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<document id="9F5E03C68A8CC7F2547D74E6D7E263A0" ID-DOI="10.5281/zenodo.5404276" ID-ISSN="0374-1036" ID-Zenodo-Dep="5404276" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630132078781" checkinUser="felipe" docAuthor="Hromádka, Lubomír" docDate="2009" docId="142187F5FFD3FF99F5E9FB26FE16FA14" docLanguage="en" docName="ActaEntMusNatPra.2009.49.1.161-190.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 49 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Philonthus limbipennis Fauvel 1905" docType="treatment" docVersion="3" lastPageNumber="183" masterDocId="E818FF8DFFC6FF8FF464FFBCFFA1FFC7" masterDocTitle="Revision of the Afrotropical species of the Philonthus caffer species group (Coleoptera: Staphylinidae: Philonthina)" masterLastPageNumber="190" masterPageNumber="161" pageNumber="182" updateTime="1699162834734" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="C8B3EEFE0C6E576A0678BC53AC3B55C3">Revision of the Afrotropical species of the Philonthus caffer species group (Coleoptera: Staphylinidae: Philonthina)</mods:title>
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<mods:namePart id="1E783A67D852709A5847E0BB53D2862E">Hromádka, Lubomír</mods:namePart>
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<mods:title id="FA1D6A012BBC89FBB8FEFB7A055D8FAD">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="8BC9D655AF8D767E6528915A21D56AF4">2009</mods:date>
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<mods:number id="33889B45F81E5048F0F01B1B04365D5F">2009-06-30</mods:number>
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<mods:number id="5E6BFDEFC1B3A61F75243B48E29CBE4B">49</mods:number>
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<mods:identifier id="6A34BD998600C7091B3BC2A25D2E1037" type="ISSN">0374-1036</mods:identifier>
<mods:identifier id="206B2705DCC181C68F8EEF2261B8E8B5" type="Zenodo-Dep">5404276</mods:identifier>
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<treatment id="142187F5FFD3FF99F5E9FB26FE16FA14" LSID="urn:lsid:plazi:treatment:142187F5FFD3FF99F5E9FB26FE16FA14" httpUri="http://treatment.plazi.org/id/142187F5FFD3FF99F5E9FB26FE16FA14" lastPageId="22" lastPageNumber="183" pageId="21" pageNumber="182">
<subSubSection id="D4926568FFD3FF9AF5E9FB26FC85FB74" box="[397,804,1178,1204]" pageId="21" pageNumber="182" type="nomenclature">
<paragraph id="9C3736E3FFD3FF9AF5E9FB26FC85FB74" blockId="21.[397,804,1178,1204]" box="[397,804,1178,1204]" pageId="21" pageNumber="182">
<heading id="C77F818FFFD3FF9AF5E9FB26FC85FB74" bold="true" box="[397,804,1178,1204]" centered="true" fontSize="8" level="2" pageId="21" pageNumber="182" reason="3">
<taxonomicName id="5B884D60FFD3FF9AF5E9FB26FC85FB74" ID-CoL="4GB66" authority="Fauvel, 1905" authorityName="Fauvel" authorityYear="1905" box="[397,804,1178,1204]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="182" phylum="Arthropoda" rank="species" species="limbipennis">
<emphasis id="AEFCEAF1FFD3FF9AF5E9FB26FC85FB74" bold="true" box="[397,804,1178,1204]" pageId="21" pageNumber="182">
<emphasis id="AEFCEAF1FFD3FF9AF5E9FB26FD2FFB74" bold="true" box="[397,654,1178,1203]" italics="true" pageId="21" pageNumber="182">Philonthus limbipennis</emphasis>
<bibRefCitation id="F8194B12FFD3FF9AF6F1FB20FC85FB74" author="FAUVEL A." box="[661,804,1179,1204]" pageId="21" pageNumber="182" pagination="149 - 184" refId="ref16362" refString="FAUVEL A. 1905: Staphylinides nouveaux de Madagascar. 2 i Partie. Revue d'Entomologie 24: 149 - 184." type="journal article" year="1905">Fauvel, 1905</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D4926568FFD3FF9AF67BFB06FD32FB0A" box="[543,659,1210,1229]" pageId="21" pageNumber="182" type="description">
<paragraph id="9C3736E3FFD3FF9AF67BFB06FD32FB0A" blockId="21.[113,1089,1210,1404]" box="[543,659,1210,1229]" pageId="21" pageNumber="182">
(
<figureCitation id="04B32A66FFD3FF9AF642FB06FD2CFB0A" box="[550,653,1210,1229]" captionStart="Figs" captionStartId="26.[151,191,1193,1212]" captionTargetBox="[160,1118,221,1161]" captionTargetId="figure-101@26.[151,1128,214,1166]" captionTargetPageId="26" captionText="Figs. 5461. 5456 Philonthus limbipennis Fauvel, 1905. 5758 P. maridadus Tottenham, 1962; 5961 P. sequens Bernhauer &amp; Schubert, 1914; 6263 P. tomicus Tottenham, 1962. 54, 57, 59 aedeagus, ventral view; 55, 58, 60 aedeagus, lateral view; 56, 61 apex of paramere with sensory peg setae, ventral view; 62 aedeagus, dorsal view; 63 paramere, ventral,view." figureDoi="http://doi.org/10.5281/zenodo.5404286" httpUri="https://zenodo.org/record/5404286/files/figure.png" pageId="21" pageNumber="182">Figs. 5456</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D4926568FFD3FF9AF415FB61FE79FB28" box="[113,472,1244,1263]" pageId="21" pageNumber="182" type="reference_group">
<paragraph id="9C3736E3FFD3FF9AF415FB61FE79FB28" blockId="21.[113,1089,1210,1404]" box="[113,472,1244,1263]" pageId="21" pageNumber="182">
<treatmentCitationGroup id="BC9811CDFFD3FF9AF415FB61FE79FB28" box="[113,472,1244,1263]" pageId="21" pageNumber="182">
<treatmentCitation id="1D2910F2FFD3FF9AF415FB61FE75FB28" author="FAUVEL A." box="[113,468,1244,1263]" page="175" pageId="21" pageNumber="182" year="1905">
<taxonomicName id="5B884D60FFD3FF9AF415FB61FE75FB28" ID-CoL="4GB66" authority="Fauvel, 1905: 175" authorityName="Fauvel" authorityPageNumber="175" authorityYear="1905" box="[113,468,1244,1263]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="182" phylum="Arthropoda" rank="species" species="limbipennis">
<emphasis id="AEFCEAF1FFD3FF9AF415FB61FE96FB28" box="[113,311,1245,1263]" italics="true" pageId="21" pageNumber="182">Philonthus limbipennis</emphasis>
<bibRefCitation id="F8194B12FFD3FF9AF558FB60FE75FB28" author="FAUVEL A." box="[316,468,1244,1263]" pageId="21" pageNumber="182" pagination="149 - 184" refId="ref16362" refString="FAUVEL A. 1905: Staphylinides nouveaux de Madagascar. 2 i Partie. Revue d'Entomologie 24: 149 - 184." type="journal article" year="1905">Fauvel, 1905:175</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="D4926568FFD3FF9AF415FB42FCEDFABB" pageId="21" pageNumber="182" type="materials_examined">
<paragraph id="9C3736E3FFD3FF9AF415FB42FDFFFAD6" blockId="21.[113,1089,1210,1404]" box="[113,606,1277,1297]" pageId="21" pageNumber="182">
<emphasis id="AEFCEAF1FFD3FF9AF415FB42FF4BFAD6" bold="true" box="[113,234,1277,1297]" pageId="21" pageNumber="182">
<typeStatus id="43338841FFD3FF9AF415FB42FF3FFAD6" box="[113,158,1278,1297]" pageId="21" pageNumber="182">Type</typeStatus>
locality.
</emphasis>
Diego-Suarez; forêt Audroy; Ambovombe.
</paragraph>
<paragraph id="9C3736E3FFD3FF9AF415FAA5FC57FA80" blockId="21.[113,1089,1210,1404]" pageId="21" pageNumber="182">
<emphasis id="AEFCEAF1FFD3FF9AF415FAA5FEF3FAEB" bold="true" box="[113,338,1304,1324]" pageId="21" pageNumber="182">Type material examined.</emphasis>
<materialsCitation id="2CE03CBEFFD3FF9AF533FAA5FD2CFA80" collectorName="Ex-Typus J &amp; Foret Audroy, R. I. Sc. N. B." country="Madagascar" determinerName="A. Fauvel'" location="Madagascar" pageId="21" pageNumber="182" specimenCount="1" typeStatus="lectotype">
<typeStatus id="43338841FFD3FF9AF533FAA5FE11FAEC" box="[343,432,1305,1324]" pageId="21" pageNumber="182" type="lectotype">LECTOTYPE</typeStatus>
(here designated):
<collectingCountry id="E49F7673FFD3FF9AF63EFAA5FD63FAEB" box="[602,706,1305,1324]" name="Madagascar" pageId="21" pageNumber="182">Madagascar</collectingCountry>
,
<collectorName id="317D5335FFD3FF9AF6AEFAA5FC90FAEA" box="[714,817,1300,1325]" pageId="21" pageNumber="182">Ex-Typus J</collectorName>
:
<collectorName id="317D5335FFD3FF9AF727FAA5FB83FAEB" box="[835,1058,1305,1324]" pageId="21" pageNumber="182">Forêt Audroy, R.I.Sc.N.B.</collectorName>
17. 479, coll. et det.
<determinerName id="69EBDA27FFD3FF9AF49BFA88FEF8FA80" box="[255,345,1332,1351]" pageId="21" pageNumber="182">A. Fauvel</determinerName>
[white oblong label, handwritten] //
</materialsCitation>
<materialsCitation id="2CE03CBEFFD3FF9AF6F6FA88FC53FA80" box="[658,1010,1332,1351]" collectionCode="MRAC" collectorName="C. E. Tottenham'" country="Madagascar" pageId="21" pageNumber="182" specimenCount="1" typeStatus="lectotype">
<typeStatus id="43338841FFD3FF9AF6F6FA88FCADFA80" box="[658,780,1332,1351]" pageId="21" pageNumber="182" type="lectotype">LECTOTYPE</typeStatus>
<collectorName id="317D5335FFD3FF9AF775FA88FC3DFA80" box="[785,924,1332,1351]" pageId="21" pageNumber="182">C.E. Tottenham</collectorName>
(
<collectionCode id="FA99AE26FFD3FF9AF7CCFA88FC4DFA80" box="[936,1004,1332,1351]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="21" pageNumber="182" type="Museum">MRAC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="9C3736E3FFD3FF9AF415FAF2FCEDFABB" blockId="21.[113,1089,1210,1404]" pageId="21" pageNumber="182">
<emphasis id="AEFCEAF1FFD3FF9AF415FAF2FF00FAA6" bold="true" box="[113,161,1358,1377]" pageId="21" pageNumber="182">Note.</emphasis>
The designation of the
<typeStatus id="43338841FFD3FF9AF50AFAF2FE1DFAA6" box="[366,444,1358,1377]" pageId="21" pageNumber="182" type="lectotype">lectotype</typeStatus>
indicated by a label attached to the specimen examined by Tottenham was never published. For this reason, I am designating the specimen as the
<typeStatus id="43338841FFD3FF9AF6A9FAD5FCBAFABB" box="[717,795,1385,1404]" pageId="21" pageNumber="182" type="lectotype">lectotype</typeStatus>
here.
</paragraph>
</subSubSection>
<subSubSection id="D4926568FFD3FF99F415FA28FE23FAF4" lastPageId="22" lastPageNumber="183" pageId="21" pageNumber="182" type="description">
<paragraph id="9C3736E3FFD3FF9AF415FA28FC4CFA6B" blockId="21.[113,1089,1428,1516]" box="[113,1005,1428,1453]" pageId="21" pageNumber="182">
<emphasis id="AEFCEAF1FFD3FF9AF415FA28FEB1FA6B" bold="true" box="[113,272,1428,1452]" pageId="21" pageNumber="182">Redescription</emphasis>
. Body length
<quantity id="5B709B06FFD3FF9AF5C3FA28FE5AFA6B" box="[423,507,1428,1452]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="21" pageNumber="182" unit="mm" value="6.7">6.7 mm</quantity>
, length of fore body (to end of elytra)
<quantity id="5B709B06FFD3FF9AF7F0FA28FC48FA6B" box="[916,1001,1428,1452]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="21" pageNumber="182" unit="mm" value="4.5">4.5 mm</quantity>
.
</paragraph>
<paragraph id="9C3736E3FFD3FF99F4F4FA08FB96FEF4" blockId="21.[113,1089,1428,1516]" lastBlockId="22.[151,1128,219,1491]" lastPageId="22" lastPageNumber="183" pageId="21" pageNumber="182">Colouration. Head and abdomen black, pronotum, scutellum and elytra black-brown, epipleura and posterior margin of elytra narrowly yellow-brown, maxillary and labial palpi brown-yellow, terminal palpomeres yellow-brown, mandibles brown, antennae dark brown, base of antennomere 2 yellow-brown. Femora and tibiae brown, medial face of all tibiae vaguely darker, tarsi yellow-brown. Head and pronotum with slight purple iridescence.</paragraph>
<paragraph id="9C3736E3FFD0FF99F4D1FE87FD18FE34" blockId="22.[151,1128,219,1491]" pageId="22" pageNumber="183">Head quadrangular, slightly wider than long (ratio 25: 21.5), parallel-sided, posterior angles conspicuously rounded, each bearing two long black bristles. Eyes small, temples much longer than eyes (ratio 12: 6). Distance between medial interocular punctures 4 times as large as distance between medial and lateral interocular punctures. Lateral interocular punctures slightly shifted anteriad. Temporal area with several punctures. Surface with very fine microsculpture consisting of transverse waves.</paragraph>
<paragraph id="9C3736E3FFD0FF99F4D1FE40FDBAFD93" blockId="22.[151,1128,219,1491]" pageId="22" pageNumber="183">Antennae stout, reaching posterior fourth of pronotum when reclined. Antennomeres 13 and 11 longer than wide, antennomeres 410 as long as wide. Relative length of antennomeres: 1 = 7; 23 = 4.5; 410 = 2.5; 11 = 4.</paragraph>
<paragraph id="9C3736E3FFD0FF99F4D1FDE0FD4EFCD4" blockId="22.[151,1128,219,1491]" pageId="22" pageNumber="183">Pronotum slightly wider than long (ratio 24.5: 23), highly convex, parallel-sided, anterior angles almost rectangular, each with several short black bristles, sides with one long black bristle in anterior third. Left dorsal row with 4 punctures, right dorsal row with 5 punctures, punctures 25 equidistant, distance between punctures 1 and 2 twice as large as distance between remaining punctures of row. Each sublateral row with 2 punctures. Surface with very fine microsculpture consisting of transverse waves.</paragraph>
<paragraph id="9C3736E3FFD0FF99F4D1FCA7FE0EFCB4" blockId="22.[151,1128,219,1491]" pageId="22" pageNumber="183">Scutellum punctate only in posterior two thirds, punctation fine and sparse, punctures somewhat larger than eye-facets, separated by a puncture diameter in transverse direction. Anterior third impunctate.</paragraph>
<paragraph id="9C3736E3FFD0FF99F4D1FCC7FD36FC34" blockId="22.[151,1128,219,1491]" pageId="22" pageNumber="183">Elytra wider than long (ratio 31: 28), vaguely widened posteriad, anterior angles of elytra each with one long black bristle. Punctation coarser and sparser, punctures somewhat larger than those of scutellum, separated by 1.52 puncture diameters in transverse direction. Surface without microsculpture; setation dark and short.</paragraph>
<paragraph id="9C3736E3FFD0FF99F4D1FC40FC78FBF3" blockId="22.[151,1128,219,1491]" pageId="22" pageNumber="183">Legs. Metatibia longer than metatarsus (ratio 17: 14). Metatarsomere 1 as long as metatarsomere 5. Relative length of metatarsomeres: 1 = 4; 2 = 2; 34 = 1.5; 5 = 4.</paragraph>
<paragraph id="9C3736E3FFD0FF99F4D1FB87FD56FB14" blockId="22.[151,1128,219,1491]" pageId="22" pageNumber="183">Abdomen wide, slightly narrowed from tergite III toward apex and base. Elevated area between two basal lines on first three visible tergites almost impunctate. Punctation of all tergites similar as on scutellum, gradually becoming finer and much sparser towards posterior margin of each tergite, separated by 2 or 3 puncture diameters in transverse direction. Surface without microsculpture; setation similar to that on elytra.</paragraph>
<paragraph id="9C3736E3FFD0FF99F4D1FB67FBC6FAD4" blockId="22.[151,1128,219,1491]" pageId="22" pageNumber="183">
Male. Protarsomeres 13 relatively slightly dilated, sub-bilobed, covered with modified pale setae ventrally, protarsomere 4 narrower than preceding ones. Aedeagus (
<figureCitation id="04B32A66FFD0FF99F7BCFB47FBFAFAD4" box="[984,1115,1275,1299]" captionStart="Figs" captionStartId="26.[151,191,1193,1212]" captionTargetBox="[160,1118,221,1161]" captionTargetId="figure-101@26.[151,1128,214,1166]" captionTargetPageId="26" captionText="Figs. 5461. 5456 Philonthus limbipennis Fauvel, 1905. 5758 P. maridadus Tottenham, 1962; 5961 P. sequens Bernhauer &amp; Schubert, 1914; 6263 P. tomicus Tottenham, 1962. 54, 57, 59 aedeagus, ventral view; 55, 58, 60 aedeagus, lateral view; 56, 61 apex of paramere with sensory peg setae, ventral view; 62 aedeagus, dorsal view; 63 paramere, ventral,view." figureDoi="http://doi.org/10.5281/zenodo.5404286" httpUri="https://zenodo.org/record/5404286/files/figure.png" pageId="22" pageNumber="183">Figs. 5456</figureCitation>
).
</paragraph>
<paragraph id="9C3736E3FFD0FF99F4D1FAA7FE23FAF4" blockId="22.[151,1128,219,1491]" box="[181,386,1307,1331]" pageId="22" pageNumber="183">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="D4926568FFD0FF99F4F3FA87FE2DFA74" pageId="22" pageNumber="183" type="diagnosis">
<paragraph id="9C3736E3FFD0FF99F4F3FA87FCE6FA54" blockId="22.[151,1128,219,1491]" pageId="22" pageNumber="183">
<emphasis id="AEFCEAF1FFD0FF99F4F3FA87FE32FA94" bold="true" box="[151,403,1339,1363]" pageId="22" pageNumber="183">Differential diagnosis.</emphasis>
<taxonomicName id="5B884D60FFD0FF99F5FFFA81FD33FA93" authorityName="Fauvel" authorityYear="1905" box="[411,658,1341,1364]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="183" phylum="Arthropoda" rank="species" species="limbipennis">
<emphasis id="AEFCEAF1FFD0FF99F5FFFA81FD33FA93" box="[411,658,1341,1364]" italics="true" pageId="22" pageNumber="183">Philonthus limbipennis</emphasis>
</taxonomicName>
is in most characters similar to
<taxonomicName id="5B884D60FFD0FF99F795FA81FBC1FA93" authorityName="Tottenham" authorityYear="1962" box="[1009,1120,1341,1364]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="183" phylum="Arthropoda" rank="species" species="tomicus">
<emphasis id="AEFCEAF1FFD0FF99F795FA81FBC1FA93" box="[1009,1120,1341,1364]" italics="true" pageId="22" pageNumber="183">P. tomicus</emphasis>
</taxonomicName>
, but differs by the shorter eyes and antennae, slight purple iridescence of head and pronotum, uniformly black-brown elytra, and by the shape of the aedeagus.
</paragraph>
<paragraph id="9C3736E3FFD0FF99F4F3FA27FE2DFA74" blockId="22.[151,1128,219,1491]" box="[151,396,1435,1459]" pageId="22" pageNumber="183">
<emphasis id="AEFCEAF1FFD0FF99F4F3FA27FEB4FA74" bold="true" box="[151,277,1435,1459]" pageId="22" pageNumber="183">Bionomics.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="D4926568FFD0FF99F4F3FA07FE16FA14" box="[151,439,1467,1491]" pageId="22" pageNumber="183" type="distribution">
<paragraph id="9C3736E3FFD0FF99F4F3FA07FE16FA14" blockId="22.[151,1128,219,1491]" box="[151,439,1467,1491]" pageId="22" pageNumber="183">
<emphasis id="AEFCEAF1FFD0FF99F4F3FA07FE8BFA14" bold="true" box="[151,298,1467,1491]" pageId="22" pageNumber="183">Distribution.</emphasis>
<collectingCountry id="E49F7673FFD0FF99F554FA07FE12FA14" box="[304,435,1467,1491]" name="Madagascar" pageId="22" pageNumber="183">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>