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<document id="7B29D7F509B3BA9C8E24D29A1C2D77CC" ID-CLB-Dataset="21944" ID-DOI="10.5852/ejt.2014.102" ID-GBIF-Dataset="6fb40ed6-8bf1-4bd2-8602-12f477b1dfad" ID-ISSN="2118-9773" ID-Zenodo-Dep="3838819" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590096682365" checkinUser="tatiana" docAuthor="Constant, Jérôme" docDate="2014" docId="64738955C410DA02D4F402B73823F86C" docLanguage="en" docName="ejt-2014-102.pdf.imf" docOrigin="European Journal of Taxonomy 102" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Belbina madagascariensis Constant 2014, comb. nov." docType="treatment" docVersion="6" lastPageNumber="27" masterDocId="984AF12DC409DA18D5330357393BFFBB" masterDocTitle="Revision of the Malagasy lanternfly genus Belbina Stål, 1863, with two new species (Hemiptera: Fulgoromorpha: Fulgoridae)" masterLastPageNumber="37" masterPageNumber="1" pageNumber="26" updateTime="1698840770983" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="7F93B2023BEA36FECBA8C73AD10A4D66">Revision of the Malagasy lanternfly genus Belbina Stål, 1863, with two new species (Hemiptera: Fulgoromorpha: Fulgoridae)</mods:title>
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<mods:namePart id="8A6981FA1A23817ECBB21EC71F456A33">Constant, Jérôme</mods:namePart>
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<mods:title id="97F45FA0102D80CB5BB3A19289F8929B">European Journal of Taxonomy</mods:title>
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<mods:date id="7E848400392095AF83D293601D503E1C">2014</mods:date>
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<treatment id="64738955C410DA02D4F402B73823F86C" ID-DOI="http://doi.org/10.5281/zenodo.3852493" ID-GBIF-Taxon="164248031" ID-Zenodo-Dep="3852493" LSID="urn:lsid:plazi:treatment:64738955C410DA02D4F402B73823F86C" httpUri="http://treatment.plazi.org/id/64738955C410DA02D4F402B73823F86C" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
<subSubSection id="A4C06BC8C410DA01D4F402B73D56FE41" box="[455,1133,480,507]" pageId="25" pageNumber="26" type="nomenclature">
<paragraph id="EC653843C410DA01D4F402B73D56FE41" blockId="25.[455,1133,480,541]" box="[455,1133,480,507]" pageId="25" pageNumber="26">
<heading id="B72D8F2FC410DA01D4F402B73D56FE41" box="[455,1133,480,507]" centered="true" fontSize="11" level="2" pageId="25" pageNumber="26" reason="2">
<taxonomicName id="2BDA43C0C410DA01D4F402B73ADFFE40" ID-CoL="L9RW" authority="(Westwood, 1851)" authorityName="Constant" authorityYear="2014" baseAuthorityName="Westwood" baseAuthorityYear="1851" box="[455,996,480,507]" class="Insecta" family="Fulgoridae" genus="Belbina" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="madagascariensis" status="comb. nov.">
<emphasis id="DEAEE451C410DA01D4F402B73A3FFE40" bold="true" box="[455,772,480,507]" italics="true" pageId="25" pageNumber="26">Belbina madagascariensis</emphasis>
(
<bibRefCitation id="884B45B2C410DA01D62002B73AE0FE40" author="Westwood J. O." box="[787,987,480,507]" pageId="25" pageNumber="26" pagination="207 - 210" refId="ref19581" refString="Westwood J. O. 1851. Description of some new species of exotic homopterous insects. Annals and Magazine of Natural History, Series 2, 7: 207 - 210. http: // biodiversitylibrary. org / page / 13787797" type="journal article" year="1851">Westwood, 1851</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="C59D592AC410DA01D6D802B73D56FE41" box="[1003,1133,480,506]" pageId="25" pageNumber="26" rank="species">comb. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A4C06BC8C410DA01D7AA01543AA0FDA6" box="[665,923,515,541]" pageId="25" pageNumber="26" type="description">
<paragraph id="EC653843C410DA01D7AA01543AA0FDA6" blockId="25.[455,1133,480,541]" box="[665,923,515,541]" pageId="25" pageNumber="26">
<figureCitation id="74E124C6C410DA01D7AA01543A21FDA6" box="[665,794,515,541]" captionStart="Figs 79. — 7" captionStartId="8.[189,241,1869,1895]" captionTargetBox="[189,1398,277,1823]" captionTargetId="figure@8.[189,1398,271,1829]" captionTargetPageId="8" captionText="Figs 79. — 7. Belbina lambertoni. 8. B. madagascariensis. 9. B. nympha. — A. Habitus, dorsal view. B. Habitus, lateral view. C. Vertex, pro- and mesonotum, dorsal view. D. Head and prothorax, lateral view. E. Frons, normal view. — Scale bar: 10 mm (CE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3838825" httpUri="https://zenodo.org/record/3838825/files/figure.png" pageId="25" pageNumber="26">Figs 8AE</figureCitation>
,
<figureCitation id="74E124C6C410DA01D61501543A49FDA6" box="[806,882,515,541]" captionStart="Figs 2429" captionStartId="12.[189,241,1928,1954]" captionTargetBox="[189,1398,498,1887]" captionTargetId="figure@12.[189,1398,498,1887]" captionTargetPageId="12" captionText="Figs 2429. Male genitalia. — 2425. Belbina lambertoni. 24. Lateral view. 25. Dorsal view. — 26 27. B. madagascariensis. 26. Lateral view. 27. Dorsal view. — 2829. B. nympha. 28. Lateral view. 29. Dorsal view. — An = anal tube, Py = pygofer, G = gonostylus. Photographs by Y. Laurent &amp; I. Bachy." figureDoi="http://doi.org/10.5281/zenodo.3838833" httpUri="https://zenodo.org/record/3838833/files/figure.png" pageId="25" pageNumber="26">2627</figureCitation>
,
<figureCitation id="74E124C6C410DA01D64D01543AA0FDA6" box="[894,923,515,541]" captionStart="Figs 4649" captionStartId="32.[189,241,1963,1989]" captionTargetBox="[297,1284,265,1915]" captionTargetId="figure@32.[296,1292,265,1921]" captionTargetPageId="32" captionText="Figs 4649. Distribution maps. — 46. Belbina bergrothi, B. bloetei and B. madagascariensis. 47. B. bourgoini sp. nov., B. falleni and B. foliacea. 48. B. laetitiae sp. nov. 49. B. lambertoni." figureDoi="http://doi.org/10.5281/zenodo.3838841" httpUri="https://zenodo.org/record/3838841/files/figure.png" pageId="25" pageNumber="26">46</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="A4C06BC8C410DA01D58E01123A94FD5C" pageId="25" pageNumber="26" type="reference_group">
<paragraph id="EC653843C410DA01D58E01123AC2FDE4" blockId="25.[189,1017,580,607]" box="[189,1017,580,607]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="CCCA1F6DC410DA01D58E01123AC2FDE4" box="[189,1017,580,607]" pageId="25" pageNumber="26">
<treatmentCitation id="6D7B1E52C410DA01D58E01123BC1FDE4" author="Westwood J. O." box="[189,762,580,607]" page="208" pageId="25" pageNumber="26" year="1851">
<taxonomicName id="2BDA43C0C410DA01D58E01123BC1FDE4" authority="Westwood, 1851: 208" authorityName="Westwood" authorityPageNumber="208" authorityYear="1851" box="[189,762,580,607]" class="Insecta" family="Fulgoridae" genus="Aphana" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="DEAEE451C410DA01D58E011238CFFDE4" box="[189,500,581,607]" italics="true" pageId="25" pageNumber="26">Aphana madagascariensis</emphasis>
<bibRefCitation id="884B45B2C410DA01D4C901123BC1FDE4" author="Westwood J. O." box="[506,762,580,607]" pageId="25" pageNumber="26" pagination="207 - 210" refId="ref19581" refString="Westwood J. O. 1851. Description of some new species of exotic homopterous insects. Annals and Magazine of Natural History, Series 2, 7: 207 - 210. http: // biodiversitylibrary. org / page / 13787797" type="journal article" year="1851">Westwood, 1851: 208</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
<typeStatus id="336186E1C410DA01D63A01123A53FDE4" box="[777,872,581,607]" pageId="25" pageNumber="26" type="neotype">neotype</typeStatus>
in MNHN).
</treatmentCitationGroup>
</paragraph>
<paragraph id="EC653843C410DA01D58E01D13D3AFD7F" blockId="25.[189,1399,646,743]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="CCCA1F6DC410DA01D58E01D13D3AFD7F" pageId="25" pageNumber="26">
<taxonomicName id="2BDA43C0C410DA01D58E01D138C8FD1B" authorityName="Westwood" authorityYear="1851" box="[189,499,646,672]" class="Insecta" family="Fulgoridae" genus="Aphana" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="DEAEE451C410DA01D58E01D138C8FD1B" box="[189,499,646,672]" italics="true" pageId="25" pageNumber="26">Aphana madagascariensis</emphasis>
</taxonomicName>
<treatmentCitation id="6D7B1E52C410DA01D73C01D13BD6FD1B" author="Walker F." box="[527,749,646,673]" page="1134" pageId="25" pageNumber="26" year="1852">
<bibRefCitation id="884B45B2C410DA01D73C01D13BD6FD1B" author="Walker F." box="[527,749,646,673]" pageId="25" pageNumber="26" pagination="1119 - 1188" refId="ref19538" refString="Walker F. 1852. Supplement. List of the Species of Homopterous Insects in the Collection of the British Museum 4: 1119 - 1188. http: // dx. doi. org / 10.5962 / bhl. title. 9063" type="journal article" year="1852">Walker 1852: 1134</bibRefCitation>
</treatmentCitation>
(listed). —
<treatmentCitation id="6D7B1E52C410DA01D64A01D13D18FD1B" author="Stal C." box="[889,1059,646,673]" page="263" pageId="25" pageNumber="26" year="1866">
<bibRefCitation id="884B45B2C410DA01D64A01D13D18FD1B" author="Stal C." box="[889,1059,646,673]" pageId="25" pageNumber="26" pagination="1 - 276" refId="ref19413" refString="Stal C. 1866. Hemiptera Homoptera Latr. Hemiptera Africana 4: 1 - 276. http: // biodiversitylibrary. org / page / 8695280" type="journal article" year="1866">Stål 1866: 263</bibRefCitation>
</treatmentCitation>
(listed). —
<treatmentCitation id="6D7B1E52C410DA01D19C01D139E1FD7F" author="Lallemand V." page="85" pageId="25" pageNumber="26" year="1959">
<bibRefCitation id="884B45B2C410DA01D19C01D139E1FD7F" author="Lallemand V." pageId="25" pageNumber="26" pagination="37 - 124" refId="ref18997" refString="Lallemand V. 1959. Revision des especes africaines de la famille Fulgoridae (super-famille Fulgoroides - sous-ordre des homopteres). Publicacoes Culturais da Companhia de Diamantes de Angola 41: 37 - 124." type="journal article" year="1959">Lallemand 1959: 85</bibRefCitation>
</treatmentCitation>
(description, comments). —
<treatmentCitation id="6D7B1E52C410DA01D70101FE3A23FD78" author="Constant J." box="[562,792,681,707]" page="31" pageId="25" pageNumber="26" year="2004">
<bibRefCitation id="884B45B2C410DA01D70101FE3A23FD78" author="Constant J." box="[562,792,681,707]" pageId="25" pageNumber="26" pagination="29 - 34" refId="ref18545" refString="Constant J. 2004 b. Fulgoridae of Madagascar: description of a new species representing a new genus, key to the genera and list of the species (Homoptera: Fulgoromorpha: Fulgoridae). Bulletin de l'Institut royal des Sciences naturelles de Belgique 74: 29 - 34." type="journal article" year="2004">Constant 2004b: 31</bibRefCitation>
</treatmentCitation>
(listed, comments).
</treatmentCitationGroup>
</paragraph>
<paragraph id="EC653843C410DA01D58E019B3A94FD5C" blockId="25.[189,1399,646,743]" box="[189,943,716,743]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="CCCA1F6DC410DA01D58E019B3A94FD5C" box="[189,943,716,743]" pageId="25" pageNumber="26">
<taxonomicName id="2BDA43C0C410DA01D58E019B3B2CFD5D" ID-CoL="F9SV" baseAuthorityName="Westwood" baseAuthorityYear="1851" box="[189,535,716,742]" class="Insecta" family="Fulgoridae" genus="Aphaenina" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="DEAEE451C410DA01D58E019B3B2CFD5D" box="[189,535,716,742]" italics="true" pageId="25" pageNumber="26">Aphaenina madagascariensis</emphasis>
</taxonomicName>
<treatmentCitation id="6D7B1E52C410DA01D706019B3A36FD5C" author="Metcalf Z. P." box="[565,781,716,743]" page="152" pageId="25" pageNumber="26" year="1947">
<bibRefCitation id="884B45B2C410DA01D706019B3A36FD5C" author="Metcalf Z. P." box="[565,781,716,743]" pageId="25" pageNumber="26" refId="ref19123" refString="Metcalf Z. P. 1947. General Catalogue of the Homoptera. Fascicle IV. Fulgoroidea. Part 9. Fulgoridae. North Carolina State College, Raleigh." type="book" year="1947">Metcalf 1947: 152</bibRefCitation>
</treatmentCitation>
(catalogued).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A4C06BC8C410DA01D58E00593D04FC26" pageId="25" pageNumber="26" type="diagnosis">
<paragraph id="EC653843C410DA01D58E005938F7FC93" blockId="25.[189,460,782,808]" box="[189,460,782,808]" pageId="25" pageNumber="26">
<heading id="B72D8F2FC410DA01D58E005938F7FC93" bold="true" box="[189,460,782,808]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="DEAEE451C410DA01D58E005938F7FC93" bold="true" box="[189,460,782,808]" pageId="25" pageNumber="26">Diagnostic characters</emphasis>
</heading>
</paragraph>
<paragraph id="EC653843C410DA01D58E006A3D04FC26" blockId="25.[189,1399,828,926]" pageId="25" pageNumber="26">
(1) disc of tegmina red or orange (
<figureCitation id="74E124C6C410DA01D747006B3BECFCEC" box="[628,727,828,855]" captionStart="Figs 79. — 7" captionStartId="8.[189,241,1869,1895]" captionTargetBox="[189,1398,277,1823]" captionTargetId="figure@8.[189,1398,271,1829]" captionTargetPageId="8" captionText="Figs 79. — 7. Belbina lambertoni. 8. B. madagascariensis. 9. B. nympha. — A. Habitus, dorsal view. B. Habitus, lateral view. C. Vertex, pro- and mesonotum, dorsal view. D. Head and prothorax, lateral view. E. Frons, normal view. — Scale bar: 10 mm (CE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3838825" httpUri="https://zenodo.org/record/3838825/files/figure.png" pageId="25" pageNumber="26">Fig. 8A</figureCitation>
); (2) cephalic process elongate and narrow, projecting anterodorsad (
<figureCitation id="74E124C6C410DA01D454003738D2FCC1" box="[359,489,864,890]" captionStart="Figs 79. — 7" captionStartId="8.[189,241,1869,1895]" captionTargetBox="[189,1398,277,1823]" captionTargetId="figure@8.[189,1398,271,1829]" captionTargetPageId="8" captionText="Figs 79. — 7. Belbina lambertoni. 8. B. madagascariensis. 9. B. nympha. — A. Habitus, dorsal view. B. Habitus, lateral view. C. Vertex, pro- and mesonotum, dorsal view. D. Head and prothorax, lateral view. E. Frons, normal view. — Scale bar: 10 mm (CE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3838825" httpUri="https://zenodo.org/record/3838825/files/figure.png" pageId="25" pageNumber="26">Fig. 8DE</figureCitation>
); (3) ground colour of tegmina brown (
<figureCitation id="74E124C6C410DA01D6F500373D1DFCC1" box="[966,1062,864,890]" captionStart="Figs 79. — 7" captionStartId="8.[189,241,1869,1895]" captionTargetBox="[189,1398,277,1823]" captionTargetId="figure@8.[189,1398,271,1829]" captionTargetPageId="8" captionText="Figs 79. — 7. Belbina lambertoni. 8. B. madagascariensis. 9. B. nympha. — A. Habitus, dorsal view. B. Habitus, lateral view. C. Vertex, pro- and mesonotum, dorsal view. D. Head and prothorax, lateral view. E. Frons, normal view. — Scale bar: 10 mm (CE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3838825" httpUri="https://zenodo.org/record/3838825/files/figure.png" pageId="25" pageNumber="26">Fig. 8A</figureCitation>
); (4) large-sized (more than
<quantity id="2B2295A6C410DA01D58E00D4382BFC26" box="[189,272,899,925]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="25" pageNumber="26" unit="mm" value="26.0">26 mm</quantity>
long); (5) black-brown spots with white center on tegmina (
<figureCitation id="74E124C6C410DA01D6E000D43D0AFC26" box="[979,1073,899,925]" captionStart="Figs 79. — 7" captionStartId="8.[189,241,1869,1895]" captionTargetBox="[189,1398,277,1823]" captionTargetId="figure@8.[189,1398,271,1829]" captionTargetPageId="8" captionText="Figs 79. — 7. Belbina lambertoni. 8. B. madagascariensis. 9. B. nympha. — A. Habitus, dorsal view. B. Habitus, lateral view. C. Vertex, pro- and mesonotum, dorsal view. D. Head and prothorax, lateral view. E. Frons, normal view. — Scale bar: 10 mm (CE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3838825" httpUri="https://zenodo.org/record/3838825/files/figure.png" pageId="25" pageNumber="26">Fig. 8A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A4C06BC8C410DA01D58E00933D89F91E" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="EC653843C410DA01D58E00933891FC65" blockId="25.[189,426,964,990]" box="[189,426,964,990]" pageId="25" pageNumber="26">
<heading id="B72D8F2FC410DA01D58E00933891FC65" bold="true" box="[189,426,964,990]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="DEAEE451C410DA01D58E00933891FC65" bold="true" box="[189,426,964,990]" pageId="25" pageNumber="26">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="EC653843C410DA01D5D900A4386AFBB6" blockId="25.[189,1399,1011,1565]" box="[234,337,1011,1037]" pageId="25" pageNumber="26">
<emphasis id="DEAEE451C410DA01D5D900A4386AFBB6" bold="true" box="[234,337,1011,1037]" pageId="25" pageNumber="26">
<typeStatus id="336186E1C410DA01D5D900A4386AFBB6" box="[234,337,1011,1037]" pageId="25" pageNumber="26" type="neotype">Neotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="EC653843C410DA01D58E07413D06FB21" blockId="25.[189,1399,1011,1565]" pageId="25" pageNumber="26">
<materialsCitation id="5CB2321EC410DA01D58E07413D06FB21" ID-GBIF-Occurrence="2621775310" collectionCode="MNHN" country="Madagascar" determinerName="Jerome Constant" location="MADAGASCAR" pageId="25" pageNumber="26" specimenCount="1" specimenCount-female="1" typeStatus="neotype">
<collectingCountry id="94CD78D3C410DA01D58E074138A9FB8B" box="[189,402,1046,1072]" name="Madagascar" pageId="25" pageNumber="26">MADAGASCAR</collectingCountry>
:
<specimenCount id="FADCF3CAC410DA01D49B07403885FB8A" box="[424,446,1047,1073]" pageId="25" pageNumber="26" type="female"></specimenCount>
,
<typeStatus id="336186E1C410DA01D4E407403B0FFB8A" box="[471,564,1047,1073]" pageId="25" pageNumber="26" type="neotype">neotype</typeStatus>
of
<taxonomicName id="2BDA43C0C410DA01D74207413DADFB8A" authority="Westwood, 1851" authorityName="Westwood" authorityYear="1851" box="[625,1174,1046,1073]" class="Insecta" family="Fulgoridae" genus="Aphana" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="DEAEE451C410DA01D74207413A88FB8B" box="[625,947,1046,1072]" italics="true" pageId="25" pageNumber="26">Aphana madagascariensis</emphasis>
<bibRefCitation id="884B45B2C410DA01D6F707413DADFB8A" author="Westwood J. O." box="[964,1174,1046,1073]" pageId="25" pageNumber="26" pagination="207 - 210" refId="ref19581" refString="Westwood J. O. 1851. Description of some new species of exotic homopterous insects. Annals and Magazine of Natural History, Series 2, 7: 207 - 210. http: // biodiversitylibrary. org / page / 13787797" type="journal article" year="1851">Westwood, 1851</bibRefCitation>
</taxonomicName>
, here designated, [Goudot Madagascar 1832] [Museum Paris] [
<emphasis id="DEAEE451C410DA01D630076E3AABFBE8" box="[771,912,1081,1107]" italics="true" pageId="25" pageNumber="26">Pterobaptes</emphasis>
. Amyot.-
<emphasis id="DEAEE451C410DA01D13B076D3DB9FBE8" box="[1032,1154,1082,1108]" italics="true" pageId="25" pageNumber="26">vari -egata</emphasis>
Spin. A. Fr. VIII. 225.-Senegal] [Neotype ♀
<taxonomicName id="2BDA43C0C410DA01D4C8070B3DBCFBCD" authority="Westwood, 1851 J. Constant" authorityName="Westwood" authorityYear="1851" box="[507,1159,1116,1143]" class="Insecta" family="Fulgoridae" genus="Aphana" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="DEAEE451C410DA01D4C8070B3A09FBCD" box="[507,818,1116,1142]" italics="true" pageId="25" pageNumber="26">Aphana madagascariensis</emphasis>
<bibRefCitation id="884B45B2C410DA01D60A070B3AC5FBCC" author="Westwood J. O." box="[825,1022,1116,1143]" pageId="25" pageNumber="26" pagination="207 - 210" refId="ref19581" refString="Westwood J. O. 1851. Description of some new species of exotic homopterous insects. Annals and Magazine of Natural History, Series 2, 7: 207 - 210. http: // biodiversitylibrary. org / page / 13787797" type="journal article" year="1851">Westwood, 1851</bibRefCitation>
J. Constant
</taxonomicName>
des. 2013] [
<taxonomicName id="2BDA43C0C410DA01D028070B3A81FB21" authority="(Westwood, 1851) Det. Jerome Constant 2013" authorityName="Det. Jerome Constant" authorityYear="2013" baseAuthorityName="Westwood" baseAuthorityYear="1851" class="Insecta" family="Fulgoridae" genus="Belbina" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="DEAEE451C410DA01D028070B38AAFB21" italics="true" pageId="25" pageNumber="26">Belbina madagascariensis</emphasis>
(
<bibRefCitation id="884B45B2C410DA01D49307283B53FB21" author="Westwood J. O." box="[416,616,1151,1178]" pageId="25" pageNumber="26" pagination="207 - 210" refId="ref19581" refString="Westwood J. O. 1851. Description of some new species of exotic homopterous insects. Annals and Magazine of Natural History, Series 2, 7: 207 - 210. http: // biodiversitylibrary. org / page / 13787797" type="journal article" year="1851">Westwood, 1851</bibRefCitation>
) Det.
<determinerName id="19B9D487C410DA01D78307D73A4EFB21" box="[688,885,1152,1178]" pageId="25" pageNumber="26">Jérôme Constant</determinerName>
2013
</taxonomicName>
] (
<collectionCode id="8ACBA086C410DA01D6E307D73D0AFB21" box="[976,1073,1152,1178]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="25" pageNumber="26">MNHN</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="EC653843C410DA01D58E07F43BCAF9A6" blockId="25.[189,1399,1011,1565]" pageId="25" pageNumber="26">
Note: A
<typeStatus id="336186E1C410DA01D41107F43844FB06" box="[290,383,1187,1213]" pageId="25" pageNumber="26" type="neotype">neotype</typeStatus>
is here designated in order to stabilise the nomenclature in the group, following rule 75.3 of the International Code of Zoological Nomenclature (
<bibRefCitation id="884B45B2C410DA01D6B007913D2BFB5B" author="ICZN" box="[899,1040,1222,1248]" pageId="25" pageNumber="26" refId="ref18755" refString="ICZN. 1999. International Code of Zoological Nomenclature. Fourth Edition. The International Trust for Zoological Nomenclature, London, U. K. http: // iczn. org / iczn / index. jsp" type="url" year="1999">ICZN 1999</bibRefCitation>
). The specimen chosen here is probably the one on which
<bibRefCitation id="884B45B2C410DA01D73507BE3BE2FABF" author="Westwood J. O." box="[518,729,1257,1284]" pageId="25" pageNumber="26" pagination="207 - 210" refId="ref19581" refString="Westwood J. O. 1851. Description of some new species of exotic homopterous insects. Annals and Magazine of Natural History, Series 2, 7: 207 - 210. http: // biodiversitylibrary. org / page / 13787797" type="journal article" year="1851">Westwood (1851)</bibRefCitation>
based his description. It was collected by Jules Prosper Goudot in 1832 and included in the collections of the MNHN in 1834 (Adeline Soulier-Perkins pers. comm.).
<bibRefCitation id="884B45B2C410DA01D41806783B39FAF1" author="Westwood J. O." box="[299,514,1327,1354]" pageId="25" pageNumber="26" pagination="207 - 210" refId="ref19581" refString="Westwood J. O. 1851. Description of some new species of exotic homopterous insects. Annals and Magazine of Natural History, Series 2, 7: 207 - 210. http: // biodiversitylibrary. org / page / 13787797" type="journal article" year="1851">Westwood (1851)</bibRefCitation>
stated that the specimen he examined was from the collections of the “Mus. Jardin des Plantes, Paris”, the name of the MNHN at that time. No specimen labelled
<taxonomicName id="2BDA43C0C410DA01D028060438AAFA2B" authorityName="Westwood" authorityYear="1851" class="Insecta" family="Fulgoridae" genus="Aphana" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="DEAEE451C410DA01D028060438AAFA2B" italics="true" pageId="25" pageNumber="26">Aphana madagascariensis</emphasis>
</taxonomicName>
was found in the collections of the MNHN, but the specimen chosen here perfectly matches the original description.
<bibRefCitation id="884B45B2C410DA01D76606CE3A10FA0F" author="Lallemand V." box="[597,811,1433,1460]" pageId="25" pageNumber="26" pagination="37 - 124" refId="ref18997" refString="Lallemand V. 1959. Revision des especes africaines de la famille Fulgoridae (super-famille Fulgoroides - sous-ordre des homopteres). Publicacoes Culturais da Companhia de Diamantes de Angola 41: 37 - 124." type="journal article" year="1959">Lallemand (1959)</bibRefCitation>
assumed that the type was lost and stated that, according to the description, the species probably belonged to
<taxonomicName id="2BDA43C0C410DA01D69706EB3AC4FA6D" authorityName="Stal" authorityYear="1863" box="[932,1023,1468,1494]" class="Insecta" family="Fulgoridae" genus="Belbina" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="DEAEE451C410DA01D69706EB3AC4FA6D" box="[932,1023,1468,1494]" italics="true" pageId="25" pageNumber="26">Belbina</emphasis>
</taxonomicName>
or
<taxonomicName id="2BDA43C0C410DA01D11A06EB3DAAFA6D" box="[1065,1169,1468,1494]" class="Insecta" family="Fulgoridae" genus="Cornelia" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="DEAEE451C410DA01D11A06EB3DAAFA6D" box="[1065,1169,1468,1494]" italics="true" pageId="25" pageNumber="26">Cornelia</emphasis>
</taxonomicName>
. As it is not totally certain that the specimen is the one on which the original description was based, I designate it here as a
<typeStatus id="336186E1C410DA01D58E05543821F9A6" box="[189,282,1539,1565]" pageId="25" pageNumber="26" type="neotype">neotype</typeStatus>
rather than recognize it as the
<typeStatus id="336186E1C410DA01D7B005543BD5F9A6" box="[643,750,1539,1565]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="EC653843C410DA01D5D9051338DAF9E5" blockId="25.[189,1399,1604,1701]" box="[234,481,1604,1630]" pageId="25" pageNumber="26">
<emphasis id="DEAEE451C410DA01D5D9051338DAF9E5" bold="true" box="[234,481,1604,1630]" pageId="25" pageNumber="26">Additional material</emphasis>
</paragraph>
<paragraph id="EC653843C410DA01D58E053F3D89F91E" blockId="25.[189,1399,1604,1701]" pageId="25" pageNumber="26">
<collectingCountry id="94CD78D3C410DA01D58E053F38A9F939" box="[189,402,1640,1666]" name="Madagascar" pageId="25" pageNumber="26">MADAGASCAR</collectingCountry>
:
<specimenCount id="FADCF3CAC410DA01D4AC053F38F6F939" box="[415,461,1640,1666]" pageId="25" pageNumber="26" type="female">1 ♀</specimenCount>
, Mad[agascar] (
<collectionCode id="8ACBA086C410DA01D7A3053F3BCBF939" box="[656,752,1640,1666]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="25" pageNumber="26">MNHN</collectionCode>
);
<specimenCount id="FADCF3CAC410DA01D635053F3A0FF939" box="[774,820,1640,1666]" pageId="25" pageNumber="26" type="female">1 ♀</specimenCount>
, Tenina, J. Vadon,
<geoCoordinate id="89EE5E84C410DA01D12905303DB8F939" box="[1050,1155,1639,1666]" degrees="15" direction="south" minutes="45" orientation="latitude" pageId="25" pageNumber="26" precision="925" value="-15.75">15°45 S</geoCoordinate>
,
<geoCoordinate id="89EE5E84C410DA01D1A1053F3DC0F939" box="[1170,1275,1640,1666]" degrees="49" direction="east" minutes="40" orientation="longitude" pageId="25" pageNumber="26" precision="925" value="49.666668">49°40 E</geoCoordinate>
(
<collectionCode id="8ACBA086C410DA01D03E053F3C51F939" box="[1293,1386,1640,1666]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="25" pageNumber="26" type="Museum">RBINS</collectionCode>
);
<specimenCount id="FADCF3CAC410DA01D58E05DC39D0F91E" box="[189,235,1675,1701]" pageId="25" pageNumber="26" type="male">1 ♂</specimenCount>
, NE Madagascar, Fampanambo,
<date id="98641E83C410DA01D74305DC3BE5F91E" box="[624,734,1674,1701]" pageId="25" pageNumber="26" value="1959-07">Jul. 1959</date>
, J. Vadon,
<geoCoordinate id="89EE5E84C410DA01D66E05DD3AFFF91E" box="[861,964,1674,1701]" degrees="15" direction="south" minutes="22" orientation="latitude" pageId="25" pageNumber="26" precision="925" value="-15.366667">15°22 S</geoCoordinate>
,
<geoCoordinate id="89EE5E84C410DA01D6E205DC3D02F91E" box="[977,1081,1675,1701]" degrees="49" direction="east" minutes="38" orientation="longitude" pageId="25" pageNumber="26" precision="925" value="49.633335">49°38 E</geoCoordinate>
(
<collectionCode id="8ACBA086C410DA01D17B05DC3D9EF91E" box="[1096,1189,1675,1701]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="25" pageNumber="26" type="Museum">MRAC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="A4C06BC8C410DA02D58E059B3A4DF9BD" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="description">
<paragraph id="EC653843C410DA01D58E059B3850F95D" blockId="25.[189,363,1740,1766]" box="[189,363,1740,1766]" pageId="25" pageNumber="26">
<heading id="B72D8F2FC410DA01D58E059B3850F95D" bold="true" box="[189,363,1740,1766]" fontSize="11" level="3" pageId="25" pageNumber="26" reason="3">
<emphasis id="DEAEE451C410DA01D58E059B3850F95D" bold="true" box="[189,363,1740,1766]" pageId="25" pageNumber="26">Redescription</emphasis>
</heading>
</paragraph>
<paragraph id="EC653843C410DA01D58E05AC3B85F8AE" blockId="25.[189,702,1786,1813]" box="[189,702,1786,1813]" pageId="25" pageNumber="26">
LT:
<specimenCount id="FADCF3CAC410DA01D5DF05AC3839F8AE" box="[236,258,1787,1813]" pageId="25" pageNumber="26" type="male"></specimenCount>
(n = 1)
<quantity id="2B2295A6C410DA01D45205AD38F6F8AE" box="[353,461,1786,1813]" metricMagnitude="-2" metricUnit="m" metricValue="2.63" pageId="25" pageNumber="26" unit="mm" value="26.3">26.3 mm</quantity>
;
<specimenCount id="FADCF3CAC410DA01D4EA05AC38D4F8AE" box="[473,495,1787,1813]" pageId="25" pageNumber="26" type="female"></specimenCount>
(n = 1)
<quantity id="2B2295A6C410DA01D77E05AC3B82F8AE" box="[589,697,1787,1813]" metricMagnitude="-2" metricUnit="m" metricValue="2.83" pageId="25" pageNumber="26" unit="mm" value="28.3">28.3 mm</quantity>
.
</paragraph>
<paragraph id="EC653843C410DA02D58E046A3DECFE34" blockId="25.[189,1399,1852,2019]" lastBlockId="26.[189,1399,267,399]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
HEAD. Yellow-brown with 2 darker patches on disc of vertex (
<figureCitation id="74E124C6C410DA01D6AB046B3D2DF8ED" box="[920,1046,1852,1878]" captionStart="Figs 79. — 7" captionStartId="8.[189,241,1869,1895]" captionTargetBox="[189,1398,277,1823]" captionTargetId="figure@8.[189,1398,271,1829]" captionTargetPageId="8" captionText="Figs 79. — 7. Belbina lambertoni. 8. B. madagascariensis. 9. B. nympha. — A. Habitus, dorsal view. B. Habitus, lateral view. C. Vertex, pro- and mesonotum, dorsal view. D. Head and prothorax, lateral view. E. Frons, normal view. — Scale bar: 10 mm (CE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3838825" httpUri="https://zenodo.org/record/3838825/files/figure.png" pageId="25" pageNumber="26">Fig. 8CE</figureCitation>
); yellow-orange patch around insertion of antennae (
<figureCitation id="74E124C6C410DA01D4F204083B24F8C2" box="[449,543,1887,1913]" captionStart="Figs 79. — 7" captionStartId="8.[189,241,1869,1895]" captionTargetBox="[189,1398,277,1823]" captionTargetId="figure@8.[189,1398,271,1829]" captionTargetPageId="8" captionText="Figs 79. — 7. Belbina lambertoni. 8. B. madagascariensis. 9. B. nympha. — A. Habitus, dorsal view. B. Habitus, lateral view. C. Vertex, pro- and mesonotum, dorsal view. D. Head and prothorax, lateral view. E. Frons, normal view. — Scale bar: 10 mm (CE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3838825" httpUri="https://zenodo.org/record/3838825/files/figure.png" pageId="25" pageNumber="26">Fig. 8D</figureCitation>
); vertex with hind margin slightly carinate and obsolete median carina on disc extending posterad beyond hind margin; sides of vertex strongly carinate, laminate above eye and extending anteriorly to apex of cephalic process; sides of vertex slightly bisinuate in lateral view (
<figureCitation id="74E124C6C410DA01D07404F1383CF858" captionStart="Figs 79. — 7" captionStartId="8.[189,241,1869,1895]" captionTargetBox="[189,1398,277,1823]" captionTargetId="figure@8.[189,1398,271,1829]" captionTargetPageId="8" captionText="Figs 79. — 7. Belbina lambertoni. 8. B. madagascariensis. 9. B. nympha. — A. Habitus, dorsal view. B. Habitus, lateral view. C. Vertex, pro- and mesonotum, dorsal view. D. Head and prothorax, lateral view. E. Frons, normal view. — Scale bar: 10 mm (CE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3838825" httpUri="https://zenodo.org/record/3838825/files/figure.png" pageId="25" pageNumber="26">Fig. 8CD</figureCitation>
); frons with 2 carinae extending ventrally on sides of process (
<figureCitation id="74E124C6C410DA01D6C5049E3D68F858" box="[1014,1107,1993,2019]" captionStart="Figs 79. — 7" captionStartId="8.[189,241,1869,1895]" captionTargetBox="[189,1398,277,1823]" captionTargetId="figure@8.[189,1398,271,1829]" captionTargetPageId="8" captionText="Figs 79. — 7. Belbina lambertoni. 8. B. madagascariensis. 9. B. nympha. — A. Habitus, dorsal view. B. Habitus, lateral view. C. Vertex, pro- and mesonotum, dorsal view. D. Head and prothorax, lateral view. E. Frons, normal view. — Scale bar: 10 mm (CE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3838825" httpUri="https://zenodo.org/record/3838825/files/figure.png" pageId="25" pageNumber="26">Fig. 8E</figureCitation>
); cephalic process about 3.2 times as long as diameter of eye, projecting anterodorsad, curved, elongate and narrow; apical half ventrally with median carina getting laminate near apex (
<figureCitation id="74E124C6C413DA02D64302793AC9FEF2" box="[880,1010,302,329]" captionStart="Figs 79. — 7" captionStartId="8.[189,241,1869,1895]" captionTargetBox="[189,1398,277,1823]" captionTargetId="figure@8.[189,1398,271,1829]" captionTargetPageId="8" captionText="Figs 79. — 7. Belbina lambertoni. 8. B. madagascariensis. 9. B. nympha. — A. Habitus, dorsal view. B. Habitus, lateral view. C. Vertex, pro- and mesonotum, dorsal view. D. Head and prothorax, lateral view. E. Frons, normal view. — Scale bar: 10 mm (CE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3838825" httpUri="https://zenodo.org/record/3838825/files/figure.png" pageId="26" pageNumber="27">Fig. 8DE</figureCitation>
); transverse wrinkles dorsally at base of process (
<figureCitation id="74E124C6C413DA02D44E020538E3FED7" box="[381,472,338,364]" captionStart="Figs 79. — 7" captionStartId="8.[189,241,1869,1895]" captionTargetBox="[189,1398,277,1823]" captionTargetId="figure@8.[189,1398,271,1829]" captionTargetPageId="8" captionText="Figs 79. — 7. Belbina lambertoni. 8. B. madagascariensis. 9. B. nympha. — A. Habitus, dorsal view. B. Habitus, lateral view. C. Vertex, pro- and mesonotum, dorsal view. D. Head and prothorax, lateral view. E. Frons, normal view. — Scale bar: 10 mm (CE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3838825" httpUri="https://zenodo.org/record/3838825/files/figure.png" pageId="26" pageNumber="27">Fig. 8C</figureCitation>
); postclypeus with sides strongly carinate and slight median carina; anteclypeus with median carina (
<figureCitation id="74E124C6C413DA02D49D02223B32FE34" box="[430,521,373,399]" captionStart="Figs 79. — 7" captionStartId="8.[189,241,1869,1895]" captionTargetBox="[189,1398,277,1823]" captionTargetId="figure@8.[189,1398,271,1829]" captionTargetPageId="8" captionText="Figs 79. — 7. Belbina lambertoni. 8. B. madagascariensis. 9. B. nympha. — A. Habitus, dorsal view. B. Habitus, lateral view. C. Vertex, pro- and mesonotum, dorsal view. D. Head and prothorax, lateral view. E. Frons, normal view. — Scale bar: 10 mm (CE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3838825" httpUri="https://zenodo.org/record/3838825/files/figure.png" pageId="26" pageNumber="27">Fig. 8E</figureCitation>
); labium very long, reaching or surpassing apex of abdomen.
</paragraph>
<paragraph id="EC653843C413DA02D58E02E03DACFD3B" blockId="26.[189,1399,438,641]" pageId="26" pageNumber="27">
THORAX. Yellow-brown with slightly paler markings (
<figureCitation id="74E124C6C413DA02D60702E13A88FE6B" box="[820,947,438,464]" captionStart="Figs 79. — 7" captionStartId="8.[189,241,1869,1895]" captionTargetBox="[189,1398,277,1823]" captionTargetId="figure@8.[189,1398,271,1829]" captionTargetPageId="8" captionText="Figs 79. — 7. Belbina lambertoni. 8. B. madagascariensis. 9. B. nympha. — A. Habitus, dorsal view. B. Habitus, lateral view. C. Vertex, pro- and mesonotum, dorsal view. D. Head and prothorax, lateral view. E. Frons, normal view. — Scale bar: 10 mm (CE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3838825" httpUri="https://zenodo.org/record/3838825/files/figure.png" pageId="26" pageNumber="27">Fig. 8CE</figureCitation>
); pronotum with strong median carina (
<figureCitation id="74E124C6C413DA02D5F5028E381FFE4F" box="[198,292,473,500]" captionStart="Figs 79. — 7" captionStartId="8.[189,241,1869,1895]" captionTargetBox="[189,1398,277,1823]" captionTargetId="figure@8.[189,1398,271,1829]" captionTargetPageId="8" captionText="Figs 79. — 7. Belbina lambertoni. 8. B. madagascariensis. 9. B. nympha. — A. Habitus, dorsal view. B. Habitus, lateral view. C. Vertex, pro- and mesonotum, dorsal view. D. Head and prothorax, lateral view. E. Frons, normal view. — Scale bar: 10 mm (CE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3838825" httpUri="https://zenodo.org/record/3838825/files/figure.png" pageId="26" pageNumber="27">Fig. 8C</figureCitation>
); carina very slightly sinuate in lateral view (
<figureCitation id="74E124C6C413DA02D67B028E3A9EFE4F" box="[840,933,473,500]" captionStart="Figs 79. — 7" captionStartId="8.[189,241,1869,1895]" captionTargetBox="[189,1398,277,1823]" captionTargetId="figure@8.[189,1398,271,1829]" captionTargetPageId="8" captionText="Figs 79. — 7. Belbina lambertoni. 8. B. madagascariensis. 9. B. nympha. — A. Habitus, dorsal view. B. Habitus, lateral view. C. Vertex, pro- and mesonotum, dorsal view. D. Head and prothorax, lateral view. E. Frons, normal view. — Scale bar: 10 mm (CE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3838825" httpUri="https://zenodo.org/record/3838825/files/figure.png" pageId="26" pageNumber="27">Fig. 8D</figureCitation>
) and with strongly impressed point on each side (
<figureCitation id="74E124C6C413DA02D47302AA389AFDAC" box="[320,417,509,535]" captionStart="Figs 79. — 7" captionStartId="8.[189,241,1869,1895]" captionTargetBox="[189,1398,277,1823]" captionTargetId="figure@8.[189,1398,271,1829]" captionTargetPageId="8" captionText="Figs 79. — 7. Belbina lambertoni. 8. B. madagascariensis. 9. B. nympha. — A. Habitus, dorsal view. B. Habitus, lateral view. C. Vertex, pro- and mesonotum, dorsal view. D. Head and prothorax, lateral view. E. Frons, normal view. — Scale bar: 10 mm (CE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3838825" httpUri="https://zenodo.org/record/3838825/files/figure.png" pageId="26" pageNumber="27">Fig. 8C</figureCitation>
); hind margin elevated above level of mesonotum medially (
<figureCitation id="74E124C6C413DA02D1BA02AA3DD0FDAC" box="[1161,1259,509,535]" captionStart="Figs 79. — 7" captionStartId="8.[189,241,1869,1895]" captionTargetBox="[189,1398,277,1823]" captionTargetId="figure@8.[189,1398,271,1829]" captionTargetPageId="8" captionText="Figs 79. — 7. Belbina lambertoni. 8. B. madagascariensis. 9. B. nympha. — A. Habitus, dorsal view. B. Habitus, lateral view. C. Vertex, pro- and mesonotum, dorsal view. D. Head and prothorax, lateral view. E. Frons, normal view. — Scale bar: 10 mm (CE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3838825" httpUri="https://zenodo.org/record/3838825/files/figure.png" pageId="26" pageNumber="27">Fig. 8D</figureCitation>
); peridiscal carina anteriorly (
<figureCitation id="74E124C6C413DA02D4A0017738C9FD81" box="[403,498,544,570]" captionStart="Figs 79. — 7" captionStartId="8.[189,241,1869,1895]" captionTargetBox="[189,1398,277,1823]" captionTargetId="figure@8.[189,1398,271,1829]" captionTargetPageId="8" captionText="Figs 79. — 7. Belbina lambertoni. 8. B. madagascariensis. 9. B. nympha. — A. Habitus, dorsal view. B. Habitus, lateral view. C. Vertex, pro- and mesonotum, dorsal view. D. Head and prothorax, lateral view. E. Frons, normal view. — Scale bar: 10 mm (CE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3838825" httpUri="https://zenodo.org/record/3838825/files/figure.png" pageId="26" pageNumber="27">Fig. 8C</figureCitation>
); sides of prothorax with oblique carina; lateral lobe of pronotum rounded posteriorly (
<figureCitation id="74E124C6C413DA02D47F011438F0FDE6" box="[332,459,579,605]" captionStart="Figs 79. — 7" captionStartId="8.[189,241,1869,1895]" captionTargetBox="[189,1398,277,1823]" captionTargetId="figure@8.[189,1398,271,1829]" captionTargetPageId="8" captionText="Figs 79. — 7. Belbina lambertoni. 8. B. madagascariensis. 9. B. nympha. — A. Habitus, dorsal view. B. Habitus, lateral view. C. Vertex, pro- and mesonotum, dorsal view. D. Head and prothorax, lateral view. E. Frons, normal view. — Scale bar: 10 mm (CE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3838825" httpUri="https://zenodo.org/record/3838825/files/figure.png" pageId="26" pageNumber="27">Fig. 8DE</figureCitation>
); mesonotum with median carina stopped at scutellum; curved peridiscal carina; short oblique carina at base of scutellum; scutellum transversely wrinkled (
<figureCitation id="74E124C6C413DA02D11E01313DB1FD3B" box="[1069,1162,614,640]" captionStart="Figs 79. — 7" captionStartId="8.[189,241,1869,1895]" captionTargetBox="[189,1398,277,1823]" captionTargetId="figure@8.[189,1398,271,1829]" captionTargetPageId="8" captionText="Figs 79. — 7. Belbina lambertoni. 8. B. madagascariensis. 9. B. nympha. — A. Habitus, dorsal view. B. Habitus, lateral view. C. Vertex, pro- and mesonotum, dorsal view. D. Head and prothorax, lateral view. E. Frons, normal view. — Scale bar: 10 mm (CE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3838825" httpUri="https://zenodo.org/record/3838825/files/figure.png" pageId="26" pageNumber="27">Fig. 8C</figureCitation>
).
</paragraph>
<paragraph id="EC653843C413DA02D58E01FF3DE7FCF4" blockId="26.[189,1399,680,847]" pageId="26" pageNumber="27">
TEGMINA (
<figureCitation id="74E124C6C413DA02D40701FF38A8FD79" box="[308,403,680,706]" captionStart="Figs 79. — 7" captionStartId="8.[189,241,1869,1895]" captionTargetBox="[189,1398,277,1823]" captionTargetId="figure@8.[189,1398,271,1829]" captionTargetPageId="8" captionText="Figs 79. — 7. Belbina lambertoni. 8. B. madagascariensis. 9. B. nympha. — A. Habitus, dorsal view. B. Habitus, lateral view. C. Vertex, pro- and mesonotum, dorsal view. D. Head and prothorax, lateral view. E. Frons, normal view. — Scale bar: 10 mm (CE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3838825" httpUri="https://zenodo.org/record/3838825/files/figure.png" pageId="26" pageNumber="27">Fig. 8A</figureCitation>
). Brown with irregular black-brown, often confluent, small spots with pale yellowbrown center; center of spots covered with white waxy secretion in fresh specimens; no spots beyond nodal line of cross-veins except at apicosutural angle; apex of clavus with large black-brown marking followed by white patch along sutural margin; costal margin straight, slightly rounded after nodal line; apical margin oblique, straight in middle and with angles rounded; sutural margin sinuate.
</paragraph>
<paragraph id="EC653843C413DA02D58E00203A9FFC08" blockId="26.[189,1398,886,948]" pageId="26" pageNumber="27">
HIND WINGS (
<figureCitation id="74E124C6C413DA02D4660021388FFC2B" box="[341,436,886,912]" captionStart="Figs 79. — 7" captionStartId="8.[189,241,1869,1895]" captionTargetBox="[189,1398,277,1823]" captionTargetId="figure@8.[189,1398,271,1829]" captionTargetPageId="8" captionText="Figs 79. — 7. Belbina lambertoni. 8. B. madagascariensis. 9. B. nympha. — A. Habitus, dorsal view. B. Habitus, lateral view. C. Vertex, pro- and mesonotum, dorsal view. D. Head and prothorax, lateral view. E. Frons, normal view. — Scale bar: 10 mm (CE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3838825" httpUri="https://zenodo.org/record/3838825/files/figure.png" pageId="26" pageNumber="27">Fig. 8A</figureCitation>
). Broad; disc largely bright red or orange; apex and sutural margin brown-black; 24 small, brown-black spots on disc; sutural margin bisinuate.
</paragraph>
<paragraph id="EC653843C413DA02D58E008C3C06FBE4" blockId="26.[189,1398,987,1119]" pageId="26" pageNumber="27">
LEGS (
<figureCitation id="74E124C6C413DA02D439008C3852FC4E" box="[266,361,987,1013]" captionStart="Figs 79. — 7" captionStartId="8.[189,241,1869,1895]" captionTargetBox="[189,1398,277,1823]" captionTargetId="figure@8.[189,1398,271,1829]" captionTargetPageId="8" captionText="Figs 79. — 7. Belbina lambertoni. 8. B. madagascariensis. 9. B. nympha. — A. Habitus, dorsal view. B. Habitus, lateral view. C. Vertex, pro- and mesonotum, dorsal view. D. Head and prothorax, lateral view. E. Frons, normal view. — Scale bar: 10 mm (CE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3838825" httpUri="https://zenodo.org/record/3838825/files/figure.png" pageId="26" pageNumber="27">Fig. 8A</figureCitation>
). Elongate and slender; pro- and mesofemora brown with 3 narrow, sinuate and often incomplete pale yellow rings; pro- and mesotibiae brown with 2 pale yellow rings; metafemora brown with imprecise yellow ring; metatibiae and metatarsi yellow-brown, tibiae darker basally; metatibiae with 5 (sometimes 4) lateral and 7 apical spines; 89 spines apicoventrally on first metatarsomere.
</paragraph>
<paragraph id="EC653843C413DA02D58E07D13A61FB1B" blockId="26.[189,858,1158,1184]" box="[189,858,1158,1184]" pageId="26" pageNumber="27">
ABDOMEN (
<figureCitation id="74E124C6C413DA02D40F07D138A2FB1B" box="[316,409,1158,1184]" captionStart="Figs 79. — 7" captionStartId="8.[189,241,1869,1895]" captionTargetBox="[189,1398,277,1823]" captionTargetId="figure@8.[189,1398,271,1829]" captionTargetPageId="8" captionText="Figs 79. — 7. Belbina lambertoni. 8. B. madagascariensis. 9. B. nympha. — A. Habitus, dorsal view. B. Habitus, lateral view. C. Vertex, pro- and mesonotum, dorsal view. D. Head and prothorax, lateral view. E. Frons, normal view. — Scale bar: 10 mm (CE not to scale)." figureDoi="http://doi.org/10.5281/zenodo.3838825" httpUri="https://zenodo.org/record/3838825/files/figure.png" pageId="26" pageNumber="27">Fig. 8A</figureCitation>
). Red dorsally, brown-black ventrally.
</paragraph>
<paragraph id="EC653843C413DA02D58E0790384BFB5A" blockId="26.[189,368,1223,1249]" box="[189,368,1223,1249]" pageId="26" pageNumber="27">
<heading id="B72D8F2FC413DA02D58E0790384BFB5A" bold="true" box="[189,368,1223,1249]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="DEAEE451C413DA02D58E0790384BFB5A" bold="true" box="[189,368,1223,1249]" pageId="26" pageNumber="27">Male genitalia</emphasis>
</heading>
</paragraph>
<paragraph id="EC653843C413DA02D58E07A13A4DF9BD" blockId="26.[189,1399,1270,1543]" pageId="26" pageNumber="27">
Very finely granulose, dark brown, paler along sides of anal tube (
<figureCitation id="74E124C6C413DA02D6D607A13D4AFAAA" box="[997,1137,1270,1297]" captionStart="Figs 2429" captionStartId="12.[189,241,1928,1954]" captionTargetBox="[189,1398,498,1887]" captionTargetId="figure@12.[189,1398,498,1887]" captionTargetPageId="12" captionText="Figs 2429. Male genitalia. — 2425. Belbina lambertoni. 24. Lateral view. 25. Dorsal view. — 26 27. B. madagascariensis. 26. Lateral view. 27. Dorsal view. — 2829. B. nympha. 28. Lateral view. 29. Dorsal view. — An = anal tube, Py = pygofer, G = gonostylus. Photographs by Y. Laurent &amp; I. Bachy." figureDoi="http://doi.org/10.5281/zenodo.3838833" httpUri="https://zenodo.org/record/3838833/files/figure.png" pageId="26" pageNumber="27">Figs 2627</figureCitation>
); pygofer higher than long and with posterior margin broadly rounded in lateral view (
<figureCitation id="74E124C6C413DA02D6F0064E3D26FA8F" box="[963,1053,1305,1332]" captionStart="Figs 2429" captionStartId="12.[189,241,1928,1954]" captionTargetBox="[189,1398,498,1887]" captionTargetId="figure@12.[189,1398,498,1887]" captionTargetPageId="12" captionText="Figs 2429. Male genitalia. — 2425. Belbina lambertoni. 24. Lateral view. 25. Dorsal view. — 26 27. B. madagascariensis. 26. Lateral view. 27. Dorsal view. — 2829. B. nympha. 28. Lateral view. 29. Dorsal view. — An = anal tube, Py = pygofer, G = gonostylus. Photographs by Y. Laurent &amp; I. Bachy." figureDoi="http://doi.org/10.5281/zenodo.3838833" httpUri="https://zenodo.org/record/3838833/files/figure.png" pageId="26" pageNumber="27">Fig. 26</figureCitation>
); anal tube slightly elongate, 1.31 times longer than broad at apex and with lateral margins sinuate in dorsal view (
<figureCitation id="74E124C6C413DA02D192066B3DC2FAED" box="[1185,1273,1340,1366]" captionStart="Figs 2429" captionStartId="12.[189,241,1928,1954]" captionTargetBox="[189,1398,498,1887]" captionTargetId="figure@12.[189,1398,498,1887]" captionTargetPageId="12" captionText="Figs 2429. Male genitalia. — 2425. Belbina lambertoni. 24. Lateral view. 25. Dorsal view. — 26 27. B. madagascariensis. 26. Lateral view. 27. Dorsal view. — 2829. B. nympha. 28. Lateral view. 29. Dorsal view. — An = anal tube, Py = pygofer, G = gonostylus. Photographs by Y. Laurent &amp; I. Bachy." figureDoi="http://doi.org/10.5281/zenodo.3838833" httpUri="https://zenodo.org/record/3838833/files/figure.png" pageId="26" pageNumber="27">Fig. 27</figureCitation>
); posterior margin obliquely rounded and underside nearly straight in lateral view (
<figureCitation id="74E124C6C413DA02D13206083D63FAC1" box="[1025,1112,1375,1402]" captionStart="Figs 2429" captionStartId="12.[189,241,1928,1954]" captionTargetBox="[189,1398,498,1887]" captionTargetId="figure@12.[189,1398,498,1887]" captionTargetPageId="12" captionText="Figs 2429. Male genitalia. — 2425. Belbina lambertoni. 24. Lateral view. 25. Dorsal view. — 26 27. B. madagascariensis. 26. Lateral view. 27. Dorsal view. — 2829. B. nympha. 28. Lateral view. 29. Dorsal view. — An = anal tube, Py = pygofer, G = gonostylus. Photographs by Y. Laurent &amp; I. Bachy." figureDoi="http://doi.org/10.5281/zenodo.3838833" httpUri="https://zenodo.org/record/3838833/files/figure.png" pageId="26" pageNumber="27">Fig. 26</figureCitation>
); gonostyli elongate, 1.6 times longer than high, not surpassing anal tube and broadly rounded at apex in lateral view (
<figureCitation id="74E124C6C413DA02D02306D53C53FA26" box="[1296,1384,1410,1437]" captionStart="Figs 2429" captionStartId="12.[189,241,1928,1954]" captionTargetBox="[189,1398,498,1887]" captionTargetId="figure@12.[189,1398,498,1887]" captionTargetPageId="12" captionText="Figs 2429. Male genitalia. — 2425. Belbina lambertoni. 24. Lateral view. 25. Dorsal view. — 26 27. B. madagascariensis. 26. Lateral view. 27. Dorsal view. — 2829. B. nympha. 28. Lateral view. 29. Dorsal view. — An = anal tube, Py = pygofer, G = gonostylus. Photographs by Y. Laurent &amp; I. Bachy." figureDoi="http://doi.org/10.5281/zenodo.3838833" httpUri="https://zenodo.org/record/3838833/files/figure.png" pageId="26" pageNumber="27">Fig. 26</figureCitation>
); ventral margin slightly sinuate on basal 2/3 (
<figureCitation id="74E124C6C413DA02D78806F13A28FA7A" box="[699,787,1446,1473]" captionStart="Figs 2429" captionStartId="12.[189,241,1928,1954]" captionTargetBox="[189,1398,498,1887]" captionTargetId="figure@12.[189,1398,498,1887]" captionTargetPageId="12" captionText="Figs 2429. Male genitalia. — 2425. Belbina lambertoni. 24. Lateral view. 25. Dorsal view. — 26 27. B. madagascariensis. 26. Lateral view. 27. Dorsal view. — 2829. B. nympha. 28. Lateral view. 29. Dorsal view. — An = anal tube, Py = pygofer, G = gonostylus. Photographs by Y. Laurent &amp; I. Bachy." figureDoi="http://doi.org/10.5281/zenodo.3838833" httpUri="https://zenodo.org/record/3838833/files/figure.png" pageId="26" pageNumber="27">Fig. 26</figureCitation>
); dorsal margin obliquely directed dorsally on basal half, then straight after angle, pointed hook-like tooth at half of basal oblique part curved lateroventrally (
<figureCitation id="74E124C6C413DA02D5F506BB3825F9BC" box="[198,286,1516,1543]" captionStart="Figs 2429" captionStartId="12.[189,241,1928,1954]" captionTargetBox="[189,1398,498,1887]" captionTargetId="figure@12.[189,1398,498,1887]" captionTargetPageId="12" captionText="Figs 2429. Male genitalia. — 2425. Belbina lambertoni. 24. Lateral view. 25. Dorsal view. — 26 27. B. madagascariensis. 26. Lateral view. 27. Dorsal view. — 2829. B. nympha. 28. Lateral view. 29. Dorsal view. — An = anal tube, Py = pygofer, G = gonostylus. Photographs by Y. Laurent &amp; I. Bachy." figureDoi="http://doi.org/10.5281/zenodo.3838833" httpUri="https://zenodo.org/record/3838833/files/figure.png" pageId="26" pageNumber="27">Fig. 26</figureCitation>
); gonostyli nearly not visible from above (
<figureCitation id="74E124C6C413DA02D62206BB3A52F9BD" box="[785,873,1516,1542]" captionStart="Figs 2429" captionStartId="12.[189,241,1928,1954]" captionTargetBox="[189,1398,498,1887]" captionTargetId="figure@12.[189,1398,498,1887]" captionTargetPageId="12" captionText="Figs 2429. Male genitalia. — 2425. Belbina lambertoni. 24. Lateral view. 25. Dorsal view. — 26 27. B. madagascariensis. 26. Lateral view. 27. Dorsal view. — 2829. B. nympha. 28. Lateral view. 29. Dorsal view. — An = anal tube, Py = pygofer, G = gonostylus. Photographs by Y. Laurent &amp; I. Bachy." figureDoi="http://doi.org/10.5281/zenodo.3838833" httpUri="https://zenodo.org/record/3838833/files/figure.png" pageId="26" pageNumber="27">Fig. 27</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A4C06BC8C413DA02D58E05793A76F8DC" pageId="26" pageNumber="27" type="discussion">
<paragraph id="EC653843C413DA02D58E05793814F9F3" blockId="26.[189,303,1582,1608]" box="[189,303,1582,1608]" pageId="26" pageNumber="27">
<heading id="B72D8F2FC413DA02D58E05793814F9F3" bold="true" box="[189,303,1582,1608]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="DEAEE451C413DA02D58E05793814F9F3" bold="true" box="[189,303,1582,1608]" pageId="26" pageNumber="27">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="EC653843C413DA02D58E050B3A77F9CD" blockId="26.[189,844,1628,1655]" box="[189,844,1628,1655]" pageId="26" pageNumber="27">Male genitalia without basodorsal process on gonostyli.</paragraph>
<paragraph id="EC653843C413DA02D58E05CA3A76F8DC" blockId="26.[189,1399,1693,1896]" pageId="26" pageNumber="27">
<taxonomicName id="2BDA43C0C413DA02D58E05CA38D1F90C" box="[189,490,1693,1719]" class="Insecta" family="Fulgoridae" genus="Belbina" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="madagascarensis">
<emphasis id="DEAEE451C413DA02D58E05CA38D1F90C" box="[189,490,1693,1719]" italics="true" pageId="26" pageNumber="27">Belbina madagascarensis</emphasis>
</taxonomicName>
can be separated (1) from
<taxonomicName id="2BDA43C0C413DA02D61105C93A94F90C" authorityName="Constant" authorityYear="2014" baseAuthorityName="Schmidt" baseAuthorityYear="1911" box="[802,943,1693,1719]" class="Insecta" family="Fulgoridae" genus="Belbina" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="bergrothi">
<emphasis id="DEAEE451C413DA02D61105C93A94F90C" box="[802,943,1693,1719]" italics="true" pageId="26" pageNumber="27">B. bergrothi</emphasis>
</taxonomicName>
by the less elongate gonostyli, without large pale marking ventrally and more broadly rounded apex, and less elongate anal tube; (2) from
<taxonomicName id="2BDA43C0C413DA02D58E05B23804F945" authorityName="Lallemand" authorityYear="1950" box="[189,319,1764,1790]" class="Insecta" family="Fulgoridae" genus="Belbina" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="foliacea">
<emphasis id="DEAEE451C413DA02D58E05B23804F945" box="[189,319,1764,1790]" italics="true" pageId="26" pageNumber="27">B. foliacea</emphasis>
</taxonomicName>
by having gonostyli more broadly rounded apically and without a strong angle above the mediodorsal tooth; (3) from
<taxonomicName id="2BDA43C0C413DA02D73A045F3BBDF89A" authorityName="Constant" authorityYear="2014" baseAuthorityName="Stal" baseAuthorityYear="1866" box="[521,646,1799,1825]" class="Insecta" family="Fulgoridae" genus="Belbina" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="nympha">
<emphasis id="DEAEE451C413DA02D73A045F3BBDF89A" box="[521,646,1799,1825]" italics="true" pageId="26" pageNumber="27">B. nympha</emphasis>
</taxonomicName>
by having the margins of the anal tube not rounded laterally and the dorsal margin of the gonostyli nearly straight on the apical half; (4) from
<taxonomicName id="2BDA43C0C413DA02D162047C3DF6F8FF" authorityName="Lallem., V. Lallemand" authorityYear="1950" box="[1105,1229,1835,1860]" class="Insecta" family="Fulgoridae" genus="Belbina" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="recurva">
<emphasis id="DEAEE451C413DA02D162047C3DF6F8FF" box="[1105,1229,1835,1860]" italics="true" pageId="26" pageNumber="27">B. recurva</emphasis>
</taxonomicName>
by having the anal tube not produced into a semi-circular lateral plate.
</paragraph>
</subSubSection>
<subSubSection id="A4C06BC8C413DA02D58E04D83823F86C" box="[189,342,1935,2007]" pageId="26" pageNumber="27" type="distribution">
<paragraph id="EC653843C413DA02D58E04D8386DF812" blockId="26.[189,342,1935,1961]" box="[189,342,1935,1961]" pageId="26" pageNumber="27">
<heading id="B72D8F2FC413DA02D58E04D8386DF812" bold="true" box="[189,342,1935,1961]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="DEAEE451C413DA02D58E04D8386DF812" bold="true" box="[189,342,1935,1961]" pageId="26" pageNumber="27">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="EC653843C413DA02D58E04EA3823F86C" blockId="26.[189,280,1981,2007]" box="[189,280,1981,2007]" pageId="26" pageNumber="27">
<figureCitation id="74E124C6C413DA02D58E04EA382FF86C" box="[189,276,1981,2007]" captionStart="Figs 4649" captionStartId="32.[189,241,1963,1989]" captionTargetBox="[297,1284,265,1915]" captionTargetId="figure@32.[296,1292,265,1921]" captionTargetPageId="32" captionText="Figs 4649. Distribution maps. — 46. Belbina bergrothi, B. bloetei and B. madagascariensis. 47. B. bourgoini sp. nov., B. falleni and B. foliacea. 48. B. laetitiae sp. nov. 49. B. lambertoni." figureDoi="http://doi.org/10.5281/zenodo.3838841" httpUri="https://zenodo.org/record/3838841/files/figure.png" pageId="26" pageNumber="27">Fig. 46</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>