237 lines
31 KiB
XML
237 lines
31 KiB
XML
<document id="C1FA93E5385A63E44537C407BF792BDB" ID-DOI="10.5281/zenodo.3726987" ID-GBIF-Dataset="80783542-5446-42b3-a9bd-177b0e4ec73a" ID-ISSN="1942-1354" ID-Zenodo-Dep="3726987" ID-ZooBank="CF6C1B9C-2EC0-4220-BED7-6402A6C9D175" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1585149460741" checkinUser="felipe" docAuthor="Zidek, Jiri & Pokorný, Svatopluk" docDate="2018" docId="1E76DE47FFE1312B52FAFB1FE839FB24" docLanguage="en" docName="InsectaMundi.0611.1-35.pdf.imf" docOrigin="Insecta Mundi 611" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Scarabaeus (Scarabaeolus) similis Zidek and Pokorny 2018, sp. n." docType="treatment" docVersion="5" lastPageNumber="11" masterDocId="E24FA63FFFEA3127523AFFD7EC13FFD4" masterDocTitle="New species of the Scarabaeus subgenus Scarabaeolus Balthasar, with a review of the subgenus (Scarabaeidae: Scarabaeinae: Scarabaeini)" masterLastPageNumber="35" masterPageNumber="1" pageNumber="10" updateTime="1698731770912" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
|
||
<mods:mods id="488CBBDB49018EB29D518BB3FB26CDB3" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="1B70BD0AEB3795F9F6534BE02876A2E4">
|
||
<mods:title id="7D97AC0B174981A18C2D3EBCFEC7FE67">New species of the Scarabaeus subgenus Scarabaeolus Balthasar, with a review of the subgenus (Scarabaeidae: Scarabaeinae: Scarabaeini)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="D63DFE0138AFD46CBE5474F63CE4F52C" type="personal">
|
||
<mods:role id="C6C0D3E8BD452607AC6A7700F1B3E13A">
|
||
<mods:roleTerm id="081616DC6B198D1DB915024D39297225">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="CC0CC229A618DA587166D9A31F2F7373">Zidek, Jiri</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="83B691493E242C3A26DDDEC00EAE1DE5" type="personal">
|
||
<mods:role id="EFC02A373338B017816171E01BF99910">
|
||
<mods:roleTerm id="8A88C6B81AABF88DEDA461DC4B4EEAC9">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="AF27517375C02E3713CB8DEBAD2FF237">Pokorný, Svatopluk</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="5BD5E60356F58389715EB840CBC334A9">text</mods:typeOfResource>
|
||
<mods:relatedItem id="1159A56628E1C10CC8CAEF0A6D83326D" type="host">
|
||
<mods:titleInfo id="1C5EDA644684735D57FF5F9E22E6B251">
|
||
<mods:title id="88EE7BD55A5740BCAEF75DC26DC9F28E">Insecta Mundi</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="1B6834E4268130018E2CE76292C8F63C">
|
||
<mods:date id="EDA83DB86E0237376C4BF3324DD13691">2018</mods:date>
|
||
<mods:detail id="BA44791D33B73C5221A766FAB493D796" type="pubDate">
|
||
<mods:number id="730F7E146988CEABBF41A8918A9E2006">2018-03-30</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="7B426E6A7722442FC6675AC92C505C81" type="issue">
|
||
<mods:number id="C103BDBAB5109187CD2EFDCCFEEC865C">611</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="FD225679161328931B6AD8B20571FE9B" unit="page">
|
||
<mods:start id="9344DCEEBE0E64FBF2C9C848D3096DF1">1</mods:start>
|
||
<mods:end id="E7EA2A3E32E892BE043C4D53953762CE">35</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="84E77BE7292A66FB0BE87ACD590696DA">journal article</mods:classification>
|
||
<mods:identifier id="10453E4DAB58D758AC2A32DFFFEFDD5D" type="DOI">10.5281/zenodo.3726987</mods:identifier>
|
||
<mods:identifier id="ECC3CB40802F98CB6779779492389C7A" type="GBIF-Dataset">80783542-5446-42b3-a9bd-177b0e4ec73a</mods:identifier>
|
||
<mods:identifier id="5641EF97415332A108347298BDF95B3C" type="ISSN">1942-1354</mods:identifier>
|
||
<mods:identifier id="EFD98D44891A7F0EDCA13D705B105ED7" type="Zenodo-Dep">3726987</mods:identifier>
|
||
<mods:identifier id="048D398AFB75A69EECA961F8BDAD89A5" type="ZooBank">CF6C1B9C-2EC0-4220-BED7-6402A6C9D175</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="1E76DE47FFE1312B52FAFB1FE839FB24" ID-DOI="http://doi.org/10.5281/zenodo.3729627" ID-GBIF-Taxon="163288773" ID-Zenodo-Dep="3729627" LSID="urn:lsid:plazi:treatment:1E76DE47FFE1312B52FAFB1FE839FB24" httpUri="http://treatment.plazi.org/id/1E76DE47FFE1312B52FAFB1FE839FB24" lastPageId="12" lastPageNumber="11" pageId="11" pageNumber="10">
|
||
<subSubSection id="DEC53CDAFFE1312C52FAFB1FEFECFB35" box="[192,1023,1224,1249]" pageId="11" pageNumber="10" type="nomenclature">
|
||
<paragraph id="96606F51FFE1312C52FAFB1FEFECFB35" blockId="11.[192,1023,1224,1281]" box="[192,1023,1224,1249]" pageId="11" pageNumber="10">
|
||
<heading id="CD28D83DFFE1312C52FAFB1FEFECFB35" bold="true" box="[192,1023,1224,1249]" fontSize="10" level="2" pageId="11" pageNumber="10" reason="3">
|
||
<emphasis id="A4ABB343FFE1312C52FAFB1FEFECFB35" bold="true" box="[192,1023,1224,1249]" pageId="11" pageNumber="10">
|
||
<taxonomicName id="51DF14D2FFE1312C52FAFB1FEFB9FB34" ID-CoL="79VTY" authority="Zidek and Pokorny" authorityName="Zidek and Pokorny" authorityYear="2018" box="[192,938,1224,1249]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="similis" status="sp. nov." subGenus="Scarabaeolus">
|
||
<emphasis id="A4ABB343FFE1312C52FAFB1FED73FB34" bold="true" box="[192,352,1224,1248]" italics="true" pageId="11" pageNumber="10">Scarabaeus</emphasis>
|
||
(
|
||
<emphasis id="A4ABB343FFE1312C5348FB1FEE3FFB34" bold="true" box="[370,556,1224,1248]" italics="true" pageId="11" pageNumber="10">Scarabaeolus</emphasis>
|
||
)
|
||
<emphasis id="A4ABB343FFE1312C5004FB1FEE89FB34" bold="true" box="[574,666,1224,1248]" italics="true" pageId="11" pageNumber="10">similis</emphasis>
|
||
Zidek and Pokorný
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="BF980E38FFE1312C5182FB1EEFECFB35" box="[952,1023,1225,1249]" pageId="11" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DEC53CDAFFE1312C52FAFB3EED52FAD5" box="[192,321,1257,1281]" pageId="11" pageNumber="10" type="description">
|
||
<paragraph id="96606F51FFE1312C52FAFB3EED52FAD5" blockId="11.[192,1023,1224,1281]" box="[192,321,1257,1281]" pageId="11" pageNumber="10">
|
||
<figureCitation id="0EE473D4FFE1312C52FAFB3EED52FAD5" box="[192,321,1257,1281]" captionStart="Figures 55–62" captionStartId="32.[192,287,1819,1841]" captionTargetBox="[212,1424,293,1748]" captionTargetPageId="32" captionText="Figures 55–62. Scarabaeus (Scarabaeolus) spp. 55–58) Scarabaeus (Scarabaeolus) similis sp. n., HTm, 14 mm. 55) Dorsal habitus. 56) Ventral habitus. 57) Aedeagus, dorsal. 58) Aedeagus, left lateral. 59–62) Scarabaeus (Scarabaeolus) megaparvulus Davis and Deschodt, 12 mm. 59) Dorsal habitus. 60) Ventral habitus. 61) Aedeagus, dorsal. 62) Aedeagus, left lateral." figureDoi="http://doi.org/10.5281/zenodo.3727003" httpUri="https://zenodo.org/record/3727003/files/figure.png" pageId="11" pageNumber="10">Fig. 55–58</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DEC53CDAFFE1312C52FAFACDE8BCFAE6" box="[192,1199,1305,1330]" pageId="11" pageNumber="10" type="distribution">
|
||
<paragraph id="96606F51FFE1312C52FAFACDE8BCFAE6" blockId="11.[192,1199,1305,1330]" box="[192,1199,1305,1330]" pageId="11" pageNumber="10">
|
||
<emphasis id="A4ABB343FFE1312C52FAFACDED68FAE5" bold="true" box="[192,379,1305,1330]" pageId="11" pageNumber="10">
|
||
<typeStatus id="4964D1F3FFE1312C52FAFACDED17FAE6" box="[192,260,1306,1330]" pageId="11" pageNumber="10">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
N.
|
||
<collectingRegion id="541BA1B3FFE1312C539CFACDEE3CFAE5" box="[422,559,1306,1330]" country="South Africa" name="Western Cape" pageId="11" pageNumber="10">Cape Prov.</collectingRegion>
|
||
, 8.5 km S Pienaarsriver, leg. C. Bellamy
|
||
<date id="E2614991FFE1312C561CFACDE8B9FAE6" box="[1062,1194,1306,1330]" pageId="11" pageNumber="10" value="1999-07-04">4.VII.1999</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DEC53CDAFFE1312C52FAFA9DEF53FAB6" box="[192,832,1354,1378]" pageId="11" pageNumber="10" type="materials_examined">
|
||
<paragraph id="96606F51FFE1312C52FAFA9DEF53FAB6" blockId="11.[192,832,1354,1378]" box="[192,832,1354,1378]" pageId="11" pageNumber="10">
|
||
<materialsCitation id="26B7650CFFE1312C52FAFA9DEE2EFAB6" ID-GBIF-Occurrence="2580374349" box="[192,573,1354,1378]" collectionCode="TMSA" pageId="11" pageNumber="10" specimenCount="1" typeStatus="type">
|
||
<emphasis id="A4ABB343FFE1312C52FAFA9DED9AFAB6" bold="true" box="[192,393,1354,1378]" pageId="11" pageNumber="10">
|
||
<typeStatus id="4964D1F3FFE1312C52FAFA9DED17FAB5" box="[192,260,1354,1377]" pageId="11" pageNumber="10">Type</typeStatus>
|
||
material.
|
||
</emphasis>
|
||
HTm—
|
||
<collectionCode id="F0CEF794FFE1312C53D0FA9DEE2EFAB6" box="[490,573,1354,1378]" collectionName="South Africa, Gauteng, Pretoria, Transvaal Museum" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="11" pageNumber="10">TMSA</collectionCode>
|
||
</materialsCitation>
|
||
. No other specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DEC53CDAFFE1312C52FAFAADEEB4FA47" box="[192,679,1402,1427]" pageId="11" pageNumber="10" type="etymology">
|
||
<paragraph id="96606F51FFE1312C52FAFAADEEB4FA47" blockId="11.[192,679,1402,1427]" box="[192,679,1402,1427]" pageId="11" pageNumber="10">
|
||
<emphasis id="A4ABB343FFE1312C52FAFAADED48FA46" bold="true" box="[192,347,1402,1426]" pageId="11" pageNumber="10">Etymology.</emphasis>
|
||
Similar, resembling (Lat.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DEC53CDAFFE1312B52FAFA7DE839FB24" lastPageId="12" lastPageNumber="11" pageId="11" pageNumber="10" type="description">
|
||
<paragraph id="96606F51FFE1312C52FAFA7DEF64FA17" blockId="11.[192,1440,1450,1928]" box="[192,887,1450,1475]" pageId="11" pageNumber="10">
|
||
<emphasis id="A4ABB343FFE1312C52FAFA7DEE1FFA17" bold="true" box="[192,524,1450,1475]" pageId="11" pageNumber="10">
|
||
Description of
|
||
<typeStatus id="4964D1F3FFE1312C53B6FA7CEE15FA17" box="[396,518,1451,1475]" pageId="11" pageNumber="10" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Length
|
||
<quantity id="5127C2B4FFE1312C5048FA7CEEDAFA16" box="[626,713,1451,1475]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="11" pageNumber="10" unit="mm" value="14.0">14 mm</quantity>
|
||
; black, glossy.
|
||
</paragraph>
|
||
<paragraph id="96606F51FFE1312C52D2FA1CEF46F952" blockId="11.[192,1440,1450,1928]" pageId="11" pageNumber="10">
|
||
<emphasis id="A4ABB343FFE1312C52D2FA1CED22FA37" box="[232,305,1483,1507]" italics="true" pageId="11" pageNumber="10">Head.</emphasis>
|
||
Clypeus with teeth upturned, spaces between them V-shaped with that between medials more open, tips of medials and right lateral broken off. Genae with anterior tips triangular but rounded, posterolateral margins setose, posterior corners obliquely curved toward eyes, posterior margin rimmed by a carina that medially branches to laterally/anterolaterally surround eye. Surface at bases of teeth and entire genae granulorugose, with a narrow glabrous medial strip from bases of teeth through frons and vertex. Antennal peduncle dark brown, club gray.
|
||
</paragraph>
|
||
<paragraph id="96606F51FFE1312C52D2F959EDFEF8F3" blockId="11.[192,1440,1450,1928]" pageId="11" pageNumber="10">
|
||
<emphasis id="A4ABB343FFE1312C52D2F959ED7AF971" box="[232,361,1678,1701]" italics="true" pageId="11" pageNumber="10">Pronotum.</emphasis>
|
||
Strongly convex and transverse, bordered all around, with front angles obtuse, lateral margins crenulate and setose, and hind corners obliquely merging into base, which lacks medial lobe. Broad, glabrous sagittal line present from front margin to near base, reminder of surface unevenly coarsely punctate, size of punctures diminishes laterally and anteriorly; punctation nearly as sparse as in
|
||
<taxonomicName id="51DF14D2FFE1312C52DAF8D8ED6BF8F3" box="[224,376,1807,1831]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="laevifrons">
|
||
<emphasis id="A4ABB343FFE1312C52DAF8D8ED6BF8F3" box="[224,376,1807,1831]" italics="true" pageId="11" pageNumber="10">S. laevifrons</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="0EE473D4FFE1312C53B2F8D8EDF3F8F3" box="[392,480,1807,1831]" captionStart="Figures 81–87" captionStartId="35.[192,287,1819,1841]" captionTargetBox="[222,1414,278,1747]" captionTargetPageId="35" captionText="Figures 81–87. Scarabaeus (Scarabaeolus) species with two mesotibial spurs. 81) S. intricatus (Fabricius), m, 16.5 mm. 82) S. inoportunus Ferreira, m, 13.5 mm. 83) S. lucidulus (Boheman), m, 13.5 mm. 84) S. laevifrons Fairmaire, m, 11.5 mm. 85) S. reichei Waterhouse, HT, 18.5 mm. 86) S. reichei HT, detail of left elytron. 87) S. fritschi Harold, detail of left elytron (from specimen in Fig. 74)." figureDoi="http://doi.org/10.5281/zenodo.3727009" httpUri="https://zenodo.org/record/3727009/files/figure.png" pageId="11" pageNumber="10">Fig. 84</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="96606F51FFE1312C52D2F8F8EDECF89C" blockId="11.[192,1440,1450,1928]" box="[232,511,1839,1864]" pageId="11" pageNumber="10">
|
||
<emphasis id="A4ABB343FFE1312C52D2F8F8ED7FF893" box="[232,364,1839,1863]" italics="true" pageId="11" pageNumber="10">
|
||
<taxonomicName id="51DF14D2FFE1312C52D2F8F8ED7BF893" box="[232,360,1839,1863]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Corynexochida" pageId="11" pageNumber="10" phylum="Arthropoda" rank="genus">Scutellum</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Not visible.
|
||
</paragraph>
|
||
<paragraph id="96606F51FFE1312B52D2F887EF70FEC6" blockId="11.[192,1440,1450,1928]" lastBlockId="12.[192,1440,217,696]" lastPageId="12" lastPageNumber="11" pageId="11" pageNumber="10">
|
||
<emphasis id="A4ABB343FFE1312C52D2F887ED2EF8BC" box="[232,317,1872,1896]" italics="true" pageId="11" pageNumber="10">Elytra.</emphasis>
|
||
With striae indistinctly, shallowly punctate, interspaces about equal puncture diameters. Second through fourth intervals medium convex and unevenly coarsely punctate, with interspaces equaling 2 to 5 puncture diameters; first and fifth through seventh intervals flat and mostly impunctate. Humeri barely indicated and confined to sixth interval.
|
||
</paragraph>
|
||
<paragraph id="96606F51FFE6312B52D2FECDE87DFEE6" blockId="12.[192,1440,217,696]" box="[232,1134,282,306]" pageId="12" pageNumber="11">
|
||
<emphasis id="A4ABB343FFE6312B52D2FECDED75FEE6" box="[232,358,282,306]" italics="true" pageId="12" pageNumber="11">
|
||
<taxonomicName id="51DF14D2FFE6312B52D2FECDED71FEE6" box="[232,354,282,306]" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageId="12" pageNumber="11" phylum="Chordata" rank="genus">Pygidium</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Semilunar, bordered all around, finely and shallowly punctate.
|
||
</paragraph>
|
||
<paragraph id="96606F51FFE6312B52D2FEECE81EFE40" blockId="12.[192,1440,217,696]" pageId="12" pageNumber="11">
|
||
<emphasis id="A4ABB343FFE6312B52D2FEECED2CFE86" box="[232,319,315,338]" italics="true" pageId="12" pageNumber="11">Venter.</emphasis>
|
||
Metasternum largely glabrous, finely punctate only lateral of anterior process and with short rusty setae on anterolateral margins; anterior process broadly lunate, with a rounded tip. Abdominal ventrites finely punctate along anterior margins, otherwise glabrous.
|
||
</paragraph>
|
||
<paragraph id="96606F51FFE6312B52D2FE4AEE8DFDA3" blockId="12.[192,1440,217,696]" pageId="12" pageNumber="11">
|
||
<emphasis id="A4ABB343FFE6312B52D2FE4AED34FE60" box="[232,295,413,436]" italics="true" pageId="12" pageNumber="11">Legs.</emphasis>
|
||
Protibia weakly bent inward between second and third teeth, devoid of setae, with lateral margin finely serrate and medial margin finely granulate throughout length; dorsomedial carina branches toward bases of second and third teeth; ventromedial carina simple and continuous throughout length. Profemur robust, proximally thickened by oblique expansion of anterior edge, anteroventral surface glabrous, posteroventral surface coarsely punctate. Mesotibia with one spur. Both meso- and metatibiae with two weak transverse carinae, meso- and metafemora antero- and posteroventrally with rows of setose punctures, setae short and gray.
|
||
</paragraph>
|
||
<paragraph id="96606F51FFE6312B52D2FDA8EEEDFD6C" blockId="12.[192,1440,217,696]" pageId="12" pageNumber="11">
|
||
<emphasis id="A4ABB343FFE6312B52D2FDA8ED77FD43" box="[232,356,639,663]" italics="true" pageId="12" pageNumber="11">Aedeagus.</emphasis>
|
||
With paramere tips in dorsal view weakly inflated, in lateral view with a low ventral tooth and tapering into long, ventrally curved beaks.
|
||
</paragraph>
|
||
<paragraph id="96606F51FFE6312B52FAFD07EEB9FC9D" blockId="12.[192,1441,720,1264]" pageId="12" pageNumber="11">
|
||
<emphasis id="A4ABB343FFE6312B52FAFD07ED7DFD3C" bold="true" box="[192,366,720,744]" pageId="12" pageNumber="11">Comparison.</emphasis>
|
||
The habitus of this new species is reminiscent of
|
||
<taxonomicName id="51DF14D2FFE6312B51EEFD07E9B3FD3C" authority="Davis and Deschodt" authorityName="Davis and Deschodt" box="[980,1440,720,744]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="megaparvulus">
|
||
<emphasis id="A4ABB343FFE6312B51EEFD07E8B0FD3C" box="[980,1187,720,744]" italics="true" pageId="12" pageNumber="11">S. megaparvulus</emphasis>
|
||
Davis and Deschodt
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="0EE473D4FFE6312B52F3FD26ED58FCDD" box="[201,331,753,777]" captionStart="Figures 55–62" captionStartId="32.[192,287,1819,1841]" captionTargetBox="[212,1424,293,1748]" captionTargetPageId="32" captionText="Figures 55–62. Scarabaeus (Scarabaeolus) spp. 55–58) Scarabaeus (Scarabaeolus) similis sp. n., HTm, 14 mm. 55) Dorsal habitus. 56) Ventral habitus. 57) Aedeagus, dorsal. 58) Aedeagus, left lateral. 59–62) Scarabaeus (Scarabaeolus) megaparvulus Davis and Deschodt, 12 mm. 59) Dorsal habitus. 60) Ventral habitus. 61) Aedeagus, dorsal. 62) Aedeagus, left lateral." figureDoi="http://doi.org/10.5281/zenodo.3727003" httpUri="https://zenodo.org/record/3727003/files/figure.png" pageId="12" pageNumber="11">Fig. 59–60</figureCitation>
|
||
) and
|
||
<taxonomicName id="51DF14D2FFE6312B53AAFD26EEB1FCDD" authority="(Boheman)" baseAuthorityName="Boheman" box="[400,674,752,777]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Corynexochida" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="funebris">
|
||
<emphasis id="A4ABB343FFE6312B53AAFD26EE07FCDC" box="[400,532,752,776]" italics="true" pageId="12" pageNumber="11">S. funebris</emphasis>
|
||
(Boheman)
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="0EE473D4FFE6312B5089FD26EF19FCDD" box="[691,778,753,777]" captionStart="Figures 63–71" captionStartId="33.[192,287,1790,1812]" captionTargetBox="[188,1440,210,1767]" captionTargetId="figure@33.[640,1002,724,1193]" captionTargetPageId="33" captionText="Figures 63–71. Scarabaeus (Scarabaeolus) species with one mesotibial spur. 63) S. funebris (Boheman), m, 17.5 mm. 64) S. parvulus (Boheman), m, 14 mm. 65) S. clanceyi Ferreira, m, 12.7 mm. 66) S. karrooensis zur Strassen, m, 13 mm. 67) S. obsoletepunctatus Balthasar, HTf, 14.5 mm. 68) Same, ventral view. 69) S. palemo Olivier, m, 14.5 mm (Benin, brownish because teneral). 70) S. morbillosus Fabricius, ST, 13.7 mm (= S. palemo Olivier). 71) Same, frontal view showing orangish profemoral pads." figureDoi="http://doi.org/10.5281/zenodo.3727005" httpUri="https://zenodo.org/record/3727005/files/figure.png" pageId="12" pageNumber="11">Fig. 63</figureCitation>
|
||
), of which the latter has not hitherto been included in the subgenus. These two species resemble
|
||
<taxonomicName id="51DF14D2FFE6312B50F1FCC6EF2DFCFD" authority="Zidek & Pokorný, 2018" authorityName="Zidek & Pokorný" authorityYear="2018" box="[715,830,785,809]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="similis" status="sp. nov.">
|
||
<emphasis id="A4ABB343FFE6312B50F1FCC6EF2DFCFD" box="[715,830,785,809]" italics="true" pageId="12" pageNumber="11">S. similis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BF980E38FFE6312B517CFCC5EF9BFCFD" box="[838,904,786,809]" pageId="12" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
|
||
chiefly due to their dorsal sculpture, but at the same time differ in several respects.
|
||
</paragraph>
|
||
<paragraph id="96606F51FFE6312B52D2FC85E9B3FC1F" blockId="12.[192,1441,720,1264]" pageId="12" pageNumber="11">
|
||
<taxonomicName id="51DF14D2FFE6312B52D2FC85EE38FCBE" box="[232,555,850,874]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="megaparvulus">
|
||
<emphasis id="A4ABB343FFE6312B52D2FC85EE38FCBE" box="[232,555,850,874]" italics="true" pageId="12" pageNumber="11">Scarabaeus megaparvulus</emphasis>
|
||
</taxonomicName>
|
||
has a well-developed transverse frontal carina; a more densely punctate pronotum without sagittal line and with a basal medial lobe; exposed
|
||
<taxonomicName id="51DF14D2FFE6312B562AFCA4E898FC5F" box="[1040,1163,883,907]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Corynexochida" pageId="12" pageNumber="10" phylum="Arthropoda" rank="genus">scutellum</taxonomicName>
|
||
; a narrow metasternal process whose triangular tip is delimited by a transverse groove; much thinner profemur; mesotibia with two spurs (only one noted in the original description); and paramere tips in lateral view truncate.
|
||
</paragraph>
|
||
<paragraph id="96606F51FFE6312B52D2FC03EF3EFB5A" blockId="12.[192,1441,720,1264]" pageId="12" pageNumber="11">
|
||
<taxonomicName id="51DF14D2FFE6312B52D2FC03EDCCFC38" box="[232,479,980,1004]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="funebris">
|
||
<emphasis id="A4ABB343FFE6312B52D2FC03EDCCFC38" box="[232,479,980,1004]" italics="true" pageId="12" pageNumber="11">Scarabaeus funebris</emphasis>
|
||
</taxonomicName>
|
||
has the entire clypeus, genae and frons granulorugose; pronotum densely punctate, a narrow sagittal line reaching basal medial lobe, and a striola of finer setose punctures paralleling the basal margin; exposed
|
||
<taxonomicName id="51DF14D2FFE6312B5038FBC2EE6EFBF9" box="[514,637,1045,1069]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Corynexochida" pageId="12" pageNumber="10" phylum="Arthropoda" rank="genus">scutellum</taxonomicName>
|
||
; and elytral intervals much more coarsely punctate. The aedeagus of
|
||
<taxonomicName id="51DF14D2FFE6312B52E7FBE2ED72FB99" box="[221,353,1077,1101]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="funebris">
|
||
<emphasis id="A4ABB343FFE6312B52E7FBE2ED72FB99" box="[221,353,1077,1101]" italics="true" pageId="12" pageNumber="11">S. funebris</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="0EE473D4FFE6312B534BFBE2EDC7FB99" box="[369,468,1077,1101]" captionStart="Figures 88–104" captionStartId="36.[192,287,1762,1784]" captionTargetBox="[263,1414,235,1714]" captionTargetPageId="36" captionText="Figures 88–104. Scarabaeus (Scarabaeolus) aedeagi in dorsal and left lateral views. 88) S. palemo Olivier (Benin, from specimen in Fig. 69). 89) S. parvulus (Boheman). 90) S. karrooensis zur Strassen. 91) S. laevifrons Fairmaire. 92) S. reichei = S. canaliculatus (det. Ferreira). 93) S. canaliculatus Fairmaire. 94) S. fritschi Harold. 95) S. damarensis Janssens. 96) S. gilleti Janssens. 97) S. kwiluensis Janssens. 98) S. kochi Ferreira. 99) S. pabulator Péringuey. 100) S. intricatus (Fabricius). 101) S. lucidulus (Boheman). 102) S. funebris (Boheman). 103) S. clanceyi Ferreira. 104) S. inoportunus Ferreira." figureDoi="http://doi.org/10.5281/zenodo.3727011" httpUri="https://zenodo.org/record/3727011/files/figure.png" pageId="12" pageNumber="11">Fig. 102</figureCitation>
|
||
) is indistinguishable from that of
|
||
<taxonomicName id="51DF14D2FFE6312B5149FBE2EFF6FB99" authority="Zidek & Pokorný, 2018" authorityName="Zidek & Pokorný" authorityYear="2018" box="[883,997,1077,1101]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="similis" status="sp. nov.">
|
||
<emphasis id="A4ABB343FFE6312B5149FBE2EFF6FB99" box="[883,997,1077,1101]" italics="true" pageId="12" pageNumber="11">S. similis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BF980E38FFE6312B51D6FBE1E83EFB99" box="[1004,1069,1078,1101]" pageId="12" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="0EE473D4FFE6312B5607FBE2E8D3FB99" box="[1085,1216,1077,1101]" captionStart="Figures 55–62" captionStartId="32.[192,287,1819,1841]" captionTargetBox="[212,1424,293,1748]" captionTargetPageId="32" captionText="Figures 55–62. Scarabaeus (Scarabaeolus) spp. 55–58) Scarabaeus (Scarabaeolus) similis sp. n., HTm, 14 mm. 55) Dorsal habitus. 56) Ventral habitus. 57) Aedeagus, dorsal. 58) Aedeagus, left lateral. 59–62) Scarabaeus (Scarabaeolus) megaparvulus Davis and Deschodt, 12 mm. 59) Dorsal habitus. 60) Ventral habitus. 61) Aedeagus, dorsal. 62) Aedeagus, left lateral." figureDoi="http://doi.org/10.5281/zenodo.3727003" httpUri="https://zenodo.org/record/3727003/files/figure.png" pageId="12" pageNumber="11">Fig. 57–58</figureCitation>
|
||
), but it can hardly be claimed to indicate a close relationship because the same applies to the aedeagus of
|
||
<taxonomicName id="51DF14D2FFE6312B56EEFB81E9B3FBBA" authority="Ferreira" authorityName="Ferreira" box="[1236,1440,1109,1134]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Corynexochida" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="kochi">
|
||
<emphasis id="A4ABB343FFE6312B56EEFB81E927FBB9" box="[1236,1332,1109,1133]" italics="true" pageId="12" pageNumber="11">S. kochi</emphasis>
|
||
Ferreira
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="0EE473D4FFE6312B52F3FBA1ED33FB5A" box="[201,288,1142,1166]" captionStart="Figures 88–104" captionStartId="36.[192,287,1762,1784]" captionTargetBox="[263,1414,235,1714]" captionTargetPageId="36" captionText="Figures 88–104. Scarabaeus (Scarabaeolus) aedeagi in dorsal and left lateral views. 88) S. palemo Olivier (Benin, from specimen in Fig. 69). 89) S. parvulus (Boheman). 90) S. karrooensis zur Strassen. 91) S. laevifrons Fairmaire. 92) S. reichei = S. canaliculatus (det. Ferreira). 93) S. canaliculatus Fairmaire. 94) S. fritschi Harold. 95) S. damarensis Janssens. 96) S. gilleti Janssens. 97) S. kwiluensis Janssens. 98) S. kochi Ferreira. 99) S. pabulator Péringuey. 100) S. intricatus (Fabricius). 101) S. lucidulus (Boheman). 102) S. funebris (Boheman). 103) S. clanceyi Ferreira. 104) S. inoportunus Ferreira." figureDoi="http://doi.org/10.5281/zenodo.3727011" httpUri="https://zenodo.org/record/3727011/files/figure.png" pageId="12" pageNumber="11">Fig. 98</figureCitation>
|
||
), whose habitus (
|
||
<figureCitation id="0EE473D4FFE6312B53CFFBA1EE59FB5A" box="[501,586,1142,1166]" captionStart="Figures 72–80" captionStartId="34.[192,287,1821,1843]" captionTargetBox="[209,1440,219,1762]" captionTargetId="figure@34.[606,996,770,1212]" captionTargetPageId="34" captionText="Figures 72–80. Scarabaeus (Scarabaeolus) species with two mesotibial spurs. 72) S. anderseni Waterhouse, HTf, length 10.5 mm. 73) S. canaliculatus Fairmaire, m, 18 mm. 74) S. fritschi Harold, m, 19 mm. 75) S. damarensis Janssens, m, 13 mm. 76) S. gilleti Janssens, HTm, 14.5 mm. 77) S. kwiluensis Janssens, m, 15 mm. 78) S. gracai Ferreira, f, 11 mm. 79) S. kochi Ferreira, PTm, 12.5 mm. 80) S. pabulator Péringuey, m, 12 mm." figureDoi="http://doi.org/10.5281/zenodo.3727007" httpUri="https://zenodo.org/record/3727007/files/figure.png" pageId="12" pageNumber="11">Fig. 79</figureCitation>
|
||
) is quite different.
|
||
</paragraph>
|
||
<paragraph id="96606F51FFE6312B52D2FB40E839FB24" blockId="12.[192,1441,720,1264]" pageId="12" pageNumber="11">
|
||
The combination of a fully concealed
|
||
<taxonomicName id="51DF14D2FFE6312B5084FB40EF25FB7B" box="[702,822,1175,1199]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Corynexochida" pageId="12" pageNumber="10" phylum="Arthropoda" rank="genus">scutellum</taxonomicName>
|
||
and absence of the pronotal basal medial lobe is unusual. Apart from
|
||
<taxonomicName id="51DF14D2FFE6312B53FAFB60EE21FB1B" authority="Zidek & Pokorný, 2018" authorityName="Zidek & Pokorný" authorityYear="2018" box="[448,562,1207,1231]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="similis" status="sp. nov.">
|
||
<emphasis id="A4ABB343FFE6312B53FAFB60EE21FB1B" box="[448,562,1207,1231]" italics="true" pageId="12" pageNumber="11">S. similis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BF980E38FFE6312B5003FB6FEE68FB1B" box="[569,635,1208,1231]" pageId="12" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
|
||
, we have found a concealed
|
||
<taxonomicName id="51DF14D2FFE6312B51EAFB60E85BFB1B" box="[976,1096,1207,1231]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Corynexochida" pageId="12" pageNumber="10" phylum="Arthropoda" rank="genus">scutellum</taxonomicName>
|
||
only in
|
||
<taxonomicName id="51DF14D2FFE6312B5692FB60ED29FB24" authority="Balthasar" authorityName="Balthasar" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Corynexochida" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="obsoletepunctatus">
|
||
<emphasis id="A4ABB343FFE6312B5692FB60E9B3FB1B" box="[1192,1440,1207,1231]" italics="true" pageId="12" pageNumber="11">S. obsoletepunctatus</emphasis>
|
||
Balthasar
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="0EE473D4FFE6312B5370FB0FEDB2FB24" box="[330,417,1240,1264]" captionStart="Figures 63–71" captionStartId="33.[192,287,1790,1812]" captionTargetBox="[188,1440,210,1767]" captionTargetId="figure@33.[640,1002,724,1193]" captionTargetPageId="33" captionText="Figures 63–71. Scarabaeus (Scarabaeolus) species with one mesotibial spur. 63) S. funebris (Boheman), m, 17.5 mm. 64) S. parvulus (Boheman), m, 14 mm. 65) S. clanceyi Ferreira, m, 12.7 mm. 66) S. karrooensis zur Strassen, m, 13 mm. 67) S. obsoletepunctatus Balthasar, HTf, 14.5 mm. 68) Same, ventral view. 69) S. palemo Olivier, m, 14.5 mm (Benin, brownish because teneral). 70) S. morbillosus Fabricius, ST, 13.7 mm (= S. palemo Olivier). 71) Same, frontal view showing orangish profemoral pads." figureDoi="http://doi.org/10.5281/zenodo.3727005" httpUri="https://zenodo.org/record/3727005/files/figure.png" pageId="12" pageNumber="11">Fig. 67</figureCitation>
|
||
), but that species does possess the basal medial lobe.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |