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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1175.102485" ID-Pensoft-Pub="1313-2970-1175-223" ID-Pensoft-UUID="BA77FE2C3D8953E49212BBE221185CC1" ID-ZooBank="A5C1D3DFFF87404A9EA04608CF89EFA7" ModsDocID="1313-2970-1175-223" checkinTime="1692351780866" checkinUser="pensoft" docAuthor="Villarreal, Osvaldo, Ahumada-C., Daniela &amp; Delgado-Santa, Leonardo" docDate="2023" docId="F4102D81CBD05AB58DDDA6E2F419A8FB" docLanguage="en" docName="ZooKeys 1175: 223-284" docOrigin="ZooKeys 1175" docPubDate="2023-08-17" docSource="http://dx.doi.org/10.3897/zookeys.1175.102485" docTitle="Phalangodus andresi Villarreal &amp; Ahumada-C. &amp; Delgado-Santa 2023, sp. nov." docType="treatment" docUuid="B451F03A-F855-4F5B-8E90-8BDAF7890A11" docUuidSource="ZooBank" docVersion="1" id="BA77FE2C3D8953E49212BBE221185CC1" lastPageNumber="223" masterDocId="BA77FE2C3D8953E49212BBE221185CC1" masterDocTitle="Mapping the distribution of armored harvestmen (Opiliones, Laniatores) in Colombia: updated list of species, taxonomic contributions, and insight of diversity in protected areas" masterLastPageNumber="284" masterPageNumber="223" pageNumber="223" updateTime="1692351780866" updateUser="pensoft">
<mods:mods id="D23B717CE73CAA8D8E3E6240BA6CAA4D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="61323162070E6ECBB0532A3E27329DAD">
<mods:title id="E61A42BCDDBD34D92B22757B50B895EE">Mapping the distribution of armored harvestmen (Opiliones, Laniatores) in Colombia: updated list of species, taxonomic contributions, and insight of diversity in protected areas</mods:title>
</mods:titleInfo>
<mods:name id="43C0062ED47EFD8274A557052F88E508" type="personal">
<mods:role id="CD6C9EA9B8A21AC5D0E01A42ACD86DD6">
<mods:roleTerm id="37C44107AF4593AF7FD20B2A6D9CE075">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7B44A91ED37B838FA3A310603B8A5C85">Villarreal, Osvaldo</mods:namePart>
<mods:nameIdentifier id="9BE287A52D63F761E7440A7EC41CFA97" type="ORCID">https://orcid.org/0000-0001-5355-3723</mods:nameIdentifier>
<mods:affiliation id="B9D01ACC2B8109ED77C74B2C35632AEA">Departamento de Invertebrados, Museu Nacional / Universidade Federal do Rio de Janeiro, Quinta da Boa Vista, Sao Cristovao, 20.940 - 040, Rio de Janeiro, RJ, Brazil &amp; Museo del Instituto de Zoologia Agricola, Facultad de Agronomia, Universidad Central de Venezuela, Maracay, Aragua, Venezuela</mods:affiliation>
<mods:nameIdentifier id="E3DA69DB93D69DABA9E8CCCFEDC465D4" type="email">osvaldovillarreal@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="816BD15E8324000EA1EB5F70490ACF86" type="personal">
<mods:role id="EEA1EB8B3781EFB8F02C08B0050F95C5">
<mods:roleTerm id="3616D85C23FD5D64E8A8A461E515630E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3B0D2B75AFF586611DC9FFF96521FDB3">Ahumada-C., Daniela</mods:namePart>
<mods:nameIdentifier id="68A6F8F5CFFF463CFC2E034D0463870E" type="ORCID">https://orcid.org/0000-0002-4182-5143</mods:nameIdentifier>
<mods:affiliation id="81D5FB87C5B969808A9CBC9EF1FD2712">Grupo de Investigacion Biologia Descriptiva y Aplicada, Programa de Biologia, Universidad de Cartagena, Cartagena de Indias, Bolivar, Colombia</mods:affiliation>
</mods:name>
<mods:name id="343CD92F8041419EC722CFC8010D7358" type="personal">
<mods:role id="1130CBEC04024E23F5CCBC8371DDE786">
<mods:roleTerm id="43CEC530D4D12E11D37AF8764B585F44">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="619D6438832A130F559B0914E2CEC647">Delgado-Santa, Leonardo</mods:namePart>
<mods:nameIdentifier id="595DFA4E53BD0BE409C9DB7ED720C92E" type="ORCID">https://orcid.org/0000-0002-9714-6924</mods:nameIdentifier>
<mods:affiliation id="EEB0187A6654D7292D9DCDCEB0EA40CF">Grupo de Investigacion Hidrobiologia, Programa de Biologia, Universidad de Cartagena, Cartagena de Indias, Bolivar, Colombia</mods:affiliation>
<mods:nameIdentifier id="CDA7910BD2CC0388947D27DD2BC9849E" type="email">leonardodelgadosanta@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="69DB722CA402F9C214372EEC62D9972F">text</mods:typeOfResource>
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<mods:title id="FC5854FFDA0FD6206D2BAD13852F9814">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="6A80B324DF37058EA1D217DE703E9072">
<mods:date id="8FC8CCD86FED9D7811DE33A0F9F1F99A">2023</mods:date>
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<mods:number id="81EEA8B376B8E0EBA0EB1424EBFF3C22">2023-08-17</mods:number>
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<mods:detail id="3DB6068AE042BF72A18140D73E6DD790" type="volume">
<mods:number id="2F12D3CD86E41EAECE72CECF6E668729">1175</mods:number>
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<mods:start id="946EA421D71EDCF6E0AF0ECA7794783E">223</mods:start>
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<mods:url id="8CFBA9E25635F962124BD44BC091F9D4">http://dx.doi.org/10.3897/zookeys.1175.102485</mods:url>
</mods:location>
<mods:classification id="D24C2EDB20FB8B0E1D36E469A8A167FD">journal article</mods:classification>
<mods:identifier id="CD2B654AE5915F809C285CBB057B3A2E" type="DOI">http://dx.doi.org/10.3897/zookeys.1175.102485</mods:identifier>
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<treatment id="F4102D81CBD05AB58DDDA6E2F419A8FB" LSID="urn:lsid:zoobank.org:act:B451F03A-F855-4F5B-8E90-8BDAF7890A11" httpUri="http://treatment.plazi.org/id/F4102D81CBD05AB58DDDA6E2F419A8FB" lastPageNumber="223" pageId="0" pageNumber="223">
<subSubSection id="A36697AACE83A77BD0C65782DBCC8ED0" pageId="0" pageNumber="223" type="nomenclature">
<paragraph id="2FE4CC85DA8831EEDA27711A0D1B6B7B" pageId="0" pageNumber="223">
<taxonomicName id="C4A020CC03A6E87F67F643522CD19F85" LSID="https://zoobank.org/B451F03A-F855-4F5B-8E90-8BDAF7890A11" authority="Villarreal &amp; Ahumada-C. &amp; Delgado-Santa, 2023" authorityName="Villarreal &amp; Ahumada-C. &amp; Delgado-Santa" authorityYear="2023" class="Arachnida" family="Cranaidae" genus="Phalangodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phalangodus andresi" order="Opiliones" pageId="0" pageNumber="223" phylum="Arthropoda" rank="species" species="andresi" status="sp. nov.">Phalangodus andresi</taxonomicName>
<taxonomicNameLabel id="7C671544E0E014AB6AB4A4C64D65C597" pageId="0" pageNumber="223">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="A38E0F5FB8DA99A35BC87ECE41CD2977" pageId="0" pageNumber="223" type="description">
<paragraph id="C2BD248275AE52A7F97F0B84191E99B0" pageId="0" pageNumber="223">
<figureCitation id="1CB1771EDB7E702AE53E7BC2757E2ED5" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phalangodus andresi sp. nov. Male holotype (ICN-Ao- 1908) A habitus, dorsal view B lateral view. Female paratype (ICN-Ao- 1909): C dorsal view D lateral view. Scale bars: 3 mm." figureDoi="10.3897/zookeys.1175.102485.figure1" httpUri="https://binary.pensoft.net/fig/893703" pageId="0" pageNumber="223">Figs 1</figureCitation>
<figureCitation id="B64745A280DCD5E6AA1A1D3141994477" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phalangodus andresi sp. nov. Male holotype (ICN-Ao- 1908) A chelicera, frontal view B right palp, femur, ectal view D right leg IV: femur distal portion and patella, prolateral view E ditto, retrolateral view F ditto, ventral view C female, paratype (ICN-Ao- 1909), right palp, femur, ectal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1175.102485.figure2" httpUri="https://binary.pensoft.net/fig/893704" pageId="0" pageNumber="223">, 2</figureCitation>
<figureCitation id="294F2E374B4C6545C2B77A2E4D1D19B5" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Phalangodus andresi sp. nov. Male holotype (ICN-Ao- 1908) A habitus, dorsal view B habitus, lateral view C chelicera, frontal view D right pedipalp: trochanter, femur, and patella, ectal view E ditto, tibia, tarsus and claw, dorsoectal view F ditto, mesal view G right leg IV, trochanter and femur, dorsal view H ditto, femur, distal portion, prolateral view I ditto, ventral view J right leg IV, patella and tibia, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1175.102485.figure3" httpUri="https://binary.pensoft.net/fig/893705" pageId="0" pageNumber="223">, 3</figureCitation>
<figureCitation id="4D1BD498A8C3121C6700E0A98B523EFD" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Phalangodus andresi sp. nov. Male paratype (ICN-Ao- 1003). Penis, apical portion A dorsal view B lateral view C ventral view. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.1175.102485.figure4" httpUri="https://binary.pensoft.net/fig/893706" pageId="0" pageNumber="223">, 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="F5C7CAF182A8CDF4B996CB4F1042067F" pageId="0" pageNumber="223" type="materials_examined">
<paragraph id="61B8D4D10FA07AEA34F6193B0C6588FA" pageId="0" pageNumber="223">Material examined.</paragraph>
<paragraph id="7DCEAE5FD006006392D71AFF89CE6309" pageId="0" pageNumber="223">
<materialsCitation id="1A334A00C82A4302F49055C0C101F762" collectingDate="2018-06-03" collectorName="Tequendama, R. N., A. Garcia, S. Galvis" country="Colombia" elevation="2250" latitude="4.562234" location="San Antonio del Tequendama" longLatPrecision="1" longitude="-74.31453" specimenCount="♂" stateProvince="Cundinamarca" typeStatus="Holotype">
•
<emphasis id="D33E61815B1CC8F6055F36BCB683377E" bold="true" italics="true" pageId="0" pageNumber="223">
<typeStatus id="AAC7274B467587690A6ECFB499D269AA">Holotype</typeStatus>
</emphasis>
:
<specimenCount id="677EF0ABFDFC7C445B6CEE82827B1C51">♂</specimenCount>
(ICN-Ao-1908),
<collectingCountry id="58D59B43DAC227FEE24EA37F82414137" name="Colombia">Colombia</collectingCountry>
,
<collectingRegion id="98AF0506BE05E52D386E79859C136002" country="Colombia" name="Cundinamarca">Cundinamarca</collectingRegion>
,
<location id="96A23EA6E1698B8094B41D634E2AC3FA" LSID="urn:lsid:plazi:treatment:F4102D81CBD05AB58DDDA6E2F419A8FB:96A23EA6E1698B8094B41D634E2AC3FA" country="Colombia" latitude="4.562234" longLatPrecision="1" longitude="-74.31453" name="San Antonio del Tequendama" stateProvince="Cundinamarca">
San Antonio del
<collectorName id="913D6C1F84BAB2CC64AF58318B7295BE">Tequendama, R.N.</collectorName>
</location>
Los Tunos (
<geoCoordinate id="A1C9CA30658C5F4B0B0351CDBB9DB689" degrees="4.562234" direction="north" orientation="latitude" precision="1" value="4.562234">4.562234</geoCoordinate>
,
<geoCoordinate id="CE17F31A49382E0BA7EA601269877F5F" degrees="74.314527" direction="west" orientation="longitude" precision="1" value="-74.31453">-74.314527</geoCoordinate>
);
<quantity id="8DA0EDB4557AB62B50C2D12997334CA5" metricMagnitude="3" metricUnit="m" metricValue="2.25" unit="m" value="2250.0">
<elevation id="1A82AB44275E67097CA4F5B9A7167173" metricMagnitude="3" metricUnit="m" metricValue="2.25" unit="m" value="2250.0">2,250 m</elevation>
</quantity>
;
<collectingDate id="E51B9820F86C942C893EE5F05C70F4ED" value="2018-06-03">3 Jun. 2018</collectingDate>
; (
<collectorName id="3BAB56F8DABDAC37FDAEE57C4744F50E">
A.
<normalizedToken id="D3BD229089E06CCCB73A0CB5D531FDA8" originalValue="García">Garcia</normalizedToken>
</collectorName>
,
<collectorName id="F3EC052145005D11B30179B4CD5DB11E">S. Galvis</collectorName>
leg.)
</materialsCitation>
. •
<materialsCitation id="48825395E881C41F588AE604F6A15B1C" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="Tequendama, R. N. &amp; A. Garcia &amp; S. Galvis" country="Colombia" elevation="2250" latitude="4.562234" location="San Antonio del Tequendama" longLatPrecision="1" longitude="-74.31453" specimenCount="3" specimenCount-female="3" stateProvince="Cundinamarca" typeStatus="Paratypes">
<emphasis id="7B93A71FF70E59F3C5BFFB62E9C46BB1" bold="true" italics="true" pageId="0" pageNumber="223">
<typeStatus id="0DADFCB07B1643294F76FADF72D123D8">Paratypes</typeStatus>
</emphasis>
: •
<specimenCount id="382246BF679D3791D419E3A819C2B67D" type="female">3 ♀♀</specimenCount>
(ICN-Ao-1909, with one used for description), same data as the holotype
</materialsCitation>
; •
<materialsCitation id="C78568A022AD8826FB2574DF8DB3D5D7" collectingDate="2006-08-28" collectorName="Tequendama, R. N., F. Borrero" country="Colombia" location="San Antonio del Tequendama" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Cundinamarca" typeStatus="paratype">
<specimenCount id="A86E0511CF9F7EBAC0665498B3F86F36" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="60AE3EE1771C5E6493D832C09E4D2A23" type="female">1 ♀</specimenCount>
(ICN-Ao-1003),
<collectingCountry id="DC5D4EC35BF87BA21D0228C8E4D4AD79" name="Colombia">Colombia</collectingCountry>
,
<collectingRegion id="0956F5BDF79FDDB7B94FEDBE7158AB24" country="Colombia" name="Cundinamarca">Cundinamarca</collectingRegion>
,
<location id="60E73B3157E60430C5BEE2A652A09767" LSID="urn:lsid:plazi:treatment:F4102D81CBD05AB58DDDA6E2F419A8FB:60E73B3157E60430C5BEE2A652A09767" country="Colombia" name="San Antonio del Tequendama" stateProvince="Cundinamarca">
San Antonio del
<collectorName id="2847309E0D479335C34961E45C27A1CF">Tequendama, R.N.</collectorName>
</location>
Los Tunos;
<collectingDate id="86167C021695356E076CC3AC5930D4A1" value="2006-08-28">28 Aug. 2006</collectingDate>
; (
<collectorName id="D83CEAF537C77005CBC63574AF637E57">F. Borrero</collectorName>
leg.)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="6972642E85F85E681D3EE02EFA0F1F4A" pageId="0" pageNumber="223" type="diagnosis">
<paragraph id="1A93EDA87EBC4D9D6D27A6EF8C0ADB25" pageId="0" pageNumber="223">Diagnosis.</paragraph>
<paragraph id="6AB172E3A123919CA0A7B0BA78A07642" pageId="0" pageNumber="223">
<taxonomicName id="BC680E556C9A0BBB985FC63A2AC52A3A" authorityName="Villarreal &amp; Ahumada-C. &amp; Delgado-Santa" authorityYear="2023" class="Arachnida" family="Cranaidae" genus="Phalangodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phalangodus andresi" order="Opiliones" pageId="0" pageNumber="223" phylum="Arthropoda" rank="species" species="andresi">
<emphasis id="8A10ABEA787B1828B6056F8B9AF15362" italics="true" pageId="0" pageNumber="223">Phalangodus andresi</emphasis>
</taxonomicName>
sp. nov. can be distinguished from all other species of the genus except
<taxonomicName id="2A81025240DC300A47C586112919EFD9" class="Arachnida" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="P. palpiconus" pageId="0" pageNumber="223" phylum="Arthropoda" rank="species" species="palpiconus">
<emphasis id="A60F3C89EF177BD6EF9A969CB97C385C" italics="true" pageId="0" pageNumber="223">P. palpiconus</emphasis>
</taxonomicName>
by the (1) presence of conspicuous granulation of mesotergal areas I -IV, lateral borders of dorsal scutum, ocularium and posterior region of the carapace (Figs
<figureCitation id="4A9A87D806C2D8F961850F93CF030A24" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phalangodus andresi sp. nov. Male holotype (ICN-Ao- 1908) A habitus, dorsal view B lateral view. Female paratype (ICN-Ao- 1909): C dorsal view D lateral view. Scale bars: 3 mm." figureDoi="10.3897/zookeys.1175.102485.figure1" httpUri="https://binary.pensoft.net/fig/893703" pageId="0" pageNumber="223">1A, B</figureCitation>
,
<figureCitation id="273DC631BFB0ED497DB581CD93FC8A59" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Phalangodus andresi sp. nov. Male holotype (ICN-Ao- 1908) A habitus, dorsal view B habitus, lateral view C chelicera, frontal view D right pedipalp: trochanter, femur, and patella, ectal view E ditto, tibia, tarsus and claw, dorsoectal view F ditto, mesal view G right leg IV, trochanter and femur, dorsal view H ditto, femur, distal portion, prolateral view I ditto, ventral view J right leg IV, patella and tibia, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1175.102485.figure3" httpUri="https://binary.pensoft.net/fig/893705" pageId="0" pageNumber="223">3A, B</figureCitation>
); (2) small size (males DSL ~ 10.7-11.9 mm), except
<taxonomicName id="F708C70A4ACC02CF3B1530DDA7C38F45" class="Arachnida" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="P. kuryi" pageId="0" pageNumber="223" phylum="Arthropoda" rank="species" species="kuryi">
<emphasis id="8FA648D0219CF5709EF40F64D48074E1" italics="true" pageId="0" pageNumber="223">P. kuryi</emphasis>
</taxonomicName>
(9.0-11.4 mm). It can be distinguished from the latter species by the ornamentation of the pedipalpal femur (with a very large ventroproximal tubercle absent in
<taxonomicName id="753AD39943995D55C803FD2BB2848BAC" class="Arachnida" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="P. kuryi" pageId="0" pageNumber="223" phylum="Arthropoda" rank="species" species="kuryi">
<emphasis id="22C5AA67397EB9D66D9EECD9B77EC50F" italics="true" pageId="0" pageNumber="223">P. kuryi</emphasis>
</taxonomicName>
) (Fig.
<figureCitation id="3BBD0B352E5F22E501743D1DE6D2CF06" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Phalangodus andresi sp. nov. Male holotype (ICN-Ao- 1908) A habitus, dorsal view B habitus, lateral view C chelicera, frontal view D right pedipalp: trochanter, femur, and patella, ectal view E ditto, tibia, tarsus and claw, dorsoectal view F ditto, mesal view G right leg IV, trochanter and femur, dorsal view H ditto, femur, distal portion, prolateral view I ditto, ventral view J right leg IV, patella and tibia, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1175.102485.figure3" httpUri="https://binary.pensoft.net/fig/893705" pageId="0" pageNumber="223">3D</figureCitation>
), and the presence of ornamentation in leg IV of the males (Fig.
<figureCitation id="3C1914BA28871595A6244E6B5D907AEA" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Phalangodus andresi sp. nov. Male holotype (ICN-Ao- 1908) A habitus, dorsal view B habitus, lateral view C chelicera, frontal view D right pedipalp: trochanter, femur, and patella, ectal view E ditto, tibia, tarsus and claw, dorsoectal view F ditto, mesal view G right leg IV, trochanter and femur, dorsal view H ditto, femur, distal portion, prolateral view I ditto, ventral view J right leg IV, patella and tibia, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1175.102485.figure3" httpUri="https://binary.pensoft.net/fig/893705" pageId="0" pageNumber="223">3G-J</figureCitation>
) (absent in
<taxonomicName id="9AE4861F5AF3983D3ECE82629B613C16" class="Arachnida" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="P. kuryi" pageId="0" pageNumber="223" phylum="Arthropoda" rank="species" species="kuryi">
<emphasis id="7BCF682831639A3F7B4B2A04A605FA2F" italics="true" pageId="0" pageNumber="223">P. kuryi</emphasis>
</taxonomicName>
), as well as the interocular distance, height of the ocularium and presence of paired tubercles near each eye (absent in
<taxonomicName id="C4E0754818663817D15C78D61AEA4DEA" class="Arachnida" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="P. kuryi" pageId="0" pageNumber="223" phylum="Arthropoda" rank="species" species="kuryi">
<emphasis id="534BD99C565AD2FAE1127290EF509B58" italics="true" pageId="0" pageNumber="223">P. kuryi</emphasis>
</taxonomicName>
). From
<taxonomicName id="8162FCEDF1BC4FE417BDFA47291B399C" class="Arachnida" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="P. palpiconus" pageId="0" pageNumber="223" phylum="Arthropoda" rank="species" species="palpiconus">
<emphasis id="6A3EF0A7F07E441CC5384F5E9C4EAD0C" italics="true" pageId="0" pageNumber="223">P. palpiconus</emphasis>
</taxonomicName>
, the most morphologically similar species, it is distinguished by the ornamentation of the femur IV of the males, having a subdistal large and curved spine and a short distal bifid tubercle (Figs
<figureCitation id="00716717722C5EE9414746D8E6B1ED9B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phalangodus andresi sp. nov. Male holotype (ICN-Ao- 1908) A chelicera, frontal view B right palp, femur, ectal view D right leg IV: femur distal portion and patella, prolateral view E ditto, retrolateral view F ditto, ventral view C female, paratype (ICN-Ao- 1909), right palp, femur, ectal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1175.102485.figure2" httpUri="https://binary.pensoft.net/fig/893704" pageId="0" pageNumber="223">2E, F</figureCitation>
,
<figureCitation id="72F578C2554846B3EEA9C040C85E4F7A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Phalangodus andresi sp. nov. Male holotype (ICN-Ao- 1908) A habitus, dorsal view B habitus, lateral view C chelicera, frontal view D right pedipalp: trochanter, femur, and patella, ectal view E ditto, tibia, tarsus and claw, dorsoectal view F ditto, mesal view G right leg IV, trochanter and femur, dorsal view H ditto, femur, distal portion, prolateral view I ditto, ventral view J right leg IV, patella and tibia, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1175.102485.figure3" httpUri="https://binary.pensoft.net/fig/893705" pageId="0" pageNumber="223">3H, I</figureCitation>
) on the prolateral face (absent in
<taxonomicName id="C6DDE03C5298E47DA297C8B7B3424E5F" class="Arachnida" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="P. palpiconus" pageId="0" pageNumber="223" phylum="Arthropoda" rank="species" species="palpiconus">
<emphasis id="676895CC2BDDA5B351B11D6F0FA981BD" italics="true" pageId="0" pageNumber="223">P. palpiconus</emphasis>
</taxonomicName>
); lack of a retrolateral subdistal spine in the same segment (present in
<taxonomicName id="6F90F8483E4ED498C8E7297DAAE08E68" class="Arachnida" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="P. palpiconus" pageId="0" pageNumber="223" phylum="Arthropoda" rank="species" species="palpiconus">
<emphasis id="2921AF36AEC94ECA343D372E85BE0F19" italics="true" pageId="0" pageNumber="223">P. palpiconus</emphasis>
</taxonomicName>
(
<bibRefCitation id="4DD8851BD3A266EE1B168ACEB94F56CD" DOI="https://doi.org/10.11646/zootaxa.3814.4.8" author="Hara, MR" journalOrPublisher="Zootaxa" pageId="0" pageNumber="223" pagination="567 - 580" refId="B25" refString="Hara, MR, Pinto-da-Rocha, R, Villarreal, O, 2014. Revision of the cranaid genera Phalangodus, Iquitosa and Aguaytiella (Opiliones: Laniatores: Gonyleptoidea). Zootaxa 3814 (4): 567 - 580, DOI: https://doi.org/10.11646/zootaxa.3814.4.8" title="Revision of the cranaid genera Phalangodus, Iquitosa and Aguaytiella (Opiliones: Laniatores: Gonyleptoidea)." url="https://doi.org/10.11646/zootaxa.3814.4.8" volume="3814" year="2014">Hara et al. 2014</bibRefCitation>
: figs 4, 5)); the shape of the VP of the penis, more elongated and with more marked medial constrictions, and the more basal position of MS-A/B groups.
</paragraph>
<caption id="E320300322202EB98E4F3D82A3F0E304" doi="10.3897/zookeys.1175.102485.figure1" httpUri="https://binary.pensoft.net/fig/893703" pageId="0" pageNumber="223" start="Figure 1" startId="F1">
<paragraph id="1D1613598EBD24CF1819F9B20322C5EF" pageId="0" pageNumber="223">
<emphasis id="8459A1FF60389FD099DA47EABAC44776" bold="true" pageId="0" pageNumber="223">Figure 1.</emphasis>
<taxonomicName id="4191797A09C3A827FA0C18A75AB9B3AC" authorityName="Villarreal &amp; Ahumada-C. &amp; Delgado-Santa" authorityYear="2023" class="Arachnida" family="Cranaidae" genus="Phalangodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phalangodus andresi" order="Opiliones" pageId="0" pageNumber="223" phylum="Arthropoda" rank="species" species="andresi">
<emphasis id="20B634F1B1881C3895E8B2F8DFE41C38" italics="true" pageId="0" pageNumber="223">Phalangodus andresi</emphasis>
</taxonomicName>
sp. nov. Male holotype (ICN-Ao-1908)
<emphasis id="C3379B2B1C986AD0FE51D7A310D09D61" bold="true" pageId="0" pageNumber="223">A</emphasis>
habitus, dorsal view
<emphasis id="5F6EBCF26FABA228792E21BF407D963A" bold="true" pageId="0" pageNumber="223">B</emphasis>
lateral view. Female paratype (ICN-Ao-1909):
<emphasis id="16B3F9BF22E11E051787C34C4D8D2153" bold="true" pageId="0" pageNumber="223">C</emphasis>
dorsal view
<emphasis id="2BFE4BE507EF6DA3303163F389154C32" bold="true" pageId="0" pageNumber="223">D</emphasis>
lateral view. Scale bars: 3 mm.
</paragraph>
</caption>
<caption id="6B4AA6EFC82F294F97054138160C03A6" doi="10.3897/zookeys.1175.102485.figure2" httpUri="https://binary.pensoft.net/fig/893704" pageId="0" pageNumber="223" start="Figure 2" startId="F2">
<paragraph id="23D915A075A1AB1AEF2EF496168B9243" pageId="0" pageNumber="223">
<emphasis id="CADA01DED8B5EFA4DE67527D6A425D8E" bold="true" pageId="0" pageNumber="223">Figure 2.</emphasis>
<taxonomicName id="91C21EE8DEE220F028E688E8F50D3D28" authorityName="Villarreal &amp; Ahumada-C. &amp; Delgado-Santa" authorityYear="2023" class="Arachnida" family="Cranaidae" genus="Phalangodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phalangodus andresi" order="Opiliones" pageId="0" pageNumber="223" phylum="Arthropoda" rank="species" species="andresi">
<emphasis id="FBED4916671E1721B8EAA6BA4EC37108" italics="true" pageId="0" pageNumber="223">Phalangodus andresi</emphasis>
</taxonomicName>
sp. nov. Male holotype (ICN-Ao-1908)
<emphasis id="8EB1D35B5095A71764BDDDE00E380772" bold="true" pageId="0" pageNumber="223">A</emphasis>
chelicera, frontal view
<emphasis id="6D6C42C255C10BCD62B12F86A5260BF2" bold="true" pageId="0" pageNumber="223">B</emphasis>
right palp, femur, ectal view
<emphasis id="D0E7805645DA0D6B6AA6840157747105" bold="true" pageId="0" pageNumber="223">D</emphasis>
right leg IV: femur distal portion and patella, prolateral view
<emphasis id="29A05BC83DF62C6A09A5E65EF6A17F49" bold="true" pageId="0" pageNumber="223">E</emphasis>
ditto, retrolateral view
<emphasis id="8A158569EF2E3DA7A5DA609EC42DE3D0" bold="true" pageId="0" pageNumber="223">F</emphasis>
ditto, ventral view
<emphasis id="764CEB723187B80388E4A0AF84DD4A72" bold="true" pageId="0" pageNumber="223">C</emphasis>
female, paratype (ICN-Ao-1909), right palp, femur, ectal view. Scale bars: 1 mm.
</paragraph>
</caption>
<caption id="23D789B7DAE342C067807430D1AC5B40" doi="10.3897/zookeys.1175.102485.figure3" httpUri="https://binary.pensoft.net/fig/893705" pageId="0" pageNumber="223" start="Figure 3" startId="F3">
<paragraph id="16150431DA51FA21B7D5CCD441C6947D" pageId="0" pageNumber="223">
<emphasis id="85255DEC91F6325562CF9E8B3E9B0E56" bold="true" pageId="0" pageNumber="223">Figure 3.</emphasis>
<taxonomicName id="0EF17FB57CF8BF5F5A955F91B10B1003" authorityName="Villarreal &amp; Ahumada-C. &amp; Delgado-Santa" authorityYear="2023" class="Arachnida" family="Cranaidae" genus="Phalangodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phalangodus andresi" order="Opiliones" pageId="0" pageNumber="223" phylum="Arthropoda" rank="species" species="andresi">
<emphasis id="35818826871810EAE83A9C7CFCC465BC" italics="true" pageId="0" pageNumber="223">Phalangodus andresi</emphasis>
</taxonomicName>
sp. nov. Male holotype (ICN-Ao-1908)
<emphasis id="05798C59F1B23417C61F3E5EDA544071" bold="true" pageId="0" pageNumber="223">A</emphasis>
habitus, dorsal view
<emphasis id="7EC436F44C425A148100EF098ABB0378" bold="true" pageId="0" pageNumber="223">B</emphasis>
habitus, lateral view
<emphasis id="3A1BBDA06ADE6C04C19AA671876AF274" bold="true" pageId="0" pageNumber="223">C</emphasis>
chelicera, frontal view
<emphasis id="E45718AB8F3A73BD037A5CBBCA9ABB37" bold="true" pageId="0" pageNumber="223">D</emphasis>
right pedipalp: trochanter, femur, and patella, ectal view
<emphasis id="369EC830409F8588E5F839A1DF4A181B" bold="true" pageId="0" pageNumber="223">E</emphasis>
ditto, tibia, tarsus and claw, dorsoectal view
<emphasis id="678A1DAE64EC1DD0E88478744FAE78A1" bold="true" pageId="0" pageNumber="223">F</emphasis>
ditto, mesal view
<emphasis id="1D666E40E732FA3D573838D93CAB7454" bold="true" pageId="0" pageNumber="223">G</emphasis>
right leg IV, trochanter and femur, dorsal view
<emphasis id="E5A7AF4F452A87F34470BA5A9143CABD" bold="true" pageId="0" pageNumber="223">H</emphasis>
ditto, femur, distal portion, prolateral view
<emphasis id="D10099E189F5C4DA7B727EF105FC7B75" bold="true" pageId="0" pageNumber="223">I</emphasis>
ditto, ventral view
<emphasis id="96D312F0D1EF7E340F6E572CD1B611B2" bold="true" pageId="0" pageNumber="223">J</emphasis>
right leg IV, patella and tibia, ventral view. Scale bars: 1 mm.
</paragraph>
</caption>
<caption id="DAD26C7936263C22BECFD56591C6BFE6" doi="10.3897/zookeys.1175.102485.figure4" httpUri="https://binary.pensoft.net/fig/893706" pageId="0" pageNumber="223" start="Figure 4" startId="F4">
<paragraph id="8DC7415F707E2AE689696895766996D9" pageId="0" pageNumber="223">
<emphasis id="72FD7813224FA37809C4CAD6FAB8D1C7" bold="true" pageId="0" pageNumber="223">Figure 4.</emphasis>
<taxonomicName id="C0548C91161623F55FFBE95CBEFDC89A" authorityName="Villarreal &amp; Ahumada-C. &amp; Delgado-Santa" authorityYear="2023" class="Arachnida" family="Cranaidae" genus="Phalangodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phalangodus andresi" order="Opiliones" pageId="0" pageNumber="223" phylum="Arthropoda" rank="species" species="andresi">
<emphasis id="EFB295BB01837967F3766176BE3968E8" italics="true" pageId="0" pageNumber="223">Phalangodus andresi</emphasis>
</taxonomicName>
sp. nov. Male paratype (ICN-Ao-1003). Penis, apical portion
<emphasis id="F1A572A43F142386DD438E3F7DCDA112" bold="true" pageId="0" pageNumber="223">A</emphasis>
dorsal view
<emphasis id="FCCF0C4FDEC6A85DB5961D6D441DA393" bold="true" pageId="0" pageNumber="223">B</emphasis>
lateral view
<emphasis id="8DE9723E9E5FF70ADD1D43EE561EF7EF" bold="true" pageId="0" pageNumber="223">C</emphasis>
ventral view. Scale bars: 0.25 mm.
</paragraph>
</caption>
<caption id="FC81EA73B23CD314B8FDA3E0E8CDA72A" doi="10.3897/zookeys.1175.102485.figure5" httpUri="https://binary.pensoft.net/fig/893707" pageId="0" pageNumber="223" start="Figure 5" startId="F5">
<paragraph id="39CB5C17DC6BC989625B19F0C46851FA" pageId="0" pageNumber="223">
<emphasis id="B4D67CD8ED12A0BC578B1A88C4B60EF5" bold="true" pageId="0" pageNumber="223">Figure 5.</emphasis>
<taxonomicName id="F4DE84DF1B8282A49074522795EC34F1" authorityName="Villarreal &amp; Ahumada-C. &amp; Delgado-Santa" authorityYear="2023" class="Arachnida" family="Cranaidae" genus="Ventrifurca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ventrifurca phallaina" order="Opiliones" pageId="0" pageNumber="223" phylum="Arthropoda" rank="species" species="phallaina">
<emphasis id="21B94255CBCD2C2988BBD00F1313FF65" italics="true" pageId="0" pageNumber="223">Ventrifurca phallaina</emphasis>
</taxonomicName>
sp. nov. Male holotype (CIUQ-020631)
<emphasis id="F6FEA72E48A318F2E8341A68AF547AD1" bold="true" pageId="0" pageNumber="223">A</emphasis>
dorsal view
<emphasis id="6C6783D616BA71C4B8A376FC944D82AD" bold="true" pageId="0" pageNumber="223">B</emphasis>
lateral view
<emphasis id="D06C29C6C17F5A49167A07BB600A2441" bold="true" pageId="0" pageNumber="223">C</emphasis>
lateral view
<emphasis id="452DE46BCB70C7656CC98581503137A8" bold="true" pageId="0" pageNumber="223">D</emphasis>
right chelicera, frontal view. Scale bars: 1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="9ED39A2EEE4597F180131A802DA61401" pageId="0" pageNumber="223" type="description">
<paragraph id="314590972E297152683F74F440E99DEB" pageId="0" pageNumber="223">Description.</paragraph>
<paragraph id="740BB19B3682890574D492944B516420" pageId="0" pageNumber="223">
<emphasis id="B86810C3B71EDF8FCC140B8971A8E3C5" bold="true" pageId="0" pageNumber="223">Measurements of body and appendages.</emphasis>
Holotype ♂ (ICN-Ao-1908): DSL = 10.8; CL = 4.5; AL = 6.3; CW = 5.6; AW = 7.4; IOD = 1.6; BaCh = 2.7; FePp = 2.1, PaPp = 1.1, TiPp = 1.6, TaPp = 1.3, ClPp = 1.1; FeL I: 6.0; FeL II = 11.4; FeL III = 8.9; FeL IV = 14.1; TiL I = 4.6; TiL II = 11.3; TiL III = 5.3; TiL IV = 8.2. • Paratypes: • (ICN-AO-1909, ICN-AO-1003, 4 ♀♀, min-max): DSL = 8.3-9.3; CL = 2.9-4.5; AL = 4.0-7.8; CW = 4.9-5.8; AW = 7.2-7.9; IOD = 1.4-2.0; BaCh = 1.4-1.9; FePp = 1.9, PaPp = 1.1, TiPp = 1.4, TaPp = 1.1, ClPp = 1.0; FeL I = 2.9-3.6; FeL II = 8.1-10.6; FeL III = 7.3-9.4; FeL IV = 8.9-14.4; TiL I = 3.8-4.5; TiL II = 7.9-9.5; TiL III = 4.9-5.7; TiL IV = 5.9-9.0. • (ICN-AO-1003; 2 ♂♂, min-max): DSL = 10.7-11.8; CL = 4.9-5.3; AL = 6.6-7.4; CW = 5.3-6.3; AW = 7.4-8.8; IOD = 1.9-2.1; BaCh = 2.6-3.5; FePp = 1.9-2.1, PaPp = 1.0-1.2, TiPp = 1.5-1.7, TaPp = 1.2-1.4, ClPp = 1.1; FeL I = 5.1-6.4; FeL II = 11.1-11.1; FeL III = 9.4-10.6; FeL IV = 12.2-16.8; TiL I = 4.3-5.0; TiL II = 10.2-10.7; TiL III = 5.6-9.3; TiL IV = 8.4-10.2.
</paragraph>
<paragraph id="00BBE1069E11CD895CA3F9D33E0DEE48" pageId="0" pageNumber="223">
<emphasis id="03A4867997C667B228E9787D0031D2DB" bold="true" pageId="0" pageNumber="223">
Male holotype (ICN-Ao-1908).
<emphasis id="8884F13D33E22B3D0A5D1C0BDC4F5DD0" italics="true" pageId="0" pageNumber="223">Dorsum</emphasis>
</emphasis>
(Figs
<figureCitation id="9659F9446B1097D2CA4FB3F2CA73F886" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phalangodus andresi sp. nov. Male holotype (ICN-Ao- 1908) A habitus, dorsal view B lateral view. Female paratype (ICN-Ao- 1909): C dorsal view D lateral view. Scale bars: 3 mm." figureDoi="10.3897/zookeys.1175.102485.figure1" httpUri="https://binary.pensoft.net/fig/893703" pageId="0" pageNumber="223">1A, B</figureCitation>
,
<figureCitation id="610C04D606DBD8E8D4F71B10A0EB3497" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Phalangodus andresi sp. nov. Male holotype (ICN-Ao- 1908) A habitus, dorsal view B habitus, lateral view C chelicera, frontal view D right pedipalp: trochanter, femur, and patella, ectal view E ditto, tibia, tarsus and claw, dorsoectal view F ditto, mesal view G right leg IV, trochanter and femur, dorsal view H ditto, femur, distal portion, prolateral view I ditto, ventral view J right leg IV, patella and tibia, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1175.102485.figure3" httpUri="https://binary.pensoft.net/fig/893705" pageId="0" pageNumber="223">3A, B</figureCitation>
). DS outline type alpha, with bulge longitudinally asymmetric widest at scutal groove II, lateral borders with granules only on the middle region. Carapace with few granules on the anterolateral and posterior region. Ocularium high, without median depression, with a paramedian pair of sharp tubercles and granules close to the eyes. Integumentary dome of ozopore raised and conspicuous. Abdominal scutum well delimited, divided into four well-marked scutal areas; I divided into left and right halves by invasion of the scutal area II; I and II granulated, with a pair of conspicuous medial tubercles, one tubercle on each side; III with a pair of paramedian acuminate spines and densely granulated; IV divided, with a row of six or seven granules on each side. Posterior border of the DS slightly convex and with a row of granules. Free tergites I-III with a row of granules.
</paragraph>
<paragraph id="7CA6DDF59A226478DAD9DB760600D571" pageId="0" pageNumber="223">
<emphasis id="A10F3D024DC9812F6BC2EFF3CE4FFBA4" bold="true" pageId="0" pageNumber="223">
<emphasis id="344C394AC2BBFA9AF5E7FC31DE566D73" italics="true" pageId="0" pageNumber="223">Venter</emphasis>
.
</emphasis>
Coxa I with a row of large tubercles of different sizes; II longer than I and III, with two median rows of low tubercles, the anterior one more conspicuous; III densely covered with irregular rows of small tubercles and with the posterior border sigmoid; IV strongly backward, with a median row of tubercles in the medium area and lateral border, and small tubercles densely distributed. Stigmatic area with a row of small tubercles on posterior border and minute granules sparsely distributed. Stigmata large, oval, and oblique. Free sternites with a row of small granules.
</paragraph>
<paragraph id="E458824EAF64E5F49AF7EA5E1F1A1887" pageId="0" pageNumber="223">
<emphasis id="D509FF00AC213E95FC22FBAC76F9D0BC" bold="true" italics="true" pageId="0" pageNumber="223">Chelicerae</emphasis>
(Figs
<figureCitation id="3D608E4197A5412D3FAC904AEE687735" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phalangodus andresi sp. nov. Male holotype (ICN-Ao- 1908) A chelicera, frontal view B right palp, femur, ectal view D right leg IV: femur distal portion and patella, prolateral view E ditto, retrolateral view F ditto, ventral view C female, paratype (ICN-Ao- 1909), right palp, femur, ectal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1175.102485.figure2" httpUri="https://binary.pensoft.net/fig/893704" pageId="0" pageNumber="223">2A</figureCitation>
,
<figureCitation id="0FD7B2D2C5D32043986AF69D9706C26F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Phalangodus andresi sp. nov. Male holotype (ICN-Ao- 1908) A habitus, dorsal view B habitus, lateral view C chelicera, frontal view D right pedipalp: trochanter, femur, and patella, ectal view E ditto, tibia, tarsus and claw, dorsoectal view F ditto, mesal view G right leg IV, trochanter and femur, dorsal view H ditto, femur, distal portion, prolateral view I ditto, ventral view J right leg IV, patella and tibia, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1175.102485.figure3" httpUri="https://binary.pensoft.net/fig/893705" pageId="0" pageNumber="223">3C</figureCitation>
). Segment I with well-defined bulla, with dorsomesal tubercles and a row of three large tubercles in the ectal region. Segment II swollen. Fixed finger with a proximal narrow and low tooth, a large gap and three subdistal median teeth. Movable finger with a proximal wide and low tooth, one large tooth and the distal inner surface irregularly dentate. Mesal side of the base of the fixed finger and near the base of the movable finger with setiferous tubercles of different sizes.
</paragraph>
<paragraph id="48D96C78F9B8338F5D027A5C3EDD8733" pageId="0" pageNumber="223">
<emphasis id="76372CEB850467077EF7CCFA00102075" bold="true" italics="true" pageId="0" pageNumber="223">Pedipalps</emphasis>
(Figs
<figureCitation id="C6D5EE993433070A18FDD3A69EC4B276" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phalangodus andresi sp. nov. Male holotype (ICN-Ao- 1908) A chelicera, frontal view B right palp, femur, ectal view D right leg IV: femur distal portion and patella, prolateral view E ditto, retrolateral view F ditto, ventral view C female, paratype (ICN-Ao- 1909), right palp, femur, ectal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1175.102485.figure2" httpUri="https://binary.pensoft.net/fig/893704" pageId="0" pageNumber="223">2B, C</figureCitation>
,
<figureCitation id="9811487BCDA207ACC12964B81BE5040B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Phalangodus andresi sp. nov. Male holotype (ICN-Ao- 1908) A habitus, dorsal view B habitus, lateral view C chelicera, frontal view D right pedipalp: trochanter, femur, and patella, ectal view E ditto, tibia, tarsus and claw, dorsoectal view F ditto, mesal view G right leg IV, trochanter and femur, dorsal view H ditto, femur, distal portion, prolateral view I ditto, ventral view J right leg IV, patella and tibia, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1175.102485.figure3" httpUri="https://binary.pensoft.net/fig/893705" pageId="0" pageNumber="223">3D-F</figureCitation>
). Trochanter with a dorsal pair of paramedian tubercles. Ventrally with large bifid tubercle in distal portion. Femur slightly compressed, dorsally curved, and ventrally straight in lateral view, with dorsoectal distal row of granules, one dorsal, and one ventral row of large forward projected tubercles (the apicalmost of the ventral row bifid and thicker than the remaining), the ventrodistal portion unarmed. Mesal and ectal faces without large tubercles. Patella short, cylindrical, and curved, with a dorsal row of short tubercles and small dorsodistal granules. Tibia dorsally granulated; tibia mesal IiIi (3&gt;1&gt;2=4), ectal IiiIi (4&gt;1&gt;5&gt;2&gt;3). Tarsus dorsally granulated, tibia with spines only in the distal portion, mesal Ii, ectal IiIiI (3&gt;1&gt;5&gt;4&gt;2). Claw proximally swollen.
</paragraph>
<paragraph id="AE02364892CBF0921BE6669E5FC2CA46" pageId="0" pageNumber="223">
<emphasis id="148B5F5F25F8E82F2AA73602F4263368" bold="true" italics="true" pageId="0" pageNumber="223">Legs</emphasis>
(Figs
<figureCitation id="244AB1D1601C272CEE02C5665F205D85" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phalangodus andresi sp. nov. Male holotype (ICN-Ao- 1908) A habitus, dorsal view B lateral view. Female paratype (ICN-Ao- 1909): C dorsal view D lateral view. Scale bars: 3 mm." figureDoi="10.3897/zookeys.1175.102485.figure1" httpUri="https://binary.pensoft.net/fig/893703" pageId="0" pageNumber="223">1A</figureCitation>
,
<figureCitation id="750FEC7AF0F3AB05E902E2426E8E2204" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phalangodus andresi sp. nov. Male holotype (ICN-Ao- 1908) A chelicera, frontal view B right palp, femur, ectal view D right leg IV: femur distal portion and patella, prolateral view E ditto, retrolateral view F ditto, ventral view C female, paratype (ICN-Ao- 1909), right palp, femur, ectal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1175.102485.figure2" httpUri="https://binary.pensoft.net/fig/893704" pageId="0" pageNumber="223">2D-F</figureCitation>
,
<figureCitation id="EA8084AE22CD37A72B88198D0F46B531" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Phalangodus andresi sp. nov. Male holotype (ICN-Ao- 1908) A habitus, dorsal view B habitus, lateral view C chelicera, frontal view D right pedipalp: trochanter, femur, and patella, ectal view E ditto, tibia, tarsus and claw, dorsoectal view F ditto, mesal view G right leg IV, trochanter and femur, dorsal view H ditto, femur, distal portion, prolateral view I ditto, ventral view J right leg IV, patella and tibia, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1175.102485.figure3" httpUri="https://binary.pensoft.net/fig/893705" pageId="0" pageNumber="223">3B</figureCitation>
,
<figureCitation id="963F3EC3CA341797D30334F72BFF79FB" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Phalangodus andresi sp. nov. Male holotype (ICN-Ao- 1908) A habitus, dorsal view B habitus, lateral view C chelicera, frontal view D right pedipalp: trochanter, femur, and patella, ectal view E ditto, tibia, tarsus and claw, dorsoectal view F ditto, mesal view G right leg IV, trochanter and femur, dorsal view H ditto, femur, distal portion, prolateral view I ditto, ventral view J right leg IV, patella and tibia, ventral view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1175.102485.figure3" httpUri="https://binary.pensoft.net/fig/893705" pageId="0" pageNumber="223">3G-J</figureCitation>
). Coxa I and III smooth; II with one dorsal tubercle; IV with one dorsodistal domed large tubercle, and a row of large tubercles in the prolateral border. Trochanters I-III unarmed; II with a prolateral row of small tubercles; III with one retrodistal tubercle and one group of small prolateral tubercles; IV with small prolateral granules sparsely distributed and one row of small tubercles in the retrolateral border. Femora I-III straight and with longitudinal rows of granules; IV sub-straight, densely granulated, ventrally with one large subdistal tubercle hook-like shaped and one distal rounded trifid tubercle. Ratio Fe IV/DSL = 1.85. Patellae I-IV granulated. Tibiae I-V straight and densely granulated, unarmed. Metatarsi I to IV with rows of small granules, I-III unarmed, IV with a ventro-distal border pair of spines. Tarsi III and IV with subparallel smooth claws and tarsal process. Tarsal counts: 8(3)-8(3), 13(3)-14(3), 8-8, 8-7.
</paragraph>
<paragraph id="2B667A4A8AA24874A6011F06E1CA0843" pageId="0" pageNumber="223">
<emphasis id="71D1E332F4A0ED76E6F9E824BBB8AC9B" bold="true" italics="true" pageId="0" pageNumber="223">Penis</emphasis>
(Fig.
<figureCitation id="0A4E4EEFEF9D23663DB2EC2B4CE9A075" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Phalangodus andresi sp. nov. Male paratype (ICN-Ao- 1003). Penis, apical portion A dorsal view B lateral view C ventral view. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.1175.102485.figure4" httpUri="https://binary.pensoft.net/fig/893706" pageId="0" pageNumber="223">4</figureCitation>
). VP subsquare, with proximal and distal portion delimited by a medial constriction, and lateral rounded lobes in the proximal portion; distal border slightly concave. Glans + Stylus columnar, glans with some folds at the base, stylus normally thickened, shorter than the glans, substraight; stylar caps ring-shaped. MS-A/B groups indistinguishable from each other, located on the proximal portion of the VP, with an increase in setae and composed of three to four pairs; MS-C/D groups located in the distal portion of the VP, separated from MS-A/B by a small gap, the setae rearranged into two irregular rows, one laterodistal and one mesodorsal. MS-E presumably present, since they are very small (as in other members of the genus), and were not observable under the magnification used.
</paragraph>
<paragraph id="9F38B623E0A50F4210E74FAEAFA73B99" pageId="0" pageNumber="223">
<emphasis id="D01EFD4C0DE4BC48E1E2C444F2E82C67" bold="true" italics="true" pageId="0" pageNumber="223">Coloration (in alcohol)</emphasis>
(Figs
<figureCitation id="26F035337479EE9EFD131D888BB3842F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phalangodus andresi sp. nov. Male holotype (ICN-Ao- 1908) A habitus, dorsal view B lateral view. Female paratype (ICN-Ao- 1909): C dorsal view D lateral view. Scale bars: 3 mm." figureDoi="10.3897/zookeys.1175.102485.figure1" httpUri="https://binary.pensoft.net/fig/893703" pageId="0" pageNumber="223">1</figureCitation>
,
<figureCitation id="1A2BA464AD4430E311135D72E65BE6DE" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Phalangodus andresi sp. nov. Male holotype (ICN-Ao- 1908) A chelicera, frontal view B right palp, femur, ectal view D right leg IV: femur distal portion and patella, prolateral view E ditto, retrolateral view F ditto, ventral view C female, paratype (ICN-Ao- 1909), right palp, femur, ectal view. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1175.102485.figure2" httpUri="https://binary.pensoft.net/fig/893704" pageId="0" pageNumber="223">2</figureCitation>
). Carapace, DS borders, posterior region and ocularium reticulated in Moderate brown 58, on background Light yellowish brown 76 (in females, it is Dark brown 59 on background Light brown 57). Abdominal scutum Light yellowish brown 76 (in females, it is moderate orange 53), with the scutal areas Moderate brown 58 (in females, it is Strong brown 55). Free tergites Dark grayish yellowish brown 81. Pedipalps, chelicerae and trochanters Strong brown 55 reticulated; remaining segment of the legs Strong brown 55 reticulated with fine mottled Deep orange 51. Stigmatic area Strong brown 55. Tip of cheliceral teeth Deep reddish brown 41.
</paragraph>
<paragraph id="611D3F7102B6819E65CC81268CC92EC3" pageId="0" pageNumber="223">
<emphasis id="D63FBD2673EDF348D72BE207BAA962FC" bold="true" pageId="0" pageNumber="223">Female (ICN-Ao-1909)</emphasis>
(Fig.
<figureCitation id="95FA0EEC9064C8BF43EF32AFF3517F6A" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phalangodus andresi sp. nov. Male holotype (ICN-Ao- 1908) A habitus, dorsal view B lateral view. Female paratype (ICN-Ao- 1909): C dorsal view D lateral view. Scale bars: 3 mm." figureDoi="10.3897/zookeys.1175.102485.figure1" httpUri="https://binary.pensoft.net/fig/893703" pageId="0" pageNumber="223">1C, D</figureCitation>
). Differing from male by: ocularium slightly narrower; carapace shorter; coda wider (DS outline type alpha-keyhole); tubercles of area III slightly higher. Chelicerae non-hypertelic, with movable finger thinner. Pedipalpal femur lower and thinner in lateral view. Stigmatic area shorter. Genital operculum wider. Trochanters III and IV narrower; femur IV thinner, without large ventral-subdistal tubercle with hook-like shape and without a ventrodistal rounded trifid tubercle. For color differences, see the color description of the male.
</paragraph>
<paragraph id="70F37044560C995F05C7ABE249C6B7C1" pageId="0" pageNumber="223">
<emphasis id="F8289BBA2EF0E6D0C0438B95D49DCA9D" bold="true" pageId="0" pageNumber="223">
<emphasis id="40C11E027759731A593586B30C53E814" italics="true" pageId="0" pageNumber="223">Ovipositor</emphasis>
.
</emphasis>
Dorsal lobes (dl) and ventral lobes (vl) rounded, with four and two pairs of large, acuminated, single-tipped setae, respectively. The dl with three pairs of dorsal setae (ds) distally located and one pair basally located. Lateral region of the ovipositor with one pair of dorso-lateral setae (dls).
</paragraph>
</subSubSection>
<subSubSection id="90CB72195AB34193F43864C44BFCCFA6" pageId="0" pageNumber="223" type="distribution">
<paragraph id="61324B8EA2AD11AE7A721A8F0D60B198" pageId="0" pageNumber="223">Distribution.</paragraph>
<paragraph id="84BE365249464ED88EDD5F62C2DD1009" pageId="0" pageNumber="223">Known only from the type locality.</paragraph>
</subSubSection>
<subSubSection id="CEE87DB73BCBE4CCCE96E6A262A85458" pageId="0" pageNumber="223" type="etymology">
<paragraph id="48BA1E79C1F62421E40A1DB5D04F0DAD" pageId="0" pageNumber="223">Etymology.</paragraph>
<paragraph id="DD2261FD69B4E6EC42FE8C88528DD5B2" pageId="0" pageNumber="223">
The species is named in honor of our colleague and friend, the arachnologist
<normalizedToken id="5BABDABBF6E95F2AF106F450A4DEB717" originalValue="Andrés">Andres</normalizedToken>
F.
<normalizedToken id="1C5188629B761B997A6B2111796D49F9" originalValue="García">Garcia</normalizedToken>
, who has greatly enriched the
<normalizedToken id="6A716CD6A37EAFCB58C4F387A33280E8" originalValue="field’s">field's</normalizedToken>
knowledge of
<taxonomicName id="696AA3172D4171C65AC1BD2A2CCDA8CF" authorityName="Roewer" authorityYear="1913" class="Arachnida" kingdom="Animalia" lsidName="" order="Opiliones" pageId="0" pageNumber="223" rank="order">Opiliones</taxonomicName>
in Colombia and described the vast majority of species within the genus
<taxonomicName id="744225A222481285851558E4DBFD5CAB" authorityName="Gervais" authorityYear="1842" class="Arachnida" family="Cranaidae" genus="Phalangodus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phalangodus" order="Opiliones" pageId="0" pageNumber="223" phylum="Arthropoda" rank="genus">
<emphasis id="CC7EE7B1C86ED29F6FBD227E776116AC" italics="true" pageId="0" pageNumber="223">Phalangodus</emphasis>
</taxonomicName>
; moreover, he was the collector of the type series for this species.
</paragraph>
</subSubSection>
</treatment>
</document>
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