treatments-xml/data/16/56/47/16564724BACB77262262B06FC0F8CFD4.xml

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<mods:title id="2BBAEAF75C407BA7860BCB01D79E81EA">Astyanax endy (Characiformes: Characidae), a new fish species from the upper Río Bermejo basin, northwestern Argentina.</mods:title>
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<mods:namePart id="7BCBDBFCDF5FCA5C32F42895765F1F21">J. M. Mirande</mods:namePart>
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<mods:namePart id="9CD9BAACC7AFB9174FF07CD98601ADD8">G. Aguilera</mods:namePart>
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<mods:namePart id="A83DC62D39A0C64661FDC37A7DE73B2D">M. D. L. M. Azpelicueta</mods:namePart>
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<treatment id="16564724BACB77262262B06FC0F8CFD4" ID-DOI="http://doi.org/10.5281/zenodo.6260743" ID-GBIF-Taxon="100122206" ID-Zenodo-Dep="6260743" LSID="urn:lsid:plazi:treatment:16564724BACB77262262B06FC0F8CFD4" httpUri="http://treatment.plazi.org/id/16564724BACB77262262B06FC0F8CFD4" lastPageNumber="66" pageNumber="59">
<subSubSection id="99B56328AE09A408F17FE393C04F629B" type="nomenclature">
<paragraph id="E721A616DCD74D2ABBA07787E952FDC1" pageNumber="59">
<taxonomicName id="D81654E03C878FC8ED4DCFA7067BBDAB" ID-CoL="J5RF" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="59" phylum="Chordata" rank="species" species="endy">Astyanax endy</taxonomicName>
, 
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</paragraph>
</subSubSection>
<subSubSection id="403CB386162B6434ACBD4AC30DC6A43A" type="description">
<paragraph id="6CEDDEAA9570BFFED87B9E4506FD5AD0" pageNumber="59">(Figs. 1-3, table 1)</paragraph>
</subSubSection>
<subSubSection id="C5295CF1282D5B34B7BFDC5789BD62E7" type="nomenclature">
<paragraph id="ADFAB8D4B83BBA7F1E0DE42FD246389E" pageNumber="59">
<materialsCitation id="CDD91B0900649D2F17B6AB1C0579723B" ID-GBIF-Occurrence="923905407" country="Argentina" stateProvince="Salta">
<typeStatus id="438FE12269CE340CF00A904A3F0BFD67">Holotype</typeStatus>
. 
<collectionCode id="B07A44BC325E71525B92D58177AC5551">CI-FML</collectionCode>
3834 male, 55.8 mm SL, 
<collectingCountry id="3BDC74D80B080101EACF2560BCADFAB3">Argentina</collectingCountry>
: 
<collectingRegion id="31273BFA26A421A600AEB7BA6223DBA3">Salta</collectingRegion>
, 
<normalizedToken id="088174BC1C38EB02018C715375550DE9" originalValue="Orán">Oran</normalizedToken>
, 
<normalizedToken id="E7282CBA27132C989181B563630A1BFB" originalValue="Río">Rio</normalizedToken>
Bermejo basin, tributary of 
<normalizedToken id="2B93D8A6E898DEAC09FA3ACDB75A67D7" originalValue="Río">Rio</normalizedToken>
Bermejo in its intersection with Ruta Provincial 19, near Estancia Santa Rosa (
<geoCoordinate id="57E260A2DCD061D220B990F251D78809">64º24’13’’W</geoCoordinate>
- 
<geoCoordinate id="FCFF5FC328844834AF4E9E60E43750AD">22º43’18’’S</geoCoordinate>
; approximately 440 meters above sea level). Coll: M. Mirande, G. Aguilera and G. Padilla, August 2003.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="B58C3C53BD816F61A76A3301C90BA3D3" type="nomenclature">
<paragraph id="91726DF76172D7950FCC85A3E7E45161" pageNumber="60">
<materialsCitation id="091B707B82459D6CBB3F02B5D3C19396" ID-GBIF-Occurrence="923905405" country="Argentina" stateProvince="Salta">
<typeStatus id="8EDBE305422E6AF4B2078F381979C002">Paratypes</typeStatus>
: 
<collectionCode id="6DB57699B1C56D82B3C17E90FBA3C7FC">CI-FML</collectionCode>
3835, 14 ex., 48.6-56.3 mm SL, collected with the holotype.
</materialsCitation>
<materialsCitation id="474D3202818E952C4FD00540C862F335" ID-GBIF-Occurrence="923905408" country="Argentina" stateProvince="Salta">
<collectionCode id="ACF7A59CBB4365A8ED11D6A86C2D4750">CI-FML</collectionCode>
3836, 13 ex., 39.3-62.0 mm SL, 
<collectingCountry id="0CA5E59D8172F2FF9D69F24CF2248240">Argentina</collectingCountry>
: 
<collectingRegion id="65D413DFCBE7C1FDDDAC234E855694F4">Salta</collectingRegion>
, 
<normalizedToken id="FE8F7E6ACFC607E9D455E47B5C353D2D" originalValue="Orán">Oran</normalizedToken>
, El Oculto, 
<normalizedToken id="1204C7DF4AF2B7F5D4060B1EB6C5421A" originalValue="Río">Rio</normalizedToken>
Bermejo basin, 
<normalizedToken id="6399E06F84C3C87528CFC558F41D4896" originalValue="Río">Rio</normalizedToken>
Anta Muerta, tributary of 
<normalizedToken id="1E14A6DA6C7B76616E3D3A9072FE4A36" originalValue="Río">Rio</normalizedToken>
Blanco. Coll: M. Mirande &amp; G. Aguilera, May 2002.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="2F675624C714FC4FDC58FD0F52030EB9" type="materials_examined">
<paragraph id="36880364C27876477E08EAB2DF6A183C" pageNumber="60">
Additional Material (not designated as paratypes): 
<materialsCitation id="A4E252EE665AF507BB3D6D6A28335240" ID-GBIF-Occurrence="923905410" country="Argentina" stateProvince="Salta">
<collectionCode id="7DEA46ACBDB63A10078F510CB0C60A6C">CI-FML</collectionCode>
3262, 23 ex., 29.4-50.7 mm, 
<collectingCountry id="4883FE67624BFB3B31CD6DE7F59906AA">Argentina</collectingCountry>
, 
<collectingRegion id="746D3B3B29CF7D95BD10C4B7B071F129">Salta</collectingRegion>
, La 
<normalizedToken id="5465D10F24DFDA48B1EBFBFC33321D4D" originalValue="Bambú">Bambu</normalizedToken>
, 
<normalizedToken id="2352D03EDE4EB3BB83BADAB035082B40" originalValue="Río">Rio</normalizedToken>
Bermejo basin, 
<normalizedToken id="07C4C3B7FF8DA175A4C06BB11F14A61C" originalValue="Río">Rio</normalizedToken>
Anta Muerta.
</materialsCitation>
<materialsCitation id="F91D221C25C1ED546DD00AB6989287DD" ID-GBIF-Occurrence="923905406" country="Argentina" stateProvince="Salta">
<collectionCode id="40653D0449735298F17E8DBC05514FC8">CI-FML</collectionCode>
3838, 4 ex., 24.2-44.6 mm, 
<collectingCountry id="15C2177BEB6CD6D0A2E8B0BD1288CC82">Argentina</collectingCountry>
, 
<collectingRegion id="882A9E2E4B8AD279DB247F456D158004">Salta</collectingRegion>
, 
<normalizedToken id="6F5F19108D47BBAEE55F52F7560F9D2E" originalValue="Orán">Oran</normalizedToken>
, Estancia Anta Muerta, 
<normalizedToken id="3618BD7ACAC38E9485F30EF259DDA34B" originalValue="Río">Rio</normalizedToken>
Bermejo basin, 
<normalizedToken id="D91CFFF2EAAA147BCA00E9CE58B37736" originalValue="Río">Rio</normalizedToken>
Pescado.
</materialsCitation>
<materialsCitation id="B3C26953558BF44AECA887C15191AF14" ID-GBIF-Occurrence="923905409" country="Argentina" stateProvince="Salta">
<collectionCode id="70CEDD7B15D6F56B487429FF4CD28C3E">CI-FML</collectionCode>
3839, 2 ex., 52.8-63.2 mm, 
<collectingCountry id="CCB0E25F0C403EB12EF05CA73C0EBCA1">Argentina</collectingCountry>
, 
<collectingRegion id="33FB9866C8CDDDEB6467A533985F4F54">Salta</collectingRegion>
, 
<normalizedToken id="7077A1D5AEC2E0B3E69701960DBACD28" originalValue="Orán">Oran</normalizedToken>
, 
<normalizedToken id="A88B76DCBFBCEC01A50BE842D439CA77" originalValue="Río">Rio</normalizedToken>
Bermejo basin, 
<normalizedToken id="34373578C5FCB3C152E79CA71843046E" originalValue="Río">Rio</normalizedToken>
Santa 
<normalizedToken id="70D758908C6C9CA3D823DD478DE7133B" originalValue="María">Maria</normalizedToken>
.
</materialsCitation>
<materialsCitation id="71C64D9588F430D2708C61D960EAC6F4" ID-GBIF-Occurrence="923905413" country="Argentina" stateProvince="Salta">
<collectionCode id="42A0B893700852BAE3E0B2E63E18F0E3">CI-FML</collectionCode>
(uncatalogued) 2 ex. (C&amp;S), 48.6-49.6 mm, same data as holotype.
</materialsCitation>
<materialsCitation id="7AA697E0A5B5E41F27AE76715C531ECA" ID-GBIF-Occurrence="923905412" country="Argentina" stateProvince="Salta">
<collectionCode id="FE5A8F2D7360ABDA8722561DBBC55A99">CI-FML</collectionCode>
(uncatalogued) 2 ex. (C&amp;S), 46.3-54.7 mm, 
<collectingCountry id="2D0549E2AF42A527C53D4F0F56F761E9">Argentina</collectingCountry>
, 
<collectingRegion id="339D18FA6B4330FED5ADB496F53B167C">Salta</collectingRegion>
, 
<normalizedToken id="34FC33ABB64DCA140CA11609D7E80A08" originalValue="Orán">Oran</normalizedToken>
, El Oculto, 
<normalizedToken id="02E89A76D10CDB49A5F169D886F5653A" originalValue="Río">Rio</normalizedToken>
Anta Muerta.
</materialsCitation>
<materialsCitation id="BA07358D3273C51EA0EE25FDFBB744FD" ID-GBIF-Occurrence="923905411" country="Argentina" stateProvince="Salta">
<collectionCode id="2F07F64B371EBE9B2CEAC1A5D871DEFD">CI-FML</collectionCode>
3848, 1 ex (C&amp;S), 43.6 mm, 
<collectingCountry id="A059CA8B2A8E367562784B432A28005D">Argentina</collectingCountry>
, 
<collectingRegion id="1AE1991D199133F0DBEA57220640942C">Salta</collectingRegion>
, 
<normalizedToken id="17D3BCB1CB243EEA5651991A9A673FCE" originalValue="Orán">Oran</normalizedToken>
, El Oculto, Arroyo El Oculto.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="10EF178118FA0E66F5F7C61BEB82E0DF" type="diagnosis">
<paragraph id="1F399D2510BFA1E9CFA258512C3BF24C" pageNumber="60">
Diagnosis: 
<taxonomicName id="02FE740F46594DF669647F856282F187" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="60" phylum="Chordata" rank="species" species="endy">Astyanax endy</taxonomicName>
is distinguished by the combination of the following characters: a markedly convex snout profile; deep body (36.6-42.2 % SL); relatively large orbital diameter (36.0-40.9 % HL); relatively short interorbital distance (31.9-37.6 % HL); deep caudal peduncle (11.7-14.1 % SL); long caudal peduncle (12.6-15.9 % SL), short upper-jaw length (34.4-40.2 % HL); maxilla scarcely reaching anterior margin of eye; maxilla meeting mouth commissure at an oblique angle; 35-37 perforated scales in lateral line; 7/6 transverse scales above and below lateral line on body; 17-24 branched anal-fin rays; tricuspidate or pentacuspidate teeth on outer premaxillary row; 1-2 tricuspidate or pentacuspidate maxillary teeth; 4 gradually decreasing dentary teeth, followed by 1 intermediate and 3-6 small tricuspidate or conical teeth; and 19-21 gill-rakers in first branchial arch.
</paragraph>
<paragraph id="4836EE61F053FBA362B917E50910AB81" pageNumber="61">
A detailed comparison of 
<taxonomicName id="7C969ACE9EC28C8040E1EFF73EB5D4C3" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="61" phylum="Chordata" rank="species" species="endy">Astyanax endy</taxonomicName>
with the other species of 
<taxonomicName id="586B8FDECD453B17427BB16CAD330EA1" LSID-ZBK="urn:lsid:zoobank.org:act:4DB39814-5BBA-4D7B-8C5D-F830AB26824D" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax Baird &amp; Girard 1854:26" order="Characiformes" pageNumber="61" phylum="Chordata" rank="genus">Astyanax</taxonomicName>
appears in the “Discussion” section later in this paper.
</paragraph>
</subSubSection>
<subSubSection id="133B529A18F2691B9368EDE31094EA9C" type="description">
<paragraph id="5D24F996235A8C3479EA107E8808BCDA" lastPageNumber="62" pageNumber="61">Description: Morphometric data for the holotype and 27 paratypes are presented in Table 1. Body rather deep, the maximum depth anterior to dorsal-fin origin. Dorsal profile of body moderate to strongly convex in snout region, straight to supraoccipital tip, convex from this point to dorsal-fin origin; slanted ventrally from dorsal-fin origin to caudal peduncle; gently concave along caudal peduncle to caudal-fin base. Ventral profile of body convex from tip of lower jaw to pelvic-fin origin, straight between pelvic and analfin origins, posterodorsally slanted from anal-fin origin to caudal peduncle, and slightly convex along caudal peduncle. Ventral portion of body between bases of pectoral and pelvic fins transversally rounded; ventral portion of body between origins of pelvic and anal fins more compressed. Dorsal-fin origin usually situated nearer base of caudal-fin rays than snout tip. Pelvic-fin origin located anteriorly to vertical through dorsal-fin origin. Anal-fin origin located posteriorly to vertical through base of last dorsal-fin ray. Tip of pectoral fin reaching pelvic-fin origin, and pelvic fin reaching anal-fin origin in larger mature males.</paragraph>
<paragraph id="B6056D646B812A6F5613356E19474ED4" pageNumber="63">Mouth terminal, placed at level of lower third of eye. Premaxilla bearing two series of teeth; a developed ascending process, with rather broad base and acute tip. Outer premaxillary tooth row with 3(9) or 4(19*) tricuspidate or pentacuspidate teeth. Inner premaxillary row with 5 teeth gently concave anteriorly; slender symphysial tooth, with 4 cusps, the second and third with 5-7 cusps; and the remaining teeth with 3-5 cusps (Fig. 2). Ascending maxillary process well developed; laminar portion of maxilla meeting mouth commissure at an oblique angle, bearing 1(23*), or 2(5) tricuspidate or pentacuspidate teeth. Dentary with 9-12 teeth: four anterior teeth decreasing gradually in size, followed by one tooth of intermediate size, and a series of 4-7 smaller conical or tricuspidate teeth (Fig. 2); two anterior teeth hexacuspidate or heptacuspidate, the third and fourth pentacuspidate to heptacuspidate, the fifth tricuspidate to pentacuspidate.</paragraph>
<paragraph id="1B6AE766A5E8175A8C6E28550F77C313" pageNumber="63">Eye of medium size. Third infraorbital reaching close to horizontal arm of preopercle latero-sensory canal, although not contacting it.</paragraph>
<paragraph id="4635100D5C9162B6B4F10A921E943B26" pageNumber="63">Dorsal fin rays iii, 9; first one readily visible in C&amp;S specimens; last unbranched and first branched rays longest. Anal fin rays iv -v, 17(1), 19(1), 20(1), 21(6*), 22(6), 23(12), or 24(1); females with last unbranched and first 5-6 branched rays forming a small lobe. Anal fin in males with straight distal margin. Males with hooks on last unbranched analfin ray and posterior branch of anterior 6-12 branched anal-fin rays. Principal caudal-fin rays i,17,i; caudal lobes of approximately same size. Pectoral fin rays i,11(2), 12(15), or 13(11*). Each pelvic fin with splint and i,7 rays; males with medially directed hooks well developed in medial branches of first to fifth or sixth branched pelvic-fin rays.</paragraph>
<paragraph id="F1165D70E00AA9DCA72E6272746010AD" pageNumber="63">Scales cycloid, with radii only on posterior field and not meeting each other anteriorly; no circuli on posterior field. Lateral-line with 35(11*), 36(13) or 37(4) perforated scales; 6(2) or 7(26*) transverse scales from dorsal-fin origin to lateral line, and 5(2), 6(24), or 7(2*) scales from lateral line to pelvic-fin origin; 15(3), 16(17*) or 17(8) scales around caudal peduncle; one row with 9-14 scales covering base of anterior seventh to fourteenth anal-fin ray; scales covering only base of caudal-fin rays.</paragraph>
<paragraph id="E0C09351D27FBCBD3CD394401FD3B28E" pageNumber="63">Lateral edge of first branchial arch with 19-21 gill rakers, placed as follows: 7-8 on epibranchial, 1 on cartilage, 10 on ceratobranchial, and 2 on hypobranchial; medial-posterior edge of epibranchial with a second row of 4-5 gill rakers. One row of 3-7 hooks on anterior edge of first ceratobranchial gill rakers; occasionally also a 1-3 hooks on posterior edge. Total vertebrae 33-35 (16-17 precaudal vertebrae and 17-18 caudal vertebrae). Pairs of ribs 11-12. Caudal fin with 9-10 dorsal and 7-9 ventral procurrent rays.</paragraph>
<paragraph id="A6E3857ED5CEC784E35E063E7810774B" pageNumber="63">Color in life: Body silvery, especially over lateral stripe and abdominal area; darker on dorsal part of body. Dark humeral spot vertically elongated. Caudal spot hardly visible, becoming evident over caudal fin to end of its medial rays. Orange to intense red on dorsal and anal fins; caudal lobes yellowish proximally and distally, with an intense red area in middle of its length. Pupil with a reddish area on upper margin and often on lower one.</paragraph>
<paragraph id="58EB7D9F259B71E8562370806BDEF582" pageNumber="63">Color in alcohol-preserved specimens: Uniformly yellowish, darker dorsally.</paragraph>
<paragraph id="B380E561266EA512199019ABE2ACB38B" pageNumber="64">First black humeral spot vertically elongated; second one faint although always evident. Lateral stripe very faint anteriorly, becoming more evident posteriorly. Caudal spot well developed, black, triangular or irregular in shape, extending to tip of middle caudal-fin rays. Dorsal, anal, pelvic and adipose fins hyaline; caudal-fin lobes hyaline. Sometimes, pupil bearing a vertical brownish stripe.</paragraph>
</subSubSection>
<subSubSection id="63D2160532D2E93490B4019798073F3C" type="distribution">
<paragraph id="6F9F6E582F798BBFC60C3AD71AAAFED0" pageNumber="64">
Distribution: Upper 
<normalizedToken id="F504100AAF64FF394094F424D8034BF7" originalValue="Río">Rio</normalizedToken>
Bermejo basin, Provincia de Salta, Argentina (Fig. 3).
</paragraph>
</subSubSection>
<subSubSection id="BB0DE56DB10D25C07E1BF2AC5D41AA01" type="biology_ecology">
<paragraph id="4C0ED0953D987729EBDC7E4636E52836" pageNumber="64">Habitat notes: In a series of collections made by the authors during different seasons over a three-year period, this species was very abundant in a broad range of environments, from rather fast-flowing streams to deep pools.</paragraph>
</subSubSection>
<subSubSection id="92BD24DB730633C8765C008617D9414A" type="etymology">
<paragraph id="3310CBCBC26B14D80988D7BD3108C8FF" pageNumber="64">Etymology: The specific epithet endy is a Chiriguano word meaning flame, in allusion to the color of the caudal-fin in life. A noun in apposition.</paragraph>
</subSubSection>
<subSubSection id="17E0B66C6B2D2B899A191ED2E27EB8DE" type="discussion">
<paragraph id="EE4F7BF57C41DFB691406B0B2F06ED23" pageNumber="65">Discussion</paragraph>
<paragraph id="87F346DEE4703F36B391C0B09B5322BC" pageNumber="65">
<taxonomicName id="5CC6715F5A9B0ED9DC333F9861A7C77E" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="endy">Astyanax endy</taxonomicName>
resembles 
<taxonomicName id="A5D57246270D4E052555489623026D6D" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="eigenmanniorum">A. eigenmanniorum</taxonomicName>
in number of lateral line scales and fin-rays, but is distinguished from that species by having a higher number of transverse scales (7/6 vs. 6/5) on the body, the maxilla scarcely reaching (vs. surpassing) the anterior margin of the eye, a lower number of gill-rakers in the first branchial arch (19-21 vs. 23-26), lesser upper-jaw length (34.4-40.2 vs. 41.0-46.3% HL), deeper body (36.6-42.2 vs. 32.1-37.9% SL), and greater postorbital distance (41.0-48.5 vs. 33.0-41.7 % HL).
</paragraph>
<paragraph id="C9B9FB3C16C51DF956E51CD07ED4F238" pageNumber="65">
The new species is distinguished from 
<taxonomicName id="13D3696C21CC3DB3B48F37F13A31ABCA" LSID-ZBK="urn:lsid:zoobank.org:act:E5863476-57AD-4DFB-9869-C74D9004AFD1" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax leonidas Azpelicueta, Casciotta &amp; Almirón 2002:245" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="leonidas">
Astyanax leonidas Azpelicueta, Casciotta &amp; 
<normalizedToken id="EF46E9878D8C8B7783FDE649DF8C2B09" originalValue="Almirón">Almiron</normalizedToken>
</taxonomicName>
; 
<taxonomicName id="1EBFF036942A3D03E45BD7C9FFF265B8" LSID-ZBK="urn:lsid:zoobank.org:act:AE44CFDC-63B3-46DF-A875-9C84D52198F0" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax ojiara Azpelicueta &amp; Garcia 2000:246" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="ojiara">
A. ojiara Azpelicueta &amp; 
<normalizedToken id="A9F8E7E5D9EABF5AD01FB0E67ED05879" originalValue="García">Garcia</normalizedToken>
</taxonomicName>
; 
<taxonomicName id="7A033501398F6F416218DA60913471F3" LSID-ZBK="urn:lsid:zoobank.org:act:290BA648-48A7-485B-AFDF-9A33A7AE0F21" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax troya Azpelicueta, Casciotta &amp; Almirón 2002:249" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="troya">
A. troya Azpelicueta, Casciotta &amp; 
<normalizedToken id="DD59F187B0C09B54B6BA10F259995823" originalValue="Almirón">Almiron</normalizedToken>
</taxonomicName>
; 
<taxonomicName id="6CAC010A757D80C602B775D84936D8C7" LSID-ZBK="urn:lsid:zoobank.org:act:96A907F5-8583-4DFB-8209-1B4852986965" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax pynandi Casciotta, Almirón, Bechara, Roux &amp; Ruiz Diaz 2003:808" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="pynandi">
A. pynandi Casciotta, 
<normalizedToken id="09815E0C7D8233C4B7D72149AFCC15F7" originalValue="Almirón">Almiron</normalizedToken>
&amp; Azpelicueta
</taxonomicName>
; 
<taxonomicName id="EF84C1CB00C6A478D11177CF22FD3FD8" LSID-ZBK="urn:lsid:zoobank.org:act:277C9FEE-EFEC-4675-8415-DC69F6205FE7" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax chico Casciotta &amp; Almirón 2004:12" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="chico">
A. chico Casciotta &amp; 
<normalizedToken id="11AE0763BFEA31B79EF0C37AA6266C7B" originalValue="Almirón">Almiron</normalizedToken>
</taxonomicName>
; 
<taxonomicName id="3C1FEB6117BA7A4BAD35D208BBBD5EE0" LSID-ZBK="urn:lsid:zoobank.org:act:15E68D20-6FA1-43D1-A098-7843F661D1CD" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax hermosus Miquelarena, Protogino &amp; López 2005:14" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="hermosus">
A. hermosus Miquelarena, Protogino &amp; 
<normalizedToken id="B3CF08AF2AAB5DEA9AC098E3C9324E11" originalValue="López">Lopez</normalizedToken>
</taxonomicName>
; and 
<taxonomicName id="4CF033EBA95C5EC6FCBD20087654FD75" LSID-ZBK="urn:lsid:zoobank.org:act:E47C3BA8-9E72-4C96-B1B3-9167C828A693" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax tumbayaensis Miquelarena &amp; Menni 2005" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="tumbayaensis">A. tumbayaensis Miquelarena &amp; Menni</taxonomicName>
by the absence of hooks on the dorsal, caudal and/or pectoral fins of males. 
<taxonomicName id="311E96230323D4CDA61534D7F13BB732" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="endy">Astyanax endy</taxonomicName>
differs from 
<taxonomicName id="CE5627EE0793A383EC20A5BB3FB6F9C7" LSID-ZBK="urn:lsid:zoobank.org:act:AE44CFDC-63B3-46DF-A875-9C84D52198F0" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax ojiara Azpelicueta &amp; Garcia 2000:246" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="ojiara">A. ojiara</taxonomicName>
, 
<taxonomicName id="A7A96DEE420F6385C1A207C2073DE9B7" LSID-ZBK="urn:lsid:zoobank.org:act:290BA648-48A7-485B-AFDF-9A33A7AE0F21" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax troya Azpelicueta, Casciotta &amp; Almirón 2002:249" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="troya">A. troya</taxonomicName>
, 
<taxonomicName id="00B597B5985B30B0C10247137464D866" LSID-ZBK="urn:lsid:zoobank.org:act:96A907F5-8583-4DFB-8209-1B4852986965" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax pynandi Casciotta, Almirón, Bechara, Roux &amp; Ruiz Diaz 2003:808" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="pynandi">A. pynandi</taxonomicName>
, 
<taxonomicName id="9CEA6211F5368DEB7ED083472AD77115" LSID-ZBK="urn:lsid:zoobank.org:act:277C9FEE-EFEC-4675-8415-DC69F6205FE7" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax chico Casciotta &amp; Almirón 2004:12" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="chico">A. chico</taxonomicName>
, and 
<taxonomicName id="0B855DB507F60F94BD86F8AC66D85D00" LSID-ZBK="urn:lsid:zoobank.org:act:0B1C3AED-1739-41BA-830F-3CAACF68E2C3" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax ita Almirón, Azpelicueta &amp; Casciotta 2002:4" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="ita">
A. ita 
<normalizedToken id="39597B4C6225BD935B92F764B93E6FA4" originalValue="Almirón">Almiron</normalizedToken>
, Azpelicueta &amp; Casciotta
</taxonomicName>
in having 4 gradually decreasing dentary teeth, 1 intermediate, and 4-7 posterior conical teeth of approximately equal size (vs. 7-10 gradually decreasing dentary teeth).
</paragraph>
<paragraph id="415FFDED7C245E0F14A3296F06FDAC25" pageNumber="65">
<taxonomicName id="B4FC34F869D719CA6F37A893B52EB4DE" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="endy">Astyanax endy</taxonomicName>
shares a similar shape of the 5 anterior dentary teeth with 
<taxonomicName id="D5189E4C484D2B1D6F73A5302A9D760B" LSID-ZBK="urn:lsid:zoobank.org:act:E5863476-57AD-4DFB-9869-C74D9004AFD1" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax leonidas Azpelicueta, Casciotta &amp; Almirón 2002:245" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="leonidas">A. leonidas</taxonomicName>
, 
<taxonomicName id="74F751EEAD6F889BE8399EEE0FDB1CA2" LSID-ZBK="urn:lsid:zoobank.org:act:15E68D20-6FA1-43D1-A098-7843F661D1CD" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax hermosus Miquelarena, Protogino &amp; López 2005:14" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="hermosus">A. hermosus</taxonomicName>
, and 
<taxonomicName id="7FCDD80CA2B8BC5683867056F6D4AA4B" LSID-ZBK="urn:lsid:zoobank.org:act:4ED3ADE0-ACC8-43D3-B38F-81A3CA580CCC" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax pampa Casciotta, Almirón &amp; Azpelicueta 2005:401" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="pampa">
A. pampa Casciotta, 
<normalizedToken id="5A24CA66F3E658CCDB12AE4E52E6B777" originalValue="Almirón">Almiron</normalizedToken>
&amp; Azpelicueta
</taxonomicName>
. It is further distinguished from 
<taxonomicName id="97F7E21CDABDBC723A341C76EA8334A8" LSID-ZBK="urn:lsid:zoobank.org:act:E5863476-57AD-4DFB-9869-C74D9004AFD1" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax leonidas Azpelicueta, Casciotta &amp; Almirón 2002:245" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="leonidas">A. leonidas</taxonomicName>
by the dorsal profile of the body (nearly straight vs. very convex), greater number of small posterior dentary teeth (4-7 vs. 3-4), greater number of anal-fin rays (usually 21-23 vs. 17-21), deeper body (36.6-42.2 vs. 30.2-35.3 % SL), greater interorbital distance (31.9-37.6 vs. 26.9-30.4 % HL), and greater number of gill rakers on first branchial arch (19-21 vs. 17-19). 
<taxonomicName id="C09F52DC50BE14E7E20C376F2B4D3BCC" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="endy">Astyanax endy</taxonomicName>
is also distinguished from 
<taxonomicName id="197A947B8FF3D2CAE8EF15FD09C042CD" LSID-ZBK="urn:lsid:zoobank.org:act:15E68D20-6FA1-43D1-A098-7843F661D1CD" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax hermosus Miquelarena, Protogino &amp; López 2005:14" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="hermosus">A. hermosus</taxonomicName>
by shape of the humeral spot (vertically elongated vs. Y-shaped), longer caudal peduncle (12.6-15.9 vs. 6.8-10.4 % SL), greater eye diameter (36.0-40.9 vs. 29.7-34.7 % HL), and shorter maxillary length (22.7-28.0 vs. 30.8-34.0 % HL). The new species is also distinguished from 
<taxonomicName id="B05CFBED8BB2FF360E172214555B4039" LSID-ZBK="urn:lsid:zoobank.org:act:4ED3ADE0-ACC8-43D3-B38F-81A3CA580CCC" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax pampa Casciotta, Almirón &amp; Azpelicueta 2005:401" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="pampa">A. pampa</taxonomicName>
by the greater number of branched anal-fin rays (usually 21-23 vs. 17-20), greater number of transverse scales (7/6 vs. 6/5), and longer caudal peduncle (12.6-15.9 vs. 10.5-12.7 % SL).
</paragraph>
<paragraph id="1EBEC49F6A6823DC1A966AC9AA4B9314" lastPageNumber="66" pageNumber="65">
<taxonomicName id="BEB76B3E496CB6C9A894902369A0BEC7" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="endy">Astyanax endy</taxonomicName>
is distinguished from 
<taxonomicName id="4AB87AD6D3CE3ACA69C589D360FE4C41" LSID-ZBK="urn:lsid:zoobank.org:act:BEC3DC67-E87A-4076-8B17-F24838734DB3" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax latens Mirande, Aguilera &amp; Azpelicueta 2004:215" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="latens">A. latens</taxonomicName>
, 
<taxonomicName id="21062D5E75338576462EC9CE0ADC612A" LSID-ZBK="urn:lsid:zoobank.org:act:1CC12BAD-F757-476E-A1E1-59006410ADDE" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax tupi Azpelicueta, Mirande, Almirón &amp; Casciotta 2003:3" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="tupi">
A. tupi Azpelicueta, Mirande, 
<normalizedToken id="A40380A431AFDBDDEC277BCC903DDE4A" originalValue="Almirón">Almiron</normalizedToken>
&amp; Casciotta
</taxonomicName>
; and 
<taxonomicName id="702DA0A26BA1D6DAAAA7E2AEF4F61609" LSID-ZBK="urn:lsid:zoobank.org:act:0D3D0B86-8534-4F8A-AB1D-E25D183E1968" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax saguazu Casciotta, Almirón &amp; Azpelicueta 2003:330" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="saguazu">
A. saguazu Casciotta, 
<normalizedToken id="60EFC82C209D4009AECC14C1176E6C18" originalValue="Almirón">Almiron</normalizedToken>
&amp; Azpelicueta
</taxonomicName>
by a lower number of branched anal-fin rays (17-24 vs. 24-29 in 
<taxonomicName id="9D2DF90E6650585FFDAE97EC6CEC6F3E" LSID-ZBK="urn:lsid:zoobank.org:act:BEC3DC67-E87A-4076-8B17-F24838734DB3" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax latens Mirande, Aguilera &amp; Azpelicueta 2004:215" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="latens">A. latens</taxonomicName>
, 24-27 in 
<taxonomicName id="24952C8A306A396BC3749EBDA3643E46" LSID-ZBK="urn:lsid:zoobank.org:act:1CC12BAD-F757-476E-A1E1-59006410ADDE" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax tupi Azpelicueta, Mirande, Almirón &amp; Casciotta 2003:3" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="tupi">A. tupi</taxonomicName>
, and 25-29 in 
<taxonomicName id="96B8DD34A8914238D0ADE75E91C64916" LSID-ZBK="urn:lsid:zoobank.org:act:0D3D0B86-8534-4F8A-AB1D-E25D183E1968" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax saguazu Casciotta, Almirón &amp; Azpelicueta 2003:330" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="saguazu">A. saguazu</taxonomicName>
). In addition, it is distinguished from 
<taxonomicName id="2CE40B3CA152C0D05EEA7D6B1FD6BEE5" LSID-ZBK="urn:lsid:zoobank.org:act:BEC3DC67-E87A-4076-8B17-F24838734DB3" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax latens Mirande, Aguilera &amp; Azpelicueta 2004:215" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="latens">A. latens</taxonomicName>
and 
<taxonomicName id="1FC21C15ED1A496A759FDFAC0E853A98" LSID-ZBK="urn:lsid:zoobank.org:act:1CC12BAD-F757-476E-A1E1-59006410ADDE" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax tupi Azpelicueta, Mirande, Almirón &amp; Casciotta 2003:3" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="tupi">A. tupi</taxonomicName>
by a lower number of gill rakers on the first branchial arch (19-21 in 
<taxonomicName id="F1B93682A2326C33884C4AB6C9CE7E3F" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="endy">A. endy</taxonomicName>
vs. 26-27 in 
<taxonomicName id="5715FC0694BCBAC726DF296EAD194EC0" LSID-ZBK="urn:lsid:zoobank.org:act:BEC3DC67-E87A-4076-8B17-F24838734DB3" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax latens Mirande, Aguilera &amp; Azpelicueta 2004:215" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="latens">A. latens</taxonomicName>
and 24-25 in 
<taxonomicName id="B4EFC0BB8FB94E41D965EC6E631FC097" LSID-ZBK="urn:lsid:zoobank.org:act:1CC12BAD-F757-476E-A1E1-59006410ADDE" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax tupi Azpelicueta, Mirande, Almirón &amp; Casciotta 2003:3" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="tupi">A. tupi</taxonomicName>
). The new species is distinguished from 
<taxonomicName id="6FE0A28D16103EF67FFAC9524A5286C8" LSID-ZBK="urn:lsid:zoobank.org:act:8BA46343-D322-47F5-BDDA-156C7CC75C6B" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax paris Azpelicueta, Almirón &amp; Casciotta 2002:1052" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="paris">
A. paris Azpelicueta, 
<normalizedToken id="837020D0B499BED9CFFC22E806D014EC" originalValue="Almirón">Almiron</normalizedToken>
&amp; Casciotta
</taxonomicName>
by the lower number of maxillary teeth (1-2 vs. 3-4). 
<taxonomicName id="1B4C15492E2EC772C7859B3FAACFA566" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="endy">Astyanax endy</taxonomicName>
lacks longitudinal stripes between the scale rows, which are found in 
<taxonomicName id="27452F8D0ABB8C90302510A21E9674BB" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="lineatus">A. lineatus</taxonomicName>
; and also lacks the horizontally oval first humeral spot found in both 
<taxonomicName id="230B53D0525C248B89E70671B0F4C30E" LSID-ZBK="urn:lsid:zoobank.org:act:F06B3814-F57B-4FE4-9FA9-A74ED072124F" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax bimaculatus asuncionensis Géry 1972:3" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="asuncionensis">A. asuncionensis</taxonomicName>
and 
<taxonomicName id="8AAE7E0A7D701735F7943AE1F9406CC1" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="abramis">A. abramis</taxonomicName>
. 
<taxonomicName id="B4EB0A92E2C0C46B05366AA073D92B37" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="endy">Astyanax endy</taxonomicName>
can be further distinguished from 
<taxonomicName id="0C30850A2CA0D34C61EF8358822CCEBD" LSID-ZBK="urn:lsid:zoobank.org:act:277C9FEE-EFEC-4675-8415-DC69F6205FE7" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax chico Casciotta &amp; Almirón 2004:12" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="chico">A. chico</taxonomicName>
by a convex (vs. straight) snout profile; an oblique (vs. nearly right) angle between maxilla and mouth commissure; cusp bases from inner premaxillary row teeth following an arch (vs. nearly aligned) in lingual view; and a longer caudal peduncle (12.6-15.9 vs. 10.1-12.0 % SL). The new species can be further distinguished from 
<taxonomicName id="DF6C5EBC9E141B14E5FFC621B73F0298" LSID-ZBK="urn:lsid:zoobank.org:act:E47C3BA8-9E72-4C96-B1B3-9167C828A693" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax tumbayaensis Miquelarena &amp; Menni 2005" order="Characiformes" pageNumber="65" phylum="Chordata" rank="species" species="tumbayaensis">A. tumbayaensis</taxonomicName>
by the absence of a reticulated pigmentation pattern; the absence of densely concentrated superficial chromatophores on the third infraorbital, opercle, premaxilla, maxilla and dentary; a greater orbital diameter (36.0-40.9 vs. 28.9- 35.0 % HL); a shorter interorbital distance (31.9-37.6 vs. 38.8-44.1 % HL); and a different distribution of anal-fin hooks (on anterior 7-12 rays vs. on anterior 16-19 rays).
</paragraph>
<paragraph id="43A52D5A3101B6977E219AC899BA6C32" pageNumber="66">
In the upper 
<normalizedToken id="BF4B5319D6573367D51D1508447F366F" originalValue="Río">Rio</normalizedToken>
Bermejo basin, the following species occur together with 
<taxonomicName id="B853313A61480D96D4AA6C9F8EB95A28" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="66" phylum="Chordata" rank="species" species="endy">Astyanax endy</taxonomicName>
: 
<taxonomicName id="DE1845F3E67DE923568AE949A47D34C1" LSID-ZBK="urn:lsid:zoobank.org:act:BEC3DC67-E87A-4076-8B17-F24838734DB3" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax latens Mirande, Aguilera &amp; Azpelicueta 2004:215" order="Characiformes" pageNumber="66" phylum="Chordata" rank="species" species="latens">A. latens</taxonomicName>
, 
<taxonomicName id="07FAD6B998563A079EF2039E38DF3DEF" LSID-ZBK="urn:lsid:zoobank.org:act:277C9FEE-EFEC-4675-8415-DC69F6205FE7" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax chico Casciotta &amp; Almirón 2004:12" order="Characiformes" pageNumber="66" phylum="Chordata" rank="species" species="chico">A. chico</taxonomicName>
, 
<taxonomicName id="41C907F769244BE67932BBDABA9BED9A" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="66" phylum="Chordata" rank="species" species="lineatus">A. lineatus</taxonomicName>
, 
<taxonomicName id="FEEA831BFC71DA690F9D677B343622E3" LSID-ZBK="urn:lsid:zoobank.org:act:F06B3814-F57B-4FE4-9FA9-A74ED072124F" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax bimaculatus asuncionensis Géry 1972:3" order="Characiformes" pageNumber="66" phylum="Chordata" rank="species" species="asuncionensis">A. asuncionensis</taxonomicName>
, 
<taxonomicName id="AD4A0C15B045AB31D7C276D7C0AA9D53" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="66" phylum="Chordata" rank="species" species="abramis">A. abramis</taxonomicName>
, and 
<taxonomicName id="210FA735C772DA7F2BD8F380BD935929" LSID-ZBK="urn:lsid:zoobank.org:act:E47C3BA8-9E72-4C96-B1B3-9167C828A693" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax tumbayaensis Miquelarena &amp; Menni 2005" order="Characiformes" pageNumber="66" phylum="Chordata" rank="species" species="tumbayaensis">A. tumbayaensis</taxonomicName>
, with the last species limited to a restricted area in the highlands of Jujuy Province, Argentina.
</paragraph>
</subSubSection>
</treatment>
</document>