treatments-xml/data/39/ED/26/39ED266850245127807C7803833D1A68.xml
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<document ID-DOI="http://dx.doi.org/10.3897/contrib.entomol.73.e101117" ID-Pensoft-Pub="2511-6428-1-31" ID-Pensoft-UUID="B18FC9724BE75A1D93667997031C092F" ID-ZooBank="70D521C7067F42A29242144AD3359645" ModsDocID="2511-6428-73-1-31" checkinTime="1689231645774" checkinUser="pensoft" docAuthor="Yokoi, Yaheita" docDate="2023" docId="39ED266850245127807C7803833D1A68" docLanguage="en" docName="ContribEntomol 73(1): 31-66" docOrigin="Contributions to Entomology 73 (1)" docPubDate="2023-07-12" docSource="http://dx.doi.org/10.3897/contrib.entomol.73.e101117" docTitle="Examnes lumawigi subsp. subrugosus Yokoi 2023, subsp. nov." docType="treatment" docUuid="7C452F51-85C5-41A8-A808-68F208A4F60E" docUuidSource="ZooBank" docVersion="1" id="B18FC9724BE75A1D93667997031C092F" lastPageNumber="31" masterDocId="B18FC9724BE75A1D93667997031C092F" masterDocTitle="Note on genitalia and taxonomy of the Callidiopini from the Philippines, with description of six new species and two subspecies (Coleoptera, Cerambycidae, Cerambycinae)" masterLastPageNumber="66" masterPageNumber="31" pageNumber="31" updateTime="1689231645774" updateUser="pensoft">
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<mods:title id="776F358E33FDEA8D937937528E6F5320">Note on genitalia and taxonomy of the Callidiopini from the Philippines, with description of six new species and two subspecies (Coleoptera, Cerambycidae, Cerambycinae)</mods:title>
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<mods:namePart id="184F7D6F8FF0F3B0CD3A10BB96E1D673">Yokoi, Yaheita</mods:namePart>
<mods:affiliation id="CC1FEFE305DDC61CDDEAFAD716F2549E">Beerenkothen 33, 40882 Ratingen, Germany</mods:affiliation>
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<mods:title id="3F050BCEE3CE89CF57DA33161D1728A4">Contributions to Entomology</mods:title>
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<mods:part id="C2BCEE78FAA3C23535721B8851B3DCD2">
<mods:date id="0B810F26EDC7D67D5D0BAFED5B0D8CE5">2023</mods:date>
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<mods:number id="25963BDCDB0F6E895EF69778E83252FD">2023-07-12</mods:number>
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<mods:number id="8E6CFD560F862A6F2F20E38C3FC4223A">73</mods:number>
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<subSubSection id="4A6E2D91CB0732BA7FD20475C08DE14C" pageId="0" pageNumber="31" type="nomenclature">
<paragraph id="B5052B53B7B3C65831DB45F0AFFCF06F" pageId="0" pageNumber="31">
<taxonomicName id="0A503FFA4B4B237C67373AF7BD4A81A9" LSID="https://zoobank.org/7C452F51-85C5-41A8-A808-68F208A4F60E" authority="Yokoi, 2023" authorityName="Yokoi" authorityYear="2023" class="Insecta" family="Cerambycidae" genus="Examnes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Examnes lumawigi subsp. subrugosus" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="lumawigi" status="subsp. nov." subSpecies="subrugosus">Examnes lumawigi subrugosus</taxonomicName>
<taxonomicNameLabel id="E9A4B721FFE6A6AB10AAC2CC5B6C0A2E" pageId="0" pageNumber="31">subsp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="B49B4E1D21C92889DD87C7EB5AF2B23F" pageId="0" pageNumber="31" type="description">
<paragraph id="64A157DC93B588205C53D30E032FD1AA" pageId="0" pageNumber="31">
<figureCitation id="D86F1F5BF6804E76E38C217007BCFBB2" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. A-F. Examnes lumawigi subrugosus subsp. nov. Holotype male. A. Habitus, dorsal view; B. Ditto, ventral; C. Head and pronotum; D. Ditto, front-lateral view; E. Prosternum and mesoventrite; F. Endophallus, with apical process." figureDoi="10.3897/contrib.entomol.73.e101117.figure13" httpUri="https://binary.pensoft.net/fig/876715" pageId="0" pageNumber="31">Figs 13A-F</figureCitation>
<figureCitation id="3239BAB82B8DFADE566588C373B4F353" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. A-L. Examnes lumawigi subrugosus subsp. nov. Holotype male. Genitalia. A. Median lobe with endophallus, dorsal view; B. Ditto, lateral; C. Ditto, ventral; D. Endophallus, apical process, dorsal view; E. Ditto, lateral; F. Ditto, ventral; G. Tegmen, dorsal view; H. Ditto, lateral; I. Ditto, ventral; J. 8 th sternite, ventral view; K. Ditto, with 8 th tergite in the background; L. 8 th tergite, ventral view. Scale bars: 0.5 mm (D-F); 1 mm (A-C, G-L)." figureDoi="10.3897/contrib.entomol.73.e101117.figure25" httpUri="https://binary.pensoft.net/fig/876727" pageId="0" pageNumber="31">, 25A-L</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="F64BC54CCE436DCD7F5737B1CB6141B7" pageId="0" pageNumber="31" type="type material">
<paragraph id="B094F1B296E60F97722C6F54F2B94B73" pageId="0" pageNumber="31">Type material.</paragraph>
<paragraph id="727B991F7B09C6938670D046C237A7DF" pageId="0" pageNumber="31">
<emphasis id="5075D7732841988F595D2DC84460CF9F" bold="true" italics="true" pageId="0" pageNumber="31">Holotype</emphasis>
♂ (SMF): &quot;Phil., E. Luzon, Sierra Madre&quot;; &quot;Dipaculao, Aurora, I-2014&quot;; &quot;Cope Collection&quot;. (former CJC).
</paragraph>
</subSubSection>
<subSubSection id="F38D3EE6099D4FAABC40D5BDF9233382" pageId="0" pageNumber="31" type="etymology">
<paragraph id="0D0996F681F80C5D3A85627351BD1CE2" pageId="0" pageNumber="31">Etymology.</paragraph>
<paragraph id="142C47C9B42224766E6B5C8D29DEEDD3" pageId="0" pageNumber="31">The name of this subspecies refers to the pronotal surface.</paragraph>
</subSubSection>
<subSubSection id="002087CB0403240EBD0A72A516397C02" pageId="0" pageNumber="31" type="diagnosis">
<paragraph id="48A929CDA7FAD4C1443C05AFCAAA82F6" pageId="0" pageNumber="31">Diagnosis.</paragraph>
<paragraph id="4DFEC1A5E0C8E4F5E1153257F5676E40" pageId="0" pageNumber="31">This subspecies differs from the nominotype in the structure and integument of pronotum.</paragraph>
</subSubSection>
<subSubSection id="A367498E0B6A1522732B6024952AEAC2" pageId="0" pageNumber="31" type="description">
<paragraph id="F7287DB02020511BFD3A476000DED99D" pageId="0" pageNumber="31">Measurements.</paragraph>
<paragraph id="607CB4578A382E76F60AC4017368468B" pageId="0" pageNumber="31">Holotype ♂: TL = 10.0 mm; EL/TL = 0.67; EL/EW = 2.85; HW/PW = 0.90; PL/PW = 1.02; PA/PW = 0.76; PB/PW = 0.92.</paragraph>
</subSubSection>
<subSubSection id="729872CA647E847BD6C89905D28FB2B8" pageId="0" pageNumber="31" type="description">
<paragraph id="92025FC10E4B3553B61E7D0894335C56" pageId="0" pageNumber="31">External characteristics: comparison with the nominotype.</paragraph>
<paragraph id="AFBF7DEFBD95CB4B89BB65FC193D2A80" pageId="0" pageNumber="31">
Body smaller. Pronotum with PL/PW = 1.02 slightly longer than wide, instead of &quot;distinctly longer than wide&quot;; surface obtusely rugose; several erect setae on each side (Fig.
<figureCitation id="BC09CA242A95406B7A280F5392018C3E" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. A-F. Examnes lumawigi subrugosus subsp. nov. Holotype male. A. Habitus, dorsal view; B. Ditto, ventral; C. Head and pronotum; D. Ditto, front-lateral view; E. Prosternum and mesoventrite; F. Endophallus, with apical process." figureDoi="10.3897/contrib.entomol.73.e101117.figure13" httpUri="https://binary.pensoft.net/fig/876715" pageId="0" pageNumber="31">13C, D</figureCitation>
). 7th sternite feebly emarginated instead of sub-truncated.
</paragraph>
<caption id="9129D8DC514042E7DC47431A5550351C" doi="10.3897/contrib.entomol.73.e101117.figure13" httpUri="https://binary.pensoft.net/fig/876715" pageId="0" pageNumber="31" start="Figure 13" startId="F13">
<paragraph id="D8AA71F1B1BF21BB8F0D23018403E896" pageId="0" pageNumber="31">
<emphasis id="254D8B88B72E622B42716F07D5DDE766" bold="true" pageId="0" pageNumber="31">Figure 13.</emphasis>
<emphasis id="40C411612E1E6FB3C7C6E4859B98BD50" bold="true" pageId="0" pageNumber="31">A-F.</emphasis>
<taxonomicName id="07DB8A35BE604AC03FCD649A4BB0B4CF" authorityName="Yokoi" authorityYear="2023" class="Insecta" family="Cerambycidae" genus="Examnes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Examnes lumawigi subsp. subrugosus" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="lumawigi" subSpecies="subrugosus">
<emphasis id="50E646B395DDA6D18873A9F32A31C6F6" italics="true" pageId="0" pageNumber="31">Examnes lumawigi subrugosus</emphasis>
</taxonomicName>
subsp. nov. Holotype male.
<emphasis id="146C405547B141B147A1922074CB65E0" bold="true" pageId="0" pageNumber="31">A.</emphasis>
Habitus, dorsal view;
<emphasis id="1188C939A676E90ACB7A3B5CF64198A0" bold="true" pageId="0" pageNumber="31">B.</emphasis>
Ditto, ventral;
<emphasis id="68E309BDB501BD35219E9535B37E8642" bold="true" pageId="0" pageNumber="31">C.</emphasis>
Head and pronotum;
<emphasis id="656E4F3F20D4809199F7748E5ADC5882" bold="true" pageId="0" pageNumber="31">D.</emphasis>
Ditto, front-lateral view;
<emphasis id="770864522A7A5A66381DDB983ED7A803" bold="true" pageId="0" pageNumber="31">E.</emphasis>
Prosternum and mesoventrite;
<emphasis id="33DEE7913EF9BE77C5A9C1272D6CEA45" bold="true" pageId="0" pageNumber="31">F.</emphasis>
Endophallus, with apical process.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="3F7F69A0D5FA9B81E6F410088E121969" pageId="0" pageNumber="31" type="description">
<paragraph id="C3B35B5B3563F60F5C635ADF622D679F" pageId="0" pageNumber="31">Description of male genitalia</paragraph>
<paragraph id="E536562B4390984FFCA6163B903647B4" pageId="0" pageNumber="31">
<emphasis id="E500B1F6E16C2AC45E8BE7D7F60B3323" bold="true" pageId="0" pageNumber="31">
(Figs
<figureCitation id="6CC0AD2E5E04DD199B47BF591B185D92" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. A-F. Examnes lumawigi subrugosus subsp. nov. Holotype male. A. Habitus, dorsal view; B. Ditto, ventral; C. Head and pronotum; D. Ditto, front-lateral view; E. Prosternum and mesoventrite; F. Endophallus, with apical process." figureDoi="10.3897/contrib.entomol.73.e101117.figure13" httpUri="https://binary.pensoft.net/fig/876715" pageId="0" pageNumber="31">13F</figureCitation>
,
<figureCitation id="DB97A4EB755EC1610C55F9EDFF46F4AF" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. A-L. Examnes lumawigi subrugosus subsp. nov. Holotype male. Genitalia. A. Median lobe with endophallus, dorsal view; B. Ditto, lateral; C. Ditto, ventral; D. Endophallus, apical process, dorsal view; E. Ditto, lateral; F. Ditto, ventral; G. Tegmen, dorsal view; H. Ditto, lateral; I. Ditto, ventral; J. 8 th sternite, ventral view; K. Ditto, with 8 th tergite in the background; L. 8 th tergite, ventral view. Scale bars: 0.5 mm (D-F); 1 mm (A-C, G-L)." figureDoi="10.3897/contrib.entomol.73.e101117.figure25" httpUri="https://binary.pensoft.net/fig/876727" pageId="0" pageNumber="31">25A-L</figureCitation>
).
</emphasis>
Median lobe about half the length of abdomen; sub-bullet-shaped in dorsal view, in lateral view strongly curved in the middle; dorsal plate dehiscent in basal 11/20; ventral plate longer, dehiscent in basal 3/4, gradually narrowed toward apex. Tegmen almost 5/6 as long as median lobe; weakly bi-sinuate in lateral view. Parameres half the length of tegmen; apical 1/6 bi-lobed; lobes each fringed with short to medium setae; venter widely emarginated. Apical process of endophallus as Figs
<figureCitation id="4655FC20F4B01F886A50C7866FEDA2C5" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. A-F. Examnes lumawigi subrugosus subsp. nov. Holotype male. A. Habitus, dorsal view; B. Ditto, ventral; C. Head and pronotum; D. Ditto, front-lateral view; E. Prosternum and mesoventrite; F. Endophallus, with apical process." figureDoi="10.3897/contrib.entomol.73.e101117.figure13" httpUri="https://binary.pensoft.net/fig/876715" pageId="0" pageNumber="31">13F</figureCitation>
,
<figureCitation id="44B95A468E4651CD3C6665BA8029C98C" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. A-L. Examnes lumawigi subrugosus subsp. nov. Holotype male. Genitalia. A. Median lobe with endophallus, dorsal view; B. Ditto, lateral; C. Ditto, ventral; D. Endophallus, apical process, dorsal view; E. Ditto, lateral; F. Ditto, ventral; G. Tegmen, dorsal view; H. Ditto, lateral; I. Ditto, ventral; J. 8 th sternite, ventral view; K. Ditto, with 8 th tergite in the background; L. 8 th tergite, ventral view. Scale bars: 0.5 mm (D-F); 1 mm (A-C, G-L)." figureDoi="10.3897/contrib.entomol.73.e101117.figure25" httpUri="https://binary.pensoft.net/fig/876727" pageId="0" pageNumber="31">25D-F</figureCitation>
; about 1/3 as long as median lobe, sub-monolithic, sub-fusiform in dorsal view, arcuate in lateral view, with a prominent, horn-shaped dorsal projection; sides thick, elongated; the middle sub-enclosed by a membranous sack; base sub-triangular-spatulate, bent upward in profile. Blade of 8th sternite transverse, cotyledonary; apex in shallow
<normalizedToken id="EB7B257617A073B3DA19E8E28AD1789E" originalValue="“V”">&quot;V&quot;</normalizedToken>
-form emarginated; extensively thinned in middle; apex and sides fringed with sporadic long hairs; apical middle clothed with additional shorter hairs; basal middle thickened near the peduncle; peduncle nearly as long as blade. 8th tergite transverse, sub-trapezoidal; corners rounded; fringed with sporadic medium to long hairs.
</paragraph>
</subSubSection>
<subSubSection id="9373DB1A9F9A2E791A4E897A3E5C51FE" pageId="0" pageNumber="31" type="distribution">
<paragraph id="08EEA245AC929E3ADCD77E1284B7FD41" pageId="0" pageNumber="31">Distribution.</paragraph>
<paragraph id="5924070A1ACCC1C87B7D1539CF11D9E9" pageId="0" pageNumber="31">Luzon, Philippines.</paragraph>
</subSubSection>
<subSubSection id="AAF93555486FF4C7FFA081255468E01E" pageId="0" pageNumber="31" type="comparisons">
<paragraph id="0FE7B2BE71F90F0AA84096D73ED9BE72" pageId="0" pageNumber="31">Comparative notes.</paragraph>
<paragraph id="84F5AF66FAEA809C1363C42CC0E19073" pageId="0" pageNumber="31">
The pronotal structure and surface of this subspecies differ substantially from those of the nominotype
<taxonomicName id="C498458F74D84F6D29A06AAEDAB4CB87" authority="lumawigi" authorityName="lumawigi Huedepohl" authorityYear="1990" class="Insecta" family="Cerambycidae" genus="Examnes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Examnes lumawigi subsp. lumawigi" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="lumawigi" subSpecies="lumawigi">
<emphasis id="42416BBC955D484404813A50F33D8F39" italics="true" pageId="0" pageNumber="31">Examnes lumawigi lumawigi</emphasis>
</taxonomicName>
(Fig.
<figureCitation id="39161A8B6B04D2261CB43C6721F2F557" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. A, B. Pronotum in lateral view, comparison. A. Examnes lumawigi lumawigi Huedepohl. Holotype male, from Mountain Province, Luzon. B. Examnes lumawigi subrugosus subsp. nov. Holotype male, from Sierra Madre, Luzon." figureDoi="10.3897/contrib.entomol.73.e101117.figure14" httpUri="https://binary.pensoft.net/fig/876716" pageId="0" pageNumber="31">14A, B</figureCitation>
). The other observed differences are not of fundamental nature. The holotype male of this new subspecies was collected in Luzon, same as that of
<taxonomicName id="518ED8D1E2A4428CD2078DAB0E1BA1AC" authority="lumawigi" lsidName="E. lumawigi subsp. lumawigi" pageId="0" pageNumber="31" rank="subSpecies" species="lumawigi" subSpecies="lumawigi">
<emphasis id="F5CC04030DA3A97D0C119972C5E81451" italics="true" pageId="0" pageNumber="31">E. lumawigi lumawigi</emphasis>
</taxonomicName>
, though in a geographically separated locality. Its final taxonomical position should be determined by comparison of male genitalia. In this publication, it is described as a new subspecies.
</paragraph>
<caption id="9650006DD6434E178C5C66A7010C9BB2" doi="10.3897/contrib.entomol.73.e101117.figure14" httpUri="https://binary.pensoft.net/fig/876716" pageId="0" pageNumber="31" start="Figure 14" startId="F14">
<paragraph id="2FEC2042BE058DA7FCBC78ED7CBD6550" pageId="0" pageNumber="31">
<emphasis id="5796A3B18A1980D4FA060E75E6EF7E6D" bold="true" pageId="0" pageNumber="31">Figure 14.</emphasis>
<emphasis id="9206885A2BF7F6F08E8FCE0A00622C20" bold="true" pageId="0" pageNumber="31">A, B.</emphasis>
Pronotum in lateral view, comparison.
<emphasis id="081480FD39AD3DB4567B2851109F7861" bold="true" pageId="0" pageNumber="31">A.</emphasis>
<taxonomicName id="87C0BB5BE21D367B6A44D16058A0183C" authority="lumawigi" authorityName="lumawigi Huedepohl" authorityYear="1990" class="Insecta" family="Cerambycidae" genus="Examnes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Examnes lumawigi subsp. lumawigi" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="lumawigi" subSpecies="lumawigi">
<emphasis id="B004DAE2C83F0ACC7112F5DC51479FCB" italics="true" pageId="0" pageNumber="31">Examnes lumawigi lumawigi</emphasis>
</taxonomicName>
<normalizedToken id="5D536DBE2673CD0585591E19765CBFA9" originalValue="Hüdepohl">Huedepohl</normalizedToken>
. Holotype male, from Mountain Province, Luzon.
<emphasis id="299A70FEDA43A6EB5E57831BC98E0615" bold="true" pageId="0" pageNumber="31">B.</emphasis>
<taxonomicName id="3E2630AE3B5FE005D68AA4CBE394D7FE" authorityName="Yokoi" authorityYear="2023" class="Insecta" family="Cerambycidae" genus="Examnes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Examnes lumawigi subsp. subrugosus" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="lumawigi" subSpecies="subrugosus">
<emphasis id="5896302CF0575330F01743DD9595E58C" italics="true" pageId="0" pageNumber="31">Examnes lumawigi subrugosus</emphasis>
</taxonomicName>
subsp. nov. Holotype male, from Sierra Madre, Luzon.
</paragraph>
</caption>
<paragraph id="4D63B214DD1410DDA051C8EF5486802B" pageId="0" pageNumber="31">
Apical process of its endophallus is essentially of similar structure as that of
<taxonomicName id="D55EA40D0602C7E67F2AD45BF4309F5C" lsidName="E. philippensis" pageId="0" pageNumber="31" rank="species" species="philippensis">
<emphasis id="D2135ED025A08B070B7DB877DE117305" italics="true" pageId="0" pageNumber="31">E. philippensis</emphasis>
</taxonomicName>
, nominotype of the genus
<taxonomicName id="8F0DB47AB23B8424825B99600B7D41CD" authorityName="Pascoe" authorityYear="1869" class="Insecta" family="Cerambycidae" genus="Examnes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Examnes" order="Coleoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="E00F403DE86B8392893619BEE11A647F" italics="true" pageId="0" pageNumber="31">Examnes</emphasis>
</taxonomicName>
. Differences are concerned with the structures of sides and base as well as of appendage (Figs
<figureCitation id="0A2C4D69A6D1AD8C94BC2D8E380EF766" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. A-L. Examnes lumawigi subrugosus subsp. nov. Holotype male. Genitalia. A. Median lobe with endophallus, dorsal view; B. Ditto, lateral; C. Ditto, ventral; D. Endophallus, apical process, dorsal view; E. Ditto, lateral; F. Ditto, ventral; G. Tegmen, dorsal view; H. Ditto, lateral; I. Ditto, ventral; J. 8 th sternite, ventral view; K. Ditto, with 8 th tergite in the background; L. 8 th tergite, ventral view. Scale bars: 0.5 mm (D-F); 1 mm (A-C, G-L)." figureDoi="10.3897/contrib.entomol.73.e101117.figure25" httpUri="https://binary.pensoft.net/fig/876727" pageId="0" pageNumber="31">25D-F</figureCitation>
,
<figureCitation id="D6F13AF663B68BE628A230B261513877" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. A-L. Examnes philippensis Newman. Male from Basilan, Philippines (New record). Genitalia. A. Median lobe with endophallus, dorsal view; B. Ditto, lateral; C. Ditto, ventral; D. Endophallus, apical process, dorsal view; E. Ditto, lateral; F. Ditto, ventral; G. Tegmen, dorsal view; H. Ditto, lateral; I. Ditto, ventral; J. 8 th sternite, ventral view; K. Ditto, with 8 th tergite in the background; L. 9 th sternite (spicum gastrale) and tergite, ventral view. Scale bars: 0.5 mm (D-F); 1 mm (A-C, G-L)." figureDoi="10.3897/contrib.entomol.73.e101117.figure27" httpUri="https://binary.pensoft.net/fig/876729" pageId="0" pageNumber="31">27D-F</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>