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<document id="33C1AA33B6941ABB6A723DC84B6D83A3" ID="10.11646/zootaxa.4281.1.13SLASH11447" ID-CLB-Dataset="28730" ID-DOI="10.11646/zootaxa.4281.1.13" ID-GBIF-Dataset="660f1f7d-2a80-4f3b-aa03-861c81fffb6c" ID-ISSN="1175-5326" ID-Zenodo-Dep="815997" ID-ZooBank="C04CCD37-B5C7-4C7A-A831-D585FFFB651E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1498121191771" checkinUser="plazi" docAuthor="Quintas, Victor, Felix, Márcio, Lima, Douglas Felipe Dos Santos &amp; Mejdalani, Gabriel" docDate="2017" docId="2E6B3A059C3B562ACDA0FD44FC8CFEA8" docLanguage="en" docName="zootaxa.4281.1.13SLASH11447.pdf" docOrigin="Zootaxa 4281 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dilobopterus nelsoni Quintas &amp; Felix &amp; Lima &amp; Mejdalani 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="130" masterDocId="D252427D9C3A5629CD37FFEBFFEDFFCC" masterDocTitle="Two new species of Dilobopterus Signoret, 1850 and the male of D. segmentalis (Signoret, 1853) from Southeastern Brazil, with taxonomic notes on the genus (Hemiptera: Cicadellidae: Cicadellini)" masterLastPageNumber="134" masterPageNumber="127" pageNumber="128" updateTime="1698448211075" updateUser="ExternalLinkService">
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<mods:title id="AE27B3D350FECE5C974073CFF2CDB7C8">Two new species of Dilobopterus Signoret, 1850 and the male of D. segmentalis (Signoret, 1853) from Southeastern Brazil, with taxonomic notes on the genus (Hemiptera: Cicadellidae: Cicadellini)</mods:title>
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<mods:namePart id="8F6EF88787FE797CD7C8C8266A6E4C0D">Quintas, Victor</mods:namePart>
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<mods:namePart id="CD54D9F7876BE5640E76038824817272">Felix, Márcio</mods:namePart>
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<mods:namePart id="91B84558371542F24EE75313F54BA302">Lima, Douglas Felipe Dos Santos</mods:namePart>
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<mods:namePart id="AEFFE07E4CE536F2D24642851A13018B">Mejdalani, Gabriel</mods:namePart>
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<paragraph id="A67D8B139C3B5628CDA0FD44FE1AFD06" blockId="1.[151,503,687,747]" box="[151,503,687,714]" pageId="1" pageNumber="128">
<heading id="FD353C7F9C3B5628CDA0FD44FE1AFD06" bold="true" box="[151,503,687,714]" fontSize="11" level="1" pageId="1" pageNumber="128" reason="1">
<emphasis id="94B657019C3B5628CDA0FD44FE1AFD06" bold="true" box="[151,503,687,714]" pageId="1" pageNumber="128">
<taxonomicName id="61C2F0909C3B5628CDA0FD44FE7FFD05" ID-CoL="365PR" authority="Quintas &amp; Felix &amp; Lima &amp; Mejdalani, 2017" authorityName="Quintas &amp; Felix &amp; Lima &amp; Mejdalani" authorityYear="2017" box="[151,402,687,713]" class="Insecta" family="Cicadellidae" genus="Dilobopterus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="128" phylum="Arthropoda" rank="species" species="nelsoni" status="sp. nov.">
<emphasis id="94B657019C3B5628CDA0FD44FE7FFD05" bold="true" box="[151,402,687,713]" italics="true" pageId="1" pageNumber="128">Dilobopterus nelsoni</emphasis>
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<taxonomicNameLabel id="8F85EA7A9C3B5628CCADFD5BFE1AFD06" box="[410,503,688,714]" pageId="1" pageNumber="128" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="A67D8B139C3B5628CDA0FD39FEB5FD27" blockId="1.[151,503,687,747]" box="[151,344,722,747]" pageId="1" pageNumber="128">
(
<figureCitation id="3EF997969C3B5628CDA9FD39FECAFD27" box="[158,295,722,747]" captionStart="FIGURES 1 8" captionStartId="2.[151,263,1323,1346]" captionTargetBox="[175,1395,312,1273]" captionTargetId="figure@2.[150,1436,301,1302]" captionTargetPageId="2" captionText="FIGURES 1 8. Dilobopterus nelsoni sp. nov. 1, head, pronotum, and mesonotum, dorsal view (male paratype). 2, forewing. 3 8, male genitalia. 3, pygofer, lateral view. 4, valve and subgenital plate, ventral view. 5, connective and style, dorsal view. 6, ejaculatory reservoir, aedeagus, and basal segment of anal tube, lateral view. 7, paraphyses, laterodorsal view. 8, paraphyses, dorsal view. Scale bars: 1 = 1 mm, 2 = 2 mm, 3 8 = 0.5 mm." httpUri="https://zenodo.org/record/815999/files/figure.png" pageId="1" pageNumber="128">Figures 18</figureCitation>
,
<figureCitation id="3EF997969C3B5628CC04FD39FEBDFD27" box="[307,336,722,747]" captionStart="FIGURES 26 29" captionStartId="6.[151,263,1653,1676]" captionTargetBox="[184,1378,948,1599]" captionTargetId="figure@6.[151,1435,916,1632]" captionTargetPageId="6" captionText="FIGURES 26 29. Body in dorsal view (antennae and legs not depicted). 26, Dilobopterus nelsoni sp. nov., male holotype (total length 6.7 mm). 27, D. sakakibara sp. nov., male holotype (t. l. 6.9 mm). 28, D. segmentalis (Signoret, 1853), male (t. l. 9.0 mm). 29, D. trinotatus (Signoret, 1853), female (t. l. 9.0 mm). The last three species belong to the trinotatus group (see text for details)." httpUri="https://zenodo.org/record/816005/files/figure.png" pageId="1" pageNumber="128">26</figureCitation>
)
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<subSubSection id="EED8D8989C3B562BCDA0FCF1FB9BF986" lastPageId="2" lastPageNumber="129" pageId="1" pageNumber="128" type="description">
<paragraph id="A67D8B139C3B5628CDA0FCF1FC35FCFF" blockId="1.[151,1437,794,2007]" box="[151,984,794,819]" pageId="1" pageNumber="128">Total length 6.7 mm (male holotype), 6.66.8 mm (male paratypes, n = 5).</paragraph>
<paragraph id="A67D8B139C3B5628CDF0FCD5FAAEFBBB" blockId="1.[151,1437,794,2007]" pageId="1" pageNumber="128">
Head and thorax. Head (
<figureCitation id="3EF997969C3B5628CCE0FCD5FDC9FC9B" box="[471,548,830,855]" captionStart="FIGURES 1 8" captionStartId="2.[151,263,1323,1346]" captionTargetBox="[175,1395,312,1273]" captionTargetId="figure@2.[150,1436,301,1302]" captionTargetPageId="2" captionText="FIGURES 1 8. Dilobopterus nelsoni sp. nov. 1, head, pronotum, and mesonotum, dorsal view (male paratype). 2, forewing. 3 8, male genitalia. 3, pygofer, lateral view. 4, valve and subgenital plate, ventral view. 5, connective and style, dorsal view. 6, ejaculatory reservoir, aedeagus, and basal segment of anal tube, lateral view. 7, paraphyses, laterodorsal view. 8, paraphyses, dorsal view. Scale bars: 1 = 1 mm, 2 = 2 mm, 3 8 = 0.5 mm." httpUri="https://zenodo.org/record/815999/files/figure.png" pageId="1" pageNumber="128">Figs. 1</figureCitation>
,
<figureCitation id="3EF997969C3B5628CF05FCD5FDA2FC9B" box="[562,591,830,855]" captionStart="FIGURES 26 29" captionStartId="6.[151,263,1653,1676]" captionTargetBox="[184,1378,948,1599]" captionTargetId="figure@6.[151,1435,916,1632]" captionTargetPageId="6" captionText="FIGURES 26 29. Body in dorsal view (antennae and legs not depicted). 26, Dilobopterus nelsoni sp. nov., male holotype (total length 6.7 mm). 27, D. sakakibara sp. nov., male holotype (t. l. 6.9 mm). 28, D. segmentalis (Signoret, 1853), male (t. l. 9.0 mm). 29, D. trinotatus (Signoret, 1853), female (t. l. 9.0 mm). The last three species belong to the trinotatus group (see text for details)." httpUri="https://zenodo.org/record/816005/files/figure.png" pageId="1" pageNumber="128">26</figureCitation>
) moderately produced anteriorly; median length of crown approximately 1/ 2 of interocular width and 3/10 of transocular width; antennal ledges, in lateral view, with anterior margin steeply declivous, slightly concave superiorly and slightly convex inferiorly; face with frons convex, swollen, muscle impressions inconspicuous; epistomal suture incomplete; clypeus, in lateral view, slightly swollen, its profile continuing contour of lower portion of frons. Thorax (
<figureCitation id="3EF997969C3B5628CE27FC25FC8CFC2B" box="[784,865,974,999]" captionStart="FIGURES 1 8" captionStartId="2.[151,263,1323,1346]" captionTargetBox="[175,1395,312,1273]" captionTargetId="figure@2.[150,1436,301,1302]" captionTargetPageId="2" captionText="FIGURES 1 8. Dilobopterus nelsoni sp. nov. 1, head, pronotum, and mesonotum, dorsal view (male paratype). 2, forewing. 3 8, male genitalia. 3, pygofer, lateral view. 4, valve and subgenital plate, ventral view. 5, connective and style, dorsal view. 6, ejaculatory reservoir, aedeagus, and basal segment of anal tube, lateral view. 7, paraphyses, laterodorsal view. 8, paraphyses, dorsal view. Scale bars: 1 = 1 mm, 2 = 2 mm, 3 8 = 0.5 mm." httpUri="https://zenodo.org/record/815999/files/figure.png" pageId="1" pageNumber="128">Figs. 1</figureCitation>
,
<figureCitation id="3EF997969C3B5628CE45FC25FC62FC2B" box="[882,911,974,999]" captionStart="FIGURES 26 29" captionStartId="6.[151,263,1653,1676]" captionTargetBox="[184,1378,948,1599]" captionTargetId="figure@6.[151,1435,916,1632]" captionTargetPageId="6" captionText="FIGURES 26 29. Body in dorsal view (antennae and legs not depicted). 26, Dilobopterus nelsoni sp. nov., male holotype (total length 6.7 mm). 27, D. sakakibara sp. nov., male holotype (t. l. 6.9 mm). 28, D. segmentalis (Signoret, 1853), male (t. l. 9.0 mm). 29, D. trinotatus (Signoret, 1853), female (t. l. 9.0 mm). The last three species belong to the trinotatus group (see text for details)." httpUri="https://zenodo.org/record/816005/files/figure.png" pageId="1" pageNumber="128">26</figureCitation>
) with lateral pronotal margins approximately parallel; posterior margin slightly concave; dorsopleural carinae strongly declivous anteriorly. Forewings (
<figureCitation id="3EF997969C3B5628C870FC19FA7AFBC7" box="[1351,1431,1010,1035]" captionStart="FIGURES 1 8" captionStartId="2.[151,263,1323,1346]" captionTargetBox="[175,1395,312,1273]" captionTargetId="figure@2.[150,1436,301,1302]" captionTargetPageId="2" captionText="FIGURES 1 8. Dilobopterus nelsoni sp. nov. 1, head, pronotum, and mesonotum, dorsal view (male paratype). 2, forewing. 3 8, male genitalia. 3, pygofer, lateral view. 4, valve and subgenital plate, ventral view. 5, connective and style, dorsal view. 6, ejaculatory reservoir, aedeagus, and basal segment of anal tube, lateral view. 7, paraphyses, laterodorsal view. 8, paraphyses, dorsal view. Scale bars: 1 = 1 mm, 2 = 2 mm, 3 8 = 0.5 mm." httpUri="https://zenodo.org/record/815999/files/figure.png" pageId="1" pageNumber="128">Figs. 2</figureCitation>
,
<figureCitation id="3EF997969C3B5628CDA0FBFDFF59FBE3" box="[151,180,1046,1071]" captionStart="FIGURES 26 29" captionStartId="6.[151,263,1653,1676]" captionTargetBox="[184,1378,948,1599]" captionTargetId="figure@6.[151,1435,916,1632]" captionTargetPageId="6" captionText="FIGURES 26 29. Body in dorsal view (antennae and legs not depicted). 26, Dilobopterus nelsoni sp. nov., male holotype (total length 6.7 mm). 27, D. sakakibara sp. nov., male holotype (t. l. 6.9 mm). 28, D. segmentalis (Signoret, 1853), male (t. l. 9.0 mm). 29, D. trinotatus (Signoret, 1853), female (t. l. 9.0 mm). The last three species belong to the trinotatus group (see text for details)." httpUri="https://zenodo.org/record/816005/files/figure.png" pageId="1" pageNumber="128">26</figureCitation>
) mostly membranous, translucent; veins mostly distinct, except on clavus. Hind legs with meron slightly exposed; femoral setal formula 2:1:1; length of first tarsomere approximately equal to combined length of second and third tarsomeres. Other features of head and thorax as in the generic description of
<bibRefCitation id="C253F6E29C3B5628C95FFBB4FAD2FBBB" author="Young" box="[1128,1343,1119,1143]" pageId="1" pageNumber="128" refString="Young, D. A. (1977) Taxonomic study of the Cicadellinae (Homoptera: Cicadellidae). Part 2. New World Cicadellini and the genus Cicadella. Bulletin of the North Carolina Agricultural Experiment Station, 239, 1 - 1135." type="journal article" year="1977">Young (1977: 104)</bibRefCitation>
.
</paragraph>
<paragraph id="A67D8B139C3B5628CDF0FB69FDD3F97B" blockId="1.[151,1437,794,2007]" pageId="1" pageNumber="128">
Color (
<figureCitation id="3EF997969C3B5628CC21FB69FE6CFB57" box="[278,385,1154,1179]" captionStart="FIGURES 1 8" captionStartId="2.[151,263,1323,1346]" captionTargetBox="[175,1395,312,1273]" captionTargetId="figure@2.[150,1436,301,1302]" captionTargetPageId="2" captionText="FIGURES 1 8. Dilobopterus nelsoni sp. nov. 1, head, pronotum, and mesonotum, dorsal view (male paratype). 2, forewing. 3 8, male genitalia. 3, pygofer, lateral view. 4, valve and subgenital plate, ventral view. 5, connective and style, dorsal view. 6, ejaculatory reservoir, aedeagus, and basal segment of anal tube, lateral view. 7, paraphyses, laterodorsal view. 8, paraphyses, dorsal view. Scale bars: 1 = 1 mm, 2 = 2 mm, 3 8 = 0.5 mm." httpUri="https://zenodo.org/record/815999/files/figure.png" pageId="1" pageNumber="128">Figs. 12</figureCitation>
,
<figureCitation id="3EF997969C3B5628CCBAFB69FE40FB57" box="[397,429,1154,1179]" captionStart="FIGURES 26 29" captionStartId="6.[151,263,1653,1676]" captionTargetBox="[184,1378,948,1599]" captionTargetId="figure@6.[151,1435,916,1632]" captionTargetPageId="6" captionText="FIGURES 26 29. Body in dorsal view (antennae and legs not depicted). 26, Dilobopterus nelsoni sp. nov., male holotype (total length 6.7 mm). 27, D. sakakibara sp. nov., male holotype (t. l. 6.9 mm). 28, D. segmentalis (Signoret, 1853), male (t. l. 9.0 mm). 29, D. trinotatus (Signoret, 1853), female (t. l. 9.0 mm). The last three species belong to the trinotatus group (see text for details)." httpUri="https://zenodo.org/record/816005/files/figure.png" pageId="1" pageNumber="128">26</figureCitation>
). Ground color of anterior dorsum (head, pronotum, and mesonotum) dark brown to black with yellow spots as follows: crown with eight (
<typeStatus id="797935B19C3B5628CF87FB4DFCF8FB73" box="[688,789,1190,1215]" pageId="1" pageNumber="128" type="holotype">holotype</typeStatus>
) small spots, three pairs around ocelli, one pair on central portion close to posterior margin (two
<typeStatus id="797935B19C3B5628CF74FB20FD43FB2F" box="[579,686,1227,1251]" pageId="1" pageNumber="128" type="paratype">paratypes</typeStatus>
with additional pair on anterior margin); pronotum (
<typeStatus id="797935B19C3B5628C9C0FB21FAB1FB2F" box="[1271,1372,1226,1251]" pageId="1" pageNumber="128" type="holotype">holotype</typeStatus>
) with four spots, pair of smaller ones on anterior third and pair of larger ones, closer to lateral margins, on median third (two
<typeStatus id="797935B19C3B5628CDE2FAF8FEADFAE7" box="[213,320,1299,1323]" pageId="1" pageNumber="128" type="paratype">paratypes</typeStatus>
without anterior pair of spots; one
<typeStatus id="797935B19C3B5628CFDFFAF8FCA4FAE7" box="[744,841,1299,1323]" pageId="1" pageNumber="128" type="paratype">paratype</typeStatus>
with five small spots near anterior margin and no posterior pair of spots; one
<typeStatus id="797935B19C3B5628CCD5FADCFDAFFA83" box="[482,578,1335,1359]" pageId="1" pageNumber="128" type="paratype">paratype</typeStatus>
with four small spots near anterior margin plus posterior pair of spots); mesonotum (
<typeStatus id="797935B19C3B5628CC1EFAB1FE63FABF" box="[297,398,1370,1395]" pageId="1" pageNumber="128" type="holotype">holotype</typeStatus>
) with pair of smaller spots laterobasally on scutum and pair of larger spots laterobasally on scutellum (in the
<typeStatus id="797935B19C3B5628CC54FA94FE3CFA5B" box="[355,465,1407,1431]" pageId="1" pageNumber="128" type="paratype">paratypes</typeStatus>
, the first pair of spots can be very small or even absent). Forewings brown to dark brown, translucent, with large vitreous area on basal half occupying much of discal and brachial cells on corium and outer area of clavus but divided diagonally by dark brown claval sulcus. Face with frons dark brown to black medially and yellow laterally; centrolateral portions of frons with alternating dark brown to black and yellow narrow transverse stripes; clypeus, much of lorum, maxillary plate, and posterior area of gena dark brown to black, anterior area of gena yellow; labrum and labium mostly yellow. Thorax laterally mostly yellow, ventrally with more extensive dark brown to black areas; legs mostly yellow. Abdomen, in ventral view, yellow, with dark brown to black marks on basal portions and central area of sternites; subgenital plates, in ventral view, brownish-yellow, apex dark brown; pygofer mostly red.
</paragraph>
<paragraph id="A67D8B139C3B562BCDF0F929FEF7FF34" blockId="1.[151,1437,794,2007]" lastBlockId="2.[151,1436,151,248]" lastPageId="2" lastPageNumber="129" pageId="1" pageNumber="128">
Male genitalia. Pygofer (
<figureCitation id="3EF997969C3B5628CCD9F929FDDBF917" box="[494,566,1730,1755]" captionStart="FIGURES 1 8" captionStartId="2.[151,263,1323,1346]" captionTargetBox="[175,1395,312,1273]" captionTargetId="figure@2.[150,1436,301,1302]" captionTargetPageId="2" captionText="FIGURES 1 8. Dilobopterus nelsoni sp. nov. 1, head, pronotum, and mesonotum, dorsal view (male paratype). 2, forewing. 3 8, male genitalia. 3, pygofer, lateral view. 4, valve and subgenital plate, ventral view. 5, connective and style, dorsal view. 6, ejaculatory reservoir, aedeagus, and basal segment of anal tube, lateral view. 7, paraphyses, laterodorsal view. 8, paraphyses, dorsal view. Scale bars: 1 = 1 mm, 2 = 2 mm, 3 8 = 0.5 mm." httpUri="https://zenodo.org/record/815999/files/figure.png" pageId="1" pageNumber="128">Fig. 3</figureCitation>
), in lateral view, strongly produced posteriorly; posterior margin convex; without processes; with numerous dispersed macrosetae absent only on basal fourth. Subgenital plates (
<figureCitation id="3EF997969C3B5628C81CF90DFA9DF933" box="[1323,1392,1766,1791]" captionStart="FIGURES 1 8" captionStartId="2.[151,263,1323,1346]" captionTargetBox="[175,1395,312,1273]" captionTargetId="figure@2.[150,1436,301,1302]" captionTargetPageId="2" captionText="FIGURES 1 8. Dilobopterus nelsoni sp. nov. 1, head, pronotum, and mesonotum, dorsal view (male paratype). 2, forewing. 3 8, male genitalia. 3, pygofer, lateral view. 4, valve and subgenital plate, ventral view. 5, connective and style, dorsal view. 6, ejaculatory reservoir, aedeagus, and basal segment of anal tube, lateral view. 7, paraphyses, laterodorsal view. 8, paraphyses, dorsal view. Scale bars: 1 = 1 mm, 2 = 2 mm, 3 8 = 0.5 mm." httpUri="https://zenodo.org/record/815999/files/figure.png" pageId="1" pageNumber="128">Fig. 4</figureCitation>
), in ventral view, triangular, not extending as far posteriorly as pygofer apex; separate throughout their length; with uniseriate macrosetae along outer margin, microsetae also present. Connective (
<figureCitation id="3EF997969C3B5628C916F8C5FB8BF88B" box="[1057,1126,1838,1863]" captionStart="FIGURES 1 8" captionStartId="2.[151,263,1323,1346]" captionTargetBox="[175,1395,312,1273]" captionTargetId="figure@2.[150,1436,301,1302]" captionTargetPageId="2" captionText="FIGURES 1 8. Dilobopterus nelsoni sp. nov. 1, head, pronotum, and mesonotum, dorsal view (male paratype). 2, forewing. 3 8, male genitalia. 3, pygofer, lateral view. 4, valve and subgenital plate, ventral view. 5, connective and style, dorsal view. 6, ejaculatory reservoir, aedeagus, and basal segment of anal tube, lateral view. 7, paraphyses, laterodorsal view. 8, paraphyses, dorsal view. Scale bars: 1 = 1 mm, 2 = 2 mm, 3 8 = 0.5 mm." httpUri="https://zenodo.org/record/815999/files/figure.png" pageId="1" pageNumber="128">Fig. 5</figureCitation>
), in dorsal view, Y-shaped; with median keel; not extending as far posteriorly as style apex. Style (
<figureCitation id="3EF997969C3B5628CEEDF8B9FBCCF8A7" box="[986,1057,1874,1899]" captionStart="FIGURES 1 8" captionStartId="2.[151,263,1323,1346]" captionTargetBox="[175,1395,312,1273]" captionTargetId="figure@2.[150,1436,301,1302]" captionTargetPageId="2" captionText="FIGURES 1 8. Dilobopterus nelsoni sp. nov. 1, head, pronotum, and mesonotum, dorsal view (male paratype). 2, forewing. 3 8, male genitalia. 3, pygofer, lateral view. 4, valve and subgenital plate, ventral view. 5, connective and style, dorsal view. 6, ejaculatory reservoir, aedeagus, and basal segment of anal tube, lateral view. 7, paraphyses, laterodorsal view. 8, paraphyses, dorsal view. Scale bars: 1 = 1 mm, 2 = 2 mm, 3 8 = 0.5 mm." httpUri="https://zenodo.org/record/815999/files/figure.png" pageId="1" pageNumber="128">Fig. 5</figureCitation>
), in dorsal view, with apophysis elongate; without conspicuous preapical lobe; apex obliquely truncate, foot-shaped. Aedeagus (
<figureCitation id="3EF997969C3B5628C873F89DFA79F843" box="[1348,1428,1910,1935]" captionStart="FIGURES 1 8" captionStartId="2.[151,263,1323,1346]" captionTargetBox="[175,1395,312,1273]" captionTargetId="figure@2.[150,1436,301,1302]" captionTargetPageId="2" captionText="FIGURES 1 8. Dilobopterus nelsoni sp. nov. 1, head, pronotum, and mesonotum, dorsal view (male paratype). 2, forewing. 3 8, male genitalia. 3, pygofer, lateral view. 4, valve and subgenital plate, ventral view. 5, connective and style, dorsal view. 6, ejaculatory reservoir, aedeagus, and basal segment of anal tube, lateral view. 7, paraphyses, laterodorsal view. 8, paraphyses, dorsal view. Scale bars: 1 = 1 mm, 2 = 2 mm, 3 8 = 0.5 mm." httpUri="https://zenodo.org/record/815999/files/figure.png" pageId="1" pageNumber="128">Fig. 6</figureCitation>
) symmetrical; shaft, in lateral view, with large dorsal lobe; without processes; gonoduct distinct; gonopore located apically. Paraphyses (
<figureCitation id="3EF997969C3B5628CCBAF855FE11F81B" box="[397,508,1982,2007]" captionStart="FIGURES 1 8" captionStartId="2.[151,263,1323,1346]" captionTargetBox="[175,1395,312,1273]" captionTargetId="figure@2.[150,1436,301,1302]" captionTargetPageId="2" captionText="FIGURES 1 8. Dilobopterus nelsoni sp. nov. 1, head, pronotum, and mesonotum, dorsal view (male paratype). 2, forewing. 3 8, male genitalia. 3, pygofer, lateral view. 4, valve and subgenital plate, ventral view. 5, connective and style, dorsal view. 6, ejaculatory reservoir, aedeagus, and basal segment of anal tube, lateral view. 7, paraphyses, laterodorsal view. 8, paraphyses, dorsal view. Scale bars: 1 = 1 mm, 2 = 2 mm, 3 8 = 0.5 mm." httpUri="https://zenodo.org/record/815999/files/figure.png" pageId="1" pageNumber="128">Figs. 78</figureCitation>
), in dorsal view, with stalk elongate and expanded apically; expanded portion of stalk bearing pair of processes directed anteriorly; rami conspicuously asymmetrical; left ramus with larger spine basally and smaller one medially; right ramus bifurcated basally into ventral and dorsal arms, both armed with inner spine.
</paragraph>
<caption id="F2BDDB9B9C38562BCDA0FAC0FD13FA52" httpUri="https://zenodo.org/record/815999/files/figure.png" pageId="2" pageNumber="129" targetBox="[175,1395,312,1273]" targetPageId="2">
<paragraph id="A67D8B139C38562BCDA0FAC0FD13FA52" blockId="2.[151,1437,1323,1438]" pageId="2" pageNumber="129">
<emphasis id="94B657019C38562BCDA0FAC0FED6FA8D" bold="true" box="[151,315,1323,1346]" pageId="2" pageNumber="129">FIGURES 18.</emphasis>
<taxonomicName id="61C2F0909C38562BCC75FAC7FDF5FA8E" authority="Quintas &amp; Felix &amp; Lima &amp; Mejdalani, 2017" authorityName="Quintas &amp; Felix &amp; Lima &amp; Mejdalani" authorityYear="2017" box="[322,536,1324,1346]" class="Insecta" family="Cicadellidae" genus="Dilobopterus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="nelsoni" status="sp. nov.">
<emphasis id="94B657019C38562BCC75FAC7FDF5FA8E" box="[322,536,1324,1346]" italics="true" pageId="2" pageNumber="129">Dilobopterus nelsoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8F85EA7A9C38562BCF17FAC7FD9CFA8D" box="[544,625,1324,1345]" pageId="2" pageNumber="129" rank="species">
<emphasis id="94B657019C38562BCF17FAC7FD9CFA8D" bold="true" box="[544,625,1324,1345]" pageId="2" pageNumber="129">sp. nov.</emphasis>
</taxonomicNameLabel>
1, head, pronotum, and mesonotum, dorsal view (male paratype). 2, forewing. 38, male genitalia. 3, pygofer, lateral view. 4, valve and subgenital plate, ventral view. 5, connective and style, dorsal view. 6, ejaculatory reservoir, aedeagus, and basal segment of anal tube, lateral view. 7, paraphyses, laterodorsal view. 8, paraphyses, dorsal view. Scale bars: 1 = 1 mm, 2 = 2 mm, 38 = 0.5 mm.
</paragraph>
</caption>
<paragraph id="A67D8B139C38562BCDF0FA2EFE71FA12" blockId="2.[151,1437,1477,2006]" box="[199,412,1477,1502]" pageId="2" pageNumber="129">
<emphasis id="94B657019C38562BCDF0FA2EFE71FA12" bold="true" box="[199,412,1477,1502]" pageId="2" pageNumber="129">Female unknown.</emphasis>
</paragraph>
<paragraph id="A67D8B139C38562BCDF0FA02FD09F9EA" blockId="2.[151,1437,1477,2006]" pageId="2" pageNumber="129">
<emphasis id="94B657019C38562BCDF0FA02FE0EF9CE" bold="true" box="[199,483,1513,1538]" pageId="2" pageNumber="129">Intraspecific variation.</emphasis>
The number and position of the yellow spots of the anterior dorsum vary intraspecifically as detailed above in the description.
</paragraph>
<paragraph id="A67D8B139C38562BCDF0F9DAFB9BF986" blockId="2.[151,1437,1477,2006]" box="[199,1142,1585,1610]" pageId="2" pageNumber="129">
<materialsCitation id="16AA814E9C38562BCDF0F9DAFB9BF986" ID-GBIF-Occurrence="1950907505" box="[199,1142,1585,1610]" country="Brazil" location="Rio de Janeiro" pageId="2" pageNumber="129" specimenCount="1" stateProvince="Rio de Janeiro">
<emphasis id="94B657019C38562BCDF0F9DAFE57F986" bold="true" box="[199,442,1585,1610]" pageId="2" pageNumber="129">Known distribution.</emphasis>
Atlantic Rainforest, Southeastern
<collectingCountry id="DED5CB839C38562BCE0BF9DAFC6EF986" box="[828,899,1585,1610]" name="Brazil" pageId="2" pageNumber="129">Brazil</collectingCountry>
,
<collectingRegion id="640645F19C38562BCEB9F9DAFBDDF986" box="[910,1072,1585,1610]" country="Brazil" name="Rio de Janeiro" pageId="2" pageNumber="129">Rio de Janeiro</collectingRegion>
State.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="EED8D8989C38562BCDF0F9BEFBB4F97A" pageId="2" pageNumber="129" type="materials_examined">
<paragraph id="A67D8B139C38562BCDF0F9BEFBB4F97A" blockId="2.[151,1437,1477,2006]" pageId="2" pageNumber="129">
<materialsCitation id="16AA814E9C38562BCDF0F9BEFE32F95E" ID-GBIF-Occurrence="1950907602" collectingDate="2002-02-26" collectionCode="MNRJ" collectorName="Angra dos Reis &amp; Usina" location="Rio de Janeiro" pageId="2" pageNumber="129" specimenCount="2" stateProvince="Rio de Janeiro" typeStatus="holotype">
<emphasis id="94B657019C38562BCDF0F9BEFE98F9A2" bold="true" box="[199,373,1621,1646]" pageId="2" pageNumber="129">
<typeStatus id="797935B19C38562BCDF0F9BEFEEFF9A2" box="[199,258,1621,1646]" pageId="2" pageNumber="129">Type</typeStatus>
material.
</emphasis>
Male
<typeStatus id="797935B19C38562BCC8AF9BEFDCEF9A2" box="[445,547,1621,1646]" pageId="2" pageNumber="129" type="holotype">holotype</typeStatus>
: “BR/RJ [
<collectingRegion id="640645F19C38562BCFA3F9BEFCD4F9A2" box="[660,825,1621,1646]" country="Brazil" name="Rio de Janeiro" pageId="2" pageNumber="129">Rio de Janeiro</collectingRegion>
State] -
<collectorName id="0B37EEC59C38562BCEADF9BDFBA7F9A2" box="[922,1098,1621,1646]" pageId="2" pageNumber="129">Angra dos Reis</collectorName>
, \
<collectorName id="0B37EEC59C38562BC952F9BEFB4BF9A2" box="[1125,1190,1621,1646]" pageId="2" pageNumber="129">Usina</collectorName>
nuclear \
<collectingDate id="C238543B9C38562BC820F9BEFA60F9A2" box="[1303,1421,1621,1646]" pageId="2" pageNumber="129" value="2002-02-26">
<date id="D27CADD39C38562BC820F9BEFA60F9A2" box="[1303,1421,1621,1646]" pageId="2" pageNumber="129" value="2002-02-26">26/II/2002</date>
</collectingDate>
\ N. Ferreira-Jr. col.” (
<collectionCode id="C0D313D69C38562BCCB1F991FE3AF95E" box="[390,471,1658,1682]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="2" pageNumber="129">MNRJ</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="16AA814E9C38562BCCDCF991FBB8F97A" ID-GBIF-Occurrence="1950907605" collectingDate="2002-05-24" collectingDateMax="2002-05-25" collectingDateMin="2002-05-24" collectionCode="MNRJ, DZRJ" collectorName="Angra dos Reis &amp; Pereque" location="Rio de Janeiro" pageId="2" pageNumber="129" specimenCount="2" stateProvince="Rio de Janeiro" typeStatus="paratype">
<typeStatus id="797935B19C38562BCCDCF991FDB1F95E" box="[491,604,1658,1682]" pageId="2" pageNumber="129" type="paratype">Paratypes</typeStatus>
: five males, four with the same data as the
<typeStatus id="797935B19C38562BC967F992FB5FF95E" box="[1104,1202,1657,1682]" pageId="2" pageNumber="129" type="holotype">holotype</typeStatus>
(
<collectionCode id="C0D313D69C38562BC9F5F991FAFCF95E" box="[1218,1297,1658,1682]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="2" pageNumber="129">MNRJ</collectionCode>
,
<collectionCode id="C0D313D69C38562BC828F991FA89F95E" box="[1311,1380,1658,1682]" pageId="2" pageNumber="129">DZRJ</collectionCode>
) and one “BR/RJ -
<collectorName id="0B37EEC59C38562BCC02F975FE09F97A" box="[309,484,1693,1718]" pageId="2" pageNumber="129">Angra dos Reis</collectorName>
, \
<collectorName id="0B37EEC59C38562BCCC9F975FDB5F97A" box="[510,600,1694,1718]" pageId="2" pageNumber="129">Perequê</collectorName>
<collectingDate id="C238543B9C38562BCF68F976FD11F97A" box="[607,764,1693,1718]" pageId="2" pageNumber="129" value="2002-05-24" valueMax="2002-05-25" valueMin="2002-05-24">
<date id="D27CADD39C38562BCF68F976FD11F97A" box="[607,764,1693,1718]" pageId="2" pageNumber="129" value="2002-05-24" valueMax="2002-05-25" valueMin="2002-05-24">24-25/V/2002</date>
</collectingDate>
\ N. Ferreira-Jr. col.” (
<collectionCode id="C0D313D69C38562BCECBF975FBA0F97A" box="[1020,1101,1694,1718]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="2" pageNumber="129">MNRJ</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="EED8D8989C38562ACDF0F92AFC8CFEA8" lastPageId="3" lastPageNumber="130" pageId="2" pageNumber="129" type="etymology">
<paragraph id="A67D8B139C38562BCDF0F92AFB4FF8EE" blockId="2.[151,1437,1477,2006]" pageId="2" pageNumber="129">
<emphasis id="94B657019C38562BCDF0F92AFEA3F916" bold="true" box="[199,334,1729,1754]" pageId="2" pageNumber="129">Etymology.</emphasis>
The new species is described in honor of our colleague and friend Prof. Dr. Nelson Ferreira Junior (Departamento de Zoologia, Instituto de Biologia, Universidade Federal do
<collectingRegion id="640645F19C38562BCECCF90EFB4EF932" box="[1019,1187,1765,1790]" country="Brazil" name="Rio de Janeiro" pageId="2" pageNumber="129">Rio de Janeiro</collectingRegion>
) in recognition of his contribution to the study of Brazilian aquatic beetles. He was the collector of the
<typeStatus id="797935B19C38562BC913F8E1FBB9F8EE" box="[1060,1108,1802,1826]" pageId="2" pageNumber="129">type</typeStatus>
series.
</paragraph>
<paragraph id="A67D8B139C38562BCDF0F8C6FA71F81A" blockId="2.[151,1437,1477,2006]" pageId="2" pageNumber="129">
<emphasis id="94B657019C38562BCDF0F8C6FE7BF88A" bold="true" box="[199,406,1837,1862]" pageId="2" pageNumber="129">Taxonomic notes.</emphasis>
Among the known
<taxonomicName id="61C2F0909C38562BCF42F8C6FCE4F889" authorityName="Signoret" authorityYear="1850" box="[629,777,1837,1861]" class="Insecta" family="Cicadellidae" genus="Dilobopterus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis id="94B657019C38562BCF42F8C6FCE4F889" box="[629,777,1837,1861]" italics="true" pageId="2" pageNumber="129">Dilobopterus</emphasis>
</taxonomicName>
species,
<taxonomicName id="61C2F0909C38562BCE58F8C5FC0EF889" authority="Quintas &amp; Felix &amp; Lima &amp; Mejdalani, 2017" authorityName="Quintas &amp; Felix &amp; Lima &amp; Mejdalani" authorityYear="2017" box="[879,995,1837,1861]" class="Insecta" family="Cicadellidae" genus="Dilobopterus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="nelsoni" status="sp. nov.">
<emphasis id="94B657019C38562BCE58F8C5FC0EF889" box="[879,995,1837,1861]" italics="true" pageId="2" pageNumber="129">D. nelsoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8F85EA7A9C38562BCEDDF8C6FBA9F88A" box="[1002,1092,1837,1862]" pageId="2" pageNumber="129" rank="species">
<emphasis id="94B657019C38562BCEDDF8C6FBA9F88A" bold="true" box="[1002,1092,1837,1862]" pageId="2" pageNumber="129">sp. nov.</emphasis>
</taxonomicNameLabel>
is most similar to
<taxonomicName id="61C2F0909C38562BC823F8C5FEB1F8A6" authority="Fabricius, 1803" authorityName="Fabricius" authorityYear="1803" class="Insecta" family="Cicadellidae" genus="Dilobopterus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="demissus">
<emphasis id="94B657019C38562BC823F8C5FA71F889" box="[1300,1436,1837,1861]" italics="true" pageId="2" pageNumber="129">D. demissus</emphasis>
(Fabricius, 1803)
</taxonomicName>
,
<taxonomicName id="61C2F0909C38562BCC5CF8B9FD7EF8A6" authority="Germar, 1821" authorityName="Germar" authorityYear="1821" box="[363,659,1873,1898]" class="Insecta" family="Cicadellidae" genus="Dilobopterus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="dispar">
<emphasis id="94B657019C38562BCC5CF8B9FE3BF8A5" box="[363,470,1873,1897]" italics="true" pageId="2" pageNumber="129">D. dispar</emphasis>
(Germar, 1821)
</taxonomicName>
, and
<taxonomicName id="61C2F0909C38562BCFE4F8B9FBE1F8A6" authority="Young, 1977" authorityName="Young, 1977" authorityYear="1977" box="[723,1036,1873,1898]" class="Insecta" family="Cicadellidae" genus="Dilobopterus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="disparulus">
<emphasis id="94B657019C38562BCFE4F8B9FC83F8A5" box="[723,878,1873,1897]" italics="true" pageId="2" pageNumber="129">D. disparulus</emphasis>
<bibRefCitation id="C253F6E29C38562BCE40F8B9FBE1F8A6" author="Young" box="[887,1036,1874,1898]" pageId="2" pageNumber="129" refString="Young, D. A. (1977) Taxonomic study of the Cicadellinae (Homoptera: Cicadellidae). Part 2. New World Cicadellini and the genus Cicadella. Bulletin of the North Carolina Agricultural Experiment Station, 239, 1 - 1135." type="journal article" year="1977">Young, 1977</bibRefCitation>
</taxonomicName>
. In these three species, the ground color of the anterior dorsum is dark brown or black with brown or yellow marks (see
<bibRefCitation id="C253F6E29C38562BC951F89EFAD7F842" author="Wilson" box="[1126,1338,1909,1934]" pageId="2" pageNumber="129" refString="Wilson, M. R., Turner, J. A. &amp; McKamey, S. H. (2009) Sharpshooter leafhoppers of the world (Hemiptera: Cicadellidae subfamily Cicadellinae). National Museum Wales. Available from: http: // naturalhistory. museumwales. ac. uk / sharpshooters / home. php (accessed 26 July 2016)" type="journal article" year="2009">
Wilson
<emphasis id="94B657019C38562BC989F89DFB15F841" box="[1214,1272,1909,1933]" italics="true" pageId="2" pageNumber="129">et al.</emphasis>
2009
</bibRefCitation>
) and the rami of the paraphyses are asymmetrical.
<taxonomicName id="61C2F0909C38562BCFB4F872FC9FF87D" authority="Quintas &amp; Felix &amp; Lima &amp; Mejdalani, 2017" authorityName="Quintas &amp; Felix &amp; Lima &amp; Mejdalani" authorityYear="2017" box="[643,882,1945,1969]" class="Insecta" family="Cicadellidae" genus="Dilobopterus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="nelsoni" status="sp. nov.">
<emphasis id="94B657019C38562BCFB4F872FC9FF87D" box="[643,882,1945,1969]" italics="true" pageId="2" pageNumber="129">Dilobopterus nelsoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8F85EA7A9C38562BCE4AF872FC36F87E" box="[893,987,1945,1970]" pageId="2" pageNumber="129" rank="species">
<emphasis id="94B657019C38562BCE4AF872FC36F87E" bold="true" box="[893,987,1945,1970]" pageId="2" pageNumber="129">sp. nov.</emphasis>
</taxonomicNameLabel>
can be distinguished from these three species, as well as from the remaining ones of the genus, by the following combination of features: (1) crown (
<figureCitation id="3EF997969C38562BC853F856FA71F81A" box="[1380,1436,1981,2006]" captionStart="FIGURES 1 8" captionStartId="2.[151,263,1323,1346]" captionTargetBox="[175,1395,312,1273]" captionTargetId="figure@2.[150,1436,301,1302]" captionTargetPageId="2" captionText="FIGURES 1 8. Dilobopterus nelsoni sp. nov. 1, head, pronotum, and mesonotum, dorsal view (male paratype). 2, forewing. 3 8, male genitalia. 3, pygofer, lateral view. 4, valve and subgenital plate, ventral view. 5, connective and style, dorsal view. 6, ejaculatory reservoir, aedeagus, and basal segment of anal tube, lateral view. 7, paraphyses, laterodorsal view. 8, paraphyses, dorsal view. Scale bars: 1 = 1 mm, 2 = 2 mm, 3 8 = 0.5 mm." httpUri="https://zenodo.org/record/815999/files/figure.png" pageId="2" pageNumber="129">Figs.</figureCitation>
</paragraph>
<paragraph id="A67D8B139C39562ACDA0FF73FC8CFEA8" blockId="3.[151,1436,151,357]" pageId="3" pageNumber="130">
1,
<figureCitation id="3EF997969C39562ACD84FF7CFF3DFF7C" box="[179,208,151,176]" captionStart="FIGURES 26 29" captionStartId="6.[151,263,1653,1676]" captionTargetBox="[184,1378,948,1599]" captionTargetId="figure@6.[151,1435,916,1632]" captionTargetPageId="6" captionText="FIGURES 26 29. Body in dorsal view (antennae and legs not depicted). 26, Dilobopterus nelsoni sp. nov., male holotype (total length 6.7 mm). 27, D. sakakibara sp. nov., male holotype (t. l. 6.9 mm). 28, D. segmentalis (Signoret, 1853), male (t. l. 9.0 mm). 29, D. trinotatus (Signoret, 1853), female (t. l. 9.0 mm). The last three species belong to the trinotatus group (see text for details)." httpUri="https://zenodo.org/record/816005/files/figure.png" pageId="3" pageNumber="130">26</figureCitation>
) with eight small yellow spots, being three pairs around the ocelli and one pair on the central portion close to the posterior margin (an additional pair can be present on the apical portion); (2) stalk of the paraphyses (
<figureCitation id="3EF997969C39562AC878FF57FA7EFF18" box="[1359,1427,188,213]" captionStart="FIGURES 1 8" captionStartId="2.[151,263,1323,1346]" captionTargetBox="[175,1395,312,1273]" captionTargetId="figure@2.[150,1436,301,1302]" captionTargetPageId="2" captionText="FIGURES 1 8. Dilobopterus nelsoni sp. nov. 1, head, pronotum, and mesonotum, dorsal view (male paratype). 2, forewing. 3 8, male genitalia. 3, pygofer, lateral view. 4, valve and subgenital plate, ventral view. 5, connective and style, dorsal view. 6, ejaculatory reservoir, aedeagus, and basal segment of anal tube, lateral view. 7, paraphyses, laterodorsal view. 8, paraphyses, dorsal view. Scale bars: 1 = 1 mm, 2 = 2 mm, 3 8 = 0.5 mm." httpUri="https://zenodo.org/record/815999/files/figure.png" pageId="3" pageNumber="130">Fig. 8</figureCitation>
) elongate, expanded apically and with a pair of processes directed anteriorly; (3) rami of the paraphyses (
<figureCitation id="3EF997969C39562AC81EFF34FA79FF34" box="[1321,1428,223,248]" captionStart="FIGURES 1 8" captionStartId="2.[151,263,1323,1346]" captionTargetBox="[175,1395,312,1273]" captionTargetId="figure@2.[150,1436,301,1302]" captionTargetPageId="2" captionText="FIGURES 1 8. Dilobopterus nelsoni sp. nov. 1, head, pronotum, and mesonotum, dorsal view (male paratype). 2, forewing. 3 8, male genitalia. 3, pygofer, lateral view. 4, valve and subgenital plate, ventral view. 5, connective and style, dorsal view. 6, ejaculatory reservoir, aedeagus, and basal segment of anal tube, lateral view. 7, paraphyses, laterodorsal view. 8, paraphyses, dorsal view. Scale bars: 1 = 1 mm, 2 = 2 mm, 3 8 = 0.5 mm." httpUri="https://zenodo.org/record/815999/files/figure.png" pageId="3" pageNumber="130">Figs. 78</figureCitation>
) strongly asymmetrical and armed with spines (left ramus with a larger spine basally and a smaller one medially; right ramus bifurcated basally into ventral and dorsal arms, both armed with an inner spine); and (4) dorsal lobe of the aedeagus (
<figureCitation id="3EF997969C39562ACC01FEA7FE94FEA9" box="[310,377,332,357]" captionStart="FIGURES 1 8" captionStartId="2.[151,263,1323,1346]" captionTargetBox="[175,1395,312,1273]" captionTargetId="figure@2.[150,1436,301,1302]" captionTargetPageId="2" captionText="FIGURES 1 8. Dilobopterus nelsoni sp. nov. 1, head, pronotum, and mesonotum, dorsal view (male paratype). 2, forewing. 3 8, male genitalia. 3, pygofer, lateral view. 4, valve and subgenital plate, ventral view. 5, connective and style, dorsal view. 6, ejaculatory reservoir, aedeagus, and basal segment of anal tube, lateral view. 7, paraphyses, laterodorsal view. 8, paraphyses, dorsal view. Scale bars: 1 = 1 mm, 2 = 2 mm, 3 8 = 0.5 mm." httpUri="https://zenodo.org/record/815999/files/figure.png" pageId="3" pageNumber="130">Fig. 6</figureCitation>
) large, without emarginations or processes.
</paragraph>
</subSubSection>
</treatment>
</document>