elevation="280" latitude="14.33389" location="West Thailand" longLatPrecision="538" longitude="98.98" municipality="Sai Yok District" pageId="13" pageNumber="130" specimenCount="8" specimenCount-worker="8" stateProvince="Kanchanaburi Province">
<collectingCountry id="CDC50F8CFF8FFFE0FF26FE75FEBDFDA7" box="[189,342,506,532]" name="Thailand" pageId="13" pageNumber="130">THAILAND</collectingCountry>
– 
<emphasis id="87A6930EFF8FFFE0FEE2FE75FDD8FDA7" bold="true" box="[377,563,506,532]" pageId="13" pageNumber="130">
<location id="B00D19C7FF8FFFE0FEE2FE75FDD8FDA7" LSID="urn:lsid:plazi:treatment:3D7BFE0AFF89FFFEFDA2F95CFAA4FA39:B00D19C7FF8FFFE0FEE2FE75FDD8FDA7" box="[377,563,506,532]" country="Thailand" latitude="14.33389" longLatPrecision="538" longitude="98.98" municipality="Sai Yok District" name="West Thailand" pageId="13" pageNumber="130" stateProvince="Kanchanaburi Province">West Thailand</location>
</emphasis>
• 
<specimenCount id="A3D48495FF8FFFE0FDD5FE75FD22FDA7" box="[590,713,506,532]" pageId="13" pageNumber="130" type="worker">8 workers</specimenCount>
; 
<collectingRegion id="771681FEFF8FFFE0FD4CFE75FC18FDA7" box="[727,1011,506,532]" country="Thailand" name="Kanchanaburi" pageId="13" pageNumber="130">Kanchanaburi Province</collectingRegion>
, 
<collectingMunicipality id="5509D566FF8FFFE0FB9AFE75FB2FFDA7" box="[1025,1220,506,532]" pageId="13" pageNumber="130">Sai Yok District</collectingMunicipality>
, 
<location id="B00D19C7FF8FFFE0FB49FE75FA99FDA7" LSID="urn:lsid:plazi:treatment:3D7BFE0AFF89FFFEFDA2F95CFAA4FA39:B00D19C7FF8FFFE0FB49FE75FA99FDA7" box="[1234,1394,506,532]" country="Thailand" latitude="14.33389" longLatPrecision="538" longitude="98.98" municipality="Sai Yok District" name="Ban Tha Sao" pageId="13" pageNumber="130" stateProvince="Kanchanaburi Province">Ban Tha Sao</location>
; 
<geoCoordinate id="D0E629DBFF8FFFE0FF26FD92FEBFFD8B" box="[189,340,541,568]" degrees="14.33389" direction="north" orientation="latitude" pageId="13" pageNumber="130" precision="1" value="14.33389">14.33389° N</geoCoordinate>
, 
<geoCoordinate id="D0E629DBFF8FFFE0FEC7FD92FE28FD8B" box="[348,451,541,568]" degrees="98.98" direction="east" orientation="longitude" pageId="13" pageNumber="130" precision="555" value="98.98">98.98° E</geoCoordinate>
; 
<elevation id="3EFFA82FFF8FFFE0FE50FD92FDA4FD84" box="[459,591,541,568]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="13" pageNumber="130" unit="m" value="280.0">
<quantity id="722AE2F9FF8FFFE0FE50FD92FDF9FD8B" box="[459,530,541,568]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="13" pageNumber="130" unit="m" value="280.0">280 m</quantity>
a.s.l.
</elevation>
; 
<date id="C16C69DCFF8FFFE0FDC2FD92FD19FD84" box="[601,754,541,568]" pageId="13" pageNumber="130" value="2018-03-11">
<collectingDate id="D1289034FF8FFFE0FDC2FD92FD19FD84" box="[601,754,541,568]" pageId="13" pageNumber="130" value="2018-03-11">11 Mar.2018</collectingDate>
</date>
; 
<collectorName id="18272ACAFF8FFFE0FD61FD91FC95FD84" box="[762,894,541,568]" pageId="13" pageNumber="130">W.Jaitrong</collectorName>
leg.; 
<collectedFrom id="06CE0CE1FF8FFFE0FC23FD91FBD7FD8B" box="[952,1084,542,568]" pageId="13" pageNumber="130">mango tree</collectedFrom>
;colony code:WJT110318- 1; 
<collectionCode id="D3C3D7D9FF8FFFE0FF40FDCEFEACFDE8" box="[219,327,577,603]" pageId="13" pageNumber="130">THNHM</collectionCode>
</materialsCitation>
• 
<materialsCitation id="05BA4541FF8FFFE0FEF9FDCFFDB4FDCD" ID-GBIF-Occurrence="3053841363" accessionNumber="MW267108, MW267110" collectedFrom="mango tree" collectingDate="2018-03-11" collectionCode="MW, ZRC" collectorName="W. Jaitrong" country="Thailand" elevation="280" latitude="14.33389" location="GenBank" longLatPrecision="538" longitude="98.98" municipality="Sai Yok District" pageId="13" pageNumber="130" specimenCode="ZRC_ENT00000931" specimenCount="8" specimenCount-worker="8" stateProvince="West Thailand">
<specimenCount id="A3D48495FF8FFFE0FEF9FDCFFE30FDE9" box="[354,475,576,602]" pageId="13" pageNumber="130" type="worker">8 workers</specimenCount>
; same collection data as for preceding; GenBank: 
<accessionNumber id="AA81D2FFFF8FFFE0FBA8FDCFFB29FDE9" box="[1075,1218,576,602]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MW267108" pageId="13" pageNumber="130">MW267108</accessionNumber>
to 
<accessionNumber id="AA81D2FFFF8FFFE0FB72FDCFFA9CFDE9" box="[1257,1399,576,602]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MW267110" pageId="13" pageNumber="130">MW267110</accessionNumber>
(3 barcoded); 
<specimenCode id="E574E767FF8FFFE0FEFCFDEBFDB4FDCD" box="[359,607,612,638]" pageId="13" pageNumber="130">ZRC_ENT00000931</specimenCode>
</materialsCitation>
• 
<materialsCitation id="05BA4541FF8FFFE0FDE6FDEBFD25FD12" ID-GBIF-Occurrence="3053841330" accessionNumber="MW267108, MW267110" collectedFrom="mango tree" collectingDate="2018-03-11" collectionCode="THNHM" collectorName="W. Jaitrong" country="Thailand" elevation="280" latitude="14.33389" location="GenBank" longLatPrecision="538" longitude="98.98" municipality="Sai Yok District" pageId="13" pageNumber="130" specimenCount="9" specimenCount-male="1" specimenCount-worker="8" stateProvince="West Thailand">
<specimenCount id="A3D48495FF8FFFE0FDE6FDEBFD1CFDCE" box="[637,759,611,638]" pageId="13" pageNumber="130" type="worker">8 workers</specimenCount>
, 
<specimenCount id="A3D48495FF8FFFE0FC9EFDEBFCD9FD33" box="[773,818,612,640]" pageId="13" pageNumber="130" type="male">1 ♂</specimenCount>
; same collection data as for preceding; 
<date id="C16C69DCFF8FFFE0FA81FDEBFF17FD12" pageId="13" pageNumber="130" value="2018-03-11">
<collectingDate id="D1289034FF8FFFE0FA81FDEBFF17FD12" pageId="13" pageNumber="130" value="2018-03-11">11 Mar. 2018</collectingDate>
</date>
; colony code: WJT110318-2; 
<collectionCode id="D3C3D7D9FF8FFFE0FDF9FD08FD25FD12" box="[610,718,647,673]" pageId="13" pageNumber="130">THNHM</collectionCode>
</materialsCitation>
• 
<materialsCitation id="05BA4541FF8FFFE0FD6AFD08FBA5FD77" ID-GBIF-Occurrence="3053841321" accessionNumber="MW267111, MW267113" collectedFrom="mango tree" collectingDate="2018-03-11" collectionCode="MW, ZRC" collectorName="W. Jaitrong" country="Thailand" elevation="280" latitude="14.33389" location="GenBank" longLatPrecision="538" longitude="98.98" municipality="Sai Yok District" pageId="13" pageNumber="130" specimenCode="ZRC_ENT00000932" specimenCount="9" specimenCount-male="1" specimenCount-worker="8" stateProvince="West Thailand">
<specimenCount id="A3D48495FF8FFFE0FD6AFD08FC82FD12" box="[753,873,647,673]" pageId="13" pageNumber="130" type="worker">8 workers</specimenCount>
, 
<specimenCount id="A3D48495FF8FFFE0FCEDFD08FC4AFD10" box="[886,929,647,675]" pageId="13" pageNumber="130" type="male">1 ♂</specimenCount>
; same collection data as for preceding; GenBank: 
<accessionNumber id="AA81D2FFFF8FFFE0FED8FD25FE3BFD77" box="[323,464,682,708]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MW267111" pageId="13" pageNumber="130">MW267111</accessionNumber>
to 
<accessionNumber id="AA81D2FFFF8FFFE0FD98FD25FD7BFD77" box="[515,656,682,708]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MW267113" pageId="13" pageNumber="130">MW267113</accessionNumber>
(3 barcoded); 
<specimenCode id="E574E767FF8FFFE0FCC9FD25FBA5FD77" box="[850,1102,682,708]" pageId="13" pageNumber="130">ZRC_ENT00000932</specimenCode>
</materialsCitation>
. – 
<materialsCitation id="05BA4541FF8FFFE0FBE7FD25FC34FCB8" ID-GBIF-Occurrence="3053841336" collectingDate="2018-03-18" collectionCode="THNHM" collectorName="W. Jaitrong" country="Thailand" elevation="89" latitude="14.67056" location="Phu Khae Botanical Garden" longLatPrecision="1" longitude="100.885" pageId="13" pageNumber="130" specimenCount="8" specimenCount-worker="8" stateProvince="Saraburi Province">
<emphasis id="87A6930EFF8FFFE0FBE7FD25FAB5FD77" bold="true" box="[1148,1374,682,708]" pageId="13" pageNumber="130">
Central 
<collectingCountry id="CDC50F8CFF8FFFE0FB70FD25FAB5FD77" box="[1259,1374,682,708]" name="Thailand" pageId="13" pageNumber="130">Thailand</collectingCountry>
</emphasis>
• 
<specimenCount id="A3D48495FF8FFFE0FF26FD42FEDEFD54" box="[189,309,717,743]" pageId="13" pageNumber="130" type="worker">8 workers</specimenCount>
; 
<collectingRegion id="771681FEFF8FFFE0FEDAFD42FDF3FD54" box="[321,536,717,743]" country="Thailand" name="Sara Buri" pageId="13" pageNumber="130">Saraburi Province</collectingRegion>
, 
<location id="B00D19C7FF8FFFE0FDB8FD42FC9BFD54" LSID="urn:lsid:plazi:treatment:3D7BFE0AFF89FFFEFDA2F95CFAA4FA39:B00D19C7FF8FFFE0FDB8FD42FC9BFD54" box="[547,880,717,743]" country="Thailand" latitude="14.67056" longLatPrecision="1" longitude="100.885" name="Phu Khae Botanical Garden" pageId="13" pageNumber="130" stateProvince="Saraburi Province">Phu Khae Botanical Garden</location>
; 
<geoCoordinate id="D0E629DBFF8FFFE0FCE0FD42FBFEFD5B" box="[891,1045,717,744]" degrees="14.67056" direction="north" orientation="latitude" pageId="13" pageNumber="130" precision="1" value="14.67056">14.67056° N</geoCoordinate>
, 
<geoCoordinate id="D0E629DBFF8FFFE0FBBBFD42FB2EFD5B" box="[1056,1221,717,744]" degrees="100.88500" direction="east" orientation="longitude" pageId="13" pageNumber="130" precision="1" value="100.885">100.88500° E</geoCoordinate>
; 
<elevation id="3EFFA82FFF8FFFE0FB4AFD42FAA7FD54" box="[1233,1356,717,744]" metricMagnitude="1" metricUnit="m" metricValue="8.9" pageId="13" pageNumber="130" unit="m" value="89.0">
<quantity id="722AE2F9FF8FFFE0FB4AFD42FAE7FD5B" box="[1233,1292,717,744]" metricMagnitude="1" metricUnit="m" metricValue="8.9" pageId="13" pageNumber="130" unit="m" value="89.0">89 m</quantity>
a.s.l.
</elevation>
; 
<date id="C16C69DCFF8FFFE0FAC2FD42FED7FCB9" pageId="13" pageNumber="130" value="2018-03-18">
<collectingDate id="D1289034FF8FFFE0FAC2FD42FED7FCB9" pageId="13" pageNumber="130" value="2018-03-18">18 Mar. 2018</collectingDate>
</date>
; 
<collectorName id="18272ACAFF8FFFE0FED3FD7EFE3AFCB9" box="[328,465,752,779]" pageId="13" pageNumber="130">W. Jaitrong</collectorName>
leg.; colony code: WJT180318-2; 
<collectionCode id="D3C3D7D9FF8FFFE0FCE8FD7EFC34FCB8" box="[883,991,753,779]" pageId="13" pageNumber="130">THNHM</collectionCode>
</materialsCitation>
• 
<materialsCitation id="05BA4541FF8FFFE0FC60FD7FFB09FC9D" ID-GBIF-Occurrence="3053841326" accessionNumber="MW267105, MW267107" collectingDate="2018-03-18" collectionCode="MW, ZRC" collectorName="W. Jaitrong" country="Thailand" elevation="89" latitude="14.67056" location="GenBank" longLatPrecision="1" longitude="100.885" pageId="13" pageNumber="130" specimenCode="ZRC_ENT00000937" specimenCount="8" specimenCount-worker="8" stateProvince="Central Thailand">
<specimenCount id="A3D48495FF8FFFE0FC60FD7FFB9EFCB9" box="[1019,1141,752,778]" pageId="13" pageNumber="130" type="worker">8 workers</specimenCount>
; same collection data as for preceding; GenBank: 
<accessionNumber id="AA81D2FFFF8FFFE0FD9DFC9CFD79FC9D" box="[518,658,787,814]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MW267105" pageId="13" pageNumber="130">MW267105</accessionNumber>
to 
<accessionNumber id="AA81D2FFFF8FFFE0FD2CFC9CFCAFFC9E" box="[695,836,787,813]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MW267107" pageId="13" pageNumber="130">MW267107</accessionNumber>
(3 barcoded); 
<specimenCode id="E574E767FF8FFFE0FC75FC9BFB09FC9D" box="[1006,1250,788,814]" pageId="13" pageNumber="130">ZRC_ENT00000937</specimenCode>
</materialsCitation>
• 
<materialsCitation id="05BA4541FF8FFFE0FB60FC9BFEACFCC7" ID-GBIF-Occurrence="3053841344" collectedFrom="in bark of mango tree" collectingDate="2018-07-07" collectionCode="THNHM" collectorName="W. Jaitrong" country="Thailand" location="Khao Yai National Park" pageId="13" pageNumber="130" specimenCount="9" specimenCount-worker="9" stateProvince="Central Thailand">
<specimenCount id="A3D48495FF8FFFE0FB60FC9BFA99FC9E" box="[1275,1394,787,814]" pageId="13" pageNumber="130" type="worker">9 workers</specimenCount>
; 
<location id="B00D19C7FF8FFFE0FF26FCB8FE3EFCE2" LSID="urn:lsid:plazi:treatment:3D7BFE0AFF89FFFEFDA2F95CFAA4FA39:B00D19C7FF8FFFE0FF26FCB8FE3EFCE2" box="[189,469,823,849]" country="Thailand" name="Khao Yai National Park" pageId="13" pageNumber="130" stateProvince="Central Thailand">Khao Yai National Park</location>
; 
<date id="C16C69DCFF8FFFE0FE7AFCB8FD8EFCE2" box="[481,613,823,849]" pageId="13" pageNumber="130" value="2018-07-07">
<collectingDate id="D1289034FF8FFFE0FE7AFCB8FD8EFCE2" box="[481,613,823,849]" pageId="13" pageNumber="130" value="2018-07-07">7 Jul. 2018</collectingDate>
</date>
; 
<collectorName id="18272ACAFF8FFFE0FDEBFCB8FD1EFCE2" box="[624,757,823,849]" pageId="13" pageNumber="130">W. Jaitrong</collectorName>
leg.; 
<collectedFrom id="06CE0CE1FF8FFFE0FCACFCB8FBDEFCE2" box="[823,1077,823,849]" pageId="13" pageNumber="130">in bark of mango tree</collectedFrom>
; colony code: WJT070718- 1; 
<collectionCode id="D3C3D7D9FF8FFFE0FF40FCD5FEACFCC7" box="[219,327,858,884]" pageId="13" pageNumber="130">THNHM</collectionCode>
</materialsCitation>
• 
<materialsCitation id="05BA4541FF8FFFE0FEFAFCD5FD8FFC24" ID-GBIF-Occurrence="3053841348" accessionNumber="MW267096, MW267098" collectedFrom="in bark of mango tree" collectingDate="2018-07-07" collectionCode="MW, ZRC" collectorName="W. Jaitrong" country="Thailand" location="GenBank" pageId="13" pageNumber="130" specimenCode="ZRC_ ENT00007610" specimenCount="5" specimenCount-worker="5" stateProvince="Central Thailand">
<specimenCount id="A3D48495FF8FFFE0FEFAFCD5FE30FCC7" box="[353,475,858,885]" pageId="13" pageNumber="130" type="worker">5 workers</specimenCount>
; same collection data as for preceding; GenBank: 
<accessionNumber id="AA81D2FFFF8FFFE0FBA9FCD5FB2BFCC7" box="[1074,1216,858,884]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MW267096" pageId="13" pageNumber="130">MW267096</accessionNumber>
to 
<accessionNumber id="AA81D2FFFF8FFFE0FB73FCD5FA9DFCC7" box="[1256,1398,858,884]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MW267098" pageId="13" pageNumber="130">MW267098</accessionNumber>
(3 barcoded); 
<specimenCode id="E574E767FF8FFFE0FEFAFCF2FD8FFC24" box="[353,612,893,919]" pageId="13" pageNumber="130">ZRC_ ENT00007610</specimenCode>
</materialsCitation>
. – 
<materialsCitation id="05BA4541FF8FFFE0FD19FCF2FB39FC6D" ID-GBIF-Occurrence="3053841324" collectedFrom="in bark of Azadirachta excelsa (Jack) Jacobs" collectingDate="2018-01-23" collectionCode="THNHM" collectorName="W. Jaitrong" country="Thailand" elevation="28" latitude="7.50833" location="Nakhaw Sia Subdistrict" longLatPrecision="1" longitude="99.71639" municipality="Nayong District" pageId="13" pageNumber="130" specimenCount="10" specimenCount-worker="8" stateProvince="Trang Province">
<emphasis id="87A6930EFF8FFFE0FD19FCF2FCAEFC24" bold="true" box="[642,837,893,919]" pageId="13" pageNumber="130">
South 
<collectingCountry id="CDC50F8CFF8FFFE0FD49FCF2FCAEFC24" box="[722,837,893,919]" name="Thailand" pageId="13" pageNumber="130">Thailand</collectingCountry>
</emphasis>
• 
<specimenCount id="A3D48495FF8FFFE0FCC6FCF2FC3FFC24" box="[861,980,893,919]" pageId="13" pageNumber="130" type="worker">8 workers</specimenCount>
, 
<specimenCount id="A3D48495FF8FFFE0FC7BFCF2FBA0FC2B" box="[992,1099,893,920]" pageId="13" pageNumber="130">2 queens</specimenCount>
; 
<collectingRegion id="771681FEFF8FFFE0FBCEFCF1FAE5FC24" box="[1109,1294,893,920]" country="Thailand" name="Trang" pageId="13" pageNumber="130">Trang Province</collectingRegion>
, 
<collectingMunicipality id="5509D566FF8FFFE0FA82FCF1FEF2FC09" pageId="13" pageNumber="130">Nayong District</collectingMunicipality>
, 
<location id="B00D19C7FF8FFFE0FEB3FC2FFDA1FC09" LSID="urn:lsid:plazi:treatment:3D7BFE0AFF89FFFEFDA2F95CFAA4FA39:B00D19C7FF8FFFE0FEB3FC2FFDA1FC09" box="[296,586,928,954]" country="Thailand" latitude="7.50833" longLatPrecision="1" longitude="99.71639" municipality="Nayong District" name="Nakhaw Sia Subdistrict" pageId="13" pageNumber="130" stateProvince="Trang Province">Nakhaw Sia Subdistrict</location>
; 
<geoCoordinate id="D0E629DBFF8FFFE0FDC3FC2FFD0CFC08" box="[600,743,928,955]" degrees="7.50833" direction="north" orientation="latitude" pageId="13" pageNumber="130" precision="1" value="7.50833">7.50833° N</geoCoordinate>
, 
<geoCoordinate id="D0E629DBFF8FFFE0FD6DFC2FFC74FC08" box="[758,927,928,955]" degrees="99.716389" direction="east" orientation="longitude" pageId="13" pageNumber="130" precision="1" value="99.71639">99.716389° E</geoCoordinate>
; 
<quantity id="722AE2F9FF8FFFE0FC35FC2FFC1BFC08" box="[942,1008,928,955]" metricMagnitude="1" metricUnit="m" metricValue="2.8" pageId="13" pageNumber="130" unit="m" value="28.0">
<elevation id="3EFFA82FFF8FFFE0FC35FC2FFC1BFC08" box="[942,1008,928,955]" metricMagnitude="1" metricUnit="m" metricValue="2.8" pageId="13" pageNumber="130" unit="m" value="28.0">28 m</elevation>
</quantity>
; 
<date id="C16C69DCFF8FFFE0FB9BFC2FFB4BFC09" box="[1024,1184,928,955]" pageId="13" pageNumber="130" value="2018-01-23">
<collectingDate id="D1289034FF8FFFE0FB9BFC2FFB4BFC09" box="[1024,1184,928,955]" pageId="13" pageNumber="130" value="2018-01-23">23 Jan. 2018</collectingDate>
</date>
; 
<collectorName id="18272ACAFF8FFFE0FB35FC2EFAD2FC09" box="[1198,1337,928,955]" pageId="13" pageNumber="130">W. Jaitrong</collectorName>
leg.; 
<collectedFrom id="06CE0CE1FF8FFFE0FF26FC4CFD05FC6E" box="[189,750,963,990]" pageId="13" pageNumber="130">
in bark of 
<taxonomicName id="72D2349FFF8FFFE0FEDDFC4CFD05FC6E" authority="(Jack) Jacobs" authorityName="Jacobs" baseAuthorityName="Jack" box="[326,750,963,990]" class="Magnoliopsida" family="Meliaceae" genus="Azadirachta" kingdom="Plantae" order="Sapindales" pageId="13" pageNumber="130" phylum="Tracheophyta" rank="species" species="excelsa">
<emphasis id="87A6930EFF8FFFE0FEDDFC4CFDD1FC6E" box="[326,570,963,989]" italics="true" pageId="13" pageNumber="130">Azadirachta excelsa</emphasis>
(Jack) Jacobs
</taxonomicName>
</collectedFrom>
; colony code: WJT230118-1; 
<collectionCode id="D3C3D7D9FF8FFFE0FBFDFC4BFB39FC6D" box="[1126,1234,964,990]" pageId="13" pageNumber="130">THNHM</collectionCode>
</materialsCitation>
• 
<materialsCitation id="05BA4541FF8FFFE0FB6EFC4BFE51FB97" ID-GBIF-Occurrence="3053841332" accessionNumber="MW267114, MW267116" collectedFrom="in bark of Azadirachta excelsa (Jack) Jacobs" collectingDate="2018-01-23" collectionCode="MW, ZRC" collectorName="W. Jaitrong" country="Thailand" elevation="28" latitude="7.50833" location="GenBank" longLatPrecision="1" longitude="99.71639" municipality="Nayong District" pageId="13" pageNumber="130" specimenCode="ZRC_ENT00000942" specimenCount="10" specimenCount-worker="8" stateProvince="South Thailand">
<specimenCount id="A3D48495FF8FFFE0FB6EFC4BFA99FC6E" box="[1269,1394,963,990]" pageId="13" pageNumber="130" type="worker">8 workers</specimenCount>
, 
<specimenCount id="A3D48495FF8FFFE0FF26FC68FEC0FBB2" box="[189,299,999,1025]" pageId="13" pageNumber="130">2 queens</specimenCount>
; same collection data as for preceding; GenBank: 
<accessionNumber id="AA81D2FFFF8FFFE0FC11FC68FBFCFBB2" box="[906,1047,999,1025]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MW267114" pageId="13" pageNumber="130">MW267114</accessionNumber>
to 
<accessionNumber id="AA81D2FFFF8FFFE0FBDBFC68FB25FBB2" box="[1088,1230,999,1025]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MW267116" pageId="13" pageNumber="130">MW267116</accessionNumber>
(3 barcoded); 
<specimenCode id="E574E767FF8FFFE0FF25FB85FE51FB97" box="[190,442,1034,1060]" pageId="13" pageNumber="130">ZRC_ENT00000942</specimenCode>
</materialsCitation>
. – 
<materialsCitation id="05BA4541FF8FFFE0FE40FB85FD79FBFB" ID-GBIF-Occurrence="3053841319" collectingDate="2018-10-21" collectionCode="THNHM" collectorName="W. Jaitrong" country="Thailand" location="Chiang Rai" pageId="13" pageNumber="130" specimenCount="15" specimenCount-worker="15" stateProvince="Chiang Rai">
<emphasis id="87A6930EFF8FFFE0FE40FB85FD27FB97" bold="true" box="[475,716,1034,1060]" pageId="13" pageNumber="130">
Northern 
<collectingCountry id="CDC50F8CFF8FFFE0FDC2FB85FD27FB97" box="[601,716,1034,1060]" name="Thailand" pageId="13" pageNumber="130">Thailand</collectingCountry>
</emphasis>
• 
<specimenCount id="A3D48495FF8FFFE0FD7EFB85FC85FB97" box="[741,878,1034,1061]" pageId="13" pageNumber="130" type="worker">15 workers</specimenCount>
; 
<collectingRegion id="771681FEFF8FFFE0FCE1FB85FBEFFB97" box="[890,1028,1034,1060]" country="Thailand" name="Chiang Rai" pageId="13" pageNumber="130">Chiang Rai</collectingRegion>
; 
<date id="C16C69DCFF8FFFE0FB8AFB85FB44FB97" box="[1041,1199,1034,1060]" pageId="13" pageNumber="130" value="2018-10-21">
<collectingDate id="D1289034FF8FFFE0FB8AFB85FB44FB97" box="[1041,1199,1034,1060]" pageId="13" pageNumber="130" value="2018-10-21">21 Oct. 2018</collectingDate>
</date>
; 
<collectorName id="18272ACAFF8FFFE0FB21FB85FAA8FB97" box="[1210,1347,1034,1060]" pageId="13" pageNumber="130">W. Jaitrong</collectorName>
leg; colony code: WJT211018-34; 
<collectionCode id="D3C3D7D9FF8FFFE0FDBDFBA1FD79FBFB" box="[550,658,1070,1096]" pageId="13" pageNumber="130">THNHM</collectionCode>
</materialsCitation>
• 
<materialsCitation id="05BA4541FF8FFFE0FD37FBA2FC5DFBD9" ID-GBIF-Occurrence="3053841328" accessionNumber="MW267099, MW267101" collectingDate="2018-10-21" collectionCode="MW, ZRC" collectorName="W. Jaitrong" country="Thailand" location="GenBank" pageId="13" pageNumber="130" specimenCode="ZRC_ ENT00007609" specimenCount="8" specimenCount-worker="8" stateProvince="Northern Thailand">
<specimenCount id="A3D48495FF8FFFE0FD37FBA2FCCDFBF4" box="[684,806,1069,1095]" pageId="13" pageNumber="130" type="worker">8 workers</specimenCount>
; same collection data as for preceding; GenBank: 
<accessionNumber id="AA81D2FFFF8FFFE0FF26FBDFFEA7FBD9" box="[189,332,1104,1130]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MW267099" pageId="13" pageNumber="130">MW267099</accessionNumber>
to 
<accessionNumber id="AA81D2FFFF8FFFE0FEE8FBDFFDE9FBD9" box="[371,514,1104,1130]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MW267101" pageId="13" pageNumber="130">MW267101</accessionNumber>
(3 barcoded); 
<specimenCode id="E574E767FF8FFFE0FD2AFBDFFC5DFBD9" box="[689,950,1104,1130]" pageId="13" pageNumber="130">ZRC_ ENT00007609</specimenCode>
</materialsCitation>
. – 
<materialsCitation id="05BA4541FF8FFFE0FC43FBDFFD7CFB02" ID-GBIF-Occurrence="3053841373" collectingDate="2018-09-13" collectionCode="THNHM" collectorName="W. Jaitrong" country="Thailand" county="Srisa Ket Province" location="Krabau Krabai" municipality="Phanom Dong Rak" pageId="13" pageNumber="130" specimenCount="19" specimenCount-queen="1" specimenCount-worker="18" stateProvince="Northeast Thailand">
<emphasis id="87A6930EFF8FFFE0FC43FBDFFB3BFBD9" bold="true" box="[984,1232,1104,1130]" pageId="13" pageNumber="130">
<collectingRegion id="771681FEFF8FFFE0FC43FBDFFB3BFBD9" box="[984,1232,1104,1130]" pageId="13" pageNumber="130">Northeast Thailand</collectingRegion>
</emphasis>
• 
<specimenCount id="A3D48495FF8FFFE0FB71FBDFFA99FBD9" box="[1258,1394,1104,1130]" pageId="13" pageNumber="130" type="worker">18 workers</specimenCount>
, 
<specimenCount id="A3D48495FF8FFFE0FF26FBFBFEC9FB3D" box="[189,290,1140,1166]" pageId="13" pageNumber="130" type="queen">1 queen</specimenCount>
; 
<collectingCounty id="5C0C3790FF8FFFE0FEA8FBFCFDF6FB3E" box="[307,541,1139,1166]" pageId="13" pageNumber="130">Srisa Ket Province</collectingCounty>
, 
<collectingMunicipality id="5509D566FF8FFFE0FDB6FBFCFCF0FB3E" box="[557,795,1139,1166]" pageId="13" pageNumber="130">Phanom Dong Rak</collectingMunicipality>
, 
<location id="B00D19C7FF8FFFE0FCB0FBFCFC09FB3E" LSID="urn:lsid:plazi:treatment:3D7BFE0AFF89FFFEFDA2F95CFAA4FA39:B00D19C7FF8FFFE0FCB0FBFCFC09FB3E" box="[811,994,1139,1165]" country="Thailand" county="Srisa Ket Province" municipality="Phanom Dong Rak" name="Krabau Krabai" pageId="13" pageNumber="130" stateProvince="Northeast Thailand">Krabau Krabai</location>
; 
<date id="C16C69DCFF8FFFE0FC68FBFBFB70FB3D" box="[1011,1179,1140,1166]" pageId="13" pageNumber="130" value="2018-09-13">
<collectingDate id="D1289034FF8FFFE0FC68FBFBFB70FB3D" box="[1011,1179,1140,1166]" pageId="13" pageNumber="130" value="2018-09-13">13 Sep. 2018</collectingDate>
</date>
; 
<collectorName id="18272ACAFF8FFFE0FB30FBFBFADCFB3E" box="[1195,1335,1139,1166]" pageId="13" pageNumber="130">W. Jaitrong</collectorName>
leg.; colony code: WJT130918-17; 
<collectionCode id="D3C3D7D9FF8FFFE0FDB0FB18FD7CFB02" box="[555,663,1175,1201]" pageId="13" pageNumber="130">THNHM</collectionCode>
</materialsCitation>
• 
<materialsCitation id="05BA4541FF8FFFE0FD2FFB18FBB7FB67" ID-GBIF-Occurrence="3053841313" accessionNumber="MW267117, MW267119" collectingDate="2018-09-13" collectionCode="MW, ZRC" collectorName="W. Jaitrong" country="Thailand" county="Srisa Ket Province" location="GenBank" municipality="Phanom Dong Rak" pageId="13" pageNumber="130" specimenCode="ZRC_ENT00007611" specimenCount="9" specimenCount-queen="1" specimenCount-worker="8" stateProvince="Northeast Thailand">
<specimenCount id="A3D48495FF8FFFE0FD2FFB18FCC5FB02" box="[692,814,1175,1201]" pageId="13" pageNumber="130" type="worker">8 workers</specimenCount>
, 
<specimenCount id="A3D48495FF8FFFE0FCA7FB18FC75FB02" box="[828,926,1175,1201]" pageId="13" pageNumber="130" type="queen">1 queen</specimenCount>
; same collection data as for preceding; GenBank: 
<accessionNumber id="AA81D2FFFF8FFFE0FEDEFB35FE38FB67" box="[325,467,1210,1236]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MW267117" pageId="13" pageNumber="130">MW267117</accessionNumber>
to 
<accessionNumber id="AA81D2FFFF8FFFE0FD91FB35FD73FB67" box="[522,664,1210,1236]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MW267119" pageId="13" pageNumber="130">MW267119</accessionNumber>
(3 barcoded); 
<specimenCode id="E574E767FF8FFFE0FCFBFB35FBB7FB67" box="[864,1116,1210,1236]" pageId="13" pageNumber="130">ZRC_ENT00007611</specimenCode>
</materialsCitation>
. – 
<materialsCitation id="05BA4541FF8FFFE0FB15FB35FEACFAA8" ID-GBIF-Occurrence="3053841333" collectingDate="2018-12-02" collectionCode="THNHM" collectorName="W. Jaitrong" country="Thailand" location="Nan Fa Sai Resort" pageId="13" pageNumber="130" specimenCount="8" specimenCount-worker="8" stateProvince="Nan Province">
<emphasis id="87A6930EFF8FFFE0FB15FB35FAB7FB67" bold="true" box="[1166,1372,1210,1236]" pageId="13" pageNumber="130">
North 
<collectingCountry id="CDC50F8CFF8FFFE0FB72FB35FAB7FB67" box="[1257,1372,1210,1236]" name="Thailand" pageId="13" pageNumber="130">Thailand</collectingCountry>
</emphasis>
• 
<specimenCount id="A3D48495FF8FFFE0FF26FB52FEDDFB44" box="[189,310,1245,1271]" pageId="13" pageNumber="130" type="worker">8 workers</specimenCount>
; 
<collectingRegion id="771681FEFF8FFFE0FEDAFB51FE0EFB44" box="[321,485,1245,1272]" country="Thailand" name="Nan" pageId="13" pageNumber="130">Nan Province</collectingRegion>
, 
<location id="B00D19C7FF8FFFE0FE6AFB51FD20FB4B" LSID="urn:lsid:plazi:treatment:3D7BFE0AFF89FFFEFDA2F95CFAA4FA39:B00D19C7FF8FFFE0FE6AFB51FD20FB4B" box="[497,715,1245,1272]" country="Thailand" name="Nan Fa Sai Resort" pageId="13" pageNumber="130" stateProvince="Nan Province">Nan Fa Sai Resort</location>
; 
<date id="C16C69DCFF8FFFE0FD4DFB52FC82FB44" box="[726,873,1245,1272]" pageId="13" pageNumber="130" value="2018-12-02">
<collectingDate id="D1289034FF8FFFE0FD4DFB52FC82FB44" box="[726,873,1245,1272]" pageId="13" pageNumber="130" value="2018-12-02">2 Dec. 2018</collectingDate>
</date>
; 
<collectorName id="18272ACAFF8FFFE0FCEFFB51FC10FB44" box="[884,1019,1245,1272]" pageId="13" pageNumber="130">W. Jaitrong</collectorName>
leg.; colony code: WJT021218- 7; 
<collectionCode id="D3C3D7D9FF8FFFE0FF40FA8EFEACFAA8" box="[219,327,1281,1307]" pageId="13" pageNumber="130">THNHM</collectionCode>
</materialsCitation>
• 
<materialsCitation id="05BA4541FF8FFFE0FEF9FA8FFD8AFA8E" ID-GBIF-Occurrence="3053841369" accessionNumber="MW267126, MW267128" collectingDate="2018-12-02" collectionCode="MW, ZRC" collectorName="W. Jaitrong" country="Thailand" location="GenBank" pageId="13" pageNumber="130" specimenCode="ZRC_ENT00007869" specimenCount="8" specimenCount-worker="8" stateProvince="North Thailand">
<specimenCount id="A3D48495FF8FFFE0FEF9FA8FFE30FAA9" box="[354,475,1280,1306]" pageId="13" pageNumber="130" type="worker">8 workers</specimenCount>
; same collection data as for preceding; GenBank: 
<accessionNumber id="AA81D2FFFF8FFFE0FBA9FA8FFB2BFAA9" box="[1074,1216,1280,1306]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MW267126" pageId="13" pageNumber="130">MW267126</accessionNumber>
to 
<accessionNumber id="AA81D2FFFF8FFFE0FB73FA8FFA9DFAA9" box="[1256,1398,1280,1306]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MW267128" pageId="13" pageNumber="130">MW267128</accessionNumber>
(3 barcoded); 
<specimenCode id="E574E767FF8FFFE0FEF2FAACFD8AFA8E" box="[361,609,1315,1341]" pageId="13" pageNumber="130">ZRC_ENT00007869</specimenCode>
</materialsCitation>
• 
<materialsCitation id="05BA4541FF8FFFE0FDE4FAABFC74FAD2" ID-GBIF-Occurrence="3053841356" collectingDate="2018-12-04" collectionCode="THNHM" collectorName="W. Jaitrong" country="Thailand" location="Puar" municipality="Puar" pageId="13" pageNumber="130" specimenCount="10" specimenCount-worker="8" stateProvince="Nan Province">
<specimenCount id="A3D48495FF8FFFE0FDE4FAABFD11FA8E" box="[639,762,1315,1342]" pageId="13" pageNumber="130" type="worker">8 workers</specimenCount>
, 
<specimenCount id="A3D48495FF8FFFE0FC92FAABFC93FA8D" box="[777,888,1316,1342]" pageId="13" pageNumber="130">2 queens</specimenCount>
; 
<collectingRegion id="771681FEFF8FFFE0FC1DFAABFBC6FA8E" box="[902,1069,1315,1342]" country="Thailand" name="Nan" pageId="13" pageNumber="130">Nan Province</collectingRegion>
, 
<collectingMunicipality id="5509D566FF8FFFE0FBA7FAABFB99FA8D" box="[1084,1138,1316,1342]" pageId="13" pageNumber="130">Puar</collectingMunicipality>
[sic] District; 
<date id="C16C69DCFF8FFFE0FABCFAABFF17FAD2" pageId="13" pageNumber="130" value="2018-12-04">
<collectingDate id="D1289034FF8FFFE0FABCFAABFF17FAD2" pageId="13" pageNumber="130" value="2018-12-04">4 Dec. 2018</collectingDate>
</date>
; 
<collectorName id="18272ACAFF8FFFE0FE92FAC8FE79FAD2" box="[265,402,1351,1377]" pageId="13" pageNumber="130">W. Jaitrong</collectorName>
leg.; colony code: WJT041218-1; 
<collectionCode id="D3C3D7D9FF8FFFE0FCA8FAC8FC74FAD2" box="[819,927,1351,1377]" pageId="13" pageNumber="130">THNHM</collectionCode>
</materialsCitation>
• 
<materialsCitation id="05BA4541FF8FFFE0FC20FAC8FA9DFA37" ID-GBIF-Occurrence="3053841350" accessionNumber="MW267120, MW267122" collectingDate="2018-12-04" collectionCode="ZRC" collectorName="W. Jaitrong" country="Thailand" location="GenBank" municipality="Puar" pageId="13" pageNumber="130" specimenCode="ZRC_ENT00007870" specimenCount="10" specimenCount-worker="8" stateProvince="North Thailand">
<specimenCount id="A3D48495FF8FFFE0FC20FAC8FBDFFAD2" box="[955,1076,1351,1377]" pageId="13" pageNumber="130" type="worker">8 workers</specimenCount>
, 
<specimenCount id="A3D48495FF8FFFE0FBD9FAC8FB44FAD2" box="[1090,1199,1351,1377]" pageId="13" pageNumber="130">2 queens</specimenCount>
; same collection data as for preceding; GenBank: 
<accessionNumber id="AA81D2FFFF8FFFE0FDDFFAE5FD38FA37" box="[580,723,1386,1412]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MW267120" pageId="13" pageNumber="130">MW267120</accessionNumber>
to 
<accessionNumber id="AA81D2FFFF8FFFE0FD62FAE5FC63FA37" box="[761,904,1386,1412]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MW267122" pageId="13" pageNumber="130">MW267122</accessionNumber>
(3 barcoded); 
<specimenCode id="E574E767FF8FFFE0FBAEFAE5FADAFA37" box="[1077,1329,1386,1412]" pageId="13" pageNumber="130">ZRC_ENT00007870</specimenCode>
; 
<collectionCode id="D3C3D7D9FF8FFFE0FAA6FAE5FA9DFA37" box="[1341,1398,1386,1412]" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="13" pageNumber="130">ZRC</collectionCode>
</materialsCitation>
• 
<materialsCitation id="05BA4541FF8FFFE0FF54FA02FD52FA78" ID-GBIF-Occurrence="3053841359" collectingDate="2018-12-04" collectionCode="THNHM" collectorName="W. Jaitrong" country="Thailand" county="Puar [sic] District" location="Sirapach Waterfall" municipality="District" pageId="13" pageNumber="130" specimenCount="14" specimenCount-queen="1" specimenCount-worker="13" stateProvince="Nan Province">
<specimenCount id="A3D48495FF8FFFE0FF54FA02FEBDFA14" box="[207,342,1421,1447]" pageId="13" pageNumber="130" type="worker">13 workers</specimenCount>
, 
<specimenCount id="A3D48495FF8FFFE0FEF8FA02FE28FA1B" box="[355,451,1421,1448]" pageId="13" pageNumber="130" type="queen">1 queen</specimenCount>
; 
<collectingRegion id="771681FEFF8FFFE0FE54FA01FD9FFA14" box="[463,628,1421,1448]" country="Thailand" name="Nan" pageId="13" pageNumber="130">Nan Province</collectingRegion>
, 
<collectingCounty id="5C0C3790FF8FFFE0FD1AFA01FCBCFA14" box="[641,855,1420,1448]" pageId="13" pageNumber="130">Puar [sic] District</collectingCounty>
, 
<location id="B00D19C7FF8FFFE0FCF8FA02FBAFFA14" LSID="urn:lsid:plazi:treatment:3D7BFE0AFF89FFFEFDA2F95CFAA4FA39:B00D19C7FF8FFFE0FCF8FA02FBAFFA14" box="[867,1092,1421,1447]" country="Thailand" county="Puar [sic] District" municipality="District" name="Sirapach Waterfall" pageId="13" pageNumber="130" stateProvince="Nan Province">Sirapach Waterfall</location>
; 
<date id="C16C69DCFF8FFFE0FBD4FA02FB08FA14" box="[1103,1251,1421,1448]" pageId="13" pageNumber="130" value="2018-12-04">
<collectingDate id="D1289034FF8FFFE0FBD4FA02FB08FA14" box="[1103,1251,1421,1448]" pageId="13" pageNumber="130" value="2018-12-04">4 Dec. 2018</collectingDate>
</date>
; 
<collectorName id="18272ACAFF8FFFE0FB74FA01FA9DFA14" box="[1263,1398,1421,1448]" pageId="13" pageNumber="130">W. Jaitrong</collectorName>
leg.; colony code: WJT041218-2; 
<collectionCode id="D3C3D7D9FF8FFFE0FDD6FA3EFD52FA78" box="[589,697,1457,1483]" pageId="13" pageNumber="130">THNHM</collectionCode>
</materialsCitation>
• 
<materialsCitation id="05BA4541FF8FFFE0FD4BFA3FFBD1FA5D" ID-GBIF-Occurrence="3053841311" accessionNumber="MW267123, MW267125" collectingDate="2018-12-04" collectionCode="MW, ZRC" collectorName="W. Jaitrong" country="Thailand" county="Puar" location="GenBank" municipality="District" pageId="13" pageNumber="130" specimenCode="ZRC_ENT00007871" specimenCount="9" specimenCount-queen="1" specimenCount-worker="8" stateProvince="North Thailand">
<specimenCount id="A3D48495FF8FFFE0FD4BFA3FFCACFA79" box="[720,839,1456,1482]" pageId="13" pageNumber="130" type="worker">8 workers</specimenCount>
, 
<specimenCount id="A3D48495FF8FFFE0FCC9FA3FFC5AFA78" box="[850,945,1456,1483]" pageId="13" pageNumber="130" type="queen">1 queen</specimenCount>
; same collection data as for preceding; GenBank: 
<accessionNumber id="AA81D2FFFF8FFFE0FEDBFA5CFE24FA5E" box="[320,463,1491,1517]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MW267123" pageId="13" pageNumber="130">MW267123</accessionNumber>
to 
<accessionNumber id="AA81D2FFFF8FFFE0FE60FA5CFD61FA5D" box="[507,650,1491,1518]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MW267125" pageId="13" pageNumber="130">MW267125</accessionNumber>
(3 barcoded); 
<specimenCode id="E574E767FF8FFFE0FCD9FA5BFBD1FA5D" box="[834,1082,1492,1518]" pageId="13" pageNumber="130">ZRC_ENT00007871</specimenCode>
</materialsCitation>
• 
<materialsCitation id="05BA4541FF8FFFE0FBC1FA5BFC97F987" ID-GBIF-Occurrence="3053841338" collectingDate="2018-10-21" collectionCode="THNHM" collectorName="W. Jaitrong &amp; Sk. Yamane" country="Thailand" county="Muang district" location="Huai Chom Koo subdistrict" municipality="Huai Chom Koo" pageId="13" pageNumber="130" specimenCount="8" specimenCount-worker="8" stateProvince="Chiang Rai">
<specimenCount id="A3D48495FF8FFFE0FBC1FA5BFB3DFA5E" box="[1114,1238,1491,1518]" pageId="13" pageNumber="130" type="worker">8 workers</specimenCount>
; 
<collectingRegion id="771681FEFF8FFFE0FB7DFA5CFA99FA5E" box="[1254,1394,1491,1517]" country="Thailand" name="Chiang Rai" pageId="13" pageNumber="130">Chiang Rai</collectingRegion>
, 
<collectingCounty id="5C0C3790FF8FFFE0FF26FA78FE98F9A2" box="[189,371,1527,1553]" pageId="13" pageNumber="130">Muang district</collectingCounty>
, 
<location id="B00D19C7FF8FFFE0FE18FA78FD31F9A2" LSID="urn:lsid:plazi:treatment:3D7BFE0AFF89FFFEFDA2F95CFAA4FA39:B00D19C7FF8FFFE0FE18FA78FD31F9A2" box="[387,730,1527,1553]" country="Thailand" county="Muang district" municipality="Huai Chom Koo" name="Huai Chom Koo subdistrict" pageId="13" pageNumber="130" stateProvince="Chiang Rai">Huai Chom Koo subdistrict</location>
, 
<location id="B00D19C7FF8FFFE0FD71FA78FC06F9A2" LSID="urn:lsid:plazi:treatment:3D7BFE0AFF89FFFEFDA2F95CFAA4FA39:B00D19C7FF8FFFE0FD71FA78FC06F9A2" box="[746,1005,1527,1553]" country="Thailand" county="Muang district" municipality="Huai Chom Koo" name="Khun Korn waterfall" pageId="13" pageNumber="130" stateProvince="Chiang Rai">Khun Korn waterfall</location>
; 
<date id="C16C69DCFF8FFFE0FC65FA78FB4FF9A2" box="[1022,1188,1527,1553]" pageId="13" pageNumber="130" value="2018-10-21">
<collectingDate id="D1289034FF8FFFE0FC65FA78FB4FF9A2" box="[1022,1188,1527,1553]" pageId="13" pageNumber="130" value="2018-10-21">21 Oct. 2018</collectingDate>
</date>
; 
<collectorName id="18272ACAFF8FFFE0FB2FFA78FAAAF9A2" box="[1204,1345,1527,1553]" pageId="13" pageNumber="130">W. Jaitrong</collectorName>
and 
<collectorName id="18272ACAFF8FFFE0FF26F995FEA5F987" box="[189,334,1562,1588]" pageId="13" pageNumber="130">Sk. Yamane</collectorName>
leg.; colony code: TH18-SKY-169; 
<collectionCode id="D3C3D7D9FF8FFFE0FC8BF995FC97F987" box="[784,892,1562,1588]" pageId="13" pageNumber="130">THNHM</collectionCode>
</materialsCitation>
• 
<materialsCitation id="05BA4541FF8FFFE0FC06F995FB44F9E4" ID-GBIF-Occurrence="3053841306" accessionNumber="MW267102, MW267104" collectingDate="2018-10-21" collectionCode="MW, ZRC" collectorName="W. Jaitrong &amp; Sk. Yamane" country="Thailand" county="Muang district" location="GenBank" municipality="Huai Chom Koo" pageId="13" pageNumber="130" specimenCode="ZRC_ENT00007884" specimenCount="8" specimenCount-worker="8" stateProvince="North Thailand">
<specimenCount id="A3D48495FF8FFFE0FC06F995FBF1F987" box="[925,1050,1562,1588]" pageId="13" pageNumber="130" type="worker">8 workers</specimenCount>
; same collection data as for preceding; GenBank: 
<accessionNumber id="AA81D2FFFF8FFFE0FE5AF9B2FDBAF9E4" box="[449,593,1597,1623]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MW267102" pageId="13" pageNumber="130">MW267102</accessionNumber>
to 
<accessionNumber id="AA81D2FFFF8FFFE0FDECF9B2FCEDF9E4" box="[631,774,1597,1623]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MW267104" pageId="13" pageNumber="130">MW267104</accessionNumber>
(3 barcoded); 
<specimenCode id="E574E767FF8FFFE0FC2FF9B2FB44F9E4" box="[948,1199,1597,1623]" pageId="13" pageNumber="130">ZRC_ENT00007884</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="FDC81C97FF8FFFFFFF26F9F1FEF8FBD2" lastPageId="18" lastPageNumber="135" pageId="13" pageNumber="130" type="description">
<paragraph id="B56D4F1CFF8FFFE0FF26F9F1FEA5F92B" blockId="13.[189,334,1662,1688]" box="[189,334,1662,1688]" pageId="13" pageNumber="130">
<heading id="EE25F870FF8FFFE0FF26F9F1FEA5F92B" bold="true" box="[189,334,1662,1688]" fontSize="11" level="3" pageId="13" pageNumber="130" reason="3">
<emphasis id="87A6930EFF8FFFE0FF26F9F1FEA5F92B" bold="true" box="[189,334,1662,1688]" pageId="13" pageNumber="130">Description</emphasis>
</heading>
</paragraph>
<paragraph id="B56D4F1CFF8FFFE0FF71F922FEA7F974" blockId="13.[189,1399,1709,1841]" box="[234,332,1709,1735]" pageId="13" pageNumber="130">
<emphasis id="87A6930EFF8FFFE0FF71F922FEA7F974" bold="true" box="[234,332,1709,1735]" pageId="13" pageNumber="130">Worker</emphasis>
</paragraph>
<paragraph id="B56D4F1CFF8FFFE0FF26F95EFB8DF882" blockId="13.[189,1399,1709,1841]" pageId="13" pageNumber="130">
Measurements. 
<typeStatus id="6A69F1BEFF8FFFE0FEECF95EFE07F958" box="[375,492,1745,1771]" pageId="13" pageNumber="130" type="lectotype">Lectotype</typeStatus>
and 
<typeStatus id="6A69F1BEFF8FFFE0FDBCF95FFD2EF959" box="[551,709,1744,1770]" pageId="13" pageNumber="130" type="paralectotype">paralectotype</typeStatus>
workers (n = 3), measurements from 
<bibRefCitation id="D14332EDFF8FFFE0FB18F95EFA98F958" author="Wang W. Y. &amp; Yong G. W. &amp; Jaitrong W." box="[1155,1395,1744,1771]" pageId="13" pageNumber="130" pagination="301 - 340" refId="ref24028" refString="Wang W. Y., Yong G. W. &amp; Jaitrong W. 2018 b. The ant genus Rhopalomastix (Hymenoptera: Formicidae: Myrmicinae) in Southeast Asia, with descriptions of four new species from Singapore based on morphology and DNA barcoding. Zootaxa 4532 (3): 301 - 340. https: // doi. org / 10.11646 / zootaxa. 4532.3.1" type="journal article" year="2018">
Wang 
<emphasis id="87A6930EFF8FFFE0FB54F95EFAE1F959" box="[1231,1290,1744,1770]" italics="true" pageId="13" pageNumber="130">et al.</emphasis>
(2018b)
</bibRefCitation>
: EL 0.08; EW 0.04–0.06; HL 0.48–0.52; HW 0.45–0.48; ML 0.48–0.50; PronW 0.28–0.32; PtH, PtL unavailable; SL 0.18–0.20; TL 1.83–1.98; CI 92–94; REL 17–18; SI 39–42.
</paragraph>
<paragraph id="B56D4F1CFF8FFFE0FF26F8D6FD61F80A" blockId="13.[189,1399,1880,1977]" pageId="13" pageNumber="130">
Non-type workers from 
<collectingCountry id="CDC50F8CFF8FFFE0FE42F8D7FDBBF8C1" box="[473,592,1880,1906]" name="Singapore" pageId="13" pageNumber="130">Singapore</collectingCountry>
(n = 9): EL 0.08–0.1; EW 0.04–0.06; HL 0.46–0.48; HW 0.41–0.44; ML 0.50–0.54; PronW 0.28–0.32; PtH 0.15–0.18; PtL 0.17–0.20; TL 1.69–2.02; SL 0.18–0.19; CI 89– 93; PtHI 87–90; REL 18–23; SI 41–44.
</paragraph>
<paragraph id="B56D4F1CFF8CFFE3FF26FE86FD61FEDA" blockId="14.[188,1398,265,362]" pageId="14" pageNumber="131">
Non-type workers from 
<collectingCountry id="CDC50F8CFF8CFFE3FE4EFE86FDD6FE90" box="[469,573,265,291]" name="Thailand" pageId="14" pageNumber="131">Thailand</collectingCountry>
(n = 13): EL 0.08–0.12; EW 0.06–0.08; HL 0.46–0.58; HW 0.46–0.56; ML 0.56–0.76; PronW 0.30–0.36; PtH 0.17–0.23; PtL 0.19–0.25; TL 2.02– 2.88; SL 0.20–0.26; CI 92– 96; PtHI 89–92; REL 17–21; SI 44–48.
</paragraph>
<paragraph id="B56D4F1CFF8CFFE2FF26FE1EFD03FDA9" blockId="14.[187,1400,401,850]" lastBlockId="15.[189,1401,265,538]" lastPageId="15" lastPageNumber="132" pageId="14" pageNumber="131">
Workers monomorphic with little intranidal variation in size. Head in full-face view subrectangular, longer than wide, posterior margin with a broad but shallow median concavity, posterolateral corners roundly convex, lateral margins of head broadly convex and weakly converging anteriorly, anterior clypeal margin less wide than posterior margin of head, with broadly and weakly convex median section (
<figureCitation id="2DE95399FF8CFFE3FF5EFD91FEF1FD8B" box="[197,282,541,568]" captionStart="Fig" captionStartId="14.[189,232,1913,1939]" captionTargetBox="[227,1361,933,1860]" captionTargetId="figure-234@14.[227,1361,933,1860]" captionTargetPageId="14" captionText="Fig. 5. Rhopalomastix javana Wheeler, 1929, non-type, worker, West Thailand (ZRC_ENT00000932). a. Head in full-face view. b. Body in dorsal view. c. Close-up of clypeus and head in profile. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610573" httpUri="https://zenodo.org/record/4610573/files/figure.png" pageId="14" pageNumber="131">Fig. 5a</figureCitation>
). Eye with 11–16 ommatidia for 
<typeStatus id="6A69F1BEFF8CFFE3FD07FD91FD31FD8B" box="[668,730,542,568]" pageId="14" pageNumber="131">types</typeStatus>
and specimens for 
<collectingCountry id="CDC50F8CFF8CFFE3FC26FD91FBDCFD8B" box="[957,1079,542,568]" name="Singapore" pageId="14" pageNumber="131">Singapore</collectingCountry>
, 13–21 ommatidia for Thai specimens. In lateral view, outline of clypeus evenly and weakly convex, projecting slightly forward from dorsal margin of head (
<figureCitation id="2DE95399FF8CFFE3FE57FDEBFDC9FDCC" box="[460,546,612,639]" captionStart="Fig" captionStartId="14.[189,232,1913,1939]" captionTargetBox="[227,1361,933,1860]" captionTargetId="figure-234@14.[227,1361,933,1860]" captionTargetPageId="14" captionText="Fig. 5. Rhopalomastix javana Wheeler, 1929, non-type, worker, West Thailand (ZRC_ENT00000932). a. Head in full-face view. b. Body in dorsal view. c. Close-up of clypeus and head in profile. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610573" httpUri="https://zenodo.org/record/4610573/files/figure.png" pageId="14" pageNumber="131">Fig. 5c</figureCitation>
). Mesosoma in lateral view box-shaped or subcylindrical, dorsal outline weakly and broadly convex, propodeal junction roundly obtuse, propodeal declivity steep and shallowly concave (
<figureCitation id="2DE95399FF8CFFE3FEB5FD25FE6DFD76" box="[302,390,682,709]" captionStart="Fig" captionStartId="14.[189,232,1913,1939]" captionTargetBox="[227,1361,933,1860]" captionTargetId="figure-234@14.[227,1361,933,1860]" captionTargetPageId="14" captionText="Fig. 5. Rhopalomastix javana Wheeler, 1929, non-type, worker, West Thailand (ZRC_ENT00000932). a. Head in full-face view. b. Body in dorsal view. c. Close-up of clypeus and head in profile. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610573" httpUri="https://zenodo.org/record/4610573/files/figure.png" pageId="14" pageNumber="131">Fig. 5d</figureCitation>
); dorsum of propodeum differentiated from posterior declivitous face by a weak but distinct marginate edge. In dorsal view, mesosoma subrectangular with lateral sides almost parallel to each other; anterior margin of pronotal disc broadly convex, sometimes angulate at its median point, humeral corners rounded and often indistinctly angulate; lateral margins of propodeum broadly convex, transitioning to and differentiated from weakly concave posterior propodeal margin at bluntly angulate posterolateral corners (
<figureCitation id="2DE95399FF8DFFE2FE57FE86FDC8FE97" box="[460,547,265,292]" captionStart="Fig" captionStartId="14.[189,232,1913,1939]" captionTargetBox="[227,1361,933,1860]" captionTargetId="figure-234@14.[227,1361,933,1860]" captionTargetPageId="14" captionText="Fig. 5. Rhopalomastix javana Wheeler, 1929, non-type, worker, West Thailand (ZRC_ENT00000932). a. Head in full-face view. b. Body in dorsal view. c. Close-up of clypeus and head in profile. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610573" httpUri="https://zenodo.org/record/4610573/files/figure.png" pageId="15" pageNumber="132">Fig. 5b</figureCitation>
); posterolateral corners slightly projecting posteriorly and differentiated from inner posterior propodeal face. Posterior face of propodeum weakly marginate, rounding into lateral propodeal face at indistinct angulate edge. In lateral view, petiole inclined posteriorly, slightly longer than high, distinctly narrower at apex than at base, anterior face weakly concave and longer than posterior face; apex roundly convex, apical dorsum rounding smoothly into steeper posterior face at indistinct rounded edge; petiole higher and longer than postpetiole, dorsal margin of postpetiole weakly convex and nearly flat (
<figureCitation id="2DE95399FF8DFFE2FE4EFE53FDC5FE44" box="[469,558,476,503]" captionStart="Fig" captionStartId="14.[189,232,1913,1939]" captionTargetBox="[227,1361,933,1860]" captionTargetId="figure-234@14.[227,1361,933,1860]" captionTargetPageId="14" captionText="Fig. 5. Rhopalomastix javana Wheeler, 1929, non-type, worker, West Thailand (ZRC_ENT00000932). a. Head in full-face view. b. Body in dorsal view. c. Close-up of clypeus and head in profile. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610573" httpUri="https://zenodo.org/record/4610573/files/figure.png" pageId="15" pageNumber="132">Fig. 5d</figureCitation>
). In dorsal view, petiole suboval, wider than long; postpetiole globular and less than twice as wide as petiole (
<figureCitation id="2DE95399FF8DFFE2FD18FE70FD31FDA9" box="[643,730,511,538]" captionStart="Fig" captionStartId="14.[189,232,1913,1939]" captionTargetBox="[227,1361,933,1860]" captionTargetId="figure-234@14.[227,1361,933,1860]" captionTargetPageId="14" captionText="Fig. 5. Rhopalomastix javana Wheeler, 1929, non-type, worker, West Thailand (ZRC_ENT00000932). a. Head in full-face view. b. Body in dorsal view. c. Close-up of clypeus and head in profile. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610573" httpUri="https://zenodo.org/record/4610573/files/figure.png" pageId="15" pageNumber="132">Fig. 5b</figureCitation>
).
</paragraph>
<caption id="E1AD1F94FF8CFFE3FF26F8F6FEA7F86A" ID-DOI="http://doi.org/10.5281/zenodo.4610573" ID-Zenodo-Dep="4610573" httpUri="https://zenodo.org/record/4610573/files/figure.png" pageId="14" pageNumber="131" startId="14.[189,232,1913,1939]" targetBox="[227,1361,933,1860]" targetPageId="14">
<paragraph id="B56D4F1CFF8CFFE3FF26F8F6FEA7F86A" blockId="14.[189,1399,1913,2009]" pageId="14" pageNumber="131">
<emphasis id="87A6930EFF8CFFE3FF26F8F6FEE0F820" bold="true" box="[189,267,1913,1939]" pageId="14" pageNumber="131">Fig. 5.</emphasis>
<taxonomicName id="72D2349FFF8CFFE3FE89F8F6FD3BF820" authority="Wheeler, 1929" authorityName="Wheeler" authorityYear="1929" box="[274,720,1913,1939]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="131" phylum="Arthropoda" rank="species" species="javana">
<emphasis id="87A6930EFF8CFFE3FE89F8F6FDF1F820" box="[274,538,1913,1939]" italics="true" pageId="14" pageNumber="131">Rhopalomastix javana</emphasis>
<bibRefCitation id="D14332EDFF8CFFE3FDB9F8F6FD3BF820" author="Wheeler W. M." box="[546,720,1913,1939]" pageId="14" pageNumber="131" pagination="95 - 101" refId="ref24237" refString="Wheeler W. M. 1929. The ant genus Rhopalomastix. Psyche 36 (2): 95 - 101." type="journal article" year="1929">Wheeler, 1929</bibRefCitation>
</taxonomicName>
, non-type, worker, West Thailand (ZRC_ENT00000932). 
<emphasis id="87A6930EFF8CFFE3FF26F813FF20F805" bold="true" box="[189,203,1948,1974]" pageId="14" pageNumber="131">a</emphasis>
. Head in full-face view. 
<emphasis id="87A6930EFF8CFFE3FE76F813FE16F805" bold="true" box="[493,509,1948,1974]" pageId="14" pageNumber="131">b</emphasis>
. Body in dorsal view. 
<emphasis id="87A6930EFF8CFFE3FC9FF813FCF2F805" bold="true" box="[772,793,1948,1974]" pageId="14" pageNumber="131">c.</emphasis>
Close-up of clypeus and head in profile. 
<emphasis id="87A6930EFF8CFFE3FB61F813FAE1F805" bold="true" box="[1274,1290,1948,1974]" pageId="14" pageNumber="131">d</emphasis>
. Body in lateral view.
</paragraph>
</caption>
<paragraph id="B56D4F1CFF8DFFE2FF26FDCEFEF0FC6D" blockId="15.[188,1400,577,990]" pageId="15" pageNumber="132">
Dorsum and lateral face of head largely finely striate and shiny, striations on posterior half of head generally weaker and more fragmented; posterolateral corners of head feebly reticulate with interspaces smooth and shining; posteriormost strip of vertexal area just above occiput in posterodorsal view largely unsculptured or with feeble striations, mostly smooth and shiny. Ventral face of posterior half of head mostly superficially reticulate with interspaces smooth and shining. Mandible mostly smooth and shining, area immediately adjacent to masticatory margin punctate (
<figureCitation id="2DE95399FF8DFFE2FB83FD7EFB84FCB8" box="[1048,1135,753,779]" captionStart="Fig" captionStartId="5.[189,232,1816,1842]" captionTargetBox="[227,1361,973,1778]" captionTargetId="figure-170@5.[227,1361,973,1778]" captionTargetPageId="5" captionText="Fig. 2. Closeups of external mandibular surface for workers of 5 Thai species of Rhopalomastix Forel, 1900 (ZRC); white arrows indicate distinct striations on the external mandibular face. a. R. impithuksai Wang &amp; Jaitrong sp. nov., paratype (ZRC_ENT00007608). b. R. robusta Wang &amp; Jaitrong sp. nov., paratype (ZRC_ENT00013949). c. R. javana Wheeler, 1929, non-type (ZRC_ENT00000932). d. R. johorensis Wheeler, 1929, non-type (ZRC_ENT00007874). e. R. parva Wang &amp; Jaitrong sp. nov., paratype (ZRC_ENT00013948)." figureDoi="http://doi.org/10.5281/zenodo.4610567" httpUri="https://zenodo.org/record/4610567/files/figure.png" pageId="15" pageNumber="132">Fig. 2c</figureCitation>
). Median clypeal face with coarse irregular striae, interspaces punctate and weakly shining (
<figureCitation id="2DE95399FF8DFFE2FC70FC9BFBD4FC9C" box="[1003,1087,788,815]" captionStart="Fig" captionStartId="14.[189,232,1913,1939]" captionTargetBox="[227,1361,933,1860]" captionTargetId="figure-234@14.[227,1361,933,1860]" captionTargetPageId="14" captionText="Fig. 5. Rhopalomastix javana Wheeler, 1929, non-type, worker, West Thailand (ZRC_ENT00000932). a. Head in full-face view. b. Body in dorsal view. c. Close-up of clypeus and head in profile. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610573" httpUri="https://zenodo.org/record/4610573/files/figure.png" pageId="15" pageNumber="132">Fig. 5a</figureCitation>
), sometimes with a central longitudinal strip more weakly striate or largely smooth and shining. Lateral face of mesosoma largely striate-reticulate with interspaces smooth and shining (
<figureCitation id="2DE95399FF8DFFE2FCC1FCD5FC5EFCC6" box="[858,949,858,885]" captionStart="Fig" captionStartId="14.[189,232,1913,1939]" captionTargetBox="[227,1361,933,1860]" captionTargetId="figure-234@14.[227,1361,933,1860]" captionTargetPageId="14" captionText="Fig. 5. Rhopalomastix javana Wheeler, 1929, non-type, worker, West Thailand (ZRC_ENT00000932). a. Head in full-face view. b. Body in dorsal view. c. Close-up of clypeus and head in profile. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610573" httpUri="https://zenodo.org/record/4610573/files/figure.png" pageId="15" pageNumber="132">Fig. 5d</figureCitation>
); dorsum of mesosoma finely striate (
<figureCitation id="2DE95399FF8DFFE2FF5DFCF1FEF4FC2B" box="[198,287,893,920]" captionStart="Fig" captionStartId="14.[189,232,1913,1939]" captionTargetBox="[227,1361,933,1860]" captionTargetId="figure-234@14.[227,1361,933,1860]" captionTargetPageId="14" captionText="Fig. 5. Rhopalomastix javana Wheeler, 1929, non-type, worker, West Thailand (ZRC_ENT00000932). a. Head in full-face view. b. Body in dorsal view. c. Close-up of clypeus and head in profile. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610573" httpUri="https://zenodo.org/record/4610573/files/figure.png" pageId="15" pageNumber="132">Fig. 5b</figureCitation>
). Lateral face of petiole superficially reticulate with interspaces smooth and shining, petiolar apex finely striate and shiny. Postpetiole and gaster superficially reticulate with interspaces smooth and shining.
</paragraph>
<paragraph id="B56D4F1CFF8DFFE2FF26FB89FD77FACC" blockId="15.[188,1401,1029,1408]" pageId="15" pageNumber="132">Dorsum of head with fairly dense though well-spaced short erect and suberect hairs interspersed with sparse long erect hairs, gena with fairly dense short appressed and decumbent hairs. Ventral face of head with fairly dense though well-spaced short suberect, decumbent and appressed hairs. Antennal funiculus mostly with fairly dense appressed and decumbent hairs. Mesosoma dorsum with sparse, scattered short standing hairs that are relatively denser closer to dorsolateral margin, paired long erect hairs sparsely present along dorsolateral margin, with one pair each flanking the anterior margin of pronotal disc, pronotum, mesonotum and propodeum. Anterior rising face of petiole with fairly dense appressed and decumbent hairs, posterior declivity with dense short erect and subdecumbent hairs; apex of petiolar node with few short erect hairs and 1–2 pairs of long erect hairs. Postpetiole with dense short erect and suberect hairs, and one pair of long erect hairs. Gaster pilose, with dense suberect and erect hairs interspersed with sparse long erect hairs.</paragraph>
<paragraph id="B56D4F1CFF8DFFE2FF26FA28FCEFF9B4" blockId="15.[189,1399,1447,1544]" pageId="15" pageNumber="132">Head, dorsum of mesosoma and gaster generally dark brown, rest of body lighter yellowish brown, dorsum of head and area around eye in some workers with blackish undertones; tips of antennae, first funicular segment and legs pale yellowish brown.</paragraph>
<paragraph id="B56D4F1CFF8DFFE2FF71F9A1FED0F9FB" blockId="15.[188,1398,1582,1715]" box="[234,315,1582,1608]" pageId="15" pageNumber="132">
<emphasis id="87A6930EFF8DFFE2FF71F9A1FED0F9FB" bold="true" box="[234,315,1582,1608]" pageId="15" pageNumber="132">Queen</emphasis>
</paragraph>
<paragraph id="B56D4F1CFF8DFFE2FF26F9DDFDB6F900" blockId="15.[188,1398,1582,1715]" pageId="15" pageNumber="132">Measurements. Non-type queens (n = 3): EL 0.15–0.16; EW 0.09–0.10; HL 0.48–0.49; HW 0.44–0.45; ML 0.70–0.72; MsW 0.36–0.40; PtH 0.21–0.22; PtL 0.25–0.26; SL 0.18–0.20; TL 2.21–2.43; CI 92–94; PtHI 85–88; REL 33–36; SI 40–45.</paragraph>
<paragraph id="B56D4F1CFF8DFFFDFF26F955FBCDFD79" blockId="15.[187,1400,1754,1992]" lastBlockId="16.[188,1401,265,714]" lastPageId="16" lastPageNumber="133" pageId="15" pageNumber="132">
Small-medium size (relative to 
<taxonomicName id="72D2349FFF8DFFE2FDA7F954FD32F947" authorityName="Wheeler" authorityYear="1929" box="[572,729,1754,1780]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="132" phylum="Arthropoda" rank="species" species="johorensis">
<emphasis id="87A6930EFF8DFFE2FDA7F954FD32F947" box="[572,729,1754,1780]" italics="true" pageId="15" pageNumber="132">R. johorensis</emphasis>
</taxonomicName>
queen). Head in full-face view subrectangular, slightly longer than wide, slightly narrower anteriorly with anterior clypeal margin distinctly less wide than posterior margin; lateral margins broadly and weakly convex, almost parallel to each other, posterolateral corners rounded, posterior margin nearly straight with a shallow median concavity (
<figureCitation id="2DE95399FF8DFFE2FB05F8CCFB1DF8ED" box="[1182,1270,1859,1886]" captionStart="Fig" captionStartId="16.[190,233,1926,1952]" captionTargetBox="[227,1361,1098,1871]" captionTargetId="figure-312@16.[227,1361,1097,1871]" captionTargetPageId="16" captionText="Fig. 6. Rhopalomastix javana Wheeler, 1929, non-type, queen, Northeast Thailand (THNHM-I-21751). a. Head in full-face view. b. Body in dorsal view. c. Close-up of mesosoma in dorsal view. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610575" httpUri="https://zenodo.org/record/4610575/files/figure.png" pageId="15" pageNumber="132">Fig. 6a</figureCitation>
). Eye very large (13–18 ommatidia in the longest axis); posteriormost point of eye located slightly below transverse midline of head in full-face view. In profile, clypeus weakly and broadly convex, sometimes almost flat (
<figureCitation id="2DE95399FF8DFFE2FF5DF822FEF4F87B" box="[198,287,1965,1992]" captionStart="Fig" captionStartId="16.[190,233,1926,1952]" captionTargetBox="[227,1361,1098,1871]" captionTargetId="figure-312@16.[227,1361,1097,1871]" captionTargetPageId="16" captionText="Fig. 6. Rhopalomastix javana Wheeler, 1929, non-type, queen, Northeast Thailand (THNHM-I-21751). a. Head in full-face view. b. Body in dorsal view. c. Close-up of mesosoma in dorsal view. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610575" httpUri="https://zenodo.org/record/4610575/files/figure.png" pageId="15" pageNumber="132">Fig. 6d</figureCitation>
). Ocelli oval in shape and generally equal in size, roughly equidistant from each other; small area immediately anterior to median ocellus gently depressed (
<figureCitation id="2DE95399FF92FFFDFC2BFE86FBCCFE97" box="[944,1063,265,292]" captionStart="Fig" captionStartId="16.[190,233,1926,1952]" captionTargetBox="[227,1361,1098,1871]" captionTargetId="figure-312@16.[227,1361,1097,1871]" captionTargetPageId="16" captionText="Fig. 6. Rhopalomastix javana Wheeler, 1929, non-type, queen, Northeast Thailand (THNHM-I-21751). a. Head in full-face view. b. Body in dorsal view. c. Close-up of mesosoma in dorsal view. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610575" httpUri="https://zenodo.org/record/4610575/files/figure.png" pageId="16" pageNumber="133">Fig. 6a–b</figureCitation>
). In lateral view, mesosoma subcylindrical, more elongate than that of worker, dorsal outline weakly and broadly convex, propodeal junction roundly obtuse and indistinct (
<figureCitation id="2DE95399FF92FFFDFD15FEC0FD0DFED9" box="[654,742,335,362]" captionStart="Fig" captionStartId="16.[190,233,1926,1952]" captionTargetBox="[227,1361,1098,1871]" captionTargetId="figure-312@16.[227,1361,1097,1871]" captionTargetPageId="16" captionText="Fig. 6. Rhopalomastix javana Wheeler, 1929, non-type, queen, Northeast Thailand (THNHM-I-21751). a. Head in full-face view. b. Body in dorsal view. c. Close-up of mesosoma in dorsal view. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610575" httpUri="https://zenodo.org/record/4610575/files/figure.png" pageId="16" pageNumber="133">Fig. 6d</figureCitation>
). Propodeal dorsum weakly convex (
<figureCitation id="2DE95399FF92FFFDFB05FEC0FB1DFED9" box="[1182,1270,335,362]" captionStart="Fig" captionStartId="16.[190,233,1926,1952]" captionTargetBox="[227,1361,1098,1871]" captionTargetId="figure-312@16.[227,1361,1097,1871]" captionTargetPageId="16" captionText="Fig. 6. Rhopalomastix javana Wheeler, 1929, non-type, queen, Northeast Thailand (THNHM-I-21751). a. Head in full-face view. b. Body in dorsal view. c. Close-up of mesosoma in dorsal view. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610575" httpUri="https://zenodo.org/record/4610575/files/figure.png" pageId="16" pageNumber="133">Fig. 6d</figureCitation>
), rounding into steep posterior declivity at indistinct rounded posterior edge; posterior propodeal face with shallow median concavity. In dorsal view, mesosoma subcylindrical, anterior margin of pronotal disc broadly convex, humeral corners obtusely angulate, mesoscutum slightly wider than long, propodeum much wider than long, posterior margin weakly concave and indistinct, lateral margin of propodeum rounding into posterior margin at indistinct rounded edge (
<figureCitation id="2DE95399FF92FFFDFC9CFE70FCB4FDA9" box="[775,863,511,538]" captionStart="Fig" captionStartId="16.[190,233,1926,1952]" captionTargetBox="[227,1361,1098,1871]" captionTargetId="figure-312@16.[227,1361,1097,1871]" captionTargetPageId="16" captionText="Fig. 6. Rhopalomastix javana Wheeler, 1929, non-type, queen, Northeast Thailand (THNHM-I-21751). a. Head in full-face view. b. Body in dorsal view. c. Close-up of mesosoma in dorsal view. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610575" httpUri="https://zenodo.org/record/4610575/files/figure.png" pageId="16" pageNumber="133">Fig. 6c</figureCitation>
). In lateral view, petiole inclined posteriorly, longer than high, anterior margin weakly concave and steep; petiolar node bell-shaped with bluntly rounded apex, apical face rounding smoothly into and not distinctly differentiated from posterior face, posterior face shorter and steeper than anterior face (
<figureCitation id="2DE95399FF92FFFDFCB1FDE6FC6AFD37" box="[810,897,617,644]" captionStart="Fig" captionStartId="16.[190,233,1926,1952]" captionTargetBox="[227,1361,1098,1871]" captionTargetId="figure-312@16.[227,1361,1097,1871]" captionTargetPageId="16" captionText="Fig. 6. Rhopalomastix javana Wheeler, 1929, non-type, queen, Northeast Thailand (THNHM-I-21751). a. Head in full-face view. b. Body in dorsal view. c. Close-up of mesosoma in dorsal view. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610575" httpUri="https://zenodo.org/record/4610575/files/figure.png" pageId="16" pageNumber="133">Fig. 6d</figureCitation>
). In lateral view, postpetiole not as long as petiole, dorsal margin broadly convex (
<figureCitation id="2DE95399FF92FFFDFD12FD03FD0AFD14" box="[649,737,652,679]" captionStart="Fig" captionStartId="16.[190,233,1926,1952]" captionTargetBox="[227,1361,1098,1871]" captionTargetId="figure-312@16.[227,1361,1097,1871]" captionTargetPageId="16" captionText="Fig. 6. Rhopalomastix javana Wheeler, 1929, non-type, queen, Northeast Thailand (THNHM-I-21751). a. Head in full-face view. b. Body in dorsal view. c. Close-up of mesosoma in dorsal view. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610575" httpUri="https://zenodo.org/record/4610575/files/figure.png" pageId="16" pageNumber="133">Fig. 6d</figureCitation>
). In dorsal view, petiole suboval, less than twice as wide as long; postpetiole globular, much larger and wider than petiole (
<figureCitation id="2DE95399FF92FFFDFC5AFD20FBF2FD79" box="[961,1049,687,714]" captionStart="Fig" captionStartId="16.[190,233,1926,1952]" captionTargetBox="[227,1361,1098,1871]" captionTargetId="figure-312@16.[227,1361,1097,1871]" captionTargetPageId="16" captionText="Fig. 6. Rhopalomastix javana Wheeler, 1929, non-type, queen, Northeast Thailand (THNHM-I-21751). a. Head in full-face view. b. Body in dorsal view. c. Close-up of mesosoma in dorsal view. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610575" httpUri="https://zenodo.org/record/4610575/files/figure.png" pageId="16" pageNumber="133">Fig. 6b</figureCitation>
).
</paragraph>
<paragraph id="B56D4F1CFF92FFFCFF26FD7EFC81FE3F" blockId="16.[186,1400,753,1026]" lastBlockId="17.[189,1399,265,397]" lastPageId="17" lastPageNumber="134" pageId="16" pageNumber="133">
Anterior half of head dorsum largely finely striate and shiny, posterior half mostly smooth and shining with scattered punctures, frons and vertex with feeble striations; clypeus coarsely striate but with a median longitudinal strip that is smooth and shining. Ventral face of head mostly smooth and shining. Mandible mostly smooth and shining, area next to masticatory margin slightly striate-punctate. Lateral face of mesosoma weakly substriate-reticulate, interspaces smooth and shining. Dorsum of mesosoma mostly finely striate and shining, median section of pronotum superficially reticulate with interspaces smooth and shining (
<figureCitation id="2DE95399FF92FFFDFE31FC4BFE16FC6C" box="[426,509,964,991]" captionStart="Fig" captionStartId="16.[190,233,1926,1952]" captionTargetBox="[227,1361,1098,1871]" captionTargetId="figure-312@16.[227,1361,1097,1871]" captionTargetPageId="16" captionText="Fig. 6. Rhopalomastix javana Wheeler, 1929, non-type, queen, Northeast Thailand (THNHM-I-21751). a. Head in full-face view. b. Body in dorsal view. c. Close-up of mesosoma in dorsal view. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610575" httpUri="https://zenodo.org/record/4610575/files/figure.png" pageId="16" pageNumber="133">Fig. 6c</figureCitation>
); lateral-most triangular area of mesoscutum immediately above parascutellar carina mostly smooth and shining or more weakly striate than rest of mesoscutum (
<figureCitation id="2DE95399FF92FFFDFB07FC68FB1FFBB1" box="[1180,1268,999,1026]" captionStart="Fig" captionStartId="16.[190,233,1926,1952]" captionTargetBox="[227,1361,1098,1871]" captionTargetId="figure-312@16.[227,1361,1097,1871]" captionTargetPageId="16" captionText="Fig. 6. Rhopalomastix javana Wheeler, 1929, non-type, queen, Northeast Thailand (THNHM-I-21751). a. Head in full-face view. b. Body in dorsal view. c. Close-up of mesosoma in dorsal view. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610575" httpUri="https://zenodo.org/record/4610575/files/figure.png" pageId="16" pageNumber="133">Fig. 6c</figureCitation>
); posterior face of propodeum mostly smooth and shining. Lateral faces of petiole and postpetiole superficially reticulate with interspaces smooth and shining; apicodorsal faces of petiole and postpetiole feebly and superficially reticulate with a central area that is smooth and shining. Gaster largely smooth and shining, area near anterior margin of tergite superficially reticulate.
</paragraph>
<caption id="E1AD1F94FF92FFFDFF25F809FEA6F855" ID-DOI="http://doi.org/10.5281/zenodo.4610575" ID-Zenodo-Dep="4610575" httpUri="https://zenodo.org/record/4610575/files/figure.png" pageId="16" pageNumber="133" startId="16.[190,233,1926,1952]" targetBox="[227,1361,1098,1871]" targetPageId="16">
<paragraph id="B56D4F1CFF92FFFDFF25F809FEA6F855" blockId="16.[188,1399,1926,2022]" pageId="16" pageNumber="133">
<emphasis id="87A6930EFF92FFFDFF25F809FEE1F813" bold="true" box="[190,266,1926,1952]" pageId="16" pageNumber="133">Fig. 6.</emphasis>
<taxonomicName id="72D2349FFF92FFFDFE8AF809FD20F813" authority="Wheeler, 1929" authorityName="Wheeler" authorityYear="1929" box="[273,715,1926,1952]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="133" phylum="Arthropoda" rank="species" species="javana">
<emphasis id="87A6930EFF92FFFDFE8AF809FDF3F813" box="[273,536,1926,1952]" italics="true" pageId="16" pageNumber="133">Rhopalomastix javana</emphasis>
<bibRefCitation id="D14332EDFF92FFFDFD85F809FD20F813" author="Wheeler W. M." box="[542,715,1926,1952]" pageId="16" pageNumber="133" pagination="95 - 101" refId="ref24237" refString="Wheeler W. M. 1929. The ant genus Rhopalomastix. Psyche 36 (2): 95 - 101." type="journal article" year="1929">Wheeler, 1929</bibRefCitation>
</taxonomicName>
, non-type, queen, Northeast Thailand (THNHM-I-21751). 
<emphasis id="87A6930EFF92FFFDFF27F826FF20F870" bold="true" box="[188,203,1961,1987]" pageId="16" pageNumber="133">a</emphasis>
. Head in full-face view. 
<emphasis id="87A6930EFF92FFFDFE68F826FDE9F870" bold="true" box="[499,514,1961,1987]" pageId="16" pageNumber="133">b</emphasis>
. Body in dorsal view. 
<emphasis id="87A6930EFF92FFFDFC8BF826FCF7F870" bold="true" box="[784,796,1961,1987]" pageId="16" pageNumber="133">c</emphasis>
. Close-up of mesosoma in dorsal view. 
<emphasis id="87A6930EFF92FFFDFB62F826FAE3F870" bold="true" box="[1273,1288,1961,1987]" pageId="16" pageNumber="133">d</emphasis>
. Body in lateral view.
</paragraph>
</caption>
<paragraph id="B56D4F1CFF93FFFCFF26FE3AFB0CFC9D" blockId="17.[188,1401,436,815]" pageId="17" pageNumber="134">
Dorsum of head with fairly dense short suberect and decumbent hairs, interspersed with very sparse long erect hairs, including one pair on median section of clypeus; lateral and ventral faces with fairly dense short appressed and decumbent hairs. Antennal scape with uniform short standing hairs and a long erect hair on basal and median points respectively. Dorsum of mesosoma with sparse short standing hairs (
<figureCitation id="2DE95399FF93FFFCFE93FDCEFE8AFDEF" box="[264,353,577,604]" captionStart="Fig" captionStartId="16.[190,233,1926,1952]" captionTargetBox="[227,1361,1098,1871]" captionTargetId="figure-312@16.[227,1361,1097,1871]" captionTargetPageId="16" captionText="Fig. 6. Rhopalomastix javana Wheeler, 1929, non-type, queen, Northeast Thailand (THNHM-I-21751). a. Head in full-face view. b. Body in dorsal view. c. Close-up of mesosoma in dorsal view. d. Body in lateral view." figureDoi="http://doi.org/10.5281/zenodo.4610575" httpUri="https://zenodo.org/record/4610575/files/figure.png" pageId="17" pageNumber="134">Fig. 6c</figureCitation>
); paired long erect hairs sparsely present along dorsolateral margin, with one pair each flanking the anterior margin of pronotal disc, pronotum, mesoscutum, and propodeum; mesoscutum and mesoscutal disc each one pair of long erect hairs. Anterior face of petiole with dense short appressed and/or decumbent hairs, posterior face with dense short erect hairs, apex with sparse short erect hairs, 1–2 pairs of long erect hairs. Postpetiole with dense short decumbent or standing hairs and two pairs of long erect hairs. Gaster pilose, with dense short appressed and/or decumbent hairs, scattered short standing hairs, and long erect hairs sparsely distributed along posterior margins of tergites.
</paragraph>
<paragraph id="B56D4F1CFF93FFFCFF26FCDAFD91FC20" blockId="17.[189,1398,853,915]" pageId="17" pageNumber="134">Body generally uniform dark brown; head, dorsum of mesosoma and gaster slightly darker brown; tibia and tarsus paler and yellowish brown.</paragraph>
<paragraph id="B56D4F1CFF93FFFCFF71FC35FEC1FC67" blockId="17.[187,1399,954,1086]" box="[234,298,954,980]" pageId="17" pageNumber="134">
<emphasis id="87A6930EFF93FFFCFF71FC35FEC1FC67" bold="true" box="[234,298,954,980]" pageId="17" pageNumber="134">Male</emphasis>
</paragraph>
<paragraph id="B56D4F1CFF93FFFCFF26FC51FA9CFBA8" blockId="17.[187,1399,954,1086]" pageId="17" pageNumber="134">
Measurements. Thai male specimens unavailable. Non-type males from 
<collectingCountry id="CDC50F8CFF93FFFCFC67FC52FB98FC44" box="[1020,1139,989,1015]" name="Singapore" pageId="17" pageNumber="134">Singapore</collectingCountry>
(n = 2): EL 0.18–0.20; EW 0.14; HL 0.39–0.41; HW 0.38–0.40; ML 0.66–0.70; MsW 0.40–0.44; PtH 0.17–0.18; PtL 0.21–
</paragraph>
<paragraph id="B56D4F1CFF93FFFCFF26FBABFC4BFB8D" blockId="17.[187,1399,954,1086]" box="[189,928,1059,1086]" pageId="17" pageNumber="134">0.22; SL 0.06; TL 1.98–1.99; CI 97–98; REL 45–53; SI 15–16.</paragraph>
<paragraph id="B56D4F1CFF93FFFCFF26FBEAFA89F995" blockId="17.[187,1399,1125,1574]" pageId="17" pageNumber="134">
Small size (relative to 
<taxonomicName id="72D2349FFF93FFFCFE21FBE9FDB9FBCC" authorityName="Wheeler" authorityYear="1929" box="[442,594,1125,1151]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="134" phylum="Arthropoda" rank="species" species="johorensis">
<emphasis id="87A6930EFF93FFFCFE21FBE9FDB9FBCC" box="[442,594,1125,1151]" italics="true" pageId="17" pageNumber="134">R. johorensis</emphasis>
</taxonomicName>
male). Head in full-face view sub-spherical, posterior margin broadly and strongly rounded (
<bibRefCitation id="D14332EDFF93FFFCFE5BFB07FD77FB11" author="Wang W. Y. &amp; Yong G. W. &amp; Jaitrong W." box="[448,668,1160,1186]" pageId="17" pageNumber="134" pagination="301 - 340" refId="ref24028" refString="Wang W. Y., Yong G. W. &amp; Jaitrong W. 2018 b. The ant genus Rhopalomastix (Hymenoptera: Formicidae: Myrmicinae) in Southeast Asia, with descriptions of four new species from Singapore based on morphology and DNA barcoding. Zootaxa 4532 (3): 301 - 340. https: // doi. org / 10.11646 / zootaxa. 4532.3.1" type="journal article" year="2018">
Wang 
<emphasis id="87A6930EFF93FFFCFD96FB06FDACFB11" box="[525,583,1160,1186]" italics="true" pageId="17" pageNumber="134">et al.</emphasis>
2018b
</bibRefCitation>
: fig. 21). Eye very large (20–25 ommatidia in the longest axis); posteriormost point of eye only slightly exceeding transverse midline of head, outline of eye bulging from lateral margin of head by a little (
<bibRefCitation id="D14332EDFF93FFFCFD06FB40FC6BFB5A" author="Wang W. Y. &amp; Yong G. W. &amp; Jaitrong W." box="[669,896,1231,1257]" pageId="17" pageNumber="134" pagination="301 - 340" refId="ref24028" refString="Wang W. Y., Yong G. W. &amp; Jaitrong W. 2018 b. The ant genus Rhopalomastix (Hymenoptera: Formicidae: Myrmicinae) in Southeast Asia, with descriptions of four new species from Singapore based on morphology and DNA barcoding. Zootaxa 4532 (3): 301 - 340. https: // doi. org / 10.11646 / zootaxa. 4532.3.1" type="journal article" year="2018">
Wang 
<emphasis id="87A6930EFF93FFFCFD70FB40FCC3FB5A" box="[747,808,1231,1257]" italics="true" pageId="17" pageNumber="134">et al.</emphasis>
2018b
</bibRefCitation>
: fig. 21). Median ocellus subcircular, less elongate and slightly smaller than lateral ocelli; small area of head dorsum immediately anterior to median ocellus weakly depressed, continuous with weak median furrow separating frontal lobes; frontal lobes strongly raised and differentiated from supraclypeal area (
<bibRefCitation id="D14332EDFF93FFFCFC2DFAB7FB72FAE1" author="Wang W. Y. &amp; Yong G. W. &amp; Jaitrong W." box="[950,1177,1336,1362]" pageId="17" pageNumber="134" pagination="301 - 340" refId="ref24028" refString="Wang W. Y., Yong G. W. &amp; Jaitrong W. 2018 b. The ant genus Rhopalomastix (Hymenoptera: Formicidae: Myrmicinae) in Southeast Asia, with descriptions of four new species from Singapore based on morphology and DNA barcoding. Zootaxa 4532 (3): 301 - 340. https: // doi. org / 10.11646 / zootaxa. 4532.3.1" type="journal article" year="2018">
Wang 
<emphasis id="87A6930EFF93FFFCFB9CFAB6FBAFFAE1" box="[1031,1092,1336,1362]" italics="true" pageId="17" pageNumber="134">et al.</emphasis>
2018b
</bibRefCitation>
: fig. 21). Posterior apex of supraclypeal area bluntly rounded. In lateral view, head broad-ovate with dorsum gently inclined posteriorly, posterodorsal outline broadly rounded with differentiation between dorsal and posterior faces indistinct, posterior margin rounding into broadly convex ventral margin forming a continuous curve with no distinct posteroventral angle (
<figureCitation id="2DE95399FF93FFFCFD25FA4AFCF8FA53" box="[702,787,1477,1504]" captionStart="Fig" captionStartId="18.[189,232,1956,1982]" captionTargetBox="[258,1330,1395,1916]" captionTargetId="figure-340@18.[258,1330,1395,1916]" captionTargetPageId="18" captionText="Fig. 7. Rhopalomastix javana Wheeler, 1929, non-type, ♂, Singapore (ZRC_HYM_0000576). a. Head and clypeus in profile. b. Close-up of genitalia in profile." figureDoi="http://doi.org/10.5281/zenodo.4610577" httpUri="https://zenodo.org/record/4610577/files/figure.png" pageId="17" pageNumber="134">Fig. 7a</figureCitation>
). In the same view, clypeus strongly projecting from dorsum of head, with distinct dorsal and anterior faces separated by a roundly angulate edge, dorsal face gently sloped downwards and nearly flat, anterior face steep and very weakly convex or flat (
<figureCitation id="2DE95399FF93FFFCFB64F984FABFF995" box="[1279,1364,1547,1574]" captionStart="Fig" captionStartId="18.[189,232,1956,1982]" captionTargetBox="[258,1330,1395,1916]" captionTargetId="figure-340@18.[258,1330,1395,1916]" captionTargetPageId="18" captionText="Fig. 7. Rhopalomastix javana Wheeler, 1929, non-type, ♂, Singapore (ZRC_HYM_0000576). a. Head and clypeus in profile. b. Close-up of genitalia in profile." figureDoi="http://doi.org/10.5281/zenodo.4610577" httpUri="https://zenodo.org/record/4610577/files/figure.png" pageId="17" pageNumber="134">Fig. 7a</figureCitation>
).
</paragraph>
<paragraph id="B56D4F1CFF93FFFFFF26F9C2FDE2FEF5" blockId="17.[189,1400,1613,1956]" lastBlockId="18.[189,1398,265,326]" lastPageId="18" lastPageNumber="135" pageId="17" pageNumber="134">
In lateral view, mesosoma sub-oblong, dorsal margin almost completely straight or very weakly and broadly convex, propodeal junction roundly obtuse; anterior face of pronotum about as high as anterodorsal face of mesoscutum (
<bibRefCitation id="D14332EDFF93FFFCFDFEF91CFCB9F91E" author="Wang W. Y. &amp; Yong G. W. &amp; Jaitrong W." box="[613,850,1683,1709]" pageId="17" pageNumber="134" pagination="301 - 340" refId="ref24028" refString="Wang W. Y., Yong G. W. &amp; Jaitrong W. 2018 b. The ant genus Rhopalomastix (Hymenoptera: Formicidae: Myrmicinae) in Southeast Asia, with descriptions of four new species from Singapore based on morphology and DNA barcoding. Zootaxa 4532 (3): 301 - 340. https: // doi. org / 10.11646 / zootaxa. 4532.3.1" type="journal article" year="2018">
Wang 
<emphasis id="87A6930EFF93FFFCFD2CF91BFD13F91E" box="[695,760,1683,1709]" italics="true" pageId="17" pageNumber="134">et al.</emphasis>
2018b
</bibRefCitation>
: fig. 20); propodeal dorsum weakly convex, rounding into steep and short posterior declivity at indistinct rounded posterior edge. In posterior view, posterior propodeal face indistinctly marginate, weakly differentiated from lateral propodeal face by indistinct rounded lateral margin. In dorsal view, mesosoma obovate, notauli absent, parapsidal line weakly present (
<bibRefCitation id="D14332EDFF93FFFCFE1CF8AFFD99F889" author="Wang W. Y. &amp; Yong G. W. &amp; Jaitrong W." box="[391,626,1824,1850]" pageId="17" pageNumber="134" pagination="301 - 340" refId="ref24028" refString="Wang W. Y., Yong G. W. &amp; Jaitrong W. 2018 b. The ant genus Rhopalomastix (Hymenoptera: Formicidae: Myrmicinae) in Southeast Asia, with descriptions of four new species from Singapore based on morphology and DNA barcoding. Zootaxa 4532 (3): 301 - 340. https: // doi. org / 10.11646 / zootaxa. 4532.3.1" type="journal article" year="2018">
Wang 
<emphasis id="87A6930EFF93FFFCFE42F8AEFDF3F889" box="[473,536,1824,1850]" italics="true" pageId="17" pageNumber="134">et al.</emphasis>
2018b
</bibRefCitation>
: figs 22–23). In lateral view, petiole inclined posteriorly, short and subtrapezoidal, longer than high, anterior margin long, steep and weakly concave, apical dorsum weakly convex, rounding into short posterior face at indistinct rounded edge; anteroventral extension of subpetiolar process shallow and subtriangular, longer at base than apex (
<bibRefCitation id="D14332EDFF93FFFCFBA8F805FAE5F817" author="Wang W. Y. &amp; Yong G. W. &amp; Jaitrong W." box="[1075,1294,1930,1956]" pageId="17" pageNumber="134" pagination="301 - 340" refId="ref24028" refString="Wang W. Y., Yong G. W. &amp; Jaitrong W. 2018 b. The ant genus Rhopalomastix (Hymenoptera: Formicidae: Myrmicinae) in Southeast Asia, with descriptions of four new species from Singapore based on morphology and DNA barcoding. Zootaxa 4532 (3): 301 - 340. https: // doi. org / 10.11646 / zootaxa. 4532.3.1" type="journal article" year="2018">
Wang 
<emphasis id="87A6930EFF93FFFCFBE5F804FB51F817" box="[1150,1210,1930,1956]" italics="true" pageId="17" pageNumber="134">et al.</emphasis>
2018b
</bibRefCitation>
: fig. 20). Dorsal outline of postpetiole weakly convex, differentiated from gaster by a weak but distinct cinctus (
<bibRefCitation id="D14332EDFF90FFFFFF5DFEA3FE4AFEF5" author="Wang W. Y. &amp; Yong G. W. &amp; Jaitrong W." box="[198,417,300,326]" pageId="18" pageNumber="135" pagination="301 - 340" refId="ref24028" refString="Wang W. Y., Yong G. W. &amp; Jaitrong W. 2018 b. The ant genus Rhopalomastix (Hymenoptera: Formicidae: Myrmicinae) in Southeast Asia, with descriptions of four new species from Singapore based on morphology and DNA barcoding. Zootaxa 4532 (3): 301 - 340. https: // doi. org / 10.11646 / zootaxa. 4532.3.1" type="journal article" year="2018">
Wang 
<emphasis id="87A6930EFF90FFFFFE89FEA2FEA6FEF5" box="[274,333,300,326]" italics="true" pageId="18" pageNumber="135">et al.</emphasis>
2018b
</bibRefCitation>
: fig. 20).
</paragraph>
<paragraph id="B56D4F1CFF90FFFFFF26FEE1FD79FD5B" blockId="18.[188,1401,366,744]" pageId="18" pageNumber="135">
Head mostly superficially substriate-reticulate with interspaces smooth and shining, and scattered punctures (
<bibRefCitation id="D14332EDFF90FFFFFEA5FE1EFDF2FE18" author="Wang W. Y. &amp; Yong G. W. &amp; Jaitrong W." box="[318,537,401,427]" pageId="18" pageNumber="135" pagination="301 - 340" refId="ref24028" refString="Wang W. Y., Yong G. W. &amp; Jaitrong W. 2018 b. The ant genus Rhopalomastix (Hymenoptera: Formicidae: Myrmicinae) in Southeast Asia, with descriptions of four new species from Singapore based on morphology and DNA barcoding. Zootaxa 4532 (3): 301 - 340. https: // doi. org / 10.11646 / zootaxa. 4532.3.1" type="journal article" year="2018">
Wang 
<emphasis id="87A6930EFF90FFFFFE11FE1DFE2EFE18" box="[394,453,401,427]" italics="true" pageId="18" pageNumber="135">et al.</emphasis>
2018b
</bibRefCitation>
: fig. 21); area of head dorsum enclosed by and around ocelli rugulose and weakly shining; median area immediately anterior to median ocellus largely smooth and shining; frontal lobes coarsely rugulose-reticulate and weakly shining; clypeus coarsely rugulose with interspaces punctate and weakly shining. Lateral face of mesosoma weakly striate-reticulate with interspaces smooth and shining (
<bibRefCitation id="D14332EDFF90FFFFFE28FD91FD7AFD8B" author="Wang W. Y. &amp; Yong G. W. &amp; Jaitrong W." box="[435,657,542,568]" pageId="18" pageNumber="135" pagination="301 - 340" refId="ref24028" refString="Wang W. Y., Yong G. W. &amp; Jaitrong W. 2018 b. The ant genus Rhopalomastix (Hymenoptera: Formicidae: Myrmicinae) in Southeast Asia, with descriptions of four new species from Singapore based on morphology and DNA barcoding. Zootaxa 4532 (3): 301 - 340. https: // doi. org / 10.11646 / zootaxa. 4532.3.1" type="journal article" year="2018">
Wang 
<emphasis id="87A6930EFF90FFFFFD9AFD91FDD7FD8B" box="[513,572,542,568]" italics="true" pageId="18" pageNumber="135">et al.</emphasis>
2018b
</bibRefCitation>
: fig. 20). In dorsal view, mesoscutum largely weakly substriate with interspaces smooth and shining, lateralmost area of mesoscutum in between parapsidal line and parascutellar carina unsculptured, largely smooth and shining; mesoscutal disc largely smooth and shining with feeble superficial striations; propodeum superficially reticulate with interspaces smooth and shining (
<bibRefCitation id="D14332EDFF90FFFFFECEFD24FDDFFD77" author="Wang W. Y. &amp; Yong G. W. &amp; Jaitrong W." box="[341,564,682,709]" pageId="18" pageNumber="135" pagination="301 - 340" refId="ref24028" refString="Wang W. Y., Yong G. W. &amp; Jaitrong W. 2018 b. The ant genus Rhopalomastix (Hymenoptera: Formicidae: Myrmicinae) in Southeast Asia, with descriptions of four new species from Singapore based on morphology and DNA barcoding. Zootaxa 4532 (3): 301 - 340. https: // doi. org / 10.11646 / zootaxa. 4532.3.1" type="journal article" year="2018">
Wang 
<emphasis id="87A6930EFF90FFFFFE38FD24FE34FD77" box="[419,479,682,708]" italics="true" pageId="18" pageNumber="135">et al.</emphasis>
2018b
</bibRefCitation>
: fig. 23). Petiole, postpetiole mostly reticulate with interspaces smooth and shining, apices smooth and shining.
</paragraph>
<paragraph id="B56D4F1CFF90FFFFFF26FC9FFF15FC6A" blockId="18.[188,1401,783,985]" pageId="18" pageNumber="135">
Dorsum of head with dense short standing hairs; lateral, ventral and posterior faces with dense short decumbent and appressed hairs (
<figureCitation id="2DE95399FF90FFFFFDA6FCBDFD78FCFE" box="[573,659,818,845]" captionStart="Fig" captionStartId="18.[189,232,1956,1982]" captionTargetBox="[258,1330,1395,1916]" captionTargetId="figure-340@18.[258,1330,1395,1916]" captionTargetPageId="18" captionText="Fig. 7. Rhopalomastix javana Wheeler, 1929, non-type, ♂, Singapore (ZRC_HYM_0000576). a. Head and clypeus in profile. b. Close-up of genitalia in profile." figureDoi="http://doi.org/10.5281/zenodo.4610577" httpUri="https://zenodo.org/record/4610577/files/figure.png" pageId="18" pageNumber="135">Fig. 7a</figureCitation>
). Dorsum of pronotum and mesoscutum with fairly dense short standing hairs; posterolateral margins of propodeum with sparse short erect hairs. Anterior rising face of petiole with few or no hairs, apicodorsal and posterior petiolar faces with short standing hairs; dorsum of postpetiole with short standing hairs. Gaster pilose with dense standing, decumbent and subdecumbent hairs.
</paragraph>
<paragraph id="B56D4F1CFF90FFFFFF26FB8FFEF8FBD2" blockId="18.[189,1398,1024,1121]" pageId="18" pageNumber="135">Body generally dark greyish-brown; head, dorsum of mesosoma and gaster dark blackish-brown; funiculus of antenna uniformly light brown; legs greyish brown with joints and tarsi pale brownish yellow.</paragraph>
</subSubSection>
<subSubSection id="FDC81C97FF90FFFFFF26FB07FCBFFA89" pageId="18" pageNumber="135" type="distribution">
<paragraph id="B56D4F1CFF90FFFFFF26FB07FDEFFB11" blockId="18.[189,516,1160,1186]" box="[189,516,1160,1186]" pageId="18" pageNumber="135">
<heading id="EE25F870FF90FFFFFF26FB07FDEFFB11" bold="true" box="[189,516,1160,1186]" fontSize="11" level="3" pageId="18" pageNumber="135" reason="3">
<emphasis id="87A6930EFF90FFFFFF26FB07FDEFFB11" bold="true" box="[189,516,1160,1186]" pageId="18" pageNumber="135">Distribution and binomics</emphasis>
</heading>
</paragraph>
<paragraph id="B56D4F1CFF90FFFFFF26FB38FCBFFA89" blockId="18.[189,1400,1207,1339]" pageId="18" pageNumber="135">
<collectingCountry id="CDC50F8CFF90FFFFFF26FB38FEC4FB62" box="[189,303,1207,1233]" name="Indonesia" pageId="18" pageNumber="135">Indonesia</collectingCountry>
(
<collectingRegion id="771681FEFF90FFFFFEDBFB38FE92FB62" box="[320,377,1207,1233]" country="Indonesia" name="Jawa Timur" pageId="18" pageNumber="135">Java</collectingRegion>
), 
<collectingCountry id="CDC50F8CFF90FFFFFE15FB38FDE3FB62" box="[398,520,1207,1233]" name="Singapore" pageId="18" pageNumber="135">Singapore</collectingCountry>
, 
<collectingCountry id="CDC50F8CFF90FFFFFD8FFB38FD97FB62" box="[532,636,1207,1233]" name="Thailand" pageId="18" pageNumber="135">Thailand</collectingCountry>
(
<figureCitation id="2DE95399FF90FFFFFD16FB38FD1EFB61" box="[653,757,1207,1234]" captionStart="Fig" captionStartId="37.[191,234,1923,1949]" captionTargetBox="[224,1364,1287,1873]" captionTargetId="figure-368@37.[224,1364,1287,1873]" captionTargetPageId="37" captionText="Fig. 16. Distribution maps of species of Rhopalomastix Forel, 1900 in Thailand. a. New species: R. impithuksai Wang &amp; Jaitrong sp. nov., R. parva Wang &amp; Jaitrong sp. nov., R. robusta Wang &amp; Jaitrong sp. nov. b. R. javana Wheeler, 1929 and R. johorensis Wheeler, 1929." figureDoi="http://doi.org/10.5281/zenodo.4610595" httpUri="https://zenodo.org/record/4610595/files/figure.png" pageId="18" pageNumber="135">Fig. 16b</figureCitation>
). As with 
<taxonomicName id="72D2349FFF90FFFFFCF5FB37FBECFB62" authorityName="Wheeler" authorityYear="1929" box="[878,1031,1207,1233]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="135" phylum="Arthropoda" rank="species" species="johorensis">
<emphasis id="87A6930EFF90FFFFFCF5FB37FBECFB62" box="[878,1031,1207,1233]" italics="true" pageId="18" pageNumber="135">R. johorensis</emphasis>
</taxonomicName>
, colonies can be usually found in (but not limited to) bark of fruit trees such as mango, located close to human dwellings. Populations appear widespread, but more sparsely distributed and less commonplace than those of 
<taxonomicName id="72D2349FFF90FFFFFB5EFB71FAB4FAA4" authorityName="Wheeler" authorityYear="1929" box="[1221,1375,1277,1303]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="135" phylum="Arthropoda" rank="species" species="johorensis">
<emphasis id="87A6930EFF90FFFFFB5EFB71FAB4FAA4" box="[1221,1375,1277,1303]" italics="true" pageId="18" pageNumber="135">R. johorensis</emphasis>
</taxonomicName>
– this could merely be an artefact of incomplete sampling.
</paragraph>
</subSubSection>
<caption id="E1AD1F94FF90FFFFFF26F82BFCB3F852" ID-DOI="http://doi.org/10.5281/zenodo.4610577" ID-Zenodo-Dep="4610577" httpUri="https://zenodo.org/record/4610577/files/figure.png" pageId="18" pageNumber="135" startId="18.[189,232,1956,1982]" targetBox="[258,1330,1395,1916]" targetPageId="18">
<paragraph id="B56D4F1CFF90FFFFFF26F82BFCB3F852" blockId="18.[189,1399,1956,2018]" pageId="18" pageNumber="135">
<emphasis id="87A6930EFF90FFFFFF26F82BFEE0F80D" bold="true" box="[189,267,1956,1982]" pageId="18" pageNumber="135">Fig. 7.</emphasis>
<taxonomicName id="72D2349FFF90FFFFFE89F82BFD3BF80D" authority="Wheeler, 1929" authorityName="Wheeler" authorityYear="1929" box="[274,720,1956,1982]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="135" phylum="Arthropoda" rank="species" species="javana">
<emphasis id="87A6930EFF90FFFFFE89F82BFDF1F80D" box="[274,538,1956,1982]" italics="true" pageId="18" pageNumber="135">Rhopalomastix javana</emphasis>
<bibRefCitation id="D14332EDFF90FFFFFDB9F82BFD3BF80D" author="Wheeler W. M." box="[546,720,1956,1982]" pageId="18" pageNumber="135" pagination="95 - 101" refId="ref24237" refString="Wheeler W. M. 1929. The ant genus Rhopalomastix. Psyche 36 (2): 95 - 101." type="journal article" year="1929">Wheeler, 1929</bibRefCitation>
</taxonomicName>
, non-type, ♂, Singapore (ZRC_HYM_0000576). 
<emphasis id="87A6930EFF90FFFFFA80F82BFAC1F80D" bold="true" box="[1307,1322,1956,1982]" pageId="18" pageNumber="135">a</emphasis>
. Head and clypeus in profile. 
<emphasis id="87A6930EFF90FFFFFE52F847FE32F851" bold="true" box="[457,473,1992,2018]" pageId="18" pageNumber="135">b</emphasis>
. Close-up of genitalia in profile.
</paragraph>
</caption>
<subSubSection id="FDC81C97FF91FFFEFF26FE86FAA4FA39" pageId="19" pageNumber="136" type="discussion">
<paragraph id="B56D4F1CFF91FFFEFF26FE86FEC4FE90" blockId="19.[189,303,265,291]" box="[189,303,265,291]" pageId="19" pageNumber="136">
<heading id="EE25F870FF91FFFEFF26FE86FEC4FE90" bold="true" box="[189,303,265,291]" fontSize="11" level="3" pageId="19" pageNumber="136" reason="3">
<emphasis id="87A6930EFF91FFFEFF26FE86FEC4FE90" bold="true" box="[189,303,265,291]" pageId="19" pageNumber="136">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="B56D4F1CFF91FFFEFF26FEB8FDECFDD8" blockId="19.[188,1400,311,619]" pageId="19" pageNumber="136">
Thai populations of 
<taxonomicName id="72D2349FFF91FFFEFE22FEB7FDC5FEE2" authorityName="Wheeler" authorityYear="1929" box="[441,558,312,337]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="136" phylum="Arthropoda" rank="species" species="javana">
<emphasis id="87A6930EFF91FFFEFE22FEB7FDC5FEE2" box="[441,558,312,337]" italics="true" pageId="19" pageNumber="136">R. javana</emphasis>
</taxonomicName>
appear to be generally larger in size compared to populations from 
<collectingCountry id="CDC50F8CFF91FFFEFF26FED5FEC4FEC7" box="[189,303,346,372]" name="Indonesia" pageId="19" pageNumber="136">Indonesia</collectingCountry>
(
<collectingRegion id="771681FEFF91FFFEFEDBFED5FE9CFEC7" box="[320,375,346,372]" country="Indonesia" name="Jawa Timur" pageId="19" pageNumber="136">Java</collectingRegion>
) and 
<collectingCountry id="CDC50F8CFF91FFFEFE27FED5FDD8FEC7" box="[444,563,346,372]" name="Singapore" pageId="19" pageNumber="136">Singapore</collectingCountry>
(Java and 
<collectingCountry id="CDC50F8CFF91FFFEFD2DFED5FCC6FEC7" box="[694,813,346,372]" name="Singapore" pageId="19" pageNumber="136">Singapore</collectingCountry>
TL 1.83–2.02; 
<collectingCountry id="CDC50F8CFF91FFFEFC73FED5FBBBFEC7" box="[1000,1104,346,372]" name="Thailand" pageId="19" pageNumber="136">Thailand</collectingCountry>
TL 2.02–2.88). In terms of variation in morphological measurement metrics, Thai populations also seem much more variable relative to the other two countries (e.g., Thai HL 0.46–0.58; Java HL 0.48–0.52; 
<collectingCountry id="CDC50F8CFF91FFFEFB1EFE2EFB17FE08" box="[1157,1276,417,443]" name="Singapore" pageId="19" pageNumber="136">Singapore</collectingCountry>
HL 0.46– 0.48), however, this might have arisen from bias in geographic range of sampling. For Thai 
<taxonomicName id="72D2349FFF91FFFEFB64FE4AFA84FE6D" authorityName="Wheeler" authorityYear="1929" box="[1279,1391,453,478]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="136" phylum="Arthropoda" rank="species" species="javana">
<emphasis id="87A6930EFF91FFFEFB64FE4AFA84FE6D" box="[1279,1391,453,478]" italics="true" pageId="19" pageNumber="136">R. javana</emphasis>
</taxonomicName>
, measurements were taken from 13 individuals from seven colonies distributed across Central, North and West 
<collectingCountry id="CDC50F8CFF91FFFEFF60FD85FE8DFD97" box="[251,358,522,548]" name="Thailand" pageId="19" pageNumber="136">Thailand</collectingCountry>
, whereas for Java and 
<collectingCountry id="CDC50F8CFF91FFFEFDF1FD85FD0FFD97" box="[618,740,522,548]" name="Singapore" pageId="19" pageNumber="136">Singapore</collectingCountry>
, mostly individuals from 1–2 colonies were measured. A combination of measurements from different countries may be a more accurate representation of actual variation within the species.
</paragraph>
<paragraph id="B56D4F1CFF91FFFEFF26FD1DFB4BFA95" blockId="19.[187,1401,658,1318]" pageId="19" pageNumber="136">
Thai colonies that could be morphologically identified as 
<taxonomicName id="72D2349FFF91FFFEFCEAFD1CFC08FD1F" authorityName="Wheeler" authorityYear="1929" box="[881,995,659,684]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="136" phylum="Arthropoda" rank="species" species="javana">
<emphasis id="87A6930EFF91FFFEFCEAFD1CFC08FD1F" box="[881,995,659,684]" italics="true" pageId="19" pageNumber="136">R. javana</emphasis>
</taxonomicName>
were extremely variable in terms of COI (313 bp), with internidal divergences spread across multiple different values from 0.0–4.5% (
<figureCitation id="2DE95399FF91FFFEFF5DFD57FEE3FD40" box="[198,264,728,755]" captionStart="Fig" captionStartId="4.[189,232,1885,1911]" captionTargetPageId="4" captionText="Fig. 1. Cluster dendrogram of COI (313 bp) barcodes of Rhopalomastix Forel, 1900 from Thailand. Objective clustering of barcodes was performed based on uncorrected p-distances using the ‘best close match’ criteria; numbers at nodes represent percentage divergences or pairwise distance thresholds at which clusters split or lump together." figureDoi="http://doi.org/10.5281/zenodo.4614563" httpUri="https://zenodo.org/record/4614563/files/figure.png" pageId="19" pageNumber="136">Fig. 1</figureCitation>
). While sequence divergence between different species existing in sympatry can be as low as 3.8% (average uncorrected 
<emphasis id="87A6930EFF91FFFEFE24FD73FE25FCA6" box="[447,462,764,789]" italics="true" pageId="19" pageNumber="136">p</emphasis>
distance) in ants, especially for recently diverged species (
<bibRefCitation id="D14332EDFF91FFFEFB11FD73FA82FCA5" author="Wang W. Y. &amp; Srivathsan A. &amp; Foo M. &amp; Yamane S. K. &amp; Meier R." box="[1162,1385,764,790]" pageId="19" pageNumber="136" pagination="490 - 501" refId="ref23961" refString="Wang W. Y., Srivathsan A., Foo M., Yamane S. K. &amp; Meier R. 2018 a. Sorting specimen-rich invertebrate samples with cost-effective NGS barcodes: validating a reverse workflow for specimen processing. Molecular Ecology Resources 18 (3): 490 - 501. https: // doi. org / 10.1111 / 1755 - 0998.12751" type="journal article" year="2018">
Wang 
<emphasis id="87A6930EFF91FFFEFB41FD73FAFDFCA5" box="[1242,1302,764,790]" italics="true" pageId="19" pageNumber="136">et al.</emphasis>
2018a
</bibRefCitation>
), higher percentage distances (&gt; 4%) between allopatric populations do not necessarily indicate different species. In the latter case, clustering based on barcodes may split allopatric or geographically distant populations of the same species, belonging to older lineages that diverged genetically back in time (
<bibRefCitation id="D14332EDFF91FFFEFAAAFCEAFED6FC10" author="Meier R. &amp; Zhang G. &amp; Ali F." pageId="19" pageNumber="136" pagination="809 - 813" refId="ref23791" refString="Meier R., Zhang G. &amp; Ali F. 2008. The use of mean instead of smallest interspecific distances exaggerates the size of the &quot; barcoding gap &quot; and leads to misidentification. Systematic Biology 57 (5): 809 - 813. https: // doi. org / 10.1080 / 10635150802406343" type="journal article" year="2008">
Meier 
<emphasis id="87A6930EFF91FFFEFF20FC06FF1EFC11" box="[187,245,904,930]" italics="true" pageId="19" pageNumber="136">et al.</emphasis>
2008
</bibRefCitation>
). Thus, in light of the two aforementioned possibilities, inferring species boundaries from the clustering pattern of Thai 
<taxonomicName id="72D2349FFF91FFFEFE74FC23FD8BFC76" authorityName="Wheeler" authorityYear="1929" box="[495,608,940,965]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="136" phylum="Arthropoda" rank="species" species="javana">
<emphasis id="87A6930EFF91FFFEFE74FC23FD8BFC76" box="[495,608,940,965]" italics="true" pageId="19" pageNumber="136">R. javana</emphasis>
</taxonomicName>
specimens is complicated. If we define allopatric colonies as those coming from different parts (i.e., North, South, East, West, Central) of 
<collectingCountry id="CDC50F8CFF91FFFEFC65FC40FB82FC5A" box="[1022,1129,975,1001]" name="Thailand" pageId="19" pageNumber="136">Thailand</collectingCountry>
, then possibly the split clusters observed (
<figureCitation id="2DE95399FF91FFFEFE01FC7DFE34FBBF" box="[410,479,1010,1036]" captionStart="Fig" captionStartId="4.[189,232,1885,1911]" captionTargetPageId="4" captionText="Fig. 1. Cluster dendrogram of COI (313 bp) barcodes of Rhopalomastix Forel, 1900 from Thailand. Objective clustering of barcodes was performed based on uncorrected p-distances using the ‘best close match’ criteria; numbers at nodes represent percentage divergences or pairwise distance thresholds at which clusters split or lump together." figureDoi="http://doi.org/10.5281/zenodo.4614563" httpUri="https://zenodo.org/record/4614563/files/figure.png" pageId="19" pageNumber="136">Fig. 1</figureCitation>
) mostly represent different lineages of the same species 
<taxonomicName id="72D2349FFF91FFFEFB1EFC7CFB1EFBBF" authorityName="Wheeler" authorityYear="1929" box="[1157,1269,1011,1036]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="136" phylum="Arthropoda" rank="species" species="javana">
<emphasis id="87A6930EFF91FFFEFB1EFC7CFB1EFBBF" box="[1157,1269,1011,1036]" italics="true" pageId="19" pageNumber="136">R. javana</emphasis>
</taxonomicName>
. However, 
<taxonomicName id="72D2349FFF91FFFEFF25FB99FED9FB9C" authorityName="Wheeler" authorityYear="1929" box="[190,306,1046,1071]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="136" phylum="Arthropoda" rank="species" species="javana">
<emphasis id="87A6930EFF91FFFEFF25FB99FED9FB9C" box="[190,306,1046,1071]" italics="true" pageId="19" pageNumber="136">R. javana</emphasis>
</taxonomicName>
colonies considered ‘sympatric’ from North and Central 
<collectingCountry id="CDC50F8CFF91FFFEFC7DFB9AFBA5FB9C" box="[998,1102,1045,1071]" name="Thailand" pageId="19" pageNumber="136">Thailand</collectingCountry>
respectively diverged at 4.5% (
<figureCitation id="2DE95399FF91FFFEFE92FBB7FEA0FBE0" box="[265,331,1080,1107]" captionStart="Fig" captionStartId="4.[189,232,1885,1911]" captionTargetPageId="4" captionText="Fig. 1. Cluster dendrogram of COI (313 bp) barcodes of Rhopalomastix Forel, 1900 from Thailand. Objective clustering of barcodes was performed based on uncorrected p-distances using the ‘best close match’ criteria; numbers at nodes represent percentage divergences or pairwise distance thresholds at which clusters split or lump together." figureDoi="http://doi.org/10.5281/zenodo.4614563" httpUri="https://zenodo.org/record/4614563/files/figure.png" pageId="19" pageNumber="136">Fig. 1</figureCitation>
), suggesting (but not ascertaining) possible newly diverged species. Taking into account the broad inter- and intranidal variation shown amongst 
<taxonomicName id="72D2349FFF91FFFEFCBDFBD3FC7CFBC6" authorityName="Wheeler" authorityYear="1929" box="[806,919,1116,1141]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="136" phylum="Arthropoda" rank="species" species="javana">
<emphasis id="87A6930EFF91FFFEFCBDFBD3FC7CFBC6" box="[806,919,1116,1141]" italics="true" pageId="19" pageNumber="136">R. javana</emphasis>
</taxonomicName>
workers, we were unable to observe any striking morphological differences between specimens from the genetically divergent clusters. Given the lack of more convincing morphological or molecular evidence supporting further species delimitation in this study, we tentatively treat these colonies as conspecific. The possibility remains for this current treatment to be revised in future, when gene flow between 
<taxonomicName id="72D2349FFF91FFFEFCE2FB66FC00FAB1" authorityName="Wheeler" authorityYear="1929" box="[889,1003,1257,1282]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="136" phylum="Arthropoda" rank="species" species="javana">
<emphasis id="87A6930EFF91FFFEFCE2FB66FC00FAB1" box="[889,1003,1257,1282]" italics="true" pageId="19" pageNumber="136">R. javana</emphasis>
</taxonomicName>
populations are more thoroughly investigated, and the relevant data from other molecular markers are made available.
</paragraph>
<paragraph id="B56D4F1CFF91FFFEFF26FAC2FAA4FA39" blockId="19.[189,1400,1357,1419]" pageId="19" pageNumber="136">
This species is most similar to 
<taxonomicName id="72D2349FFF91FFFEFDA9FAC1FC44FAD4" authority="Wang &amp; Jaitrong" authorityName="Wang &amp; Yong &amp; Jaitrong" authorityYear="2021" box="[562,943,1357,1384]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="136" phylum="Arthropoda" rank="species" species="impithuksai" status="sp. nov.">
<emphasis id="87A6930EFF91FFFEFDA9FAC1FD30FAD4" box="[562,731,1357,1383]" italics="true" pageId="19" pageNumber="136">R. impithuksai</emphasis>
Wang &amp; Jaitrong
</taxonomicName>
<taxonomicNameLabel id="9C952E75FF91FFFEFC2CFAC1FBFFFADB" box="[951,1044,1358,1384]" pageId="19" pageNumber="136" rank="species">sp. nov.</taxonomicNameLabel>
and 
<emphasis id="87A6930EFF91FFFEFBCAFAC1FB19FAD4" box="[1105,1266,1357,1383]" italics="true" pageId="19" pageNumber="136">
<taxonomicName id="72D2349FFF91FFFEFBCAFAC1FB06FAD4" authorityName="Wheeler" authorityYear="1929" box="[1105,1261,1357,1383]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="136" phylum="Arthropoda" rank="species" species="johorensis">R. johorensis</taxonomicName>
,
</emphasis>
but can be differentiated by characters described in detail under the Remarks for each of the latter two species.
</paragraph>
</subSubSection>
</treatment>
</document>