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<mods:title id="16C004EE0BE15ACC1E76E0D10143409C">Revision of the elusive ant genus Rhopalomastix (Hymenoptera, Formicidae, Myrmicinae) in Thailand based on morphology and DNA barcodes, with descriptions of three new species</mods:title>
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<mods:affiliation id="44040FE67C93524B8ECFE8A281985262">Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, Singapore 117377. Department of Biological Sciences, Faculty of Science, National University of Singapore, 16 Science Drive 4, Singapore 117558. Natural History Museum, National Science Museum, Technopolis, Khlong 5, Khlong Luang, Pathum Thani, 12120 Thailand.</mods:affiliation>
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Genus
<taxonomicName id="72D2349FFF81FFEEFD3BF923FC0EF974" ID-CoL="79CY" authority="Forel, 1900" authorityName="Forel" authorityYear="1900" box="[672,997,1708,1735]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="120" phylum="Arthropoda" rank="genus">
<emphasis id="87A6930EFF81FFEEFD3BF923FCBCF974" bold="true" box="[672,855,1708,1735]" italics="true" pageId="3" pageNumber="120">Rhopalomastix</emphasis>
<bibRefCitation id="D14332EDFF81FFEEFCC5F922FC0EF974" author="Forel A." box="[862,997,1709,1735]" pageId="3" pageNumber="120" pagination="24 - 26" refId="ref23556" refString="Forel A. 1900. Un nouveau genre et une nouvelle espece de Myrmicide. Annales de la Societe entomologique de Belgique 44: 24 - 26." type="journal article" year="1900">Forel, 1900</bibRefCitation>
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<treatmentCitation id="3473690DFF81FFEEFF27F961FDC0F8BB" author="Forel A." box="[188,555,1774,1800]" page="24" pageId="3" pageNumber="120" year="1900">
<taxonomicName id="72D2349FFF81FFEEFF27F961FDC0F8BB" ID-CoL="79CY" authority="Forel, 1900: 24" authorityName="Forel" authorityPageNumber="24" authorityYear="1900" box="[188,555,1774,1800]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="120" phylum="Arthropoda" rank="genus">
<emphasis id="87A6930EFF81FFEEFF27F961FE87F8BB" box="[188,364,1774,1800]" italics="true" pageId="3" pageNumber="120">Rhopalomastix</emphasis>
<bibRefCitation id="D14332EDFF81FFEEFEEFF961FDC0F8BB" author="Forel A." box="[372,555,1774,1800]" pageId="3" pageNumber="120" pagination="24 - 26" refId="ref23556" refString="Forel A. 1900. Un nouveau genre et une nouvelle espece de Myrmicide. Annales de la Societe entomologique de Belgique 44: 24 - 26." type="journal article" year="1900">Forel, 1900: 24</bibRefCitation>
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.
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<subSubSection id="FDC81C97FF81FFEEFDADF960FAD8F8BA" box="[566,1331,1774,1801]" pageId="3" pageNumber="120" type="type_taxon">
<paragraph id="B56D4F1CFF81FFEEFDADF960FAD8F8BA" blockId="3.[188,1331,1774,1801]" box="[566,1331,1774,1801]" pageId="3" pageNumber="120">
<typeStatus id="6A69F1BEFF81FFEEFDADF960FD9BF8BA" box="[566,624,1775,1801]" pageId="3" pageNumber="120">Type</typeStatus>
species:
<taxonomicName id="72D2349FFF81FFEEFD47F961FB6EF8BB" ID-CoL="4SKR2" authority="Forel, 1900" authorityName="Forel" authorityYear="1900" box="[732,1157,1774,1800]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="120" phylum="Arthropoda" rank="species" species="rothneyi">
<emphasis id="87A6930EFF81FFEEFD47F961FC1FF8BB" box="[732,1012,1774,1800]" italics="true" pageId="3" pageNumber="120">Rhopalomastix rothneyi</emphasis>
<bibRefCitation id="D14332EDFF81FFEEFC60F961FB6EF8BB" author="Forel A." box="[1019,1157,1774,1800]" pageId="3" pageNumber="120" pagination="24 - 26" refId="ref23556" refString="Forel A. 1900. Un nouveau genre et une nouvelle espece de Myrmicide. Annales de la Societe entomologique de Belgique 44: 24 - 26." type="journal article" year="1900">Forel, 1900</bibRefCitation>
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, by monotypy.
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<treatmentCitationGroup id="95C26832FF81FFEEFF27F8A0FC3DF8F9" box="[188,982,1839,1866]" pageId="3" pageNumber="120">
<taxonomicName id="72D2349FFF81FFEEFF27F8A0FE87F8FA" ID-CoL="79CY" authorityName="Forel" authorityYear="1900" box="[188,364,1839,1865]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="120" phylum="Arthropoda" rank="genus">
<emphasis id="87A6930EFF81FFEEFF27F8A0FE87F8FA" box="[188,364,1839,1865]" italics="true" pageId="3" pageNumber="120">Rhopalomastix</emphasis>
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in
<taxonomicName id="72D2349FFF81FFEEFE09F8A0FDCFF8FA" ID-CoL="JY2" authorityName="Lepeletier de Saint-Fargeau" authorityYear="1835" box="[402,548,1839,1865]" class="Insecta" family="Formicidae" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="120" phylum="Arthropoda" rank="subFamily" subFamily="Myrmicinae">Myrmicinae</taxonomicName>
,
<taxonomicName id="72D2349FFF81FFEEFDABF8A0FD3AF8FA" ID-CoL="L8M" authorityName="Lepeletier de Saint-Fargeau" authorityYear="1835" box="[560,721,1839,1865]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="120" phylum="Arthropoda" rank="tribe" tribe="Solenopsidini">Solenopsidini</taxonomicName>
–
<treatmentCitation id="3473690DFF81FFEEFD75F8A0FC39F8F9" author="Wheeler W. M." box="[750,978,1839,1866]" page="140" pageId="3" pageNumber="120" year="1910">
<bibRefCitation id="D14332EDFF81FFEEFD75F8A0FC39F8F9" author="Wheeler W. M." box="[750,978,1839,1866]" pageId="3" pageNumber="120" refId="ref24166" refString="Wheeler W. M. 1910. Ants: their Structure, Development and Behavior. Columbia University Press, New York." type="book" year="1910">Wheeler 1910: 140</bibRefCitation>
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.
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<treatmentCitationGroup id="95C26832FF81FFEEFF27F8DCFD2FF823" pageId="3" pageNumber="120">
<taxonomicName id="72D2349FFF81FFEEFF27F8DCFE87F8DE" ID-CoL="79CY" authorityName="Forel" authorityYear="1900" box="[188,364,1875,1901]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="120" phylum="Arthropoda" rank="genus">
<emphasis id="87A6930EFF81FFEEFF27F8DCFE87F8DE" box="[188,364,1875,1901]" italics="true" pageId="3" pageNumber="120">Rhopalomastix</emphasis>
</taxonomicName>
in
<taxonomicName id="72D2349FFF81FFEEFE0CF8DCFDC2F8DE" ID-CoL="JY2" authorityName="Lepeletier de Saint-Fargeau" authorityYear="1835" box="[407,553,1875,1901]" class="Insecta" family="Formicidae" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="120" phylum="Arthropoda" rank="subFamily" subFamily="Myrmicinae">Myrmicinae</taxonomicName>
,
<taxonomicName id="72D2349FFF81FFEEFDA3F8DCFD35F8DE" ID-CoL="KZ7" authorityName="Lepeletier de Saint-Fargeau" authorityYear="1835" box="[568,734,1875,1901]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="120" phylum="Arthropoda" rank="tribe" tribe="Melissotarsini">Melissotarsini</taxonomicName>
–
<treatmentCitation id="3473690DFF81FFEEFC9AF8DCFC2CF8DE" author="Emery C." box="[769,967,1875,1901]" page="40" pageId="3" pageNumber="120" year="1914">
<bibRefCitation id="D14332EDFF81FFEEFC9AF8DCFC2CF8DE" author="Emery C." box="[769,967,1875,1901]" pageId="3" pageNumber="120" pagination="29 - 42" refId="ref23474" refString="Emery C. 1914. Intorno alla classificazione dei Myrmicinae. Rendiconti delle Sessioni della Reale Accademia delle Scienze dell'Istituto di Bologna. Classe di Scienze Fisiche (n. s.) 18: 29 - 42." type="journal article" year="1914">Emery 1914: 40</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="3473690DFF81FFEEFC4DF8DCFBBAF8DE" author="Emery C." box="[982,1105,1875,1901]" page="118" pageId="3" pageNumber="120" year="1922">
<bibRefCitation id="D14332EDFF81FFEEFC4DF8DCFBBAF8DE" author="Emery C." box="[982,1105,1875,1901]" pageId="3" pageNumber="120" pagination="95 - 206" refId="ref23513" refString="Emery C. 1922. Hymenoptera. Fam. Formicidae. Subfam. Myrmicinae. [part]. In: Wytsmann P. (Dir.) Genera Insectorum 174 B: 95 - 206. L. Desmet-Verteneuil, Brussels." type="book chapter" year="1922">1922: 118</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="3473690DFF81FFEEFB1DF8DCFAA0F8DE" author="Forel A." box="[1158,1355,1875,1901]" page="242" pageId="3" pageNumber="120" year="1917">
<bibRefCitation id="D14332EDFF81FFEEFB1DF8DCFAA0F8DE" author="Forel A." box="[1158,1355,1875,1901]" pageId="3" pageNumber="120" pagination="229 - 253" refId="ref23584" refString="Forel A. 1917. Cadre synoptique actuel de la faune universelle des fourmis. Bulletin de la Societe Vaudoise des Sciences naturelles 51: 229 - 253. https: // doi. org / 10.5281 / zenodo. 25600" type="journal article" year="1917">Forel 1917: 242</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="3473690DFF81FFEEFF71F8F9FE25F823" author="Wheeler W. M." box="[234,462,1910,1936]" page="661" pageId="3" pageNumber="120" year="1922">
<bibRefCitation id="D14332EDFF81FFEEFF71F8F9FE25F823" author="Wheeler W. M." box="[234,462,1910,1936]" pageId="3" pageNumber="120" pagination="631 - 710" refId="ref24189" refString="Wheeler W. M. 1922. Ants of the American Museum Congo Expedition. A contribution to the myrmecology of Africa. VII. Keys to the genera and subgenera of ants. Bulletin of the American Museum of Natural History 45: 631 - 710." type="journal article" year="1922">Wheeler 1922: 661</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="3473690DFF81FFEEFE65F8F9FD2BF823" author="Bolton B." box="[510,704,1910,1936]" page="72" pageId="3" pageNumber="120" year="2003">
<bibRefCitation id="D14332EDFF81FFEEFE65F8F9FD2BF823" author="Bolton B." box="[510,704,1910,1936]" pageId="3" pageNumber="120" pagination="1 - 370" refId="ref23366" refString="Bolton B. 2003. Synopsis and classification of Formicidae. Memoirs of the American Entomological Institute 71: 1 - 370." type="journal article" year="2003">Bolton 2003: 72</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="B56D4F1CFF81FFEEFF27F816FBC6F807" blockId="3.[188,1398,1839,1972]" box="[188,1069,1945,1972]" pageId="3" pageNumber="120">
<treatmentCitationGroup id="95C26832FF81FFEEFF27F816FBC6F807" box="[188,1069,1945,1972]" pageId="3" pageNumber="120">
<taxonomicName id="72D2349FFF81FFEEFF27F816FE87F800" ID-CoL="79CY" authorityName="Forel" authorityYear="1900" box="[188,364,1945,1971]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="120" phylum="Arthropoda" rank="genus">
<emphasis id="87A6930EFF81FFEEFF27F816FE87F800" box="[188,364,1945,1971]" italics="true" pageId="3" pageNumber="120">Rhopalomastix</emphasis>
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in
<taxonomicName id="72D2349FFF81FFEEFE09F816FDCFF800" ID-CoL="JY2" authorityName="Lepeletier de Saint-Fargeau" authorityYear="1835" box="[402,548,1945,1971]" class="Insecta" family="Formicidae" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="120" phylum="Arthropoda" rank="subFamily" subFamily="Myrmicinae">Myrmicinae</taxonomicName>
,
<taxonomicName id="72D2349FFF81FFEEFDABF816FD06F800" ID-CoL="KR4" authorityName="Lepeletier de Saint-Fargeau" authorityYear="1835" box="[560,749,1945,1971]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="120" phylum="Arthropoda" rank="tribe" tribe="Crematogastrini">Crematogastrini</taxonomicName>
–
<treatmentCitation id="3473690DFF81FFEEFC91F816FC17F800" author="Ward P. S. &amp; Brady S. G. &amp; Fisher B. L. &amp; Schultz T. R." box="[778,1020,1945,1972]" page="71" pageId="3" pageNumber="120" year="2015">
<bibRefCitation id="D14332EDFF81FFEEFC91F816FC17F800" author="Ward P. S. &amp; Brady S. G. &amp; Fisher B. L. &amp; Schultz T. R." box="[778,1020,1945,1972]" pageId="3" pageNumber="120" pagination="61 - 81" refId="ref24098" refString="Ward P. S., Brady S. G., Fisher B. L. &amp; Schultz T. R. 2015. The evolution of myrmicine ants: phylogeny and biogeography of a hyperdiverse ant clade (Hymenoptera: Formicidae). Systematic Entomology 40 (1): 61 - 81. https: // doi. org / 10.1111 / syen. 12090" type="journal article" year="2015">
Ward
<emphasis id="87A6930EFF81FFEEFCCBF815FC60F800" box="[848,907,1945,1971]" italics="true" pageId="3" pageNumber="120">et al.</emphasis>
2015: 71
</bibRefCitation>
</treatmentCitation>
, 77.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="E1AD1F94FF86FFE9FF26F8D2FD98F853" ID-DOI="http://doi.org/10.5281/zenodo.4614563" ID-Zenodo-Dep="4614563" httpUri="https://zenodo.org/record/4614563/files/figure.png" pageId="4" pageNumber="121" targetBox="[532,997,139,1022]" targetPageId="4">
<paragraph id="B56D4F1CFF86FFE9FF26F8D2FD98F853" blockId="4.[189,1399,1885,2017]" pageId="4" pageNumber="121">
<emphasis id="87A6930EFF86FFE9FF26F8D2FEE5F8C4" bold="true" box="[189,270,1885,1911]" pageId="4" pageNumber="121">Fig. 1.</emphasis>
Cluster dendrogram of COI (313 bp) barcodes of
<taxonomicName id="72D2349FFF86FFE9FCEEF8D2FB51F8C4" authority="Forel, 1900" authorityName="Forel" authorityYear="1900" box="[885,1210,1885,1911]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="121" phylum="Arthropoda" rank="genus">
<emphasis id="87A6930EFF86FFE9FCEEF8D2FBCEF8C4" box="[885,1061,1885,1911]" italics="true" pageId="4" pageNumber="121">Rhopalomastix</emphasis>
<bibRefCitation id="D14332EDFF86FFE9FBB4F8D2FB51F8C4" author="Forel A." box="[1071,1210,1885,1911]" pageId="4" pageNumber="121" pagination="24 - 26" refId="ref23556" refString="Forel A. 1900. Un nouveau genre et une nouvelle espece de Myrmicide. Annales de la Societe entomologique de Belgique 44: 24 - 26." type="journal article" year="1900">Forel, 1900</bibRefCitation>
</taxonomicName>
from Thailand. Objective clustering of barcodes was performed based on uncorrected
<emphasis id="87A6930EFF86FFE9FC60F80EFBE2F829" box="[1019,1033,1921,1946]" italics="true" pageId="4" pageNumber="121">p</emphasis>
-distances using the ‘best close match’ criteria; numbers at nodes represent percentage divergences or pairwise distance thresholds at which clusters split or lump together.
</paragraph>
</caption>
<subSubSection id="FDC81C97FF87FFE8FF26FE85FB99FEF4" pageId="5" pageNumber="122" type="diagnosis">
<paragraph id="B56D4F1CFF87FFE8FF26FE85FB99FEF4" blockId="5.[189,1397,265,327]" pageId="5" pageNumber="122">
For diagnosis of the workers in the genus
<taxonomicName id="72D2349FFF87FFE8FD01FE86FCA1FE90" authorityName="Forel" authorityYear="1900" box="[666,842,265,291]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="87A6930EFF87FFE8FD01FE86FCA1FE90" box="[666,842,265,291]" italics="true" pageId="5" pageNumber="122">Rhopalomastix</emphasis>
</taxonomicName>
, refer to
<bibRefCitation id="D14332EDFF87FFE8FC29FE86FBBEFE90" author="Bolton B." box="[946,1109,265,291]" pageId="5" pageNumber="122" pagination="1 - 370" refId="ref23366" refString="Bolton B. 2003. Synopsis and classification of Formicidae. Memoirs of the American Entomological Institute 71: 1 - 370." type="journal article" year="2003">Bolton (2003)</bibRefCitation>
and
<bibRefCitation id="D14332EDFF87FFE8FB11FE85FA9AFE97" author="Wang W. Y. &amp; Yong G. W. &amp; Jaitrong W." box="[1162,1393,265,292]" pageId="5" pageNumber="122" pagination="301 - 340" refId="ref24028" refString="Wang W. Y., Yong G. W. &amp; Jaitrong W. 2018 b. The ant genus Rhopalomastix (Hymenoptera: Formicidae: Myrmicinae) in Southeast Asia, with descriptions of four new species from Singapore based on morphology and DNA barcoding. Zootaxa 4532 (3): 301 - 340. https: // doi. org / 10.11646 / zootaxa. 4532.3.1" type="journal article" year="2018">
Wang
<emphasis id="87A6930EFF87FFE8FB49FE85FAE0FE90" box="[1234,1291,265,291]" italics="true" pageId="5" pageNumber="122">et al.</emphasis>
(2018b)
</bibRefCitation>
. For diagnoses of the queen and male of the same genus, see
<bibRefCitation id="D14332EDFF87FFE8FC1BFEA2FB85FEF4" author="Wang W. Y. &amp; Yong G. W. &amp; Jaitrong W." box="[896,1134,300,327]" pageId="5" pageNumber="122" pagination="301 - 340" refId="ref24028" refString="Wang W. Y., Yong G. W. &amp; Jaitrong W. 2018 b. The ant genus Rhopalomastix (Hymenoptera: Formicidae: Myrmicinae) in Southeast Asia, with descriptions of four new species from Singapore based on morphology and DNA barcoding. Zootaxa 4532 (3): 301 - 340. https: // doi. org / 10.11646 / zootaxa. 4532.3.1" type="journal article" year="2018">
Wang
<emphasis id="87A6930EFF87FFE8FC50FEA2FBEDFEF5" box="[971,1030,300,326]" italics="true" pageId="5" pageNumber="122">et al.</emphasis>
(2018b)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="FDC81C97FF87FFE8FF26FEE2FD16FDD4" pageId="5" pageNumber="122" type="synonymic_list">
<paragraph id="B56D4F1CFF87FFE8FF26FEE2FD86FE34" blockId="5.[189,621,365,391]" box="[189,621,365,391]" pageId="5" pageNumber="122">
<heading id="EE25F870FF87FFE8FF26FEE2FD86FE34" bold="true" box="[189,621,365,391]" fontSize="11" level="3" pageId="5" pageNumber="122" reason="3">
<emphasis id="87A6930EFF87FFE8FF26FEE2FD86FE34" bold="true" box="[189,621,365,391]" pageId="5" pageNumber="122">
Synoptic list of species in
<collectingCountry id="CDC50F8CFF87FFE8FE60FEE2FD86FE34" box="[507,621,365,391]" name="Thailand" pageId="5" pageNumber="122">Thailand</collectingCountry>
</emphasis>
</heading>
</paragraph>
<paragraph id="B56D4F1CFF87FFE8FF26FE13FCC5FE04" blockId="5.[189,814,412,615]" box="[189,814,412,439]" pageId="5" pageNumber="122">
<taxonomicName id="72D2349FFF87FFE8FF26FE13FD26FE05" authority="Wang &amp; Jaitrong" authorityName="Wang &amp; Yong &amp; Jaitrong" authorityYear="2021" box="[189,717,412,439]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="122" phylum="Arthropoda" rank="species" species="impithuksai" status="sp. nov.">
<emphasis id="87A6930EFF87FFE8FF26FE13FE16FE05" box="[189,509,412,438]" italics="true" pageId="5" pageNumber="122">Rhopalomastix impithuksai</emphasis>
Wang &amp; Jaitrong
</taxonomicName>
<taxonomicNameLabel id="9C952E75FF87FFE8FD4FFE12FCC5FE04" box="[724,814,413,439]" pageId="5" pageNumber="122" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="B56D4F1CFF87FFE8FF26FE30FD9CFE6A" blockId="5.[189,814,412,615]" box="[189,631,447,473]" pageId="5" pageNumber="122">
<taxonomicName id="72D2349FFF87FFE8FF26FE30FD9CFE6A" authority="Wheeler, 1929" authorityName="Wheeler" authorityYear="1929" box="[189,631,447,473]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="122" phylum="Arthropoda" rank="species" species="javana">
<emphasis id="87A6930EFF87FFE8FF26FE30FE2FFE6A" box="[189,452,447,473]" italics="true" pageId="5" pageNumber="122">Rhopalomastix javana</emphasis>
<bibRefCitation id="D14332EDFF87FFE8FE50FE30FD9CFE6A" author="Wheeler W. M." box="[459,631,447,473]" pageId="5" pageNumber="122" pagination="95 - 101" refId="ref24237" refString="Wheeler W. M. 1929. The ant genus Rhopalomastix. Psyche 36 (2): 95 - 101." type="journal article" year="1929">Wheeler, 1929</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="B56D4F1CFF87FFE8FF26FE6DFD4BFE4E" blockId="5.[189,814,412,615]" box="[189,672,482,509]" pageId="5" pageNumber="122">
<taxonomicName id="72D2349FFF87FFE8FF26FE6DFD4BFE4E" authority="Wheeler, 1929" authorityName="Wheeler" authorityYear="1929" box="[189,672,482,509]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="122" phylum="Arthropoda" rank="species" species="johorensis">
<emphasis id="87A6930EFF87FFE8FF26FE6DFE06FE4F" box="[189,493,482,508]" italics="true" pageId="5" pageNumber="122">Rhopalomastix johorensis</emphasis>
<bibRefCitation id="D14332EDFF87FFE8FE6FFE6DFD4BFE4E" author="Wheeler W. M." box="[500,672,482,509]" pageId="5" pageNumber="122" pagination="95 - 101" refId="ref24237" refString="Wheeler W. M. 1929. The ant genus Rhopalomastix. Psyche 36 (2): 95 - 101." type="journal article" year="1929">Wheeler, 1929</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="B56D4F1CFF87FFE8FF78FD89FD3CFD93" blockId="5.[189,814,412,615]" box="[227,727,518,544]" pageId="5" pageNumber="122">
=
<taxonomicName id="72D2349FFF87FFE8FF60FD89FD3CFD93" authority="Donisthorpe, 1936" authorityName="Donisthorpe" authorityYear="1936" box="[251,727,518,544]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="122" phylum="Arthropoda" rank="species" species="janeti">
<emphasis id="87A6930EFF87FFE8FF60FD89FE1EFD93" box="[251,501,518,544]" italics="true" pageId="5" pageNumber="122">Rhopalomastix janeti</emphasis>
<bibRefCitation id="D14332EDFF87FFE8FE67FD89FD3CFD93" author="Donisthorpe H." box="[508,727,518,544]" pageId="5" pageNumber="122" pagination="55 - 56" refId="ref23430" refString="Donisthorpe H. 1936. Rhopalomastix janeti (Hym. Formicidae) a species of ant new to science. Entomologist's Record and Journal of Variation 48: 55 - 56. https: // doi. org / 10.5281 / zenodo. 26445" type="journal article" year="1936">Donisthorpe, 1936</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="B56D4F1CFF87FFE8FF26FDA6FD02FDF0" blockId="5.[189,814,412,615]" box="[189,745,553,579]" pageId="5" pageNumber="122">
<taxonomicName id="72D2349FFF87FFE8FF26FDA6FD63FDF0" authority="Wang &amp; Jaitrong" authorityName="Wang &amp; Yong &amp; Jaitrong" authorityYear="2021" box="[189,648,553,579]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="122" phylum="Arthropoda" rank="species" species="parva" status="sp. nov.">
<emphasis id="87A6930EFF87FFE8FF26FDA6FE53FDF0" box="[189,440,553,579]" italics="true" pageId="5" pageNumber="122">Rhopalomastix parva</emphasis>
Wang &amp; Jaitrong
</taxonomicName>
<taxonomicNameLabel id="9C952E75FF87FFE8FD14FDA6FD02FDF0" box="[655,745,553,579]" pageId="5" pageNumber="122" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="B56D4F1CFF87FFE8FF26FDC3FD16FDD4" blockId="5.[189,814,412,615]" box="[189,765,588,615]" pageId="5" pageNumber="122">
<taxonomicName id="72D2349FFF87FFE8FF26FDC3FD77FDD5" authority="Wang &amp; Jaitrong" authorityName="Wang &amp; Yong &amp; Jaitrong" authorityYear="2021" box="[189,668,588,615]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="122" phylum="Arthropoda" rank="species" species="robusta" status="sp. nov.">
<emphasis id="87A6930EFF87FFE8FF26FDC3FE26FDD5" box="[189,461,588,614]" italics="true" pageId="5" pageNumber="122">Rhopalomastix robusta</emphasis>
Wang &amp; Jaitrong
</taxonomicName>
<taxonomicNameLabel id="9C952E75FF87FFE8FD38FDC2FD16FDD4" box="[675,765,589,615]" pageId="5" pageNumber="122" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="FDC81C97FF87FFE8FF26FD01FD11FD1B" box="[189,762,653,680]" pageId="5" pageNumber="122" type="discussion">
<paragraph id="B56D4F1CFF87FFE8FF26FD01FD11FD1B" blockId="5.[189,762,653,680]" box="[189,762,653,680]" pageId="5" pageNumber="122">
<heading id="EE25F870FF87FFE8FF26FD01FD11FD1B" bold="true" box="[189,762,653,680]" fontSize="11" level="3" pageId="5" pageNumber="122" reason="3">
<emphasis id="87A6930EFF87FFE8FF26FD01FD11FD1B" bold="true" box="[189,762,653,680]" pageId="5" pageNumber="122">Key to Thai species based on the worker caste</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FDC81C97FF87FFEBFF26FD33FA9CFBE1" lastPageId="6" lastPageNumber="123" pageId="5" pageNumber="122" type="key">
<key id="C143182DFF87FFEBFF26FD33FA9CFBE1" lastPageId="6" lastPageNumber="123" pageId="5" pageNumber="122">
<keyStep id="0E2657B9FF87FFE8FF26FD33FA9CFC35" pageId="5" pageNumber="122">
<paragraph id="B56D4F1CFF87FFE8FF26FD33FA9CFCAE" blockId="5.[189,1400,700,903]" pageId="5" pageNumber="122">
<keyLead id="0E23EC29FF87FFE8FF26FD33FA9CFCAE" pageId="5" pageNumber="122">
1. External face of mandible other than narrow strip next to masticatory margin with distinct striations (
<figureCitation id="2DE95399FF87FFE8FF68FD50FE8CFD49" box="[243,359,735,762]" captionStart="Fig" captionStartId="5.[189,232,1816,1842]" captionTargetBox="[227,1361,973,1778]" captionTargetId="figure-170@5.[227,1361,973,1778]" captionTargetPageId="5" captionText="Fig. 2. Closeups of external mandibular surface for workers of 5 Thai species of Rhopalomastix Forel, 1900 (ZRC); white arrows indicate distinct striations on the external mandibular face. a. R. impithuksai Wang &amp; Jaitrong sp. nov., paratype (ZRC_ENT00007608). b. R. robusta Wang &amp; Jaitrong sp. nov., paratype (ZRC_ENT00013949). c. R. javana Wheeler, 1929, non-type (ZRC_ENT00000932). d. R. johorensis Wheeler, 1929, non-type (ZRC_ENT00007874). e. R. parva Wang &amp; Jaitrong sp. nov., paratype (ZRC_ENT00013948)." figureDoi="http://doi.org/10.5281/zenodo.4610567" httpUri="https://zenodo.org/record/4610567/files/figure.png" pageId="5" pageNumber="122">Fig. 2a–b</figureCitation>
); extent and intensity of striations variable. Median clypeal area carinulate, supraclypeal area with dark-pigmented longitudinal carinae ................................................................................ 2
</keyLead>
</paragraph>
<paragraph id="B56D4F1CFF87FFE8FF5AFCA9FA9CFC35" blockId="5.[189,1400,700,903]" pageId="5" pageNumber="122">
<keyLead id="0E23EC29FF87FFE8FF5AFCA9FA9CFC35" pageId="5" pageNumber="122">
– External face of mandible other than narrow strip next to masticatory margin largely smooth and shiny (
<figureCitation id="2DE95399FF87FFE8FEA5FCC6FE58FCD0" box="[318,435,841,867]" captionStart="Fig" captionStartId="5.[189,232,1816,1842]" captionTargetBox="[227,1361,973,1778]" captionTargetId="figure-170@5.[227,1361,973,1778]" captionTargetPageId="5" captionText="Fig. 2. Closeups of external mandibular surface for workers of 5 Thai species of Rhopalomastix Forel, 1900 (ZRC); white arrows indicate distinct striations on the external mandibular face. a. R. impithuksai Wang &amp; Jaitrong sp. nov., paratype (ZRC_ENT00007608). b. R. robusta Wang &amp; Jaitrong sp. nov., paratype (ZRC_ENT00013949). c. R. javana Wheeler, 1929, non-type (ZRC_ENT00000932). d. R. johorensis Wheeler, 1929, non-type (ZRC_ENT00007874). e. R. parva Wang &amp; Jaitrong sp. nov., paratype (ZRC_ENT00013948)." figureDoi="http://doi.org/10.5281/zenodo.4610567" httpUri="https://zenodo.org/record/4610567/files/figure.png" pageId="5" pageNumber="122">Fig. 2c–e</figureCitation>
). Median clypeal area and supraclypeal area with varying forms of sculpture but never with multiple dark-pigmented longitudinal carinae ................................................................ 3
</keyLead>
</paragraph>
</keyStep>
<caption id="E1AD1F94FF87FFE8FF26F897FDD0F850" ID-DOI="http://doi.org/10.5281/zenodo.4610567" ID-Zenodo-Dep="4610567" httpUri="https://zenodo.org/record/4610567/files/figure.png" pageId="5" pageNumber="122" startId="5.[189,232,1816,1842]" targetBox="[227,1361,973,1778]" targetPageId="5">
<paragraph id="B56D4F1CFF87FFE8FF26F897FDD0F850" blockId="5.[189,1399,1816,2019]" pageId="5" pageNumber="122">
<emphasis id="87A6930EFF87FFE8FF26F897FEE0F881" bold="true" box="[189,267,1816,1842]" pageId="5" pageNumber="122">Fig. 2.</emphasis>
Closeups of external mandibular surface for workers of 5 Thai species of
<taxonomicName id="72D2349FFF87FFE8FBE1F897FF12F8E5" ID-CoL="79CY" authority="Forel, 1900" authorityName="Forel" authorityYear="1900" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="87A6930EFF87FFE8FBE1F897FAC1F881" box="[1146,1322,1816,1842]" italics="true" pageId="5" pageNumber="122">Rhopalomastix</emphasis>
<bibRefCitation id="D14332EDFF87FFE8FAAAF897FF12F8E5" author="Forel A." pageId="5" pageNumber="122" pagination="24 - 26" refId="ref23556" refString="Forel A. 1900. Un nouveau genre et une nouvelle espece de Myrmicide. Annales de la Societe entomologique de Belgique 44: 24 - 26." type="journal article" year="1900">Forel, 1900</bibRefCitation>
</taxonomicName>
(ZRC); white arrows indicate distinct striations on the external mandibular face.
<emphasis id="87A6930EFF87FFE8FB2BF8B3FB54F8E5" bold="true" box="[1200,1215,1852,1878]" pageId="5" pageNumber="122">a</emphasis>
.
<taxonomicName id="72D2349FFF87FFE8FB56F8B3FE67F8CA" authority="Wang &amp; Jaitrong" authorityName="Wang &amp; Yong &amp; Jaitrong" authorityYear="2021" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="122" phylum="Arthropoda" rank="species" species="impithuksai" status="sp. nov.">
<emphasis id="87A6930EFF87FFE8FB56F8B3FA9CF8E5" box="[1229,1399,1852,1878]" italics="true" pageId="5" pageNumber="122">R. impithuksai</emphasis>
Wang &amp; Jaitrong
</taxonomicName>
<taxonomicNameLabel id="9C952E75FF87FFE8FE0DF8D0FE1DF8CA" box="[406,502,1887,1913]" pageId="5" pageNumber="122" rank="species">sp. nov.</taxonomicNameLabel>
, paratype (ZRC_ENT00007608).
<emphasis id="87A6930EFF87FFE8FC0BF8D0FC4BF8CA" bold="true" box="[912,928,1887,1913]" pageId="5" pageNumber="122">b</emphasis>
.
<taxonomicName id="72D2349FFF87FFE8FC29F8EFFAE3F8CA" authority="Wang &amp; Jaitrong" authorityName="Wang &amp; Yong &amp; Jaitrong" authorityYear="2021" box="[946,1288,1887,1913]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="122" phylum="Arthropoda" rank="species" species="robusta" status="sp. nov.">
<emphasis id="87A6930EFF87FFE8FC29F8EFFBC4F8CA" box="[946,1071,1887,1913]" italics="true" pageId="5" pageNumber="122">R. robusta</emphasis>
Wang &amp; Jaitrong
</taxonomicName>
<taxonomicNameLabel id="9C952E75FF87FFE8FA89F8D0FA99F8CA" box="[1298,1394,1887,1913]" pageId="5" pageNumber="122" rank="species">sp. nov.</taxonomicNameLabel>
, paratype (ZRC_ENT00013949).
<emphasis id="87A6930EFF87FFE8FDC0F80DFD83F82F" bold="true" box="[603,616,1922,1948]" pageId="5" pageNumber="122">c</emphasis>
.
<taxonomicName id="72D2349FFF87FFE8FD1FF80CFC3BF82F" authority="Wheeler, 1929" authorityName="Wheeler" authorityYear="1929" box="[644,976,1922,1948]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="122" phylum="Arthropoda" rank="species" species="javana">
<emphasis id="87A6930EFF87FFE8FD1FF80CFCEAF82F" box="[644,769,1923,1948]" italics="true" pageId="5" pageNumber="122">R. javana</emphasis>
<bibRefCitation id="D14332EDFF87FFE8FC8EF80DFC3BF82F" author="Wheeler W. M." box="[789,976,1922,1948]" pageId="5" pageNumber="122" pagination="95 - 101" refId="ref24237" refString="Wheeler W. M. 1929. The ant genus Rhopalomastix. Psyche 36 (2): 95 - 101." type="journal article" year="1929">Wheeler, 1929</bibRefCitation>
</taxonomicName>
, non-type (ZRC_ENT00000932).
<emphasis id="87A6930EFF87FFE8FF26F82AFF26F80C" bold="true" box="[189,205,1957,1983]" pageId="5" pageNumber="122">d</emphasis>
.
<taxonomicName id="72D2349FFF87FFE8FF47F829FDC0F80C" authority="Wheeler, 1929" authorityName="Wheeler" authorityYear="1929" box="[220,555,1957,1983]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="122" phylum="Arthropoda" rank="species" species="johorensis">
<emphasis id="87A6930EFF87FFE8FF47F829FE9EF80C" box="[220,373,1957,1983]" italics="true" pageId="5" pageNumber="122">R. johorensis</emphasis>
<bibRefCitation id="D14332EDFF87FFE8FEE6F82AFDC0F80C" author="Wheeler W. M." box="[381,555,1957,1983]" pageId="5" pageNumber="122" pagination="95 - 101" refId="ref24237" refString="Wheeler W. M. 1929. The ant genus Rhopalomastix. Psyche 36 (2): 95 - 101." type="journal article" year="1929">Wheeler, 1929</bibRefCitation>
</taxonomicName>
, non-type (ZRC_ENT00007874).
<emphasis id="87A6930EFF87FFE8FC24F82AFC38F80C" bold="true" box="[959,979,1957,1983]" pageId="5" pageNumber="122">e.</emphasis>
<taxonomicName id="72D2349FFF87FFE8FC41F829FAE5F80C" authority="Wang &amp; Jaitrong" authorityName="Wang &amp; Yong &amp; Jaitrong" authorityYear="2021" box="[986,1294,1957,1984]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="122" phylum="Arthropoda" rank="species" species="parva" status="sp. nov.">
<emphasis id="87A6930EFF87FFE8FC41F829FBD4F80C" box="[986,1087,1958,1983]" italics="true" pageId="5" pageNumber="122">R. parva</emphasis>
Wang &amp; Jaitrong
</taxonomicName>
<taxonomicNameLabel id="9C952E75FF87FFE8FA8EF829FA99F873" box="[1301,1394,1958,1984]" pageId="5" pageNumber="122" rank="species">sp. nov.</taxonomicNameLabel>
, paratype (ZRC_ENT00013948).
</paragraph>
</caption>
<keyStep id="0E2657B9FF84FFEBFF26FE86FA9CFDA9" pageId="6" pageNumber="123">
<paragraph id="B56D4F1CFF84FFEBFF26FE86FA9CFE3E" blockId="6.[189,1399,265,1106]" pageId="6" pageNumber="123">
<keyLead id="0E23EC29FF84FFEBFF26FE86FA9CFE3E" pageId="6" pageNumber="123">
2. Striations on dorsum of head uniformly dense from frontal lobes to posterior margin; relatively large species (TL 2.24–3.16), total length of smallest worker&gt;
<quantity id="722AE2F9FF84FFEBFC02FEA3FC09FEF4" box="[921,994,300,327]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="123" unit="mm" value="2.0">2 mm</quantity>
. Large worker with head blackish brown, and either as long as wide or wider than long (CI 100–103) .................................................. ...................................................................................................
<taxonomicName id="72D2349FFF84FFEBFC53FEFCFAFEFE3F" authority="Wang &amp; Jaitrong" authorityName="Wang &amp; Yong &amp; Jaitrong" authorityYear="2021" box="[968,1301,370,397]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="robusta" status="sp. nov.">
<emphasis id="87A6930EFF84FFEBFC53FEFCFBADFE3E" bold="true" box="[968,1094,370,397]" italics="true" pageId="6" pageNumber="123">R. robusta</emphasis>
Wang &amp; Jaitrong
</taxonomicName>
<taxonomicNameLabel id="9C952E75FF84FFEBFA87FEFCFA9CFE3E" box="[1308,1399,371,397]" pageId="6" pageNumber="123" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="B56D4F1CFF84FFEBFF5AFE19FA9CFDA9" blockId="6.[189,1399,265,1106]" pageId="6" pageNumber="123">
<keyLead id="0E23EC29FF84FFEBFF5AFE19FA9CFDA9" pageId="6" pageNumber="123">
<emphasis id="87A6930EFF84FFEBFF5AFE19FF3BFE03" bold="true" box="[193,208,406,432]" pageId="6" pageNumber="123">–</emphasis>
Striations on dorsum of head weaker and more fragmented in posterior half; relatively smaller species (TL 1.84–2.93), total length of smallest worker &lt;
<quantity id="722AE2F9FF84FFEBFC1DFE36FC27FE60" box="[902,972,441,467]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="123" unit="mm" value="2.0">2 mm</quantity>
. Worker (of all sizes) with head dark brown, but never blackish, always longer than wide (CI 94–96) ....................................................... ............................................................................................
<taxonomicName id="72D2349FFF84FFEBFC0FFD8FFAFEFDAA" authority="Wang &amp; Jaitrong" authorityName="Wang &amp; Yong &amp; Jaitrong" authorityYear="2021" box="[916,1301,511,538]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="impithuksai" status="sp. nov.">
<emphasis id="87A6930EFF84FFEBFC0FFD8FFBAEFDA9" bold="true" box="[916,1093,511,538]" italics="true" pageId="6" pageNumber="123">R. impithuksai</emphasis>
Wang &amp; Jaitrong
</taxonomicName>
<taxonomicNameLabel id="9C952E75FF84FFEBFA87FD8FFA9CFDA9" box="[1308,1399,512,538]" pageId="6" pageNumber="123" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="0E2657B9FF84FFEBFF26FDC7FA9CFD5C" pageId="6" pageNumber="123">
<paragraph id="B56D4F1CFF84FFEBFF26FDC7FA9CFD1A" blockId="6.[189,1399,265,1106]" pageId="6" pageNumber="123">
<keyLead id="0E23EC29FF84FFEBFF26FDC7FA9CFD1A" pageId="6" pageNumber="123">
3. Head distinctly much longer than wide (CI 84–87); anterior margin of pronotal disc (excluding the collar) strongly convex with roundly angulate median point. Relatively small species (TL 1.85– 2.25) .............................................................................................
<taxonomicName id="72D2349FFF84FFEBFC44FD00FAFEFD1A" authority="Wang &amp; Jaitrong" authorityName="Wang &amp; Yong &amp; Jaitrong" authorityYear="2021" box="[991,1301,655,681]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="parva" status="sp. nov.">
<emphasis id="87A6930EFF84FFEBFC44FD00FBAEFD1A" bold="true" box="[991,1093,655,681]" italics="true" pageId="6" pageNumber="123">R. parva</emphasis>
Wang &amp; Jaitrong
</taxonomicName>
<taxonomicNameLabel id="9C952E75FF84FFEBFA87FD00FA9CFD1A" box="[1308,1399,655,681]" pageId="6" pageNumber="123" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="B56D4F1CFF84FFEBFF5AFD3DFA9CFD5C" blockId="6.[189,1399,265,1106]" pageId="6" pageNumber="123">
<keyLead id="0E23EC29FF84FFEBFF5AFD3DFA9CFD5C" pageId="6" pageNumber="123">
<emphasis id="87A6930EFF84FFEBFF5AFD3DFF3BFD7F" bold="true" box="[193,208,690,716]" pageId="6" pageNumber="123">–</emphasis>
Head not distinctly longer than wide (CI 87–96); anterior margin of pronotal disc (excluding the collar) broadly and often weakly convex with no angular median point ......................................... 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="0E2657B9FF84FFEBFF26FC91FA9CFBE1" pageId="6" pageNumber="123">
<paragraph id="B56D4F1CFF84FFEBFF26FC91FA9CFC11" blockId="6.[189,1399,265,1106]" pageId="6" pageNumber="123">
<keyLead id="0E23EC29FF84FFEBFF26FC91FA9CFC11" pageId="6" pageNumber="123">
4. Propodeal junction in lateral view distinct and roundly obtuse; propodeal dorsum differentiated from posterior declivity by weakly marginate angular edge. Posterolateral corner of propodeal dorsum angulate; posterior propodeal face overall weakly marginate. Anterior clypeal face in profile weakly and broadly convex .....................................................................................
<taxonomicName id="72D2349FFF84FFEBFBCBFC07FA9CFC11" authority="Wheeler, 1929" authorityName="Wheeler" authorityYear="1929" box="[1104,1399,904,930]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="javana">
<emphasis id="87A6930EFF84FFEBFBCBFC07FB28FC11" bold="true" box="[1104,1219,904,930]" italics="true" pageId="6" pageNumber="123">R. javana</emphasis>
<bibRefCitation id="D14332EDFF84FFEBFB50FC07FA9CFC11" author="Wheeler W. M." box="[1227,1399,904,930]" pageId="6" pageNumber="123" pagination="95 - 101" refId="ref24237" refString="Wheeler W. M. 1929. The ant genus Rhopalomastix. Psyche 36 (2): 95 - 101." type="journal article" year="1929">Wheeler, 1929</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B56D4F1CFF84FFEBFF5AFC24FA9CFBE1" blockId="6.[189,1399,265,1106]" pageId="6" pageNumber="123">
<keyLead id="0E23EC29FF84FFEBFF5AFC24FA9CFBE1" pageId="6" pageNumber="123">
– Propodeal junction in lateral view indistinct and broadly rounded, propodeal dorsum rounding smoothly into and not differentiated from posterior declivity by marginate angular edge. Posterolateral corner of propodeal dorsum usually indistinct, sometimes bluntly and shallowly rounded; posterior propodeal face overall incompletely marginate with obscure dorsal edge. Anterior clypeal face in profile mostly flat or weakly and broadly convex ................................
<taxonomicName id="72D2349FFF84FFEBFBBFFBB7FA9CFBE1" authority="Wheeler, 1929" authorityName="Wheeler" authorityYear="1929" box="[1060,1399,1079,1106]" class="Insecta" family="Formicidae" genus="Rhopalomastix" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="123" phylum="Arthropoda" rank="species" species="johorensis">
<emphasis id="87A6930EFF84FFEBFBBFFBB7FB2FFBE1" bold="true" box="[1060,1220,1079,1106]" italics="true" pageId="6" pageNumber="123">R. johorensis</emphasis>
<bibRefCitation id="D14332EDFF84FFEBFB50FBB7FA9CFBE1" author="Wheeler W. M." box="[1227,1399,1080,1106]" pageId="6" pageNumber="123" pagination="95 - 101" refId="ref24237" refString="Wheeler W. M. 1929. The ant genus Rhopalomastix. Psyche 36 (2): 95 - 101." type="journal article" year="1929">Wheeler, 1929</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>
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