treatments-xml/data/6B/77/85/6B7785212A51FFD92BC1C425FCBEFE6A.xml

<document id="01BF737A056BC9335AAB64DCDE30ED31" ID-CLB-Dataset="27816" ID-DOI="10.5852/ejt.2019.508" ID-GBIF-Dataset="6e176ff1-c6ce-429c-8b36-9d32c3cdc5c9" ID-ISSN="2118-9773" ID-Zenodo-Dep="2612875" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1553714929206" checkinUser="plazi" docAuthor="Rahayu, Sri &amp; Rodda, Michele" docDate="2019" docId="6B7785212A51FFD92BC1C425FCBEFE6A" docLanguage="en" docName="ejt-508_rahayu_rodda.pdf.imf" docOrigin="European Journal of Taxonomy 508" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Hoya danumensis subsp. amarii Rahayu &amp; Rodda 2019, subsp. nov." docType="treatment" docVersion="9" lastPageNumber="7" masterDocId="974EFD592A52FFDF2A5FC529FF8CFF8F" masterDocTitle="Hoya of Sumatra, an updated checklist, three new species, and a new subspecies" masterLastPageNumber="23" masterPageNumber="1" pageNumber="4" updateTime="1698715017593" updateUser="ExternalLinkService">
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<mods:title id="33C4DFF192C933AB4D068BFFB212EE28">Hoya of Sumatra, an updated checklist, three new species, and a new subspecies</mods:title>
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<treatment id="6B7785212A51FFD92BC1C425FCBEFE6A" ID-DOI="http://doi.org/10.5281/zenodo.3489034" ID-GBIF-Taxon="154516739" ID-Zenodo-Dep="3489034" LSID="urn:lsid:plazi:treatment:6B7785212A51FFD92BC1C425FCBEFE6A" httpUri="http://treatment.plazi.org/id/6B7785212A51FFD92BC1C425FCBEFE6A" lastPageId="6" lastPageNumber="7" pageId="3" pageNumber="4">
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<paragraph id="E36134372A51FFDC2BC1C425FB1AFEA9" blockId="3.[414,1174,267,366]" box="[414,1174,267,294]" pageId="3" pageNumber="4">
<taxonomicName id="24DE4FB42A51FFDC2BC1C425FB87FEAA" ID-CoL="7ZBVZ" authority="S.Rahayu &amp; Rodda" authorityName="Rahayu &amp; Rodda" authorityYear="2019" box="[414,1035,267,294]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="subSpecies" species="danumensis" status="subsp. nov." subSpecies="amarii">
<emphasis id="D1AAE8252A51FFDC2BC1C425FDF4FEA9" bold="true" box="[414,632,267,294]" italics="true" pageId="3" pageNumber="4">Hoya danumensis</emphasis>
subsp. 
<emphasis id="D1AAE8252A51FFDC288EC422FCADFEAA" bold="true" box="[721,801,267,293]" italics="true" pageId="3" pageNumber="4">amarii</emphasis>
S.Rahayu &amp; Rodda
</taxonomicName>
<taxonomicNameLabel id="CA99555E2A51FFDC2E4DC422FB1AFEA9" box="[1042,1174,267,294]" pageId="3" pageNumber="4" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="E36134372A51FFDC2818C419FC61FEC5" blockId="3.[414,1174,267,366]" box="[583,1005,304,330]" pageId="3" pageNumber="4">urn:lsid:ipni.org:names:77194999-1</paragraph>
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<subSubSection id="ABC467BC2A51FFDC28F0C47DFC08FEE1" box="[687,900,340,366]" pageId="3" pageNumber="4" type="description">
<paragraph id="E36134372A51FFDC28F0C47DFC08FEE1" blockId="3.[414,1174,267,366]" box="[687,900,340,366]" pageId="3" pageNumber="4">
<figureCitation id="7BE528B22A51FFDC28F0C47DFD7FFEE1" box="[687,755,340,366]" captionStart="Fig" captionStartId="4.[189,236,1843,1872]" captionTargetBox="[290,1290,265,1798]" captionTargetId="figure@4.[287,1300,265,1803]" captionTargetPageId="4" captionText="Fig. 1. Flowers. A, C, E. Top view of the flower. B, D, F. Side view of the flower, with part of corolla removed. A–B. H. danumensis subsp. amarii S.Rahayu &amp; Rodda subsp. nov. (Amar Husein Sitompul s.n. leg., BO). C–D. H. solokensis S.Rahayu &amp; Rodda sp. nov. (Fadly s.n. leg., BO). E–F. H. rigidifolia S.Rahayu &amp; Rodda sp. nov. (S. Rahayu 865 leg., BO). Drawing: X. Y. Loh." httpUri="https://zenodo.org/record/2612877/files/figure.png" pageId="3" pageNumber="4">Figs 1</figureCitation>
A–B, 2A–B
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<subSubSection id="ABC467BC2A51FFDC2AE2C4BEFB54FE6F" box="[189,1240,407,481]" pageId="3" pageNumber="4" type="etymology">
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<heading id="B829835B2A51FFDC2AE2C4BEFEC8FE3E" bold="true" box="[189,324,407,433]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="D1AAE8252A51FFDC2AE2C4BEFEC8FE3E" bold="true" box="[189,324,407,433]" pageId="3" pageNumber="4">Etymology</emphasis>
</heading>
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<paragraph id="E36134372A51FFDC2AE2C4EFFB54FE6F" blockId="3.[189,1240,454,481]" box="[189,1240,454,481]" pageId="3" pageNumber="4">The new species is named after Amar Husein Sitompul, who collected the type specimen.</paragraph>
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<subSubSection id="ABC467BC2A51FFDC2AE2C720FE47FD14" pageId="3" pageNumber="4" type="materials_examined">
<paragraph id="E36134372A51FFDC2AE2C720FEE0FDAC" blockId="3.[189,364,521,547]" box="[189,364,521,547]" pageId="3" pageNumber="4">
<heading id="B829835B2A51FFDC2AE2C720FEE0FDAC" bold="true" box="[189,364,521,547]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="D1AAE8252A51FFDC2AE2C720FEE0FDAC" bold="true" box="[189,364,521,547]" pageId="3" pageNumber="4">Type material</emphasis>
</heading>
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<paragraph id="E36134372A51FFDC2AB5C710FE47FD14" blockId="3.[189,1398,569,668]" pageId="3" pageNumber="4">
<materialsCitation id="53B63E6A2A51FFDC2AB5C710FE47FD14" ID-GBIF-Occurrence="2029332660" collectingDate="2014-09-16" collectionCode="BO" country="Indonesia" elevation="900" location="Padang Sidempuan" pageId="3" pageNumber="4" specimenCount="1" stateProvince="Sumatera Utara" typeStatus="holotype">
<emphasis id="D1AAE8252A51FFDC2AB5C710FED0FDDC" bold="true" box="[234,348,569,595]" pageId="3" pageNumber="4">
<typeStatus id="3C658A952A51FFDC2AB5C710FED0FDDC" box="[234,348,569,595]" pageId="3" pageNumber="4" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="9BC974A72A51FFDC2AE2C774FED7FDF8" box="[189,347,605,631]" name="Indonesia" pageId="3" pageNumber="4">INDONESIA</collectingCountry>
• 
<collectingRegion id="211AFAD52A51FFDC2B2EC774FE5AFDF8" box="[369,470,605,631]" country="Indonesia" name="Sumatera Utara" pageId="3" pageNumber="4">Sumatra</collectingRegion>
, 
<collectingRegion id="211AFAD52A51FFDC2B80C774FD02FDF8" box="[479,654,605,631]" country="Indonesia" name="Sumatera Utara" pageId="3" pageNumber="4">North Sumatra</collectingRegion>
, 
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; ca 
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a.s.l.
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; 
<date id="976012F72A51FFDC2E67C774FB5EFDF8" box="[1080,1234,605,631]" pageId="3" pageNumber="4" value="2014-09-16">
<collectingDate id="8724EB1F2A51FFDC2E67C774FB5EFDF8" box="[1080,1234,605,631]" pageId="3" pageNumber="4" value="2014-09-16">16 Sep. 2014</collectingDate>
</date>
; Amar Husein Sitompul s.n. leg.; 
<collectionCode id="85CFACF22A51FFDC2BC4C7A8FE4BFD14" box="[411,455,641,667]" country="Indonesia" lsid="urn:lsid:biocol.org:col:15186" name="Herbarium Bogoriense" pageId="3" pageNumber="4" type="Herbarium">BO</collectionCode>
.
</materialsCitation>
</paragraph>
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<emphasis id="D1AAE8252A51FFDC2AE2C7EAFEC2FD52" bold="true" box="[189,334,707,733]" pageId="3" pageNumber="4">Description</emphasis>
</heading>
</paragraph>
<paragraph id="E36134372A51FFDC2AE2C7DAFB5EF916" blockId="3.[189,1399,755,1690]" pageId="3" pageNumber="4">
Epiphytic shrub, with white latex in all vegetative parts. Roots only basal, no adventitious roots observed. Stems erect or spreading, 
<quantity id="242699D22A51FFDC283BC63EFD49FCBD" box="[612,709,791,818]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="3" pageNumber="4" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
in diameter, bright green, sparsely pubescent when young; older stems grey, glabrous; internodes 2–4(6) cm. Leaf blades 5–12 × 
<quantity id="242699D22A51FFDC2E4EC615FB08FCD9" box="[1041,1156,828,855]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" metricValueMax="5.0" metricValueMin="2.5" pageId="3" pageNumber="4" unit="cm" value="3.75" valueMax="5.0" valueMin="2.5">2.5–5 cm</quantity>
, thin, not succulent, chartaceous when dry, broadly lanceolate to oblong, base rounded, apex acuminate or caudate, mid green on adaxial surface, pale green on abaxial surface, glabrous (sparsely pubescent when young), venation pinnate, midrib slightly depressed on adaxial surface, convex on abaxial surface, secondary veins 3–7 pairs, basal colleters absent; petioles 5–10 × 
<quantity id="242699D22A51FFDC2918C6E4FC20FC67" box="[839,940,973,1000]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="3" pageNumber="4" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
, terete, channelled above, twisted, mid green, sparsely pubescent when young, otherwise glabrous. Inflorescences sciadioidal, convex, of 5–15 flowers, with peduncles extra-axillary, positively geotropic, unbranched, one at each node, producing flowers subsequently a few times, terete, 10–15 × 
<quantity id="242699D22A51FFDC2942C113FC3CFBDA" box="[797,944,1082,1109]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="3" pageNumber="4" unit="mm" value="2.0" valueMax="2.5" valueMin="1.5">1.5–2.5 mm</quantity>
, green, sparsely pubescent when very young. Flowers with terete pedicels 
<quantity id="242699D22A51FFDC2821C177FD7DFBF6" box="[638,753,1118,1145]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="3" pageNumber="4" unit="cm" value="2.75" valueMax="3.0" valueMin="2.5">2.5–3 cm</quantity>
× 
<quantity id="242699D22A51FFDC2946C177FC40FBF6" box="[793,972,1118,1145]" metricMagnitude="-5" metricUnit="m" metricValue="7" metricValueMax="8.0" metricValueMin="6.0" pageId="3" pageNumber="4" unit="mm" value="0.07" valueMax="0.08" valueMin="0.06">0.06–0.08 mm</quantity>
, pale green, glabrous. Calyx lobes triangular, apex round, 1.5–2 × 
<quantity id="242699D22A51FFDC2864C1AAFD42FB12" box="[571,718,1155,1182]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="3" pageNumber="4" unit="mm" value="0.55" valueMax="0.6" valueMin="0.5">0.5–0.6 mm</quantity>
, sparsely pubescent outside or glabrous, inside glabrous, ciliate; basal colleters 0.15–0.25 × 
<quantity id="242699D22A51FFDC280EC18EFD63FB4D" box="[593,751,1191,1218]" metricMagnitude="-4" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="3" pageNumber="4" unit="mm" value="0.175" valueMax="0.2" valueMin="0.15">0.15–0.2 mm</quantity>
, one at each calyx sinus, ovoid. Buds globose, 5-ridged, dark brown or deep purple when young, turning green. Corolla shallowly campanulate, 
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in diameter; tube 
<quantity id="242699D22A51FFDC2B33C1D9FE56FA85" box="[364,474,1264,1290]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="3" pageNumber="4" unit="mm" value="10.0" valueMax="11.0" valueMin="9.0">9–11 mm</quantity>
long, creamy white, glabrous outside, pilose inside, basally densely pubescent; lobes 2–3 × 
<quantity id="242699D22A51FFDC2B0CC03DFE59FAA0" box="[339,469,1300,1327]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="3" pageNumber="4" unit="mm" value="11.5" valueMax="12.0" valueMin="11.0">11–12 mm</quantity>
, very broadly triangular, valvate in bud, creamy white, glabrous outside, very sparsely pubescent inside, tip glabrous. Corona staminal 
<quantity id="242699D22A51FFDC2931C011FC5FFADC" box="[878,979,1336,1363]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="3" pageNumber="4" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
high, 
<quantity id="242699D22A51FFDC2E7CC010FB29FADC" box="[1059,1189,1337,1363]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="3" pageNumber="4" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">10–12 mm</quantity>
in diameter, stiff and waxy-looking, white; lobes 4.5–5 × 
<quantity id="242699D22A51FFDC28EDC075FCCAFAF8" box="[690,838,1372,1399]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="2.5" metricValueMin="2.3" pageId="3" pageNumber="4" unit="mm" value="2.4" valueMax="2.5" valueMin="2.3">2.3–2.5 mm</quantity>
, spreading, ovate, attached at the back of the anthers, inner process apiculate, not touching in the middle, outer process acute with a rounded tip, lobes with revolute margin underneath. Anthers ca 0.7 × 
<quantity id="242699D22A51FFDC297DC08CFC0CFA4F" box="[802,896,1445,1472]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="4" unit="mm" value="0.5">0.5 mm</quantity>
, ovate, with apical round membranaceous appendage covering the style-head apex. Pollinia 550–650 × 220–270 μm, oblong, with a round base and an obliquely truncated apex; pellucid margin present all along the outer edge; caudicles ca 280 × 180 μm, broadly triangular, almost transparent; corpusculum 300–350 × 100–120 μm, oblong; style-head 5-angled in cross section, with five spreading lobes alternating with the stamens, style-head depressed with a central raised conical apex ca 0.5 × 
<quantity id="242699D22A51FFDC28E9C372FC9DF9FA" box="[694,785,1627,1654]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="4" unit="mm" value="0.5">0.5 mm</quantity>
broad at the base, apex rounded; ovary 
<quantity id="242699D22A51FFDC2EB6C372FAFBF9FA" box="[1257,1399,1627,1653]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="3" pageNumber="4" unit="mm" value="1.5" valueMax="1.6" valueMin="1.4">1.4–1.6 mm</quantity>
high, 
<quantity id="242699D22A51FFDC2B5FC356FE01F915" box="[256,397,1663,1690]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="3" pageNumber="4" unit="mm" value="0.35" valueMax="0.4" valueMin="0.3">0.3–0.4 mm</quantity>
in diameter, narrowly conical, apex acute. Fruit and seed not observed.
</paragraph>
</subSubSection>
<subSubSection id="ABC467BC2A51FFDC2AE2C3EBFE6BF8BF" pageId="3" pageNumber="4" type="distribution">
<paragraph id="E36134372A51FFDC2AE2C3EBFE62F953" blockId="3.[189,494,1730,1756]" box="[189,494,1730,1756]" pageId="3" pageNumber="4">
<heading id="B829835B2A51FFDC2AE2C3EBFE62F953" bold="true" box="[189,494,1730,1756]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="D1AAE8252A51FFDC2AE2C3EBFE62F953" bold="true" box="[189,494,1730,1756]" pageId="3" pageNumber="4">Distribution and habitat</emphasis>
</heading>
</paragraph>
<paragraph id="E36134372A51FFDC2AE2C3DBFE6BF8BF" blockId="3.[189,1398,1777,1840]" pageId="3" pageNumber="4">
<taxonomicName id="24DE4FB42A51FFDC2AE2C3DBFDBBF884" authority="Rahayu &amp; Rodda, 2019" authorityName="Rahayu &amp; Rodda" authorityYear="2019" box="[189,567,1777,1803]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="subSpecies" species="danumensis" status="subsp. nov." subSpecies="amarii">
<emphasis id="D1AAE8252A51FFDC2AE2C3DBFE1CF884" box="[189,400,1777,1803]" italics="true" pageId="3" pageNumber="4">Hoya danumensis</emphasis>
subsp. 
<emphasis id="D1AAE8252A51FFDC2BB6C3DBFDBBF884" box="[489,567,1778,1803]" italics="true" pageId="3" pageNumber="4">amarii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA99555E2A51FFDC281FC3D8FD48F883" box="[576,708,1777,1804]" pageId="3" pageNumber="4" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
is only known from the type locality in northern 
<collectingRegion id="211AFAD52A51FFDC2F52C3DBFAFEF883" box="[1293,1394,1778,1804]" country="Indonesia" name="Sumatera Utara" pageId="3" pageNumber="4">Sumatra</collectingRegion>
, 
<collectingCountry id="9BC974A72A51FFDC2AE2C23FFEBEF8BF" box="[189,306,1814,1840]" name="Indonesia" pageId="3" pageNumber="4">Indonesia</collectingCountry>
, ca 
<quantity id="242699D22A51FFDC2B00C23FFE25F8BF" box="[351,425,1814,1840]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="4" unit="m" value="900.0">900 m</quantity>
a.s.l.
</paragraph>
</subSubSection>
<subSubSection id="ABC467BC2A51FFD92AE2C271FE8FFEAA" lastPageId="6" lastPageNumber="7" pageId="3" pageNumber="4" type="description">
<paragraph id="E36134372A51FFDC2AE2C271FE39F8FC" blockId="3.[189,437,1880,1907]" box="[189,437,1880,1907]" pageId="3" pageNumber="4">
<heading id="B829835B2A51FFDC2AE2C271FE39F8FC" bold="true" box="[189,437,1880,1907]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="D1AAE8252A51FFDC2AE2C271FE39F8FC" bold="true" box="[189,437,1880,1907]" pageId="3" pageNumber="4">Conservation status</emphasis>
</heading>
</paragraph>
<paragraph id="E36134372A51FFDC2AE2C2A0FE2BF864" blockId="3.[189,1399,1928,2027]" pageId="3" pageNumber="4">
Known only from a single collection and lacking information on the distribution area, the population size and the possible threats to the habitat, 
<taxonomicName id="24DE4FB42A51FFDC28EEC284FB89F849" authorityName="Rahayu &amp; Rodda" authorityYear="2019" box="[689,1029,1964,1990]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="subSpecies" species="danumensis" subSpecies="amarii">
<emphasis id="D1AAE8252A51FFDC28EEC284FCECF849" box="[689,864,1964,1990]" italics="true" pageId="3" pageNumber="4">H. danumensis</emphasis>
subsp. 
<emphasis id="D1AAE8252A51FFDC29E8C284FB89F849" box="[951,1029,1965,1990]" italics="true" pageId="3" pageNumber="4">amarii</emphasis>
</taxonomicName>
is considered as Data Deficient (DD) (
<bibRefCitation id="874F49C62A51FFDC2B55C2F8FE17F864" author="IUCN &amp; Cambridge, UK" box="[266,411,2001,2027]" pageId="3" pageNumber="4" refId="ref8900" refString="IUCN 2012. IUCN Red List Categories and Criteria (version 3.1, 2 nd edition). IUCN, Gland, Switzerland and Cambridge, UK." type="book" year="2012">IUCN 2012</bibRefCitation>
).
</paragraph>
<caption id="B7A164BF2A56FFDB2AE2C21AFF75F865" httpUri="https://zenodo.org/record/2612877/files/figure.png" pageId="4" pageNumber="5" startId="4.[189,236,1843,1872]" targetBox="[290,1290,265,1798]" targetPageId="4">
<paragraph id="E36134372A56FFDB2AE2C21AFF75F865" blockId="4.[189,1399,1843,2026]" pageId="4" pageNumber="5">
<emphasis id="D1AAE8252A56FFDB2AE2C21AFE9FF8DE" bold="true" box="[189,275,1843,1873]" pageId="4" pageNumber="5">Fig. 1.</emphasis>
Flowers. 
<emphasis id="D1AAE8252A56FFDB2BC9C21AFE21F8DF" bold="true" box="[406,429,1843,1872]" pageId="4" pageNumber="5">A</emphasis>
, 
<emphasis id="D1AAE8252A56FFDB2BE0C21AFE5AF8DF" bold="true" box="[447,470,1843,1872]" pageId="4" pageNumber="5">C</emphasis>
, 
<emphasis id="D1AAE8252A56FFDB2BB7C21DFE71F8DF" bold="true" box="[488,509,1844,1872]" pageId="4" pageNumber="5">E</emphasis>
. Top view of the flower. 
<emphasis id="D1AAE8252A56FFDB2916C21DFCD2F8DF" bold="true" box="[841,862,1844,1872]" pageId="4" pageNumber="5">B</emphasis>
, 
<emphasis id="D1AAE8252A56FFDB292EC21DFC04F8DF" bold="true" box="[881,904,1844,1872]" pageId="4" pageNumber="5">D</emphasis>
, 
<emphasis id="D1AAE8252A56FFDB29C5C21DFC22F8DF" bold="true" box="[922,942,1844,1872]" pageId="4" pageNumber="5">F</emphasis>
. Side view of the flower, with part of corolla removed. 
<emphasis id="D1AAE8252A56FFDB2B9EC273FE70F8F9" bold="true" box="[449,508,1882,1911]" pageId="4" pageNumber="5">A–B</emphasis>
. 
<taxonomicName id="24DE4FB42A56FFDB2852C272FB08F8F8" authority="S.Rahayu &amp; Rodda" authorityName="Rahayu &amp; Rodda" authorityYear="2019" box="[525,1156,1882,1911]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="subSpecies" species="danumensis" status="subsp. nov." subSpecies="amarii">
<emphasis id="D1AAE8252A56FFDB2852C272FD47F8F8" box="[525,715,1882,1911]" italics="true" pageId="4" pageNumber="5">H. danumensis</emphasis>
subsp. 
<emphasis id="D1AAE8252A56FFDB2972C272FC0EF8F8" box="[813,898,1883,1911]" italics="true" pageId="4" pageNumber="5">amarii</emphasis>
S.Rahayu &amp; Rodda
</taxonomicName>
<taxonomicNameLabel id="CA99555E2A56FFDB2ED4C273FA97F8F8" box="[1163,1307,1882,1911]" pageId="4" pageNumber="5" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
(Amar Husein Sitompul s.n. leg., BO). 
<emphasis id="D1AAE8252A56FFDB280DC2A9FD02F812" bold="true" box="[594,654,1920,1949]" pageId="4" pageNumber="5">C–D</emphasis>
. 
<taxonomicName id="24DE4FB42A56FFDB28C2C2A8FBCFF812" authority="S.Rahayu &amp; Rodda" authorityName="Rahayu &amp; Rodda" authorityYear="2019" box="[669,1091,1920,1950]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="species" species="solokensis" status="sp. nov.">
<emphasis id="D1AAE8252A56FFDB28C2C2A8FCCBF812" box="[669,839,1920,1949]" italics="true" pageId="4" pageNumber="5">H. solokensis</emphasis>
S.Rahayu &amp; Rodda
</taxonomicName>
<taxonomicNameLabel id="CA99555E2A56FFDB2E15C2A8FB27F812" box="[1098,1195,1921,1949]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
(Fadly s.n. leg., BO). 
<emphasis id="D1AAE8252A56FFDB2B5BC28EFEB0F84C" bold="true" box="[260,316,1959,1987]" pageId="4" pageNumber="5">E–F</emphasis>
. 
<taxonomicName id="24DE4FB42A56FFDB2B14C281FD7FF84B" authority="S.Rahayu &amp; Rodda" authorityName="Rahayu &amp; Rodda" authorityYear="2019" box="[331,755,1959,1988]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="species" species="rigidifolia" status="sp. nov.">
<emphasis id="D1AAE8252A56FFDB2B14C281FE7FF84B" box="[331,499,1959,1988]" italics="true" pageId="4" pageNumber="5">H. rigidifolia</emphasis>
S.Rahayu &amp; Rodda
</taxonomicName>
<taxonomicNameLabel id="CA99555E2A56FFDB28A5C28EFCD0F84C" box="[762,860,1959,1987]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
(S. Rahayu 865 leg., BO). Drawing: X. Y. Loh.
</paragraph>
</caption>
<caption id="B7A164BF2A57FFDA2AE2C203FBAAF86E" httpUri="https://zenodo.org/record/2612879/files/figure.png" pageId="5" pageNumber="6" startId="5.[189,236,1834,1863]" targetBox="[290,1298,265,1795]" targetPageId="5">
<paragraph id="E36134372A57FFDA2AE2C203FBAAF86E" blockId="5.[189,1399,1834,2017]" pageId="5" pageNumber="6">
<emphasis id="D1AAE8252A57FFDA2AE2C203FE9BF8C8" bold="true" box="[189,279,1834,1863]" pageId="5" pageNumber="6">Fig. 2.</emphasis>
Inflorescence, pollinarium, and leaf. 
<emphasis id="D1AAE8252A57FFDA2949C203FCA1F8C8" bold="true" box="[790,813,1834,1863]" pageId="5" pageNumber="6">A</emphasis>
, 
<emphasis id="D1AAE8252A57FFDA291CC203FCD6F8C8" bold="true" box="[835,858,1834,1863]" pageId="5" pageNumber="6">C</emphasis>
, 
<emphasis id="D1AAE8252A57FFDA292FC203FC09F8C9" bold="true" box="[880,901,1834,1862]" pageId="5" pageNumber="6">E</emphasis>
. Inflorescences. 
<emphasis id="D1AAE8252A57FFDA2E35C203FBF3F8C9" bold="true" box="[1130,1151,1834,1862]" pageId="5" pageNumber="6">B</emphasis>
, 
<emphasis id="D1AAE8252A57FFDA2EC9C203FB21F8C9" bold="true" box="[1174,1197,1834,1862]" pageId="5" pageNumber="6">D</emphasis>
, 
<emphasis id="D1AAE8252A57FFDA2E9CC203FB50F8C8" bold="true" box="[1219,1244,1834,1863]" pageId="5" pageNumber="6">G</emphasis>
. Pollinaria. 
<emphasis id="D1AAE8252A57FFDA2AE2C278FF5DF8E2" bold="true" box="[189,209,1873,1901]" pageId="5" pageNumber="6">F</emphasis>
. Leaf. 
<emphasis id="D1AAE8252A57FFDA2B6BC279FEE2F8E2" bold="true" box="[308,366,1872,1901]" pageId="5" pageNumber="6">A–B</emphasis>
. 
<taxonomicName id="24DE4FB42A57FFDA2BDCC27BFB9DF8E2" authority="S.Rahayu &amp; Rodda" authorityName="Rahayu &amp; Rodda" authorityYear="2019" box="[387,1041,1872,1902]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="subSpecies" species="danumensis" status="subsp. nov." subSpecies="amarii">
<emphasis id="D1AAE8252A57FFDA2BDCC27BFDC9F8E1" box="[387,581,1873,1902]" italics="true" pageId="5" pageNumber="6">H. danumensis</emphasis>
subsp. 
<emphasis id="D1AAE8252A57FFDA28F0C27BFC88F8E1" box="[687,772,1874,1902]" italics="true" pageId="5" pageNumber="6">amarii</emphasis>
S.Rahayu &amp; Rodda
</taxonomicName>
<taxonomicNameLabel id="CA99555E2A57FFDA2E42C279FB3CF8E2" box="[1053,1200,1872,1901]" pageId="5" pageNumber="6" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
(Amar Husein Sitompul s.n. leg., BO). 
<emphasis id="D1AAE8252A57FFDA2BA8C25EFDB8F81C" bold="true" box="[503,564,1911,1940]" pageId="5" pageNumber="6">C–D</emphasis>
. 
<taxonomicName id="24DE4FB42A57FFDA281AC251FC7FF81B" authority="S.Rahayu &amp; Rodda" authorityName="Rahayu &amp; Rodda" authorityYear="2019" box="[581,1011,1911,1940]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="solokensis" status="sp. nov.">
<emphasis id="D1AAE8252A57FFDA281AC251FD7DF81B" box="[581,753,1911,1940]" italics="true" pageId="5" pageNumber="6">H. solokensis</emphasis>
S.Rahayu &amp; Rodda
</taxonomicName>
<taxonomicNameLabel id="CA99555E2A57FFDA29A4C251FBD2F81B" box="[1019,1118,1912,1940]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
(Fadly s.n. leg., BO). 
<emphasis id="D1AAE8252A57FFDA2AE2C2B7FF76F835" bold="true" box="[189,250,1949,1978]" pageId="5" pageNumber="6">E–G</emphasis>
. 
<taxonomicName id="24DE4FB42A57FFDA2B57C2B7FD20F835" authority="S.Rahayu &amp; Rodda" authorityName="Rahayu &amp; Rodda" authorityYear="2019" box="[264,684,1949,1979]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="rigidifolia" status="sp. nov.">
<emphasis id="D1AAE8252A57FFDA2B57C2B7FE3CF835" box="[264,432,1949,1978]" italics="true" pageId="5" pageNumber="6">H. rigidifolia</emphasis>
S.Rahayu &amp; Rodda
</taxonomicName>
<taxonomicNameLabel id="CA99555E2A57FFDA28EDC2B7FC98F835" box="[690,788,1950,1978]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
(S. Rahayu 865 leg., BO). Photos: Amar Husein Sitompul (A), Fadly (C), S. Rahayu (E 
<emphasis id="D1AAE8252A57FFDA28F3C2EFFD37F86F" bold="true" box="[684,699,1990,2016]" pageId="5" pageNumber="6">–</emphasis>
F), and M. Rodda (B, D, G).
</paragraph>
</caption>
<paragraph id="E36134372A54FFD92AE2C422FE8FFEAA" blockId="6.[189,259,267,293]" box="[189,259,267,293]" pageId="6" pageNumber="7">
<heading id="B829835B2A54FFD92AE2C422FE8FFEAA" bold="true" box="[189,259,267,293]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="D1AAE8252A54FFD92AE2C422FE8FFEAA" bold="true" box="[189,259,267,293]" pageId="6" pageNumber="7">Notes</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="ABC467BC2A54FFD92AE2C412FCBEFE6A" pageId="6" pageNumber="7" type="reference_group">
<paragraph id="E36134372A54FFD92AE2C412FCBEFE6A" blockId="6.[189,1406,314,486]" pageId="6" pageNumber="7">
<taxonomicName id="24DE4FB42A54FFD92AE2C412FDB1FEDB" ID-CoL="7ZBVZ" authority="Rahayu &amp; Rodda, 2019" authorityName="Rahayu &amp; Rodda" authorityYear="2019" box="[189,573,314,340]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="subSpecies" species="danumensis" status="subsp. nov." subSpecies="amarii">
<emphasis id="D1AAE8252A54FFD92AE2C412FE1DFEDB" box="[189,401,314,340]" italics="true" pageId="6" pageNumber="7">Hoya danumensis</emphasis>
subsp. 
<emphasis id="D1AAE8252A54FFD92BB0C412FDB1FEDB" box="[495,573,315,340]" italics="true" pageId="6" pageNumber="7">amarii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA99555E2A54FFD92818C413FD41FEDA" box="[583,717,314,341]" pageId="6" pageNumber="7" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
can be separated from 
<taxonomicName id="24DE4FB42A54FFD929B6C412FE7EFEF7" ID-CoL="7ZBW2" authority="Rodda &amp; Nyhuus (2009) subsp. danumensis" authorityName="Rodda &amp; Nyhuus" authorityYear="2009" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="subSpecies" species="danumensis" subSpecies="danumensis">
<emphasis id="D1AAE8252A54FFD929B6C412FB17FEDB" box="[1001,1179,314,340]" italics="true" pageId="6" pageNumber="7">H. danumensis</emphasis>
<bibRefCitation id="874F49C62A54FFD92EFBC413FE87FEF6" author="Rodda M. &amp; Nyhuus T." pageId="6" pageNumber="7" pagination="163 - 167" refId="ref9664" refString="Rodda M. &amp; Nyhuus T. 2009. Hoya danumensis, a new species of Hoya (Apocynaceae, Asclepiadoideae) from Borneo. Webbia 64: 163 - 167. https: // doi. org / 10.1080 / 00837792.2009.10670856" type="journal article" year="2009">Rodda &amp; Nyhuus (2009)</bibRefCitation>
subsp. 
<emphasis id="D1AAE8252A54FFD92B38C477FE7EFEF7" box="[359,498,350,376]" italics="true" pageId="6" pageNumber="7">danumensis</emphasis>
</taxonomicName>
because of the shape and size of the corolla that is shallowly campanulate, 
<quantity id="242699D22A54FFD92AE2C4AAFEB0FE12" box="[189,316,387,413]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" metricValueMax="2.3" metricValueMin="2.0" pageId="6" pageNumber="7" unit="mm" value="21.5" valueMax="23.0" valueMin="20.0">20–23 mm</quantity>
in diameter and deeply campanulate, vs 
<quantity id="242699D22A54FFD92943C4ABFC17FE12" box="[796,923,386,413]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="6" pageNumber="7" unit="mm" value="30.0" valueMax="35.0" valueMin="25.0">25–35 mm</quantity>
in the latter; in the shape and size of the corona lobes that are ovate-oblong and 5.5–6 × 
<quantity id="242699D22A54FFD9295FC48EFC1CFE4E" box="[768,912,423,449]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.6" metricValueMin="2.4" pageId="6" pageNumber="7" unit="mm" value="2.5" valueMax="2.6" valueMin="2.4">2.4–2.6 mm</quantity>
in 
<taxonomicName id="24DE4FB42A54FFD929E5C481FADFFE4E" ID-CoL="7ZBW2" authority="subsp. danumensis" box="[954,1363,423,449]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="6" pageNumber="7" phylum="Tracheophyta" rank="subSpecies" species="danumensis" subSpecies="danumensis">
<emphasis id="D1AAE8252A54FFD929E5C481FBE7FE4E" box="[954,1131,423,449]" italics="true" pageId="6" pageNumber="7">H. danumensis</emphasis>
subsp. 
<emphasis id="D1AAE8252A54FFD92E97C48EFADFFE4E" box="[1224,1363,423,449]" italics="true" pageId="6" pageNumber="7">danumensis</emphasis>
</taxonomicName>
vs ovate and 4.5–5 × 
<quantity id="242699D22A54FFD92BC8C4E2FDA8FE69" box="[407,548,459,486]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="2.5" metricValueMin="2.3" pageId="6" pageNumber="7" unit="mm" value="2.4" valueMax="2.5" valueMin="2.3">2.3–2.5 mm</quantity>
in the new subspecies.
</paragraph>
</subSubSection>
</treatment>
</document>