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<document id="415B7432C8CAB88F5D51C4C9B980A66B" ID-DOI="10.37520/aemnp.2021.006" ID-ISSN="1804-6487" ID-Zenodo-Dep="5037544" ID-ZooBank="7A927D20-F3B3-4611-BDC3-7EC99A131B69" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624896857361" checkinUser="carolina" docAuthor="Anichtchenko, Alexander, Minkina, Łukasz, Vasiljeva, Anastasija &amp; Medina, Milton Norman D." docDate="2021" docId="03AA81215331FFD1508C6A30B86DF834" docLanguage="en" docName="ActaEntMusNatPra.61.1.99-111.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 61 (1)" docSource="http://dx.doi.org/10.37520/aemnp.2021.006" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Rhyparus philippinensis Arrow 1905" docType="treatment" docVersion="3" lastPageNumber="109" masterDocId="FF93F9595338FFDB50746D24BA16FFBC" masterDocTitle="A review of the genus Rhyparus in the Philippines, with descriptions of two new species from Mindanao (Coleoptera: Scarabaeidae: Aphodiinae)" masterLastPageNumber="111" masterPageNumber="99" pageNumber="108" updateTime="1698977438183" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-3.0">
<mods:mods id="A545BF89BB501047F106579AD956F1CE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="22EEA713208E554FEC483E16D76E7D1F">
<mods:title id="65383F0EAA3187476488A62049398F94">A review of the genus Rhyparus in the Philippines, with descriptions of two new species from Mindanao (Coleoptera: Scarabaeidae: Aphodiinae)</mods:title>
</mods:titleInfo>
<mods:name id="9FBCF131242F05066498AC822C12DE3D" type="personal">
<mods:role id="52D7FB8C297AAD2FB1EB817F6064A154">
<mods:roleTerm id="9AA065FD28CC30EC2AEB145860C40A3A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C9D79B744AA6BCC845F5C50C7DC22D30">Anichtchenko, Alexander</mods:namePart>
<mods:affiliation id="69207A8412CACC0FC4076613845EE659">Institute of Life Sciences and Technologies, Daugavpils University, Daugavpils, Latvia</mods:affiliation>
<mods:nameIdentifier id="B1FADF13C8E1733A5D28A9DBBA69FDA5" type="email">beetl2000@mail.ru</mods:nameIdentifier>
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<mods:roleTerm id="092EBE0C44ABBEA9E424BC85B93E0208">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="95F8D22316FC278571BC6C9AD04B57FD">Minkina, Łukasz</mods:namePart>
<mods:affiliation id="7C729AB5A58167742FAF4F9AAB697243">os. Polana Szalfarska</mods:affiliation>
</mods:name>
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<mods:namePart id="295FD5AD6C2D72EFF4B55D4AB646E926">Vasiljeva, Anastasija</mods:namePart>
<mods:affiliation id="40049A3B4851DEA2684D0C5AA8ACEF66">Institute of Life Sciences and Technologies, Daugavpils University, Daugavpils, Latvia</mods:affiliation>
</mods:name>
<mods:name id="FAB103AD9F4E75F262AA596E113E7E61" type="personal">
<mods:role id="D5952C14DC5A3C33B6FEEC2478BB93C1">
<mods:roleTerm id="9FDC8192E95646392B84DB5F818753CE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9F7A33B3D96B5B761239BBB36D768E7A">Medina, Milton Norman D.</mods:namePart>
<mods:affiliation id="77DD223B5ED028BFBDCD89E877D6F8D4">Institute of Biodiversity and Environment, Coleoptera Research Center, University of Mindanao, Davao City, Philippines</mods:affiliation>
</mods:name>
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<mods:title id="25360E6F314AAB7B6263AA0B536860CF">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="7994B13A0D15F643BCBC352A849998E5">
<mods:date id="D6DCA2B20A475FDAB2BC6ABF7B933098">2021</mods:date>
<mods:detail id="06C01C79564E15BB04C448B60141A6C3" type="pubDate">
<mods:number id="C909C668E40993B4929F551F24EC7FFE">2021-03-16</mods:number>
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<mods:number id="A2596320F80D96EF799230155BA06180">61</mods:number>
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<mods:number id="6C6032CD9A3262C7CBEB98ED7E459805">1</mods:number>
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<mods:url id="B9171400103AE2875D0108736284E51F">http://dx.doi.org/10.37520/aemnp.2021.006</mods:url>
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<mods:identifier id="47BEB59F7F39E483238200A5C6D69A22" type="ISSN">1804-6487</mods:identifier>
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<treatment id="03AA81215331FFD1508C6A30B86DF834" ID-DOI="http://doi.org/10.5281/zenodo.5037528" ID-Zenodo-Dep="5037528" LSID="urn:lsid:plazi:treatment:03AA81215331FFD1508C6A30B86DF834" httpUri="http://treatment.plazi.org/id/03AA81215331FFD1508C6A30B86DF834" lastPageId="10" lastPageNumber="109" pageId="9" pageNumber="108">
<subSubSection id="C31963BC5331FFD2508C6A30B880F890" box="[248,662,1812,1836]" pageId="9" pageNumber="108" type="nomenclature">
<paragraph id="8BBC30375331FFD2508C6A30B880F890" blockId="9.[248,662,1812,1836]" box="[248,662,1812,1836]" pageId="9" pageNumber="108">
<heading id="D0F4875B5331FFD2508C6A30B880F890" box="[248,662,1812,1836]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="108" reason="3">
<taxonomicName id="4C034BB45331FFD2508C6A30B880F890" authority="Arrow, 1905" authorityName="Arrow" authorityYear="1905" box="[248,662,1812,1836]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="108" phylum="Arthropoda" rank="species" species="philippinensis">
<emphasis id="B977EC255331FFD2508C6A30B880F890" bold="true" box="[248,662,1812,1836]" pageId="9" pageNumber="108">
<emphasis id="B977EC255331FFD2508C6A30B812F890" bold="true" box="[248,516,1812,1836]" italics="true" pageId="9" pageNumber="108">Rhyparus philippinensis</emphasis>
<bibRefCitation id="EF924DC65331FFD2527E6A31B880F890" author="ARROW G. J." box="[522,662,1813,1836]" pageId="9" pageNumber="108" pagination="534 - 540" refId="ref6410" refString="ARROW G. J. 1905: On some Oriental aphodiid Coleoptera of the Rhyparus Group, with description of a new genus. Annals and Magazine of Natural History, Series 7 15: 534 - 540." type="journal article" year="1905">Arrow, 1905</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31963BC5331FFD2515E6A0BB872F8FA" box="[298,612,1839,1862]" pageId="9" pageNumber="108" type="description">
<paragraph id="8BBC30375331FFD2515E6A0BB872F8FA" blockId="9.[151,760,1839,2064]" box="[298,612,1839,1862]" pageId="9" pageNumber="108">
(
<figureCitation id="13382CB25331FFD251456A0BBB8AF8FA" box="[305,412,1839,1862]" captionStart="Fig" captionStartId="7.[150,181,1259,1278]" captionTargetBox="[172,1389,192,1237]" captionTargetId="figure-366@7.[150,1435,163,1251]" captionTargetPageId="7" captionText="Fig. 6. Habitus of Rhyparus philippinensis Arrow, 1905, syntypes: a – dorsal view; b – right dorsolateral view; c – right lateral view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5037558" httpUri="https://zenodo.org/record/5037558/files/figure.png" pageId="9" pageNumber="108">Figs 6a–c</figureCitation>
,
<figureCitation id="13382CB25331FFD251D36A0BBBD2F8FA" box="[423,452,1839,1862]" captionStart="Figs 7–12" captionStartId="8.[151,188,1611,1630]" captionTargetBox="[165,1410,204,1585]" captionTargetId="figure-191@8.[151,1436,189,1595]" captionTargetPageId="8" captionText="Figs 7–12. Head of Rhyparus species, frontal view: 7 – R. barclayi Masumoto, Ochi &amp; Ho, 2019; 8 – R. flavohirtus sp. nov.; 9 – R. gracilis Arrow, 1905; 10 – R. helophoroides Fairmaire, 1893; 11 – R. mindanaoensis sp. nov.; 12 – R. philippinensis Arrow, 1905. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5037560" httpUri="https://zenodo.org/record/5037560/files/figure.png" pageId="9" pageNumber="108">12</figureCitation>
,
<figureCitation id="13382CB25331FFD251BB6A0BBBFAF8FA" box="[463,492,1839,1862]" captionStart="Figs 13–18" captionStartId="9.[150,187,1526,1545]" captionTargetBox="[158,1411,197,1485]" captionTargetId="figure-259@9.[150,1435,161,1515]" captionTargetPageId="9" captionText="Figs 13–18. Epipharynx of Rhyparus species: 13 – R. barclayi Masumoto, Ochi &amp; Ho, 2019; 14 – R. flavohirtus sp. nov.; 15 – R. gracilis Arrow, 1905; 16 – R. helophoroides Fairmaire, 1893; 17– R. mindanaoensis sp. nov.; 18 – R. philippinensis Arrow, 1905. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5037562" httpUri="https://zenodo.org/record/5037562/files/figure.png" pageId="9" pageNumber="108">18</figureCitation>
,
<figureCitation id="13382CB25331FFD251836A0BB802F8FA" box="[503,532,1839,1862]" captionStart="Figs 19–24" captionStartId="10.[151,188,1324,1343]" captionTargetBox="[164,1415,209,1277]" captionTargetId="figure-243@10.[151,1436,189,1299]" captionTargetPageId="10" captionText="Figs 19–24. Elytra of Rhyparus species, oblique apico-dorsolateral view: 19 – R. barclayi Masumoto, Ochi &amp; Ho, 2019; 20 – R. flavohirtus sp. nov.; 21 – R. gracilis Arrow, 1905; 22 – R. helophoroides Fairmaire, 1893; 23 – R. mindanaoensis sp. nov.; 24 – R. philippinensis Arrow, 1905. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5037564" httpUri="https://zenodo.org/record/5037564/files/figure.png" pageId="9" pageNumber="108">24</figureCitation>
,
<figureCitation id="13382CB25331FFD2526B6A0BB82CF8FA" box="[543,570,1839,1862]" captionStart="Figs 25–30" captionStartId="11.[150,187,1486,1505]" captionTargetBox="[207,1392,193,1454]" captionTargetId="figure-302@11.[151,1436,180,1472]" captionTargetPageId="11" captionText="Figs 25–30.Aedeagus of Rhyparus species, a – lateral view, b – dorsal view. 25 – R.barclayi Masumoto, Ochi &amp; Ho, 2019; 26 – R. flavohirtus sp. nov.; 27 – R. gracilis Arrow, 1905; 28 – R. helophoroides Fairmaire, 1893; 29 – R.mindanaoensis sp. nov.; 30 – R.philippinensis Arrow, 1905. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5037566" httpUri="https://zenodo.org/record/5037566/files/figure.png" pageId="9" pageNumber="108">30</figureCitation>
a-b)
</paragraph>
</subSubSection>
<subSubSection id="C31963BC5331FFD250E36A79B9F8F891" pageId="9" pageNumber="108" type="materials_examined">
<paragraph id="8BBC30375331FFD250E36A79BB4EF7AC" blockId="9.[151,760,1839,2064]" pageId="9" pageNumber="108">
<emphasis id="B977EC255331FFD250E36A79BB36F8CC" bold="true" box="[151,288,1885,1904]" pageId="9" pageNumber="108">
<typeStatus id="54B88E955331FFD250E36A79BAD2F8CC" box="[151,196,1885,1904]" pageId="9" pageNumber="108">Type</typeStatus>
material.
</emphasis>
Sඒඇඍඒඉൾඌ: 1 J, labelled: ʻPhil. Isl.ʼ [round label] // ʻSyntype ʼ [round label] // ʻNHMUK 014189207ʼ (
<collectionCode id="ED12A8F25331FFD2523E6A53B899F836" box="[586,655,1911,1930]" pageId="9" pageNumber="108">BMNH</collectionCode>
);
<specimenCount id="9D05FBBE5331FFD252E96A53B8ABF836" box="[669,701,1905,1930]" pageId="9" pageNumber="108" type="female">1 ♀</specimenCount>
, labelled: ʻPhil. Islʼ [round label] // ʻ45 49ʼ [round label] // ʻSyntypeʼ [round label] // ʻNHMUK 014189209ʼ (
<collectionCode id="ED12A8F25331FFD251D36A89BBFAF87C" box="[423,492,1965,1984]" pageId="9" pageNumber="108">BMNH</collectionCode>
);
<specimenCount id="9D05FBBE5331FFD2518C6A89B801F803" box="[504,535,1958,1984]" pageId="9" pageNumber="108" type="female">1 ♀</specimenCount>
, labelled: ʻPhil. Islʼ [round label] // ʻSyntypeʼ [round label] // ʻNHMUK 014189208ʼ (
<collectionCode id="ED12A8F25331FFD252F66AE3B8D1F866" box="[642,711,1991,2010]" pageId="9" pageNumber="108">BMNH</collectionCode>
);
<specimenCount id="9D05FBBE5331FFD252A06AE3B8E5F866" box="[724,755,1985,2010]" pageId="9" pageNumber="108" type="female">1 ♀</specimenCount>
, labelled: ʻPhil. Islʼ [round label] // ʻSyntypeʼ [round label] // ʻNHMUK 013906231ʼ (
<collectionCode id="ED12A8F25331FFD2517E6AD9BB59F7AC" box="[266,335,2045,2064]" pageId="9" pageNumber="108">BMNH</collectionCode>
).
</paragraph>
<paragraph id="8BBC30375331FFD253486B5EB9F8F891" blockId="9.[828,1437,1652,1837]" pageId="9" pageNumber="108">
<emphasis id="B977EC255331FFD253486B5EBF7CF931" bold="true" box="[828,1386,1658,1677]" pageId="9" pageNumber="108">
Additional material examined.
<collectingCountry id="F31470A75331FFD2540F6B5EBF07F931" box="[1147,1297,1658,1677]" name="Philippines" pageId="9" pageNumber="108">PHILIPPINES</collectingCountry>
: Lඎඓඈඇ:
</emphasis>
<specimenCount id="9D05FBBE5331FFD255066B5EBF81F931" box="[1394,1431,1652,1677]" pageId="9" pageNumber="108" type="female">1 ♀</specimenCount>
, Lagonoy,
<collectingRegion id="49C7FED55331FFD253E16BB1BE0FF914" box="[917,1049,1685,1704]" country="Philippines" name="Camarines Sur" pageId="9" pageNumber="108">Camarines Sur</collectingRegion>
, Bicol,
<date id="FFBD16F75331FFD254156BB1BEBDF914" box="[1121,1195,1685,1704]" pageId="9" pageNumber="108" value="2018-07">vii.2018</date>
, local coll. (
<collectionCode id="ED12A8F25331FFD255696BB1BF75F914" box="[1309,1379,1685,1704]" pageId="9" pageNumber="108">CNCW</collectionCode>
);
<specimenCount id="9D05FBBE5331FFD255006BB1BF81F91B" box="[1396,1431,1678,1704]" pageId="9" pageNumber="108" type="female">1 ♀</specimenCount>
,
<collectingRegion id="49C7FED55331FFD253486B94B99CF97F" box="[828,906,1712,1731]" country="Philippines" name="Cagayan" pageId="9" pageNumber="108">Cagayan</collectingRegion>
, Santa Ana,
<date id="FFBD16F75331FFD253896B94BE5BF97F" box="[1021,1101,1712,1731]" pageId="9" pageNumber="108" value="2018-08">viii.2018</date>
, local coll. (
<collectionCode id="ED12A8F25331FFD254CA6B94BEE9F97F" box="[1214,1279,1712,1731]" pageId="9" pageNumber="108">DUBC</collectionCode>
); 1 J, Barlig Mt. Province,
<date id="FFBD16F75331FFD253E06BEEB9C8F961" box="[916,990,1738,1757]" pageId="9" pageNumber="108" value="2018-12">xii.2018</date>
, local coll. (
<collectionCode id="ED12A8F25331FFD2543D6BEEBE9BF961" box="[1097,1165,1738,1757]" pageId="9" pageNumber="108">LMCN</collectionCode>
).
<emphasis id="B977EC255331FFD254EE6BEEBF11F961" bold="true" box="[1178,1287,1738,1757]" pageId="9" pageNumber="108">Mංඇൽൺඇൺඈ:</emphasis>
<specimenCount id="9D05FBBE5331FFD2557F6BEEBF3DF961" box="[1291,1323,1732,1757]" pageId="9" pageNumber="108" type="female">1 ♀</specimenCount>
, New
<collectingRegion id="49C7FED55331FFD255176BEEBF8FF961" box="[1379,1433,1738,1757]" country="Philippines" name="Albay" pageId="9" pageNumber="108">Albay</collectingRegion>
,
<collectingRegion id="49C7FED55331FFD253486BC1B9F7F944" box="[828,993,1765,1784]" country="Philippines" name="Compostela" pageId="9" pageNumber="108">Compostela Valley</collectingRegion>
,
<date id="FFBD16F75331FFD2539C6BC1BE24F944" box="[1000,1074,1765,1784]" pageId="9" pageNumber="108" value="2018-07">vii.2018</date>
, local coll. (
<collectionCode id="ED12A8F25331FFD254EE6BC1BEF6F944" box="[1178,1248,1765,1784]" pageId="9" pageNumber="108">CNCW</collectionCode>
); 1 J, Wao,
<collectingRegion id="49C7FED55331FFD255336BC1B94DF8AF" country="Philippines" name="Lanao del Sur" pageId="9" pageNumber="108">Lanao del Sur</collectingRegion>
,
<date id="FFBD16F75331FFD253176A24B9B6F8AF" box="[867,928,1792,1811]" pageId="9" pageNumber="108" value="2018-10">x.2018</date>
, local coll. (
<collectionCode id="ED12A8F25331FFD254736A24BE5BF8AF" box="[1031,1101,1792,1811]" pageId="9" pageNumber="108">CNCW</collectionCode>
);
<specimenCount id="9D05FBBE5331FFD2542F6A24BE6CF8AE" box="[1115,1146,1785,1811]" pageId="9" pageNumber="108" type="female">1 ♀</specimenCount>
, Alamada,
<collectingRegion id="49C7FED55331FFD254AF6A24BF45F8AF" box="[1243,1363,1792,1811]" country="Philippines" name="Lanao del Sur" pageId="9" pageNumber="108">Lanao del Sur</collectingRegion>
,
<date id="FFBD16F75331FFD2552F6A24BF8EF8AF" box="[1371,1432,1792,1811]" pageId="9" pageNumber="108" value="2018-10">x.2018</date>
, local coll. (
<collectionCode id="ED12A8F25331FFD253EA6A3EB9F2F891" box="[926,996,1818,1837]" pageId="9" pageNumber="108">CNCW</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C31963BC5331FFD153486A62BBDCF8D4" lastPageId="10" lastPageNumber="109" pageId="9" pageNumber="108" type="diagnosis">
<paragraph id="8BBC30375331FFD153486A62B855F974" blockId="9.[828,1437,1862,2077]" lastBlockId="10.[151,760,1585,1928]" lastPageId="10" lastPageNumber="109" pageId="9" pageNumber="108">
<emphasis id="B977EC255331FFD253486A62B9A7F8E1" bold="true" box="[828,945,1862,1885]" pageId="9" pageNumber="108">Diagnosis.</emphasis>
Medium-sized species: body length 5.0–
<quantity id="4CFB9DD25331FFD255086A62B970F8C1" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="9" pageNumber="108" unit="mm" value="6.5">6.5 mm</quantity>
(
<figureCitation id="13382CB25331FFD253076A42B9F6F8C1" box="[883,992,1894,1917]" captionStart="Fig" captionStartId="7.[150,181,1259,1278]" captionTargetBox="[172,1389,192,1237]" captionTargetId="figure-366@7.[150,1435,163,1251]" captionTargetPageId="7" captionText="Fig. 6. Habitus of Rhyparus philippinensis Arrow, 1905, syntypes: a – dorsal view; b – right dorsolateral view; c – right lateral view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5037558" httpUri="https://zenodo.org/record/5037558/files/figure.png" pageId="9" pageNumber="108">Figs 6a–c</figureCitation>
). Head as depicted in
<figureCitation id="13382CB25331FFD254BE6A42BF0EF8C1" box="[1226,1304,1894,1917]" captionStart="Figs 7–12" captionStartId="8.[151,188,1611,1630]" captionTargetBox="[165,1410,204,1585]" captionTargetId="figure-191@8.[151,1436,189,1595]" captionTargetPageId="8" captionText="Figs 7–12. Head of Rhyparus species, frontal view: 7 – R. barclayi Masumoto, Ochi &amp; Ho, 2019; 8 – R. flavohirtus sp. nov.; 9 – R. gracilis Arrow, 1905; 10 – R. helophoroides Fairmaire, 1893; 11 – R. mindanaoensis sp. nov.; 12 – R. philippinensis Arrow, 1905. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5037560" httpUri="https://zenodo.org/record/5037560/files/figure.png" pageId="9" pageNumber="108">Fig. 12</figureCitation>
; epipharynx as in
<figureCitation id="13382CB25331FFD253026AA2B9D3F821" box="[886,965,1926,1949]" captionStart="Figs 13–18" captionStartId="9.[150,187,1526,1545]" captionTargetBox="[158,1411,197,1485]" captionTargetId="figure-259@9.[150,1435,161,1515]" captionTargetPageId="9" captionText="Figs 13–18. Epipharynx of Rhyparus species: 13 – R. barclayi Masumoto, Ochi &amp; Ho, 2019; 14 – R. flavohirtus sp. nov.; 15 – R. gracilis Arrow, 1905; 16 – R. helophoroides Fairmaire, 1893; 17– R. mindanaoensis sp. nov.; 18 – R. philippinensis Arrow, 1905. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5037562" httpUri="https://zenodo.org/record/5037562/files/figure.png" pageId="9" pageNumber="108">Fig. 18</figureCitation>
. Intermediate lobes of pronotum as large as anterior one; median intercosta with dense punctation; second and third intercostae with dense punctation basally. All intercostae of elytra with two rows of medium-sized punctures. Internal and median protrusion of caudal bulbs connected (
<figureCitation id="13382CB25332FFD1517A6B15BB4DF9F4" box="[270,347,1585,1608]" captionStart="Figs 19–24" captionStartId="10.[151,188,1324,1343]" captionTargetBox="[164,1415,209,1277]" captionTargetId="figure-243@10.[151,1436,189,1299]" captionTargetPageId="10" captionText="Figs 19–24. Elytra of Rhyparus species, oblique apico-dorsolateral view: 19 – R. barclayi Masumoto, Ochi &amp; Ho, 2019; 20 – R. flavohirtus sp. nov.; 21 – R. gracilis Arrow, 1905; 22 – R. helophoroides Fairmaire, 1893; 23 – R. mindanaoensis sp. nov.; 24 – R. philippinensis Arrow, 1905. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5037564" httpUri="https://zenodo.org/record/5037564/files/figure.png" pageId="10" pageNumber="109">Fig. 24</figureCitation>
); surface between external protrusion of caudal bulb of elytra and lateral margin of elytra distinctly sinuate – external protrusion more or less developed; however, median and internal ones always with the same proportions. Aedeagus as in
<figureCitation id="13382CB25332FFD151B26B95B829F974" box="[454,575,1713,1736]" captionStart="Figs 25–30" captionStartId="11.[150,187,1486,1505]" captionTargetBox="[207,1392,193,1454]" captionTargetId="figure-302@11.[151,1436,180,1472]" captionTargetPageId="11" captionText="Figs 25–30.Aedeagus of Rhyparus species, a – lateral view, b – dorsal view. 25 – R.barclayi Masumoto, Ochi &amp; Ho, 2019; 26 – R. flavohirtus sp. nov.; 27 – R. gracilis Arrow, 1905; 28 – R. helophoroides Fairmaire, 1893; 29 – R.mindanaoensis sp. nov.; 30 – R.philippinensis Arrow, 1905. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5037566" httpUri="https://zenodo.org/record/5037566/files/figure.png" pageId="10" pageNumber="109">Figs 30a–b</figureCitation>
.
</paragraph>
<caption id="DF7C60BF5332FFD150E36808BF88FAE5" ID-DOI="http://doi.org/10.5281/zenodo.5037564" ID-Zenodo-Dep="5037564" httpUri="https://zenodo.org/record/5037564/files/figure.png" pageId="10" pageNumber="109" startId="10.[151,188,1324,1343]" targetBox="[164,1415,209,1277]" targetPageId="10">
<paragraph id="8BBC30375332FFD150E36808BF88FAE5" blockId="10.[151,1438,1324,1369]" pageId="10" pageNumber="109">
Figs 19–24. Elytra of
<taxonomicName id="4C034BB45332FFD1513A6808BBF2FA83" box="[334,484,1324,1343]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="109" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B977EC255332FFD1513A6808BBB6FA83" box="[334,416,1324,1343]" italics="true" pageId="10" pageNumber="109">Rhyparus</emphasis>
species
</taxonomicName>
, oblique apico-dorsolateral view: 19 –
<taxonomicName id="4C034BB45332FFD1535F6808BE93FA83" authority="Masumoto, Ochi &amp; Ho, 2019" authorityName="Masumoto, Ochi &amp; Ho" authorityYear="2019" box="[811,1157,1324,1343]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="109" phylum="Arthropoda" rank="species" species="barclayi">
<emphasis id="B977EC255332FFD1535F6808B99EFA83" box="[811,904,1324,1343]" italics="true" pageId="10" pageNumber="109">R. barclayi</emphasis>
<bibRefCitation id="EF924DC65332FFD153F96808BE93FA83" author="MASUMOTO K. &amp; OCHI T. &amp; HO B. H." box="[909,1157,1324,1343]" pageId="10" pageNumber="109" pagination="13 - 18" refId="ref6667" refString="MASUMOTO K., OCHI T. &amp; HO B. H. 2019: A new Rhyparus species from Taiwan (Scarabaeidae, Aphodiinae, Rhyparini). Kogane 22: 13 - 18." type="journal article" year="2019">Masumoto, Ochi &amp; Ho, 2019</bibRefCitation>
</taxonomicName>
; 20 –
<taxonomicName id="4C034BB45332FFD154C26808BF3FFA83" authority="Anichtchenko &amp; Minkina &amp; Vasiljeva &amp; Medina, 2021" authorityName="Anichtchenko &amp; Minkina &amp; Vasiljeva &amp; Medina" authorityYear="2021" box="[1206,1321,1324,1343]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="109" phylum="Arthropoda" rank="species" species="flavohirtus" status="sp. nov.">
<emphasis id="B977EC255332FFD154C26808BF3FFA83" box="[1206,1321,1324,1343]" italics="true" pageId="10" pageNumber="109">R. flavohirtus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A244515E5332FFD155596808BF67FA83" box="[1325,1393,1324,1343]" pageId="10" pageNumber="109" rank="species">sp. nov.</taxonomicNameLabel>
; 21 –
<taxonomicName id="4C034BB45332FFD150E36862BB76FAE5" authority="Arrow, 1905" authorityName="Arrow" authorityYear="1905" box="[151,352,1350,1369]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="109" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B977EC255332FFD150E36862BAF9FAE5" box="[151,239,1350,1369]" italics="true" pageId="10" pageNumber="109">R. gracilis</emphasis>
<bibRefCitation id="EF924DC65332FFD150876862BB76FAE5" author="ARROW G. J." box="[243,352,1350,1369]" pageId="10" pageNumber="109" pagination="534 - 540" refId="ref6410" refString="ARROW G. J. 1905: On some Oriental aphodiid Coleoptera of the Rhyparus Group, with description of a new genus. Annals and Magazine of Natural History, Series 7 15: 534 - 540." type="journal article" year="1905">Arrow, 1905</bibRefCitation>
</taxonomicName>
; 22 –
<taxonomicName id="4C034BB45332FFD151E56862B8B9FAE5" authority="Fairmaire, 1893" authorityName="Fairmaire" authorityYear="1893" box="[401,687,1350,1369]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="109" phylum="Arthropoda" rank="species" species="helophoroides">
<emphasis id="B977EC255332FFD151E56862B836FAE5" box="[401,544,1350,1369]" italics="true" pageId="10" pageNumber="109">R. helophoroides</emphasis>
Fairmaire, 1893
</taxonomicName>
; 23 –
<taxonomicName id="4C034BB45332FFD152AB6862B960FAE5" authority="Anichtchenko &amp; Minkina &amp; Vasiljeva &amp; Medina, 2021" authorityName="Anichtchenko &amp; Minkina &amp; Vasiljeva &amp; Medina" authorityYear="2021" box="[735,886,1350,1369]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="109" phylum="Arthropoda" rank="species" species="mindanaoensis" status="sp. nov.">
<emphasis id="B977EC255332FFD152AB6862B960FAE5" box="[735,886,1350,1369]" italics="true" pageId="10" pageNumber="109">R. mindanaoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A244515E5332FFD1530E6862B9A8FAE5" box="[890,958,1350,1369]" pageId="10" pageNumber="109" rank="species">sp. nov.</taxonomicNameLabel>
; 24 –
<taxonomicName id="4C034BB45332FFD153996862BEFBFAE5" authority="Arrow, 1905" authorityName="Arrow" authorityYear="1905" box="[1005,1261,1350,1369]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="109" phylum="Arthropoda" rank="species" species="philippinensis">
<emphasis id="B977EC255332FFD153996862BE6AFAE5" box="[1005,1148,1350,1369]" italics="true" pageId="10" pageNumber="109">R. philippinensis</emphasis>
<bibRefCitation id="EF924DC65332FFD154F46862BEFBFAE5" author="ARROW G. J." box="[1152,1261,1350,1369]" pageId="10" pageNumber="109" pagination="534 - 540" refId="ref6410" refString="ARROW G. J. 1905: On some Oriental aphodiid Coleoptera of the Rhyparus Group, with description of a new genus. Annals and Magazine of Natural History, Series 7 15: 534 - 540." type="journal article" year="1905">Arrow, 1905</bibRefCitation>
</taxonomicName>
. Scale bar = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BBC30375332FFD150E36BF5BBDCF8D4" blockId="10.[151,760,1585,1928]" pageId="10" pageNumber="109">
<emphasis id="B977EC255332FFD150E36BF5BB00F954" bold="true" box="[151,278,1745,1768]" pageId="10" pageNumber="109">Variability.</emphasis>
Lateral lobes of pronotum slightly variable in size and proportions. Punctures of intercostae of pronotum and elytra can be proportionally variable in size between the specimens. Costae of pronotum and elytra are slightly variable in height and width.
</paragraph>
</subSubSection>
<subSubSection id="C31963BC5332FFD150E36A55B86DF834" box="[151,635,1905,1928]" pageId="10" pageNumber="109" type="distribution">
<paragraph id="8BBC30375332FFD150E36A55B86DF834" blockId="10.[151,760,1585,1928]" box="[151,635,1905,1928]" pageId="10" pageNumber="109">
<emphasis id="B977EC255332FFD150E36A55BB3CF834" bold="true" box="[151,298,1905,1928]" pageId="10" pageNumber="109">Distribution.</emphasis>
<collectingCountry id="F31470A75332FFD151446A55BBBAF834" box="[304,428,1905,1928]" name="Philippines" pageId="10" pageNumber="109">Philippines</collectingCountry>
: Luzon, Mindanao.
</paragraph>
</subSubSection>
</treatment>
</document>
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