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<document id="B6C531BA0966BB2AEE43F9EBCCC23740" ID-CLB-Dataset="27816" ID-DOI="10.5852/ejt.2019.508" ID-GBIF-Dataset="6e176ff1-c6ce-429c-8b36-9d32c3cdc5c9" ID-ISSN="2118-9773" ID-Zenodo-Dep="2612875" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1553714929206" checkinUser="plazi" docAuthor="Rahayu, Sri &amp; Rodda, Michele" docDate="2019" docId="6B7785212A5FFFD1284EC132FAFBFA3C" docLanguage="en" docName="ejt-508_rahayu_rodda.pdf.imf" docOrigin="European Journal of Taxonomy 508" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Hoya solokensis Rahayu &amp; Rodda 2019, sp. nov." docType="treatment" docVersion="9" lastPageNumber="15" masterDocId="974EFD592A52FFDF2A5FC529FF8CFF8F" masterDocTitle="Hoya of Sumatra, an updated checklist, three new species, and a new subspecies" masterLastPageNumber="23" masterPageNumber="1" pageNumber="14" updateTime="1698715017593" updateUser="ExternalLinkService">
<mods:mods id="A6B6C1D0326AA6121FFE5D313BBE6DA2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B8E30E7D9815517420CD1A8FF661E964">
<mods:title id="BC71277AFA1D4AABCE2EAD15E3BBF652">Hoya of Sumatra, an updated checklist, three new species, and a new subspecies</mods:title>
</mods:titleInfo>
<mods:name id="615A1E4517938B62F45F7B018A7753D0" type="personal">
<mods:role id="D335D75B35365D73E1E8A17B1A5644DC">
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<mods:namePart id="3B54CEF5DCF313498FE0D6EF11311EE4">Rahayu, Sri</mods:namePart>
</mods:name>
<mods:name id="396E8C04CE4D7AB52010121C0CEDE6FF" type="personal">
<mods:role id="597FBDBF2FC21D7054A9883AD3BCEA08">
<mods:roleTerm id="8CD77F992547445C69E45C14307274E2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="74B775ED818E9D093045EC68822578FC">Rodda, Michele</mods:namePart>
</mods:name>
<mods:typeOfResource id="3B4D4B35A059B3545493A4C4E5E4F5C3">text</mods:typeOfResource>
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<mods:titleInfo id="E6E08CCA2F6AD87A7E7B8C648ABB6137">
<mods:title id="BB965E4A76D68A1C17AA1B899995F814">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="30F25875CF65E3E8FFBF76DBF9622606">
<mods:date id="EE859CF36421A6604AC23511CAEF3EC3">2019</mods:date>
<mods:detail id="06978586D4E9D9D6D5B68BE4BEF5A19D" type="pubDate">
<mods:number id="EF186C39E4A129A73763732CC381736D">2019-03-23</mods:number>
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<mods:detail id="10F582667E0682FE3D8A4857754A8737" type="volume">
<mods:number id="0DB689334A0DAFA5EB170CFD058F032B">508</mods:number>
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<mods:start id="EB33DA89656B47EF9B173B81F03F0767">1</mods:start>
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<mods:classification id="28EFB5F0D35E028F38CF7C10AB5AA1DC">journal article</mods:classification>
<mods:identifier id="A6A1093CB8864BA24DA6B6102ECCAFCA" type="CLB-Dataset">27816</mods:identifier>
<mods:identifier id="965FC8144514FF92670BB78AC4DD87CE" type="DOI">10.5852/ejt.2019.508</mods:identifier>
<mods:identifier id="2DFD31C2D7A64A1077CFC779EFD1342E" type="GBIF-Dataset">6e176ff1-c6ce-429c-8b36-9d32c3cdc5c9</mods:identifier>
<mods:identifier id="2115FFDB12BBCAA764E8BF189D68158A" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="2389857B817A3D0F04B6D7D3A6EAE2DA" type="Zenodo-Dep">2612875</mods:identifier>
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<treatment id="6B7785212A5FFFD1284EC132FAFBFA3C" ID-DOI="http://doi.org/10.5281/zenodo.3489092" ID-GBIF-Taxon="154516707" ID-Zenodo-Dep="3489092" LSID="urn:lsid:plazi:treatment:6B7785212A5FFFD1284EC132FAFBFA3C" httpUri="http://treatment.plazi.org/id/6B7785212A5FFFD1284EC132FAFBFA3C" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<subSubSection id="ABC467BC2A5FFFD2284EC132FC61FBD6" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="E36134372A5FFFD2284EC132FBAEFBBA" blockId="13.[529,1058,1050,1149]" box="[529,1058,1050,1077]" pageId="13" pageNumber="14">
<heading id="B829835B2A5FFFD2284EC132FBAEFBBA" box="[529,1058,1050,1077]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="14" reason="2">
<taxonomicName id="24DE4FB42A5FFFD2284EC132FC4DFBBA" ID-CoL="7WNGY" authority="S.Rahayu &amp; Rodda" authorityName="Rahayu &amp; Rodda" authorityYear="2019" box="[529,961,1050,1077]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="species" species="solokensis" status="sp. nov.">
<emphasis id="D1AAE8252A5FFFD2284EC132FD5AFBBA" bold="true" box="[529,726,1050,1077]" italics="true" pageId="13" pageNumber="14">Hoya solokensis</emphasis>
S.Rahayu &amp; Rodda
</taxonomicName>
<taxonomicNameLabel id="CA99555E2A5FFFD22997C132FBAEFBBA" box="[968,1058,1051,1077]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="E36134372A5FFFD22818C117FC61FBD6" blockId="13.[529,1058,1050,1149]" box="[583,1005,1086,1113]" pageId="13" pageNumber="14">urn:lsid:ipni.org:names:77195001-1</paragraph>
</subSubSection>
<subSubSection id="ABC467BC2A5FFFD228F0C14BFC08FBF2" box="[687,900,1122,1149]" pageId="13" pageNumber="14" type="description">
<paragraph id="E36134372A5FFFD228F0C14BFC08FBF2" blockId="13.[529,1058,1050,1149]" box="[687,900,1122,1149]" pageId="13" pageNumber="14">
<figureCitation id="7BE528B22A5FFFD228F0C14BFD7FFBF2" box="[687,755,1122,1149]" captionStart="Fig" captionStartId="4.[189,236,1843,1872]" captionTargetBox="[290,1290,265,1798]" captionTargetId="figure@4.[287,1300,265,1803]" captionTargetPageId="4" captionText="Fig. 1. Flowers. A, C, E. Top view of the flower. B, D, F. Side view of the flower, with part of corolla removed. A–B. H. danumensis subsp. amarii S.Rahayu &amp; Rodda subsp. nov. (Amar Husein Sitompul s.n. leg., BO). C–D. H. solokensis S.Rahayu &amp; Rodda sp. nov. (Fadly s.n. leg., BO). E–F. H. rigidifolia S.Rahayu &amp; Rodda sp. nov. (S. Rahayu 865 leg., BO). Drawing: X. Y. Loh." httpUri="https://zenodo.org/record/2612877/files/figure.png" pageId="13" pageNumber="14">Figs 1</figureCitation>
C–D, 2C–D
</paragraph>
</subSubSection>
<subSubSection id="ABC467BC2A5FFFD22AE2C18CFB98FB61" box="[189,1044,1189,1262]" pageId="13" pageNumber="14" type="etymology">
<paragraph id="E36134372A5FFFD22AE2C18CFEC8FB30" blockId="13.[189,324,1189,1215]" box="[189,324,1189,1215]" pageId="13" pageNumber="14">
<heading id="B829835B2A5FFFD22AE2C18CFEC8FB30" bold="true" box="[189,324,1189,1215]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="D1AAE8252A5FFFD22AE2C18CFEC8FB30" bold="true" box="[189,324,1189,1215]" pageId="13" pageNumber="14">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="E36134372A5FFFD22AE2C1FDFB98FB61" blockId="13.[189,1044,1236,1262]" box="[189,1044,1236,1262]" pageId="13" pageNumber="14">
The new species is named after the collection locality in Solok,
<collectingRegion id="211AFAD52A5FFFD229F4C1FDFB9DFB61" box="[939,1041,1236,1262]" country="Indonesia" name="Sumatera Utara" pageId="13" pageNumber="14">Sumatra</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="ABC467BC2A5FFFD22AE2C03FFB01FAD0" box="[189,1165,1302,1376]" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="E36134372A5FFFD22AE2C03FFEA3FABF" blockId="13.[189,303,1302,1328]" box="[189,303,1302,1328]" pageId="13" pageNumber="14">
<heading id="B829835B2A5FFFD22AE2C03FFEA3FABF" bold="true" box="[189,303,1302,1328]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="D1AAE8252A5FFFD22AE2C03FFEA3FABF" bold="true" box="[189,303,1302,1328]" pageId="13" pageNumber="14">
<typeStatus id="3C658A952A5FFFD22AE2C03FFEA3FABF" box="[189,303,1302,1328]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph id="E36134372A5FFFD22AE2C06CFB01FAD0" blockId="13.[189,1165,1349,1376]" box="[189,1165,1349,1376]" pageId="13" pageNumber="14">
<materialsCitation id="53B63E6A2A5FFFD22AE2C06CFB01FAD0" ID-GBIF-Occurrence="2029332046" box="[189,1165,1349,1376]" collectingDate="2016-10-10" collectionCode="BO" country="Indonesia" elevation="1000" location="Solok" pageId="13" pageNumber="14" specimenCount="1" stateProvince="Sumatera Utara" typeStatus="holotype">
<collectingCountry id="9BC974A72A5FFFD22AE2C06CFED7FAD0" box="[189,347,1349,1375]" name="Indonesia" pageId="13" pageNumber="14">INDONESIA</collectingCountry>
•
<collectingRegion id="211AFAD52A5FFFD22B2BC06CFE55FAD0" box="[372,473,1349,1375]" country="Indonesia" name="Sumatera Utara" pageId="13" pageNumber="14">Sumatra</collectingRegion>
,
<location id="E60162EC2A5FFFD22BBBC06CFDA0FAD0" LSID="urn:lsid:plazi:treatment:6B7785212A5FFFD1284EC132FAFBFA3C:E60162EC2A5FFFD22BBBC06CFDA0FAD0" box="[484,556,1349,1375]" country="Indonesia" name="Solok" pageId="13" pageNumber="14" stateProvince="Sumatera Utara">Solok</location>
; ca
<elevation id="68F3D3042A5FFFD22805C06CFD7FFAD0" box="[602,755,1349,1376]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="14" unit="m" value="1000.0">
<quantity id="242699D22A5FFFD22805C06CFD3FFAEF" box="[602,691,1349,1376]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="14" unit="m" value="1000.0">1000 m</quantity>
a.s.l.
</elevation>
;
<date id="976012F72A5FFFD2295FC06CFC10FAD0" box="[768,924,1349,1375]" pageId="13" pageNumber="14" value="2016-10-10">
<collectingDate id="8724EB1F2A5FFFD2295FC06CFC10FAD0" box="[768,924,1349,1375]" pageId="13" pageNumber="14" value="2016-10-10">10 Oct. 2016</collectingDate>
</date>
; Fadly s.n. leg.;
<collectionCode id="85CFACF22A5FFFD22E02C06CFB05FAD0" box="[1117,1161,1349,1375]" country="Indonesia" lsid="urn:lsid:biocol.org:col:15186" name="Herbarium Bogoriense" pageId="13" pageNumber="14" type="Herbarium">BO</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="ABC467BC2A5FFFD12AE2C0AEFB86FDD5" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="description">
<paragraph id="E36134372A5FFFD22AE2C0AEFEC2FA2E" blockId="13.[189,334,1415,1441]" box="[189,334,1415,1441]" pageId="13" pageNumber="14">
<heading id="B829835B2A5FFFD22AE2C0AEFEC2FA2E" bold="true" box="[189,334,1415,1441]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="D1AAE8252A5FFFD22AE2C0AEFEC2FA2E" bold="true" box="[189,334,1415,1441]" pageId="13" pageNumber="14">Description</emphasis>
</heading>
</paragraph>
<paragraph id="E36134372A5FFFD12AE2C09FFB86FDD5" blockId="13.[189,1399,1462,2027]" lastBlockId="14.[189,1399,267,603]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
Epiphytic shrub, with white latex in all vegetative parts. Roots only basal, no adventitious roots observed. Stems erect or spreading,
<quantity id="242699D22A5FFFD22BB8C0F3FDD1FA7A" box="[487,605,1498,1525]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="5.0" metricValueMin="2.5" pageId="13" pageNumber="14" unit="mm" value="3.75" valueMax="5.0" valueMin="2.5">2.5–5 mm</quantity>
in diameter, dull green, sparsely pubescent when young; older stems grey-brown, glabrous; internodes 1–2(5) cm. Leaf blades 5–10(12) ×
<quantity id="242699D22A5FFFD22E52C0D7FB0DF996" box="[1037,1153,1534,1561]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" metricValueMax="4.0" metricValueMin="2.5" pageId="13" pageNumber="14" unit="cm" value="3.25" valueMax="4.0" valueMin="2.5">2.5–4 cm</quantity>
, thin, not succulent, chartaceous when dry, elliptic, base and apex acute or acuminate, mid green on adaxial surface, pale green on abaxial surface, glabrous, venation pinnate, midrib slightly depressed on adaxial surface, convex on abaxial surface, with 4–7 pairs of secondary veins, basal colleters absent; petioles 5–10 ×
<quantity id="242699D22A5FFFD22AE2C3A7FEB5F927" box="[189,313,1678,1704]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="2.0" metricValueMin="1.7" pageId="13" pageNumber="14" unit="mm" value="1.85" valueMax="2.0" valueMin="1.7">1.7–2 mm</quantity>
, terete channeled above, mid green, sparsely pubescent. Inflorescences sciadioidal, convex, of 3–10 flowers; peduncles 8–12 ×
<quantity id="242699D22A5FFFD22821C39BFC98F943" box="[638,788,1714,1740]" metricMagnitude="-3" metricUnit="m" metricValue="1.2999999999999998" metricValueMax="1.4" metricValueMin="1.2" pageId="13" pageNumber="14" unit="mm" value="1.2999999999999998" valueMax="1.4" valueMin="1.2">1.2–1.4 mm</quantity>
, extra-axillary, positively geotropic or horizontal, unbranched, one at each node, producing flowers only once, terete, green, sparsely pubescent just below the rachis, otherwise glabrous. Flowers with terete pedicels 5–8 ×
<quantity id="242699D22A5FFFD22E53C3D0FB0AF89B" box="[1036,1158,1785,1812]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="13" pageNumber="14" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 mm</quantity>
, pale green, sparsely pubescent. Calyx lobes ovate- round, apex round, 1.4–2.2 ×
<quantity id="242699D22A5FFFD22926C234FB85F8B7" box="[889,1033,1821,1848]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.6" metricValueMin="1.2" pageId="13" pageNumber="14" unit="mm" value="1.4" valueMax="1.6" valueMin="1.2">1.2–1.6 mm</quantity>
, light green, pubescent outside, glabrous inside, ciliate; basal colleters 1.5–2.3 ×
<quantity id="242699D22A5FFFD22959C268FC14F8D3" box="[774,920,1857,1884]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="13" pageNumber="14" unit="mm" value="1.4" valueMax="1.5" valueMin="1.3">1.3–1.5 mm</quantity>
, 1–3 at each calyx sinus, ovoid. Corolla tube basally bulbous, tightly enveloping the lower half of the corona, with a contracted throat, a short funnel-shaped tube and free, lanceolate, spreading lobes, white; basal bulbous part
<quantity id="242699D22A5FFFD22EEAC2A0FABCF82C" box="[1205,1328,1929,1956]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="13" pageNumber="14" unit="mm" value="2.75" valueMax="3.0" valueMin="2.5">2.5–3 mm</quantity>
high,
<quantity id="242699D22A5FFFD22AE2C284FEBAF848" box="[189,310,1965,1992]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="13" pageNumber="14" unit="mm" value="3.25" valueMax="3.5" valueMin="3.0">3–3.5 mm</quantity>
in diameter; free lobes 5–6 ×
<quantity id="242699D22A5FFFD228C0C284FC97F848" box="[671,795,1965,1992]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="13" pageNumber="14" unit="mm" value="2.75" valueMax="3.0" valueMin="2.5">2.5–3 mm</quantity>
, narrowly triangular, valvate in bud, with recurved edges and tip, tube pubescent inside with retrorse hairs, sparsely pubescent outside with spreading hairs, lobes sparsely pubescent inside and outside with spreading hairs. Corona staminal
<quantity id="242699D22A5CFFD12EDCC422FB76FEA9" box="[1155,1274,267,294]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="5.0" metricValueMin="4.5" pageId="14" pageNumber="15" unit="mm" value="4.75" valueMax="5.0" valueMin="4.5">4.5–5 mm</quantity>
high,
<quantity id="242699D22A5CFFD12F1CC422FE8EFEC3" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="14" pageNumber="15" unit="mm" value="2.75" valueMax="3.0" valueMin="2.5">2.5– 3 mm</quantity>
in diameter, stiff and waxy-looking, white; lobes 4.5–5 ×
<quantity id="242699D22A5CFFD129E6C41BFBB8FEC3" box="[953,1076,306,332]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="14" pageNumber="15" unit="mm" value="1.1" valueMax="1.2" valueMin="1.0">1–1.2 mm</quantity>
, erect, oblong and sinuose, attached at the back of the anthers, upper apicis rounded, touching in the middle, outer tips obtuse, with a narrow revolute margin.Anthers ca 0.5 ×
<quantity id="242699D22A5CFFD128EEC456FC81FE15" box="[689,781,383,410]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="15" unit="mm" value="0.3">0.3 mm</quantity>
, ovate, with apical round membranaceous appendage just covering the style-head apex. Pollinia 470–520 × 200–240 μm, oblong, with a round base and apex; pellucid margin missing; caudicles ca 130 × 70 μm, ovoid, almost transparent; corpusculum 300–320 × 160–180 μm, ovoid; style-head 5-angled in cross section, with five spreading lobes alternating with the stamens, style-head apex conical,
<quantity id="242699D22A5CFFD12817C730FD32FDBB" box="[584,702,537,564]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="14" pageNumber="15" unit="mm" value="1.1" valueMax="1.2" valueMin="1.0">1–1.2 mm</quantity>
high,
<quantity id="242699D22A5CFFD12959C730FC1EFDBB" box="[774,914,537,564]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="14" pageNumber="15" unit="mm" value="1.0" valueMax="1.1" valueMin="0.9">0.9–1.1 mm</quantity>
in diameter at the base, apex acute; ovary
<quantity id="242699D22A5CFFD12AE2C769FEC6FDD5" box="[189,330,576,603]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="14" pageNumber="15" unit="mm" value="1.6" valueMax="1.7" valueMin="1.5">1.5–1.7 mm</quantity>
high, conical, with apex acute. Fruit and seed not observed.
</paragraph>
</subSubSection>
<subSubSection id="ABC467BC2A5CFFD12AE2C7ADFB47FD78" pageId="14" pageNumber="15" type="distribution">
<paragraph id="E36134372A5CFFD12AE2C7ADFE62FD11" blockId="14.[189,494,644,671]" box="[189,494,644,671]" pageId="14" pageNumber="15">
<heading id="B829835B2A5CFFD12AE2C7ADFE62FD11" bold="true" box="[189,494,644,671]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="D1AAE8252A5CFFD12AE2C7ADFE62FD11" bold="true" box="[189,494,644,671]" pageId="14" pageNumber="15">Distribution and habitat</emphasis>
</heading>
</paragraph>
<paragraph id="E36134372A5CFFD12AE2C79EFB47FD78" blockId="14.[189,1398,694,760]" pageId="14" pageNumber="15">
<taxonomicName id="24DE4FB42A5CFFD12AE2C79EFE0DFD5F" authority="Rahayu &amp; Rodda, 2019" authorityName="Rahayu &amp; Rodda" authorityYear="2019" box="[189,385,694,720]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="solokensis" status="sp. nov.">
<emphasis id="D1AAE8252A5CFFD12AE2C79EFE0DFD5F" box="[189,385,694,720]" italics="true" pageId="14" pageNumber="15">Hoya solokensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA99555E2A5CFFD12BD2C79EFE60FD5E" box="[397,492,695,721]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
is only known from the type locality in Solok,
<collectingRegion id="211AFAD52A5CFFD12E1DC79EFB2BFD5E" box="[1090,1191,695,721]" country="Indonesia" name="Sumatera Utara" pageId="14" pageNumber="15">Sumatra</collectingRegion>
,
<collectingCountry id="9BC974A72A5CFFD12EE9C79FFAA7FD5F" box="[1206,1323,694,720]" name="Indonesia" pageId="14" pageNumber="15">Indonesia</collectingCountry>
, at ca
<quantity id="242699D22A5CFFD12AE2C7F4FE9AFD77" box="[189,278,733,760]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="15" unit="m" value="1000.0">1000 m</quantity>
a.s.l. It was collected in an evergreen forest, where it was growing epiphytically.
</paragraph>
</subSubSection>
<subSubSection id="ABC467BC2A5CFFD12AE2C60BFE8FFB8F" pageId="14" pageNumber="15" type="description">
<paragraph id="E36134372A5CFFD12AE2C60BFE39FCB3" blockId="14.[189,437,802,828]" box="[189,437,802,828]" pageId="14" pageNumber="15">
<heading id="B829835B2A5CFFD12AE2C60BFE39FCB3" bold="true" box="[189,437,802,828]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="D1AAE8252A5CFFD12AE2C60BFE39FCB3" bold="true" box="[189,437,802,828]" pageId="14" pageNumber="15">Conservation status</emphasis>
</heading>
</paragraph>
<paragraph id="E36134372A5CFFD12AE2C67DFEEEFC34" blockId="14.[189,1398,852,955]" pageId="14" pageNumber="15">
Known only from a single collection and lacking information on the distribution area, the population size and the possible threats to the habitat,
<taxonomicName id="24DE4FB42A5CFFD128E5C652FCDBFC1B" authority="Rahayu &amp; Rodda, 2019" authorityName="Rahayu &amp; Rodda" authorityYear="2019" box="[698,855,890,916]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="solokensis" status="sp. nov.">
<emphasis id="D1AAE8252A5CFFD128E5C652FCDBFC1B" box="[698,855,890,916]" italics="true" pageId="14" pageNumber="15">H. solokensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA99555E2A5CFFD12900C652FC36FC1A" box="[863,954,891,917]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
is considered as Data Deficient (DD) (
<bibRefCitation id="874F49C62A5CFFD12A9AC688FEDAFC34" author="IUCN &amp; Cambridge, UK" box="[197,342,929,955]" pageId="14" pageNumber="15" refId="ref8900" refString="IUCN 2012. IUCN Red List Categories and Criteria (version 3.1, 2 nd edition). IUCN, Gland, Switzerland and Cambridge, UK." type="book" year="2012">IUCN 2012</bibRefCitation>
).
</paragraph>
<paragraph id="E36134372A5CFFD12AE2C6CFFE8FFB8F" blockId="14.[189,259,998,1024]" box="[189,259,998,1024]" pageId="14" pageNumber="15">
<heading id="B829835B2A5CFFD12AE2C6CFFE8FFB8F" bold="true" box="[189,259,998,1024]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="D1AAE8252A5CFFD12AE2C6CFFE8FFB8F" bold="true" box="[189,259,998,1024]" pageId="14" pageNumber="15">Notes</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="ABC467BC2A5CFFD12AE2C131FAFBFA3C" pageId="14" pageNumber="15" type="reference_group">
<paragraph id="E36134372A5CFFD12AE2C131FAFBFA3C" blockId="14.[189,1399,1047,1460]" pageId="14" pageNumber="15">
<taxonomicName id="24DE4FB42A5CFFD12AE2C131FE0CFBBE" ID-CoL="7WNGY" authority="Rahayu &amp; Rodda, 2019" authorityName="Rahayu &amp; Rodda" authorityYear="2019" box="[189,384,1047,1073]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="solokensis" status="sp. nov.">
<emphasis id="D1AAE8252A5CFFD12AE2C131FE0CFBBE" box="[189,384,1047,1073]" italics="true" pageId="14" pageNumber="15">Hoya solokensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA99555E2A5CFFD12BD6C131FE6AFBBD" box="[393,486,1048,1074]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
is one of the few species of
<taxonomicName id="24DE4FB42A5CFFD12916C131FC05FBBE" ID-CoL="4Z2L" box="[841,905,1048,1073]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="genus">
<emphasis id="D1AAE8252A5CFFD12916C131FC05FBBE" box="[841,905,1048,1073]" italics="true" pageId="14" pageNumber="15">Hoya</emphasis>
</taxonomicName>
that exhibits a non-climbing habit but is instead an epiphytic shrub. It is morphologically similar to
<taxonomicName id="24DE4FB42A5CFFD129CFC116FE88FBF0" ID-CoL="3MRXY" authority="Rodda (Rodda &amp; Ercole 2014)" authorityName="Rodda (Rodda &amp; Ercole" authorityYear="2014" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="papaschonii">
<emphasis id="D1AAE8252A5CFFD129CFC116FBCAFBD7" box="[912,1094,1086,1112]" italics="true" pageId="14" pageNumber="15">H. papaschonii</emphasis>
<bibRefCitation id="874F49C62A5CFFD12E0EC117FE88FBF0" author="Rodda M. &amp; Ercole E." pageId="14" pageNumber="15" pagination="97 - 106" refId="ref9614" refString="Rodda M. &amp; Ercole E. 2014. Hoya papaschonii (Apocynaceae: Asclepiadoideae), a new species from southern Thailand with a peculiar corona. Phytotaxa 175 (2): 97 - 106. https: // doi. org / 10.11646 / phytotaxa. 175.2.4" type="journal article" year="2014">Rodda (Rodda &amp; Ercole 2014)</bibRefCitation>
</taxonomicName>
, a species only found in southern
<collectingCountry id="9BC974A72A5CFFD128E9C14DFCAEFBF1" box="[694,802,1124,1150]" name="Thailand" pageId="14" pageNumber="15">Thailand</collectingCountry>
. Both species share a shrubby habit, short-lived peduncles, and flowers with a tubular corolla. However,
<taxonomicName id="24DE4FB42A5CFFD12907C1A5FB87FB2A" ID-CoL="3MRXY" authorityName="Rodda (Rodda &amp; Ercole" authorityYear="2014" box="[856,1035,1163,1189]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="papaschonii">
<emphasis id="D1AAE8252A5CFFD12907C1A5FB87FB2A" box="[856,1035,1163,1189]" italics="true" pageId="14" pageNumber="15">H. papaschonii</emphasis>
</taxonomicName>
has the free part of the corolla lobes mostly held upright, while in
<taxonomicName id="24DE4FB42A5CFFD1282BC19AFC98FB43" ID-CoL="7WNGY" authority="Rahayu &amp; Rodda, 2019" authorityName="Rahayu &amp; Rodda" authorityYear="2019" box="[628,788,1202,1228]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="solokensis" status="sp. nov.">
<emphasis id="D1AAE8252A5CFFD1282BC19AFC98FB43" box="[628,788,1202,1228]" italics="true" pageId="14" pageNumber="15">H. solokensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA99555E2A5CFFD12940C19BFCF1FB43" box="[799,893,1202,1228]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
the lobes are spreading; furthermore, the corona of
<taxonomicName id="24DE4FB42A5CFFD12B65C1F0FE7CFB7D" ID-CoL="3MRXY" authorityName="Rodda (Rodda &amp; Ercole" authorityYear="2014" box="[314,496,1240,1266]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="papaschonii">
<emphasis id="D1AAE8252A5CFFD12B65C1F0FE7CFB7D" box="[314,496,1240,1266]" italics="true" pageId="14" pageNumber="15">H. papaschonii</emphasis>
</taxonomicName>
has both staminal and interstaminal elements while
<taxonomicName id="24DE4FB42A5CFFD12E30C1F0FA83FB7D" ID-CoL="7WNGY" authority="Rahayu &amp; Rodda, 2019" authorityName="Rahayu &amp; Rodda" authorityYear="2019" box="[1135,1295,1240,1266]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="solokensis" status="sp. nov.">
<emphasis id="D1AAE8252A5CFFD12E30C1F0FA83FB7D" box="[1135,1295,1240,1266]" italics="true" pageId="14" pageNumber="15">H. solokensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA99555E2A5CFFD12F46C1F0FAFBFB7C" box="[1305,1399,1241,1267]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
only has a staminal corona. Another species with a tubular corolla is
<taxonomicName id="24DE4FB42A5CFFD129BFC029FABAFA95" ID-CoL="3MS4N" authority="Omlor (1996)" authorityName="Omlor" authorityYear="1996" box="[992,1334,1279,1306]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="telosmoides">
<emphasis id="D1AAE8252A5CFFD129BFC029FB02FA96" box="[992,1166,1279,1305]" italics="true" pageId="14" pageNumber="15">H. telosmoides</emphasis>
<bibRefCitation id="874F49C62A5CFFD12ECAC1D6FABAFA95" author="Omlor R." box="[1173,1334,1279,1306]" pageId="14" pageNumber="15" pagination="288 - 294" refId="ref9328" refString="Omlor R. 1996. Notes on Marsdenieae (Asclepiadaceae): A new, unusual species of Hoya from Northern Borneo. Novon 6: 288 - 294. https: // doi. org / 10.2307 / 3392096" type="journal article" year="1996">Omlor (1996)</bibRefCitation>
</taxonomicName>
from Borneo, that is, however, a climber. The flowers of
<taxonomicName id="24DE4FB42A5CFFD1294FC00FFC20FAB0" ID-CoL="7WNGY" authority="Rahayu &amp; Rodda, 2019" authorityName="Rahayu &amp; Rodda" authorityYear="2019" box="[784,940,1317,1343]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="solokensis" status="sp. nov.">
<emphasis id="D1AAE8252A5CFFD1294FC00FFC20FAB0" box="[784,940,1317,1343]" italics="true" pageId="14" pageNumber="15">H. solokensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA99555E2A5CFFD129EDC00FFB80FACF" box="[946,1036,1318,1344]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
superficially resemble those of the Bornean
<taxonomicName id="24DE4FB42A5CFFD12B0BC064FB5AFAE9" ID-CoL="3MRRB" authority="A. L. Lamb, Gavrus, Emoi &amp; Gokusing (Lamb et al. 2014)" authorityName="A. L. Lamb, Gavrus, Emoi &amp; Gokusing" authorityYear="2014" box="[340,1238,1356,1382]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="hamiltoniorum">
<emphasis id="D1AAE8252A5CFFD12B0BC064FDAAFAE9" box="[340,550,1356,1382]" italics="true" pageId="14" pageNumber="15">H. hamiltoniorum</emphasis>
A.L.
<bibRefCitation id="874F49C62A5CFFD12801C065FB5AFAE9" author="Lamb A. &amp; Gavrus A. &amp; Emoi B. &amp; Gokusing L." box="[606,1238,1356,1382]" pageId="14" pageNumber="15" pagination="1 - 89" refId="ref9187" refString="Lamb A., Gavrus A., Emoi B. &amp; Gokusing L. 2014. The Hoyas of Sabah, a commentary with seven new species and a new subspecies. Sandakania 19: 1 - 89." type="journal article" year="2014">
Lamb, Gavrus, Emoi &amp; Gokusing (Lamb
<emphasis id="D1AAE8252A5CFFD12E10C064FB06FAE9" box="[1103,1162,1356,1382]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
2014)
</bibRefCitation>
</taxonomicName>
because both species have the free part of the corolla lobes spreading, however
<taxonomicName id="24DE4FB42A5CFFD1299EC05AFB1EFA03" ID-CoL="3MRRB" box="[961,1170,1394,1420]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="hamiltoniorum">
<emphasis id="D1AAE8252A5CFFD1299EC05AFB1EFA03" box="[961,1170,1394,1420]" italics="true" pageId="14" pageNumber="15">H. hamiltoniorum</emphasis>
</taxonomicName>
is a climber and its inner apex of the corona lobe is bifid, while the apex of the inner lobes of
<taxonomicName id="24DE4FB42A5CFFD12E4EC0B3FB21FA3C" ID-CoL="7WNGY" authority="Rahayu &amp; Rodda, 2019" authorityName="Rahayu &amp; Rodda" authorityYear="2019" box="[1041,1197,1433,1459]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="solokensis" status="sp. nov.">
<emphasis id="D1AAE8252A5CFFD12E4EC0B3FB21FA3C" box="[1041,1197,1433,1459]" italics="true" pageId="14" pageNumber="15">H. solokensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA99555E2A5CFFD12EECC0B3FA80FA3B" box="[1203,1292,1434,1460]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
is entire.
</paragraph>
</subSubSection>
</treatment>
</document>
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