treatments-xml/data/A6/45/89/A6458955FFDB6624D1DDFE4BFE8677E7.xml

<document ID-DOI="http://doi.org/10.5281/zenodo.5872987" ID-GBIF-Dataset="6bc8f93f-8227-4361-9bef-0976c7102fe7" ID-GBIF-Taxon="192411996" ID-ISSN="1639-4798" ID-Zenodo-Dep="5872987" checkinTime="1629118158663" checkinUser="carolina" docAuthor="Rivero-Guerra, Aixa O." docDate="2012" docId="A6458955FFDB6624D1DDFE4BFE8677E7" docLanguage="en" docName="Adansonia.34.1.133-154.pdf" docOrigin="Adansonia (3) 34 (1)" docSource="http://dx.doi.org/10.5252/a2012n1a16" docStyle="DocumentStyle:B98EC8A323C0F63E1C55336AD58D9273.1:Adansonia.2011-2014.journal_article" docStyleId="B98EC8A323C0F63E1C55336AD58D9273" docStyleName="Adansonia.2011-2014.journal_article" docStyleVersion="1" docTitle="Santolina heterophylla Willk. &amp; Cut. (Willkomm 1859" docType="treatment" docVersion="4" lastPageId="5" lastPageNumber="138" masterDocId="5A7CF12DFFD86621D142FF9EFFD0734E" masterDocTitle="Santolina orocarpetana sp. nov. (Asteraceae: Anthemideae), a new species from the Iberian Peninsula. Revision of the lectotype of S. oblongifolia Boiss." masterLastPageNumber="154" masterPageNumber="133" pageId="3" pageNumber="136" updateTime="1642515947841" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title>Santolina orocarpetana sp. nov. (Asteraceae: Anthemideae), a new species from the Iberian Peninsula. Revision of the lectotype of S. oblongifolia Boiss.</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rivero-Guerra, Aixa O.</mods:namePart>
<mods:affiliation>Universidad de Sevilla, Facultad de Biología, Departamento de Biología Vegetal y Ecología, avda. Reina Mercedes, E- 41012 Sevilla (Spain) &amp; Centro de Investigação en Biodiversidade e Recursos Genéticos, Campus Agrario de Vairão, rua Padre Armando Quintas-Crasto, 4485 - 661 Vairão (Portugal)</mods:affiliation>
<mods:nameIdentifier type="email">rivero-guerra@hotmail.com</mods:nameIdentifier>
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<mods:titleInfo>
<mods:title>Adansonia</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="series">
<mods:title>3</mods:title>
</mods:detail>
<mods:detail type="pubDate">
<mods:number>2012-06-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>34</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>133</mods:start>
<mods:end>154</mods:end>
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<mods:url>http://dx.doi.org/10.5252/a2012n1a16</mods:url>
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<mods:identifier type="DOI">10.5252/a2012n1a16</mods:identifier>
<mods:identifier type="GBIF-Dataset">6bc8f93f-8227-4361-9bef-0976c7102fe7</mods:identifier>
<mods:identifier type="ISSN">1639-4798</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5872987" ID-GBIF-Taxon="192411996" ID-Zenodo-Dep="5872987" LSID="urn:lsid:plazi:treatment:A6458955FFDB6624D1DDFE4BFE8677E7" httpUri="http://treatment.plazi.org/id/A6458955FFDB6624D1DDFE4BFE8677E7" lastPageId="5" lastPageNumber="138" pageId="3" pageNumber="136">
<subSubSection box="[159,600,469,496]" pageId="3" pageNumber="136" type="nomenclature">
<paragraph blockId="3.[147,614,469,559]" box="[159,600,469,496]" pageId="3" pageNumber="136">
2. 
<taxonomicName authority="Willk. &amp; Cut." authorityName="Willk. &amp; Cut. (Willkomm" authorityYear="1859" box="[187,596,469,496]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="species" species="heterophylla">
<emphasis bold="true" box="[187,432,469,496]" italics="true" pageId="3" pageNumber="136">Santolina heterophylla</emphasis>
Willk. &amp; Cut.
</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="136" type="multiple">
<paragraph blockId="3.[147,614,469,559]" box="[253,507,501,527]" pageId="3" pageNumber="136">
<emphasis box="[253,292,502,527]" italics="true" pageId="3" pageNumber="136">non</emphasis>
<taxonomicName authority="Benth." authorityName="Benth." box="[299,503,501,527]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="species" species="pectinata">
<emphasis box="[299,420,501,527]" italics="true" pageId="3" pageNumber="136">S. pectinata</emphasis>
Benth.
</taxonomicName>
,
</paragraph>
<paragraph blockId="3.[147,614,469,559]" box="[147,614,532,559]" pageId="3" pageNumber="136">
<emphasis box="[147,186,533,558]" italics="true" pageId="3" pageNumber="136">non</emphasis>
<taxonomicName authority="DC" authorityName="DC" box="[193,489,532,559]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="species" species="heterophylla">
Lag. var. β 
<emphasis box="[314,438,532,558]" italics="true" pageId="3" pageNumber="136">heterophylla</emphasis>
DC
</taxonomicName>
, 
<emphasis box="[500,614,532,558]" italics="true" pageId="3" pageNumber="136">nom inval.</emphasis>
</paragraph>
</subSubSection>
<subSubSection box="[136,527,589,614]" pageId="3" pageNumber="136" type="reference_group">
<paragraph blockId="3.[114,648,589,772]" box="[136,527,589,614]" pageId="3" pageNumber="136">
<treatmentCitationGroup box="[136,524,589,614]" pageId="3" pageNumber="136">
<treatmentCitation author="Willk. &amp; Cut." box="[136,524,589,614]" journal="Linnaea" page="106" pageId="3" pageNumber="136" score="4.864797368871433" volume="30" year="1859">
<collectorName box="[136,274,590,614]" pageId="3" pageNumber="136">In Willkomm</collectorName>
, 
<emphasis box="[285,362,590,613]" italics="true" pageId="3" pageNumber="136">Linnaea</emphasis>
30: 106 (1859)
</treatmentCitation>
</treatmentCitationGroup>
.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="136" type="materials_examined">
<paragraph blockId="3.[114,648,589,772]" pageId="3" pageNumber="136">
— 
<materialsCitation ID-GBIF-Occurrence="3446332324" collectingDate="1855-06" collectionCode="COI-WILLK" collectorName="Ysern" country="Spain" location="Sierra de Gredos" pageId="3" pageNumber="136" specimenCode="COI-WILLK 00035956" specimenCount="1" typeStatus="lectotype">
Type (as given in protologue): “In Hispania centrali regione montana: in monte Sierra de Gredos et in provinciae Avilensi prope Servanillos, Cutanda et Ysern! (1857). Floret Julio”. — 
<typeStatus box="[287,386,696,720]" pageId="3" pageNumber="136">Lectotype</typeStatus>
(here designated): 
<collectingCountry box="[583,642,695,719]" name="Spain" pageId="3" pageNumber="136">Spain</collectingCountry>
, 
<location LSID="urn:lsid:plazi:treatment:A6458955FFDB6624D1DDFE4BFE8677E7:2B336E98FFDB6622D131FD4CFEF171A4" box="[115,289,722,746]" country="Spain" municipality="Ysern" name="Sierra de Gredos" pageId="3" pageNumber="136">Sierra de Gredos</location>
, 
<date box="[301,389,722,746]" pageId="3" pageNumber="136" value="1855-06">
<collectingDate box="[301,389,722,746]" pageId="3" pageNumber="136" value="1855-06">VI.1855</collectingDate>
</date>
, 
<emphasis box="[402,468,723,746]" italics="true" pageId="3" pageNumber="136">
<collectorName box="[402,455,723,746]" pageId="3" pageNumber="136">Ysern</collectorName>
s
</emphasis>
. 
<emphasis box="[475,488,723,746]" italics="true" pageId="3" pageNumber="136">n</emphasis>
. (
<specimenCode collectionCode="COI-WILLK" pageId="3" pageNumber="136">COI-WILLK 00035956</specimenCode>
; 
<figureCitation box="[229,287,748,772]" captionStart="FIG" captionStartId="4.[351,362,1680,1697]" captionTargetBox="[160,1163,215,1629]" captionTargetId="figure-26@4.[160,1163,215,1629]" captionTargetPageId="4" captionText="FIG 2. — Lectotype of Santolina heterophylla Willk. (COI-WILLK 00035956)." figureDoi="http://doi.org/10.5281/zenodo.5205805" httpUri="https://zenodo.org/record/5205805/files/figure.png" pageId="3" pageNumber="136">Fig. 2</figureCitation>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="138" pageId="3" pageNumber="136" type="discussion">
<paragraph blockId="3.[111,648,818,1700]" box="[113,210,818,844]" pageId="3" pageNumber="136">REMARKS</paragraph>
<paragraph blockId="3.[111,648,818,1700]" pageId="3" pageNumber="136">
<bibRefCitation author="CANDOLLE A. P. DE" box="[113,347,849,876]" firstAuthor="De Candolle" pageId="3" pageNumber="136" refId="ref9290" refString="CANDOLLE A. P. DE. 1837. - Prodromus systematis naturalis regni vegetabilis. Vol. VI. Sumptibus Sociorum Treuttel et Wurtz, Paris, 687 p." type="book" year="1837">De Candolle (1837)</bibRefCitation>
included 
<taxonomicName box="[465,595,850,876]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="species" species="canescens">
<emphasis box="[465,595,850,876]" italics="true" pageId="3" pageNumber="136">S. canescens</emphasis>
</taxonomicName>
and 
<taxonomicName box="[116,216,881,907]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="species" species="viscosa">
<emphasis box="[116,216,881,907]" italics="true" pageId="3" pageNumber="136">S. viscosa</emphasis>
</taxonomicName>
into the circumscription of the genus 
<taxonomicName box="[113,250,913,939]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="genus">
<emphasis box="[113,216,913,939]" italics="true" pageId="3" pageNumber="136">Santolina</emphasis>
L.
</taxonomicName>
, and reduced 
<taxonomicName authority="Lag." authorityName="Lag." box="[412,596,913,939]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="species" species="pectinata">
<emphasis box="[412,539,913,939]" italics="true" pageId="3" pageNumber="136">S. pectinata</emphasis>
Lag.
</taxonomicName>
as a variety of 
<taxonomicName authority="Benth." authorityName="Benth." box="[218,413,944,971]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="species" species="pectinata">
<emphasis box="[218,336,945,971]" italics="true" pageId="3" pageNumber="136">S. pectinata</emphasis>
Benth.
</taxonomicName>
(
<emphasis box="[426,548,944,970]" italics="true" pageId="3" pageNumber="136">nom. illegit.</emphasis>
). The author retained the epithet “ 
<taxonomicName box="[387,481,977,1002]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="species" species="pectinata">
<emphasis box="[387,481,977,1002]" italics="true" pageId="3" pageNumber="136">pectinata</emphasis>
</taxonomicName>
” for a variety 
<emphasis box="[626,639,978,1005]" italics="true" pageId="3" pageNumber="136">α</emphasis>
, whereas he conserved the 
<typeStatus box="[391,437,1008,1034]" pageId="3" pageNumber="136">type</typeStatus>
of 
<taxonomicName authority="Lag." authorityName="Lag." box="[472,647,1008,1034]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="species" species="pectinata">
<emphasis box="[472,593,1008,1034]" italics="true" pageId="3" pageNumber="136">S. pectinata</emphasis>
Lag.
</taxonomicName>
for a variety 
<emphasis box="[256,269,1042,1069]" italics="true" pageId="3" pageNumber="136">β</emphasis>
. He published var. 
<emphasis box="[491,504,1042,1069]" italics="true" pageId="3" pageNumber="136">β</emphasis>
. 
<taxonomicName box="[521,646,1039,1065]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="species" species="heterophylla">
<emphasis box="[521,646,1039,1065]" italics="true" pageId="3" pageNumber="136">heterophylla</emphasis>
</taxonomicName>
based on plants from the 
<emphasis box="[389,527,1071,1097]" italics="true" pageId="3" pageNumber="136">locus classicus</emphasis>
of 
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="species" species="pectinata">
<emphasis italics="true" pageId="3" pageNumber="136">S. pectinata</emphasis>
</taxonomicName>
. He also cited 
<taxonomicName authority="Lag." authorityName="Lag." box="[321,496,1103,1129]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="species" species="pectinata">
<emphasis box="[321,442,1103,1129]" italics="true" pageId="3" pageNumber="136">S. pectinata</emphasis>
Lag.
</taxonomicName>
as a synonym of this name, adding a short note as diagnosis “ 
<emphasis italics="true" pageId="3" pageNumber="136">foliis infimis et junioribus linearibus indivisis margine tuberculatis</emphasis>
”. 
<taxonomicName authority="Benth." authorityName="Benth." box="[258,540,1198,1224]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="species" species="pectinata">
<emphasis box="[258,458,1198,1224]" italics="true" pageId="3" pageNumber="136">Santolina pectinata</emphasis>
Benth.
</taxonomicName>
, non 
<taxonomicName authority="DC." authorityName="DC." class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="species" species="heterophylla">
Lag. var. 
<emphasis box="[157,170,1232,1259]" italics="true" pageId="3" pageNumber="136">β</emphasis>
. 
<emphasis box="[182,306,1229,1255]" italics="true" pageId="3" pageNumber="136">heterophylla</emphasis>
DC.
</taxonomicName>
, is an invalid name(ICBN: arts 53.1, 26.2; 
<bibRefCitation author="Mcneill" box="[292,501,1261,1288]" etAl="et al." firstAuthor="Mcneill" pageId="3" pageNumber="136" refId="ref9601" refString="MCNEILL J., BARRIE F. R., BURDET H. M., DE- MOULIN V., HAWKSWORTH D. L., MARHOLD K., NICOLSON D. H., PRADO J., SILVA P. C., SKOG J. E., WIERSEMA J. H. &amp; TURLAND N. J. 2006. - International Code of Botanical Nomenclature (Vienna Code) adopted by the Seventeenth International Botanical Congress Vienna. Austria, July 2005. A. R." type="proceedings" year="2006">
Mcneill 
<emphasis box="[382,425,1261,1287]" italics="true" pageId="3" pageNumber="136">et al</emphasis>
. 2006
</bibRefCitation>
).
</paragraph>
<paragraph blockId="3.[111,648,818,1700]" lastBlockId="3.[674,1212,216,1699]" pageId="3" pageNumber="136">
Furthermore,Willkomm &amp; Cutanda 
<emphasis box="[509,530,1294,1319]" italics="true" pageId="3" pageNumber="136">in</emphasis>
<bibRefCitation author="Willkomm" firstAuthor="Willkomm" pageId="3" pageNumber="136" pagination="106 - 107" refId="ref9993" refString="WILLKOMM M. 1859. - Pugillus plantarum novarum peninsulae pyrenaicae. Linnaea 30: 106 - 107." type="journal article" year="1859">Willkomm (1859)</bibRefCitation>
, and Willkomm 
<emphasis box="[386,408,1325,1350]" italics="true" pageId="3" pageNumber="136">in</emphasis>
<bibRefCitation author="Willkomm &amp; Lange" firstAuthor="Willkomm" pageId="3" pageNumber="136" refId="ref10035" refString="WILLKOMM M. &amp; LANGE J. 1865. - Prodromus Florae Hispanicae II. Typis e sumtibus librariae E. Schweizerbart, Stuttgartiae, 680 p." type="book" year="1865">Willkomm &amp; Lange (1865)</bibRefCitation>
ignored the variety 
<taxonomicName box="[393,538,1356,1385]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="species" species="heterophylla">
<emphasis box="[393,404,1358,1385]" italics="true" pageId="3" pageNumber="136">β</emphasis>
. 
<emphasis box="[416,538,1356,1382]" italics="true" pageId="3" pageNumber="136">heterophylla</emphasis>
</taxonomicName>
published by 
<bibRefCitation author="CANDOLLE A. P. DE" box="[143,362,1388,1415]" firstAuthor="De Candolle" pageId="3" pageNumber="136" refId="ref9290" refString="CANDOLLE A. P. DE. 1837. - Prodromus systematis naturalis regni vegetabilis. Vol. VI. Sumptibus Sociorum Treuttel et Wurtz, Paris, 687 p." type="book" year="1837">De Candolle (1837)</bibRefCitation>
. Willkomm &amp; Cutanda 
<emphasis box="[623,645,1389,1414]" italics="true" pageId="3" pageNumber="136">in</emphasis>
<bibRefCitation author="Willkomm" box="[112,305,1419,1446]" firstAuthor="Willkomm" pageId="3" pageNumber="136" pagination="106 - 107" refId="ref9993" refString="WILLKOMM M. 1859. - Pugillus plantarum novarum peninsulae pyrenaicae. Linnaea 30: 106 - 107." type="journal article" year="1859">Willkomm (1859)</bibRefCitation>
published a new taxon, 
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="species" species="heterophylla">
<emphasis italics="true" pageId="3" pageNumber="136">S. heterophylla</emphasis>
</taxonomicName>
, based on a specimen from Sierra de Gredos. Willkomm 
<emphasis box="[235,257,1484,1509]" italics="true" pageId="3" pageNumber="136">in</emphasis>
<bibRefCitation author="Willkomm &amp; Lange" box="[261,555,1483,1510]" firstAuthor="Willkomm" pageId="3" pageNumber="136" refId="ref10035" refString="WILLKOMM M. &amp; LANGE J. 1865. - Prodromus Florae Hispanicae II. Typis e sumtibus librariae E. Schweizerbart, Stuttgartiae, 680 p." type="book" year="1865">Willkomm &amp; Lange (1865)</bibRefCitation>
reduced 
<taxonomicName authority="Wilk. &amp; Cut." authorityName="Willk. &amp; Cut. (Willkomm" authorityYear="1859" box="[112,417,1515,1541]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="species" species="heterophylla">
<emphasis box="[112,263,1515,1541]" italics="true" pageId="3" pageNumber="136">S. heterophylla</emphasis>
Wilk. &amp; Cut.
</taxonomicName>
to a variety of 
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="species" species="rosmarinifolia">
<emphasis italics="true" pageId="3" pageNumber="136">S. rosmarinifolia</emphasis>
</taxonomicName>
, but he omitted Cutanda as author of the epithet. He cited 
<taxonomicName authority="Lag." authorityName="Lag." box="[334,503,1578,1605]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="species" species="pectinata">
<emphasis box="[334,452,1578,1604]" italics="true" pageId="3" pageNumber="136">S. pectinata</emphasis>
Lag.
</taxonomicName>
as a synonym of “ 
<emphasis box="[154,238,1610,1636]" italics="true" pageId="3" pageNumber="136">variat 1</emphasis>
” of 
<taxonomicName authority="Willk." authorityName="Willk. &amp; Cut. (Willkomm" authorityYear="1859" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="species" species="rosmarinifolia">
<emphasis box="[289,464,1610,1636]" italics="true" pageId="3" pageNumber="136">S. rosmarinifolia</emphasis>
L. var. 
<emphasis box="[552,565,1612,1639]" italics="true" pageId="3" pageNumber="136">β</emphasis>
<emphasis italics="true" pageId="3" pageNumber="136">heterophylla</emphasis>
Willk.
</taxonomicName>
, whereas 
<bibRefCitation author="Willkomm" box="[352,551,1641,1668]" firstAuthor="Willkomm" pageId="3" pageNumber="136" refId="ref10012" refString="WILLKOMM M. 1893. - Supplementum Prodromi Florae Hispanicae. Schweizerbart (E. Koch), Stuttgartiae, 370 p." type="book" year="1893">Willkomm (1893)</bibRefCitation>
changed the original concept and accepted 
<taxonomicName authority="Willk." authorityName="Willk. &amp; Cut. (Willkomm" authorityYear="1859" box="[475,948,216,1700]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="species" species="rosmarinifolia">
<emphasis box="[475,488,1673,1699]" italics="true" pageId="3" pageNumber="136">S</emphasis>
. 
<emphasis box="[500,646,1673,1699]" italics="true" pageId="3" pageNumber="136">rosmarinifolia</emphasis>
var. 
<emphasis box="[723,736,218,245]" italics="true" pageId="3" pageNumber="136">β</emphasis>
<emphasis box="[745,869,216,242]" italics="true" pageId="3" pageNumber="136">heterophylla</emphasis>
Willk.
</taxonomicName>
, f. 
<emphasis box="[983,997,216,242]" italics="true" pageId="3" pageNumber="136">1</emphasis>
Willk. as synonym of 
<taxonomicName box="[707,832,248,274]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="species" species="pectinata">
<emphasis box="[707,720,248,274]" italics="true" pageId="3" pageNumber="136">S</emphasis>
. 
<emphasis box="[737,832,248,273]" italics="true" pageId="3" pageNumber="136">pectinata</emphasis>
</taxonomicName>
. He placed this 
<emphasis box="[1017,1080,248,273]" italics="true" pageId="3" pageNumber="136">variat</emphasis>
in the 
<typeStatus box="[1163,1209,248,274]" pageId="3" pageNumber="136">type</typeStatus>
locality of 
<taxonomicName box="[797,923,280,306]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="species" species="pectinata">
<emphasis box="[797,923,280,306]" italics="true" pageId="3" pageNumber="136">S. pectinata</emphasis>
</taxonomicName>
, and in Sierra de la Sagra (COI-WILLK 00035964, MPU, P 00752621 from “La Puebla de Don Fadrique et montem La Sagra”).
</paragraph>
<paragraph blockId="3.[674,1212,216,1699]" pageId="3" pageNumber="136">
Willkomm misspelled the locality for the 
<typeStatus box="[1115,1208,374,400]" pageId="3" pageNumber="136">lectotype</typeStatus>
of 
<taxonomicName authorityName="Willk. &amp; Cut. (Willkomm" authorityYear="1859" box="[704,855,406,432]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="species" species="heterophylla">
<emphasis box="[704,855,406,432]" italics="true" pageId="3" pageNumber="136">S. heterophylla</emphasis>
</taxonomicName>
; the correct name is “Serranillos” (
<bibRefCitation author="Cerveira Rodrigues Gomes" box="[684,1046,437,464]" firstAuthor="Cerveira Rodrigues Gomes" pageId="3" pageNumber="136" refId="ref9387" refString="CERVEIRA RODRIGUES GOMES C. I. 2000. - Estudo nomenclatural e taxonomico de ejemplares tipo do herbario de Willkomm. Universidade de Coimbra, Departamento de Botanica. Jardim Botanico, Coimbra." type="book" year="2000">Cerveira Rodrigues Gomes 2000</bibRefCitation>
), and it will be used hereafter in this paper.
</paragraph>
<paragraph blockId="3.[674,1212,216,1699]" pageId="3" pageNumber="136">
Two collections of Isern were found. In the first one, labelled with the name “ 
<taxonomicName box="[986,1133,532,559]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="species" species="oblongifolia">
<emphasis box="[986,1133,532,559]" italics="true" pageId="3" pageNumber="136">S. oblongifolia</emphasis>
</taxonomicName>
”, from Puerto del Pico (
<accessionNumber box="[845,980,564,591]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/MA126799" pageId="3" pageNumber="136">MA 126799</accessionNumber>
), the date is only indicated as“August”.The other specimen (COI-WILLK 00035956), labelled by Willkomm with the names 
<taxonomicName box="[675,967,659,688]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="species" species="oblongifolia">
<emphasis box="[675,967,659,688]" italics="true" pageId="3" pageNumber="136">S. oblongifolia β ceratophylla</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Willk. &amp; Cut. (Willkomm" authorityYear="1859" box="[1017,1166,659,686]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="species" species="heterophylla">
<emphasis box="[1017,1166,659,686]" italics="true" pageId="3" pageNumber="136">S. heterophylla</emphasis>
</taxonomicName>
was gathered by Isern in 
<date box="[890,993,691,718]" pageId="3" pageNumber="136" value="1855-07">
July 
<quantity box="[937,993,691,718]" metricMagnitude="1" metricUnit="m" metricValue="4.7116999999999996" pageId="3" pageNumber="136" unit="in" value="1855.0">1855</quantity>
</date>
in Sierra de Gredos. Presumably, Willkomm changed 1855 into 1857 during transcription of the original label.
</paragraph>
<paragraph blockId="3.[674,1212,216,1699]" lastBlockId="5.[111,649,216,1193]" lastPageId="5" lastPageNumber="138" pageId="3" pageNumber="136">
The 
<typeStatus box="[748,844,786,812]" pageId="3" pageNumber="136">lectotype</typeStatus>
(COI-WILLK 00035956; 
<figureCitation box="[1138,1202,786,812]" captionStart="FIG" captionStartId="4.[351,362,1680,1697]" captionTargetBox="[160,1163,215,1629]" captionTargetId="figure-26@4.[160,1163,215,1629]" captionTargetPageId="4" captionText="FIG 2. — Lectotype of Santolina heterophylla Willk. (COI-WILLK 00035956)." figureDoi="http://doi.org/10.5281/zenodo.5205805" httpUri="https://zenodo.org/record/5205805/files/figure.png" pageId="3" pageNumber="136">Fig. 2</figureCitation>
) has six flowering stems and various sterile stems. The sheet has two labels in the centre of the lower margin with the following notation in black ink: 
<emphasis bold="true" box="[678,758,913,939]" pageId="3" pageNumber="136">Label 1</emphasis>
: “Herbarium Willkommi [printed] // 
<taxonomicName authority="Mill." authorityName="Mill." class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="species" species="rosmarinifolia">
<emphasis italics="true" pageId="3" pageNumber="136">Santolina rosmarinifolia</emphasis>
Mill.
</taxonomicName>
[underlined] var?. 
<taxonomicName box="[1145,1209,945,970]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="species" species="sericea">
<emphasis box="[1145,1209,945,970]" italics="true" pageId="3" pageNumber="136">sericea</emphasis>
</taxonomicName>
[underlined] Wk.= 
<taxonomicName authority="Wk." authorityName="Willk. &amp; Cut. (Willkomm" authorityYear="1859" box="[881,1075,976,1003]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="species" species="heterophylla">
<emphasis box="[881,1024,976,1003]" italics="true" pageId="3" pageNumber="136">S. heterophylla</emphasis>
Wk.
</taxonomicName>
[underlined] et Cut. // Sierra de Gredos [Willkomm’s handwritten notation] // Legit [printed] Ysern // 
<date box="[1098,1209,1039,1066]" pageId="3" pageNumber="136" value="1855-07">Julio 1855</date>
[Willkomm’s handwritten notation]”; 
<emphasis bold="true" box="[1064,1141,1071,1097]" pageId="3" pageNumber="136">Label 2</emphasis>
: “ 
<taxonomicName authority="Boiss." authorityName="Boiss." class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="species" species="oblongifolia">
<emphasis italics="true" pageId="3" pageNumber="136">Santolina oblongifolia</emphasis>
Boiss.
</taxonomicName>
[underlined] 
<emphasis box="[1072,1210,1103,1132]" italics="true" pageId="3" pageNumber="136">β ceratophylla</emphasis>
[underlined] Wk. [Willkomm’s handwritten notation]”. This specimen has two revision labels by D. Sánchez-Mata and S. Sardinero with the following notation in black ink: 
<emphasis bold="true" box="[903,981,1229,1256]" pageId="3" pageNumber="136">Label 1</emphasis>
: “ 
<taxonomicName authority="Willk." authorityName="Willk. (Willkomm &amp; Lange" authorityYear="1865" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="variety" species="oblongifolia" variety="ceratophylla">
<emphasis italics="true" pageId="3" pageNumber="136">Santolina oblongifolia</emphasis>
Boiss var. 
<emphasis box="[815,934,1261,1287]" italics="true" pageId="3" pageNumber="136">ceratophylla</emphasis>
Willk.
</taxonomicName>
<emphasis box="[1014,1036,1262,1287]" italics="true" pageId="3" pageNumber="136">in</emphasis>
Willk. &amp; Lange // Typus [handwritten] // 
<collectingRegion box="[950,1045,1293,1319]" country="Portugal" name="Coimbra" pageId="3" pageNumber="136">Coimbra</collectingRegion>
[printed] 
<date pageId="3" pageNumber="136" value="1993-09">1993 IX</date>
[handwritten]”; 
<emphasis bold="true" box="[882,962,1324,1351]" pageId="3" pageNumber="136">Label 2</emphasis>
: “ 
<taxonomicName authority="Boiss." authorityName="Boiss." class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="species" species="oblongifolia">
<emphasis box="[987,1210,1324,1350]" italics="true" pageId="3" pageNumber="136">Santolina oblongifolia</emphasis>
Boiss.
</taxonomicName>
[handwritten] // 
<collectingRegion box="[920,1015,1356,1382]" country="Portugal" name="Coimbra" pageId="3" pageNumber="136">Coimbra</collectingRegion>
[printed] 
<date box="[1121,1209,1356,1383]" pageId="3" pageNumber="136" value="1993-09">1993 IX</date>
[handwritten]”. This specimen displays all the characteristics listed in the protologue (Appendix 1). The 
<typeStatus box="[677,772,1451,1477]" pageId="3" pageNumber="136">lectotype</typeStatus>
clearly shows intermediate characteristics between 
<taxonomicName box="[770,915,1483,1510]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="species" species="oblongifolia">
<emphasis box="[770,783,1483,1509]" italics="true" pageId="3" pageNumber="136">S</emphasis>
. 
<emphasis box="[797,915,1483,1509]" italics="true" pageId="3" pageNumber="136">oblongifolia</emphasis>
</taxonomicName>
and 
<taxonomicName authority="subsp. rosmarinifolia" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="subSpecies" species="rosmarinifolia" subSpecies="rosmarinifolia">
<emphasis box="[968,981,1483,1509]" italics="true" pageId="3" pageNumber="136">S</emphasis>
. 
<emphasis box="[995,1137,1483,1509]" italics="true" pageId="3" pageNumber="136">rosmarinifolia</emphasis>
subsp. 
<emphasis box="[677,820,1515,1541]" italics="true" pageId="3" pageNumber="136">rosmarinifolia</emphasis>
</taxonomicName>
. The relationship between 
<taxonomicName authorityName="Willk. &amp; Cut. (Willkomm" authorityYear="1859" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="species" species="heterophylla">
<emphasis italics="true" pageId="3" pageNumber="136">S. heterophylla</emphasis>
</taxonomicName>
, 
<taxonomicName authority="subsp. rosmarinifolia" box="[755,1160,1546,1573]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="subSpecies" species="rosmarinifolia" subSpecies="rosmarinifolia">
<emphasis box="[755,930,1546,1573]" italics="true" pageId="3" pageNumber="136">S. rosmarinifolia</emphasis>
subsp. 
<emphasis box="[1014,1160,1546,1572]" italics="true" pageId="3" pageNumber="136">rosmarinifolia</emphasis>
</taxonomicName>
and 
<taxonomicName box="[678,823,1578,1604]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="species" species="oblongifolia">
<emphasis box="[678,823,1578,1604]" italics="true" pageId="3" pageNumber="136">S. oblongifolia</emphasis>
</taxonomicName>
has been cited by 
<bibRefCitation author="Willkomm" box="[1015,1211,1578,1605]" firstAuthor="Willkomm" pageId="3" pageNumber="136" pagination="106 - 107" refId="ref9993" refString="WILLKOMM M. 1859. - Pugillus plantarum novarum peninsulae pyrenaicae. Linnaea 30: 106 - 107." type="journal article" year="1859">Willkomm (1859)</bibRefCitation>
in the protologue: “ 
<emphasis box="[876,1024,1610,1636]" italics="true" pageId="3" pageNumber="136">Specie proxima</emphasis>
<taxonomicName authority="Mill." authorityName="Mill." class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="136" phylum="Tracheophyta" rank="species" species="rosmarinifoliae">
S. rosmarinifoliae 
<emphasis box="[677,731,1641,1667]" italics="true" pageId="3" pageNumber="136">Mill.</emphasis>
</taxonomicName>
<emphasis italics="true" lastPageId="5" lastPageNumber="138" pageId="3" pageNumber="136">, a cujus formis numerosis foliis eximie heteromorphis planis sericeis statim distinguitur. Calathiorum florumque structura a</emphasis>
<taxonomicName box="[345,521,216,242]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="138" phylum="Tracheophyta" rank="species" species="rosmarinifolia">S. rosmarinifolia</taxonomicName>
<emphasis italics="true" pageId="5" pageNumber="138">non differre videtur, sed propter foliorum formam ab omnibus</emphasis>
Santolinis 
<emphasis italics="true" pageId="5" pageNumber="138">valde discrepantem hanc stirpem ad illam speciem referre non audeo.</emphasis>
<taxonomicName authority="Boiss." authorityName="Boiss." box="[379,594,311,337]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="138" phylum="Tracheophyta" rank="species" species="oblongifolia">
S. oblongifolia 
<emphasis box="[538,594,312,337]" italics="true" pageId="5" pageNumber="138">Boiss.</emphasis>
</taxonomicName>
<emphasis italics="true" pageId="5" pageNumber="138">foliis planis sericeis pariter gaudens ramis sterilibus elongatis, foliis multo minus difformibus aliaque ratione formatis a nostra differre videtur</emphasis>
[
<emphasis box="[364,392,407,432]" italics="true" pageId="5" pageNumber="138">…</emphasis>
]”.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5205805" ID-Zenodo-Dep="5205805" box="[351,974,1680,1697]" httpUri="https://zenodo.org/record/5205805/files/figure.png" pageId="4" pageNumber="137" startId="4.[351,362,1680,1697]" targetBox="[160,1163,215,1629]" targetPageId="4">
<paragraph blockId="4.[351,974,1680,1697]" box="[351,974,1680,1697]" pageId="4" pageNumber="137">
FIG 2. — Lectotype of 
<taxonomicName authority="Willk." authorityName="Willk. &amp; Cut. (Willkomm" authorityYear="1859" box="[535,771,1680,1697]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="137" phylum="Tracheophyta" rank="species" species="heterophylla">
<emphasis box="[535,719,1680,1697]" italics="true" pageId="4" pageNumber="137">Santolina heterophylla</emphasis>
Willk.
</taxonomicName>
(COI-WILLK 00035956).
</paragraph>
</caption>
<paragraph blockId="5.[111,649,216,1193]" pageId="5" pageNumber="138">
The sheet A 00251592 has two labels in the lower left margin: 
<emphasis bold="true" box="[252,337,469,495]" pageId="5" pageNumber="138">Label 1</emphasis>
: “HERBARIUM GRAELSIANUM // PLANTAE CASTELLANAE [printed] // 
<taxonomicName authority="Wk." authorityName="Willk. &amp; Cut. (Willkomm" authorityYear="1859" box="[187,502,533,559]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="138" phylum="Tracheophyta" rank="species" species="heterophylla">Santolina heterophylla Wk.</taxonomicName>
// in saxosis Sierra de Gredos et Puerto del Pico prope riventos // Julio [Graells’ handwritten notation]”; and 
<emphasis bold="true" box="[565,644,596,622]" pageId="5" pageNumber="138">Label 2</emphasis>
(printed): “HERBARIUM OF GEORGE CURLING JOAD. PRESENTED BY THE ROYAL GARDENS, KEW, 1882”. The labels do not show the collector and year. The notation of the labels does not match with the 
<typeStatus box="[391,492,754,780]" pageId="5" pageNumber="138">lectotype</typeStatus>
, however, this specimen is the counterpart of the type material of this name. 
<bibRefCitation author="Carrasco" box="[273,518,818,844]" etAl="et al." firstAuthor="Carrasco" pageId="5" pageNumber="138" pagination="93 - 103" refId="ref9321" refString="CARRASCO M. A., PEREA D., ARAGON C. F. &amp; GARCIA- CAMACHO R. 2002. - La coleccion &quot; Plantes de la province de Granade &quot; de Pedro del Campo, en el herbario del Real Colegio Alfonso XII de San Lorenzo de El Escorial, Madrid (Espana). Botanica Complutensis 26: 93 - 103." type="journal article" year="2002">
Carrasco 
<emphasis box="[379,434,818,844]" italics="true" pageId="5" pageNumber="138">et al.</emphasis>
(2002)
</bibRefCitation>
discuss the value of the collections of the 
<emphasis italics="true" pageId="5" pageNumber="138">Real Colegio Alfonso XII</emphasis>
historical herbarium of 
<emphasis italics="true" pageId="5" pageNumber="138">San Lorenzo del Escorial</emphasis>
(
<collectingRegion box="[165,251,913,939]" country="Spain" name="Madrid" pageId="5" pageNumber="138">Madrid</collectingRegion>
, 
<collectingCountry box="[263,328,913,940]" name="Spain" pageId="5" pageNumber="138">Spain</collectingCountry>
). They argue that the Graells’ collection was used by Willkomm &amp; Lange as part of the material examined for the 
<emphasis italics="true" pageId="5" pageNumber="138">Prodromus florae hispanicae</emphasis>
. However, 
<taxonomicName authorityName="Willk. &amp; Cut. (Willkomm" authorityYear="1859" box="[341,491,1008,1034]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="138" phylum="Tracheophyta" rank="species" species="heterophylla">
<emphasis box="[341,491,1008,1034]" italics="true" pageId="5" pageNumber="138">S. heterophylla</emphasis>
</taxonomicName>
was published several years before the publication of the species corology of the genus 
<taxonomicName box="[344,443,1071,1097]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="138" phylum="Tracheophyta" rank="genus">
<emphasis box="[344,443,1071,1097]" italics="true" pageId="5" pageNumber="138">Santolina</emphasis>
</taxonomicName>
in 
<emphasis italics="true" pageId="5" pageNumber="138">Prodromus florae hispanicae</emphasis>
(1865). There is not enough evidence to justify the choice of this material as original material of 
<taxonomicName authorityName="Willk. &amp; Cut. (Willkomm" authorityYear="1859" box="[184,335,1166,1193]" class="Magnoliopsida" family="Asteraceae" genus="Santolina" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="138" phylum="Tracheophyta" rank="species" species="heterophylla">
<emphasis box="[184,335,1166,1193]" italics="true" pageId="5" pageNumber="138">S. heterophylla</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>