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<document id="46FAB164045EFFB4E0D71A8317D83540" ID="10.11646/zootaxa.4270.1.1SLASH11140" ID-CLB-Dataset="32975" ID-DOI="10.5281/zenodo.583183" ID-GBIF-Dataset="6c479acc-8b18-4f0b-a6e5-85bcd6d7b6b7" ID-ISSN="1175-5326" ID-Zenodo-Dep="583183" ID-ZooBank="2D00AFF5-4FE2-4EC1-A328-C8670CFB8D6D" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1495696370209" checkinUser="plazi" docAuthor="Jong, Rienk De" docDate="2017" docId="03AA87D32859FFCEF7F0FCA7FB86B581" docLanguage="en" docName="zootaxa.4270.1.1SLASH11140.pdf" docOrigin="Zootaxa 4270 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Archaeolycorea ferreirai" docType="treatment" docVersion="9" lastPageNumber="26" masterDocId="FF93FFAB2840FFD7F767FF87FFA3B03E" masterDocTitle="Fossil butterflies, calibration points and the molecular clock (Lepidoptera: Papilionoidea)" masterLastPageNumber="63" masterPageNumber="1" pageNumber="26" updateTime="1698439901205" updateUser="ExternalLinkService">
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<mods:title id="CC31EA1440D24516FD8CD470DB1A1CCD">Fossil butterflies, calibration points and the molecular clock (Lepidoptera: Papilionoidea)</mods:title>
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<mods:namePart id="9E5EB113EFCC97699AE9BE852699D888">Jong, Rienk De</mods:namePart>
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<mods:date id="308D85D96EF60D1E83BC10CCF53AFAB0">2017</mods:date>
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<treatment id="03AA87D32859FFCEF7F0FCA7FB86B581" ID-DOI="http://doi.org/10.5281/zenodo.6046952" ID-GBIF-Taxon="130844921" ID-Zenodo-Dep="6046952" LSID="urn:lsid:plazi:treatment:03AA87D32859FFCEF7F0FCA7FB86B581" httpUri="http://treatment.plazi.org/id/03AA87D32859FFCEF7F0FCA7FB86B581" lastPageNumber="26" pageId="25" pageNumber="26">
<subSubSection id="C319654E2859FFCEF7F0FCA7FF7CB364" pageId="25" pageNumber="26" type="nomenclature">
<paragraph id="8BBC36C52859FFCEF7F0FCA7FC8FB304" blockId="25.[151,812,800,859]" box="[151,812,800,826]" pageId="25" pageNumber="26">
<heading id="D0F481A92859FFCEF7F0FCA7FC8FB304" bold="true" box="[151,812,800,826]" fontSize="11" level="1" pageId="25" pageNumber="26" reason="1">
<emphasis id="B977EAD72859FFCEF7F0FCA7FC8FB304" bold="true" box="[151,812,800,826]" pageId="25" pageNumber="26">
<emphasis id="B977EAD72859FFCEF7F0FCA7FEA0B304" bold="true" box="[151,259,800,826]" italics="true" pageId="25" pageNumber="26">
<taxonomicName id="4C034D462859FFCEF7F0FCA7FF5CB304" box="[151,255,800,826]" class="Insecta" family="Nymphalidae" genus="Archaeolycorea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="ferreirai">ferreirai</taxonomicName>
.
</emphasis>
<taxonomicName id="4C034D462859FFCEF66DFCA7FC8FB304" authority="Martins Neto, 1989" authorityName="Martins Neto" authorityYear="1989" box="[266,812,800,826]" class="Insecta" family="Nymphalidae" genus="Archaeolycorea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="ferreirai">
<emphasis id="B977EAD72859FFCEF66DFCA7FD96B304" bold="true" box="[266,565,800,826]" italics="true" pageId="25" pageNumber="26">Archaeolycorea ferreirai</emphasis>
<bibRefCitation id="EF924B342859FFCEF55BFCA7FC8FB304" author="Martins" box="[572,812,800,826]" pageId="25" pageNumber="26" refString="Martins Neto, R. G. (1989) Novos insetos terciarios do Estado de Sao Paulo. Revista Brasileira de Geociencias, 19, 375 - 386." type="journal article" year="1989">Martins Neto, 1989</bibRefCitation>
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<paragraph id="8BBC36C52859FFCEF7F0FCC5FF7CB364" blockId="25.[151,812,800,859]" box="[151,223,834,859]" pageId="25" pageNumber="26">Fig. 7.</paragraph>
</subSubSection>
<subSubSection id="C319654E2859FFCEF7F0FC0DFB86B581" pageId="25" pageNumber="26" type="description">
<paragraph id="8BBC36C52859FFCEF7F0FC0DFE2FB39D" blockId="25.[151,1436,906,1471]" box="[151,396,906,931]" pageId="25" pageNumber="26">Family incertae sedis.</paragraph>
<paragraph id="8BBC36C52859FFCEF7A0FC29FE9CB3D5" blockId="25.[151,1436,906,1471]" pageId="25" pageNumber="26">
<materialsCitation id="3B6B3C982859FFCEF7A0FC29FE9CB3D5" ID-GBIF-Occurrence="1503739219" country="Brazil" location="Sao Paulo" pageId="25" pageNumber="26" specimenCount="1" stateProvince="Sao Paulo">
Tremembé Formation, Taubaté Basin, State of
<collectingRegion id="49C7F8272859FFCEF5B0FC28FCEFB3F9" box="[727,844,942,967]" country="Brazil" name="Sao Paulo" pageId="25" pageNumber="26">São Paulo</collectingRegion>
,
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; (Rupelian)—(Aquitanian); late Oligoceneearly Miocene.
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</paragraph>
<paragraph id="8BBC36C52859FFCEF7A0FC71FD38B431" blockId="25.[151,1436,906,1471]" box="[199,667,1014,1039]" pageId="25" pageNumber="26">Depository: IGEO (holotype, no. 5618-I).</paragraph>
<paragraph id="8BBC36C52859FFCEF7A0FB9DFD41B40D" blockId="25.[151,1436,906,1471]" box="[199,738,1050,1075]" pageId="25" pageNumber="26">
Published figure: Martins-Neto (1989:
<figureCitation id="13382A402859FFCEF51BFB9DFD1DB40D" box="[636,702,1050,1075]" captionStart="FIGURES 3 4" captionStartId="10.[151,264,1801,1823]" captionTargetBox="[165,1429,840,1774]" captionTargetId="figure@10.[154,1438,828,1788]" captionTargetPageId="10" captionText="FIGURES 3 4. Wing venation of (A) Pareronia Bingham, 1907 (Pieridae), and (B) Hasora Moore, 1881 (Hesperiidae). See text for explanation." httpUri="https://zenodo.org/record/583186/files/figure.png" pageId="25" pageNumber="26">Fig. 4</figureCitation>
A).
</paragraph>
<paragraph id="8BBC36C52859FFCEF7A0FBB8FB86B581" blockId="25.[151,1436,906,1471]" pageId="25" pageNumber="26">
Part of a forewing of which the distal area, about 1/4 of the wing surface, is missing. The reconstruction of the termen in
<figureCitation id="13382A402859FFCEF66AFBE5FEF2B445" box="[269,337,1122,1147]" captionStart="FIGURES 3 4" captionStartId="10.[151,264,1801,1823]" captionTargetBox="[165,1429,840,1774]" captionTargetId="figure@10.[154,1438,828,1788]" captionTargetPageId="10" captionText="FIGURES 3 4. Wing venation of (A) Pareronia Bingham, 1907 (Pieridae), and (B) Hasora Moore, 1881 (Hesperiidae). See text for explanation." httpUri="https://zenodo.org/record/583186/files/figure.png" pageId="25" pageNumber="26">Fig. 4</figureCitation>
A of the publication and with that, the shape of the wing, is, therefore, uncertain. The venation shows a most unusual arrangement: M1 branches off M2 shortly before the middle of the latter. In many butterfly genera (particularly in
<taxonomicName id="4C034D462859FFCEF6FCFB2DFE5AB4FD" box="[411,505,1194,1219]" class="Insecta" family="Pieridae" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="family">Pieridae</taxonomicName>
) M1 branches off R5, but I have not yet found such an arrangement illustrated in any extant butterfly genus. It could be an incorrect interpretation of the fossil, but apart from being clear in the figure, it is also explicitly described so in the text. The remainder part of the venation is not helpful. Since it is not clear in the figure where exactly is the distal end of the discoidal cell, either R3 and R4+5 have a common stem (an arrangement commonly found in butterflies), or R3 is separate and R4 and R5 have a common stem, an unusual arrangement in butterflies, but at least found in
<taxonomicName id="4C034D462859FFCEF5DEFAD9FC43B549" authority="Drury" authorityName="Drury" box="[697,992,1374,1399]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="antimachus">
<emphasis id="B977EAD72859FFCEF5DEFAD9FC36B549" box="[697,917,1374,1399]" italics="true" pageId="25" pageNumber="26">Papilio antimachus</emphasis>
Drury
</taxonomicName>
from the Afrotropics. In summary, the taxonomic position of this fossil is unclear and the suggestion implied by the name that it is in the ancestral line of the extant genus
<taxonomicName id="4C034D462859FFCEF633FA20FE0EB580" box="[340,429,1447,1470]" class="Insecta" family="Nymphalidae" genus="Lycorea" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="B977EAD72859FFCEF633FA20FE0EB580" box="[340,429,1447,1470]" italics="true" pageId="25" pageNumber="26">Lycorea</emphasis>
</taxonomicName>
(
<taxonomicName id="4C034D462859FFCEF6DBFA21FDFBB581" box="[444,600,1446,1471]" class="Insecta" family="Nymphalidae" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="family">Nymphalidae</taxonomicName>
,
<taxonomicName id="4C034D462859FFCEF504FA21FD6DB581" box="[611,718,1446,1471]" class="Insecta" family="Danainae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="family">Danainae</taxonomicName>
) is not supported by evidence.
</paragraph>
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